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https://doi.org/10.24199/j.mmv.1977.38.01 4 April 1977
A TAXONOMIC AND ZOOGEOGRAPHIC STUDY OF TASMANIAN CADDIS-FLIES (INSECTA: TRICHOPTERA) By Arturs Neboiss
Curator of Insects, National Museum of Victoria
Abstract The systematics of the Trichoptera of Tasmania (excluding the two island groups in Bass Strait) is examined. For easier assessment the island is subdivided into seven provinces, the boundaries of which are based on a combination of landforms, geological features, vegetation and climatic conditions. About 18 000 specimens, most of which were collected by the author between the years 1965 and 1974, were examined. There are now 157 species recognized in Tasmania, which are placed in 66 genera and 21 families, of which 16 genera and 64 species are described as new. Due to changes in higher classification, several new family names appear, and three families are recorded from the state for the first time—families Stenopsychidae, Kokiriidae and Oeconesidae, the latter not being known from the Australian mainland. Of the total caddis-fly fauna, 74% of the species appear to be endemic to Tasmania. The proportion of endemic species varies throughout the island, being highest in the two western provinces (73%), but lowest in the eastern provinces (50%). The fauna shows a high proportion of trans-antarctic elements, exhibiting close relationships with New Zealand and South American species.
description of a mollusc, which proved to be Introduction an insect living in a snail-like case, made from The objective of this study is to determine sand grains (Dyer, 1879). Undoubtedly, this the composition of the Tasmanian trichopteran note refers to the larvae of the family Helicop- fauna, which until recent years was very poorly sychidae. It was not until Mosely added further known. Undoubtedly, further intensive collect- species between 1933 and 1936, that the num- ing will reveal more, yet unknown species and ber of species increased and reached the total the knowledge of the distribution of the pre- of 18, placed in seven families. In the publica- sently known species will be greatly expanded. tion by Mosely and Kimmins (1953), the Information on immature stages is limited, but number of species was increased to 58, but once studied in detail, will clarify many out- with other publications, which appeared in the standing questions on the relationships between following years (Neboiss, 1959, 1962 and Jac- species, genera and families, particularly those quemart, 1965a and b), the total number of with a trans-antarctic distribution. species recorded rose, and stood at 85 at the The first three species of Trichoptera from time when this study was started. Tasmania were recorded by Francis Walker Most of the material was collected by the (1852), when he described Leptocerus magnus author during seven extensive collecting ex- (now Triplectides) and Leptocerus oppositus peditions between 1965 and 1974, when more (now Symphitoneuria); as a third species he than 18 000 specimens were collected. Ad- listed a variety of Plectrotarsus gravenhorsti ditionally, Dr E. F. Riek, of Canberra, made Kolenati, which later was recognized and de- available his collection containing about 2000 scribed by Mosely as a separate species Plec- specimens. Smaller numbers were obtained trotarsus tasmanicus. This first record was fol- from other collectors. About 16 000 specimens lowed by nearly 80 years of almost complete of the available material have been identified to silence, with only an occasional reference to species and constitute the basis of this study. Trichoptera of the island. One such note ap- The remaining specimens are mainly females, peared in the meeting notices of the Royal So- which could be identified only to genus, or re- ciety of Tasmania, regarding the erroneous ferred to species with some doubt.
J ARTURS NEBOISS
Altogether 66 genera, containing 157 species The author gratefully acknowledges the grant and placed in 21 families are dealt with in this received from the Australian Biological Re-
study; of these, 16 genera and 64 species are search Study, Interim Council, which made it described as new. It was found that the avail- possible to extend investigations to additional able material contained all but eight species: localities and aided the printing of this volume; the unique types of three were available for the Tasmanian National Parks and Wildlife study, but for each of the remaining five, only a Service for permission to collect specimens brief description is given. within the National Parks, and the respective Locality and collecting data are listed for rangers for their personal interest and help; Dr each species following the description, and dis- P. S. Lake and Dr P. Taylor of the University tribution is shown on the accompanying maps. of Tasmania, Hobart, for their co-operation; Depository institutions are indicated in the Dr E. F. Riek, CSIRO Division of Entomology, text by the following abbreviations: Canberra, for the loan of his extensive collec- ANIC— Australian National Insect Collec- tion and valuable discussions, which helped to tion, Canberra clarify many taxonomic problems; Prof G. Mar- BMNH— British Museum (Natural History), lier and Dr S. Jacquemart of the Institute Royal London des Sciences Naturelles de Belgique, Brussels, IRScNB— Institut Royal des Sciences Natu- for the loan of type material in their custody; relles de Belgique, Brussels Dr P. C. Barnard of Department of Ento- LM— Zoological Museum, Leningrad mology, British Museum Natural History, Lon- MCZ— Museum of Comparative Zoology, don, Dr V. Ziltzova, of Zoological Museum, Harvard University, Cambridge Leningrad, and Mr A. G. McFarlane of Can- NMV— National Museum of Victoria, Mel- terbury Museum, Christchurch, New Zealand, bourne for loan of types and other specimens; Dr H. H. NRS— Naturhistoriska Riksmuseet, Stock- Ross of the University of Georgia, Athens, holm Georgia, USA, Prof. J. lilies of Limnologische QM— Queensland Museum, Brisbane Flusstation, Schlitz, Germany, and Dr H. QU— Queensland University, Brisbane Malicky of Biologische Station Lunz, Lunz am SAM— South Australian Museum, Ade- See, Austria, for helpful discussions and advice laide on taxonomic problems. TM— Tasmanian Museum and Art Gal- Equally important and greatly appreciated is lery, Hobart. the generous technical help received from Mr V. Salinitri, Zoology Acknowledgements Department, Monash Uni- versity, Clayton. A study of this nature could not be accom- Sincere thanks are due to Mr I. Dimits for plished without the assistance of many institutes the permission to use his excellent photographs and persons, therefore to all of them I wish to to illustrate some of the most interesting locali- extend gratitude my and sincere thanks. ties. Especially I wish to thank Professor J. W. Finally, and most of all, I would like Warren, Chairman Department of Zoology, to thank my wife, Austra, for her patience, Monash University, Clayton, and Mr J. Mc- never-ending encouragement and support Nally, Director National Museum of Victoria, throughout the years of study and for typing for making this study possible; to Dr G. Etter- the manuscript. Without her help the comple- shank, Department of Zoology, Monash Uni- tion of this study would have been so much versity, for his constructive criticism and en- more difficult. couragement throughout the preparation of this Study Area work; and to Charles Mr McCubbin, Mel- 1. DESCRIPTION bourne, who with his skill and enthusiasm or- The present study deals with material col- ganized the first two collecting expeditions to lected in Tasmania, although occasionally Southwest Tasmania. it has been necessary to include information on TASMANIAN CADDIS-FLIES specimens from other Australian localities. coastal belt around the island. Maximum tem- Physically, Tasmania is part of the East- peratures reach as high as in the South- Australian 40°C Highlands, but it is separated from east (Derwent River valley), and drop as low the mainland by the relatively shallow Bass as — 12°C on the central high plateau. The Strait. It is an island of 2 67 340 km , located be- island is mostly mountainous and very little of tween 40° and 44° south latitude and 144°30' its surface is classified as coastal lowlands. and 148°30' east longitude. This is an area For the purpose of analysing the composition dominated by strong westerly winds, but with of Tasmanian Trichoptera, it was necessary to an otherwise temperate oceanic climate. The subdivide the island into seven provinces, the mean summer (December-February) tempera- boundaries of which are based on a combina- ture ranges between 11°C on the highlands to tion of landforms (elevations, watersheds, etc.), 18°C on the Eastern coastal regions. The mean geological features, vegetation and climatic con- winter (June-August) temperatures are as low ditions (particularly rainfall and tempera- as 2°C on the highlands to 9°C on a narrow ture) —see Map 1.
Map 1 —Boundaries of provinces. ARTURS NEBOISS
occupying the The two island groups, including King and Northwestern province (NW) line through Mt Sorell-Franklin Flinders islands, just north of Tasmania, are area north of a west of the Cen- excluded from the present survey, due to River-Lake King William and River. Vegetation limited funds and time. tral Plateau along the Mersey type, with some sedge- References to the provinces throughout the is dominantly rain-forest heaths, interrupted by tracts text are made in their abbreviated form by lands and coastal yellow podzolic soils. capital letters shown in parentheses below. of cleared land; mainly with Northern province (N) includes the coastal Relief hilly to rugged, drainage westerly and Pieman heathlands along the NE coast, but otherwise two major stream systems—Arthur Rainfall ranging from high in the it is dominated by dry sclerophyll forest and ex- Rivers. in tensive areas of cleared, cultivated land. Relief southern part of the province to moderate mainly undulating, drainage northerly, main the north. Temperatures cool to moderate. the line streams being rivers Forth, Mersey, South and Southwestern province (SW) west of MuelJer-Hythe; vegeta- North Esk, St. Patricks and Great Forester. Lake King William-Mt Rainfall low along the coast, increasing to tion with extensive sedgelands (often referred moderate towards the mountains; temperatures to as buttongrass plains from the dominant mild to warm. buttongrass Gymnoschoenus sphaerocephalus) Northeastern province (NE), a small, iso- and tracts of rainforests; skeletal soils and moor lated highland area of the Ben Lomond/Mt podzol peats. Relief dominated by rugged Barrow massif, and the mountains to the North- mountain ranges and swampy plains; drainage east. Vegetation varies from rainforest to sclero- north-westerly by Gordon River system and phyll forest, cut by sections of cleared land; it south-easterly by Huon River. High to very includes a small area of moorland on the higher high rainfall—over 2500 mm p.a. Tempera- altitudes of Ben Lomond. Relief rugged, drain- tures vary from warm to cold. age in all directions; rainfall moderate, tempera- Southeastern province (SE) includes most of tures cool to moderate. the lower section of Derwent River basin; Eastern province (E) extends along the east vegetation mainly sclerophyll forests with ex- coast from Eddystone Point in the North, to tensive areas of cleared land and some rain- Marion Bay in the South and includes hill forests in the western part of the province. country east of the slopes of the Central pro- Relief characterized by hilly terrain, river val- vince. It is dominantly dry sclerophyll forests leys and some rugged mountain ranges; drain- with large sections of cleared land. Relief hilly, age south-westerly. Rainfall rapidly decreases drainage easterly (George, Scamander, Swan from 1500 mm p.a. in the west, to less than and Prosser Rivers), and northerly (South Esk 800 mm p.a. in the east. Temperatures vary and Macquarie Rivers). Rainfall low—below from mild to hot. 700 mm p.a., with a high fluctuation level in stream flows; temperatures mild to very warm, 2. DISTRIBUTION OF LOCALITIES with low temperatures in winter. There are 130 localities for which either Central province (C) structurally part of the exact or at least approximate location is Central Plateau, with altitudes above 900 m, is known; all these are marked and numbered on dominated by alpine humus soils, and being in the accompanying map (Map 2). Data of lo- a rain-shadow area, has rather poor vegeta- cality labels have been given in their original tion—partly moorlands, partly dry sclerophyll form, and distances, were given in miles, were forests. Relief high plateau, in parts with rugged not converted into kilometres. One locality, if areas; drainage mainly southern to the Derwent labelled differently by several collectors, or River tributaries with Lake River to the North. two localities being very close to each other, Natural stream flow severely disrupted by a are marked by only one number on the map. highly developed hydro-electric industry, which The locality 'Great Lake' is on all occasions re- also controls water level in lakes and lagoons. garded as the area near Miena; 'Lake Pedder' Low average temperatures. refers to boundaries of the natural lake and the 51
