Annals of Agrarian Science 15 (2017) 252e268

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Annals of Agrarian Science

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Updated checklist of scale insects (Hemiptera: Coccomorpha)in Sakartvelo ()

* M. Batsankalashvili a, M.B. Kaydan b, c, G. Kirkitadze a, G. Japoshvili a, a Institute of Entomology, Agricultural University of Georgia, 240, Agmashenebeli Alley, , 0159, Georgia b Imamoglu Vocational School, Cukurova University, Adana, 01330, Turkey c Biotechnology Research and Application Center, Çukurova University, Adana, 01330, Turkey article info abstract

Article history: The Scale Insects, infraorder Coccomorpha (Hemiptera: Sternorryncha) includes many agricultural pests. Received 22 December 2016 The latest check list of scale insects of Sakartvelo (Republic of Georgia) was published in 2008, but it Accepted 12 February 2017 contained species mostly from the city of Tbilisi. In this study an updated check list of scale insects of Available online 9 May 2017 Sakartvelo with 242 species is presented. Fifteen scale insect families from 30 extant families in the world are presented for Sakartvelo. Most species-rich families are: Diaspididae with 90 species belonging Keywords: to 38 genera, Pseudococcidae with 72 species belonging to 31 genera and Coccidae with 43 species Hadzibeyli belonging to 23 genera. Other families are: Acanthococcidae (5 genera, 10 species), Asterolecaniidae (3 Sternorryncha Coccoidea genera, 5 species), Cryptococcidae (2 genera, 3 species), Eriococcidae (1 genus, 2 species), Kermesidae (1 Coccids genus, 3 species), Kerriidae (1 genus, 1 species), Kuwaniidae (1 genus, 1 species), Marchalinidae (1 genus, Pests 1 species), Margarodidae (2 genera, 3 species), Monophlebidae (2 genera, 2 species), Ortheziidae (3 Transcaucasia genera, 5 species), Putoidae (1 genus, 1 species). Anophococcus agropyri was found in the collection of Z. Hadzibeyli and it is herein recorded for the first time from Georgia. Distribution and host plant data for each species is given. © 2017 Agricultural University of Georgia. Production and hosting by Elsevier B.V. This is an open access article under the CC BY-NC-ND license (http://creativecommons.org/licenses/by-nc-nd/4.0/).

Introduction hermaphroditism; (iii) they have simple pathogen-like life cycles, in which dispersal is achieved by first instar nymphs blown by the The Scale insects (Hemiptera: Coccomorpha) are pests of many wind; and (iv) they have variable host specificities [4]. ornamental and agricultural plants. They cause damage by feeding In Sakartvelo (Georgia) the first study about scale insects was on plant sap, reducing vigor and producing chlorotic areas and published in 1934, by N. S. Borchsenius [5]. The basis for studying sooty mould at feeding locations, premature leaf drop and dis- the scale insect fauna, their taxonomy, morphology, biology, eco- torting stems and bark. Large populations of coccoids can cause the nomic aspects and natural enemies in Sakartvelo were provided death of their host plant [1]. Scale insects aren't all harmful and mainly by Hadzibeyli (1941e1983), Yasnosh (1957e1998) and some of them are used as a main source of income for humans, such Japoshvili (1996e2008). In Hadzibeyli's monograph [6] and publi- as producing natural dyes and some other products, in Mexico, cations of Hadzibeyli [7e15] it was recorded about 220 species of Peru, Brazil, Ethiopia, Australia, Chile, India and other countries [2]. Scale Insects in Sakartvelo. The latest publications on coccoids in Honeydew is an important food source for many birds and mam- Sakartvelo were published in 2008 [16,17]. mals and especially other insects [3]. Scale Insects are interesting to In this paper, we aimed to update the list of scale insect species evolutionary biologists for a number of reasons: (i) they have in Sakartvelo, considering modern taxonomic changes. intimate relationships with host plants, endosymbionts and social Hymenoptera; (ii) they have diverse genetic systems including thelytoky, arrhenotoky, paternal genome elimination, and Material and methods

All existed material of references had been analyzed and addi-

* Corresponding author. tionally dry and slide collection of Dr. Hadzibeyli were studied, this E-mail address: [email protected] (G. Japoshvili). collection is housed in the Entomology collection of the Agricul- Peer review under responsibility of Journal Annals of Agrarian Science. tural University of Georgia. Species names are listed following the http://dx.doi.org/10.1016/j.aasci.2017.05.002 1512-1887/© 2017 Agricultural University of Georgia. Production and hosting by Elsevier B.V. This is an open access article under the CC BY-NC-ND license (http:// creativecommons.org/licenses/by-nc-nd/4.0/). M. Batsankalashvili et al. / Annals of Agrarian Science 15 (2017) 252e268 253 nomenclature used in the scale insect database ScaleNet [4]. List of Distribution worldwide: Holarctic Region [4,21]. host plants are shown according to Georgian records. 12. Asterodiaspis quercicola (Bouche) Host plants. Quercus sp. [6], Q. iberica, Q. glauca (Fagaceae) [20]. Georgian localities. Tbilisi, , , , Sur- Results ami [1,6,20], Kvemo Chala (Dry material). Distribution worldwide: Afrotropical Region, Australian Re- Family e Acanthococcidae gion, Holarctic Region [4,21]; Genus e Acanthococcus Signoret 13. Asterodiaspis variolosa (Ratzeburg) 1. Acanthococcus aceris Signoret Host plants. Quercus hartwissiana (Fagaceae) [7]. Host plants. Acer campestre (Sapindaceae) [6]. Georgian localities. Tbilisi Botanical Garden [7]. Georgian localities. Tbilisi botanical garden [1]. Distribution worldwide: Holarctic Region, Australian Region, Distribution worldwide: Palaearctic Region [4]. Neotropical Region [4,21]. 2. Acanthococcus roboris (Goux) Genus e Planchonia Signoret Host plants. Pterocarya pterocarpa (Juglandaceae), Quercus 14. Planchonia arabidis Signoret imeretina, Castanea sativa (Fagaceae) [6]. Host plants. Thymus sp., T. transcaucasicus (Lamiaceae), Hedera Georgian localities. Tbilisi, Tbilisi botanical garden, , helix, H. pastuchovii, H. colchica (Araliaceae), Dianthus sp. (Car- , Matsimi (Dry material). yophyllaceae), Jasminum fruticans, Ligustrum vulgare (Oleaceae), Distribution worldwide: Palaearctic Region [4]. Teucrium polium (Lamiaceae), Cistus salviifolius (Cistaceae), 3. Acanthococcus spiraeae Borchsenius Andropogon virginicus (Poaceae) [6], Hieracium sp. (Compositae) Host plants. Spiraea hypericifolia, S. crenata (Rosaceae) [6]. (Dry material). Georgian localities. East Georgia [6], Tbilisi (Dry material). Georgian localities. Tbilisi, Borjomi, [6,7,20], Tbilisi Distribution worldwide: Palaearctic Region [4]. Botanical Garden, Koda (Dry material). Genus e Anophococcus (Signoret) Distribution worldwide: Holarctic Region, Australian Region 4. Anophococcus inermis (Green) [4,21]. Host plants. Poaceae Genus e Pollinia Targioni Tozzetti Georgian localities. Georgia [18]. 15. Pollinia pollini (Costa) Distribution worldwide: Europe [4]. Host plants. Olea europaea (Oleaceae) [6]. 5. Anophococcus adzharicus (Hadzibeyli) Georgian localities. Eastern Georgia [6]. Host plants. Vulpia unilateralis (Poaceae) [15]. Distribution worldwide: Holarctic Region [4,21]. Georgian localities. West Georgia, Batumi Botanical Garden [6], [15], Kolkheti (Slide and Dry material). FamilyeCoccidae Distribution worldwide: Georgia [4]. Genus e Acanthopulvinaria Borchsenius 6. Anophococcus agropyri (Borchsenius) 16. Acanthopulvinaria orientalis (Nasonov) Host plants. Ryegrass (Dry material). Host plants. Salsola ericoides, Kochia sp. (Amaranthaceae), Georgian localities. Tbilisi (Dry material). Rheum sp. (Polygonaceae), Reaumuria sp. (Tamarikaceae) [6]. Distribution worldwide: Palaearctic Region [4]. Georgian localities. South-Eastern part of East Georgia, Shiraqi, 7. Anophococcus insignis (Newstead) Gareji, Tbilisi [6]. Host plants. Poaceae [6]. Distribution worldwide: Palaearctic Region [4,21]. Georgian localities. Georgia [6]. Genus e Anapulvinaria Borchsenius Distribution worldwide: Holarctic Region [4]. 17. Anapulvinaria pistaciae (Bodenheimer) Genus e Gossyparia Signoret Host plants. Pistacia vera, Pistacia atlantica (Anacardiaceae) [6]. 8. Gossyparia spuria (Modeer) Georgian localities. East Georgia, Shiraqi [6], (Dry Host plants. Ulmus sp., U. glabra, U. minor, U. elliptica (Ulmaceae) material). [1,6,20]. Distribution worldwide: Palaearctic Region [4,21]. Georgian localities. Tbilisi, Kojori, Borjomi [1,6,20], , Genus e Ceroplastes Gray (Dry material). 18. Ceroplastes japonicus Green Distribution worldwide: Holarctic Region [4]. Host plants. Diospyros kaki (Ebenaceae), Punica granatum Genus e Greenisca Borchsenius (Lythraceae), Morus sp. (Moraceae), Chaenomeles sinensis 9. Greenisca gouxi (Balachowsky) (Rosaceae), Camellia sinensis (Theaceae) [6], Ilex aquifolium Host plants. Cypreaceae and Poaceae [4]. (Aquifoliaceae), Prunus laurocerasus (Rosaceae), Osmanthus fra- Georgian localities. East Georgia [18,20]. grans (Oleaceae), Hedera sp. (Araliaceae), Diospyros sp. (Ebena- Distribution worldwide. Palaearctic region [4]. ceae), Salix alba (Salicaceae) [1], Laurus nobilis (Lauraceae), Genus e Rhizococcus (Signoret) Eriobotrya japonica (Rosaceae) [1,6]. 10. Rhizococcus saxatilis (Kiritchenko) Georgian localities. West Georgia, Tbilisi, , [1,6], Host plants. Fragaria vesca (Rosaceae), Teucrium chamaedrys Kheta (), Sokhumi, Eshera (Dry material). (Lamiaceae) [6]. Distribution worldwide: Palaearctic Region [4,21]. Georgian localities. Adjara (Dry material). 19. Ceroplastes sinensis Del Guercio Distribution worldwide: Palaearctic Region [4]. Host plants. Citrus sp., C. nobilis, (Rutaceae), Diospyros kaki (Ebenaceae), Cinnamomum camphora, Laurus nobilis (Lauraceae), Family -Asterolecaniidae Punica granatum (Lythraceae), Camellia sinensis (Theaceae), Acca Genus e Asterodiaspis Signoret sellowiana (Myrtaceae), Cedrus deodara (Pinaceae), Ilex colchica 11. Asterodiaspis minor (Russell) (Aquifoliaceae), Magnolia grandiflora (Magnoliaceae), Spiraea Host plants. Quercus sp. [6], Q. hartwissiana, Q. macranthera sp., Crataegus sp. (Rosaceae) [6]. (Fagaceae), (Dry material). Georgian localities. West Georgia, Batumi [6]. Georgian localities. Tbilisi, , Batumi [6,20], Kvemo Chala, (Dry material). 254 M. Batsankalashvili et al. / Annals of Agrarian Science 15 (2017) 252e268

