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Zeitschrift/Journal: Beiträge zur Paläontologie
Jahr/Year: 1999
Band/Volume: 24
Autor(en)/Author(s): Summesberger Herbert, Wagreich Michael, Tröger Karl-Armin, Jagt John W. M.
Artikel/Article: Integrated biostratigraphy of the Santonian/Campanian Gosau Group of the Gams Area (Upper Cretaceous; Styria, Austria) 155- 205 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
Beitr. Paläont. 24:155-205, Wien 1999
Integrated biostratigraphy of the Santonian/Campanian Gosau Group of the Gams Area (Upper Cretaceous; Styria, Austria)
by
Herbert Summesberger 1, Michael W agreich 1 2, Karl-Armin T röger3 & JohnW.M. Jagt 4
S ummesberger , H., W agreich , M , T röger , K.-A. & Jagt , 1999. Integrated biostratigraphy of the Santonian/ Campanian Gosau Group of the Gams Area (Late Cretaceous; Styria, Austria). — Beitr. Paläont., 24:153-205, 12 Figures, 15 Tables and 12 Plates, Wien.
Contents I with Hauericeras cf. pseudogardeni (S chlüter ), P achy discus (P.) cf. launayi , Selenoceramus inflexus Abstract, Zusammenfassung ...... 155 (B eyenburg ) and Sphenoceramus aff. angustus (B ey Introduction...... 156 enburg ) is Early Campanian ( bidorsatum Zone, patoo- Lithostratigraphy of the Krimpenbach Formation 157 tensiformis Zone = inoceramid assemblage Zone 29, Systematic Palaeontology 158 CC17-CC19a, Elevata Zone). Wentneralm II yielded Ammonidea...... 158 an inoceramid rich fauna (four taxa), with few ammo Inoceramidae...... 163 nites (three taxa) and echinoids (three taxa) in a green Foraminifera...... '.... 171 ish-grey matrix. Inoceramus cf. bosenbergensis W alas - Conclusions...... 171 zczyk , Inoceramus planus M ünster , Inoceramus aff. Calcareous nannofossils 172 borilensis JolkiCev, Cataceramus balticus cf. halde Biostratigraphical conclusions ...... 174 mensis (G iers ) and the ammonites P achy discus (P) References 176 haldemsis (S chlüter ) and Pachydiscus (P.) tweenianus Abstract (S toliczka ) indicate a Late Campanian age ( Phale- ratum-Polyplocum Zone, Vulgaris-Polyplocum Zone The outcrops Krimpenbach, Wentneralm I and II in the CC19-7CC21, Elevata-Ventricosa Zone). A Cam Gosau Group Gams area (Steiermark, Austria; Fig. 1) panian age for the Wentneralm I and II sections is cor were sampled and studied for their fossil content and roborated by holasteroid and micrasterid echinoids, stratigraphic correlation. The Krimpenbach site yielded with records of Echinocorys sp. 1, Echinocorys sp. 2 three inoceramid taxa: Cordiceramus muelleri cf. (cf. fonticola A rnaud ), Echinocorys gr. subglobosa germanicus (H ein z ), Selenoceramus cf. inflexus (G öldfuss ), Micraster glyphus S chlüter and Micraster (B eyenburg ), Cataceramus balticus (J. B öhm) (subsp. gr. fastigatus/stolleyi. However, separation of the lat indet. and Cataceramus balticus cf. haldemensis ter two species is difficult in the limited material avail (G iers ) indicating a latest Santonian to earliest Cam able. Despite these difficulties the assemblage appears panian age. This is corroborated by a poor nannoflora to support an Early Campanian age for Wentneralm I indicating a Late Santonian to early Early Campanian and a Late Campanian date for Wentneralm II. The age (subzone CC 17b). Wentneralm I yielded inocera- Krimpenbach site yielded no echinoids. mids (four taxa), ammonites (four taxa) and echinoids (three taxa) in a reddish matrix. The age of Wentneralm Zusammenfassung
1 Naturhistorisches Museum Wien, Burgring 7, Die Aufschlüsse Krimpenbach, Wentneralm I und II A-1014 Wien, Austria in der Gosau Gruppe von Gams (Steiermark, Öster e-mail: herbert.summesberger@ nhm-wien.ac. at reich) wurden auf ihren Fossilgehalt und ihre strati 2 Institut für Geologie, Universität Wien, Geozentrum graphische Position hin untersucht. Die Fundstelle A-1090 Wien Althanstraße 14, Austria e-mail: [email protected] Krimpenbach lieferte drei Taxa Inoceramen ( Cordi 3 Bergakademie Freiberg (Technische Universität) ceramus muelleri cf. germanicus (H einz ), Seleno Geologisches Institut, Zeunerstr. - Meißer-Bau ceramus cf. inflexus (B eyenburg ), Cataceramus balti D-09596 Freiberg/Sachsen, Germany cus (J. B öhm) subsp. indet. and Cataceramus balticus 4 Natuurhistorisch Museum Maastricht cf. haldemensis (G iers )) aus dem höchsten Santon und P.O. Box 882, NL-6200 AW Maastricht basalen Campan in grüngrauer Matrix. Die verarmte The Netherlands, E-mail: [email protected] Nannoflora weist auf ein Obersanton- bis tiefes Unter- ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
156 Beitr. Paläont. 24, Wien 1999
Figure 1: Sketch map of the Gams area with the localities Wentneralm I (WI) and Wentneralm II (WII) and Krimpenbach (K). YJ T is the Cretaceous/Palaeogene boundary site of Knappengraben. campan-Alter (Subzone CC 17b) hin. Wentneralm I Lokalität Krimpenbach hat keine Echinodermen ge enthielt Inoceramen (vier Taxa), Ammoniten (vier liefert. Taxa) und Echinoidea (drei Taxa) in einer rötlichen Matrix. Wentneralm I mit Hauericeras cf. pseudo- Introduction gardeni (S chlüter ), Pachydiscus (P) cf. launayi und Selenoceramus inflexus (B eyenburg ), Sphenoceramus The Gosau Group in the Gams area comprises a Late aff. angustus (B eyenburg ) wird in das untere Unter- Turonian to Eocene succession (K ollm ann 1964; campan ( Bidorsatum Zone, Patootensiformis Zone = K ollmann & S ummesberger 1982; K ristan -T ollmann Inoceramen Assemblage Zone 29, CC17-CC19a, & T ollmann 1976, S ummesberger & K ennedy 1996, elevata Zone) eingestuft. Wentneralm II enthielt eine S ummesberger [in:] S chultz & P aunovic 1997). To the reiche Fauna mit Inoceramen (vier Taxa), Ammoniten south of the Gams area ammonite and inoceramid (drei Taxa) und Echiniden (drei Taxa) in einer grünlich bearing sediments were found along the forest road grauen Matrix. Mit Inoceramus cf. bosenbergensis from Gamsforst/Bachler to the Wentneralm (Fig. 1). W alaszczyk , Inoceramus planus MÜNSTER, Inocera These deposits (Krimpenbach Formation) comprise a mus aff. borilensis JolkiCev, Cataceramus balticus cf. southerly neritic facies of the Gams area, subsequently haldemensis (G ier s ), Pachydiscus (P) tweenianus overthrusted towards the northwest onto Eocene (S toliczka ) und Pachydiscus (P) haldemsis S chlüter ) turbidites of the basin fill (Zwieselalm Formation). The ist Obercampan nachgewiesen ( Phaleratum-Polyplo - Late Cretaceous sediments of the Wentneralm area rest cum Zone, Vulgaris-Polyplocum Zone, CC19-7CC21, unconformably upon Triassic carbonates of the Göller Elevata-Ventricosa-Zone). Das campane Alter für die nappe system (lower part of the Aibelmauer slice, Profilabschnitte Wentneralm I und Wentneralm II wird K ollmann 1964:139) and are overthrusted by Triassic durch holasteroide and micrasteride Echinoidea: dolomites of the main body of the Aibelmauer slice. Echinocorys sp. 1, Echinocorys sp. 2 (cf . fonticola Exposures in the Late Cretaceous succession are faulted A r n a u d ), Echinocorys gr. subglobosa (G oldfuss ), and tectonically separated due to overthrusting and later Micraster glyphus S chlüter und Micraster gr. fasti- strike-slip movements along the Ennstal-Puchberg- gatus/stolleyi gestützt. Trotz der an dem begrenzten Mariazell fault and the Göstling fault (e.g. D ecker et Material schwierigen Unterscheidbarkeit der beiden al. 1994, N emes et al. 1995). A more complete and less letzten Spezies scheinen die Echinodermenfaunen das deformed neritic succession is known from the untercampane Alter von Wentneralm I und das ober- Krimpenbach area at the eastern end of the Gams area campane Alter von Wentneralm II zu bekräftigen. Die (K ollmann 1964; W agreich 1995, Figs. 1, 2). ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
S ummesberger , H., et al., Integrated biostratigraphy ... 157
KRIM PEN BACH Field work and collecting was done by W agreich and Summesberger , lithostratigraphy, nanno- and micro Z 0 < 1-- z y-j fossils by W agreich , inoceramids by Troger , echinoids X cr o o by Jagt , ammonites by S ummesberger , who also acted o:
•ft . V V 1 * -v. * it
Figure 3: Wentneralm I. View of the outcrop in 1994. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
158 Beitr. Palaont. 24, Wien 1999
Figure 4: Wentneralm II. View of the outcrop in 1994.
ate content is a mixture of variable amounts of bioclasts Systematic Palaeontology (mostly red algae, echinoderm debris, benthic forami- nifera and bryozoans) and a significant terrigenous Phylum Mollusca component, transported into the area by shelf currents Order Ammonoidea Zittel 1884 with relatively low velocity. Primary stratification was Suborder Ammonitina H yatt 1889 largely destroyed by bioturbation. Superfamily Desmocerataceae Z ittel 1895 Above this red interval, and probably also as a lateral Family Desmoceratidae Zittel 1895 equivalent to it, grey silty to sandy marlstones to marly Subfamily Desmoceratinae Z ittel 1895 siltstones are exposed. In the Krimpenbach section they attain a thickness of up to 30 m and are overlain by Genus Desmophyllites Spath 1929 deep-water marls of the Nierental Formation. In con Type species: Desmoceraslarteti Seunes ; 1892 trast, in the Wentneralm section a second red siltstone by subsequent designation of Spath (1921). interval may be present at the top. Shell layers of ino- ceramids are a conspicuous feature of the grey marl- Desmophyllites sp. indet. juv. stone and locality Wentneralm II is situated within this (PI. 1, Figs. 2, 3) interval. While other macrofossils are. very rare, the Material: SCHU 71/52 b, a single specimen from prominent ichnofauna consists of silicified burrows of Wentneralm I Thalassinoides and rare large Zoophycos, a typical Description: The minute specimen is deform pelagic ichnofossil. The foraminiferal assemblages (p. ed into an ellipse (elongate axis 26 mm). The internal 171) indicate a deeper, outer neritic depositional envi mould is preserved in reddish matrix without adherent ronment, based upon the higher diversity of the benthos shell. The specimen is very involute. Due to poor pres and higher plankton content. Also the nannoflora (p. ervation the degree of inflation cannot be observed. 172) is relatively well developed. This grey, outer With the exception of two constrictions the surface neritic facies is coeval with the pelagic to deep-water seems to have been smooth. turbiditic Nierental Formation in the “Gams basin” Discussion: Distortion and small size make iden (K ollmann 1964:97 ff., W agreich & K renmayr tification difficult. Restored shape of the shell and the 1993:72 ff). visible constrictions suggests this specimen to be a ju Abbreviations venile representantative of the genus Desmophyllites S path . NHMW Museum of Natural History Vienna Occurrence: The known stratigraphical range of SCHU Ing. Lambert S chussler Collection, Leoben the genus (Santonian to Maastrichtian) provides no RE Ruhrland Museum Essen, Germany useful stratigraphical information. Data from associ ated ammonites and other fossil groups show the site Wentneralm I at Gams to be Early Campanian. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
S ummesberger , H., et al., Integrated biostratigraphy 159
Subfamily Hauericeratinae M atsumoto 1938 make it difficult to refer SCHÜ 71/58 to one of the Genus Hauericeras de G rossouvre 1894 high-whorled and apparently closely related species H. Type species Ammonites pseudogardeni pseudogardeni (S chlüter ), H. welschi de Grossouvre Schlüter 1872:54, pl. 16, figs. 3-6. or H. sulcatum (Kner ). H. sulcatum is an Early Maas- trichtian species and known from the Ukraine (Kenne c f (S chlüter 1872) Hauericeras . pseudogardeni dy & S ummesberger 1987:28), Bulgaria (T zankov (Pl. 1, Fig. 1) 1982:29) and Poland (B laszkiewicz 1980:41). H. Compare welschi occurs in the Santonian (L ommerzheim ,
1872 Ammonites pseudogardeni Schlüter : 54, pl. 16, figs. 1995:51). 3-6. Kennedy & Kaplan (1995, p. 18, 20) considered H. 1995 Hauericeras (Hauericeras) pseudogardeni (S chlüter , buszi to be a synonym of H. pseudogardeni ; however, 1872); Kennedy & Kaplan : 18; pls. 1-4; pl. 5, figs. 1,2 (only); pl. 6, figs. 1, 7; pl. 7, fig. 3 (only) (with syn Lommerzheim (1995:52) treated it as a distinct species onymy). ranging from the Santonian to lowest Campanian in 1995 Hauericeras pseudogardeni (S chlüter , 1872); Lom- the Münster basin (e.g. Olfen, Datteln, Recklinghausen; merzheim : 53; pl. 2, figs. 5, 6 (with additional syn W egner 1905:209). onymy). 1997 Hauericeras (Hauericeras) pseudogardeni (S chlüter , H. (Gardeniceras) gardeni (B aily ), H. angustum (Y a - 1872); Kennedy & Christensen : 85; figs. 5 E, 6. be), H. madagascariense Collignon , H.fayoli de Gros