TASMANIAN CADDIS-FLIES surrounding National Park, where the specimens 33 Apsley River nr. Bicheno were collected before its flooding by the Hydro- 34 Macquarie River, 8 km W of Campbell electric development scheme, converting it into Town a greatly enlarged water reservoir. Incompletely 35 Isis River nr. Auburn specified localities, such as 'Tasmania', T>er- 36 Lake Leake went River' or similar, have for obvious rea- 37 Tooms Lake sons been omitted from the map altogether. 38 Buxton River nr. Mayfield 39 Prosser River, Orford N Province 40 Prosser River, 2 km W of Buckland 1 Wilson Creek nr. Hellyer (Pebbly Bay) 41 Andover, York Rivulet 2 Burnie 42 Oatlands, small creek 5 km W 3 Guide River Falls nr. Ridgley 4 Ulverstone, 4 km NW, small waterfalls C Province on the coast 43 Blackman River, 15 km NW of Oatlands 5 Leven River nr. Heka 44 Interlaken, canal and Lake Sorell 6 Wilmot River, 10 km S of Forth 45 Clyde River, nr. Interlaken 7 Wilmot and Forth Falls 46 Lagoon of Islands 8 Sheffield 47 Arthurs Lake 9 Minnow River nr. Paradise 48 Penstock Lagoon, Waddamana 10 Mersey River, Liena 49 Great Lake, Miena, Shannon Lagoon 1 Small creek, 4 km E of Liena and Shannon River 12 Creek nr. Marakoopa Caves 50 Ouse River, 8 km W of Miena; (also 5 13 Sassafras Creek, 4 km W of Mole Creek mis W of Miena) 14 Rubicon River, 8 km SE of Sassafras 51 Lake Augusta, Howe Lagoon and 1 Meander River, Deloraine Western Lakes 16 Quamby Brook, 1 km E of Golden Val- 52 Pine Creek, 5 km N of Breona ley 53 Little Pine Lagoon and Little Pine 17 Liffey River, 5 km W of Liffey River 18 Saxon Creek, 10 km NW of Frankford 54 Nive River, 2 km W of Bronte 19 Franklin River, Frankford 55 Bronte Lagoon and Bronte-Bradys Canal 20 Meander River, 3 km N of Westbury 56 Bradys Lake 21 Lake River, 5 km SW of Delmont 57 Small creek 5 km W of Bronte 22 South Esk River, Evandale 58 Clarence River, 9 km E of Derwent 23 Lilydale, a small creek 2 km N of town Bridge 24 St. Patricks River, Targa 59 Derwent River, 2 km NW of Derwent 25 Scottsdale Bridge 26 Great Forester River, 5 km NW of For- 60 Lake St. Clair, Derwent basin, Cynthia ester Bay 27 Waterhouse estate, 25 mis NE of Scotts- 61 Hugel River, Cuvier River and Mt Ru- dale fus, West of Lake St. Clair 28 Launceston, Cataract Gorge 62 Snake Creek, Fisher River Road 63 Fisher River, Pencil Pine Grove below NE Province Lake Mackenzie dam
29 Mt Barrow, 2500 ft. NW Province 30 North Esk River nr. Blessington or 20 64 Mersey River trib. 4 km of Liena mis E of Launceston W 65 Bulls Creek, Cradle Mtn. Road; also 15 31 St. Columba Falls, Pyengana mis S of Wilmot E Province 66 Iris River tributary, 15 km N of Cradle 32 Scamander River, Upper Scamander Mtn. 1 4
ARTURS NEBOISS
67 Pencil Pine River, 6 km N of Cradle 103 Bushy Park, Derwent River Mtn. 104 Plenty River, 6 km E of Moogara 68 Waldheim, Cradle Mtn. National Park 105 Derwent River, 3 km W of New Nor- 69 Dove River, Cradle Mtn. National Park folk and New Norfolk
70 Lake Lilla, Cradle Mtn. National Park 1 06 Bridgewater, Derwent River 7 Murchison River, Murchison h-way 107 Sorell River, 3 km N of Sorell 72 Mackintosh River, Murchison h-way 108 Port Arthur 73 Farm Creek, 4 km N of Tullah 109 Strickland Ave., Hobart 74 5 mis S of Waratah, and 3 mis E of 110 Mt Wellington Waratah 111 Creekton River nr. Dover 75 Fossey River, 10 mis S of Hellyer River 112 Hythe Gorge 76 Hellyer River Gorge, Waratah h-way SW Province bridge 113 Damper Inn, Port Davey Track, Mt 77 Flowerdale River, Meunna Mueller area 78 Dip River Falls, 10 km S of Mawbanna 1 1 Wedge River, 30 mis W of Maydena 79 River, Duck 6 km SW of Roger River 115 40 mis W of Maydena (Strathgordon 80 Eckberg Creek, 12 km SW of Roger Road) River 116 Gordon River (now Strathgordon) 50 81 Arthur River bridge, 15 km SW of Roger mis W of Maydena River 117 Huon River Crossing 82 Bluff Hill creek, 12 km S of Marrawah 118 Condominion Creek nr. Mt Eliza 83 Sundown Creek, 25 km S of Marrawah 119 LakePedder 84 Rupert Point, 3 mis N of Pieman River 120 Huon Plains, nr. Scotts Peak 85 Corinna 121 West Arthur Plains 86 Hogarth Falls, Strahan 122 Junction Creek, West Arthur Plains 87 10 mis E of Strahan 123 Spring River 88 1 km SW of Queenstown 124 Cracroft River 89 Henty River, 12 km NW Queenstown 125 Huon River nr. Blakes Opening 90 King River, Lyell h-way bridge 126 Port Davey track 4 km W of Picton River 91 Collingwood River bridge, Lyell h-way 127 Huon-Picton River junction (30 km E of Gormanston) 128 Arve River, 10 km W Geeveston 92 Franklin River, 20 km SW of Derwent 129 Hartz Mtn. National Park Bridge 130 Hot Springs Creek nr. Hastings Caves 93 Arrowsmith Creek, 18 km SW of Der- went Bridge Methods 94 Navarre River, Lyell h-way COLLECTION AND PRESERVATION To obtain material used in this study, several SE Province collecting methods were employed. During the 95 Black Bobs Creek, 15 km NW of Ouse day and in the early hours of evening the cus- 96 Dee River, 8 km NW of Ouse tomary hand-net was used to capture in- 97 Cashion Creek Cave, Florentine Valley dividuals in flight, or by sweeping vegetation in 98 Ellendale the vicinity and along the banks of fresh-water 99 Russell Falls, National Park and Tyenna bodies. River Searching on the underside of bridges and 1 00 Styx River, Westerway the interior surfaces of culverts under roads 101 Lake Dobson, also Broad River, Mt proved to be very productive and was widely Field Nat. Park used while travelling. Nevertheless, the number 102 Florentine River, 7 mis W of Maydena of specimens obtained by these methods was TASMANIAN CADDIS-FLIES
Map 2—Distribution of localities. ARTURS NEBOISS never as large as that obtained by collecting The material preserved and permanendy with a mercury-vapour light source during stored in ethanol, was placed in small glass evening and night. vials, with one or several specimens in each locality labels were A portable generator of 240V 600W or vial. Written or printed within individual vials, to which the 1000W output was used as the power unit for placed labels were added when the ma- 150W to 500W blended or clear glass lamps. identification terial finally sorted. The individual vials The type used depended on their availability. was placed into During rainy weather a clear polythene sheet were then sorted by taxa and larger jars for permanent storage. was suspended above the lamp in a tent-like fashion to protect the globe from cracking. PREPARATION AND ILLUSTRATION OF On some occasions a 15W black light fluores- SPECIMENS cent tube was used, but with much less success. Description of species usually was prepared There was no attempt made to compare the from specimens preserved in ethanol with ad- attractive power of the various types of lamps ditional details, particularly, colour pattern used. of wings, taken from dried specimens when- The specimens were collected from twilight ever they were available. Although colouration
to midnight, or sometimes even later. Collect- is of limited importance, because it varies con- ing was done by hand from two white sheets, siderably with the age of the specimens, it is one hung vertically, facing the body of water still useful for correct correlation of sexes, and close behind the lamp, the other horizontally to a certain degree, for identification of species below the lamp on the ground. when the specimens are preserved dry and the During unfavourable weather conditions, hairs are not denuded. when only small numbers of specimens were at- Association of males and females was either tracted to the light, all of them were collected assured when pairs in copula were obtained in and preserved. When large numbers were at- the field and preserved separately, or by care- tracted, a selective collection of specimens fully matching specimens from one locality. was made. To obtain the largest possible num- The wings from the right side of the body ber of species, the observation of difference in were detached and used for illustrations. They size, colour, behaviour, etc. was of utmost im- were removed from the specimens, preserved portance. in ethanol, transferred to a microscope slide and orientated; after adding a small drop of The number of specimens attracted to the glycerol, a cover-glass was placed over the light varied considerably, depending on the sea- specimen. Wing drawings were made with the son and weather conditions. Collecting usually aid of a camera lucida on a stereo binocular came to a complete standstill when the even- microscope. The same method of drawing was ing temperature dropped below 8°C. The oc- applied for head and thoracic parts, using currence of cool evening breezes or sudden specimens in ethanol. The hyaline areas of the wind gusts reduced activity also, and speci- anterior wings have characteristic shape and mens already on sheets became entirely motion- position which is diagnostic to individual less. Light rain did not interrupt flying, so long families. These areas are indicated in the wing as the temperature stayed favourable. No drawings by oblique line shading. The termino- samples were collected with artificial light dur- logy for wing venation is the same as used by ing the dark hours immediately before sunrise. Mosely and Kimmins (1953). Most specimens were killed and preserved in Examination and drawing of male and fe- ethanol, and only 80% a small number were male genitalia was done after the entire abdo- killed in ethyl-acetate or cyanide vapour for men had been cleared in boiling KOH, washed dry preservation as panned specimens. This and replaced in glycerol. Genitalia drawings method was employed to help in identification were made with the aid of a camera lucida on a of colour pattern on wings and body. monocular microscope. —
TASMANIAN CADDIS-FLIES
The two Australian mainland species Plec- 8. Anterior wings with R r not forked trocnemia australica Banks and Anisocentropus at apex Ecnomidae bicoloratus ( Martynov ) are described and — . Anterior wings with not forked R t figured, but not numbered. at apex 9 9. Mesoscutum with a pair of warts .... 10 KEY TO THE FAMILIES — . Mesoscutum without
1. Scutellum flat, triangular, pointed pos- warts Hydropsychidae teriorly, margins vertical; mesoscutum 10. Mesoscutal warts separated by anteriorly without warts; wings narrow, slender, extended section of more or less pointed apically; fringes very scutellum Stenopsychidae long, those of posterior wings usually — . Mesoscutal warts not separated by scutel- longer than the width of the wing. Maxil- lum Polycentropodidae lary palpi 5-segmented in both sexes. Size 11. Maxillary palpi six segmented (in both small, length of anterior wing less than sexes) Calamoceratidae 4 mm Hydroptilidae — . Maxillary palpi one to five segmented in males, always 5-segmented —. Scutellum rounded posteriorly, convex, without vertical margins; wings propor- in females 12 12. tionally broad, fringes usually short, less Maxillary palpi segment 1 short with more than the width of the wing. Maxillary pal- or less distinct mesal nodule pi with variable number of segments. Size near apex Philorheithridae generally larger, length of anterior wing — . Maxillary palpi segment 1 without mesal more than 4 mm 2 nodule 13 2. Ocelli present 3 13. Antennae distinctly longer than anterior wings, particularly in males; mesoscutum —. Ocelli absent 7 with groups 3. Maxillary palpi 5-segmented in both two of more or less parallel sexes 4 rows of setiferous punctures 14
— . Antennae usually as long as anterior — . Maxillary palpi 3-segmented in males, wings, seldom slightly 5-segmented in females 6 longer; mesoscutum either with warts or entirely without warts 4. Maxillary palpi with segment 5 barely or setiferous punctures 15 longer than segment 4, not flexible . . 5 14. Posterior wing with a row of macrotrichia —. Maxillary palpi with segment 5 dis- along basal half of the costal tinctly longer than segment 4, flex- margin Odontoceridae ible Philopotamidae
— . Posterior wing with a short row or rows of 5. Second segment of maxillary palpi elon- curved macrotrichia along costal margin gate, longer than about opposite the discoidal segment 1 Rhyacophilidae cell Leptoceridae short, — . Second segment of maxillary palpi
15. Mesoscutum with a pair of warts , , 16 not longer than segment 1, extended la- — . Mesoscutum without warts 19 terally to globular 16. Mesoscutal elongate, projection Glossosomatidae warts narrow . . 17
. — Mesoscutal warts short, rounded . . . . 18 6. Scutum with posterolateral warts level with scutellum Plectrotarsidae 17. Scutellum with a pair of elongate warts, posterolateral warts maxillary palpi in male — . Scutum without Limnephilidae 3-segmented Kokiriidae
— . Scutellum with a single median wart, 7. Terminal segment of maxillary palpi longer maxillary palpi in male single segmented than the preceding segment, flexible . 8 maxillary palpi about Oeconesidae — . Terminal segment of the same length as the preceding seg- 18. First antennal segment very long, exceed- ment 11 ing the length of the head; basal half of —
10 ARTURS NEBOISS
the costal margin in posterior wings with Following the revision of Australian Hydro- extensive a row of hamuli; maxillary palpi 2-scg- biosinae by Neboiss (1962), more collecting has continued and new material ac- mented in males . . . . Helicopsychidae cumulated not only from Tasmania, but also — . First antennal segment about as long as the head; posterior wings with bristle-like from other Australian localities. The informa- hairs along the costal margin; maxillary tion available from the analysed Tasmanian specimens indicate that the taxonomic group- palpi 4-scgmented in males . . Tasimiidae 19. Pronotum with two pairs of warts—the ing could not be regarded as fully established middle pair small, rounded, the lateral pair and further changes are imminent.
