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Ischigualasto Formation. the Second Is a Sile- Diversity Or Abundance, but This Result Was Based on Only 19 of Saurid, Ignotosaurus Fragilis (Fig
This article was downloaded by: [University of Chicago Library] On: 10 October 2013, At: 10:52 Publisher: Taylor & Francis Informa Ltd Registered in England and Wales Registered Number: 1072954 Registered office: Mortimer House, 37-41 Mortimer Street, London W1T 3JH, UK Journal of Vertebrate Paleontology Publication details, including instructions for authors and subscription information: http://www.tandfonline.com/loi/ujvp20 Vertebrate succession in the Ischigualasto Formation Ricardo N. Martínez a , Cecilia Apaldetti a b , Oscar A. Alcober a , Carina E. Colombi a b , Paul C. Sereno c , Eliana Fernandez a b , Paula Santi Malnis a b , Gustavo A. Correa a b & Diego Abelin a a Instituto y Museo de Ciencias Naturales, Universidad Nacional de San Juan , España 400 (norte), San Juan , Argentina , CP5400 b Consejo Nacional de Investigaciones Científicas y Técnicas , Buenos Aires , Argentina c Department of Organismal Biology and Anatomy, and Committee on Evolutionary Biology , University of Chicago , 1027 East 57th Street, Chicago , Illinois , 60637 , U.S.A. Published online: 08 Oct 2013. To cite this article: Ricardo N. Martínez , Cecilia Apaldetti , Oscar A. Alcober , Carina E. Colombi , Paul C. Sereno , Eliana Fernandez , Paula Santi Malnis , Gustavo A. Correa & Diego Abelin (2012) Vertebrate succession in the Ischigualasto Formation, Journal of Vertebrate Paleontology, 32:sup1, 10-30, DOI: 10.1080/02724634.2013.818546 To link to this article: http://dx.doi.org/10.1080/02724634.2013.818546 PLEASE SCROLL DOWN FOR ARTICLE Taylor & Francis makes every effort to ensure the accuracy of all the information (the “Content”) contained in the publications on our platform. However, Taylor & Francis, our agents, and our licensors make no representations or warranties whatsoever as to the accuracy, completeness, or suitability for any purpose of the Content. -
Studies on Continental Late Triassic Tetrapod Biochronology. I. the Type Locality of Saturnalia Tupiniquim and the Faunal Succession in South Brazil
Journal of South American Earth Sciences 19 (2005) 205–218 www.elsevier.com/locate/jsames Studies on continental Late Triassic tetrapod biochronology. I. The type locality of Saturnalia tupiniquim and the faunal succession in south Brazil Max Cardoso Langer* Departamento de Biologia, FFCLRP, Universidade de Sa˜o Paulo (USP), Av. Bandeirantes 3900, 14040-901 Ribeira˜o Preto, SP, Brazil Received 1 November 2003; accepted 1 January 2005 Abstract Late Triassic deposits of the Parana´ Basin, Rio Grande do Sul, Brazil, encompass a single third-order, tetrapod-bearing sedimentary sequence that includes parts of the Alemoa Member (Santa Maria Formation) and the Caturrita Formation. A rich, diverse succession of terrestrial tetrapod communities is recorded in these sediments, which can be divided into at least three faunal associations. The stem- sauropodomorph Saturnalia tupiniquim was collected in the locality known as ‘Waldsanga’ near the city of Santa Maria. In that area, the deposits of the Alemoa Member yield the ‘Alemoa local fauna,’ which typifies the first association; includes the rhynchosaur Hyperodapedon, aetosaurs, and basal dinosaurs; and is coeval with the lower fauna of the Ischigualasto Formation, Bermejo Basin, NW Argentina. The second association is recorded in deposits of both the Alemoa Member and the Caturrita Formation, characterized by the rhynchosaur ‘Scaphonyx’ sulcognathus and the cynodont Exaeretodon, and correlated with the upper fauna of the Ischigualasto Formation. Various isolated outcrops of the Caturrita Formation yield tetrapod fossils that correspond to post-Ischigualastian faunas but might not belong to a single faunal association. The record of the dicynodont Jachaleria suggests correlations with the lower part of the Los Colorados Formation, NW Argentina, whereas remains of derived tritheledontid cynodonts indicate younger ages. -