Lectotype: Ammonites pseudo-gardeni S chlüter souvre and H. rembda (Forbes ) can be distinguished 1872, pl. 16, figs.5,6 by subsequent designation of by lower whorl height and wider umbilicus. M atsumoto [in:] M atsumoto et al. (1990:440). Occurrence: The present species was described Material: A single specimen (SCHÜ/71/58) from originally from the Early Campanian of Dülmen Wentneralm I in the Gams area. (L ommerzheim 1995:54). Co-occurrence with Pla- Description: The specimen is a slightly dis centiceras bidorsatum (A. Roemer ), Scaphites bino- torted internal mould of a very large phragmocone, dosus A. Roemer (K ennedy & Kaplan 1995:20,36, preserved in reddish matrix. The specimen is flattened Lommerzheim 1995:54) and the absence of Marsupites by post-mortem crushing to a certain degree. A part of testudinarius (Kennedy & Kaplan 1995:10) confirms the last whorl is restored artificially. The body cham the Early Campanian age of the type locality. Data on ber is absent. The coiling is rather involute, two thirds the the vertical range of H. pseudogardeni vary accord of the preceding whorl being covered. Due to distor ing to the authors. After Kennedy & K aplan 1995 tion coiling changes to about 50 % of the preceding (Fig. 3, p. 20) showed it to range from the Late whorl on the last portion preserved. The flanks are Santonian granulata Zone to the Early Campanian compressed, the whorl section is lanceolate, the keel quadrata Zone. Contradictory are the authors’ state is broken away. The umbilicus is shallow, the umbili ments that H. pseudogardeni appears below Marsupites cal wall is steep and apparently has not been vertical (p. 20): „an der Basis des Ober-Santon -.... unterhalb originally. The umbilical edge is narrowly rounded. der Marsupites Total-Range-Zone“ and (p. 13): „Ob No ribs, tubercles, or constrictions are visible. The ab Hauericeras (Hauericeras) pseudogardeni bereits im sence of constrictions may be due to preservation. Un- Ober-Santon einsetzt muß ohne Revision des Origi nalmaterials fraglich bleiben.“ Lommerzheim (1995:54) D Wh Wb U U% showed H. pseudogardeni to appear first a few metres 406 165 62 107 26,4 aboveMarsupites testudinarius and to be confined to T a b le 1. Dimensions of Hauericeras cf. pseudogardeni the lower part of the Early Campanian (l.c., Figs. 4-6, (Schlüter ), SCHÜ 71/58 from Wentneralm I. 11). Nannostratigraphically the indications given (Lom merzheim 1995) are also contradictory: CC16 in Figs. decipherable fragments of the suture are visible up to 5,7 and CC18 in Fig. 10. We provisionally follow the the end of the last whorl preserved. macropalaeontological data based upon several bore Discussion The lectotype of Hauericeras holes in the Münsterland basin given by L ommerzheim pseudogardeni measures 237 mm in diameter. The larg (l.c.). est specimen figured by Kennedy & Kaplan (1995) Geographical distribution: NW Germany, Sweden and had an estimated diameter of 440 mm. The present specimen is even larger, and would have been about England (M üller & W ollemann 1906; Kennedy & 600 mm, inclusive the body chamber. The specimens Kaplan 1995; Lommerzheim 1995). The occurrence at described by Lommerzheim (1995:53) are much smaller. Gams constitutes the first record from the Tethyan The absence of constrictions and the difference in size Realm. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
160 Beitr. Paläont. 24, Wien 1999
Family Pachydiscidae Spath 1922 project forward over the venter. No branching of the Genus Pachydiscus Z ittel 1884 ribs can be observed. Subgenus Pachydiscus Zittel 1884 Discussion: Pachydiscus tweenianus (Stoliczka ) is distinguished from all other Campanian Pachy Type species Ammonites neubergicus H auer discidae by its characteristic coarse ribbing, which dis 1858:12; pi. 2, figs. 1-4; pi. 3, figs. 1-2, by the subse appears on the body-chamber. quent designation of de G rossouvre 1894:177. P perfidus de G rossouvre (1894:213; pi. 34, fig. 1) from the Late Campanian is a closely related species Pachydiscus (Pachydiscus) tweenianus which differs in having more distant and coarser ribs, (S toliczka 1865) arising from an elongate bulla. (PI. 2, Fig. 2; PI. 3, Fig. 1) P. haldemsis (S chlüter ) (see below), predominantly a Synonymy: Late Campanian species (Kennedy & S ummesberger Kennedy & Jagt 1865 Ammonites tweenianus Stoliczka : 107, pi. 55, fig. 1 1984; 1998), is distinguished by its (only). narrower, somewhat irregular and occasionally branch 1898 Pachydiscus tweenianus Stoliczka ; Kossmat :102. ing ribs. It was decribed in detail by K ennedy & 1925 Pachydiscus tweenianus Stoliczka ; D iener :109. S ummesberger 1984 (p. 158 ff.). 1932 Parapachydiscus twenianus Stoliczka (sic!); Colli- P subrobustus S eunes (1892, pl.4, fig.l), another al gnon :27; pi. 8, figs. 3, 3a. lied Late Campanian species, is distinguished by its ? 1938 Pachydiscus aff. tw ea n i STOL. (sic!); Collignon :61; narrower, bipartite and tripartite ribs. pi. 1, figs. 3, 3a. P. cf. subrobustus S eunes (K ennedy & S ummesberger 1955 Pachydiscus twenianus Stoliczka (sic!); Collignon : 1984:161, pi. 8, fig. 4) from the Late Campanian of 83. the Gschliefgraben (Austria) is very close in style of ? 1984 Pachydiscus cf. subrobustus S eunes ; Kennedy & S ummesberger :161, pi. 8, fig. 4. ribbing and coiling and might be conspecific with P (P) 1996 P. (P .) s u b ro b u stu s S eunes ; Kaplan , Kennedy & tweenianus. Ernst :3 (the discussed specimen from the Gschlief- Eupachydiscus levyi (de Grossouvre (1894:178, pi. 21; graben only). pi. 30, fig. 1) is an Early Campanian ally, differing by its more widely spaced primaries arising from a dis Holotype: by monotypy the original of Stoliczka 1865:107, pi. 55, fig.l from the Arialoor group, NW tinct bulla and strengthening with increasing size. of Arialoor (Tamil Nadu, southern India). P. launayi de Grossouvre (1894:184, pi. 19) from the Material: 2 specimens: SCHU 71/20, SCHU 71/54 Early Campanian (base de 1’assise P1 “de M.A rnaud ”) from the greenish-grey sand-siltstone of Wentneralm n. is stouter and more evolute and has regularly branch Description Both specimens are internal ing ribs which become rursiradiate on the body-cham moulds with corroded surface, preserved in hard sili ber. ceous silty to sandy greenish-grey matrix. SCHU 71/ P. ambiguus de Grossouvre (1894:198, pi. 29, fig. 3) from the assise P3, is a microconch of P. (P) haldemsis 54 (D 211,7, WH 93,6, U 51,6, U% 24,4) shows a sur (Schlüter ) (Kennedy 1986: 45, 50; Text-fig. 11). face partially brownish coloured by a rusty limonitic P lundgreni de G rossouvre (1894:198), (S chlüter crust. SCHU 71/20 (D 172,9, Wb 45) is partially pre 1872:56, pi. 17, figs. 4-7) from the Campanian of NW- served in dark flint connecting the specimen with the Germany differs by its smooth flanks. Ribbing is re adherent sediment. The specimen is coated by a bright duced to the venter. greenish crust. The part of the body-chamber of this P duelmensis S chlüter ; (de G rossouvre 1894:199, specimen is broken and displaced. Coiling is moder pi. 20; 200; K aplan & Kennedy 1995, p. 27, pis. 9- ately involute, with about 60 percent of the preceding 12) from the Early Campanian differs in having a more whorl covered. The whorl height increases rapidly pos inflated section, narrower umbilicus and in having a sibly exaggerated by lateral compaction. The whorl denser and finer ribbing. section appears to have been compressed with conver P. soyaensis M atsumoto & M iyauchi from the Cam gent inner and outer flanks, maximum breadth being panian of Japan has a narrower umbilicus and more mid-flanks or just below mid-flanks. The umbilicus is narrowly spaced, finer ribs. P. sahekii M atsumoto & shallow with a low vertical to subvertical wall and a M iyauchi is also distinguished by a narrower umbili gently rounded umbilical shoulder. Ornament consists cus and more and much more narrowly spaced ribs. A of about 22-24 distant broad and low primary ribs. few primary ribs arise from a distinct bulla at the um They arise without a distinct bulla at the umbilical bilicus. On the adult body-chamber the primaries per shoulder passing over the flank in a slight concavity. sist, while intercalated ribs disappear. About midflank one or two intercalated ribs occur. All E. isculensis (Redtenbacher ) is distinguished by its ribs weaken and finally disappear on the body-cham narrower umbilicus and more inflated flanks and ber, first the intercalated ones, then the primaries. The rounded section. It occurs in the Late Santonian of the youngest part of the body-chamber is smooth. All ribs Austrian Gosau Group (S ummesberger 1979) and has ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
S ummesberger , H., et al., Integrated biostratigraphy 161
also been described from the Early Campanian ( de D Wh Wb U U% G r o sso uv r e 1894:185,187). SC H Ü 71/53 151,5 63,4 — 38,6 25,5 Occurrence: Pachydiscus tweenianus is recorded herein from outside India for the first time. Associated Table 2. Measurements of Pachydiscus (P.) haldemsis (Schlü P. haldemsis, inoceramid bivalves (p. 163), echinoids ter ) from the Late Campanian of Wentneralm II. and nannoflora (CC19 - ?CC21; p. 172) indicate an Late Campanian age. According to poor data it seems Discussion: The present species differs from co to be a Tethyan species. occurring Pachydiscus (P) tweenianus by its finer and more densely spaced, slightly falcoid ribs. Pachydiscus (Pachydiscus) haldemsis Occurrence: P (P) haldemsis is a widely dis (SCHLÜTER 1867) tributed species predominantly known from the North (PI. 2, Figs.l, 3) ern Temperate Realm (K e n n e d y & S ummesberger , 1984:160). The present record is the first from the Synonymy: Tethyan Realm. The stratigraphic range of P (P) 1867 Ammonites haldemsis Schlüter : 19, fíg. 1. koeneni de G r o sso uv r e , which according to K e n n e d y 1984 Pachydiscus haldemsis (S chlüter ); Kennedy & Sum - & S ummesberger (1984) is a synonym of P. (P) mesberger : p. 158; pi. 4, figs. 1-5; pi. 5, fig.l; pi. 6, haldemsis , is the Late Campanian Phaleratum to Poly- fig. 2; pi. 7, figs. 1-11; pi. 13, fig. 1; pi. 14, fig. 2 (With synonymy). plocum Zones (B laszkiewicz 1980:42, table 1) in the 1986 Pachydiscus haldemsis (S chlüter ); Kennedy :45; pl.4, Vistula River valley, where it is best dated. H anco ck figs. 1-3; pi. 5, figs. 7-14; text-figs. 11 A-D , F, G; 17 et al. (1993:143, text-fig. 4) showed this to be approxi (With additional synonymy). mately Belemnitella minor Zone and nannozones 1996 Pachydiscus (Pachydiscus) haldemsis Schlüter ; San CC19/CC20. In Belgium it occurs in the Late Cam tamaría Zabala :7; pi. 1, figs. 1 ,4 , 5. panian Zeven Wegen Member (Gulpen Fm.), together 1997 Pachydiscus (Pachydiscus) haldemsis (Schlüter 1867); (G riepenkerl ) (K en Kennedy & K aplan : p. 40, pi. 4, figs. 5-8; pi. 5, fig. 4; with Neancyloceras ? phaleratum pi. 6, figs. 1, 2; pi. 7, figs. 2, 3; pis. 8,9; pi. 10, figs. 5, n e d y & Jagt 1998). 8 (With additional synonymy). 1998 Pachydiscus (Pachydiscus) haldemsis (S chlüter ); Pachydiscus (Pachydiscus) cf. launayi Kennedy & Jagt :158, pi. 1, figs. 2-4. de G r o sso uvre 1894 Lectotype: The lectotype of Ammonites haldemsis (PI. 3, Fig. 2; PI. 4, Fig. 1; PL 5, Fig.l) designated by Kennedy & S ummesberger 1984:158 is Compare: the original of S chlüter 1867 (pi. 3, fig. 1) from the Late Campanian of Haldem, Westphalia, refigured by 1894 Pachydiscus launayi de G rossouvre : 184, pi. 19. ? 1955 Anapachydiscusfranciscae Collignon : 53; pi. 15, figs. Kennedy & S ummesberger (1984: 14, 2) pi. fig. 1-3. Material: a single specimen, SCHÜ 71/53 from 1986 Pachydiscus (Pachydiscus) launayi de G rossouvre ; Gams, Wentneralm II. Kennedy :38; pis. 2, figs. 1, 2; pi. 7, figs. 6,7; pi. 10, Description: The adult specimen is a deformed fig. 15; pi. 13, fig. 2, 3, 6; text-figs. 4C, 5B (With internal mould slightly elongated to an ellipse. The synonymy). surface is abraded by tectonic activity and partially 1989 Pachydiscus (Pachydiscus) aff. la u n a yi de G ross ouvre ; Jagt :8; pi. 6, figs. 2-5. coated by a limonitic brown crust. Coiling is moder 1995 Pachydiscus (Pachydiscus) cf. la u n a yi de G ross ately involute, with about 60 percent of the preceding ouvre ; W ippich :52; pi. 2, figs. 3,4. whorl covered. The whorl height increases moderately, 1996 Pachydiscus (Pachydiscus) launayi (de G rossouvre , possibly exaggerated by lateral compaction. The whorl 1894); Santamaria Zabala :7; pl.l, fig. 3. section appears to have been compressed with conver 1998 Pachydiscus (Pachydiscus) launayi de G rossouvre ; gent inner and outer flanks, maximum thickness being Kennedy & Jagt : 159, pi. 6, figs.l, 2. mid-flanks. The umbilicus is shallow with a low verti Material: 3 large specimens, SCHÜ 71/56, SCHÜ cal to subvertical wall and an abruptly rounded um 71/46, NHMW 1997/zl27/l and a large fragment, bilical shoulder. Ornament consists of about 25 distant NHMW 1997/z 130/1. broad and low primary ribs and approximately 40 in Description: all specimens are internal moulds, tercalated ones. The primary ribs arise at or close to preserved in reddish matrix, diagenetically flattened, the umbilical shoulder, first curve backward and flex laterally compacted and distorted (e.g. NHMW 1997/ forward at mid-flank, projecting forwards again over zl27/l) by tectonical stressing. This is indicated also the venter. The distances between the primaries are by slicken-side striation and calcitic crusts at the sur slightly irregular becoming larger towards the end of face. Nevertheless observations are still possible and the body-chamber. The short and regularly spaced in- justify a description and tentative interpretation. tercalatories arise close to the ventrolateral shoulder Coiling seems to have been moderately involute with recalling the ventral ribbing of P (P) neubergicus. The rapidly increasing whorl-height and with about 60 or sutur is not visible. more % of the preceding whorl being covered. The ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