Distribution worldwide: Australian Region, Holarctic Region, Populus deltoides (Salicaceae), Diospyros kaki (Ebenaceae), Acer Neotropicalal Region [4,21]. sp. (Sapindaceae) [6,11]. Genus e Coccus Linnaeus Georgian localities. , Kolkheti lowland, [6,11]. 20. Coccus hesperidum Linnaeus Distribution worldwide: Mediterranean Region (Georgia, Host plants. Pinus halepensis, P. canadensis, Picea glauca (Pina- Turkey) [4]. ceae), Viscum album (Santalaceae), Populus suaveolens (Salica- 27. Eulecanium sericeum (Lindinger) ceae), Rubus sp. (Rosaceae), Vitis vinifera (Vitaceae), Cyclamen sp. Host plants. Abies nordmanniana (Pinaceae) [6,11]. (Primulaceae) [6], Citrus limon, C. aurantium, C. nobilis (Ruta- Georgian localities. Upper and lower , Bakhmaro, ceae), Phoenix sp. (Arecaceae), Laurus nobilis (Lauraceae) [7], Shaori, Oni [11,26], (Dry material). (Araliaceae) Hedera colchica, H. helix, Rubus caucasicus, R. hirtus, Distribution worldwide: Palaearctic Region [4,21]. R. sanctus (Rosaceae), Buxus sempervirens (Buxaceae), Pinus syl- 28. Eulecanium tiliae (Linnaeus) vestris (Pinaceae), Acer campestre (Sapindaceae) [8], Artemisia Host plants. Juglans regia (Juglandaceae), Quercus iberica vulgaris (Compositae) [6,8], Acacia sp. (Fabaceae), Eriobotrya (Fagaceae) [15], Parrotia persica (Hamamelidaceae), Crataegus japonica (Rosaceae), Ficus sp. (Moraceae), Pinus sp. (Pinaceae), microphylla, C. kyrtostyla, Prunus caspica, P. cerasifera, Rosa sp., Hedera sp. (Araliaceae), Elaeagnus sp. (Elaeagnaceae) [1]. R. canina, Cydonia vulgaris (Rosaceae), Tilia sp., T. caucasica, Georgian localities. Batumi, Tbilisi and Sokhumi Botanical T. cordata (Malvaceae) [6], Quercus robur subsp. imeretina Garden, Tbilisi, Batumi, , Sokhumi, Kolkheti, (Banoja, (Fagaceae) (Slide material). Sataplia), Lagodekhi (Matsimi), , , Ingiri, Georgian localities. East and West Georgia, Shiraqi, Eldari, Mtskheta, Telavi [1,6,8]. Kavtiskhevi, Borjomi, Tbilisi, , Sokhumi [6,11], East Distribution worldwide: Afrotropical Region, Australian Re- Georgia [6], Adjameti (slide). gion, Holarctic Region, Neotropical Region, Oriental Region Distribution worldwide: Holarctic Region [4,21]. [4,21]. 29. Eulecanium transcaucasicum Borchsenius 21. Coccus pseudomagnoliarum (Kuwana) Host plants. Ulmus sp. (Ulmaceae), Cydonia vulgaris (Rosaceae) Host plants. Acca sellowiana (Myrtaceae), Punica granatum [11]. (Lythraceae), Rhamnus pallasii (Rhamnaceae), Juglans regia Georgian localities. Tbilisi, [11]. (Juglandaceae), Celtis australis (Cannabaceae), Ulmus sp. Distribution worldwide: (, Georgia) [4]. (Ulmaceae) [6]. Citrus limon, C. aurantium, C. nobilis (Rutaceae) Genus e Exaeretopus Newstead [7], Citrus trifoliata (Rutaceae), Berberis sp. (Berberidaceae), 30. Exaeretopus agropyri (Hadzibeyli) Tamarix sp. (Tamaricaceae) [1]. Host plants. Elymus caucasicus [6,11,22], Elytrigia sp. (Poaceae) Georgian localities. Tbilisi, , Shiraqi, Gareji [1,6,7], (slide). Makhinjauri, Chakvistavi (Dry material). Georgian localities. East Georgia [15], Tbilisi, Mtskheta [22], Distribution worldwide: Holarctic and Neotropical Regions Tbilisi Botanical Garden (Slide material). [4,21]. Distribution worldwide: Mediterranean Region (Georgia, Genus eEriopeltis Signoret Turkey) [4]. 22. Eriopeltis festucae (Boyer de Fonscolombe) Genus eHadzibeyliaspis Koteja Host plants. Arrhenatherum elatius, Elymus caucasicus, Lolium 31. Hadzibeyliaspis stipae (Hadzibeyli) rigidum [6], Brachypodium silvaticum [22], Festuca pratensis Host plants. Stipa holosericea [6], S. caucasica [6,11,15], (Poaceae) [7]. S. pulcherrima (Poaceae) (Slide material). Georgian localities. Tbilisi, Kheltubani, Gori, Mtskheta, Georgian localities. Tbilisi [6,22]. Natakhtari, Saguramo, Krtsanisi, , , Chaqvi, Distribution worldwide: Palaearctic Region [4]. Banoja, Makharadze, Darchi [7,10,22], Banoja (Dry material). Genus e Lichtensia Signoret Distribution worldwide: Holarctic Region [4,21]. 32. Lichtensia viburni Signoret 23. Eriopeltis lichtensteini Signoret Host plants. Hedera colchica (Araliaceae), Viburnum tinus Host plants. Festuca sp., Phleum pratense (Poaceae) [10]. (Adoxaceae), Ilex colchica (Aquifoliaceae), Eurya japonica (Pen- Georgian localities. Tbilisi, Gori [10]. taphylacaceae) [6]. Distribution worldwide: Palaearctic Region [4]. Georgian localities. West Georgia [6]. Genus e Eucalymnatus Cockerell in Cockerell and Parrott Distribution worldwide: Palaearctic Region [4,21]. 24. Eucalymnatus tessellatus (Signoret) Genus eLuzulaspis Targioni Tozzetti Host plants. Laurus nobilis (Lauraceae) [6], Linum grandiflorum 33. Luzulaspis grandis Borchsenius (Linaceae) [7]. Host plants. Stachys sylvatica (Lamiaceae), Poa sp. (Poaceae) Georgian localities. Tbilisi, Batumi and Sokhumi Botanical [10,22]. Garden [1,6,7]. Georgian localities. Abkhazeti [6], Tbilisi, Gori, Batumi, Distribution worldwide: Afrotropical Region, Australian Re- Mtskheta, Tskneti [10,22]. gion, Holarctic Region, Neotropical Region, Oriental Region Distribution worldwide: Palaearctic Region [4,21]. [4,21]. 34. Luzulaspis luzulae (Dufour) Genus eEulecanium Cockerell Host plants. Unknown. 25. Eulecanium caraganae Borchsenius Georgian localities. Tbilisi [1]. Host plants. Caragana sp. [11], Caragana grandiflora (Fabaceae) Distribution worldwide: Palaearctic Region [4,21]. [6]. Genus eNemolecanium Borchsenius Georgian localities. Shiraqi [6,11]. 35. Nemolecanium abietis Borchsenius Distribution worldwide: Palaearctic Region [4,21]. Host plants. Abies nordmanniana (Pinaceae) [6]. 26. Eulecanium nocivum Borchsenius Georgian localities. Borjomi, Tsagveri [6]. Host plants. Liquidambar sp., L. stiraciflua (Altingiaceae), Pyrus Distribution worldwide: Mediterranean Region [4]. sp., Cydonia vulgaris, Cotoneaster sp., Crataegus sp. (Rosaceae), Genus e Neopulvinaria Hadzibeyli 36. Neopulvinaria innumerabilis (Rathvon) M. Batsankalashvili et al. / Annals of Agrarian Science 15 (2017) 252e268 255

Host plants. Vitis vinifera (Vitaceae), Diospyros kaki, D. lotus Host plants. Picea orientalis, P. abies, P. engelmannii, Tsuga can- (Ebenaceae), Platanus orientalis (Platanaceae), Juglans regia adensis (Pinaceae) [6]. (Juglandaceae), Malus sp., Pyrus sp., Prunus sp., P. persica, Georgian localities. Tbilisi, Batumi, [1,6]. P. cerasifera, Chaenomeles sinensis, Crataegus sp., Rosa canina Distribution worldwide: Holarctic Region [4,21]. (Rosaceae), Citrus trifoliata (Rutaceae), Punica granatum 43. Physokermes piceae (Schrank) (Lythraceae), Populus sp. (Salicaceae), Acer sp., A. platanoides Host plants. Picea orientalis (Pinaceae) [6,20]. (Sapindaceae), Tilia sp., T. rubra, T. cordata (Malvaceae), Acacia Georgian localities. Batumi Botanical Garden, Borjomi, Bakur- sp., Phaseolus vulgaris (Fabaceae), Cornus sp. (Cornaceae), iani, , , Tbilisi [20,25], Georgia [6]. Cucurbita pepo (Cucurbitaceae) [6], Cydonia oblonga (Rosaceae), Distribution worldwide: Palaearctic Region [4,21]. Platanus sp. (Platanaceae), Fraxinus sp. (Oleaceae) [1]. Genus e Pulvinaria Targioni Tozzetti Georgian localities. West and east Georgia, , Tbilisi, 44. Pulvinaria aurantii (Cockerell) Argveti [1,6,23], , Sakara (Dry material), Khashuri, Host plants. Citrus reticulata, C. limon, C. sinensis, C. trifoliata, C. Kvatsikhe, Kutaisi, Khotevi, Gardabani, Telavi region (Slide aurantium, C. paradisi, C. maxima (Rutaceae), Eriobotrya japonica material). (Rosaceae), Laurus nobilis (Lauraceae), Diospyros kaki, D. lotus Distribution worldwide: Holarctic Region [4,21]. (Ebenaceae), Pittosporum tobira (Pittosporaceae), Nerium Genus ePalaeolecanium Sulc oleander (Apocynaceae), Fatsia japonica (Araliaceae) [6]. 37. Palaeolecanium bituberculatum (Signoret) Georgian localities. Abkhazeti [6], Sokhumi, Gulripshi (Dry Host plants. Malus domestica, M. orientalis, Cydonia vulgaris, material). Pyrus communis, Prunus sp., P. laurocerasus [6], P. spinosa, Cra- Distribution worldwide: Palaearctic Region [4]. taegus sp. [6,20], Malus sylvestris (Rosaceae) [7]. 45. Pulvinaria floccifera (Westwood) Georgian localities. West and East Georgia, Tbilisi, Akhaltsikhe, Host plants. Ilex colchica, I. aquifolium (Aquifoliaceae), Taxus , Gori [1,6,7,20], Gremi (Dry material). baccata, Cephalotaxus fortunei (Taxaceae), Camellia sinensis, Distribution worldwide: Palaearctic Region [4,21]. C. japonica, C. sasanqua (Theaceae), Eurya japonica (Pentaphy- Genus eParthenolecanium Sulc lacaceae), Euonymus japonicus (Celastraceae), Pittosporum tobira 38. Parthenolecanium corni (Bouche) (Pittosporaceae), Hedera helix (Araliaceae), Cycas revoluta Host plants. Cistus salviifolius (Cistaceae), Albizia julibrissin, (Cycadaceae), Osmanthus fragrans (Oleaceae), Citrus reticulata Cajanus cajan (Fabaceae), Plectranthus sp., P. scutellarioides (Rutaceae) [6]. (Lamiaceae), Celtis australis subsp. caucasica (Canabaceae), Georgian localities. Tbilisi Botanical Garden, Mtsvane Kontskhi, Urtica sp., U. urens (Urticaceae) [6], Morus alba (Moraceae), Ingiri [1,6], , (Dry material). Carpinus orientalis (Betulaceae), Ulmus minor (Ulmaceae) (Dry Distribution worldwide: Holarctic Region, Australian Region, material). Afrotropical Region, Oriental Region, Neotropical Region [4,21]. Georgian localities. All subtropical zone of Georgia, Tbilisi, 46. Pulvinaria juglandii Hadzibeyli Akhaltsikhe, Tetritskaro, Agara, Kurta, , Borjomi, Host plants. Juglans regia, Pterocarya pterocarpa (Juglandaceae) Bakuriani, , Telavi, Vardzia, Adjara, Keda [1,6,7,26]. [6,27]. Distribution worldwide: Australian Region, Holarctic Region, Georgian localities. East Georgia, Mukhrani, Kavtiskhevi [6,27], Neotropical Region, Oriental Region [4,21]. Basiani (Dry material). 39. Parthenolecanium fletcheri (Cockerell) Distribution worldwide: Caucasus [4]. Host plants. Platycladus orientalis (Cupressaceae) [6]. 47. Pulvinaria peregrina Ben-Dov Georgian localities. Tbilisi [6]. Host plants. Diospyros kaki, D. lotus (Ebenaceae), Acer sp., Distribution worldwide: Palaearctic Region, Neotropical Re- A. japonicum (Sapindaceae), Citrus trifoliata (Rutaceae), Celtis gion, Nearctic Region [4]. sinensis (Cannabaceae), Pyrus caucasica, P. communis (Rosaceae), 40. Parthenolecanium persicae Kawecki Abelmoschus manihot (Malvaceae) [6], Acer palmatum (Sapin- Host plants. Vitis vinifera (Vitaceae), Citrus sp. (Rutaceae), Dio- daceae) (Dry material). spyros kaki, D. lotus (Ebenaceae), Punica granatum (Lythraceae), Georgian localities. Tbilisi, Batumi Botanical Garden, Qedi, Berberis vulgaris (Berberidaceae), Prunus laurocerasus, Spiraea Ozurgeti, Kobuleti [1,6]; Makhuntseti (Dry material). hypericifolia (Rosaceae), Morus alba (Moraceae) [6], Citrus aur- Distribution worldwide: Palaearctic Region [4]. antium (Rutaceae) (Dry material). 48. Pulvinaria vini Hadzibeyli Georgian localities. All subtropical zone of Georgia [6], Host plants. Vitis vinifera (Vitaceae) [6]. Makhinjauri, Tbilisi, Mukhrani (Dry material). Georgian localities. West Georgia [6], Kvatsikhe (Dry material), Distribution worldwide: Palaearctic Region, Neotropical Re- Khotevi (Ambrolauri), Sakara, Shrosha (Slide material). gion, Australian Region [4]. Distribution worldwide: Georgia [4]. 41. Parthenolecanium rufulum (Cockerell) 49. Pulvinaria vitis (Linnaeus) Host plants. Ulmus sp. (Ulmaceae), Quercus sp., Fagus orientalis Host plants. Fraxinus sp. (Oleaceae), Corylus sp., Carpinus sp., (Fagaceae), Carpinus betulus, C. orientalis (Betulaceae), Diospyros Alnus sp. (Betulaceae), Cydonia oblonga, Pyrus sp., Malus sp. kaki (Ebenaceae) [6], Castanea sativa (Fagaceae), Rosa gallica, (Rosaceae) [6], Acer sp. (Sapindaceae), Populus sp. (Salicaceae), Rubus sp. (Rosaceae) [20], Quercus iberica (Fagaceae) (Dry Pyrus caucasica (Rosaceae) [20], Vitis vinifera (Vitaceae) [6,7], material). Populus gracilis (Salicaceae) [1]. Georgian localities. All subtropical zone of Georgia, Tbilisi, Georgian localities. All subtropical zone of Georgia, Adigeni, Kojori, Manglisi, Surami, Tetri Tskaro, Natakhtari, Mtskheta, Akhaltsikhe, Bakuriani, Kutaisi, Zestafoni, Sakara, Sviri, Orta- Akhaltsikhe, Safari, Borjomi, , Batumi, Kobuleti, Makhar- chala, Tbilisi [1,6,7,20,23], Shulaveri, Telavi, Mukhrani (Dry adze, , Chiatura, Zestaponi, [6,20,24]. material). Distribution worldwide: Palaearctic Region [4,21]. Distribution worldwide: Holarctic Region, Neotropical Region, Genus e Physokermes Targioni Tozzetti Australian Region [4,21]. 42. Physokermes hemicryphus (Dalman) Genus e Rhizopulvinaria Borchsenius 50. Rhizopulvinaria artemisiae (Signoret) 256 M. Batsankalashvili et al. / Annals of Agrarian Science 15 (2017) 252e268