The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
The Sauropodomorph Biostratigraphy of the Elliot Formation of Southern Africa: Tracking the Evolution of Sauropodomorpha Across the Triassic–Jurassic Boundary
Editors' choice The sauropodomorph biostratigraphy of the Elliot Formation of southern Africa: Tracking the evolution of Sauropodomorpha across the Triassic–Jurassic boundary BLAIR W. MCPHEE, EMESE M. BORDY, LARA SCISCIO, and JONAH N. CHOINIERE McPhee, B.W., Bordy, E.M., Sciscio, L., and Choiniere, J.N. 2017. The sauropodomorph biostratigraphy of the Elliot Formation of southern Africa: Tracking the evolution of Sauropodomorpha across the Triassic–Jurassic boundary. Acta Palaeontologica Polonica 62 (3): 441–465. The latest Triassic is notable for coinciding with the dramatic decline of many previously dominant groups, followed by the rapid radiation of Dinosauria in the Early Jurassic. Among the most common terrestrial vertebrates from this time, sauropodomorph dinosaurs provide an important insight into the changing dynamics of the biota across the Triassic–Jurassic boundary. The Elliot Formation of South Africa and Lesotho preserves the richest assemblage of sauropodomorphs known from this age, and is a key index assemblage for biostratigraphic correlations with other simi- larly-aged global terrestrial deposits. Past assessments of Elliot Formation biostratigraphy were hampered by an overly simplistic biozonation scheme which divided it into a lower “Euskelosaurus” Range Zone and an upper Massospondylus Range Zone. Here we revise the zonation of the Elliot Formation by: (i) synthesizing the last three decades’ worth of fossil discoveries, taxonomic revision, and lithostratigraphic investigation; and (ii) systematically reappraising the strati- graphic provenance of important fossil locations. We then use our revised stratigraphic information in conjunction with phylogenetic character data to assess morphological disparity between Late Triassic and Early Jurassic sauropodomorph taxa. Our results demonstrate that the Early Jurassic upper Elliot Formation is considerably more taxonomically and morphologically diverse than previously thought. -
Gondwana Vertebrate Faunas of India: Their Diversity and Intercontinental Relationships
438 Article 438 by Saswati Bandyopadhyay1* and Sanghamitra Ray2 Gondwana Vertebrate Faunas of India: Their Diversity and Intercontinental Relationships 1Geological Studies Unit, Indian Statistical Institute, 203 B. T. Road, Kolkata 700108, India; email: [email protected] 2Department of Geology and Geophysics, Indian Institute of Technology, Kharagpur 721302, India; email: [email protected] *Corresponding author (Received : 23/12/2018; Revised accepted : 11/09/2019) https://doi.org/10.18814/epiiugs/2020/020028 The twelve Gondwanan stratigraphic horizons of many extant lineages, producing highly diverse terrestrial vertebrates India have yielded varied vertebrate fossils. The oldest in the vacant niches created throughout the world due to the end- Permian extinction event. Diapsids diversified rapidly by the Middle fossil record is the Endothiodon-dominated multitaxic Triassic in to many communities of continental tetrapods, whereas Kundaram fauna, which correlates the Kundaram the non-mammalian synapsids became a minor components for the Formation with several other coeval Late Permian remainder of the Mesozoic Era. The Gondwana basins of peninsular horizons of South Africa, Zambia, Tanzania, India (Fig. 1A) aptly exemplify the diverse vertebrate faunas found Mozambique, Malawi, Madagascar and Brazil. The from the Late Palaeozoic and Mesozoic. During the last few decades much emphasis was given on explorations and excavations of Permian-Triassic transition in India is marked by vertebrate fossils in these basins which have yielded many new fossil distinct taxonomic shift and faunal characteristics and vertebrates, significant both in numbers and diversity of genera, and represented by small-sized holdover fauna of the providing information on their taphonomy, taxonomy, phylogeny, Early Triassic Panchet and Kamthi fauna. -