larger, somewhat elongate (except Caeno- It was observed that the ridge on the sternite ta males, which have only one pair of 5 in the females has two basic positions. In elongate warts) Calocidae one group the ridge curves upwards and termin-
— . Pronotum with only one pair of elongate ates at the anterior margin of the sternite, in warts 20 the second group this ridge terminates at the 20. Maxillary palpi 5-segmented lateral margin. The first position occurs only in in males Helicophidae two genera A psilochorema and A llochore- — . Maxillary palpi 1-3-segmented ma—indicating their close affinity. The an- in males Conoesucidae terior wing in both genera with the cross-vein SUPERFAMILY RHYACOPHILOIDEA C-Sc present, fork 1 either short, or absent in some A psilochorema species. The males of the Wing venation entire; maxillary palpi with genus Apsilochorema have a more or less de- terminal segment simple, cylindrical. Larvae veloped, oblique pouch at the middle of the campodeiform, head prognathous, either free anterior wing. It has been recorded (Ulmer, living or constructing portable cases. 1957; Neboiss, 1962; Lepneva, 1964) that the 1 Family RHYACOPHILIDAE Stephens larvae of Apsilochorema have simple and not (1836) chelate anterior legs (Fig. 17). The larvae of the genus Allochorema are not known. These Family diagnosis: Ocelli present. Antennae two genera are now placed in a new subfamily slender, as long as or slightly shorter than an- Apsilochoreminae. terior wing; basal segment stout, shorter than head. Maxillary palpi 5-segmented, alike in The subfamily Hydrobiosinae is arranged in both sexes; first two segments very short, the two tribes—the Hydrobiosini and Psyllobetini. following ones long, cylindrical. Wings elon- The genera with anterior wings evenly pubes- gate, vestiture variable; discoidal cell in both cent, without hair tufts, open discoidal cell in wings either open or closed, in posterior wing the male anterior wings and the abdomen ter- minating sometimes absent. Anterior wing with forks 1, in a long, slightly upcurved oviscapt in the females, are placed in a new tribe Psyllo- 2, 3, 4 and 5 present; R x usually forked at apex; thyridial cell always present; additional betini. To this tribe belong genus Psyllobetina cross-veins C-Sc and Sc-Rj sometimes present. Banks, distributed from Queensland to Vic- Posterior wing shorter, forks variable, but at toria, genus Allobiosis Mosely, known from least 2 and 5 present. New South Wales only, genus Moruya Neboiss Spurs 1:4:4; 2:4:4 or 3:4:4. endemic to Tasmania, and the New Zealand The family is here divided into three sub- endemic genus Tiphobiosis Tillyard. All other families. Subfamily Rhyacophilinae with spurs genera are placed in the tribe Hydrobiosini. 3:4:4 has not yet been recorded from the Aus- The species originally placed in the genus tralian region. The Australian species have Taschorema, exhibited a number of morpho- been placed in two subfamilies—Hydrobiosinae logical differences, including the position of and Apsilochoreminae, both with spurs 2:4:4, warts and oscelli on the head, shape of terminal except in genus Ulmerochorema (Hydrobio- segments of the female abdomen, presence or sinae) which has spurs 1:4:4. absence of lateral filaments on abdominal seg- —
TASMANIAN CADDIS-FLIES 11
footstalk merit 5 in the males, various hair-structures on — . Posterior wing — fork 2 with male wings and wing venation in general. Sup- longer than fork (except U. breve) at least ported by the newly discovered species from twice as long as the width of the fork; dis- Tasmania, this group now has been divided coidal cell in anterior wing closed in both into three separate general Taschorema (sensu sexes Ulmerochorema stricto), Ethochorema and Ptychobiosis. Fur- 7. Anterior wing with discoidal cell open ther research on Australian mainland species (except in females of the genus Moruya); might call for even further changes. females with abdomen terminating in a The free living larvae of Hydrobiosinae and long, upturned oviscapt 12 Apsilochoreminae are predacious, mainly on cell closed; — . Anterior wing with discoidal other aquatic insects. They are slender, pale- females with abdomen terminating yellowish or greenish in colour, with only the bluntly 8 head and pronotum sclerotized. The larvae 8. In posterior wing R x merges with Sc of cemented sand make a pupal chamber shortly before reaching grains, within which they spin a smooth, cylin- margin Ptychobiosis drical cocoon, secured at both ends to the either connected to — . In posterior wing R t chamber. Sc with short cross vein or completely Australian genera A new generic key for all separate 9 replaces the one published by Neboiss ( 1962). 9. In anterior wing — fork 1 sessile . . 10
— . In anterior wing — fork 1 with footstalk Ethochorema KEY TO AUSTRALIAN GENERA 10. Posterior wing in males with elongate cell-
1. Posterior wing with irregular venation in like structure between Cu2 and Ai, andro- it for both sexes; Ri joins Sc, runs with a conia on A 2 and A 3 short distance, then diverges and connects present Taschorema wing margin without cell-like with R2+ 3 before reaching — . Posterior wings in males
Koetonga structure and without androconia . . 11 with regular venation, Sc — . Posterior wing 11. Lateral ocelli touching anterior with and Ri entirely separate, connected warts Tanjilana short cross vein or joined together shortly anterior — . Lateral ocelli not touching before the wing margin 2 warts Megogata 2. Posterior wing — fork 2 with footstalk 12. In posterior wing — fork 1 present, and very short) 3 (sometimes with footstalk Moruya fork 2 sessile .... 7 — . Posterior wing — 1 absent . . — . In posterior wing — fork 13 3. Anterior wing — fork 1 short .... 4 13. In anterior wing — fork 2 very long, about fork 1 long 5 — . Anterior wing — twice as long as fork 3 . . . . Psyllobetina 4. Posterior wing fork 2 longer than its — short, about as — . In anterior wing — fork 2 footstalk Apsilochorema long as fork 3 Allobiosis fork 2 about as long or — . Posterior wing —
shorter than its footstalk . . Allochorema as 5. Anterior wing — fork 4 about as long Subfamily APSILOCHOREMINAE Austrochorema fork 3 subfam. n. fork 4 distinctly longer — . Anterior wing — females the ridge on sternite 5 than fork 3 6 Diagnosis: In footstalk terminates at the anterior margin (Figs. 5 and 6. Posterior wing — fork 2 with The larvae with pro-sternum membraneous; very short (about as long as the width of 6). anterior legs with trochanter and femur fused fork); discoidal cell in anterior wing open and greatly enlarged, tibia and tarsus short, in male, usually closed in female Ipsebiosis subequal, claw long and thin (Fig. 17). 12 ARTURS NEBOISS
Genus Apsilochorema Ulmer Length of anterior wing: $ 7-8 5 mm; 2 Apsilochorema Ulmer, 1907:206; Ross and King, 8-9 5 mm. 1951:503; Ross, 1956:124; Kimmins, 1960:184; Neboiss, 1962:526. Apsilochorema obliquum Bachorema Mosely in Mosely and Kimmins, 1953: 493; Neboiss, 1957:83. Type species: Psilochorema indicum Ulmer, 1905. Wing venation differing in sexes, regular in females, irregular in males; anterior wing with
forks 1, 2, 3, 4 and 5 present, in males an oblique pouch along footstalk between base of
fork 4 and Rs. Cu2 sometimes joined with Ai just above arculus; posterior wings similar in both sexes, forks 1, 2, 3 and 5, or only 2, 3 and 5 present, all with footstalks. In male the lateral filament* on sternite 5 present; in fe- male the ridge on sternite 5 terminates at the anterior margin (Figs. 5 and 6). Ventral pro- cesses short or moderately long on sternites 6 and 7, or 7 only in males; on sternite 5 and 6 in females. Spurs 2:4:4. Genus distributed from Ceylon and India to Japan, Siberia and south to Fiji, New Guinea and Australia. Two species in Tasmania.
KEY FOR SEPARATING TASMANIAN Type material: Type & National Park, Qld., SPECIES 26 Dec. 1921 (BMNH). Type not seen. 1. Posterior wing with apical forks 1, 2, 3 Material examined: Tasmania—2 ? St. Patricks and 5 present obliquum River, Targa, 22 Feb. 1971; 2 ? St. Columba Falls,
— . Posterior wing with apical forks 2, 3 and Pyengana, 21 Feb. 1971; 2 $ Russell Falls, National 5 only gisbum Park, 15 Nov. 1972; 1 $ Corinna, 5 Nov. 1972; 1
Material examined: Tasmania 16
Allochorema Mosely in Mosely and Kimmins, 1953: 491; Ross, 1956:124; Neboiss, 1962:529. Type species: Allochorema tasmanica Mosely, 1953. Wing venation similar in both sexes; an- terior wing with discoidal cell open, forks 1, 2 and 3 short with long footstalks, forks 4 and 5 long, about equal in length; in posterior wing discoidal cell open, fork 1 absent, fork 2 with long footstalk. The lateral filament on sternite 5 in male two-segmented, base somewhat elon- gate, bulbous; in female the ridge on sternite 5 terminates at the anterior margin of the sternite. Only one species in Tasmania. Spurs 2:4:4.