162 Beitr. Palâont. 24, Wien 1999 umbilicus is shallow with oblique umbilical wall and ribs per whorl and 20 intercalated ones at a diameter distinct umbilical shoulders. The ornament is preserved of 85 mm. The primaries arise at the umbilical shoul in places. Strong and relatively narrowly spaced ribs der, the intercalatories at about mid flank. The ribs arise at the umbilical shoulder, are straight or slightly become more widely spaced at the beginning of the prorsiradiate on the flanks, fading out somewhat in the body-chamber, being rursiradiate at the smaller diam outer third of the flanks and crossing the venter in a eter and becoming straight towards the final portion. distinct convexity. There is a distinct change of orna ment already on the phragmocone: ribs become indis D Wh Wb U U% tinct undulations or efface almost completely. Partially SCHÜ 71/52 a 103,6 43 — 28,5 27,5 visible sutures (NHMW 1997/z/127/1 ) are undecipher able. Table 4. Measurements of Pachydiscus (P.) sp. indet. juv. of Discussion: It seems most likely that the large SCHU 71/52 a from Wentneralm I. pachydiscids from the Wentneralm I belong to Pachy- Discussion: Pachydiscus (Pachydiscus) sp. indet. discus launayi de G r o sso uv r e . In view of the poor pre juv. is very close to Pachydiscus (Pachydiscus) servation, open nomenclature is used. They differ from haldemsis as described by K e n n e d y & S ummesberger co-occurring Pachydiscus (Pachydiscus) sp. indet. juv. (1984), de G rosso uvre (1894; P. (P.) koeneni ) and B la in having more distinct and partially rursiradiate ribs szkiew icz (1980; P. (P.) koeneni). It differs from these and the gently rounded umbilical shoulder of the latt in having a flatter umbilical wall, straighter ribs and er. lower whorl height. Open nomenclature is preferred; Occurrence: Pachydiscus (Pachydiscus) launayi other Early Campanian congeners are less close. The de G rossouvre is definitely an Early Campanian spe material is too poor for the erection of a new species. cies known from the French Aquitaine basin in Unit C Occurrence: A sample taken from the specimen I (N eumann & P latel 1983:117, tab. 1), or the assises yielded a poor nannoflora (p. 173) indicating nanno- P 1 and P2 of A rnaud (K ennedy 1986:17, table 2), from zones CC17-CC22b, with Micula cf. praemurus sug Madagascar (Collignon 1955:36; pi. 5, figs, la-b) and gesting a Late Santonian to Late Campanian age. from Liège and Limburg (Belgium) (Jagt 1989, Kenne dy & Jagt 1998). The co-occurrence ofP(P.)cf. launayi Superfamily Hoplitaceae H. DOUVILLE 1890 and H. cf. pseudogardeni is a strong indication that the Family Placenticeratidae H yatt 1900 Wentneralm I site is situated below the first occurrence Genus Placenticeras MEEK 1876 of Eupachydiscus levyi (de G rossouvre ) in the Gonio- teuthis quadrata Zone sensu B laszkiewicz (1980, ta Placenticeras spec, indet. juv. ble 1) and in unit C II (N eumann & Platel 1983:117, table 1) in the traditional Early Campanian Placenti- Material 1 specimen (NHMW 1997z/0138/1) ceras bidorsatum Zone. from the Campanian of Krimpenbach, collected in 1996. Description: The specimen is a badly worn and D Wh Wb U U% elongated inner mould of a juvenile whorl, preserved SC H Ü 71/56 291 138,7 52,4 62,8 21,6 in greenish-grey sandy to silty matrix. The shape is SC H Ü 71/46 210,9 85,6 41,7 59,5 28,2 NHM W 97/Z127/1 170 78 26,5 37 21,7 typical of juveniles of the genus: a high lanceolate whorl section, the umbilical shoulder gently rounded
Table 3. Measurement of Pachydiscus (P.) cf. la u n a yi de with an oblique umbilical wall. The venter is very nar G rossouvre from the Early Campanian of Wentneralm I. The row and concave. The double keel is entire. The maxi values of SCHÜ 71/46 and NHMW 1997/zl27/l have been mum width of the whorl is in the inner third of the influenced by distortion. flanks. The flanks gently converge towards the venter and apparently were ornamented by delicate falcoid Pachydiscus (Pachydiscus) sp. indet. juv. ribs, nearly invisible now by post-diagenetic corrosion. (PI. 4, Fig. 2) The umbilicus is covered by sediment, which makes it Material: a single specimen, SCHÜ 71/52 a, from impossible to determine whether there were umbilical Wentneralm I. tubercles or not. Maximum length of the elongated di Description: SCHÜ 71/52 a is a relatively small ameter is 54 mm, width is about 7 mm. No suture is internal mould of a phragmocone with partially pre visible. served body-chamber. It is preserved in a reddish ma Discussion Open nomenclature is preferred, trix without adherent shell. The relatively slender gen since juveniles of the Late Santonian polyopsis group eral shape seems to have been slightly exaggerated by or the Early Campanian milleri-bidorsatum group are lateral compaction. The umbilicus is relatively shal indistinguishable. low with an oblique umbilical wall and a gently Occurrence: Krimpenbach, Steiermark. Co-oc rounded umbilical shoulder. It has about 30 primary curring inoceramids of the muelleri group (p. 164) ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
S ummesberger , H., et al., Integrated biostratigraphy 163 make a latest Santonian age likely, i.e. Paraplanum Class Bivalvia Zone in ammonite terms. Supraorder Pteriomorphia B eurlen 1944 Order Pteroidea N ew ell 1965 The Campanian Ammonite Faunas of the Gosau Family Inoceramidae G iebel 1852 Group in the Gams area
1. Krimpenbach (greenish-grey sediment; Late San Genus Cataceramus Cox 1969 tonian): (ex H einz nomen nudum) Placenticeras spec, indet. juv. 2. Wentneralm I (reddish sediment, Early Campanian): Type species Inoceramusgoldfussianus d ’O r - Desmophyllites sp. indet. juv. big n y 1846 (= Inoceramus balticus B öhm 1907) Hauericeras c f. pseudogardeni (S chlüter 1872) Cataceramus balticus (B ö h m ) subsp. indet. P achy discus (Pachydiscus) cf. launayi de G ro sso u - (PI. 8, Fig.6 ; Text-fig. 6) vre 1984 Pachydiscus (Pachydiscus) sp. indet. juv. Lectotype: the original ofB öhm (1909: pi. 11, 3. Wentneralm II (grey sediment, Late Campanian): fig. 2) by subsequent designation ofG iers (1964). Pachydiscus (Pachydiscus) haldemsis (S chlüter Compare 1867) Pachydiscus (Pachydiscus) tweenianus (S toliczka 1997 Cataceramus balticus (B öhm); W alaszczyk : 8; pi. 12, 1865) figs. 1, 2, 4, ?5 (With complete synonymy). Material Internal moulds of two right (NHMW Palaeogeographic and palaeoecologic conclusion 1997/z/158/3,8) and two left valves (NHMW 1997/z/ Wentneralm I indicates palaeogeographical connections 158/1,12) from Wentneralm I in the Gams area. with the NW European Cretaceous, Pachydiscus (Pa Distance D U A DUB DUC chydiscus) haldemsis from Wentneralm II being a from UP widely occurring species in NW Europa and Poland 10-30 mm 4,87 mm 2,76 mm (K en n e d y & S ummesberger 1984) here recorded from 30-50 mm 5,00 mm 5,70 mm 5,10 mm the Tethyan Realm for the first time. Its occurrence in 50-70 mm the Helvetic unit of the Gschliefgraben as well (K e n ne Table 5. Average undulation intervals (DU) in distances from dy & S ummesberger 1984), intermediate between the the umbonal pole (U P). A = N HM W 1997/z/158/l, B = NHM W Northern Temperate and the Tethys Realm, underscores 1997/z/158/3, C = NHM W 1997/z/158/8, all from Wentneralm I. the importance of the Gschliefgraben for palaeogeo graphical reconstructions. All ammonites from the Wentneralm are relatively large or very large. There is no known locality in the Cam panian of the Gosau Group with comparable faunas or comparable size of the specimens. The mechanism behind the possible selection by transportation are un known, local environmental conditions possibly re sponsible for extreme growth are not known either. Echinoids are the only epi/endobenthic dwellers, in- oceramids the only bivalve group present. Predomi nance of the latter two invertebrate groups leads to the assumption that prevailing environmental conditions Figure 5: Diagram showing comparison of H/L ratio at were close to those of the Gschliefgraben (K e n n e d y & Cataceramus balticus (B öhm) subsp. indet. from Wentneralm I
S ummesberger , T röger , S ummesberger & S ko um al , with H/L ratio of C. balticus balticus (B öhm) from the Early Campanian of the Subhercynian Cretaceous Basin. Ja g t , this volume): an open marine environment, rela tively deep water, muddy seafloor, precluding the growth/occurrence of a diverse benthic or sessile fauna. Remarks In shape and H/L-ratios there are dis tinct differences between Cataceramus balticus B öhm subsp. indet. and Cataceramus balticus balticus J. B öhm including Cataceramus balticus haldemensis (G iers ). The shape, development of the umbonal re gion and the total angle of Cataceramus balticus marcki (G iers ) are comparable. In contrast to Cataceramus balticus (J. B öhm ) subsp.indet. in Cataceramus balticus ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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marcki (G iers ) the H/L-ratio shows strong increases of the anterior margin are missing in all specimens. (170-180). NHMW 1997/z/l59/5 without beak. All specimens are Occurrence: Cataceramus balticus was first de preserved in a greenish-grey matrix. scribed (B öhm 1909) from the Early Campanian Description Medium- to large-sized; inequila Dülmener Schichten. It occurs in the European North teral; shape. C. balticus- like. Beak not separated from ern Temperate Realm. Its occurrence in Romania (L upu the wing. Anterior margin straight to convex at the & S ornay 1978 has recently been confirmed by W al - umbonal pole. Geniculations with a change of the un aszczyk (1997). C. balticus occurs in several subspe dulation shape and distance at all specimens. The H/L cies (C. b. balticus (B öhm), C. b. ellipticus (G iers ), C. ratio is comparable with that of the holotype and a b. aff. haldemensis (G iers ), C. balticus (B öhm) subsp. population of this species from Libya (see Text-fig.6). indet.) in the Campanian of the Austrian Gschliefgra- Total angles: 95-125° (pin part affected by compac ben (T röger et al. 1999). W alaszczyk (1997) showed tion). Distance of the undulations varies between 2,5- Cataceramus balticus to range from Early Campanian 5mm. Endocostea scar in specimen NMHW 1997/z/ to the early Late Campanian. C. b. marcki (G iers ) 159/8. ranges from the latest Santonian to the earliest Late Remarks: The specimens from Wentneralm agree Campanian (Polyplocum Zone). with Cataceramus balticus haldemensis (G iers ) in gen eral shape, shape of the umbonal pole, H/L ratio and Cataceramus balticus cf. haldemensis (G iers ), 1964 geniculations with a population from Libya. The H/L (PI. 9, Figs. 2, 4, 5; PI. 10, Fig. 1; Text-fig. 6) ratio in the holotype is higher than that in the speci mens described (Text-fig.6). Synonymy: For complete synonymy see W alasz Stratigraphic distribution: Cataceramus balticus hal czyk (1997). Following G iers (1964) we consider ,In- demensis (G iers ) is the index fossil of the haldemensis oceramus ‘ haldemensis G iers to be a representatative Zone of the Late Campanian (W alaszczyk 1997). of the Cataceramus balticus group, unlike W alasz Geographic distribution: Europe, N Africa czyk (1997). Material: 4 internal moulds of left valves (NHMW Cataceramus ex gr. balticus (BÓHM) 1997/z/159/4, 5,6,13?) and 3 internal moulds of right (PI. 7, Figs. 1, 3) valves NHMW 1997/z/l 59/ 8,9,10) from Wentneralm II, Gams area. Material: Two badly preserved internal moulds, Preservation: All specimens are incomplete and a single right (NHMW 1997/z/144/3) and a left valve deformed by compaction (NHMW 1997/z/l59/ (NHMW 1997/z/144/5) from Krimpenbach. 4, 8, 9, 10) with radial cracks. Portions of the wing and Preservation: The specimens are deformed by compaction and incomplete. Parts of the wings and the central margins are missing. Description: Mediumsized, inequilateral, shape balticus-like.
1 2
Height (incomplete): 60,0 mm 65,0 mm Length (incomplete): 60,0 mm 93,5 mm Hinge line (length): 30,2 mm 38,8 mm Anterior margin: 10,8 mm 10,4 mm Total angle: 120°
Table 6. Measurements of Cataceramus ex gr. balticus', 1 is NHMW 1997/z/144/3, 2 is NHMW 1997/z/0144/5.