Host plants. Gypsophila elegans, Dianthus capitatus, Family e Cryptococcidae D. plumarius, Herniaria incana (Caryophyllaceae), Jasminum Genus e Cryptococcus Douglas officinale, J. fruticans (Oleaceae), Galium sp. (Rubiaceae), Anthe- 59. Cryptococcus aceris Borchsenius mis sp. (Compositae) [6], Artemisia fragrans (Compositae) [7], Host plants. Acer pseudoplatanus (Sapindaceae), Tilia rubra Trifolium sp. (Fabaceae) [10], Dianthus orientalis (Car- subsp. caucasica (Malvaceae) [6]. Acer sp [20]. Pyrus caucasica yophyllaceae) (Dry material). (Rosaceae) [6,20]. Georgian localities. Tbilisi Botanical Garden, Akhaltsikhe [7,10]. Georgian localities. Lagodekhi, Borjomi, Bakuriani [20]. Distribution worldwide: Palaearctic Region [4]. Abkhazeti, Shiraqi (Dry material). Genus eRhodococcus Borchsenius Distribution worldwide: Palaearctic Region [4,21]. 51. Rhodococcus spiraeae (Borchsenius) 60. Cryptococcus fagisuga Lindinger Host plants. Spiraea hypericifolia, S. crenata [6], S. altaica Host plants. Fagus orientalis (Fagaceae) [6,20]. (Rosaceae), Rhamnus pallasii (Rhamnaceae) [1]. Georgian localities. Signagi, Lagodekhi, Bakuriani [6,20]. Georgian localities. East Georgia, Tbilisi [1,6]. Distribution worldwide: Holarctic Region [4,21]. Distribution worldwide: Palaearctic Region [4]. Genus e Pseudochermes Nitsche 52. Rhodococcus turanicus (Archangelskaya) 61. Pseudochermes fraxini (Kaltenbach) Host plants. Rhamnus pallasii (Rhamnaceae) [6], Cotoneaster sp. Host plants. Fraxinus excelsior (Oleaceae) [6,20]. (Rosaceae) [1], Amorpha sp. (Fabaceae), Rhamnus cathartica Georgian localities. Borjomi [20]. (Rhamnaceae) [20]. Distribution worldwide: Palaearctic Region [4]. Georgian localities. Southeastern part of Georgia, Lagodekhi, Family eDiaspididae , Tsagveri, Tbilisi [1,6,20], Rustavi, Tbilisi Botanical Genus e Aonidia Targioni Tozzetti Garden (Dry material). 62. Aonidia lauri (Bouche) Distribution worldwide: Palaearctic Region [4]. Host plants. Laurus nobilis (Lauraceae) [6]. Genus eSaissetia Deplanche Georgian localities. Tbilisi [1,6]. 53. Saissetia coffeae (Walker) Distribution worldwide: Holarctic Region, Neotropical Region, Host plants. Coffea arabica (Rubiaceae), Phoenix sp., Oriental Region [4,21]. P. canariensis, Arecaceae sp. (Arecaceae), Orchis sp. (Orchid- 63. Aonidia mediterranea (Lindinger) aceae), Cycas revoluta (Cycadaceae), Asparagus officinalis Host plants. Juniperus foetidissima (Cupressaceae) [6]. (Asparagaceae), Solanum tuberosum (Solanaceae), Nephrolepis Georgian localities. East Georgia, Shiraqi, Eldari [6]. sp. (Nephrolepis scottii) [6], Filicineae, Orchidaceae, Linum Distribution worldwide: Palaearctic Region [4]. grandiflorum (Linaceae) [7]. Genus e Aonidiella Berlese and Leonardi Georgian localities. Batumi, Tbilisi [1,6e8]. 64. Aonidiella aurantii (Maskell) Distribution worldwide: Afrotropical Region, Australian Re- Host plants. Agave sp. (Asparagaceae) [6]. gion, Palaearctic Region, Nearctic Region, Neotropical Region, Georgian localities. Batumi Botanical Garden [6]. Oriental Region, [4,21]. Distribution worldwide: Afrotropical Region, Australian Re- 54. Saissetia oleae (Olivier) gion, Palaearctic Region, Nearctic Region, Neotropical Region, Host plants. Olea europaea, Nerium oleander, Albizia julibrissin, Oriental region [4,21]. Cajanus cajan, Machaonia grandiflora (Rubiaceae), Citrus nobilis, 65. Aonidiella citrina (Coquillett) C. sinensis (Rutaceae) [6]. Host plants. Citrus reticulata, C. limon, C. sinensis, C. paradisi, Georgian localities. Coast, , Tbilisi [1,6e8]. C. trifoliata (Rutaceae), Agave sp. (Asparagaceae), Ilex colchica Distribution worldwide: Afrotropicalal Region, Australian Re- (Aquifoliaceae), Camellia japonica, C. sasanqua (Theaceae), Buxus gion, Holarctic Region, Neotropical Region, Oriental Region sempervirens (Buxaceae) [6]. [4,21]. Georgian localities. West Georgia: Sokhumi, Batumi, , Genus eScythia Kiritchenko Makhinjauri, Tbilisi [1,6,7,29], Tsvermagala (Dry material). 55. Scythia stipae Hadzibeyli Distribution worldwide: Palaearctic Region, Neotropical Re- Host plants. Stipa holosericea [6], Festuca rupicola (Poaceae) gion, Australian Region, Nearctic Region, Oriental Region, Afro- [6,28]. tropical Region, [4,21]. Georgian localities. Shiraqi, Eldari [28]. 66. Aonidiella taxus Leonardi Distribution worldwide: Palaearctic Region [4]. Host plants. Cephalotaxus fortunei (Taxaceae), Podocarpus sp., Genus e Sphaerolecanium Sulc P. macrophyllus (Podocarpaceae) [1]. 56. Sphaerolecanium prunastri (Boyer de Fonscolombe) Georgian localities. West Georgia, Adjara, Abkhazeti [1], Tbilisi Host plants. Prunus domestica, P. divaricata [1], P. spinosa [1,5,12], Tbilisi, and Sokhumi Botanical Gardens [37]. (Rosaceae) [1,6]. Distribution worldwide: Palaearctic Region, Nearctic Region, Georgian localities. all subtropical zone of Georgia, Tbilisi Neotropical Region, Oriental Region [4,21]. [1,6,7], Khashuri, Saguramo, Tkibuli (Dry material). Genus e Aspidiotus Bouche Distribution worldwide: Holarctic Region [4,21]. 67. Aspidiotus destructor Signoret Genus e Stotzia (Marchal) Host plants. Camellia sinensis, C. japonica, C. sasanqua (The- 57. Stotzia ephedrae (Newstead) aceae), Ilex colchica (Aquifoliaceae), Eurya japonica (Pentaphy- Host plants. Ephedra major (Ephedraceae) [6]. lacaceae), Ophiopogon japonicus (Asparagaceae), Solanum Georgian localities. East Georgia [6]. tuberosum (Solanaceae) [6]. Distribution worldwide: Palaearctic Region [4]. Georgian localities. Abkhazeti: Gagra, SokhumiAdjara, Poti, 58. Stotzia maxima (Borchsenius) Zugdidi, Kutaisi, Tbilisi [1,6,37], Salibauri, , Host plants. Ephedra major (Ephedraceae) [6]. Makhinjauri, Chakvi (Dry material). Georgian localities. Vashlovani (Dry material). Distribution worldwide: Palaearctic Region [4]. M. Batsankalashvili et al. / Annals of Agrarian Science 15 (2017) 252e268 257

Distribution worldwide: Afrotropical Region, Palaearctic Re- Georgian localities. Avadkhara, Lagodekhi, Manglisi, Bakuriani gion, Nearctic Region, Neotropical Region, Australian Region, [9], Sapara (Akhaltsikhe) (Dry material), Manglisi, Lagodekhi Oriental Region [4,21]. reserve (Slide material). 68. Aspidiotus nerii Bouche Distribution worldwide. Holarctic Region, Oriental Region Host plants. Agave sp., Asparagus sp., A. plumosus (Aspar- [4,21]. agaceae), Billbergia sp. (Bromeliaceae), Cajanus cajan (Fabaceae), Genus e Chortinaspis Ferris Citrus limon (Rutaceae), Hedera helix, H. pastuchovii (Araliaceae), 77. Chortinaspis subterranea Lindinger Prunus laurocerasus (Rosaceae), Rhododendron ponticum (Erica- Host plants. Elymus repens (Poaceae) [6]. ceae), Saxifraga sp. (Saxifragaceae), Vernicia fordii (Euphorbia- Georgian localities. East Georgia, Tbilisi, District of Gori, Skra ceae) [6]. [6,7,29]. Georgian localities. Tbilisi, Makhinjauri [1,6,7], Batumi, , Distribution worldwide. Palaearctic Region [4,21]. Kakhaberi (Dry material). Genus e Chrysomphalus Ashmead Distribution worldwide: Afrotropical Region, Oriental Region, 78. Chrysomphalus dictyospermi (Morgan) Palaearctic Region, Nearctic Region, Neotropical Region, Host plants. Buxus sempervirens (Buxaceae), Camellia sinensis Australian Region [4,21]. (Theaceae), Chamaerops humilis (Arecaceae), Citrus sinensis, 69. Aspidiotus pothos Takagi C. limon, C. paradisi (Rutaceae), Cinnamomum camphora Laurus Host plants. Camellia sinensis (Theaceae) nobilis, (Lauraceae), Cornus mas (Cornaceae), Diospyros lotus Georgian localities. Adjara (Chakvi, Batumi, Makhinjauri), Tbi- (Ebenaceae), Euonymus japonicus (Celastraceae), Hedera helix lisi [29]. (Araliaceae), Hibiscus syriacus (Malvaceae), Phytolacca ameri- Distribution worldwide. Palaearctic Region [4]. cana (Phytolaccaceae), Prunus laurocerasus (Rosaceae) [6]. Genus eAulacaspis Cockerell Georgian localities. Tbilisi, West Georgia: Gagra, Batumi [1,6,7], 70. Aulacaspis rosae (Bouche) Sokhumi [29]. Host plants. Rosa sp., R. canina, Rubus sp. (Rosaceae) [6]. Distribution worldwide. Palaearctic Region, Neotropical Re- Georgian localities. Tbilisi, all subtropical zone of Georgia gion, Australian Region, Afrotropical Region, Oriental Region, [1,6,7]. Nearctic Region [4,21]. Distribution worldwide: Afrotropical Region, Australian Re- Genus e Comstockaspis MacGillivray gion, Palaearctic Region, Nearctic Region, Neotropical Region, 79. Comstockaspis perniciosa (Comstock) Oriental Region [4,21]. Host plants. Malus sp. (Rosaceae), Citrus trifoliata (Rutaceae), Genus eCarulaspis MacGillivray Quercus hartwissiana (Fagaceae), Salix babylonica, S. matsudana 71. Carulaspis minima (Signoret) (Salicaceae), Vitis vinifera (Vitaceae) [6]. Host plants. Juniperus foetidissima, J. oxycedrus, Cryptomeria Georgian localities. Tbilisi, Zestafoni, Adjameti, Khashuri, Var- japonica (Cupressaceae), Taxus baccata (Taxaceae) [1], Cupressus iani, Zestafoni, Samtredia [6,7]. sempervirens, Platycladus orientalis [1,2,4], Juniperus virginiana Distribution worldwide. Afrotropical Region, Palaearctic Re- (Cupressaceae) [4]. gion, Australian Region, Nearctic Region, Oriental Region, Georgian localities. Tbilisi, Tsinandali, Agudzera, Gori [1,2,4,12]. Neotropical Region [4,21]. Distribution worldwide. Afrotropical Region, Australian Re- Genus e Contigaspis (MacGillivray) gion, Palaearctic Region, Nearctic Region, Neotropical Region 80. Contigaspis farsetiae (Hall) [4,21]. Host plants. Artemisia fragrans (Artemisia meyeriana) (Compo- 72. Carulaspis visci (Schrank) sitae) [6]. Host plants. Juniperus sp., Platycladus orientalis, Cupressus Georgian localities. East Georgia [6]. sempervirens (Cupressaceae) [20]. Distribution worldwide. Palaearctic Region [4]. Georgian localities. Tbilisi, Mtskheta, Gori, Borjomi [20]. Genus e Diaspidiotus Berlese Distribution worldwide. Palaearctic Region [4,21]. 81. Diaspidiotus caucasicus (Borchsenius) Genus eChionaspis Signoret Host plants. Populus sp., P. gracilis, P. hybrida (Salicaceae) [6]. 73. Chionaspis dryina (Takahashi) Georgian localities. Tbilisi, Gori [1,6,20]. Host plants. Enkianthus perulatus (Ericaceae) [6]. Distribution worldwide. Palaearctic Region [4,21]. Georgian localities. Batumi Botanical Garden, Mtsvane Konts- 82. Diaspidiotus degeneratus (Cockerell) khi [6]. Host plants. Camellia japonica, C. sasanqua, C. sinensis (Thea Distribution worldwide. Palaearctic Region [4,21]. sinensis) (Theaceae), Ilex colchica (Aquifoliaceae), Eurya japonica 74. Chionaspis etrusca Leonardi (Pentaphylacaceae) [6]. Host plants. Tamarix sp. (Tamaricaceae) [6]. Georgian localities. West Georgia: Makharadze, Batumi, Chakvi Georgian localities. East Georgia, Shiraqi, Rustavi [6]. [6,29]. Distribution worldwide. Palaearctic Region, Oriental Region, Distribution worldwide. Holarctic Region [4,21]. Saudi Arabia, Mauritania, Nearctic Region [4,21]. 83. Diaspidiotus elaeagni (Borchsenius) 75. Chionaspis lepineyi Balachowsky Host plants. Elaeagnus angustifolia, E. rhamnoides (Elaeagna- Host plants. Quercus sp., Castanea sativa (Fagaceae) [6]. ceae) [6]. Georgian localities. East and West Georgia, Abkhazeti [6,29], Georgian localities. Tbilisi, Gardabani [1,6,7,20]. Kutaisi (Dry material). Distribution worldwide. Palaearctic Region [4,21]. Distribution worldwide. Palaearctic Region [4,21]. 84. Diaspidiotus gigas (Thiem and Gerneck) 76. Chionaspis salicis (Linnaeus) Host plants. Populus sp [6]. P. gracilis (Salicaceae) [20]. Host plants. Salix sp., Populus sp. (Salicaceae), Fraxinus excelsior Georgian localities. Tbilisi, Akhaltsikhe, Gori, Zugdidi, (Oleaceae), Euonymus latifolius (Celastraceae), Acer sp., A. [1,6,20,29]. negundo, A. heldreichii subsp. trautvetteri (Sapindaceae), Cytisus Distribution worldwide. Holarctic Region, Oriental Region ruthenicus (Fabaceae), Rhamnus cathartica, Frangula grandiflora [4,21]. (Rhamnaceae), Cornus mas (Cornaceae) [6]. 85. Diaspidiotus lenticularis Lindinger 258 M. Batsankalashvili et al. / Annals of Agrarian Science 15 (2017) 252e268