The Anatomy and Phylogenetic Relationships of Antetonitrus Ingenipes (Sauropodiformes, Dinosauria): Implications for the Origins of Sauropoda
THE ANATOMY AND PHYLOGENETIC RELATIONSHIPS OF ANTETONITRUS INGENIPES (SAUROPODIFORMES, DINOSAURIA): IMPLICATIONS FOR THE ORIGINS OF SAUROPODA Blair McPhee A dissertation submitted to the Faculty of Science, University of the Witwatersrand, in partial fulfilment of the requirements for the degree of Master of Science. Johannesburg, 2013 i ii ABSTRACT A thorough description and cladistic analysis of the Antetonitrus ingenipes type material sheds further light on the stepwise acquisition of sauropodan traits just prior to the Triassic/Jurassic boundary. Although the forelimb of Antetonitrus and other closely related sauropododomorph taxa retains the plesiomorphic morphology typical of a mobile grasping structure, the changes in the weight-bearing dynamics of both the musculature and the architecture of the hindlimb document the progressive shift towards a sauropodan form of graviportal locomotion. Nonetheless, the presence of hypertrophied muscle attachment sites in Antetonitrus suggests the retention of an intermediary form of facultative bipedality. The term Sauropodiformes is adopted here and given a novel definition intended to capture those transitional sauropodomorph taxa occupying a contiguous position on the pectinate line towards Sauropoda. The early record of sauropod diversification and evolution is re- examined in light of the paraphyletic consensus that has emerged regarding the ‘Prosauropoda’ in recent years. iii ACKNOWLEDGEMENTS First, I would like to express sincere gratitude to Adam Yates for providing me with the opportunity to do ‘real’ palaeontology, and also for gladly sharing his considerable knowledge on sauropodomorph osteology and phylogenetics. This project would not have been possible without the continued (and continual) support (both emotionally and financially) of my parents, Alf and Glenda McPhee – Thank you. -
The Triassic Insect Fauna from the Los Rastros Formation (Bermejo Basin), La Rioja Province (Argentina): Their Con- Text, Taphonomy and Paleobiology
AMEGHINIANA (Rev. Asoc. Paleontol. Argent.) - 44 (2): 000-000. Buenos Aires, 30-6-2007 ISSN 0002-7014 The Triassic insect fauna from the Los Rastros Formation (Bermejo Basin), La Rioja Province (Argentina): their con- text, taphonomy and paleobiology Adriana C. MANCUSO1, Oscar F. GALLEGO2 and Rafael G. MARTINS-NETO3 Abstract. In the Bermejo Basin, the Los Rastros Formation bears an abundant insect fauna, mainly with terrestrial adult winged organisms related to the Blattoptera and the Coleoptera orders. The insect re- mains are found in the black shales of the offshore lacustrine facies and the insect taphonomic features suggest that the specimens were allochthonous to the lake. The individuals appear to arrived alive to the lake and suffered a rapid fall through the water column thus, preserving them intact, and some of them suffered fragmentation in air transportation and by biological attack during long periods of flotation. Resumen. LA FAUNA DE INSECTOS TRIÁSICOS DE LA FORMACIÓN LOS RASTROS (CUENCA BERMEJO), PROVINCIA DE LA RIOJA (ARGENTINA): SU CONTEXTO, TAFONOMÍA Y PALEOBIOLOGÍA. En la Cuenca Bermejo, la Formación Los Rastros es portadora de una abundante fauna de insectos, principalmente organismos adultos terrestres y alados pertenecientes a los órdenes Blattoptera y Coleoptera. Los restos de insectos son encontrados en las pelitas negras de la facies de lago abierto. Las características tafonómicas de los insectos sugieren que los especímenes son alóctonos al lago. Los individuos pudieron llegar vivos al lago y sufrir una rápida caída a través de la columna de agua, preservándose intactos, o sufrieron fragmentación en el transporte aéreo y por ataques biológicos durante largos períodos de flotación. -
A Non-Mammaliaform Cynodont from the Upper Triassic of South Africa: a Therapsid Lazarus Taxon?