3 Allochorema tasmanicum Mosely Figures 19-22
Allochorema tasmanica Mosely in Mosely and Kim-
mins, 1953:491; xRoss, 1956:124; Jacquemart, 1965b:36. Allochorema tasmanicum, Neboiss, 1962:530. 14 ARTURS NEBOISS
Insect brownish of medium size, wing vena- 14 Dec. 1974, A. Neboiss (NMV); 4 o* 1 ? 10 mis 1 E Strahan, 6 Feb. 1967, E. F. Riek (ANIC); 1 d tion as described in the generic description; same loc. 20 Feb. 1963, I. F. B. Common and M. S. the species is distinguished by the male geni- Upton (ANIC). talia. Other recorded localities: Cradle Mtn. ( Jacque- $ genitalia with segment 10 wide at the mart, 1965). base, deeply and roundly excised at the apex, Distribution: Tasmania—NW, SW, C and SE apices covered with short, stout teeth; superior provinces. appendages short, stout; inferior appendages large, slightly constricted at distal third, apices Subfamily HYDROBIOSINAE Ulmer (1905) dilated and covered with short spines on the Diagnosis: In females the ridge on sternite 5 inner surface. A short, blunt ventral process on terminates at the lateral margin. The larvae sternite 6. with prosternum heavily sclerotized; anterior 9 abdomen terminates bluntly; a pair of legs enlarged, chelate; the chela formed by a small cerci at the apex; strong ridge on sternite distal process from the femur articulating with 5, which terminates at the anterior margin, but the combined tibia-tarsus-claw. ventrally goes into strong ventral process, which has a few stout spines; ventral process on ster- Tribe Hydrobiosini nite 7 short. Genus Austrachorema Mosely Length of anterior wing: g 6-7 mm; 9 5*5- Austrochorema Mosely in Mosely and Kimmins, 6-5 mm. 1953:481; Neboiss, 1962:532. Type species: Austrochorema wenta Mosely, 1953. Wing venation similar in both sexes, and dis- coidal cell open in both wings; anterior wing with forks 1 and 2 long, sessile, others with
footstalks; posterior wing with forks 1, 2, 3 and 5 present, all with footstalks. Lateral fila- ment on sternite 5 in males long, with thicker proximal, and thinner, more transparent distal section. The ridge on sternite 5 in females terminates with a small loop which extends above the lateral margin. Posterior tibia in both sexes covered with long hairs. Spurs 2:4:4. In general appearance all species of the genus appear to be very similar, uniformly greyish-brown in colour and of medium size; they are separated on the differences in genitalia structures. In both, the male and female, genitalia has two basic types. The males with a short coxo- podite of the inferior appendage correspon- Type material: Type g Mt Wellington, Tas. dingly has females with short segments 9 and 3000 ft., Dec. 1937, J. W. Evans (BMNH). 1(X This group is known by six species, all of Type not seen. which, except one, occur only on the Australian Material Examined: Tasmania—5 c? 4 ? Junction mainland, the exception being the endemic Tas- Creek, West Arthur Plains, 6 Feb. 1966, A. Neboiss manian species pegidion. The other group is (NMV); 4
KEY TO THE TASMANIAN SPECIES (Males only)
1. Inferior appendages with harpago shorter than coxopodite 2
— . Inferior appendages with harpago longer than coxopodite pegidion 2. Superior appendages about as long as the inferior appendages 3
— . Superior appendages distinctly shorter
than inferior appendages . . . . lepnevae 3. Segment 9 in ventral view with sides gradually curved to slightly incised apex complexa
— . Segment 9 in ventral view abruptly nar-
rowed in upcurved apical section . . 4 4. Segment 9 upcurved terminal section with flattened and bi-pointed, inwardly directed plate evansi
— . Segment 9 with upcurved terminal section, rounded apically crinitum
4 Austrochorema pegidion Neboiss Type material: Holotype $ Broad River nr.
Lake Dobson, Mt Field National Park, Tas. s Figures 23-31 6 Nov. 1955, T. Woodward (QM). Type seen. Austrochorema pegidion Neboiss, 1962:535 Material examined: Tasmania— 1 S 1 $ Russell This is the only species which belongs to Falls, National Park, 20 Feb. 1971; 1 <$ Strickland 1 Condominion the group of Australian mainland species as Ave., Hobart, 22 Feb. 1967; c? Creek, 15 Feb. 1971; 2 cf 1 2 Ulverstone 4 km NW, described above. small waterfalls, 18 Nov. 1972; 5 o* Dip River Falls, $ genitalia: segment 10 semi-membraneous; 10 km S of Mawbanna, 1 Dec. 1974; 1
9 abdomen with the last three segments dis- tinctly flattened dorso-ventrally.
Length of anterior wing: $ 5-5-5 mm; 9 5-5-5 mm.
Type material: Type $ Tasmania, without exact locality. J. W. Evans (BMNH). Type not seen.
Material examined: Tasmania— 1
TASMANIAN CADDIS-FLIES 17
Type material; Holotype & (T4805), allotype 2 unknown. A female from Waldheim, 2 (T4806) Guide River Falls nr. Ridgley, Cradle Mtn. Nat. Park, 7 Feb. 1971, A. Ne- Tas., 18 Nov. 1972, A. Neboiss (NMV); 1 $ boiss, is doubtfully referred to this species and 2 2 paratypes (T4807-T4809) Dip River illustrated (Fig. 48). Falls 10 km S of Mawbanna, Tas., 1 Dec. Length of anterior wing: 6 5 5 mm. 1974, A. Neboiss (NMV). Type material: Holotype 6 Cradle Mtn., Tas., Distribution: Tasmania—N and NW provinces. 27 Jan. 1923, A. Tonnoir (IRScNB), dissected and mounted on three microscope slides. Type 7 Austrochorema lepnevae Jacquemart seen. Figures 47-48 No new material has been available for study. Austrochorema lepnevae Jacquemart, 1965b:43. The type has been dissected and, unfor- Distribution: Tasmania—NW province (known tunately, the parts of genitalia are so distorted from type locality only). that their original position could not be fully reconstructed. Relying on fiigure 36c of Jac- 8 A ustrochorema complexa Jacquemart quemart (1965b), the main difference between this species and crinitum described above, is Figures 45; 49-53 in the shape of the process arising on either Austrochorema complexa Jacquemart, 1965b:41. side of the base of segment 10. In lepnevae the S genitalia with segment 10 broad, some- upper angle of this process is short acute, and what rectangular, lower lateral margins ter- in crinitum long and curved. The superior ap- minating into a curved point; at the base below pendages in the original figure of lateral view segment 10 there is a chitinous plate which ends are not shown. Figure in the 36d same pub- in several processes as shown in the drawing. lication is incomplete, as it does not show the Superior appendage slender, about as long as thin distal section of the filament sternite on 5. segment 10. Inferior appendage slender, coxo- podite long, with small finger-like process at margin; short, Austrochorema lepnevae the inner apical harpago robust,
\ slightly curved inward. Lateral filament on sternite 5 long, directed anteriorly; thin, trans- 1 parent distal section short, curved. Posterior tibia covered with sparse long hairs and dense short pubescence. 1 *" cover of J'\ J The figures and description are prepared *-* VX^Y^C^ NW ( from a Lake Pedder specimen. v Jl7ne- r V 9 unknown. V / J anterior —\ x Length of wing: $ 6 mm. \ *v i*? vC" Type material: Holotype $ , Cradle Mtn. Tas., H 10 Jan. 1923, A. Tonnoir (IRScNB). Two 1.A p t \
B \ 1 I f male specimens have been used for the descrip- tion, which were mounted on 8 microscope
\ J ' slides without indication \ which part belongs to )p which specimen. The one abdomen, with the - \ SW \. genitalia intact, is labelled 'Holotype'. Type K "''^\ ' f- Uo^- j\ I seen. Material examined: Tasmania— 1 <7 Lake Pedder, p 10 Mar. 1972, A. Neboiss (NMV). Distribution: Tasmania NW and SW pro- 90 •0 80km 7 — vinces. 18 ARTURS NEBOISS
rounded chitinized lateral flaps near the apex, on either side at the base an elongate claw-like process. Superior appendages long and slender. Inferior appendages 2-segmcnted, coxopodite long, harpago short, apically rounded with small, acute projection at the inner apical mar- gin. Two long, upward directed and curved pro- cesses arise from the lower inner margin of seg- ment 9. Short, broad, laterally flattened and apically hooked lateral filament arises from the lateral margin of sternite 5. 9 abdomen terminates bluntly, segment 9 with distinct, rounded lateral flaps; ventral pro- cesses on segments 5 and 6, a fringe of long hairs along the posterior margin of sternite 7. Length of anterior wing: 6 8 5-9 mm; $ 9-9-5 mm.
Genus Ipsebiosis gen. n.