Host plants. Fagus sp. (Fagaceae) [29]. Georgian localities. Tbilisi, Qeda, Abastumani, Borjomi, Tsag- Georgian localities. West Georgia: Gagra [29]. veri, Manglisi [1,20,29], Dry material. Distribution worldwide. Australian Region, Palaearctic Region Distribution worldwide. Holarctic Region [4,21]. [4]. 96. Dynaspidiotus britannicus (Newstead) 86. Diaspidiotus marani (Zahradnik) Host plants. Prunus laurocerasus (Rosaceae), Hedera colchica, H. Host plants. Fraxinus excelsior (Oleaceae), Pyrus sp., Malus sp. helix (Araliaceae), Buxus sempervirens (Buxaceae), Ilex colchica (Rosaceae) [6]. (Aquifoliaceae), Ruscus aculeatus (Asparagaceae), Cinnamomum Georgian localities. Tbilisi [6]. camphora, Laurus nobilis (Lauraceae) [6]. Distribution worldwide. Palaearctic Region [4,21]. Georgian localities. East and West Georgia, Tbilisi [1,6], Kutaisi 87. Diaspidiotus ostreaeformis (Curtis) [29], (Dry material). Host plants. Malus domestica, Prunus domestica [1], Malus ori- Distribution worldwide. Palaearctic Region, Nearctic Region, entalis (Rosaceae), Populus gracilis (Salicaceae), Acer sp. (Sapin- Neotropical Region [4,21]. daceae) [20]. 97. Dynaspidiotus ephedrarum (Lindinger) Georgian localities. Tbilisi, Gori, Akhaltsikhe, Skra [1,6,7,20]. Host plants. Ephedra major subsp. procera (Ephedraceae) [6,20]. Distribution worldwide. Palaearctic Region, Neotropical Re- Georgian localities. East Georgia, Mtskheta [6,20], Tbilisi [29]. gion, Australian Region, Nearctic Region, Oriental Region [4,21]. (Dry material). 88. Diaspidiotus prunorum (Laing) Distribution worldwide. Afrotropical Region, Palaearctic Re- Host plants. Prunus dulcis, Amygdalus nairica, Prunus domestica, gion, Oriental Region [4,21]. P. cerasifera, P. spinosa (Rosaceae) [6,30]. Genus e Epidiaspis Cockerell Georgian localities. Tbilisi, Shiraqi, Kakheti, Eldari [1,3,6,7], 98. Epidiaspis leperii (Signoret) Tbilisi Botanical Garden (Dry material). Host plants. Malus orientalis, Prunus domestica, Sorbus sp. Distribution worldwide. Palaearctic Region [4,21,30]. (Rosaceae), Olea europaea (Oleaceae), Acer platanoides, A. 89. Diaspidiotus pyri (Lichtenstein) negundo (Sapindaceae) [6], Pyrus caucasica [6,20], Malus syl- Host plants. Prunus sp., P. domestica, P. avium, Malus sp., Pyrus vestris [7], Pyrus salicifolia (Rosaceae) [1]. sp. (Rosaceae), Fraxinus excelsior (Oleaceae) [6]. Georgian localities. Tbilisi, Kobuleti, Tbilisi, Bakuriani, Borjomi, Georgian localities. Tbilisi [1,6]. Lagodekhi [1,6,7,20]. Distribution worldwide. Australian Region, Palaearctic Region Distribution worldwide. Neotropical Region, Nearctic Region, [4,21]. Palaearctic Region [4,21]. 90. Diaspidiotus zonatus (Frauenfeld) GenuseFiorinia Targioni Tozzetti Host plants. Quercus sp., Q. longipes, Fagus orientalis, Castanea 99. Fiorinia fioriniae Targioni Tozzetti sativa (Fagaceae) [6]. Host plants. Arecaceae sp [6]. Georgian localities. Tbilisi, Makharadze, Keda, Borjomi, Lago- Georgian localities. Abkhazeti, Adjara, Batumi and Sokhumi dekhi [1,6,20,29]. Botanical Gardens [6]. Distribution worldwide. Palaearctic Region [4,21]. Distribution worldwide. Palaearctic Region, Neotropical Re- Genus e Diaspis Costa gion, Australian Region, Oriental Region, Nearctic Region, Afro- 91. Diaspis boisduvalii Signoret tropical Region [4,21]. Host plants. Chamaerops humilis, Phoenix sp., P. reclinata (Are- Genus eFroggattiella Leonardi caceae), Phormium tenax (Xanthorrhoeaceae) [6]. 100. Froggattiella penicillata Green Georgian localities. West Georgia: Sokhumi [6]. Host plants. Bambusa sp. (Poaceae) [6]. Distribution worldwide. Palaearctic Region, Neotropical Re- Georgian localities. West Georgia, Batumi Botanical Garden [6]. gion, Australian Region, Nearctic Region, Afrotropical Region, Distribution worldwide. Palaearctic Region, Oriental Region, Oriental Region [4,21]. Nearctic Region [4,21]. 92. Diaspis bromeliae Kerner Genus eHemiberlesia Cockerell Host plants. Chamaerops humilis (Arecaceae), Ananas comosus, 101. Hemiberlesia cyanophylli (Signoret) Billbergia sp. (Bromeliaceae) [29]. Host plants. Camellia sinensis (Theaceae), Ilex colchica (Aquifo- Georgian localities. Greenhouse in Batumi [29]. liaceae) [6]. Distribution worldwide. Cosmopolitan [4]. Georgian localities. West Georgia: Adjara: Salibauri, Peria. 93. Diaspis echinocacti (Bouche) Anaseuli [6]. Host plants. Opuntia humifusa, Melocactus sp. (Cactaceae) [6]. Distribution worldwide. Afrotropical Region, Neotropical Re- Georgian localities. subtropical zone of West Georgia, Sokhumi, gion, Australian Region, Palaearctic Region, Nearctic Region, Tbilisi [1,6,7,29]. Oriental Region [4,21]. Distribution worldwide. Palaearctic Region, Neotropical Re- Comments. According Danzig (1993), this species was recorded gion, Australian Region, Afrotropical Region, Nearctic Region, by Borchsenius and Hadzibeili as Dynaspidiotus degeneratus Oriental Region [4,21]. (Cockerell), this statement needs confirmation. Genus e Duplachionaspis (MacGillivray) 102. Hemiberlesia lataniae (Signoret) 94. Duplachionaspis noaeae (Hall) Host plants. Phyllocactus hybridus (Cactaceae), Hoya carnosa Host plants. Noaea mucronata (Amaranthaceae) [6]. (Apocynaceae), Prosthechea fragrans (Orchidaceae) [6]. Phoenix Georgian localities. East Georgia: Tbilisi Vashlovani National sp. (Arecaceae) [7]. Park, Dzegvi [6,29]. Georgian localities. Batumi, Tbilisi, Sokhumi [6], Tbilisi Botan- Distribution worldwide. Palaearctic Region [4]. ical Garden [1,7]. Genus e Dynaspidiotus Thiem and Gerneck Distribution worldwide. Afrotropical Region, Australian Re- 95. Dynaspidiotus abietis (Schrank) gion, Palaearctic Region, Nearctic Region, Neotropical Region, Host plants. Pinus sylvestris var. hamata, Cedrus deodara [6], Oriental Region [4,21]. Abies nordmanniana, Pinus hamata [20], Picea orientalis [7,20], 103. Hemiberlesia rapax (Comstock) Pinus sylvestris (Pinaceae) [7]. M. 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Host plants. Albizia julibrissin (Fabaceae), Broussonetia papy- Georgian localities. Tbilisi, Mtskheta, Shiraqi, Telavi, Tskhinvali rifera (Moraceae), Citrus trifoliata (Rutaceae), Cortaderia selloana [1,6,7,20]. (Poaceae), Elaeagnus angustifolia (Elaeagnaceae), Fatsia sp. Distribution worldwide. Palaearctic Region [4,21]. (Araliaceae), Haworthia sp., H. fasciata (Xanthorrhoeaceae), 112. Lepidosaphes malicola (Borchsenius) Laurus nobilis (Lauraceae), Lonicera japonica (Caprifoliaceae), Host plants. Populus gracilis (Salicaceae), Quercus robur subsp. Osmanthus fragrans (Oleaceae), Vitis vinifera (Vitaceae) [6]. pedunculiflora, Q. petraea subsp. Iberica (Fagaceae), Juglans regia Georgian localities. Abkhazeti, Eshera, Adjara [6]. (Juglandaceae), Vitis vinifera (Vitaceae) [6,31], Rosa canina Distribution worldwide. Palaearctic Region, Afrotropical Re- (Rosaceae), Populus tremula (Salicaceae) [18], Taxus baccata gion, Neotropical Region, Australian Region, Oriental region, (Slide), Cephalotaxus harringtonii (Taxaceae). Nearctic Region [4,21]. Georgian localities. Borjomi, Gori, Tbilisi, Kakabeti, Surami, Genus eIschnaspis Douglas Karaleti, Saguramo, Mtskheta, Skra [1,6,31]. Lanchkhuti (slide). 104. Ischnaspis longirostris (Signoret) Distribution worldwide. Palaearctic Region, Oriental Region Host plants. Phoenix sp. (Arecaceae), Musa sp. (Musaceae) [6]. [4,21]. Georgian localities. Batumi and Sokhumi Botanical Garden [6]. 113. Lepidosaphes pallida Green Distribution worldwide. Afrotropical Region, Neotropical Re- Host plants. Cryptomeria japonica (Cupressaceae), Sciadopitys gion, Australian Region, Nearctic Region, Oriental Region, verticillata (Sciadopityaceae), Pinus sp. (Pinaceae), Taxus sp. Palaearctic Region [4,21]. (Taxaceae) [6,29,31]. Genus e Kuwanaspis MacGillivray Georgian localities. Abkhazeti, Batumi, Kobuleti, Ingiri, 105. Kuwanaspis howardi (Cooley) Tsetskhlauri, Sokhumi, Gulripshi, Mtsvane Kontskhi, Guria Host plants. Phyllostachys bambusoides, P. viridiglaucescens, P. [6,31], Samtredia, Batumi botanical garden (Dry material). mitis, P. puberula (Poaceae) [1]. Distribution worldwide. Australian Region, Palaearctic Region, Georgian localities. Tbilisi, Telavi, Batumi [1,6,8]. Oriental Region [4]. Distribution worldwide. Holarctic Region [4]. 114. Lepidosaphes pistaciae Archangelskaya 106. Kuwanaspis pseudoleucaspis (Kuwana) Host plants. Pistacia atlantica [6,31], P. vera (Anacardiaceae) [6]. Host plants. Pseudosasa japonica [7,31], Sasa sp., Arundinaria sp., Georgian localities. Kakheti, Shiraqi, Vashlovani [6]. Bambusa sp. (Poaceae) [1]. Distribution worldwide. Palaearctic Region, Oriental Region Georgian localities. Tbilisi, Batumi, Kolkheti [1,6,7,31]. [4,21]. Distribution worldwide. Palaearctic Region, Nearctic Region, 115. Lepidosaphes ulmi (Linnaeus) Australian Region, Neotropical Region, Oriental region [4,21]. Host plants. Juglans regia, Pterocarya pterocarpa (Juglandaceae), Genus eLepidosaphes Shimer Cornus mas (Cornaceae), Taxus baccata (Taxaceae), Tilia cordata, 107. Lepidosaphes beckii (Newman) T. rubra subsp. caucasica (Malvaceae), Castanea sativa, Quercus Host plants. Citrus nobilis, C. sinensis, C. limon (Rutaceae) [6]. imeretina, Fagus orientalis (Fagaceae), Clematis vitalba, C. ori- Georgian localities. Guria, Adjara [6], Chakvi, Makhinjauri, entalis (Ranunculaceae), Malus domestica, M. orientalis, Cratae- Batumi [29]. Batumi Botanical Garden, (Dry material). gus sp., Cotoneaster melanocarpus, Rosa sp., R. canina, Prunus Distribution worldwide. Afrotropical Region, Australian Re- laurocerasus (Rosaceae), Fraxinus excelsior (Oleaceae), Acer gion, Palaearctic Region, Nearctic Region, Neotropical Region, tataricum (Sapindaceae), Urtica dioica (Urticaceae), Vaccinium Oriental Region [4,21]. arctostaphylos, Rhododendron ponticum (Ericaceae), Zelkova 108. Lepidosaphes conchiformis (Gmelin) carpinifolia (Ulmaceae) [6,31], Platanus orientalis (Platanaceae), Host plants. Ficus carica (Moraceae), Ulmus minor (Ulmaceae), Populus gracilis [19], Populus sp. (Salicaceae), Malus sylvestris Carpinus betulus [6], C. orientalis (Betulaceae) [6,20]. (Rosaceae), Gleditsia triacanthos (Fabaceae), Quercus hartwissi- Georgian localities. Batumi, Chiatura, Eshera, Gagra, Gardabani, ana (Fagaceae) [7]. Gombori, Gulripshi, Lagodekhi, Makhinjauri, Melusheti, South- Georgian localities. Alvani, Akhaltsikhe, Avadhara, Bakuriani, ern parts of Adjara, Tbilisi [1,6,8,20,32], Batumi Botanical Gar- Batumi, Betania, Borjomi, Chakvi, Chiatura, Gagra, Gombori, den, Abkhazeti (Dry material). Gudamakari, , Java, Kakabeti, Kavtiskhevi, , Kiketi, Distribution worldwide. Palaearctic Region, Neotropical Re- Kojori, Manglisi, Melusheti, Mukhrani, Oni, Pasanauri, Qeda, gion, Nearctic Region, Afrotropical Region, Oriental Region Ritsa, Rtskhilati, , Surami, Tbilisi, Telavi, Tkibuli, [4,21]. , Tkviavi, , Tsalka,Tsinandali, Tskhinvali, 109. Lepidosaphes gloverii (Packard) Tskneti, Tsunda, Variani, Zestafoni [1,6,7,19,31], Adjara, Host plants. Citrus limon, C. reticulata, C. sinensis (Rutaceae) [6]. Makhinjauri, Batumi Botanical Garden (Dry material). Georgian localities. Abkhazeti, Adjara, Guria, Makhinjauri, Distribution worldwide. Australian Region, Palaearctic Region, Samtredia, Batumi, Poti, Tsetskhlauri, Mukhaestate, , Nearctic Region, Neotropical Region, Oriental Region [4,21]. Sokhumi, Eshera, Gagra [6,31]. Genus e Lecanopsis Targioni Tozzetti Distribution worldwide. Afrotropical Region, Australian Re- 116. Lecanopsis turcica (Bodenheimer) gion, Palaearctic Region, Nearctic Region, Neotropical Region, Host plants. Undetermined species (Poaceae) [6]. Oriental Region [4,21]. Georgian localities. Georgia [6]. 110. Lepidosaphes granati Koroneos Distribution worldwide. Palaearctic Region [4]. Host plants. Punica granatum (Lythraceae) [1,6], Celtis australis Genus eLeucaspis Signoret subsp. caucasica (Cannabaceae) [6]. 117. Leucaspis lowi Colvee Georgian localities. Southeastern part of East Georgia, Tbilisi, Host plants. Pinus sosnowsky [6], P. hamata [19], P. sylvestris, P. Batumi [1,6,20]. ayacahuite (Pinaceae). Distribution worldwide. Palaearctic Region [4,21]. Georgian localities. All subtropical zones of Georgia, Tbilisi, 111. Lepidosaphes juniperi Lindinger Manglisi, Surami, Abastumani, Mtskheta [1,6,19]. Host plants. Juniperus foetidissima [6], Juniperus oxycedrus Distribution worldwide. Palaearctic Region [21]. [6,20]. Platycladus orientalis (Cupressaceae) [6,7,20]. 118. Leucaspis pini (Hartig) Host plants. Pinus sp. (Pinaceae) [6]. 260 M. Batsankalashvili et al. / Annals of Agrarian Science 15 (2017) 252e268