View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by Wits Institutional Repository on DSPACE A non-mammaliaform cynodont from the Upper Triassic of South Africa: a therapsid Lazarus taxon? Fernando Abdala1*, Ross Damiani2, Adam Yates1 & Johann Neveling3 1Bernard Price Institute for Palaeontological Research, School of Geosciences, University of the Witwatersrand, Private Bag 3, WITS, 2050 South Africa 2Staatliches Museum für Naturkunde Stuttgart, Rosenstein 1, D-70191, Stuttgart, Germany 3Council for Geoscience, Private Bag X112, Pretoria, 0001 South Africa Received 20 January 2006. Accepted 10 January 2007 The tetrapod record of the ‘Stormberg Group’, including the Lower Elliot Formation, in the South African Karoo is widely dominated by archosaurian reptiles, contrasting with the therapsid dominion of the subjacent Beaufort Group. The only therapsids represented by skeletal remains in the Upper Triassic Lower Elliot Formation are the large traversodontid cynodont Scalenodontoides macrodontes and the recently described tritheledontid cynodont Elliotherium kersteni. Here we present a fragmentary lower jaw that provides evidence of a third type of cynodont for the Upper Triassic of South Africa. The fossil is tentatively assigned to the Diademodontidae. The latter representative of this family is known from the Late Anisian, and its tentative record in the Norian Lower Elliot Formation, if confirmed, will represent a case of Lazarus taxon. Thus, Diademodontidae apparently disappeared from the fossil record by the end of the Anisian and then reappeared in the Norian of South Africa, a stratigraphic interval of some 21 million years. This new cynodont record, together with the recently described Tritheledontidae, show that cynodonts are now the second most diverse tetrapod group in the Lower Elliot fauna. -
From the Upper Triassic Ischigualasto Formation of Northwestern Argentina
A peer-reviewed open-access journal ZooKeys 63:A 55–81 new (2010)herrerasaurid (Dinosauria, Saurischia) from the Upper Triassic Ischigualasto... 55 doi: 10.3897/zookeys.63.550 RESEARCH ARTICLE www.pensoftonline.net/zookeys Launched to accelerate biodiversity research A new herrerasaurid (Dinosauria, Saurischia) from the Upper Triassic Ischigualasto Formation of northwestern Argentina Oscar A. Alcober†, Ricardo N. Martinez‡ Museo de Ciencias Naturales, San Juan 5400, Argentina † urn:lsid:zoobank.org:author:B78F72E4-F21D-40AF-AC9D-4F1866A49FE8 ‡ urn:lsid:zoobank.org:author:94DF5C23-A12C-4F01-B845-05634636FBC0 Corresponding author: Ricardo N. Martinez ( [email protected] ) Academic editor: Hans-Dieter Sues | Received 11 July 2010 | Accepted 23 September 2010 | Published 19 October 2010 urn:lsid:zoobank.org:pub:FB2AE660-C3EE-4348-BF9F-F4311C47E853 Citation: Alcober OA, Martinez RN (2010) A new herrerasaurid (Dinosauria, Saurischia) from the Upper Triassic Ischigualasto Formation of northwestern Argentina. ZooKeys 63 : 55 – 81 . doi: 10.3897/zookeys.63.550 Abstract Herrerasauridae comprises a basal clade of dinosaurs best known from the Upper Triassic of Argentina and Brazil, which have yielded remains of Herrerasaurus ischigualastensis and Staurikosaurus pricei, respectively. Systematic opinion regarding the position of Herrerasauridae at the base of Dinosauria has varied. Here we describe a new herrerasaurid, Sanjuansaurus gordilloi gen. n., sp. n., based on a partial skeleton from Carnian-age strata of the the Upper Triassic Ischigualasto Formation of northwestern Argentina. Th e new taxon is diagnosed by numerous features, including long, band-shaped and posterolaterally oriented transverse process on the posterior cervical vertebrae; neural spines of the sixth to eighth dorsal vertebrae, at least, bearing acute anterior and posterior processes; scapula and coracoid with everted lateral margins of the glenoid; and short pubis (63% of the femoral length). -
Miospores and Chlorococcalean Algae from the Los Rastros Formation, Middle to Upper Triassic of Central-Western Argentina
AMEGHINIANA (Rev. Asoc. Paleontol. Argent.) - 42 (2): 347-362. Buenos Aires, 30-06-2005 ISSN 0002-7014 Miospores and chlorococcalean algae from the Los Rastros Formation, Middle to Upper Triassic of central-western Argentina Eduardo G. OTTONE, Adriana C. MANCUSO and Magdalena RESANO Abstract. Lacustrine strata of the Los Rastros Formation (Middle to Upper Triassic) at Río Gualo section (La Rioja province), yield a distinctive palynological assemblage of miospores and chlorococcalean algae. The miospore association is characterized by a relative abundance of corystosperm pollen grains with sub- ordinate inaperturates, diploxylonoid disaccates, spores, monocolpates, monosaccates and striate pollen grains. The phytoplankton are mostly represented by Botryococcus but also by Plaesiodictyon, a form prob- ably related to the Hydrodictyaceae. Geological data and variations in phytoplankton