The umbo is slightly bent to the anterior margin. It slightly rises above the hinge line, the beak is not sepa rated from the wing. Hinge line straight. The growth lines are bent to the beak at the wing (125-130°). Growth axis slightly prosocline (25-50° in NHMW 1997/z/144/3. H/L ratio in NHMW 1997/z/144/3 as cending from 70 to 83 %. Undulations toprounded. No „Anwachsschnittreifen“ - see Cordiceramus muel-
Figure6: Diagram showing H/L ratio of Cataceramus balticus leri germanicus (H einz ). A small Endocostea scar is cf. haldemensis (G iers ) from Wentneralm II and the presence visible in NHMW 1997/z/144/3. of geniculations and of E n d o co stea scars. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
S ummesberger , H., et al., Integrated biostratigraphy 165
Distance from UP DU Distance DU A DU B 10-30 mm 2,41 mm from UP (mm) 30-50 mm 3,86 mm 10-30 mm 3,51mm 3,60 mm 50-70 mm 5,20 mm 30-50 mm ------
Table 8. Average undulation intervals (DU) in distances from Table 7. Average undulation intervals (DU) in different umbonal pole(UP). A = NHMW 1997/z/1958, B =d ’O rbigny distances from the umbonal pole (UP) ascending in NHMW collection no. 7592 A . 1997/z/144/3. Total angle (94-110°, lectotype:102°). Remarks according to the shape both specimens belong to the balticus group. The similar Late Santonian Remarks: The specimens from Wentneralm agree cordiceramids can be excluded by lack of the ,,An- in shape and development of the beak with the lecto- wachsschnittreifen“ But in the Early Campanian In- type. Endocostea scars are absent. The H/L ratio of the oceramus balticus balticus (J. BOHM) the H/L-ratio Wentneralm specimens differs remarkably from that is higher. of the lectotype. It is possible, that they belong to Early Campanian precursors of the Endocostea impressa Genus Endocostea W hitfield , 1877 group. Type species Inoceramus (Endocostea) typi- Genus Selenoceramus H einz , 1932 cus W hitfield Type species Inoceramus selenae S eitz 1967 Endocostea aff. impressa (d ’O rbigny ) (PI. 8, Figs.l, 2; Text-fig. 7) Selenoceramus inflexus (B eyenburg ) (PI. 7, Figs. 2, 4 (cf.); PI. 8, Figs. 3, 4; PI. 10, Synonymy Figs. 2, 3; Text-fig. 8) 1842 Inoceramus impressus d ’O rbigny : 515, pl.409. Synonymy * 1957 Inoceramus impressus d ’O rbigny -S ornay : Paleont.
Univers. figs. 4, 5; N.S., no. 129, figs. 1-5. 1936 Endocostea inflexa Beyenburg : 295, pi. 11, figs. 1-3; pi. 12, figs. 1 ,3 . Remarks According to S ornay (1957), the lecto 1967 Inoceramus (Selenoceramus) inflexus B eyenburg ; type of Endocostea impressa d ’O rbigny is no.7592 A Seitz: 98-101; pi. 12, fig. 5; pi. 19, figs. 3, 4; pi. 20, in d ’O rbigny ’s Collection (refigured by S ornay (1957, figs. 1-3; text-figs. 4a, b, 18. fig.4) from the Campanian/Maastrichtian boundary. Material Internal moulds of left (NHMW 1997/ Material 2 internal moulds of right valves z/158/2,10) and right valves (NHMW 1997/z/158/ (NHMW 1997/z/158/9,11) from Wentneralm I. 5,6,7) from Wentneralm I. Internal moulds of left (cf.- Preservation Both specimens are incomplete. NHMW 1997/z/144/4) and right valves (cf. NHMW Parts of the wing and the ventral margin are missing. 1997/z/144/9) from Krimpenbach. Description Small to medium-sized, inequi Preservation All specimens are incomplete lateral. Shape elongated arcuate. Beak vaulted, dis internal moulds deformed by compaction especially in tinctly separated from the wing, prosogyrate. Hinge the umbonal regions. Parts of the wing and the ventral line straight. Anterior margin slightly concave at the margin are missing. umbonal region. Undulations (concentric ribs) follow Description Valve medium-sized, inequilateral. ing the shape outline, top-rounded. The undulations Shape axe like. Anterior margin convex. No anterior are curved to umbonal pole (angle 115-120°). Total auricle. Beak terminal to subterminal, slightly proso angles: 94°, 110°(lectotype 102°). gyrate and slightly differentiated from the wing. Gen- iculations are present. The interval between the undu lations (concentric ribs) strongly increases from the geniculation to the ventral margin. The valve is even near the ventral margin in some cases. The undulations are top-rounded.The average undulation intervals fluc tuate between 2,0-4,8 mm (mainly 3,0 mm) on the umbonal regions (10-30 mm from UP). Total angle: 95-145° (mainly 120-130°). The trend in of the L/H- ratios (Text-fig.7) agrees with those described by S eitz (1967) for Selenoceramus inflexus. Remarks Selenoceramus inflexus differs from Figure 7: Diagram showing H/L ratio in E n d o c o stea aff. members of the Cataceramus balticus group includ im pressa (d ’O rbigny ) from Wentneralm I. Lectotype: no.7592 ing Endocostea barabini (M orton ), which are similiar A in d ’O rbigny ’s collection, r = right valve. in shape, particularly by the strong decrease in the H/ ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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Figure 8: Diagram showing L/H ratio in Selenoceramus inflexus (B eyenburg ) from Wentneralm I. + geniculation; r = right valve, 1 = left valve.
Figure 9: Diagram showing Na/Ha- and Vo/Ha ratio in S ph e L-ratio between H = 10-30 mm and the shape of the n oceram u s aff. an gu stu s (B eyenburg ) from Wentneralm I. r = cross section. right valve, 1 = left valve. Stratigraphic distribution: Early Campanian, patoo- tensiformis beds (S eitz 1967). Stratigraphic distribution: Early Campanian. Accord Geographic distribution: W Europe (Münsterland). ing to S eitz (1965) upper portion of the patootensi- formis-beds. Genus Sphenoceramus J.Böhm, 1915 Geographic distribution: Europe and W Asia (V oigt 1996) Type species Inoceramuscardissoides G old - fuss , 1922 Genus Inoceramus J.Sowerby 1814
Sphenoceramus aff. angustus (B eyenburg 1936) Type species: Inoceramus cuvieri J. S owerby (PI. 8, Fig. 5; Text-fig. 9) (Cox 1969, p. N.315 by subsequent designation of Cox (1969:N315). Remarks For complete synonymy see S eitz (1965). The lectotype of Sphenoceramus angustus is Inoceramus cf. bosenbergensis W alaszczyk 1997 the specimen of Inoceramus lobatus M ünster figured (PI. 9, Fig. 3; Text-fig. 10) by W egner (1905:164, text-fig.7). Material: A single internal mould of a right valve H olotype is RE A 1438, the original of W alasz (NHMW 1997/z/158/13) from Wentneralm I. czyk (1997, pi. 27, fig. 1) (Ruhrland Museum, Essen, Preservation: Poorly preserved (without shell) Germany). and incomplete. Portions of the wing and the posterior Synonymy: For complete synonymy see W alasz margin are missing. Flattened by compression; radial czyk (1997). cracks especially at the anterior margin. Material: 3 internal moulds of left valves (NHMW Description: Medium-sized; shape rhomboidal 1997/z/159/2,3,7 from Wentneralm II, Gams area. to arcuate; inequilateral. Height (incomplete): 52.5 mm; Preservation: All specimens are incomplete. length (incomplete): 50,8 mm. Total angle deformed Portions of the wing, of the posterior margin and the by compaction: 110°. Anterior margin convex, com ventral margins are missing. The umbo is not preserved pletely deformed by compaction. Hinge line straight. in NHMW 1997/z/159/2. All specimens are slightly Undulations poorly preserved, growth lines not visible. deformed by compaction pressure, partly with radial Growth axis prosocline (angles: 53-57°). Average un cracks. dulation distances: 10 mm (30-50 mm from the Description: Medium-sized, inequilateral, shape umbonal pole). For Na/Ha and Vo/Ha ratio see Text- subtrapezoidal. Beaks slightly diffentiated from the fig. 9. wing, prosogyrate. Anterior margins straight to con Remarks Based on shape, a comparison with cave immediately at the umbonal pole. Geniculation Sphenoceramus angustus is possible. However the with a change of undulation shape und intervals at H= absence of the greatest part of the wing precludes a 52,5 mm (NHMW 1997/z/159/7). The undulations are more precise identification. flexure-like bent at the corner hinge line/posterior ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
S ummesberger , H., et al., Integrated biostratigraphy 167
Figure 11: Diagram showing H/L ratio in Inoceramus planus M ünster from Wentneralm II compared to Giers ’s (1964, table Figure 10: Diagram showing H/L ratio in In o cera m u s cf. 3, fig. 2) type specimen from the P o lyp lo cu m Zone of Haldem bosenbergensis W alaszczyk from Wentneralm II. r = right (Germany), r = right valve. valve, 1 = left valve. Text-fig.ll. Small geniculation at the anterior margin. Distances from UP A B Remarks: The present specimen is comparable to 10-30 mm ------3,13 mm 30-50 mm 4,75 mm 4,75 mm the one figured by G iers (1964: pi. 3, fig. 2) and W a laszczyk (1997, pi. 30, fig. 4). Table 9. Average undulation intervals (DU) in different Stratigraphic distribution: Late Campanian ( stobaei/ distances from the umbonal pole (UP). A = NHMW 1979/z/ basiplana-polyplocum Zones). 159/2, B = N H M W 1979/z/159/3. Paleogeographic distribution: W and E Europe (W a laszczyk 1997). margin at the specimen NHMW 1997/z/159/7. The H/ L ratio (see Text-fig. 10) is comparable to that of speci Inoceramus aff. borilensis JolkiCev mens from the type locality Bosenberg in Westphalia. (PI. 10, Fig. 4) Total angles: 110°, 128°. Growth axis prosocline, Compare: slightly bent to the hinge line (40-56°). 1997 In oceram u s aff. b o rilen sis JolkiCev; W alaszczyk , p. 37; Stratigraphic distribution: Late Campanian ( basiplana/ pi. 28, figs. 1-5 (With synonymy). stobaei-vulgaris basiplana Zones) according to W a Material A single internal mould (NHMW 1997/ laszczyk (1997). z/159/13) of a poorly preserved left valve from the Geographic distribution: W Europe, Westphalia (W a Wentneralm II. laszczyk 1997). Preservation: Incomplete. The wing, posterior Inoceramus planus M ünster 1836 margin, ventral margin and large portions of the ante (PI. 9, Fig. 1; Text-fig. 11) rior margin are not preserved, thus precluding an ex act identification. For synonymy see W alaszczyk (1997). Description Tall. Shape of the umbonal region Lectotype: Specimen figured by Goldfuss (1834 balticum- like. The geniculation in the region of the -40), pi. 113, fig. lb, according to G iers (1964). Material: Internal mould of a single right valve umbo allows a comparison with two specimens of In (NHMW 1997/z/l 59/1) from Wentneralm II, Gams oceramus aff. borilensis JolkiCev, described by W al area. aszczyk 1997 (pi. 28, figs. 1, 5) from the Vorhelm Preservation Preserved in a greenish-grey Schichten (Late Campanian stobaei/basiplana-vulga- matrix; incomplete. Portions of the anterior margin and ris/basiplana Zone) of the Münsterland (Germany). of the wing are missing. Flattened by compaction pres sure. Genus Cordiceramus H einz 1932 Description Medium-sized; inequilateral. Type species: Inoceramus cordiformis SOW- Shape subcircular. Beak prosogyrate and slightly di ERBY 1825 vided from the wing. Hingeline straight. Anterior mar gin convex. Total angle: 130° specimen of Giers 1964: Cordiceramus muelleri cf. germanicus (H einz ) 130°). Delta increasing from 32° to 50°. Average dis (PI. 6, Figs. 1-3; Text-fig. 12) tance of the undulations between 10-30 from the Synonymy umbonal pole 2,13 mm (specimen of G iers 1964: 3,61 1928 Inoceramus germanicus Heinz : 82. mm) and 30-50 mm from the umbonal pole 2,76 mm 1933 Germanoceramus germanicus Heinz ; Heinz : 250, pi. 21, fig. 2. (specimen of G iers 1964:4,14 mm). For H/L ratio see ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
168 Beitr. Paläont. 24, Wien 1999
shape. Corners K 1-3 (see Seitz 1961) rounded. So- called „Diagonalleiste“ developed in NHMW 1997/z/ 144/6 (pi. 6, fig. 3 - arrow). Growth axis prosocline, slightly concave toward hinge line. All growth lines cut the undulation under pointed angles (pi. 6, figs. 1- 3). For Na/Ha- and Vo/Ha ratio see Text-fig. 12. Remarks: The Na/Ha ratio of the present speci mens is comparable to that of Cordiceramus muelleri germanicus (H einz ) described by S eitz (1961) - see Text-fig. 12. In the holotype and specimens figured by Seitz (1961:134) the average intervals of the undula tions in 10-30 mm and 30-50 mm distance from the umbonal pole are larger (Text-fig. 12). Total angles: 132-148°. Stratigraphic distribution: Late Santonian (S eitz 1961). Geographic distribution: W Europe
Biostratigraphic conclusion The inoceramid assemblages at the three localities are different. Krimpenbach of latest Santonian to earliest Campanian based on the occurrence of Cordiceramus muelleri cf. germanicus (H einz ). Selenoceramus cf. inflexus (BEYENBURG)and Cataceramus aff. balticus (J. B öhm) may testify to earliest Campanian age. Went- neralm I of Early Campanian age (patootensiformis Zone) is based on co-occurence of sphenoceramids and Selenoceramus inflexus. The inoceramid assemblage Figure 12: Diagram showing H/L-, S/L and Vo/L ratio in of Wentneralm II is typical of the Late Campanian in Cordiceramus muelleri cf. g erm an icu s (Heinz ) from Krimpen- terval agdjakendsis/vorhelmensis - haldemensis Zones bach. r = right valve, 1 = left valve. of W alaszczyk (1997), which correlate approximately