Georgian localities. Georgia [6]. Distribution worldwide. Palaearctic Region, Australian Region, Distribution worldwide. Palaearctic Region [4]. Nearctic Region, Oriental Region, Neotropical Oceania, Afro- 119. Leucaspis pusilla Low tropical Region [4,21]. Host plants. Pinus sp. [6], P. hamata [19], P. nigra, P. halepensis Genus eParlatoria Targioni Tozzetti [7], P. ayacahuite (Pinaceae). 128. Parlatoria asiatica Borchsenius Georgian localities. Subtropical zones of west and east Georgia, Host plants. Ephedra major subsp. procera (Ephedraceae) [6]. Tbilisi, Borjomi, Surami, Abastumani, Gulripshi, Agudzera Georgian localities. South-Eastern part of East Georgia [6], [1,6,7,19]. Tbilisi (Dry material). Distribution worldwide. Palaearctic Region, Neotropical Re- Distribution worldwide. Palaearctic Region [4,21]. gion, Oriental Region [4,21]. 129. Parlatoria camelliae Comstock 120. Leucaspis signoreti Signoret Host plants. Camellia japonica, C. sasanqua (Theaceae), Prunus Host plants. Pinus sp. (Pinaceae) [6]. laurocerasus (Rosaceae) [6]. Georgian localities. All subtropical zones [6]. Georgian localities. West Georgia: Sokhumi, Batumi [6,29]. Distribution worldwide: Palaearctic Region [4]. Distribution worldwide. Neotropical Region, Australian Region, Genus eLineaspis MacGillivray Palaearctic Region, Oriental Region, Nearctic Region, Afro- 121. Lineaspis striata (Newstead) tropical Region [4,21]. Host plants. Iris foetidissima (Iridaceae) [6], Juniperus sp. 130. Parlatoria oleae (Colvee) (Cupressaceae) (Dry material). Host plants. Olea europaea, Fraxinus excelsior, Jasminum fruti- Georgian localities. East Georgia: Vashlovani National park cans, Syringa vulgaris (Oleaceae), Mespilus germanica, Rosa can- [6,29], Shiraqi, Signagi (Dry material). ina, Crataegus sp., Cotoneaster multiflorus, Eriobotrya japonica, Distribution worldwide. Palaearctic Region [4,21]. Malus orientalis, Malus domestica, Prunus cerasifera, P. spinosa, Genus eLopholeucaspis Balachowsky P. domestica, P. dulcis, P. persica, P. mahaleb, P. cerasus (Rosaceae), 122. Lopholeucaspis japonica (Cockerell) Cistus sp. (Cistaceae), Ulmus minor, U. glabra, Zelkova carpinifolia Host plants. Alnus glutinosa, Carpinus betulus (Betulaceae), (Ulmaceae), Catalpa ovata, C. bignonioides (Bignoniaceae), Coti- Cinnamomum camphora, Laurus nobilis (Lauraceae), Diospyros nus coggygria, Pistacia atlantica (Anacardiaceae), Vaccinium arc- lotus (Ebenaceae), Juglans regia (Juglandaceae), Malus orientalis, tostaphylos (Ericaceae), Diospyros kaki, D. lotus (Ebenaceae), M. domestica, Prunus cerasifera, P. domestica, Pyrus caucasica, Sedum maximum (Crassulaceae), Ribes rubrum (Grossulariaceae), Rosa sp., R. canina, R. chinensis (Rosaceae), Populus alba, P. gra- Rhamnus pallasii (Rhamnaceae), Albizia julibrissin (Fabaceae), cilis, P. deltoides, Salix babylonica (Salicaceae), Citrus sp., C. trifo- Cornus mas (Cornaceae), Lonicera iberica (Caprifoliaceae), Vitis liata (Rutaceae), Quercus sp. (Fagaceae), Camellia sinensis vinifera (Vitaceae), Juglans regia (Juglandaceae), Berberis vulgaris (Theaceae), Tilia rubra (Malvaceae), Vitis vinifera (Vitaceae) [6]. (Berberidaceae), Cycas revoluta (Cycadaceae), Vernicia fordii Georgian localities. Abkhazeti, Tbilisi, Rustavi, Batumi, Kutaisi, (Euphorbiaceae), Tilia rubra (Malvaceae), Viscum album (Santa- Lanchkhuti [6], Supsa, Gulripshi, Kakhaberi (Dry material). laceae) [6], Rosa gallica (Rosaceae), Rhamnus cathartica (Rham- Distribution worldwide. Australian Region, Palaearctic Region, naceae) [19]. Nearctic Region, Oriental Region [4,21]. Georgian localities. Tbilisi, Lagodekhi, Dedoplistskaro, Shiraqi, Genus e Mercetaspis Gomez-Menor Ortega Vashlovani, National Park, Kutaisi, Bolnisi, Sokhumi, Akhali 123. Mercetaspis halimodendronis (Borchsenius and Matesova) Atoni, Kardenakhi [1,6,7,19,29],Zestaphoni (Dry material). Host plants. Fabaceae [4]. Distribution worldwide. Afrotropical Region, Australian Re- Georgian localities. Vashlovani National Park [29]. gion, Palaearctic Region, Nearctic Region, Neotropical Region, Distribution worldwide. Palaearctic Region [4]. Oriental Region [4,21]. 124. Mercetaspis halli (Green) 131. Parlatoria parlatoriae (Sulc) Host plants. Amygdalus sp., Prunus dulcis [6], Pyrus salicifolia Host plants. Picea orientalis, Abies nordmanniana [1], Picea sp. (Rosaceae) [1]. (Pinaceae) [1]. Georgian localities. South- East part of East Georgia, Tbilisi Georgian localities. Tbilisi botanical garden [1], Tsikhisjvari, [1,6]. Manglisi (Dry material). Distribution worldwide. Holarctic Region [4,21]. Distribution worldwide. Palaearctic Region [4,21]. 125. Mercetaspis sureyana (Bodenheimer, 1941) 132. Parlatoria pergandii Comstock Host plants. Astracantha caucasica, Astragalus sp. (Fabaceae) [6]. Host plants. Citrus sinensis, C. limon, C. nobilis (Rutaceae), Dio- Georgian localities. Tbilisi (Karsani), Akhaltsikhe [6]. spyros kaki (Ebenaceae), Prunus laurocerasus (Rosaceae) [6], Distribution worldwide. Palaearctic Region [4,21]. Citrus reticulata (Dry material). Genus e Oceanaspidiotus Takagi Georgian localities. Batumi Botanical Garden, Adjara, Abkhazeti 126. Oceanaspidiotus spinosus (Comstock) [6]. Host plants. Buxus sempervirens (Buxaceae), Laurus nobilis Distribution worldwide. Afrotropical Region, Australian Re- (Lauraceae), Camellia japonica, C. sasanqua (Theaceae), Ilex gion, Palaearctic Region, Nearctic Region, Neotropical Region, colchica (Aquifoliaceae) [6]. Oriental Region [4,21]. Georgian localities. Tbilisi botanical garden [6], Batumi Botan- 133. Parlatoria proteus (Curtis) ical Garden, Makhinjauri, Abkhazeti [29]. Host plants. Camellia japonica (Theaceae), Laurus nobilis (Laur- Distribution worldwide. Palaearctic Region, Neotropical Re- aceae), Eurya japonica (Pentaphylacaceae) [6]. gion, Nearctic Region, Oriental Region, Afrotropical Region, Georgian localities. West Georgia [6]. Australian Region [4,21]. Distribution worldwide. Afrotropical Region, Australian Re- Genus e Odonaspis Leonardi gion, Palaearctic Region, Nearctic Region, Neotropical Region, 127. Odonaspis secreta Cockerell Oriental Region [4,21]. Host plants. Sasa sp., Bambusa sp., Arundinaria sp. (Poaceae) [6]. 134. Parlatoria theae Cockerell Georgian localities. Batumi Botanical Garden [6]. Host plants. Camellia sinensis, C. japonica, C. sasanqua (The- aceae), Diospyros kaki, D. lotus (Ebenaceae), Eurya japonica M. Batsankalashvili et al. / Annals of Agrarian Science 15 (2017) 252e268 261

