231 Genus Euphaedra Huebner
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AFROTROPICAL BUTTERFLIES 17th edition (2018). MARK C. WILLIAMS. http://www.lepsocafrica.org/?p=publications&s=atb Genus Euphaedra Hübner, [1819] In: Hübner, [1816-[1826]. Verzeichniss bekannter Schmettlinge 39 (432 + 72 pp.). Augsburg. Type-species: Papilio cyparissa Cramer, by subsequent designation (Scudder, 1875. Proceedings of the American Academy of Arts and Sciences 10: 172 (91-293).). = Najas Hübner, 1807 in Hübner, [1806-19]. Sammlung exotischer Schmetterlinge 1: pl. 60 ([vi] pp., 213 pls.). Augsberg. Type-species: Najas themis Hübner, by monotypy. [Placed on the Official List of Rejected and Invalid Names in Zoology; Opinion 789, 1966. Bulletin of Zoological Nomenclature 23: 213-220.]. = Romaleosoma Blanchard, 1840. Histoire naturelle des Insectes. Orthoptères, Néuroptères, Hémiptères, Hyménoptères, Lépidotères et Diptères 3: 448 (672 pp.). Paris. Type-species: Papilio eleus Drury, by original designation. = Romalaeosoma Röber, 1888 in Staudinger and Schatz, 1884-8. Exotischer Schmetterlinge 1: 162 (333 pp.). Bayern. [An unjustified emendation of Romaleosoma Blanchard.] The genus Euphaedra belongs to the Family Nymphalidae Rafinesque, 1815; Subfamily Limenitidinae Behr, 1864; Tribe Adoliadini Doubleday, 1845; Subtribe Bebearina Hemming, 1960. The other genera in the Subtribe Bebearina in the Afrotropical Region are Evena, Aterica, Pseudargynnis, Cynandra, Eutyphura, Euryphaedra, Euryphurana, Harmilla, Euriphene, Bebearia, Euptera and Pseudathyma. Euphaedra (Foresters) is the most speciose genus in the Afrotropical Region and comprises 245 described species. Generic review by Hecq, 1997. Key to subgenera in Hecq, 1997 (Union des Entomologistes Belges, Tervuren: 1-117). Order of subgenera and species, below, as arranged in this publication. Relevant literature: Hecq, 2006b [New species]. Oremans, 2006 [New species]. Molleman & Hecq, 2005 [Host plants and early stages]. Amiet, 2004 [Diagnostic characters]. Subgenus Proteuphaedra Hecq, 1976 Lambillionea 76: 53 (38-48, 53-63). Type-species: Romalaeosoma [sic] luperca Hewitson, by original designation. 1 *Euphaedra (Proteuphaedra) rubrocostata (Aurivillius, 1897) Euryphene rubrocostata Aurivillius, 1897. Öfversigt af Kongl. Vetenskaps-Akademiens Förhandlingar. Stockholm 54: 279 (279- 286). Type locality: [Democratic Republic of Congo]: “Congogebiet: Beni-Bendi”. Distribution: Cameroon, Gabon, Central African Republic, Democratic Republic of Congo. Early stages: Nothing published. Larval food: Nothing published. Euphaedra (Proteuphaedra) rubrocostata rubrocostata (Aurivillius, 1897) Euryphene rubrocostata Aurivillius, 1897. Öfversigt af Kongl. Vetenskaps-Akademiens Förhandlingar. Stockholm 54: 279 (279- 286). Type locality: [Democratic Republic of Congo]: “Congogebiet: Beni-Bendi”. Distribution: Cameroon, Central African Republic, Democratic Republic of Congo. Specific localities: Central African Republic – Dzanga (Noss, 1998). Democratic Republic of Congo – Beni-Bendi (TL). Euphaedra (Proteuphaedra) rubrocostata generosa Hecq, 1987 Euphaedra rubrocostata generosa Hecq, 1987. Lambillionea 87: 105 (103-105). Type locality: Democratic Republic of Congo: “Mbandaka”. First description of male by Oremans, 1996 (Entomologia Africana 1: 14). Distribution: Gabon, Democratic Republic of Congo. Specific localities: Gabon – Camp Nouna (Vande weghe, 2010). Democratic Republic of Congo – Mbandaka (TL). *Euphaedra (Proteuphaedra) aubergeri Hecq, 1977 Auberger’s Forester Euphaedra aubergeri Hecq, 1977. Lambillionea 76: 82 (82-84). Type locality: Ivory Coast: “Danane, Côte d’Ivoire”. Distribution: Ivory Coast (north-west). Specific localities: Ivory Coast – Danane (TL). Habitat: Forest. Early stages: Nothing published. Larval food: Nothing published. 2 *Euphaedra (Proteuphaedra) symphona Bethune-Baker, 1908 Euphaedra symphona Bethune-Baker, 1908. Annals and Magazine of Natural History (8) 2: 472 (469-482). Type locality: [Democratic Republic of Congo]: “Beni-Mawambe, Congo Free State”. Distribution: Gabon, Democratic Republic of Congo (central, Uele, north Kivu). Early stages: Nothing published. Larval food: Nothing published. Euphaedra (Proteuphaedra) symphona symphona Bethune-Baker, 1908 Euphaedra symphona Bethune-Baker, 1908. Annals and Magazine of Natural History (8) 2: 472 (469-482). Type locality: [Democratic Republic of Congo]: “Beni-Mawambe, Congo Free State”. Distribution: Gabon, Democratic Republic of Congo (central, Uele, north Kivu). Specific localities: Gabon – Bitam (Vande weghe, 2010); camp Nouna (Vande weghe, 2010). Democratic Republic of Congo – Beni-Mawambe (TL). medonoides Wichgraf, 1913 (as sp. of Euphaedra). Internationale Entomologische Zeitschrift 7: 9 (9-10, 13- 14, 21-22). Democratic Republic of Congo: “Kongo”. Euphaedra (Proteuphaedra) symphona affabilis Hecq, 1996 Euphaedra symphona affabilis Hecq, 1996. Lambillionea 96 (1) (Tome I): 126 (126-128). Type locality: [Democratic Republic of Congo]: “Zaïre, Eala, VIII.1935. (J. Ghesquière).” Holotype in MRAC, Tervuren. Distribution: Democratic Republic of Congo. Specific localities: Democratic Republic of Congo – Eala (TL). *Euphaedra (Proteuphaedra) adolfifriderici Schultze, 1920 Euphaedra adolfifriderici Schultze, 1920. Ergebnisse der Zweiten Deutschen Zentral-Afrika-Expedition 1 (14): 746 (639-829). Type locality: Cameroon: “Süd-Kamerun: Endendum”. Distribution: Cameroon, Gabon, Democratic Republic of Congo (south-west). Specific localities: Cameroon – Endendum (TL). Gabon – Kinguele (Vande weghe, 2010); Tchimbele (Vande weghe, 2010); Waka (Vande weghe, 2010); Mikongo (Vande weghe, 2010); Bitam (Vande weghe, 2010); Kongou (Vande weghe, 2010); Langoue (Vande weghe, 2010); camp Nouna (Vande weghe, 2010); Bakouaka (Vande weghe, 2010). Early stages: Nothing published. Larval food: Nothing published. 3 *Euphaedra (Proteuphaedra) luperca (Hewitson, [1864]) Nigerian Blue Forester Romalaeosoma [sic] luperca Hewitson, [1864] in Hewitson, [1862-6]. Illustrations of new species of exotic butterflies 3: 33 (124 pp.). London. Type locality: Nigeria: “Calibar”. Distribution: Nigeria (south), Cameroon (west). Specific localities: Nigeria – Calabar (TL); Ore (Larsen, 2005a); Okomu (Larsen, 2005a); Sapoba (Larsen, 2005a); Port Harcourt (Larsen, 2005a); Cross River loop (Oban Hills) (Larsen, 2005a). Cameroon – Korup (Larsen, 2005a). Habitat: Forest. Habits: An uncommon butterfly that is also very wary (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published. *Euphaedra (Proteuphaedra) fucora Hecq, 1979 Oban Blue Forester Euphaedra luteofasciata fucora Hecq, 1979. Lambillionea 79: 48 (47-48). Euphaedra fucora Hecq, 1979. Ackery et al., 1995: 409. Type locality: Nigeria: “Nigeria”. Distribution: Nigeria (south-east), Cameroon. Specific localities: Nigeria – Okwangwo (Larsen, 2005a); Oban Hills (Larsen, 2005a). Cameroon – Korup (Larsen, 2005a). Habitat: Forest in good condition (Larsen, 2005a). Habits: A scarce species that is wary but comes to feed on fallen fruit (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published. Note: Raised from Euphaedra luteofasciata fucora (according to Ackery et al., 1995: 409) or from Euphaedra luperca fucora (according to the Zoological Records). *Euphaedra (Proteuphaedra) luteofasciata Hecq, 1979 Euphaedra luteofasciata Hecq, 1979. Lambillionea 79: 48 (47-48). Type locality: Cameroon: “Cameroun”; Democratic Republic of Congo: “Mayumbe”. Distribution: Cameroon, Gabon, Democratic Republic of Congo (Mayumbe). Specific localities: Cameroon – Bipindi (Bartel, 1905). Gabon – Lake Azingo (Rothschild, 1918); Kinguele (Vande weghe, 2010); Tchimbele (Vande weghe, 2010); Lake Evaro (Vande weghe, 2010); Waka (Vande weghe, 2010); Lope N.P. (Vande weghe, 2010); Bitam (Vande weghe, 2010); Ipassa (Vande weghe, 2010); Langoue (Vande weghe, 2010); Kongou (Vande weghe, 2010); camp Nouna (Vande weghe, 2010). Democratic Republic of Congo – Mayumbe (TL). Early stages: Nothing published. Larval food: Nothing published. 4 luteofasciata Bartel, 1905 (as ab. of Euphaedra luperca). Novitates Zoologicae 12: 143 (129-152). Cameroon: “Süd-Kamerun (Bipindi, Urwaldweg)”. variegata Aurivillius, 1912 in Seitz, 1908-25 (as ab. of Euphaedra luperca). Die Gross-Schmetterlinge der Erde, Stuttgart (2) 13 Die Afrikanischen Tagfalter: 320 (614 pp.). “Congo”. suffusa Rothschild, 1918 (as ab. of Euphaedra luperca). Novitates Zoologicae 25: 338 (338-345). Gabon: “Lake Azingo, Gaboon”. variegata d’Abrera, 1980 (as ssp. of Euphaedra luperca). Butterflies of the Afrotropical region 320 (593 pp.). Melbourne. Gabon/Democratic Republic of Congo: “Gabon and south-western Zaire”. *Euphaedra (Proteuphaedra) marginalis Hecq, 1979 Euphaedra luperca marginalis Hecq, 1979. Lambillionea 79: 47 (47-48). Euphaedra luperca marginalis Hecq, 1979. Ackery et al., 1995: 408. Euphaedra marginalis Hecq, 1979. Hecq, 1997. Union des Entomologistes Belges, Tervuren: 22 (1-117). Type locality: Democratic Republic of Congo. Distribution: Democratic Republic of Congo (Ubangi, Mongala, Uele). Early stages: Nothing published. Larval food: Nothing published. *Euphaedra (Proteuphaedra) lupercoides Rothschild, 1918 Euphaedra lupercoides Rothschild, 1918. Novitates Zoologicae 25: 338 (338-345). Type locality: [Democratic Republic of Congo]: “Luebo, Kassai River”. Distribution: Congo, Central African Republic, Democratic Republic of Congo. Specfic localities: Central African Republic – Dzanga (Noss, 1998). Democratic Republic of Congo – Luebo, Kassai River (TL). Early stages: Nothing published. Larval food: Nothing published.