Type species: Ipsebiosis spicula gen. et sp. n. This genus occupies a position between Aus- trochorema and Ulmerochorema. There are some characteristics from one or the other genus, particularly in the wing venation; how- ever, their combination supported by details of other structures warranted separation. Anterior wing with Rs forked apically; forks
1, 2, 3, 4 and 5 present; forks 1 and 2 very long; fork 3 shorter than fork 4, which is about as long as fork 5; discoidal cell open in male, usually closed in female; a long bristle on anal margin in male. Posterior wing with forks 1, 2, 3 and 5 present, fork 2 with very short foot- stalk, discoidal cell open. Lateral filament on sternite 5 in the male present. Spurs 2:4:4. Type material: Holotype $ (T4810), allotype $ (T4811) Lake Dobson, Tas., 20 Feb. 1967, 9 Ipsebiosis spicula sp. n. A. Neboiss (NMV); 15 paratypes— 1 $ 1 9 Figures 54-61 (T4812-T4813) Waldheim, Cradle Mtn. Nat. Moderately large species with mottled brown Park, Tas., 7 Feb. 1971, A. Neboiss (NMV); and yellowish wings. A rounded patch of short, 1 s (T 4814) Lake Pedder, Tas., 1 Feb. vertical rods on the upper surface of posterior 1965, A. Neboiss (NMV); 1 $ (T4815) Con- wing in male. dominion Creek, Tas., 15 Feb. 1971, A. Ne- $ genitalia: segment 10 semi-membraneous, boiss (NMV); 1 $ (T4816) St. Patricks elongate, gradually narrowed distally, small, River, Targa, Tas., 22 Feb. 1971, A. Neboiss TASMANIAN CADDIS-FLIES 19
(NMV); 1 $ (T4817) Huon Plains nr. Scotts — . Fork 1 in posterior wing sessile .... 4 Peak, Tas., 2 Feb. 1965, A. Neboiss (NMV); 3. Male genitalia with lateral plates of seg- 1 9 (T4818) Huon River Crossing, Tas.! ment 10 broad, distal margin with 3 to 4 16 Feb. 1971, A. Neboiss (NMV); 1 9 claw-like extensions; in female a pair of (T4819) Derwent River 2 km NW Derwent dark, roof-like plates meso-ventrally on Bridge, Tas., 12 Feb. 1971, A. Neboiss segment 10 onychion (NMV); 2 $ Lake Dobson, Tas., 20 Feb. — . Male genitalia with lateral plates of seg- 1967, E. F. Riek (ANIC); 1 $ Derwent ment 10 pointed, upcurved distally; in Bridge, Tas., 12 Feb. 1967, E. F. Riek female no dark, roof-like plates meso- (ANIC) ; 2 s 10 mis E Strahan, Tas., 20 Feb. ventrally on segment 10 lentum 1963, I. F. B. Common and M. S. Upton 4. Fork 1 in posterior wing broad (ANIC; NMV); 1 $ 1 9 3 mis E Waratah, at base seona Tas., 17 Feb. 1963, 1. F. B. Common and S. M. — . Fork 1 in posterior wing tapered to a Upton (ANIC). point at base 5 Distribution: Tasmania—N, C, NW, SE and 5. In male genitalia the inferior appendage provinces. SW with both apical branches of approxi- mately Genus Ulmerochorema Mosely the same size; female-lateral pockets between segments 8 and 9 Ulmerochorema Mosely in Mosely and Kimmins, deep 1953:432; Ross, 1956:124: Neboiss, 1962:538. rubiconum Anachorema Mosely in Mosely and Kimmins, 1953: — . In male genitalia the inferior appendage 453. with upper apical branch smaller than the Type species: Hydrobiosis stigma Ulmer, 1916. lower; female-lateral pockets between seg- Anterior wing with discoidal cell closed; ments 8 and 9 shallow .... tasmanicum forks 1, 2, 3, 4 and 5 present, forks 1 and 2 sessile, others with footstalk. Posterior wing with discoidal cell open, forks 1, 2, 3 and 5 10 Ulmerochorema breve (Mosely) present, fork 1 either sessile or with short Figures 62-66 footstalk, others with footstalk. Lateral filament Anachorema brevis Mosely in Mosely and Kimmins, on sternite 5 in male arising from the lateral 1953:456; Jacquemart, 1965b:36. margin; posterior margin of segment 9 usually Ulmerochorema breve, Neboiss, 1962:539. with long, bifurcate process. In female often a The wing venation differs from other species dark, circular patch consisting of a mass of dark of the genus by the long fork 2 in posterior hairs located near the apex of posterior wing; wing, the footstalk being only one quarter of abdomen terminates bluntly; lateral pockets the length of the fork. formed between segments 8 and 9, and some- $ genitalia segment 10 broad at base, times covered with short setae. — curved gradually towards the apex, Spurs 1:4:4. lateral margins fringed with fine hairs, at the base on Of the six species known to occur in Tas- either side a pair of acutely pointed lobes; su- mania, three appear to be endemic. The general perior appendages long and slender; inferior ap- appearance of specimens, particularly of those pendages broad, tri-lobed distally, the upper preserved in alcohol, is so similar, that no des- lobe broad, obliquely truncate, the other two cription of colour is included. slender. A long, upcurved process arises from KEY TO THE TASMANIAN SPECIES the mid-posterior margin of sternite 9, divided nearly to the base into 1. Fork 2 in posterior wing longer than its two branches, each with footstalk breve a smaller, pointed lateral branch arising about midway and directed distally. Lateral filament — . Fork 2 in posterior wing shorter than its footstalk 2 on sternite 5 short, apex rounded. Ventral pro- cesses absent. 2. Fork 1 in posterior wing with short foot-
stalk 3 $ wing venation similar to that in $ , but 20 ARTURS NEB01SS
posterior wing with circular patch of dark hairs brane between the base of fork 1 and footstalk on lower surface of posterior wing between of fork 2.
forks 1 and 3. $ genitalia with lateral process arising be- Length of anterior wing: & 7 mm; 2 7 mm. low the base of segment 10 gradually tapering to slightly upcurved apex; superior appendages long and slender; inferior appendages two-seg- mented, proximal segment long, curved, ter- minating with inwardly directed acute point, distal segment short, obliquely truncate. From the centre of sternite 9 arise two long, slightly
curved spines, wr hich sometime show very fine dentation on ventral surface. $ abdomen terminates in blunt, rounded apex; lateral pockets between segments 8 and 9 small, covered with fine setae.
Length of anterior wing: 3 5-7 5 mm; 9- 6-8 mm.
Type material: Type $ , Cradle Mtn. Tas., 18 Jan. 1917, R. J. Tillyard (BMNH). Type not seen.
Material examined: Tasmania—2
11 Ulmerochorema seona (Mosely) Figures 67-69 Type material: Type $ River Ouse, Tas., 4 Anachorema seona Mosely in Mosely and Kimmins Feb. 1933, R. J. Tillyard 1953:458. (BMNH). Type seen. Ulmerochorema soena, Neboiss, Material 1962:545. examined: Tasmania—29 S 19 $? Derwent Anachorema seona (sic!) Kimmins, River Jacquemart, 2 km NW Derwent Bridge, 12 Feb. 1971; 10 TASMANIAN CADDIS-FLIES 21 1972; 6 o* 11 ? Huon River nr. Scotts Peak, 9 Feb. 1965; 1 River ? Huon Crossing, 16 Feb. 1971; 1 $ Cracroft River, 8 Feb. 1966; 1 12 Vlmerochorema lentum Neboiss Figures 70-72 Vlmerochorema lentum Neboiss, 1962:539. In posterior wing fork 1 with short footstalk; in females, although there is no definite circular patch, some dark hairs are present on the underside of the wing between forks 1 and 3. 13 Vlmerochorema onychion sp. n. Lateral filament on sternite 5 in male short. Figures 73-79 8 genitalia: segment 10 broad at base, from about the middle gradually tapered to a narrow, Anterior wings dark brown with mottled rounded apex; below the segment arise a pair appearance; in posterior wing fork 1 with short of plates, broad at base, upcurved distally, footstalk. The male with lateral filament on tapering to acute point. Superior appendages sternite 5 present; the inside of front tibia slender, slightly dilated distally. Phallus slender covered with a row of short spines, spur often with fringed, obliquely twisted apex. Inferior very small. The posterior wing in female with appendages laterally somewhat flattened, proxi- some dark hairs present between forks 1 and 3, mal two-thirds broad, distal one-third narrow, but there is no definite circular patch. small finger-like process on the inside near the S genitalia: segment 10 semi-membraneous, distal end. The posterior margin of sternite 9 short, apically truncate, on either side with in the middle with a pair of small protuberances strongly chitinized, large, downturned hook, be- only. low that is a transverse plate with finely dentate lower margin and a large, four-pronged claw 9 abdomen terminates bluntly; lateral laterally; an upcurved, finger-like process just pockets between segments 8 and 9 short. inward from superior appendage which is Length of anterior wing: <$ 8 mm; $ 8-9 mm. long and slender. The larger proximal section of Type material: Holotype $ , allotype 9 , inferior appendage as well as posterior part of Clunes, Vic, 9 Jul. 1953 ex pupa 12 Jul. 1953, sternite 9 densely covered with long, heavy, A. Neboiss (ANIC). Type seen. dark hairs; the smaller distal section consists of Material examined: 2 <$ 17 South Esk Tasmania ? two flat plates, lying close to each other, the River nr. Evandale, 1 Mar. 1967, A. Neboiss (NMV). lower one terminating with inwardly directed 1 o* Sorell River 3 km N of Sorell, 8 Dec. 1974, A. Neboiss (NMV). acute corner. A pair of slender, upward di- Distribution: Tasmania—N and SE provinces; rected processes arise from the middle of pos- New South Wales; Victoria. terior margin of sternite 9. 22 ARTURS NEBOISS 9 abdomen terminates bluntly, ventral plate Anterior wings blackish-brown in colour; broadly truncate; mcso-ventral surface of seg- posterior wings with fork 1 sessile, but not as ment 10 with a pair of roof-like, dark, squarish broadly as in U. seona; in female a distinct cir- plates. cular patch of dark hairs between forks 1 and Length of anterior wing: $ 6-7-5 mm; $ 6 5- 3. The lateral filament on sternite 5 in male 8 nun. present. $ genitalia rather slender. The plate at the base of segment 10 with inner margin produced in small, rounded process; the lower margin of the plate as seen from the side, is produced downward in a triangular keel. Inferior ap- pendage very long, curving slightly upward, with middle section of the inner surface widened and covered with short spines. Centre of the apical margin of sternite 9 produced into a long, bifurcate spine. Phallus straight, mem- braneous, with a pair of heavily fringed, down- ward curved plates. 9 abdomen terminates bluntly, but ventral plate narrow and more elongate than in U. len- tum. Length of anterior wing: 8 6-8 mm; 5 6-5-8 5 mm. Ulmerochorema tasmanicum Type material: Holotype <$ (T4820), allotype 9 (T4821), 15 $ 15 $ paratypes (T4822- T4851), South Esk River nr. Evandale, Tas., 1 Mar. 1967, A. Neboiss (NMV); 10 6 5 14 Ulmerochorema tasmaniewn (Mosely) Figures 80-81 Type material: Type s New Norfolk, Tas., Mar. Anachorema tasmanica Mosely in Mosely and Kim- 1938, J. W. Evans (BMNH). Type seen. mins, 1953:453. Material examined: Tasmania Ulmerochorema — 1 o* Derwent River tasmanicum, Neboiss, 1962:543. 