(Pentaphylacaceae), Viburnum orientale (Adoxaceae), Acer sp., Distribution worldwide. Afrotropical Region, Australian Re- A. japonicum (Sapindaceae), Cistus creticus (Cistaceae), Euon- gion, Palaearctic Region, Nearctic Region, Neotropical Region, ymus japonicus (Celastraceae), Ilex colchica (Aquifoliaceae), Oriental Region [4,21]. Malus domestica, Pyrus communis, P. caucasica, Prunus domestica, 142. Pseudaulacaspis prunicola (Maskell) P. cerasus (Rosaceae), Magnolia grandiflora (Magnoliaceae), Hi- Host plants. Prunus domestica, P. cerasus, P. avium, P. persica biscus syriacus (Malvaceae), Vaccinium arctostaphylos, Rhodo- (Rosaceae), Syringa sp. (Oleaceae), Jasminum sp [29]. dendron luteum (Ericaceae), Cornus mas (Cornaceae), Laurus Georgian localities. Adjara, Sokhumi [29]. nobilis (Lauraceae) [6], Viburnum tinus,V. lantana (Adoxaceae) Distribution worldwide. Cosmopolitan [4]. [1]. Genus e Pseudoparlatoria Cockerell Georgian localities. Batumi Botanical Garden, Tbilisi Botanical 143. Pseudoparlatoria parlatorioides (Comstock) Garden [1,6]. Host plants. Almost poliphagous Distribution worldwide. Australian Region, Nearctic Region, Georgian localities. Apkhazeti [29]. Palaearctic Region, Oriental Region [4,21]. Distribution worldwide. Cosmopolitan [4]. 135. Parlatoria ziziphi Lucas Genus e Rhizaspidiotus (MacGillivray) Host plants. Citrus limon, C. reticulata, C. sinensis, C. aurantium 144. Rhizaspidiotus canariensis (Rutaceae) [6]. Host plants. Euphorbia sp [29]. Georgian localities. East and West Georgia, Tbilisi, Abkhazeti, Georgian localities. South Oseti [29]. Adjara [1,6,29], Tsikhisdziri (Dry matrial). Distribution worldwide. Palaearctic Region [4]. Distribution worldwide. Afrotropical Region, Australian Re- 145. Rhizaspidiotus caraganae (Kiritchenko) gion, Palaearctic Region, Nearctic Region, Neotropical Region, Host plants. Caragana grandiflora (Fabaceae), Rhamnus pallasii Oriental Region [4,21]. (Rhamnaceae) [6,29]. Genus ePinnaspis Cockerell Georgian localities. East Georgia: Tbilisi, Eldari lowland [6,29]. 136. Pinnaspis aspidistrae (Signoret) Distribution worldwide. Palaearctic Region [4]. Host plants. Undetermined species (Orchidaceae) [6], Aspidistra Genus eSalicicola Lindinger elatior (Asparagaceae) [7]. 146. Salicicola archangelskyae (Archangelskaya) Georgian localities. Batumi Botanical Garden, Tbilisi [6,7]. Host plants. Malus domestica [6], Pyrus communis (Rosaceae) Distribution worldwide. Afrotropical Region, Australian Re- (Dry material). gion, Palaearctic Region, Nearctic Region, Neotropical Region, Georgian localities. South-Eastern part of East Georgia [6], Oriental region [4,21]. Tbilisi (Dry material). 137. Pinnaspis strachani (Cooley) Distribution worldwide. Palaearctic Region [4]. Host plants. Ophiopogon japonicus (Asparagaceae) [6]. 147. Salicicola kermanensis (Lindinger) Georgian localities. Batumi Botanical Gardens, Mtsvane Host plants. Populus alba, P. gracilis, P. deltoides (Salicaceae), Kontskhi, Abkhazeti, Adjara, Guria [6,8], Tbilisi, Batumi (Dry Rhamnus pallasii (Rhamnaceae), Malus domestica (Rosaceae) material). [1,6,19]. Distribution worldwide. Afrotropical Region, Australian Re- Georgian localities. Tbilisi, Marneuli, Lagodekhi [1,6,19]. gion, Palaearctic Region, Nearctic Region, Neotropical Region, Distribution worldwide. Afrotropical Region, Australian Re- Oriental Region [4,21]. gion, Palaearctic Region, Nearctic Region, Neotropical Region, Comments. According to Danzig (1993), this species recorded Oriental Region [4,21]. by Borchsenius and Hadzibeyli as Pinnaspis aspidistrae (Signo- Genus e Targionia Signoret ret), this statement needs confirmation. 148. Targionia arthrophyti (Archangelskaya) Genus e Prodiaspis Young in Young and Wang Host plants. Salsola nodulosa (Amaranthaceae) [29]. 138. Prodiaspis tamaricicola (Malenotti) Georgian localities. Vashlovani National Park [29]. Host plants. Tamarix sp. [6], Tamarix laxa (Tamaricaceae) (Dry Distribution worldwide. Palaearctic Region [4,29]. material). 149. Targionia porifera (Borchsenius) Georgian localities. Tbilisi, Shiraqi, Eldari, Pantishara, Sokhumi Host plants. Salsola dendroides (Amaranthaceae) [6]. [1,6]. Georgian localities. East Georgia, Vashlovani, Eldari lowland Distribution worldwide. Palaearctic Region [4,21]. [6,29]. Genus e Pseudaonidia Cockerell Distribution worldwide. Palaearctic Region [4]. 139. Pseudaonidia duplex (Cockerell) 150. Targionia vitis Signoret Host plants. Almost poliphagous [4]. Host plants. Quercus sp., Quercus robur subsp. imeretina (Faga- Georgian localities. Sokhumi [29]. ceae) [6]. Distribution worldwide. Holarctic [29]. Georgian localities. West and East Georgia, Tbilisi Botanical 140. Pseudaonidia paeoniae (Cockerell) Garden, Borjomi, Surami [1,6,19], Makharadze (Dry material). Host plants. Camellia sinensis, C. japonica (Theaceae), Ilex Distribution worldwide. Palaearctic Region, Neotropical Region colchica (Aquifoliaceae) [6]. [4]. Georgian localities. West Georgia [6], Batumi, Peria (Dry Genus e Unaspis McGillivray material). 151. Unaspis euonymi (Comstock) Distribution worldwide. Palaearctic Region, Nearctic Region, Host plants. Viscum album (Santalaceae) on Carpinus sp. Oriental Region [4,21]. (Betulaceae), Euonymus europaeus, E. japonicus, E. latifolius, Genus ePseudaulacaspis MacGillivray E. verrucosus (Celastraceae), Fraxinus excelsior (Oleaceae), Hedera 141. Pseudaulacaspis pentagona (Targioni Tozzetti) colchica, H. helix (Araliaceae) [6,7], Cotinus coggygria (Ana- Host plants. Prunus sp., P. cerasus (Rosaceae), Syringa vulgaris, cardiaceae) [1]. Jasminum sp. (Oleaceae) [6]. Georgian localities. Tbilisi, Batumi, Kobuleti, Makhinjauri, Poti, Georgian localities. West Georgia: Adjara, district of Khelva- , Lagodekhi, Kojori, Manglisi, Borjomi, Zugdidi, Sokhumi, chauri, Gonio [6,29]. Ozurgeti, Gagra [1,6e8], Tbilisi Botanical Garden (Dry material) 262 M. Batsankalashvili et al. / Annals of Agrarian Science 15 (2017) 252e268

Distribution worldwide. Palaearctic Region, Nearctic Region, Georgian localities. East Georgia, Field of Shiraqi, Tbilisi Neotropical Region, Oriental Region [4,21]. botanical garden [6,12,33], Shiraqi, Pantishara (Dry material), Tbilisi (Dabakhana gorge), Benara (Slide-mounted material). Family e Eriococcidae Distribution worldwide. Palaearctic Region [4]. Genus e Eriococcus Targioni Tozzetti 161. Neomargarodes setosus Borchsenius 152. Eriococcus buxi (Boyer de Fonscolombe) Host plants. Cleistogenes serotina, Festuca ovina [10], F. rupicola Host plants. Buxus sempervirens (Buxaceae) [6]. (Poaceae) [6,10]. Georgian localities. Tbilisi, Tbilisi botanical garden [6], Ureki Georgian localities. West Georgia [6], Tbilisi, Akhaltsikhe, Adi- (Dry material). geni, Borjomi, Khashuri, Regions of Gori, Okrokhana, Tskneti, Distribution worldwide. Palaearctic Region [4,21]. Kiketi, Shiraqi [10,33], Pantishara (Dry material). 153. Eriococcus williamsi Danzig Distribution worldwide: Georgia [4]. Host plants. Buxus sp. (Buxaceae) [1]. Genus e Porphyrophora Brandt in Brandt and Ratzeburg Georgian localities. Tbilisi Botanical Garden [1]. 162. Porphyrophora monticola Borchsenius Distribution worldwide. Palaearctic Region [4]. Host plants. Dorycnium graecum, D. pentaphyllum [6,33], Family e Kermesidae Securigera varia (Fabaceae) (Dry material). Genus eKermes Boitard Georgian localities. West Georgia, Racha, Shaori, Chiatura 154. Kermes corticalis (Nasonov) [6,33]. Host plants. Quercus sp. (Fagaceae) [6]. Distribution worldwide. Caucasus (Armenia and Georgia) [4]. Georgian localities. West Georgia [6]. Family eMonophlebidae Distribution worldwide. Mediterranean Region (Italy, Ukraine) Genus e Gueriniella (Targioni Tozzetti) [4]. 163. Gueriniella serratulae (Fabricius) 155. Kermes quercus (Linnaeus) Host plants. Salsola dendroides (Amaranthaceae) [6]. Host plants. Quercus sp., Q. petraea subsp. iberica (Fagaceae) Georgian localities. East Georgia [6]. (Dry material). Distribution worldwide. Palaearctic Region [4]. Georgian localities. Natakhtari, West Georgia [6]. Genus e Icerya Signoret Distribution worldwide. Palaearctic Region [4]. 164. Icerya purchasi Maskell 156. Kermes roboris (Geoffroy in Fourcroy) Host plants. Acacia sp., A. dealbata, Cajanus cajan, Albizia juli- Host plants. Quercus sp., Q. petraea subsp. iberica (Fagaceae) brissin, Amorpha fruticosa (Fabaceae), Citrus sp., C. limon, C. [6,19]. nobilis (Rutaceae), Laurus nobilis (Lauraceae), Vitis vinifera Georgian localities. Tbilisi, Borjomi, Bakuriani [1,6,19]. (Vitaceae) [6,7]. Distribution worldwide. Palaearctic Region, Oriental Region Georgian localities. Tbilisi, Botanical garden of Batumi [1,6,7], [4]. Gulripshi, Tsvermagala (Dry material). Family e Kerriidae Distribution worldwide. Afrotropical Region, Australian Re- Genus e Kerria Oken gion, Nearctic Region, Neotropical Region, Oriental Region, 157. Kerria lacca Chamberlin Palaearctic Region [4,21]. Host plants. Ficus carica (Moraceae), Albizia julibrissin, Cajanus Family e Ortheziidae cajan (Fabaceae), Carya illinoinensis (Juglandaceae), Salix bab- Genus e Arctorthezia Cockerell ylonica (Salicaceae), Diospyros kaki (Ebenaceae), Tilia rubra 165. Arctorthezia cataphracta (Shaw) (Malvaceae), Corylus avellana (Betulaceae), Pterocarya pter- Host plants. Rhododendron sp. (Ericaceae) in association with ocarpa (Juglandaceae) [6]. Sphagnum sp. (Sphagnaceae) [6]. Georgian localities. Gulripshi, Eshera, Tbilisi [1,6]. Georgian localities. East Georgia, Mta-, Omalo [6,34]. Distribution worldwide. Palaearctic Region, Oriental Region, Distribution worldwide. Holarctic Region [4,21]. Neotropical Region [4]. 166. Arctorthezia vardziae Hadzibeyli Family e Kuwaniidae Host plants. Grass, unidentified plant (slide). Genus e Kuwania Cockerell in Fernald Georgian localities. East Georgia, Vardzia [6], [34], 158. Kuwania minuta Borchsenius Vardzia (slide). Host plants. Quercus hartwissiana (Fagaceae) [1,6]. Distribution worldwide. Georgia [4]. Georgian localities. Tbilisi Botanical Garden [1,6]. Genus eNewsteadia (Green) Distribution worldwide. Palaearctic Region [4]. 167. Newsteadia floccosa De Geer Host plants. Unknown. Family e Marchalinidae Georgian localities. Georgia [6]. Genus eMarchalina Vayssiere Distribution worldwide. Palaearctic Region [4]. 159. Marchalina hellenica (Gennadius) Genus eOrthezia Bosc Host plants. Picea orientalis, Abies nordmanniana (Pinaceae) [6]. 168. Orthezia urticae (Linnaeus) Georgian localities. Tsagveri [19], Bakuriani, Borjomi [14], Host plants. Spiraea sp., S. hypericifolia (Rosaceae), Festuca Chiatura, Tsikhisjvari [25], Avadkhara (slide-mounted material), rupicola [7], Agrostis capillaris (Poaceae) [10]. Abkhazeti, Bakhmaro (Dry material). Georgian localities. Georgia [6], Tbilisi, Akhaltsikhe, Gori, Bor- Distribution worldwide. Palaearctic Region, Australian Region jomi, Adigeni, Qeda, Tbilisi botanical garden [1,7,10,34], Vash- [21]. lovani (Dry material). Family e Margarodidae Distribution worldwide. Palaearctic Region [4,21]. Genus e Neomargarodes Green 169. Orthezia shirakensis Hadzibeyli 160. Neomargarodes festucae Archangelskaya Host plants. Artemisia fragrans [6,34], A. szowitziana (Compo- Host plants. Cleistogenes serotina, Festuca rupicola, Stipa sp. sitae) (slide). (Poaceae) [6,12,33]. Georgian localities. East Georgia, Vashlovani [6,34]. M. Batsankalashvili et al. / Annals of Agrarian Science 15 (2017) 252e268 263