Distance from UP DU A DU B DU C with the traditional vulgaris- polyplocum Zone. The three localities yielded the following inoceramid 10-30 mm 2,57 mm 2,75 mm 3,3 mm 30-50 mm 3,92 mm 4,60 mm 6,0 mm assemblages: 50-70 mm 6,63 mm - 10,0 mm KRIMPENBACH (greenish grey matrix): Cordiceramus muelleri cf. germanicus (H einz ) Table 10. Average undulation intervals (DU) in different Selenoceramus cf. inflexus (B eyenburg ) distances from the umbonal pole (UP). A = NHMW 1997/z/ Cataceramus aff. balticus (J. B öhm) 144/6, B = N H M W 1997/z/144/14, C = holotype. Total angles: 132-148°. WENTNERALM I (reddish matrix): Selenoceramus inflexus (B eyenburg ) 1961 Inoceramus (Cordiceramus) mtilleri germanicus Heinz ; Seitz: 131, pi. 7, fig. 6; pi. 8, figs. 1, 6, 7; pi. 15, fig. 1; Sphenoceramus aff. angustus (B eyenburg ) text-figs. 29, 30. Cataceramus balticus (B öhm) subsp.indet. Endocostea aff. impressa (d ’O rbigny ) Holotype according to Seitz (1961:131) is Hg.57 in the Museum of Liinen (H einz 1933, pi. 21, fig. 2). WENTNERALM II (greenish-grey matrix): Material: 2 internal moulds of left valves (NHMW Cataceramus balticus cf. haldemensis (G iers ) 1997/z/144/6,14) and one internal mould of a right Cataceramus sp. valve (NHMW 1997/z/144/1) from the Krimpenbach Inoceramus cf. bosenbergensis W alaszczyk area. Inoceramus aff. borilensis JolkiCev Preservation Incomplete. Portions of the Inoceramus planus M ünster wings, of the anterior margins and the ventral margins are missing. The specimens are flattened by compac Echinoidea tion. The holasteroid genus Echinocorys L eske 1778 and the Description: Medium-sized,inequilateral,shape micrasterid Micraster L. A gassiz 1836 are amongst the elongated subtrapezoidal. Beak not separated from the most characteristic irregular echinoid taxa of the boreal/ wing. Anterior margin slightly convex to straight. Hinge temperate North European Province and the Northern line straight. Course of the undulations follows the Transitional Subprovince of northern Spain. It has been ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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shown that their various species can be used to corre to the turrita group, which according to Lambert (1903) late faunal events or migrations within a sequence is restricted to the earliest quadrata Zone (Early Cam stratigraphic scheme (Ernst & Wood 1996). Of Echi- panian). However, H ancock et al. (1993) considered nocorys it has often been claimed that numerous sp e E. turrita in southern England to characterise the cies* (or rather, morphotypes) are good marker fossils youngest part of the Early Campanian (i.e. top of for various portions of the European Late Cretaceous quadrata Zone), which corresponds to what Kuchler (Wright & Smith 1987; H ancock et al. 1993). How & Kutz (1989) recorded for northern Spain. ever, taxonomy still lacks stability and is based mainly The present specimen cannot be confused with repre on largely typological interpretations of authors (e.g. sentatives of the conica group, which in NW Germany Lambert 1903; Smiser 1935). A modem revision of the (Ernst 1972, 1975; Schulz et al. 1984) ranges from various ,species4 groups is called for. the papillosa Zone (Early Campanian) to the conica / The present material comprises three specimens from mucronata Zone (late Campanian), with a distinct acme the collections of the Naturhistorisches Museum Wien in the upper gracilis/mucronata Zone (latest Early and eleven specimens from the L. Schussler Collec Campanian). For the time being, it must remain in open tion (Leoben), which can be assigned to at least six nomenclature. species. These compare fairly well with elements of Early/Late Campanian echinoid faunas from northwest Echinocorys sp. 2 (cf. fonticola A rnaud 1897) Germany, northeast Belgium and Poland. (PI. 11, Fig. 3) Material and discussion:Three specimens in the Class Echinoidea NHMW collections (NHMW 1997/128/1; NHMW Order Holasteroida Durham & M elville 1957 1997/129/1 a,b; from Wentneralm I, Gams/ Hieflau Family Echinocoridae Lambert 1917 (Steiermark), preserved in reddish matrix, are crushed and heavily distorted. Despite this poor state of pres Echinocorys sp. 1 ervation these specimens can be seen to differ from (PI. 11, Figs. 1,2) Echinocorys sp. 1 (see above). The least ill-preserved Material: The Schussler collection includes a sin specimen appears to have been large (> 100 mm in gle internal mould, SCHU 71/33, preserved in reddish length), inflated, rounded and depressed and with a matrix, from Reitergraben which correlates with short apical system. This set of features characterises Wentneralm I, measuring 82.2 mm in overall length, the fonticola group, as interpreted by H ancock et al. 75.1 mm in width and 56.5 mm in height. (1993). Those authors recorded typical E. fonticola Discussion: To a certain extent, this specimen from the Early Campanian of southwest France, and a resembles a few tests in ,populations4 of E. gr. conoidea closely similar form from the earliest quadrata Zone from northeast Belgium. This group is here interpreted (Early Campanian) of southern England. on the basis of the type material: two specimens in the Because of the poor preservation of the Gams speci Goldfuss Collection at the Rheinische Friedrich- mens this identification is tentative at best. Wilhelms-Universitat Bonn (registration no.343a, b), one of them preserving the test, the other being an in Echinocorys gr. subglobosa (Goldfuss 1829) ternal flint mould. The test specimen clearly shows the (PI. 11, Fig. 5) type of preservation seen in populations4 from the 1903 Echinocorys subglobosus, Goldfuss , 1826; Lambert : 62, Lixhe 1 Member (Gulpen Formation) and is thus of pi. 2, figs. 7, 8. Late Maastrichtian age. Smiser (1935), on the other 1935 Echinocorys subglobosus Goldfuss (A nanchytes ); Smiser : hand, considered E. conoidea to be (mostly) of Cam 15, fig. 3. panian age (see also Kuchler & Kutz 1989). Material: The Schussler collection includes two There is also a superficial resemblance to some tests specimens, SCHU 71/6 [Reitergraben] and SCHU 71/ usually assigned to the pyramidata group (non pyrami- 38 [ForststraBe west Rabenmauer bzw. Rodlsteinsattel], both from greenish- matrix corresponding with Went dalis Smiser 1935), which typically is almost symmetri neralm II. cally conical and which appears to characterise the Discussion: The smaller specimen (SCHU 71/ late(st) Campanian and Early (earliest) Maastrichtian 38) measures 86.3 mm in overall length, is fairly well throughout NW Europe. Ernst (1972: pl.6, fig.4) re preserved, and closely resembles specimens from the corded a broadly similar, conical form from the Early older part of the late Campanian ( conica/mucronata Campanian (pilula Zone) of the Hannover area, but and basiplana/spiniger Zones) in northeast Belgium this differs from the Gams area specimen in being bet (see also Kuchler & Kutz 1989 for northern Spain). ter rounded. M aczynska ’s (1989) illustration of E. The other specimen is less well preserved; it measures pyramidata from the Polish Campanian appears closer more than 105 mm in overall length and compares well ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
170 Beitr. Paläont. 24, Wien 1999 with material from NW Germany (especially from the Discussion: The species is here interpreted on basiplana/spiniger Zone). Although depressed, this the basis of the type material in the Schlüter collec specimen is close to some forms assigned to E. ovata tion (no. 11a, lib ; Institut für Paläontologie, Rhei (Leske 1778) in the literature (e.g. Lambert 1903; Smi- nische Friedrich-Wilhelms-Universität Bonn), and of ser 1935), which H ancock et al. (1993) recorded from the literature (Ernst 1970b, 1972; Stokes 1975). In the Early Maastrichtian (? and latest Campanian). terestingly, the latter author (Stokes 1975: 71) men Ernst (1975) recorded the gibba/marginata group to tioned a unique specimen from the Gosau Formation range from the middle lingua/quadrata to the latest of Austria in the J. Lambert Collection (Paris) under conica/papillosa Zones (Early Campanian) in the Han the name of Micraster gosaviensis Lambert (a manu nover area, and subglobosa from the lower papillosa script name ?) which he thought, est vraisem (Early Campanian) into the vulgaris/basiplana (= blablement un M. glyphus .4 roemeri) Zones, with a distinct maximum in the stobaei/ Despite the fact that the present specimens are poorly basiplana (= basiplana/spiniger ) Zone (see also to moderately preserved, they all show the test fea Christensen et al., 1975; Schulz et al., 1984; Schön tures that characterise representatives of the Micras feld et al., 1996). ter schroederi/glyphus lineage. In size, ambital out Remarks: The Schüssler collection includes two line and features of the labrum they compare well with more tests, SCHÜ 71/59, labelled: Forststraße vom late Campanian »populations4, in particular from north Rödlstein zur Wentneralm, which equals Wentneralm ern Germany, and are here referred to M. glyphus. I and SCHÜ 71/9 from Reitergraben, in grey matrix In the late Cretaceous sections in the Hannover area equalling Wentneralm II, assignable to the genus Echi- and in the standard section for the NW German white nocorys. They are both poorly preserved, one speci chalk facies, the M. schroederi/glyphus group first ap men lacking more than half the test, the other being pears in the pilula Zone (Early Campanian) and ranges strongly laterally deformed, and cannot be identified at least into the upper basiplana/spiniger Zone, per to species/group. haps even higher into the late Campanian (Ernst , 1975; Schulz et al., 1984; Schönfeld et al., 1996). Order Spatangoida Claus 1876 Suborder Micrasterina Fischer 1966 There are comparable records of the schroederi/gly Family Micrasteridae Lambert 1920 phus group from the Liège-Limburg basin (Belgium, the Netherlands), where the first representatives oc Micraster glyphus Schlüter , 1869 cur at the base of the late Campanian Zeven Wegen (PI. 12, Figs. 1,3,5) Member (Gulpen Formation). Small- to medium-sized specimens, including forma planus M aczynska 1968 1869 Micraster glyphus Schlüter : 235, pi. 1, fig. 2. tokes 1970b Micraster (Micraster) glyphus Schlüter 1869; Ernst : (see also S , 1975), occur throughout this mem pi. 17, fig. 4. ber, which has been shown to be correlatable with the 1972 M ic ra ste r m.f. schroederi/glyphus; Ernst : pi. 5, fig. 3. NW German conica/mucronata, basiplana/ spiniger 1975 M icraster glyphus Schlüter 1869; Stokes:70, fig. 291 and roemeri Zones, and phaleratum Zone in ammo (with additional synonymy). nite terms (Kennedy & Jagt 1998). The group appears to range into the latest Campanian (,langeï Zone) specimen length width height Beutenaken Member (Gulpen Formation) of this area. S C H Ü 71/3 84.8 85.7 29.7 S C H Ü 71/37 93.0 85.3 41.2 Micraster gr. fastigatus/stolleyi S C H Ü 71/57 82.8 84.8 46.7 (PI. 11, Figs. 4, 6; PI. 12, Figs. 2, 4, 6)