2 km NW Derwent Bridge, 12 Feb. 1971; 4 £ 4 ? TASMANIAN CADDIS-FLIES 23 South Esk River nr. Evandale, 1 Mar. 1967; 1 c? Mer- sey River, Liena, 16 Nov. 1972; 3 c? Leven River nr. Heka, 17 Nov. 1972; 3 ? Macquarie River 8 km W Campbell Town, 9 Nov. 1972; 185 c? 153 ? Huon- Picton River junction, 18 Feb. 1967, 3 o* same loc, 15 Nov. 1972; 5J4? Huon River nr. Blakes Opening, 9 Feb. 1966; 2 ? Huon River nr. Scotts Peak, 9 Feb. 1965; 3 o* Wedge River, 17 Feb. 1971; 1 15 Ulmerochorema rubiconum Neboiss Figure 82 Ulmerochorema rubiconum Neboiss, 1962:542. General appearance, colour and wing vena- tion similar to that of U. tasmanicum, but can Material examined: Tasmania—55 c? 78 ? St. Co- be separated by the differences in the male and lumba Falls, Pyengana, 21 Feb. 1971; 8 1 9 St. Patrick's River, Targa, 22 Feb. 1971; 29 S £ genitalia: below segment 10 there are two 13 9 Derwent River 2 km NW Derwent Bridge, 7 processes on either side, the dorsal one longer, Nov. 1972; 13 <$ 2 ? Franklin River 20 km SW Der- Flowerdale River, depressed laterally, slightly produced down- went Bridge, 20 Feb. 1971; 4 d* 4 ? Meunna, 4 Nov. 1972; 12 1 Condominion Creek, 15 Feb. 1971; 7 sternite 9, flattened dorso-ventrally, bifurcate 4 c? $ 6 Wedge River, 17 Feb. 1971; 1 <$ 1 ? 10 mis W of at apex, tapering to pointed apex. Heavily Maydena, 25 Feb. 1967; 1 rf Ouse River 5 mis W of fringed plate below phallus. Miena, 28 Feb. 1967; 1 c? Duck River 6 km SW of Roger River, 29 Nov. 1974; 3 Genus Ethochorema gen. n. 5. Ventral process on segment 6 larger than Type species: Taschorema nesydrion Neboiss, the short and blunt process on 1962. segment 7 . . (Qld., N.S.W.) ochraceum — . Ventral process on segment 6 absent, This genus has been erected to take a group process on segment 7 slender, acutely of species previously included in genus Tascho- pointed (Tas.) nesydrion rema and having fork 1 with footstalk in the anterior wing. At present seven species ochra- 6. The processes on either side of phallus ceum, brunneum, turbidum, nesydrion, hes- without lateral spines perium, secutum and kelion are included, of (Qld., N.S.W., Vic.) brunneum which the two latter ones are described as new. — . The processes on either side of phallus Three species are known from Tasmania and with one large and several small lateral they all appear to be endemic. spines (N.S.W., Vic.) turbidum Anterior wing with Rs forked apically, forks 1, 2, 3, 4 and 5 present; fork 1 with footstalk, 16 Ethochorema secutum sp. n. fork 2 sessile, forks 3, 4 and 5 with footstalks; Figures 83-85 discoidal cell closed, elongate, moderately long, some species with a pair of long bristles on the Anterior wings brown, unicolorous; males anal margin. Posterior wing with Sc and R x with long fold along C-Sc extending to almost ending separately at the wing margin; forks 1, the full length of the wing, densely covered with 2, 3 and 5 present, fork 2 sessile, others with long hairs; anal margin without a pair of long footstalks; discoidal cell open; in males a nar- bristles. Posterior wings without cell-like struc- row, elongate cell-like structure between Cu 2 ture between Cu2 and A x and without andro- and Ai, androconia on A 2 and As either conia. present or absent, depending on species. Lateral $ genitalia with segment 9 narrow dorsally; filament on sternite 5 present in male; the ridge segment 10 broad, parallel-sided, somewhat on sternite 5 in female terminates at lateral truncate apically, on either side at the base a margin. pair of short, apically rounded processes. Su- Spurs 2:4:4. perior appendages slender. Phallus membrane- ous, bearing two groups of outwards directed KEY FOR SEPARATING THE SPECIES spines at apex, and a pair of dark, strongly (Males only) sclerotized plates mesally. Inferior appendages gradually 1. Posterior wings with cell-like structure curved inwards, slightly widened basally; a pair of finger-like processes arising between Cu 2-A x present 2 from mesal plate, each bearing small, — . Posterior wings without cell-like a acute structure secutum tubercle on ventral margin near the apex. Lateral filament on sternite 5 short, apex 2. Anterior wing with footstalk of fork 1 rounded; there are no ventral processes on short 3 either sternite 6 or 7. — . Anterior wing with footstalk of fork 1 long, at least 3 times longer than cross- $ unknown. vein closing discoidal cell 4 Length of anterior wing: $ 8-5-9 mm. 3. In male the anal margin of anterior wing Type material: Holotype $ 10 mis E of Stra- with han, two long bristles; cell-like structure 6 Feb. 1967, E. F. Riek (ANIC); 3 $ in posterior wing large (Vic.) hesperium paratypes and 1 $ pupa (fully developed), In the same — . male anal margin of anterior wing date (ANIC; NMV—T5136, T5595); without bristles; cell-like structure in pos- 6 s same loc. 20 Feb. 1963, 1. F. B. Common terior wing small (Tas.) kelion and M. S. Upton (ANIC; NMV—T5 137, 4. Anterior wing with costal fold ... 5 T5138). . Anterior wing without — costal . fold . 6 Distribution: Tasmania—NW province. TASMANIAN CADDIS-FLIES process. Phallus robust, curved downwards. Lateral filament on stcrnite 5 with apex rounded; a small, acutely pointed, ventral pro- cess on sternite 7. S abdomen with fringe of long hairs along posterior margin of sternite 7; a small, acutely pointed, ventral process on sternite 6. Length of anterior wing: 6 8-9*5 mm; 9 8- 10 5 mm. 17 Ethochorema nesydrion (Neboiss) Figures 86-95 Taschorema nesydrion Neboiss, 1962:562. Originally described from a single female, this species has been found to be rather com- mon and occurs throughout Tasmania; it is not known from the Australian mainland. The mid-anterior wart on the head divided the male with central line. Anterior wing in Type material: Holotype 2 Cradle Mtn., Tas., with dense, long with C-Sc folded and covered 16 Jan. 1917, R. J. Tillyard (BMNH). Type without hairs on the upper surface, anal margin seen. the pair of long bristles; posterior wing with Material examined: Tasmania— 10 c? 12 2 Wald- cell-like structure small. heim, Cradle Mtn. Nat. Park, 7 Feb. 1971; 3 Iris River tributary 15 km N of Cradle Mtn., 13 Dec. River, 1974; 1 c? 10 ? Duck River 6 km SW of Roger of 29 Nov. 1974; 6 c? 2 ? Dip River Falls 10 km S Mawbanna, 29 Nov. 1974; 2 ? Collingwood River bridge, Lyell h-way, 9 Dec, 1974; 1 ? Arrowsmitri (reek, IK km SW Derwcnt Bridge, 9 Dec. 1974. All above specimens collected by A. Neboiss (NMV); Riek L E. F. Riek (ANIC); 1 ? Lake Dobson, 20 Feb. 1967, Feb. E. F. Riek (ANIC); 1 Distribution: Tasmania- -all provinces except E province. 18 Ethochorema kelion sp. n. Figures 96-98 Anterior wing brown, distinctly and irregu- larly mottled with pale-yellowish spots, traces of mottling also on distal section of the posterior wing. The anterior wings of the male without a fold along C-Sc, a pair of bristles on the anal margin present. Posterior wing with cell-like structure small, androconia on A- present, but 1 paratype Mur- there are only long hairs on As- The antero- 1971, A. Neboiss (NMV); 6 mcsal wart on the head not divided by central chison River, Tas., 5 Feb. 1967, E. F. Riek line; front ocellus large. (ANIC). $ genitalia: segment 10 upcurved, widened Distribution: Tasmania—NW and SW pro- apically, two fingcrlike processes at the base. vinces. Superior appendage stout; inferior appendage Genus Taschorema Mosely large, robust, widened at the base, bearing a group of spines on the inner surface. Phallus Taschorema Mosely, 1936a:422; Ross and King, 1951:503; Moselv and Kimmins, 1953:434; Kim- with proximal half wide, gradually tapering, mins, 1960:184; Neboiss, 1962:552. distal half very slender; a pair of robust, api- Notiohiosis Banks, 1939:499; Mosely and Kimmins, cally downturned parameres. Lateral filament 1953:444; Ross, 1956:125. on sternite 5 slightly curved, apex rounded; Type species: Taschorema asmana Mosely, ventral process on sternite 6 stout, with a 1936. few strong spines at the apex; ventral process After removing seven species from the genus on sternite 7 small. and placing them into two new genera, only 9 abdomen tapers gradually to a narrow, seven other species remained, and even these rounded apex; the ridge on sternite 5 ventrally are not all acceptable without certain objec- produced to a moderately slender ventral pro- tions, but being from localities outside Tas- cess, a few strong spines at the apex; ventral mania, further analysis of their characters is process on segment 6 small; posterior margin beyond the scope of this work. The genus now on sternite 7 slightly produced and covered with contains a total of nine species, two of them group of long hairs. being described as new, and all but three, are Length of anterior wing: <$ 8-5-10-5 mm; 9 from Tasmania. 1 1 mm. The genus is characterized by having forks 1 sessile, Type material: Holotype <$ (T4852), allotype and 2 in the anterior wing long and 9 (T4853) Condominion Creek, Tas., 15 Feb. the former usually very narrow; posterior wing TASMANIAN CADDIS-FLIES 27 with cell-like structure between Cu 2 and A x in structure on posterior wing between Cu 2 and A x male. Other characters as described previously. long; androconia on A and present. It 2 A ;I can Spurs 2:4:4. easily be separated by the distinctive male geni- KEY TO SPECIES OF TASCHOREMA talia as shown in the illustrations. (Males only) $ genitalia with the plates at the base on segment 10 with apices 1. Radius forked distally 2 pointed and curved up- — Radius ward and a small wart on proximal section. In- . not forked . . , (Vic.) rugulum ferior appendages stout, broad at base, 2. Fork 1 in posterior wing narrow- ing slightly to obliquely truncate absent „ (W.A.) pallescens apices, inner angles triangular, apices acute; — Fork 1 in proximal half . posterior wing present, with of inner margins bearing long, coarse, basally footstalk 3 directed spines. Phallus very long, curved down- 3. Anterior wing with two long bristles at the wards. Ventral process on sternite 6 short, anal margin 5 ro- Anterior bust, bearing a few stout spines near apex; pro- — . wing without bristles at the anal cess on sternite 7 very long, margin 4 slightly upcurved. 4. Anterior wing folded 9 abdomen terminates into a blunt, rounded along C-Sc (Vic.) kimminsi apex; the ridge on sternite 5 terminates at the lateral margin, but vent-rally extends to — . Anterior wing without a fold along a strong ventral C-Sc ... (N.S.W., Vic, Tas.) evansi process; ventral process on sternite 6 smaller than 5. Cell-like structure in posterior wing be- that on sternite 5; posterior margin of sternite 7 produced into a central tween Cu 2 and At long 7 triangular Cell-like projection, covered with short, dense pubes- — . structure in posterior wing be- cence, on either side of which there tween Qla and Aj short, rather incon- are some- what spicuous 6 squarish lobes. 