Distribution worldwide. Georgia [4]. Host plants. Salvia verticillata (Lamiaceae), Onobrychis cyri [6], Medicago sativa, M. falcata, M. glomerata, Trifolium sp. (Fabaceae) Family e Pseudococcidae [10]. Genus e Antonina Signoret Georgian localities. Tbilisi, Gori [10]. 170. Antonina crawi Cockerell Distribution worldwide. Palaearctic Region [37], Australian Host plants. Phyllostachys nigra, P. bambusoides, Region [4]. P. viridiglaucescens, P. aurea, Phragmites australis (Poaceae) [6,7]. 182. Dysmicoccus walkeri (Newstead) Georgian localities. Mtsvane kontskhi, Batumi Botanical Gar- Host plants. Lolium sp. (Poaceae) [10]. den, Tbilisi [6e8]. Georgian localities. Gardabani, Tbilisi [10]. Distribution worldwide. Australian Region, Palaearctic Region, Distribution worldwide. Palaearctic Region [4]. Nearctic Region, Neotropical Region, Oriental Region [4,21]. Genus e Erimococcus Ezzat Genus eAtrococcus Goux 183. Erimococcus salviae (Hadzibeyli) 171. Atrococcus achilleae (Kiritchenko) Host plants. Salvia verticillata (Lamiaceae) [9]. Host plants. Achillea sp. (Compositae), Salvia sp. (Lamiaceae) [6]. Georgian localities. Meskheti, Vardzia [9], Kodi (Slide). Georgian localities. East Georgia, Tbilisi [6]. Distribution worldwide. Mediterranean Region (Georgia, Distribution worldwide. Palaearctic Region [21]. Turkey) [4]. 172. Atrococcus colchicus (Hadzibeyli) 184. Erimococcus tritubulatus (Kiritshenko) Host plants. Vulpia unilateralis (Poaceae) [15]. Host plants. Salsola dendroides (Amaranthaceae) [6]. Georgian localities. Adjara [15], Batumi Botanical Garden Georgian localities. East Georgia [6]. (slide). Distribution worldwide. Palaearctic Region [4]. Distribution worldwide. Georgia [4]. Genus eFonscolombia Lichtenstein 173. Atrococcus paludinus (Green) 185. Fonscolombia abdita (Borchsenius) Host plants. Trifolium sp. (Fabaceae) [6,10]. Host plants. Poaceae [4]. Georgian localities. East Georgia [6], Tbilisi, Khashuri, Borjomi Georgian localities. Georgia [36]. [10]. Distribution worldwide. Palaearctic Region [4]. Distribution worldwide. Palaearctic Region [4], 186. Fonscolombia amnicola Borchsenius Genus e Balanococcus Williams Host plants. Cleistogenes serotina, Stipa bromoides [6], Avena 174. Balanococcus diminutus (Leonardi) sativa, Lolium sp. (Poaceae) [10]. Host plants. Phormium tenax (Xanthorrhoeaceae) [6]. Georgian localities. Tbilisi [10]. Georgian localities. Batumi Botanical Garden [6]. Distribution worldwide: Palaearctic Region [4]. Distribution worldwide. Australian Region, Palaearctic Region, 187. Fonscolombia europaea (Newstead) Nearctic Region [4]. Host plants. Dactylis glomerata (Poaceae) [6]. Genus e Ceroputo Sulc Georgian localities. East Georgia, Signagi [6]. 175. Ceroputo pilosellae (Kiritshenko) Distribution worldwide. Palaearctic Region [4]. Host plants. Fragaria vesca (Rosaceae) [6]. 188. Fonscolombia tomlinii Newstead Georgian localities. West Georgia, Banoja [6]. Host plants. Agrostis capillaris (Poaceae) [6]. Distribution worldwide. Palaearctic Region [4]. Georgian localities. East Georgia, Bakuriani [6,10]. Genus eChaetococcus Maskell Distribution worldwide. Palaearctic Region [4]. 176. Chaetococcus phragmitis (Marchal) Genus eHeliococcus Sulc Host plants. Phragmites australis (Poaceae) [6].  189. Heliococcus bohemicus (Sulc) Georgian localities. Tbilisi [7], Shiraqi (Dry material). Host plants. Rubus sp. (Rosaceae) [6]. Distribution worldwide. Holarctic Region [4]. Georgian localities. West Georgia [6]. 177. Chaetococcus sulcii (Green) Distribution worldwide: Palaearctic Region [4]. Host plants. Festuca rupicola [6], F. ovina (Poaceae) [10]. 190. Heliococcus caucasicus Borchsenius Georgian localities. Gori, Borjomi, Bakuriani, Akhaltsikhe, Adi- Host plants. Achillea millefolium (Compositae) [6]. geni [10], Tbilisi. Georgian localities. East Georgia [6]. Distribution worldwide. Palaearctic [4]. Distribution worldwide. Palaearctic Region [4]. Genus e Coccidohystrix Lindinger 191. Heliococcus glacialis (Newstead) 178. Coccidohystrix artemisiae (Kiritshenko) Host plants. Lathyrus pratensis (Fabaceae) [6]. Host plants. Asteraceae [4]. Georgian localities. Qeda [10]. Georgian localities. Vashlovani [35]. Distribution worldwide. Palaearctic Region [4,21]. Distribution worldwide. Palaearctic [4]. 192. Heliococcus destructor Borchsenius Genus e Coccura (Lichtenstein) Host plants. Robinia pseudoacacia (Fabaceae) [6]. 179. Coccura circumscripta (Kiritchenko) Georgian localities. East Georgia, Tbilisi [6,8]. Host plants. Atraphaxis caucasica (Polygonaceae) [6]. Distribution worldwide. Palaearctic Region [4]. Georgian localities. East Georgia, Tbilisi [6]. Genus eHeterococcus Ferris Distribution worldwide. Palaearctic Region [4]. 193. Heterococcus nudus (Green) 180. Coccura comari (Kunow) Host plants. Undetermined species (Poaceae) [6]. Host plants. Crassulaceae, Fabaceae, Rosaceae [4]. Georgian localities. East Georgia, Signagi [6,10]. Georgian localities. Georgia [36]. Distribution worldwide. Holarctic Region [4,21]. Distribution worldwide. Palaearctic Region [4]. Genus e Kiritshenkella Borchsenius Genus e Dysmicoccus Ferris 194. Kiritshenkella sacchari Green 181. Dysmicoccus angustifrons (Hall) Host plants. Saccharum ravennae (Poaceae) [6]. 264 M. Batsankalashvili et al. / Annals of Agrarian Science 15 (2017) 252e268

Georgian localities. Pantishara [6,40], Shiraqi (slide). Host plants. Teucrium polium (Lamiaceae) [6]. Distribution worldwide. Palaearctic Region, Oriental Region Georgian localities. East Georgia [6]. [4]. Distribution worldwide. Palaearctic Region [21]. Genus e Longicoccus Danzig Genus e Pelionella Kaydan 195. Longicoccus clarus (Borchsenius) 207. Pelionella manifecta (Borchsenius) Host plants. Festuca rupicola (Poaceae) [6], Alopecurus textilis Host plants. Cirsium arvense (Compositae) [6]. (Poaceae) [9]. Georgian localities. East Georgia [6]. Georgian localities. East Georgia, Shiraqi [6,9]. Distribution worldwide. Palaearctic Region [4]. Distribution worldwide. Palaearctic Region [4]. 208. Pelionellatritubulatus (Kiritshenko) Genus eMetadenopus Sulc Host plants. Salsola dendroides (Amaranthaceae) [6]. 196. Metadenopus festucae Sulc Georgian localities. East Georgia [6]. Host plants. Festuca rupicola (Poaceae) [6]. Distribution worldwide. Palaearctic Region [4]. Georgian localities. Borjomi, Akhaltsikhe, Adigeni [10]. Genus e Phenacoccus Cockerell Distribution worldwide. Palaearctic Region [4]. 209. Phenacoccus aceris (De Geer) Genus eMirococcopsis Borchsenius Host plants. Fraxinus excelsior (Oleaceae), Prunus persica, 197. Mirococcopsis nagyi Kozar P. domestica, P. armeniaca, Cydonia oblonga, Malus orientalis, Host plants. Stipae [37]. M. domestica (Rosaceae), Quercus hartwissiana, Q. petraea subsp. Georgian localities. East Georgia [37]. iberica, Q. longipes (Fagaceae), Acer sp. (Sapindaceae), Carpinus Distribution worldwide. Palaearctic Region [4]. betulus (Betulaceae), Hedera colchica, H. pastuchovii (Araliaceae) 198. Mirococcopsis subterranea (Newstead) [6,7,15,19]. Host plants. Undetermined species (Poaceae) [6]. Georgian localities. Adigeni, Akhaltsikhe, Bakuriani, Lagodekhi Georgian localities. East Georgia [6]. national park, Tbilisi botanical garden, Telavi, Tbilisi, Zestafoni Distribution worldwide. Palaearctic Region [4]. [6,7,19], Tbilisi Botanical Garden [15]. Genus e Mirococcus (Borchsenius) Distribution worldwide. Palaearctic Region [21]. 199. Mirococcus carthalinus Hadzibeyli 210. Phenacoccus daganiae (Bodenheimer) Host plants. Alopecurus textilis (Poaceae) [9]. Host plants. Cleistogenes serotina, Phleum sp., Bromus bieber- Georgian localities. East Georgia [9]. steinii, Bromus sp. (Poaceae) [6,10]. Distribution worldwide. Georgia [4]. Georgian localities. East Georgia, Tbilisi, Mtskheta, Oqrokhana 200. Mirococcus inermis (Hall, 1925) [6,10]. Host plants. Achillea pubescens (Compositae) [6]. Distribution worldwide. Palaearctic Region [4,21]. Georgian localities. Shiraqi [6]. 211. Phenacoccus interruptus Green Distribution worldwide. Palaearctic Region [4]. Host plants. Phalaris sp., Agropyron sp., Bromus biebersteinii Genus e Neotrionymus Borchsenius (Poaceae) [10]. 201. Neotrionymus monstatus Borchsenius Georgian localities. Borjomi, Akhaltsikhe and Khashuri region Host plants. Phalaris arundinacea [6,15], Phragmites australis [10]. (Poaceae) [15]. Distribution worldwide. Palaearctic Region [4]. Georgian localities. East Georgia [6,15], Shiraqi (slide). 212. Phenacoccus montanus (Hadzibeyli) Distribution worldwide. Palaearctic Region [4]. Host plants. Picea orientalis (Pinaceae) [6]. Genus e Nipaecoccus Sulc Georgian localities. Manglisi (Dry material), Bakuriani Abkha- 202. Nipaecoccus nipae (Maskell) zeti, Avadkhara, Kvatakhevi, Manglisi, Sapara, Tsikhisjvari Host plants. Phoenix sp., Chamaerops sp., Butia capitata (Areca- (Slide-mounted material). ceae), Annona sp. (Annonaceae), Carludovica sp. (Cyclanthaceae), Distribution worldwide. Georgia [4]. Persea sp. (Lauraceae) [6]. 213. Phenacoccus interruptus Green Georgian localities. Sokhumi, Batumi and Tbilisi Botanical Host plants. Phalaris sp., Agropyron sp., Bromus biebersteinii Gardens [6]. (Poaceae) [10]. Distribution worldwide. Australian Region, Palaearctic Region, Georgian localities. Borjomi, Akhaltsikhe and Khashuri region Nearctic Region, Neotropical Region, Oriental Region [4,21]. [10]. Genus e Peliococcopsis (Borchsenius) Distribution worldwide. Palaearctic Region [21]. 203. Peliococcopsis priesneri (Laing) 214. Phenacoccus phenacoccoides (Kiritshenko) Host plants. Undetermined species (Poaceae) [6]. Host plants. Amaranthaceae, Asteraceae, Poaceae [4]. Georgian localities. East Georgia, Signagi [6]. Georgian localities. West Georgia [6,36]. Distribution worldwide. Palaearctic Region [4]. Distribution worldwide. Palaearctic Region [4]. Genus e Peliococcus Borchsenius 215. Phenacoccus piceae Low 204. Peliococcus marrubii (Kiritchenko) Host plants. Pinaceae and Taxaceae [4], Picea orientalis [36] Host plants. Hyssopus officinalis (Lamiaceae) [6]. Georgian localities. Bakuriani, Abkhazeti [19,36]. Georgian localities. Georgia [6]. Distribution worldwide. Palaearctic [4]. Distribution worldwide. Palaearctic Region [4]. 216. Phenacoccus pumilus Kiritshenko 205. Peliococcus morrisoni Kiritshenko Host plants. Medicago sativa, M. caerulea, M. falcata, Host plants. Carpinus betulus, C. orientalis, Corylus colchica M. polychroa, Pisum sativum, Trifolium arvense, T. medium, (Betulaceae) [6]. T. repens, Onobrychis cyri, O. viciifolia (Fabaceae) [10], Teucrium Georgian localities. Tbilisi Botanical Garden, Lagodekhi [6,19]. polium (Lamiaceae) [7], Melilotus sp. (Fabaceae), Daucus carota Distribution worldwide. Palaearctic Region [4]; (Apiaceae), Chrysanthemum indicum (Compositae) [6]. 206. Peliococcus turanicus (Kiritshenko) M. Batsankalashvili et al. / Annals of Agrarian Science 15 (2017) 252e268 265