Table 11. Dimensions of M ic ra ste r glyph us Schlüter from the 1970b Micraster (Gibbaster) gibbus (Lamarck 1816); Ernst : Late Campanian of Wentneralm II. Data in mm; all measure pi. 18, fig. 5. ments are approximate on account of varying degrees of com 1970b M icraster (Isomicraster) stolleyi L ambert 1901; paction and distortion). Ernst : pi. 18, figs. 6, 7. 1970c Micraster (Isomicraster) stolleyi Lambert ; E rnst : pi. 5, figs. 7, 8. Material: The Schüssler collection includes three 1972 Micraster (Gibbaster) gibbus (Lamarck ); Ernst : pi. specimens, SCHÜ 71/3 [Reitergraben], SCHÜ 71/37 2, fig 7; pi. 4, fig. 5. [,aus dem Anstehenden, oberhalb des Reitergraben 1972 Micraster (Isomicraster) stolleyi Lambert ; Ernst : pi. 2, fig. 8; pi. 4, fig. 6. Wasserfalles4], and SCHÜ 71/57 [westlich Rödelstein 1975 Micraster fastigatus Gauthier 1887; Stokes:69, fig. sattel (Forststraße Böschung)], all from the Wentner 29j, pl.4, figs. 7-9; pi. 5, figs 1, 2 (with additional alm II. synonymy). Dimensions (in mm; all measurements are ap 1975 M icraster stolleyi Lambert 1901 ; Stokes:79, fig. 30f proximate on account of varying degrees of compac (with additional synonymy). tion and distortion). Material: The Schüssler collection includes three ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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specimen length width height As the Wentneralm I specimens cannot be confidently S C H Ü 71/15 63.2 65.2 41.9 assigned to either M. fastigatus or M. stolleyi, they are S C H Ü 71/47 63.1 64.6 47.8 here, together with the Wentneralm II specimen, re S C H Ü 71/60 56.4 62.9 40.4 ferred to as M. gr. fastigatus/stolleyi. Christensen et al. (1975), who revised the definition Table 12. Dimensions of M icraster gr. fastigatus/stolleyi from of the important coleoid cephalopod taxon Belemnitella Wentneralm (I, II); in mm; all approximate. m. mucronata (von S chlotheim 1813) from the Late Campanian, listed M. stolleyi amongst the other macro specimens, SCHÜ 71/47 [zwischen Rödlstein Sattel fossil taxa from that part of the German, unteres Ober- und Wentner-Alm (aus dem Anstehenden), Forststraße campan'. It falls within the Hoplitoplacenticeras vari Böschung], SCHÜ 71/60 [zwischen Rödlsteinsattel und Zone, and the scaphitid Trachyscaphites spiniger, one Wentner-Alm (aus dem Anstehenden), Forststraße of the characteristic zonal species, allows trans-Atlan Böschung) and SCHÜ 71/15 [Reitnergraben]. The first tic correlations. two specimens are from Wentneralm I, the last-named In the Liege-Limburg basin (NE Belgium, SE Nether is from Wentneralm II. lands), M. stolleyi ranges from the basiplana/spiniger Discussion; In the literature it has been pointed to equivalents of the roemeri Zone (Jagt in prep.). out that the distinction between M. fastigatus and M. Amongst the ammonite species recorded from the late stolleyi can occasionally be difficult, which holds true Campanian Zeven Wegen Member (Gulpen Formation) for the present specimens as well. are Pachydiscus haldemsis (Schlüter 1867) and Stokes (1975) is followed here in the interpretation of Neancyloceras ? phaleratum (Griepenker I 1889) (Ken both species; he demonstrated that his Micraster nedy & Jagt 1998), documenting the phaleratum Zone. fastigatus was the same species as M. gibbus of Ger The Vaals Formation in northeast Belgium, which is man authors (Ernst 1970a-c, 1972, 1975). The true Early (but not earliest) Campanian in ammonite terms, Micraster gibbus is a Coniacian-Santonian species has yielded, amongst other species, Pachydiscus lau- de rossouvre (Stokes 1975, 1976, 1977; Fouray 1981), which ap nayi G , 1894. However, it has not yielded pears to be restricted to the Anglo-Paris province. Typi any representatives of the genus Micraster, but the cal M. fastigatus have an anal fasciole, which, how externally similar brissid genus Diplodetus (Stokes ever, is often diffuse or poorly developed, and differ 1979) is represented. from M. stolleyi by a more convex upper side and periproct situated at 35-55% of total test height. Typi Conclusions cal M. stolleyi have periproct situated at 25-45% of total test height, and the anal fasciole is generally miss For Wentneralm I an Early Campanian age is favoured on the basis of Echinocorys sp. 2 (cf. fonticola) and ing. Stokes (1975) remarked that, as evolved speci mens of M. fastigatus are difficult to distinguish from Micraster fastigatus, whereas E. gr. subglobosa, M. early representatives of M. stolleyi, the former is more glyphus and M. stolleyi suggest a middle Late Cam panian age for Wentneralm II. or less arbitrarily considered to be of early Campanian, and the latter of late Campanian age. Küchler & Kutz ’s WENTNERALM I (reddish matrix) (1989) record of, M. (Isomicraster) aff. stolleyi‘ from Echinocorys sp. 1 the latest Campanian of northern Spain appears to be Echinocorys sp. 2 (cf. fonticola A rnaud 1897) one of the youngest representatives of this group. Micraster gr. fastigatus/stolleyi The Gams area specimens are comparatively large in WENTNERALM II (greenish-grey matrix) comparison with »populations' from the Hannover area Echinocorys gr. subglobosa (Goldfuss , 1829) (Germany; Frerichs 1989), and in this respect are closer Micraster glyphus Schlüter , 1869 to Polish material (Maczynska 1968). Of the two speci Micraster gr. fastigatus/stolleyi mens from Wentneralm I, SCHÜ 71/60 is compara KRIMPENBACH (greenish-grey matrix) tively well preserved. In the absence of an anal fasciole, no echinoids and in having a markedly smaller anal angle, this speci men differs from most of the specimens of M. fastigatus Foraminifera from the Hannover area, and is closer to M. stolleyi Krimpenbach from the same area. On the other hand, the posterior no data concerning foraminifera portion of the labrum is narrow, as are the sternal plates, and these characters compare more favourably with Wentneralm I (red siltstones) M. fastigatus than with M. stolleyi. The Wentneralm II Data on foraminifera from the red siltstones stem from specimen (SCHÜ 71/15) shows a wide labrum and wide a few thinsections only. Sample GAM 230 from the sternal plates, lacks an anal fasciole and closely matches base of the Rödlstein section yielded the following the larger specimens in »populations' of M. stolleyi from planktonic marker species: the Hannover area. Globotruncana cf. area (Cushman ) ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
172 Beitr. Palaont. 24, Wien 1999
Globotruncana linneiana (d ’O rbigny ) patelliformis is relatively weakly defined, but normally Globotruncanita cf. elevata (B rotzen ) indicates assemblages clearly above the base of the Globotruncanita stuartiformis (D albiez ) Campanian (Robaszynski et al. 1984). The Bolivinoides Rositafornicata (P lummer ) lineage, including B. decoratus, documents a Cam R. fornicata and G. linneiana both first occur in the panian age (K och 1977). B. cf. draco miliaris even (Late) Santonian to Early Campanian, whereas the FO indicates late Late Campanian to Early Maastrichtian of G. elevata and G. stuartiformis marks a level near based on correlations with the Lagerdorf section in the base of the Campanian (Robaszynski et al. 1984; northern Germany (S ch Onfeld 1988: Polyplocum to G ale et al. 1995). This assemblage is indicative for Langei Zones), but only a single small specimen was the Elevata Zone of the Early Campanian in terms of found, which might represent an earlier morphotype planktonic foraminiferal zonations (e.g. Robaszynski of this species. et al. 1984; Caron 1985). A Late Santonian to early Several samples taken from K ollmann (1964) have Early Campanian age is very unlikely based on the been re-evaluated during this study. Samples 1027 and absence of Dicarinella concavata/asymetrica, although 1190 from grey marlstones yielded the marker species: it cannot be excluded in view of the limited number of Globotruncana area (C ushman ) thin sections studied. Globotruncana linneiana ( d ’O rbigny ) The integration of foraminiferal data of K ollmann Globotruncana linneiana obliqua H erm (1964) from a larger sample area around the Went- Globotruncanita cf. elevata (B rotzen ) neralm indicates a wider age range for red siltstones to Globotruncanita stuartiformis (D albiez ) marlstones. Samples 1026 and 1030 contained G. Rosita fornicata (P lummer ) elevata only and are consistent with the results above. Stensioeina pommerana B rotzen On the other hand, samples 1010 and 1024 yielded a These samples again indicate the Elevata Zone of the rich planktonic microfauna: Early Campanian in terms of planktonic foraminiferal Robaszynski Caron Globotruncana area (C ushman ) zonations (e.g. et al. 1984; 1985), Globotruncana ventricosa W hite where G. ventricosa marks the base of the Middle Cam Globotruncana linneiana (d ’O rbigny ) panian. Also the benthic foraminifer St. pommerana Globotruncanita cf. elevata (B rotzen ) starts in the latest Santonian and is typical of the Cam Globotruncanita subspinosa (P essagno ) panian. A Late Santonian to early Early Campanian Globotruncanita stuartiformis (D albiez ) age can be excluded by the absence of Dicarinella Rugoglobigerina cf. rotunda (B ronnimann ) concavata/asymetrica. Ventilabrella glabrata Cushman The other samples (1049, 1065, 1189) appear to indi cate a higher level because of the presence of G. ven This assemblage with G. ventricosa, G. subspinosa and tricosa and B. draco miliaris together with the above- R. cf. rotunda can be assigned to the upper part of the mentioned foraminifera. The FO of G. ventricosa de Ventricosa Zone of the „Middle Campanian“, below fines the base of the Ventricosa- Zone (e.g. Caron , the Calcarata Zone (e.g. Robaszynski et al. 1984; 1985), for which a „Middle Campanian“ age may be Caron 1985). These samples therefore document the assumed following the suggestions of the Subcommiss- presence of a considerably younger red siltstone facies sion on Cretaceous Stratigraphy (L amolda , 1995). than that documented from locality Wentneralm I. Based on foraminiferal assemblages the total possible Wentneralm II (grey siltstones) stratigraphical range of the grey siltstones to silty marls is late Early Campanian to Early Maastrichtian. One sample (GAM 183) from the grey interval, taken near the inoceramid locality Wentneralm I, yielded a Calcareous Nannofossils relatively rich microfauna, including important plank Krimpenbach tonic and benthic marker species: Only one of four nannofossil samples from the Krim Archaeoglobigerina blowi (P essagno ) penbach locality contained calcareous nannofossils. Globotruncana linneiana (d ’O rbigny ) Although the assemblage is poor and only moderately Globotruncanita stuartiformis (D albiez ) preserved, it can be dated according to standard Rosita fornicata (P lummer ) nannofossil zonations for the Late Cretaceous (impor Rosita patelliformis (G andolfi ) tant species marked with *). Bolivinoides decoratus (Jones ) NHM KRIMP 1997/144/3 Bolivinoides cf. draco miliaris H iltermann & K och Arkhangelskiella sp. (small forms) R. fornicata and G. linneiana both first occur in the * Calculites obscurus (D eflandre 1959) P rins & (Late) Santonian to Early Campanian, whereas the FO S issingh 1977 of G. stuartiformis marks a level near the base of the Calculites ovalis (S tradner 1963) Prins & S issingh Campanian (Robaszynski et al. 1984). The FO of R. 1977 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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Chiastozygus litterarius (G orka 1957) M anivit 1971 Prediscosphaera cretacea (A rkhangelsky 1912) Cretarhabdus crenulatus B ramlette & M artini 1964 G artner 1968 Cribrosphaerella ehrenbergii (A rkhangelsky 1912) Watznaueria barnesae (B lack 1959) P erch -N iel D eflandre 1952 sen 1968 Cylindralithus sp. The biostratigraphic interpretation of this poor Eprolithus sp. nannofossil assemblage indicates a Campanian age. In Eiffellithus eximius (S tover 1966) P erch -N ielsen terms of the standard nannofossil zonation (S issingh 1968 1977; Perch -N ielsen 1985) CC17 (FO of C. obscurus, Eiffellithus turriseiffelii (D eflandre & F ert 1954) Late Santonian) up to CC22b (LO of E. eximius, Late Reinhardt 1965 Campanian) are possible. M. cf. praemurus is com Glaukolithus diplogrammus (D eflandre 1954) mon from CC21 onwards in the Nierental Formation Reinhardt 1964 of the „Gams basin“ (W agreich & K renmayr 1993), Lucianorhabdus cayeuxii D eflandre 1959 but only one poorly preserved specimen was found. * Lucianorhabdus cayeuxii D eflandre 1959 (ssp. B) Lucianorhabdus maleformis Reinhardt 1966 Wentneralm II (grey siltstones) Micula decussata V ekshina 1959 Nannofossils in samples from the grey siltstones are Prediscosphaera cretacea (A rkhangelsky 1912) not abundant, but assemblages were rich enough for G artner 1968 biostratigraphic interpretations. Nannofossil assem Reinhardtites sp. blages from two representative samples are given here: Tranolithus orionatus (Reinhardt 1966) Perch -N iel Sample GAM 183 was taken from a marly layer in the sen 1968 lower outcrop area below the Rodlstein (contained also Watznaueria barnesae (B lack 1959) P erch -N ielsen a significant microfauna, see above) and sample GAM 1968 123 (important species marked with *): The biostratigraphic interpretation of this sample indi cates a Late Santonian to early Early Campanian age. GAM 183