6. Inferior appendages Length of anterior wing: $ 10-12 mm; 9 11- long 12 5 mm. • - (Tas.) viridarium — . Inferior appendages short (Tas.) pedunculaium 7. Processes at the base of segment 10 short and broad 8 — . Processes at the base of segment 10 long and slender (Tas.) jerulum 8. Anterior wing uniform in colour (Tas.) asmanum — . Anterior wing with basal half dark, distal half lighter in colour . . (Tas.) apobamum 19 Taschorema asmanum Mosely Figures 99-102 Taschorema asmana Mosely, ]936a:422; Mosely and Kimmins, 1953:434. Taschorema asmanum, Neboiss, 1962:564. Similar to the two other closely related spe- cies apobamum and jerulum described in this paper, asmanum is characterized by a narrow sessile fork 1 on the anterior wing; in the male the lateral filament on sternite 5 terminates with a small apical hook; two long bristles on the anal margin of the anterior wing; cell-like 28 ARTURS NEBOISS 5 ends at the lateral margin, Type material: Type $ Great Lake, Miena, ridge on sternite to a strong ventral pro- las., Jan. 1931 (BMNH). Type not seen. and ventrally extends process on m- cess; very small pointed ventral Material examined: Tasmania— 1 c? 3 5 St. Colli 1 posterior margin of sternite 7 in the ba Falls, Pyengana, 21 Feb. 1971; 1 d 4 km F of sternite 6; Liena, small creek, 17 Nov. 1972; 2 <$ 1 9 Waldhcim, middle extends to a broad, triangular projec- Cradle Mtn. Nat. Park, 7 Feb. 1971; 2 c? 20 9 Dove tion, which is covered with short, dense pubes- River, Cradle Mtn. Nat. Park, 14 Dec. 1974; 3 c? 2 9 Dcrwcnt River 2 km NW Derwent Bridge, 12 Feb. cence, on either side of it, there is a broad, dis- 1 20 Derwent Bridge, 1971; 15 Feb. 1971; 1 c? West Arthur Plains, 7 Feb. 1965; 2 9 Russell Falls, Nat. Park, 15 Nov. 1972; 1 ? Cashion Creek Cave, Florentine Valley, 29 Dec. 1964, T. Goede; 1 c? 7 9 nr. Marakoopa Caves, 15 Dec. 1974; 2 9 Hot Springs Creek, Hastings Caves, 14 Nov. 1972; 1 g Lilydale, creek 2 km N, 16 Dec. 1974. All specimens collected by A. Neboiss, unless staled otherwise (NMV). 1 S Fossey River 10 mis S Hellycr Gorge, 5 Feb. 1967, E. F. Riek (AN1C); 2 o* 2 9 Forth Falls, 28 Jan. 1967, F. F. Riek (AN1C); 7 c? 8 ? Lake Dobson, 20 Feb. 1967, F. F. Riek (ANIC): 1 2 10 mis E Strahan, 6 Feb. 1967, E. F. Riek (ANIC). Other recorded localities: Derwent Bridge. Distribution: Tasmania—NE, C, N, NW, SW and SE provinces. 20 Tascliorema apobamitm sp. n. Figures 103-109 This is one of the largest Rhyacophilid spe- cies not only in Tasmania, but also in Australia. The two tone anterior wings in male with basal half rich, dark-brown and paler light-brown distal half, makes the species easily recogniz- able; two long bristles on anal margin present, the cell-like structure on posterior wing be- tween Cu- and Ai long; androconia on A^ and Type material: Holotype 6 (T4901), allo- A.i present, lateral filament on sternite 5 with type 9 (T4902) 6 & 6 9 paratypes (T4903- small apical hook. The anterior wings in female T49 14) St. Patricks River, Targa, Tas., 22 Feb. are concolorous rich brown. 1971, A. Neboiss (NMV). $ genitalia of similar plan to armarium but Other material examined: Tasmania—4 9 Great differs in details. The plates at the base of seg- Forester River 5 km NW Forester, 11 Nov. 1972; ment 10 pointed apically with small peglike 1 £ 2 •$ St. Coltimha Falls, Pyengana, 21 Feb. 1971; Bridge, wart on proximal section. Inferior appendages 2 9 Derwent River 2 km NW Derwent 12 Feb. 1971; 14 r? 3 9 Franklin River 20 km SW Der- long, curved, broad at base, constricted in the wen' Bridge, 11 Feb. 1971; 5 9 Mersey River, Liena, middle and widened to a truncate apex; a few 16 Nov. 1972; 2 9 Russell Fills, Nat. Park. 15 Nov. 1972; 1 9 Waldheim, Cradle Mtn. Nat. Park, 7 Feb. inward and basally directed long spines arise 1971; 31 9 Hellver River Gorge. 9 Feb. 1971: 2 r? 5 9 from the upper inner margin of the basal sec- Huon River Crossing. 16 Feb. 1971; 44 c? 98 9 Huon- tion. Phallus long, curved downward. Short Picton River junction, 18 Feb. 1967; 2 9 Huon Plains nr. Scotts Peak. 2 Feb. 1965; 5 9 Huon River nr. ventral process on sternite 6 with few strong, Blakes Opening, 9 Feb. 1966; 1 9 West Arthur Plains, apical spines; ventral process on sternite 7 6 Feb. 1965; 28 9 Cracroft River, 8 Feb. 1966; 1 S long, curved upwards. (pupa) Styx River. Westerwav, 23 Nov. 1972, J. Blyth; I 9 Collingwood River 30 km E of Gormans- gradually tapers 9 abdomen to a blunt apex, ton, 6 Nov. 1972; 23 9 Collingwood River bridge, TASMANIAN CADDIS-FLIES Lyell h-way, 9 Dec. 1974; 4 ? Dove River, Cradle Pa DeC " H 1974; X ' ? National Park, 20 pFebk io& , oH 1967; 5 ? Leven River, Heka, 17 Nov. 1977- 5 Q P Rl F 1,S X Dec 1974 ?' ' AH ' - specimens A xfu ? collected by A. Neboiss unless stated otherwise (NMV) 1 r? North Esk River 20 mis E Launceston, 1 Mar. 'l967, <* °use River 5 mIs W Miena oi Febp k ^oi^Wi I, 28 1967; 7 c? 8 $ Franklin River, 10 Feb. 1967 "• §«* (ANIC); 1 ? 10 mis E. Strahan, 6 Feb! 967 E F R ek (ANIC); ] 4 Fort Falls, ',o o o <$ ? " 28 Jan 1967,^ E, F.; Riek (ANIC); 10 rf 21 ? Huon-Pic- er 3 c Ion 17 Feb 1967 E F ' " > - - Riek (ANIC); V^^ n » f. y or Feb 1967 * R F Riek &> - t&Jr^yl( ?\ orFDerwent i- ' h- ^ rlvl£j ? R«ver, 12 Feb. 1967, E. F. ( A 3 ; 13 9 Murchison River, 5 Feb. , ,^ ^71967, E.* TOF. Riek (ANIC). Distribution; Tasmania—all provinces except E province. 2 1 Taschorema jerulum sp. n. Figures 110-114 This species is very similar to asmanum but can be separated by the details of the male genitalia as illustrated. At the base of segment 10, instead of a short, pointed plate, there is a long, finger-like process, widened at the base, from which arises a long, thin, slightly down- curved rod. Inferior appendages short, broad Neboiss (NMV); 1 $ paratype (T4929) at base, in lateral view gradually tapering to- Wedge River, Tas., 17 Feb. 1971, A. Neboiss wards the apex; inner margin flattened, curved (NMV); 1 s paratype (T4930) North Esk and terminating with River, inwardly directed blunt Blessington, Tas., 1 Mar., 1967 (with angle. Phallus long, curved downward. Lateral pupal and larval skin), A. Neboiss (NMV). filament on sternite 5 with small apical Other hook, material examined: Tasmania— 1 °_ Derwent River 2 km small ventral process on sternite 6, a long one NW Derwent Bridge, 12 Feb. 1971; 16 $> Leven River, Heka, 17 Nov. 1972; 3 $ Dip River on sternite 7. Anterior wing with two long Falls, 1 Dec. 1974; 23 ? Scamander River, Upper bristles on anal margin. Posterior wing with Scamander, 9 Nov. 1972; 10 J Mersey River, Liena, cell-like structure long, androeonia on and 16 Nov. 1972; 2 ? Huon-Picton River junction, 18 A 2 Feb. 1967; 2 ? same loc, 15 Nov. 1972; 1 ? Hellyer A 3 present. River Gorge, 9 Feb. 1971; 1 $ Ouse River 8 km W 9 abdomen terminates into a broad, blunt of Miena, 28 Feb. 1967. All specimens collected by 1 A. Neboiss (NMV). 1 Bronte Lagoon, apex, the ridge on sternite 5 terminates at the J 15 Feb 1967, E. F. Riek (ANIC); 1 <$ Bronte—Bradys lateral margin, but ventrally extends to a strong Canal, 27 Feb. 1967, E. F. Riek (ANIC); 7 ? Forth ventral process; ventral process on sternite 6 Falls, 28 Jan. 1967, E. F. Riek (ANIC); 17 $ Evandale, 1 Mar. 1967, E. F. Riek (ANIC); small; sternite 7 with deep, rounded depression 3 c? 26 § Derwent Bridge, 12 Feb. 1967, E. F. Riek (ANIC)- in the middle, distal half of the sternite broad, 1 J Ouse River 8 km W Miena, 28 Feb. 1967 E F ' Riek (ANIC). ' ' produced posteriorly, fringed with fine hair and Distribution; Tasmania all provinces a pair of small, rounded lobes distally. — except SE province. Length of anterior wing: $ 10-12 mm; 9. 11-13 mm. 22 Taschorema viridarium Neboiss Type material: Holotype s (T4915), allo- Figures 115-117 type 9 (T4916),2 s 8 $ paratypes (T4917- Taschorema viridarium Neboiss, 1962: 563. T4926) South Esk River, Evandale, Tas., 1 This is one of the smallest species in the Mar. 1967, A. Neboiss (NMV); 2 3 para- genus, in general appearance light yellowish- types (T4927-T4928) Derwent River 2 km brown. The anterior wing in male with two NW Derwent Bridge, Tas., 12 Feb. 1971, A. long bristles on the anal margin; posterior wing 30 ARTURS NEBOISS with short, cell-like structure between Cu 2 and Type material: Holotype $ , allotype 9 Broad A], androconia on A2 and A 3 present. River nr. Lake Dobson, Mt Field Nat. Park, 6 genitalia characterized by having the Tas., 6 Nov. 1955, T. E. Woodward (QM). plates at the base of segment 10 partly fused Type seen. with it, thus forming a pair of broad lobes. In- Material examined: Tasmania— 1 c? St. Patricks ferior appendages in lateral view broad at River, Targa, 22 Feb. 1971; 15 6 1 ? Huon-Picton River junction, 18 Feb. 1967; 1 1 3 Wedge River, proximal half, from about the middle gradually c? 9 17 Feb. 1971; 17 c? Huon River Crossing, 16 Feb. tapering apically, dorso-ventrally flattened and 1971; 2 r? Condominion Creek, 15 Feb. 1971; 1 c? curved inward to a blunt angle; inner surface West Arthur Plains, 3 Feb. 1965; 2 o* Junction Creek, West Arthur Plains, 7 Feb. 1966; 2 Dove River, with basally directed stout spines. Phallus slen- c? Cradle Mtn. Nat. Park, 14 Dec. 1974; 1 c? Dip River der, straight, with lower margin apically form- Falls 10 km S of Mawbanna, 1 Dec. 1974. All speci- ing an upcurved hook, upper margin terminat- mens collected by A. Neboiss (NMV). 1 c? Murchi- son River, 5 Feb. 1967, E. F. Riek CANIC); 1 either side of phallus are stout, rod-like para- (ANIC); 2 o* Lake Dobson, 20 Feb. 1967, E. F. Riek (ANIC). meres, turned downwards at distal end, upper angle with a pair of long, stiff bristles and a Other recorded localities: Tycnna River nr. pair of shorter ones near apex. Very long ven- Tyenna; Dove River, Cradle Mtn. tral process on sternite 7, a short and stout one Distribution: Tasmania—N, NW, SW and SE on sternite 6. Lateral filament on sternite 5 provinces. terminates with a minute point. 9 abdomen with ridge on sternite 5 ter- 23 Taschorema pedunculatum Jacquemart minating at the lateral margin, ventrally de- Figures 118-122 veloped into a strong ventral process, a smaller Taschorema pedunculata Jacquemart, 1965b: 35. one on sternite 6; posterior margin of sternite 7 in the middle broadly produced. Irrespective of the discrepancies between the original illustrations and the figures Length of anterior wing: $ 6-7 5 mm; 9 6-8 presented mm. here from specimens collected at Cradle Mtn. and Condominion Creek, it was found that they agree with the holotype and represent the same species. The most important deviations are the shape of the phallus, the dorsal view of seg- ments 9 and 10; the shape and position of hya- line areas and the omission of one of the anal veins in the anterior wing from the original drawing (Jacquemart 1965a, fig. 27D). This rather small, dark-brown species appears to be closely related to T. viridarium, particu- larly so, the structure of phallus, parameres and the small, cell-like structure on the pos- terior wing; androconia on A2 and A 3 present. 