Georgian localities. Tbilisi botanical garden, Tbilisi (Makhata, Distribution worldwide. Palaearctic Region, Australian Region, Mtatsminda), Gori, Khashuri, Borjomi, Lagodekhi, Qedi, Khulo Nearctic Region, Neotropical Region, Oriental Region, Afro- [7,10]. tropical Region [4,21]. Distribution worldwide. Palaearctic Region [4,21]. 226. Pseudococcus maritimus (Ehrhorn) 217. Phenacoccus querculus (Borchsenius) Host plants. Citrus japonica, C. limon, C. nobilis (Rutaceae) [7]. Host plants. Quercus sp. (Fagaceae) [6]. Georgian localities. Tbilisi [6,7]. Georgian localities. East Georgia [6]. Distribution worldwide. Nearctic Region, Neotropical Region, Distribution worldwide. Palaearctic Region [4]. Palaearctic Region [4,21]. 218. Phenacoccus transcaucasicus Hadzibeyli 227. Pseudococcus viburni (Signoret) Host plants. Lonicera iberica (Caprifoliaceae) [6,38]. Host plants. Diospyros kaki (Ebenaceae), Citrus reticulata, Georgian localities. Tbilisi [6,38], Shiraqi, Vashlovani, Tbilisi C. limon, C. aurantium, C. paradisi (Rutaceae), Vernicia cordata, V. botanical garden, Pantishara (slide) fordii (Euphorbiaceae), Laurus nobilis (Lauraceae), Prunus laur- Distribution worldwide. Palaearctic Region [4]. ocerasus (Rosaceae), Vitis vinifera (Vitaceae), Albizia sp. (Faba- Genus e Planococcus Ferris ceae) [6]. 219. Planococcus citri (Risso) Georgian localities. Gulripshi [6], Makhinjauri (Dry material). Host plants. Citrus sp. (Rutaceae) [6]. Distribution worldwide. Afrotropical Region, Australian Re- Georgian localities. West Georgia [6]. gion, Nearctic Region, Neotropical Region, Palaearctic Region, Distribution worldwide. Afrotropical Region, Australian Re- Oriental Region [4,21]. gion, Palaearctic Region, Nearctic Region, Neotropical Region, Genus e Rhodania Goux Oriental Region [4,21]. 228. Rhodania festucae Hadzibeyli 220. Planococcus ficus (Signoret) Host plants. Festuca rupicola (Poaceae) (slide). Host plants. Vitis vinifera (Vitaceae), Ficus carica (Moraceae), Georgian localities. Shiraqi, Meskheti, and Punica granatum (Lythraceae), Platanus orientalis (Platanaceae), Mountain ridge [12], Sapara (slide). Solanum lycopersicum, S. tuberosum (Solanaceae), Beta vulgaris Distribution worldwide. Georgia [4]. (Amaranthaceae) [1]. 229. Rhodania porifera Goux Georgian localities. Gulripshi [6], , Tbilisi. Host plants. Festuca rupicola (Poaceae) [6]. Distribution worldwide. Afrotropical Region, Palaearctic Re- Georgian localities. Tbilisi, Akhaltsikhe, Akhalqalaqi [7,10], gion, Neotropical Region, Oriental Region, Nearctic Region Tabatskuri Lake (slide). [4,21]. Distribution worldwide. Palaearctic Region [4,21]. 221. Planococcus vovae (Nasonov) Genus e Ritsemia Lichtenshtein Host plants. Juniperus foetidissima [6], J. oxycedrus, Cupressus 230. Ritsemia pupifera Lichtenshtein sempervirens (Cupressaceae) [19]. Host plants. Ulmus sp. [6], U. minor [39], U. laevis (Ulmaceae) Georgian localities. East Georgia [6], Tbilisi, Mtskheta [19]. (Dry material). Distribution worldwide. Palaearctic Region, Nearctic Region, Georgian localities. East Georgia [6], Gardabani [39], Tbilisi Neotropical Region [4,21]. (Dry material, slide), Shiraqi (Slide). Genus e Polystomophora Borchsenius Distribution worldwide. Palaearctic Region [4,21]. 222. Polystomophora ostiaplurima (Kiritchenko) Genus e Spilococcus Ferris Host plants. Acer campestre (Sapindaceae) [6]. 231. Spilococcus halli (McKenzie and Williams) Georgian localities. Georgia [6]. Host plants. Sacharum sp. (Poaceae) (slide). Distribution worldwide. Palaearctic Region [4]. Georgian localities. Pantishara gorge [8], Shiraqi (slide). Genus e Pseudococcus Westwood Distribution worldwide. Palaearctic Region [4,21]. 223. Pseudococcus calceolariae (Maskell) 232. Spilococcus mamillariae (Bouche) Host plants. Citrus sp. (Rutaceae) [6]. Host plants. Rebutia sp., Phyllocactus sp., Mammillaria sp. (Cac- Georgian localities. Sokhumi, Gagra, Gudauta, Abkhazeti [6]. taceae) [6]. Distribution worldwide. Palaearctic Region, Australian Region, Georgian localities. Tbilisi [6]. Nearctic Region, Neotropical Region, Oriental Region, Afro- Distribution worldwide: Australian Region, Neotropicalal Re- tropical Region [4,21]. gion, Nearctic Region, Palaearctic Region [4]. 224. Pseudococcus comstocki (Kuwana) 233. Spilococcus furcatispinus (Borchsenius) Host plants. Morus alba, Ficus carica (Moraceae), Albizia juli- Host plants. Lolium rigidum, Elymus repens (Poaceae) [6]. brissin (Fabaceae) [6]. Georgian localities. East Georgia, Tbilisi [6]. Georgian localities. East and West Georgia [6]. Distribution worldwide. Palaearctic Region [4]. Distribution worldwide. Afrotropical Region, Australian Re- 234. Spilococcus vashlovanicus (Danzig) gion, Palaearctic Region, Nearctic Region, Neotropical Region, Host plants. Amaranthaceae, Asteraceae, Brassicaceae [4]. Oriental Region [4,21]. Georgian localities. Georgia [37]. 225. Pseudococcus longispinus (Targioni Tozzetti) Distribution worldwide. Turkey, Georgia [4]. Host plants. Cyclamen sp. (Primulaceae), Chamaedorea elegans, Genus e Trabutina Marchal C. elatior, Metroxylon sp., Phoenix canariensis, Trachycarpus sp. 235. Trabutina crassispinosa Borchsenius (Arecaceae), Trichocereus sp. (Cactaceae), Cycas revoluta (Cyca- Host plants. Tamarix smyrnensis (Tamaricaceae) [6]. daceae), Begonia sp. (Begoniaceae), Orchis sp. (Orchidaceae), Georgian localities. Shiraqi [6]. Citrus sp. (Rutaceae), Asparagus sp. (Asparagaceae) [6], unde- Distribution worldwide. Palaearctic Region [4]. termined species (Cactaceae), Urtica sp. (Urticaceae) [7]. 236. Trabutina mannipara Hemprich and Ehrenberg Georgian localities. Tbilisi and Batumi Botanical Garden [6e8]. Host plants. Tamarix sp., T. smyrnensis (Tamaricaceae) [6]. Tbilisi [Dry material]. Georgian localities. South-Eastern part of East Georgia [6]. 266 M. Batsankalashvili et al. / Annals of Agrarian Science 15 (2017) 252e268

Distribution worldwide. Palaearctic Region [4,21]. Distribution worldwide. Palaearctic Region [4]. Genus e Trionymus Berg 237. Trionymus aberrans Goux Host plants. Undetermined species (Poaceae) [6], Elymus repens Discussion (Poaceae) Georgian localities. East Georgia [6]. We did not include species which identification was only to the Distribution worldwide. Palaearctic Region [4]. genus level as a morphospecies in the list. Also we could not allo- 238. Trionymus diminutus (Leonardi) Synonym: Balanococcus cate the following species: Pseudococcus phlei Borchsenius [10], diminutus Lepidosaphes caucasicus (nomen nudum) [4], Phenacoccus pseudo- Host plants. Phormium tenax (Xanthorrhoeaceae) [6]. pumilus (slide in the collection, needs reidentification), Rhizococcus Georgian localities. Batumi Botanical Garden [6]. bodbiensis [9], Rhizecus agropyri (Dry material in the collection) (we Distribution worldwide. Australian Region, Palaearctic Region, think that Rhizecus agropyri must be Anophococcus agropyri, Nearctic Region [4]. however it needs confirmation according material), this can be a 239. Trionymus isfarensis (Borchsenius) spelling error by Hadzibeyli and the species can belong to Ano- Host plants. Brachypodium sp. (Poaceae) [6]. phococcus (Rhizococcus) agropyri, Mirococcus tbilisiensis (slide in the Georgian localities. West Georgia [6]. collection, needs reidentification), Diaspidiotus buxii, Lepidosaphes Distribution worldwide. Palaearctic Region [4]. colchicum (nomen nudum) [4], Peliococcus tbilisiensis (nomen 240. Trionymus newsteadi (Green) nudum) [4], Asterolecanium repugnans [4], and Phenacoccus carpini Host plants. Fagus orientalis (Fagaceae) [6]. (nomen nudum) [4]. Georgian localities. Georgia [6]. Distribution worldwide. Palaearctic Region [4]. 241. Trionymus perrisii (Signoret) Conclusion Host plants. Dactylis glomerata (Poaceae) [6]. Georgian localities. East Georgia, Batumi Botanical garden [6]. Our study revealed that 242 species are recorded in Sakartvelo Distribution worldwide. Palaearctic Region [4]. (Georgia). Anophococcus agropyri was found in the collection of Z. Hadzibeyli and it is the first record from Sakartvelo of the species. Family e Putoidae The paper represents second summarizing paper on different insect Genus ePuto Signoret superfamilies for this journal, previously was done Apoidea [41]. 242. Puto caucasicus Hadzibeyli We presented here species numbers according families (Fig. 1) Host plants. Picea orientalis [6,40], Abies nordmanniana (Pina- and their zoogeographical distribution (Fig. 2). Most species ceae) [40]. constituting fauna of scale insects of Sakartvelo (Georgia) belong to Georgian localities. Abastumani, Avadkhara, Bakhmaro, Palaearctic species, with 119 species (47%), cosmopolitan species Bakuriani, Borjomi, Kurtskhana, Meskheti, Racha, Shovi, Tsi- are on the second place with 87 species (37%) (Fig. 2). Caucasian khisjvari, Upper Imereti, Upper Adjara [40]. endemics are 3 species (1%): Eulecanium transcaucasicum,

Fig. 1. Species number according scale insect families. M. Batsankalashvili et al. / Annals of Agrarian Science 15 (2017) 252e268 267

Fig. 2. Zoogeographical distribution of scale insect species.

Pulvinaria juglandii, Porphyrophora monticola and 9 species (4%): [17] G. Japoshvili, N. Gabroshvili, N. Chakvetadze, Eriococcus spurius (Hemiptera, Anophococcus adzharicus, Arctorthezia vardzia, Atrococcus colchicus, Eriococcidae) and its natural enemies in Tbilisi, in: Proceedings of the XI In- ternational Symposium on Scale Insect Studies, 2008, pp. 169e170. Neomargarodes setosus, Orthezia shirakensis, Phenacoccus montanus, [18] F. Kozar, M.B. Kaydan, Z.K. Benedicty, E. Szita, Acantococcidae and Related Pulvinaria vini, Rhodania festucae, Mirococcus carthalinus Hadzibeyli Families of the Palaearctic Region, Hungarian Academy of Sciences, Budapest, are Georgian endemics. 2013. [19] Z. Hadzibeyli, Materials for Coccids (Coccoidea) distributed in eastern Geor- gian forests fauna, Proc. Inst. Plant Prot. Ga. SSR 7 (1950) 255e263 (in References Georgian). [20] M. Kosztarab, F. Kozar, Scale Insects of Central Europe, Akademiai Kiado, Budapest, 1988. [1] G. Japoshvili, N. Gabroshvili, B. 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