The marker species of nannofossil standard zone CC * Arkhangelskiella cymbiformis V ekshina 1959 17 (S issingh 1977; P erch -N ielsen 1985), C. obscurus Biscutum constans (G orka 1957) B lack 1959 is present together with curved Lucianorhabdus Biscutum sp. cayeuxii, defining subzone CC 17b (W agreich 1992). * Broinsonia parca constricta H attner , W ind & W ise Additional indicators of a Santonian - Early Campanian 1980 age are Eprolithus sp., Lucianorhabdus maleformis, and * Broinsonia parca parca (S tradner 1963) B ukry Reinhardtites sp., which are very rare or absent in Late 1969 Campanian samples from the Gosau Group (W agreich * Calculites obscurus (D eflandre 1959) P rins & 1988,1992;W agreich & K renmayr 1993). According S issingh 1977 to the zonations by B urnett (1998) the sample has a Chiastozygus litterarius (G orka 1957) M anivit 1971 wider age range from U C llc to UC12 (Late Coniac- Cretarhabdus crenulatus B ramlette & M artini 1964 ian to early Early Campanian). Cribrosphaerella ehrenbergii (A rkhangelsky 1912) D eflandre 1952 Wentneralm I (red siltstones) * Eiffellithus eximius (S tover 1966) P erch -N ielsen Nannofossil samples taken from the red siltstones and 1968 samples taken from ammonite individuals of fauna Eiffellithus turriseiffelii (D eflandre & F ert 1954) Wentneralm I yielded very poor results. Only a single Reinhardt 1965 sample taken from Pachydiscus (P.) sp. indet. juv. Gartnerago obliquum (S tradner 1963) N oel 1970 (SCHU 71/52, pi. 4, fig. 2) contained a significant Helicolithus trabeculatus (G orka 1957) V erbeek nannofossil assemblage. 1977 Glaukolithus diplogrammus (D efla n d r e 1954) Calculites obscurus (D eflandre 1959) P rins & Reinhardt 1964 S issingh 1977 Glaukolithus spiralis B ramlette & M artini 1964 Calculites ovalis (S tradner 1963) P rins & S issingh 1977 Lucianorhabdus cayeuxii D eflandre 1959 (A) Lithraphidites carniolensis carniolensis D eflandre Cretarhabdus crenulatus B ramlette & M artini 1964 1963 Manivitella pemmatoidea (D eflandre in M anivit Eiffellithus eximius (S tover 1966) P erch -N ielsen 1968 1965) T hierstein 1971 Lucianorhabdus cayeuxii D eflandre 1959 Micula decussata V ekshina 1959 Micula decussata V ekshina 1959 Ottavianus giannus RISATTI 1973 Micula cf. praemurus (B ukry 1973) Stradner & Prediscosphaera cretacea (A rkhangelsky 1912) S teinmetz 1984 G artner 1968 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
174 Beitr. Paläont. 24, Wien 1999
* Quadrum aff. gartneri Prins & P erch -N ielsen in Sample GAM 123 additionally contains the marker M anivit et al. 1977 (small morphotype) species C. aculeus and therefore indicates nannofossil Russellia multiplus (Perch -N ielsen 1973) W ind & zone CC20. Zone CC21 cannot be excluded, although W ise 1977 the marker species for the base of CC21, Quadrum Tranolithus phacelosus Stover 1966 sissinghi is not present. The abundance of nannofos- Vekshinella stradneri Rood et al. 1971 sils in the samples is low, therefore this rare species Watznaueria bamesae (B lack 1959) Perch -N ielsen may have been overlooked. 1968 Zeugrhabdotus embergeri (N oel 1959) Perch -N iel sen 1984 Biostratigraphical conclusions GAM 123 Late Santonian/Early Campanian age of the Krimpen- * Arkhangelskiella cymbiformis V ekshina 1959 bach Formation at the Krimpenbach locality, dated by Bipodorhabdus cf. brooksi (B ukry 1969) Crux 1982 nannofossils and inoceramids, leads to the conclusion, Biscutum constans (G orka 1957) B lack 1959 that the base of the Krimpenbach Formation must be Braarudosphaera bigelowi (G ran & B raarud 1935) D eflandre 1959 older than Late Santonian. Microfaunal and nannofossil * Broinsonia parca constricta H attner , W ind & W ise data for the red sandstones to siltstones of locality 1980 Wentneralm I are scarce. Rare planktonic foraminif- * Broinsonia parca parca (S tradner 1963) B ukry era indicate the Elevata Zone of the Early Campanian 1969 (Robaszynski et al. 1984; G ale et al. 1995). The poor Calculites obscuras (D eflandre 1959) P rins & Sis- nannofossil assemblage is in accordance with these SINGH 1977 data. Based on data from the overlying grey siltstones, * Ceratolithoides aculeus (S tradner 1961) Prins & the red siltstones of locality Wentneralm I most prob S issingh in Sissingh 1977 Cretarhabdus crenulatus B ramlette & M artini 1964 * Eiffellithus eximius (S tover 1966) Perch -N ielsen MAASTRICHTIAN W entneralm I W entneralm II 1968 (D eflandre Fert 1954) Eiffellithus turriseiffelii & H yalti Zone Reinhardt 1965 Glaukolithus diplogrammus (D eflandre 1954) Reinhardt 1964 D onezianum Zone Glaukolithus spiralis B ramlette & M artini 1964 Helicolithus trabeculatus (G orka 1957) V erbeek Polyplocum Zone 1977 LATE CAMPANIAN
Lucianorhabdus cayeuxii D eflandre 1959 CAMPANIAN Phaleratum Zone Lucianorhabdus cayeuxii D eflandre 1959 (ssp. B - curved morphotype) Q uadrata Zone Lithraphidites carniolensis camiolensis D eflandre I ■ S ■ 1963 < u * Lucianorhabdus maleformis Reinhardt 1966 Granulaia Zone si LATE < Micula decussata V ekshina 1959 UJ SANTONIAN Prediscosphaera cretácea (A rkhangelsky 1912) Gartner 1968 Table 13. Biostratigraphical range of the fossil sites Wentneralm Quadrum aff. gartneri Prins & Perch -N ielsen in I and Wentneralm II in terms of ammonite stratigraphy (with M anivit et al. 1977 (small morphotype) alterations after B laszkiewicz 1980). Quadrum cf. gothicum (D eflandre 1959) Prins & Perch -N ielsen in M anivit et al. 1977 ably represent a level within the interval from nanno Rucinolithus sp. (8 rays) fossil zones CC17 to CC18/19a. Vekshinella stradneri Rood et al. 1971 An important indication for the minimum age of the Watznaueria bamesae (B lack 1959) Perch -N ielsen grey siltstones and thus for the locality Wentneralm II 1968 is given by sample GAM 222 from overlying soft Zeugrhabdotus embergeri (N oel 1959) Perch -N iel pelagic to turbiditic marls in the Krimpenbach section sen 1984 (Text-fig. 2). The nannofossil assemblage documents Sample GAM 183 may be assigned to nannofossil zone nannofossil zone CC22a/b, portions of which can be CC19, probably zone CC19b based on Perch -N ielsen ’s correlated to the Calcarata Zone of planktonic forami- zonation (1985). The presence of B. parca parca , B. niferal zonations (e.g. W agreich & K renmayr 1993), parca constricta and large A. cymbiformis (10 (|l) and which indicates a level in the Late Campanian (S ch On - the absence of Marthasterites furcatus (LO on top of feld & B urnett 1991). AMaastrichtian age can there zone CC19a) and C. aculeus (FO base of zone CC20) fore be excluded for the grey siltstones. Based on our indicates an Early Campanian age (e.g. Sissingh 1977). data, the grey siltstones of the Wentneralm area com- ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
S ummesberger , H., et al., Integrated biostratigraphy 175 Biostratigraphical range of the sites Krimpenbach, Wentneralm I and Wentneralm II. Inoceramids. T ab le 14. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
176 Beitr. Paläont. 24, Wien 1999
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L ommerzheim , A.J., 1995. Stratigraphie und Ammoniten P etrascheck , W, 1906. Über Inoceramen aus der Gosau faunen des Santons und Campans im Münsterländer und dem Flysch der Nordalpen.— Jahrb. k.k. geol. Becken (NW-Deutschland). — Geol. Paläont. West Reichsanst., 56:155-168, 1 pl., 4 figs., Wien. falen, 40:97 pp., 19 text-figs., 8 pis., Münster. Redtenbacher , A., 1873. Die Cephalopodenfauna der Gosauschichten in den nordöstlichen Alpen. — Abh. L upu , D. & S ornay , J., 1978. Noi date biostratigrafice asupra — Senonianului din regiunea Vidra (Muntii k.k. geol. Reichsanst., 5:91-140, pis. 22-30, Wien. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
S ummesberger , H., et al., Integrated biostratigraphy 179
Riedel , L., 1931. Zur Stratigraphie und Faciesbildung im of NW Germany (Lägerdorf Kronsmoor-Hemmoor): Oberemscher und Untersenon am Südrande des Bek- Definitions and proposals. — Bull. Geol. Soc. Den kens von Münster. — Jahrb. Preuß. Geol. Landes- mark, 33:203-215, Copenhagen. anst., 51:605-713, pls. 72-79, 6 text-figs., Berlin. S chulz , M.- G. & S chmid , E, 1983. Die Belemniten der Robaszynski , E , Caron , M., G onzales D onoso , J.M. & Inoceramen-Mergel (Buntmergelserie, Ultrahelve- W onders , A.A.H. (Eds.), 1984. Atlas of Late Cre tikum, Unt.-Maastricht) des Moos-Grabens SE taceous Globotruncanids. — Revue Micropaleont., Siegsdorf (Oberbayern) und ihre stratigraphische 26:145-305, Paris. Bedeutung. — Zitteliana 10:653-661, 7 text-figs., S antamaria Z aba l a , R., 1996. Los Ammonites del München.
Campaniense de la Provincia de Alava. Sistematica S eitz, O., 1961. Die Inoceramen des Santons von e bioestratigrafia. — Est. Mus. Cienc. Nat de Alava Nordwestdeutschland. I.Teil:(Die Untergattungen (1995-1996), 10-11:5-25, Alava. Platyceramus, Cladoceramus und Cordiceramus ). — Schlüter , C , 1869. Fossile Echinodermen des nördlichen Beih. Geol. Jahrb., H. 46:186 pp., 15 pls., 39 text- Deutschlands. — Verh. naturhist. Ver. preuß. Rhein figs., Hannover. lands und Westfalens, 26: 225-253, pls. 1-3, Bonn. S eitz, O., 1965. Die Inoceramen des Santon und Unter- S chlüter , C., 1972-1876. Cephalopoden der oberen Campan von Nordwestdeutschland. n.TeikBiometrie, deutschen Kreide. — Palaeontographica 24:1-144, Dimorphismus und Stratigraphie der Untergattung pls. 36-55, Cassel. Sphenoceramus. — Beih. Geol. Jahrb., 69: 194 pp., S chönfeld , J., 1988. Zur Stratigraphie und Ökologie 26 pls., 11 text-figs., Hannover.
benthischer Foraminiferen im Schreibkreide-Richt- S eitz, O., 1967. Die Inoceramen des Santon und Unter- profil von Lägerdorf/Holstein. — Thesis Mathem.- Campan von Nordwestdeutschland. III.Teil: Taxo Naturwiss. Fak. Univ. Kiel, 1-93, 21 figs., 7 tabl., 6 nomie und Stratigraphie der Untergattungen Endo- pls., Kiel. costea, Haenleinia, Platyceramus, Cladoceramus, Schönfeld , J. & B urnett , J., 1991. Biostratigraphical Selenoceramus und Cordiceramus mit besonderer correlation of the Campanian-Maastrichtian bound Berücksichtigung des Parasitismus bei diesen Unter ary: Lägersdorf-Hemmoor (northwestern Germany), gattungen. — Beih. Geol. Jahrb., H. 75:171, 27 text- DSDP Sites 548A, 549, and 551 (eastern North At figs., 8 tab., 27 pis., Hannover. lantic) with paleobiogeographical and paleoceano- S iegl-Farkas , A. & W agreich , M., 1997. Correlation of graphical implications. — Geol. Mag., 128:479-503, palyno- (spores, pollen, dinoflagellates) and calc 11 figs., 2 tabl., London. areous nannofossil zones in the Late Cretaceous of S chönfeld , J., S chulz , M.-G., M c A rthur , J.M., B urnett , the Northern Calcareous Alps (Austria) and Trans- J., G ale , A ., H ambach , U., H ansen , H.J., Kennedy , danubian Central Range (Hungary). — Jubiläums W.J., Rasmussen , K.L., Thirlwall , M.F. & W ray , schrift 20 Jahre Zusammenarbeit Österreich-Ungarn, D.S., 1996. New results on biostratigraphy, palaeo- 3, Wien. magnetism, geochemistry and correlation from the standard section for the Upper Cretaceous white chalk S issingh , W., 1977. Biostratigraphy of Cretaceous calc of northern Germany (Lägerdorf-Kronsmoor-Hem- areous nannoplankton. — Geol. Mijnbouw, 56: 37- moor). [in:] S paeth , C. (ed.). Jost W iedmann Memo 56, Amsterdam. rial Volume. New developments in Cretaceous re S miser , J.S., 1935. A revision of the echinoid genus search topics. Proceedings of the 4th International Echinocorys in the Senonian of Belgium. — Mém. Cretaceous Symposium, Hamburg 1992. — Mitt. Mus. royal d’Hist. nat. Belgique, 67:1-52, pls. 1-3. Geol.-Paläont. Inst. Univ. Hamburg, 77:545-575, S ornay , J., 1957. Inoceramus impressus d ’O rbigny . — Hamburg. Paleontología universalis, N.S. 129, Paris. Schultz , O. & P aunoviö , M., 1997. Der Nachweis von Sornay , J., 1962. Etude d’une faune d’inocerames du Coelodus (Osteichthyes, Pycnodontidae) imTuronien Senonien supérieur des Charentes et description d’une (Oberkreide) von Gams, Steiermark, Österreich und espece nouvelle du Senonien de Madagascar. — Bull, aus der Oberkreide von Kroatien und Italien with a soc. géol. France, 4:118-122, Paris. contribution of H. S ummesberger on the stratigraphy of the Gams basin. — Ann. Naturhist. Mus. Wien, 98 S ornay , J., 1968. Inocerames Senonien du Sud-Ouest de A:73-14, Wien. Madagascar. — Ann. Paléont. (Invertébrés), 54:25- 47, pis. A-H, Paris. S chulz , M.-G., 1979. Morphometrisch-variationsstati stische Untersuchungen zur Phylogenie der Belem- S ornay , J., 1975. Trois especes nouvelles d’inocerames niten-Gattung Belemnella im Unter-Maastricht du Senonien de Madagascar. — Ann. de Paléont. Westeuropas. — Geol. Jahrb., Reihe A, 47:160 pp., (Invertebres) 61, fase. 1:19-29, 6 pis., Paris.
66 text-figs., 12 pls., Hannover. S ornay , J., 1976. La Faune d’inocerames de Dau (Region S chulz , M.-G., E rnst , G., E rnst , H. & S chmid , E , 1984. de Royan, Charente Maritime) et Remarques sur deux Coniacian to Maastrichtian stage boundaries in the Especes de d ’O rbigny : I. regularis et /. Goldfussi .— Standard section for the Upper Cretaceous white chalk Ann. de Paléont. 62, fase. 1:1-18, Paris. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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du Campanien et Maastrichtien de la Belgique. Bull. Tzankov , V, 1982. Les Fossiles de Bulgarie. Va. Crétacé Inst. roy. Sei. nat. de Belgique, Sci.Terre 54/4:1-15, Supérieur, Cephalopoda, (Nautiloidea, Ammonoidea) 3 text-figs., 4 pis, Bruxelles. et Echinodermata (Echinoidea), 136 pp., 50 pis., Stokes, R.B., 1975. Royaumes et provinces fauniques du Sofia.