content indicate that the lacustrine system probably evolved from a stretcht of freshwater with eutrophic conditions, into a body with oligotrophic conditions through the middle and upper part of the Río Gualo section. The genus Variapollenites is emended in order to amplify its original diagnosis. Resumen. MIOSPORAS Y ALGAS CHLOROCOCCALES DE LA FORMACIÓN LOS RASTROS, TRIÁSICO MEDIO A SUPERIOR DEL CENTRO-OESTE DE ARGENTINA. El estudio de los niveles lacustres de la Formación Los Rastros (Triásico Medio a Superior) en la sección de Río Gualo (provincia de La Rioja), incluye una interesante palinoflora compuesta por miosporas y algas Chlorococcales. Entre las miosporas abundan los granos de polen de Corystospermales, con presencia subordinada de inaperturados, disacados diploxilonoides, esporas, monocolpados, monosacados y polen estriado. En el fitoplancton se destaca Botryococcus, pero también se observa Plaesiodictyon, que es una forma probablemente relacionada con las Hydrodictyaceae. -
Quebrada De La Sal Magnetoestratigraphic Section, Los Colorados Formación, Upper Triassic Ischigualasto-Villa Unión Basin, Argentina
Latinmag Letters, Volume 1, Special Issue (2011), B15, 1-7. Proceedings Tandil, Argentina Quebrada de la Sal magnetoestratigraphic section, Los Colorados Formación, Upper Triassic Ischigualasto-Villa Unión basin, Argentina. Paula Santi Malnis1,2, Dennis V. Kent3,4, Carina E. Colombi1,2, Silvana E. GEUNA1,5 1 CONICET 2Instituto y Museo de Ciencias Naturales, UNSJ; 3 Lamont Doherty Earth Observatory, Columbia University, NY-US 4Earth and Planetary Sciences, Rutgers University, NJ-US 5INGEODAV, Dep. de Ciencias Geológicas, Facultad de Ciencias Exactas y Naturales, UBA Abstract: Ischigualasto-Villa Union basin is a continental rift developed in southwestern quarter of Pangea throughout the Triassic. Los Colorados Formation, its youngest stratigraphic unit, is formed by a continuously succession of red beds that host a paleovertebrate assemblage of biostratigraphic importance. In this work are presented the results of the first magnetostratigraphic study applied on Los Colorados Formation with the aim of global correlations. For this, a full section of 600 meters were sampled. The paleomagnetic results show two components, recognized as “A” and “C”. Component C has unblocking temperature from 650°C, presents inter-site consistency and was identify as the carrier of the primary remanent magnetization with an SSW-down (reverse polarity) or NNE-up (normal polarity) orientation in different sites. The main magnetic mineral carrier is hematite and only approaches reaches saturation from 2.0 T. The magnetostratigraphic column of Los Colorados presents 16 magnetozones of normal and reverse polarity. In conjunction with published volcanic ash dates, the preferred polarity correlation for Los Colorados Formation magnetostratigraphy is between magnetochrons E7r (227 Ma) and E14n (216- 215 Ma) from Newark Basin GPTS, or early to middle Norian. -
Supplementary Information
SUPPLEMENTARY INFORMATION The oldest known communal latrines provide evidence of gregarism in Triassic megaherbivores Lucas E. Fiorelli*, Martín D. Ezcurra, E. Martín Hechenleitner, Eloisa Argañaraz, Jeremías R. A. Taborda, M. Jimena Trotteyn, M. Belén von Baczko & Julia B. Desojo *To whom correspondence should be addressed. E-mail: [email protected] 1. Provenance, authenticity, geological setting and stratigraphy of the communal latrines of the Chañares Formation 2. Depositional setting 3. Taphonomy 4. Statistics 5. Age of the Chañares Formation 6. Fossil tetrapods from the Chañares Formation 7. Dinodontosaurus body size 8. Dinodontosaurus as a gregarious megaherbivore 9. References 1. Provenance, authenticity, geological setting and stratigraphy of the communal latrines of the Chañares Formation. Several communal latrines were found in successive palaeontological field works conducted in 2011 and 2012 in outcrops of the Chañares Formation situated in the Talampaya National Park, La Rioja Province, northwestern Argentina (Supplementary Figure 1a). The Chañares Formation 1 crops out as part of the Ischigualasto-Villa Unión Basin, which represents a succession of continental deposits composed of 4,000 metres of alluvial, fluvial and lacustrine sediments 2,3 . The basin contains the reddish Talampaya and Tarjados formations as its lower- most units and corresponds to the Synrift 1 tectonic phase. The Talampaya Formation is dated as Induan/Olenekian (Early Triassic) and the Tarjados Formation as Anisian (early Middle Triassic) according to some authors 3,4 . The lower section of the Talampaya Formation is represented by alluvian fan deposits followed by fluvial and playa lake deposits in the middle and upper sections 4. The Tarjados Formation has aerealy extensive outcrops in the Talampaya National Park but at the moment no significant fossil vertebrate remains were reported.