Crétacé établis sur la base d’une étude systématique U lbrich , H,, 1971. Mitteilungen zur Biostratigraphie des du genre Micraster. — Mém. Mus. national d’Hist. Santon und Campan des mittleren Teils der Sub- nat, C (Sei. de la Terre), 31:1-94, pis. 1-12, Paris. herzynen Kreidemulde. — Freiberger Forschungsh. S tokes, R.B., 1976. Distinction between sympatric C 267:47-60, 5 pis., Leipzig.
species of Micraster (Echinoidea) from the English V oigt, S.,1996. Paläobiogeographie oberkretazischer Chalk. Palaeontology, 19(4):689-697, London. Inoceramen und Rudisten - Ozeanographische und Stokes, R.B., 1977. The echinoids Micraster and Epiaster klimatologische Konsequenzen einer neuen Paläo- from the Turonian and Senonian of England. Palae geographie. — Münchner Geowissenschaftliche ontology, 20 (4):805-821, pis. 106-109, London. Abhandlungen, Reihe A (Geologie und Paläon
Stokes, R.B., 1979. The echinoid genus Diplodetus from tologie), 31:102 pp., 53 text-figs, München. the Santonian to Danian of Northern Europe. — N. W agreich , M., 1988. Nannoplankton- und Foraminiferen Jahrb. Geol. Paläont., Mh„ 1979 (10):619-630, - Feinstratigraphie des Santon-Untercampans der Stuttgart. Gosauschichtgruppe von Gosau-Rußbach (Ober-
S toliczka , F., 1865. The fossil Cephalopoda of the österr.-Salzburg). — Mitt. Ges. Geol. Bergbaustud. Cretaceous Rocks of Southern India. A mmonitidae , Österr., 34/35:279-294, 3 figs., Wien. with revision of Nautilidae, & c. — Mem. geol. Surv. W agreich , M., 1992. Correlation of Late Cretaceous India (1), Palaeont. Indica, 3/1:1-56, pis.26-31 calcareous nannofossil zones with ammonite zones (1863); 2-5:57,106, pis. 32-54 (1864); 6-9:107-154, and planktonic foraminifera: the Austrian Gosau pis. 55-80(1865); 10-13:155-216, pis. 81-94(1866), sections. — Cretaceous Research, 13:505-516, 5 Calcutta. figs., London.
S ummesberger , H., 1979. Eine obersantone Ammo- W agreich , M., 1993. Bericht 1992 über geologische nitenfauna aus dem Becken von Gosau (Ober Aufnahmen in Oberkreide- und Alttertiärablage österreich). — Ann. Naturhist. Mus. Wien, 82:109- rungen auf den Blättern 99 Rottenmann, 100 Hieflau 176, 15 pis, Wien. und 101 Eisenerz. — Jahrb. Geol. Bundesanst., 136:
S ummesberger , H., 1997. The age of the Gosautrans- 586-587, Wien. gression in the Gams basin. — [in:] S chultz , O. & W agreich , M., 1994. Bericht 1993 über geologische PAUNO Vid, M. (1997). Der Nachweis von Coelodus Aufnahmen in Oberkreide- und Tertiärsedimenten der (Osteichthyes, Pycnodontidae) im Turonien (Ober Nördlichen Kalkalpen auf den Blättern 100 Hieflau kreide) von Gams, Steiermark, Österreich und aus und 101 Eisenerz. — Jahrb. Geol. Bundesanst., der Oberkreide von Kroatien und Italien. — Ann. 137:477-478, Wien. Naturhist. Mus. Wien, 98 A:73-141, Wien. W agreich , M., 1995. Bericht 1994 über geologische S ummesberger , H. & Kennedy , W.J., 1996. Turonian Aufnahmen in Oberkreide- und Tertiärsedimenten der Ammonites from the Gosau Group (Upper Creta Nördlichen Kalkalpen auf Blatt 101 Eisenerz. — ceous; Northern Calcareous Alps; Austria) with a Jahrb. geol. Bundesanst., 138:503, Wien. revision of Barroisiceras haberfellneri (H auer 1866). — Beitr. Paläont. Österr., 21:1-75, 23 text-figs., 18 W agreich , M., B öhm, F. & L obitzer , H., 1996. Sedi- pis., 3 tabl., Wien. mentologie des kalkalpinen Mesozoikums in Salz burg und Oberösterreich (Jura, Kreide). — Berichte S ummesberger , H., S anders , D. & K ollmann , H.A. (in der geol. Bundesanst., 33/B 1:1-57, Wien. press). Deciphering the Gosau Group - a Challenge for Cretaceous Palaeontology, [in:] N eubauer , F. W agreich , M. & Krenmayr , H.-G., 1993. Nannofossil Geology of Austria (Springer). biostratigraphy of the Late Cretaceous Nierental Formation, Northern Calcareous Alps (Bavaria, Tröger , K.-A. & Röhlich , R, 1982. Zur Variabilität von Inoceramus balticus haldemensis Giers .- Freiberg. Austria). — Zitteliana, 20:67-77, München. Forschungsh. C 375:101-111, 9 figs., 1 pi., Leipzig. W alaszczyk , I., 1997. Bioastratigraphie und Inoceramen
T röger , K.-A. & Röhlich , P., 1992. Campanian- des oberen Unter-Campan und unteren Ober-Campan Maastrichtian Inoceramid (Bivalvia) Assemblages Norddeutschlands. — Geol. u. Paläont. in Westfalen, from NW-Libya. [in:] The Geology of Libya. Vol. 49:111 pp., 21 text-figs., 32 pis., Münster. V:1357-1381, 19 text-figs., 5 pis., Tripoli. W egner , T, 1905. Die Granulatenkreide des westlichen Tröger , K.-A., S ummesberger , H. & S koumal , P. 1999. Münsterlandes. — Zs. dt. geol. Ges., 57:112-232, pis. Inoceramidae from the Campanian (Upper Creta 7-10, 20 text-figs., Berlin. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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WiPPiCH, M.G.E., 1995. Ammoniten aus dem Unter- W right , C.W. with Callomon , J.H. & H owarth , M.K., campan des westlichen Münsterlandes. — Geol. 1996. Cretaceous Ammonoidea. [in:] Kaesler , R.L. Paläont. Westfalen, 38:43-87, 11 pls., 8 text-figs., (ed.), Treatise on Invertebrate Paleontology. Part L Münster. — Mollusca 4; Revised, xx + 362 pp., Boulder and
W oods , H., 1899-1913. A Monograph of the Cretaceous Lawrence. Lamellibranchia of England. — Monogr. Paleontogr. W right , C.W. & Smith, A.B., 1987. Echinoderms, 201- Soc. Vol.I:232 pp., 42 pis., 7 text-figs. Vol.II:473 pp., 237, pis. 44-53. [in:] Owen , E. (comp.) & Smith, A.B. 62 pis., 252 text-figs., London. (ed.). Fossils of the Chalk [Field Guides to Fossils
W right , C.W., 1957. Cretaceous Ammonoidea [in:] 2], London (Palaeont.Ass.). M oore , R.C. (ed.), Treatise on Invertebrate Paleon tology. Part L, — Mollusca 4: xxii + 490 pp., New York and Lawrence. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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PLATE 1
Fig. 1: Hauericeras cf. pseudogardeni (S chlüter ); SCHÜ 71/58; Lower Campanian, Wentneralm I, Gams/Hieflau, Steiermark; x 0,5. Figs. 2, 3: Desmophyllites sp. indet. juv., SCHÜ 71/52 b; Lower Campanian, Wentneralm I, Gams/Hieflau, Steiermark; x 2. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
Summesberger , H., et al., Integrated biostratigraphy 183
PLATE 1 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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PLATE 2
Figs. 1, 3: Pachydiscus (Pachydiscus) haldemsis (Schlüter ); SCHÜ 71/53; Upper Campanian, Wentneralm II, Gams/ Hieflau, Steiermark; x 1.
Fig. 2: Pachydiscus (Pachydiscus) tweenianus (Stoliczka ); SCHÜ 71/54; Upper Campanian, Wentneralm II, Gams/Hieflau, Steiermark; x 0,5. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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PLATE 2 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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PLATE 3
Fig. 1: Pachydiscus (Pachydiscus) tweenianus (S toliczka ); SCHU 71/20; Upper Campanian, Wentneralm II, Gams/ Hieflau, Steiermark; x 0,5.
Fig. 2: Pachydiscus (Pachydiscus) cf. launayi de G rossouvre ; NHMW1997/127/1; Lower Campanian, Wentneralm I, Gams/Hieflau, Steiermark; x 1. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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PLATE 3 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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PLATE 4
Fig. 1: Pachydiscus (Pachydiscus) cf. launayi de Grossouvre ; SCHÜ 71/56; Lower Campanian, Wentneralm I, Gams/Hieflau, Steiermark; x 0,5. Fig. 2: Pachydiscus (Pachydiscus) spec, indet. juv.; SCHÜ 71/52 a; Lower Campanian, Wentneralm I, Gams/Hieflau, Steiermark; x 1. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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PLATE 4 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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PLATE 5
Fig. 1: Pachydiscus (Pachydiscus) cf. launayi de G rossouvre ; SCHÜ 71/46; Lower Campanian, Wentneralm I, Gams/Hieflau, Steiermark; x 1. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
S ummesberger , H., et al., Integrated biostratigraphy 191
PLATE 5 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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PLATE 6
Fig. 1: Cordiceramus muelleri cf. germanicus (Heinz ) (NHMW 1997/z/144/1)
Fig. 2: Cordiceramus muelleri cf. germanicus (Heinz ) (NHMW 1997/z/144/14)
Fig. 3: Cordiceramus muelleri cf. germanicus (H einz ) (NHMW 1997/z/144/6)
All specimens are from the Krimpenbach site near Wildalpen, Styria, Gams area; all are uppermost Santonian-lowest Campanian. All figures are x 1. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
S ummesberger , H., et al., Integrated biostratigraphy ...
PLATE 6 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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PLATE 7
Fig. 1: Cataceramus ex gr. balticus (B öhm) (NHMW 1997/z/144/5)
Fig. 2: Selenoceramus cf. inßexus (B eyenburg ) (NHMW 1997/z/144/9)
Fig. 3: Cataceramus ex gr, balticus (Böhm) (NHMW 1997/z/144/3)
Fig. 4: Selenoceramus cf. inflexus (Beyenburg ) (NHMW 1997/z/144/4) All specimens are from the Krimpenbach site near Wildalpen, Styria, Gams area; all are uppermost Santonian - lowest Campanian. All figures are x 1. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
S ummesberger , H., et al., Integrated biostratigraphy 195
PLATE 7 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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PLATE 8
Fig. 1: Endocostea aff. impressa (d ’Orbigny ) (NHMW 1997/z/158/9)
Fig. 2: Endocostea aff. impressa (d ’Orbigny ) (NHMW 1997/z/158/ll)
Fig. 3: Selenoceramus inflexus (B eyenburg ) (NHMW 1997/z/158/10)
Fig. 4: Selenoceramus inflexus (B eyenburg ) (NHMW 1997/z/158/6)
Fig. 5: Sphenoceramus aff. angustus (B eyenburg ) (NHMW 1997/z/158/13)
F ig . 6: Cataceramus balticus (B ö h m ) subsp.indet. (NHMW 1997/z/158/12) All specimens are from the Wentneralm I site near Gams, Gams area, Styria; Krimpenbach Formation, all are Lower Campanian. All figures are x 1. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
S ummesberger , H., et al., Integrated biostratigraphy 197
PLATE 8 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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PLATE 9
Fig. 1: Inoceramus planus M unster (NHMW 1997/z/159/1)
Fig. 2: Cataceramus balticus cf. haldemensis (Geers ) with geniculation, deformed by compaction. (NHMW 1997/ z! 159/4)
Fig. 3: Inoceramus cf. bosenbergensis W alaszczyk (NHMW 1997/z/159/7)
Fig. 4: Cataceramus balticus cf. haldemensis (Giers ) (NHMW 1997/z/159/8)
Fig. 5a, b: Cataceramus balticus cf. haldemensis (Giers ) (NHMW 1997/z/159/6) All specimens are from the Wentneralm II site near Gams, Gams area, Styria; Krimpenbach Formation, all are lower Upper Campanian. All figures are x 1. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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PLATE 9 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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PLATE 10
Fig. 1: Cataceramus balticus cf. haldemensis (G iers ) with geniculation(NHMW 1997/z/159/9)
Fig. 2: Selenoceramus inflexus (Beyenburg ) with Endocostea scar (NHMW 1997/z/158/7)
Fig. 3: Selenoceramus inflexus (B eyenburg ) with geniculation, highly deformed by compaction (NHMW 1997/z/ 158/5)
Fig. 4: Inoceramus aff. borilensis JolkiCev (NHMW 1997/z/159/13)
All specimens are from the Krimpenbach Formation, figs. 1,4 are from Wentneralm II site; figs. 2,3 are from Wentneralm I site; figs. 1,4 are lower Upper Campanian, figs. 2, 3 are Lower Campanian. All figures are x 1. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
S ummesberger , H., et al., Integrated biostratigraphy 201
PLATE 10 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
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PLATE 11
Fig. 1, 2: Echinocorys sp. 1; Wentneralm I, Lower Campanian. Reitergraben; SCHÜ 71/33.
Fig. 3: Echinocorys sp. 2 (cf .fonticola A rnaud , 1897); forest road, Wentneralm I; NHMW 1997/128/1. Fig. 4, 6: Micraster gr.fastigatus/stolleyv, Wentneralm II; Upper Campanian; Reitergraben; SCHÜ 71/15.
Fig. 5: Echinocorys gr. subglobosa (Goldfuss ); Wentneralm II, Upper Campanian; Reitergraben; SCHÜ 71/38.
All specimens are from Gams near Hieflau from the area around Wentneralm. All figures are x 1. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
S ummesberger , H., et al., Integrated biostratigraphy 203
PLATE 11 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
204 Beitr. Paläont. 24, Wien 1999
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Fig. 1: Micraster glyphus Schlüter ; Wentneralm II, Upper Campanian; Reitergraben; SCHÜ 71/37. Fig. 2, 4, 6: Micraster gr. fastigatus/stolleyv, Wentneralm I; Lower Campanian, Rödlsteinsattel; SCHÜ 71/60.
Fig. 3, 5: Micraster glyphus Schlüter ; Wentneralm II, Upper Campanian; W of Rödlsteinsattel; SCHÜ 71/57.
All specimens are from Gams near Hieflau from the area around Wentneralm. All figures are x 1. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien
S ummesberger , H., et al., Integrated biostratigraphy 205
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