3 genitalia with segment 10 fused with elon- gate basal plates, which are indicated by sclero- tized bands and somewhat indistinct lateral lobes. Inferior appendages short, broad, curved inward, bilobed apically, group of several stout spines on the inner surface near the apex. Superior appendages longer than the inferior ones. Phallus with lower margin ending with an upcurved hook, upper margin with a pair of straight projections. The phallus is flanked on TASMANTAN CADDIS-FLIES 31 either side with strong, apically downturned sternite 5 terminates with a minute point; ven- parameres. Lateral filament on sternite 5 ter- tral processes on sternites 6 and 7, the former minates with tiny hook; a small ventral process slightly larger and with a few stout apical spines. on sternite 6, a long one on sternite 7. In the male the anterior wing without bristles on 9 unknown. anal margin; posterior wing with cell-like struc- Length of anterior wing: $ 6-7 mm. ture between Cu and 2 A x ; androconia on A2 and A . Taschorema pedunculatum 3 9 abdomen terminates with darkly pigmen- ted tergites 8 and 9. Length of anterior wing: 3 8-10 mm; 9 8-11 mm. Type material: Holotype $ Cradle Mtn. Tas., 12 Jan. 1923 (IRScNB). Holotype dissected and mounted on four microscope slides. Type seen. Material examined: Tasmania—2 o* Waldheim Cradle Mtn. Nat. Park, 7 Feb. 1971; 2 <$ Con- Type material: Type Lake Leake, dommion Creek, 14 Feb. 1971, A. Neboiss (NMV). $ Tas., Feb. 1937 (BMNH). Type not seen. Distribution: Tasmania—NW and SW pro- Material vinces. examined: Tasmania—5 <$ 39 ? St Patricks River, Targa, 22 Feb. 1971; 2 d1 7 9 Great Forester River 5 km NW Forester, 11 Nov. 24 Taschorema evansi Mosely 1972; 2 9 Scamander River, Upper Scamander, 9 Nov. 1972* Figures 123-125 2 $ Leven River, Heka, 17 Nov. 1972; 1 <$ 2 ? Flower- dale River, Meunna, 4 Nov. 1972; 3 c? 4 ? Derwent Taschorema evansi Mosely in Mosely and Kimmins. River 2 km NW of Derwent Bridge, 12 Feb. 1971; 1953:440; Neboiss, 1962:561. 1 National o* Park, Tyenna River, 19 Feb. 1971; 2 $ Corinna, 5 Nov. A dark brown, medium size species with 1972; 2 ? same loc, 13 Mar. 1971 G. Minko; 1 c? 8 $ Huon River Crossing, 16 Feb' widespread distribution. 1971; 1 ? same loc., 8 Feb. 1965; 1 ? Huon River nr $ genitalia characterized by the large in- Scotts Peak, 8 Feb. 1965; 3 ? Lake Pedder, 31 Jan. 1965; 3 <$ Condominion Creek, ferior appendage, widened toward the apex; 15 Feb. 1971; 6 $ Collingwood River bridge, Lyell h-way, 9 Dec 1974* on the inner surface of which there is a ridge Lilydale, 9 J a creek 2 km N, 16 Dec. 1974- 4 $ Hellyer River with strong spines and another group of spines Gorge, 9 Feb. 1971; 1 $ same loc., 12 Dec. 1974; 5 ? Duck River 6 km SW of Roger on the upper distal margin. Lateral filament on River 29 Nov. 1974; 1 cf 1 ? Hogarth Falls, Strahan, 10 — ARTURS NEBOISS Dec. 1974; 1 $ Lake St. Clair, Derwent Basin, 6 Dec. teral filament on sternite 5 absent; strong ven- 1974; 4 cJ Tooms Lake, 4 Dec. 1974; 8 $ 7 ? Dip River tral process on sternite 6, a long, rather broad Falls 10 km S of Mawbanna, 1 Dec. 1974; 1 $> Wedge River, 17 Feb. 1971; 2 2 Cracroft River, 8 and somewhat flattened ventral process on Feb. 1966; 1 ? Mersey River, Liena, 16 Nov. 1972; 1 sternite 7. 2 St. Columba Falls, Pyengana, 21 Feb. 1971. All specimens collected by A. Neboiss, unless stated $ abdomen terminates with downsloping otherwise (NMV). 4 <$ Derwent Bridge, 12 Feb. 1967, truncate apex; sternite 8 extends posteriorly E. F. Riek (ANIC); 1 <$ Bronte Lagoon, 15 Feb. almost to the end of abdomen. 1967, E. F. Riek (ANIC); 8 ^ 12 ? Forth Falls, 28 Jan. Length 1967, E. F. Riek (ANIC); 1 o* 3 $ Hellyer of anterior wing: g 9-12 mm; $ 10-13 Gorge, 4 Feb. 1967, E. F. Riek (ANIC). mm. Distribution: Tasmania— all provinces; Vic- toria; South Australia. Genus Ptychobiosis gen. n. Type species: Notiobiosis nigrita Banks, 1939. Among the species previously included in the genus Taschorema, two species nigrita and rieki showed similarities not found among the others, and therefore they are now segregated to form a new genus. Ocelli present; anterior wings with radius straight, not forked distally; forks 1, 2, 3, 4 and 5 present, forks 1 and 2 long, sessile, others with footstalks; discoidal cell closed; in pos- terior wing Sc merges with Rs shortly before wing margin. In male a large pouch at the proximal half of A 2 in the posterior wing. In females the ridge on sternite 5 terminates at the lateral margin; mid-posterior margin of ster- nite 8 extended almost to the end of abdomen. Spurs 2:4:4. Only one species in Tasmania. 25 Ptychobiosis nigrita (Banks) Figures 126-130 Type material: Type g Mt Spurgeon, N Qld. 1100-1200 m; 26 Jul. (Darlington). Originally Notiobiosis nigrita Banks, 1939:500; Mosely and Kimmins, 1953:446; Ross, 1956:114, 125. deposited in MCZ, later transferred to ANIC. Taschorema nigra Mosely in Mosely and Kimmins Type not seen. 1953:436. Taschorema nigrita, Kimmins, 1960: 1 84; Neboiss Type s of Taschorema nigra 1962:565. Mosely, Na- tional Park, Qld. 900 m, 21 Dec, This rather large, dark blackish-brown spe- 1921 (BMNH). Type not seen. cies is widely distributed throughout the Aus- tralian Material examined: eastern states as far as North Queens- Tasmania— 1 River, 12 Feb. 1967, E. F. Riek (ANIC); 1 <$ Hellyer Gorge, 4 Feb. 1967, E. F. Riek (ANIC); 1 Genus Koetonga Neboiss Koetonga Neboiss, 1962:567. Type species: Koetonga clivicola Neboiss, 1962. Wing venation regular in anterior wing, with forks 1,2,3,4 and 5 present; forks 1 and 2 very long, sessile, others with footstalks; discoidal cell closed; an additional oblique cross- vein lo- cated between Aj and A 2 ; venation in posterior wing irregular and differing in sexes. Ventral process in male on sternites 6 and 7; in female on sternites 5 and 6; the ridge on sternite 5 ter- minates at the lateral margin. Spurs 2:4:4. 26 Koetonga clivicola Neboiss Figures 131-133 Anterior wings densely covered with fine Koetonga clivicola Neboiss, 1962:567. hairs, but the coarse erect hairs absent; forks Insect robust, anterior wings dark-brown 1, 2, 3, 4 and 5 present, all with footstalks, with yellowish mottling, posterior wings con- except fork 2; footstalk of fork 1 very short; colorous yellowish with aberrant venation in discoidal cell open in male, closed in female; sexes as figured. The Tasmanian specimens posterior wing with forks 1, 2, 3 and 5 present, agree well with original description. fork 2 long, sessile, others with footstalks; dis- Length of anterior wing: £ 11-12 mm; ? 12- coidal cell open in both sexes; cross-vein m-cu 14 mm. strongly convex. The abdomen in females ter- minates in a long, slender, upcurved oviscapt, a Type material: Holotype £ , allotype $ Tim- bertop nr. Merrijig, Vic, 5 June 1958, I. F. pair of small cerci at apex; male genitalia with Edwards (ANIC). Type seen. segment 10 long and slender, inferior appen- dages two-segmented, coxopodite stout, longer Material examined: Tasmania—6 <$ Huon River Crossing, 16 Feb. 1971; 2 ? West Arthur Plains, 3 than harpago; lateral filament on sternite 5 Caves, Feb. 1965; 1 o* Hot Springs Creek nr. Hastings present. 14 Nov. 1972; 7 c? 1 S Duck River 6 km SW of Roger Spurs 2:4:4. River, 29 Nov. 1974; 5 <$ nr. Marakoopa Caves, 15 Dec. 1971; 13 o* 1 ? Lilydale, a creek 2 km N, 16 Dec. 1974; 1 c? Dove River, Cradle Mtn. Nat. Park, KEY TO SPECIES OF MORUYA 14 Dec. 1974. All specimens collected by A. Neboiss (Males only) (NMV). ] 6* 10 mis E Strahan, 6 Feb. 1967, E. F. Riek (ANIC). 1. The processes arising at the base of seg- Distribution: Tasmania—N, NW and SW pro- ment 10 asymmetric charadra Victoria. vinces; New South Wales; — . The processes arising at the base of seg- ment 10 symmetric 2 Tribe Psyllobetini trib. n. 2. The short upper process at the base of Genus Moruya Neboiss segment 10 simple, the lower one long, Moruya Neboiss, 1962: 569. with apex turned slightly Type species: Moruya charadra Neboiss, 1962. upward tasmanica 34 ARTURS NEBOTSS — . The short upper process at the base of seg- ment 10 two-branched, the lower one long, with apex turned downward .... opora 27 Moraya charadra Neboiss Figures 134-139 Moray a charadra Neboiss, 1962:570. Anterior wing in male with long, distally directed hairs along the costa; posterior wing with large area of short, upright setae on dorsal surface; lateral filament on sternite 5 stout, abruptly tapered to fine pointed, apex; ventral process on sternite 7 short, flat. S genitalia with segment 10 long, rounded apically; at the base arise two pairs of assy- metric processes, a short upper one, two- branched, and a long lower one. The entire structure is twisted to the right side where both processes are longer. Inferior appendage with coxopodite robust, an elongate group of spines on the inner surface near the base; harpago short, somewhat twisted. Phallus long, slender, with an upward directed, slightly curved spur All specimens collected by A. Neboiss (NMV). 1 c? in the middle. Murchison River, 5 Feb. 1967, E. F. Riek (ANIC); 1 Hellyer Gorge. 4 Feb. 9 abdomen terminates into a slender, up- d 1967, E. F. Riek (ANIC); 2 c? 7 ? Lake Dobson, 20 Feb. 1967, E. F. Riek curved oviscapt. The outline of the inner struc- (ANIC). ture near the base visible in cleared prepara- Other recorded localities; Dove River nr. Lake tions, is helpful in species identification, but Dove; Hugel River, Lake St. Clair. otherwise the smaller size and fork 5 in an- Distribution; Tasmania—NE, C, NW, SW and terior wing, more or less gradually widened to SE provinces. full width, separate the female of this species from opora. 28 Moraya opora Neboiss Figures Length of anterior wing; $ 5-6-5 mm; $ 6-7 140-145 mm. Moraya opora Neboiss, 1962: 571. Type material; This species is very closely Holotype $ , allotype 9 Broad related to M. River nr. Lake Dobson, Tas., 6 Nov. 1955, charadra and differs only by its slightly larger T. E. Woodward (QM). Type seen. size and details of genitalia structure, and by the large area with short, upright setae present on Material examined; Tasmania—3 <$ 1 $ Lake Dob- son, 20 Feb. 1967; 1 <$ Henty River 12 km NW the posterior wing. Queenstown, 10 Feb. 1971; 2 t? 1 2 Wedge River 17 6 genitalia symmetric; the upper process at Feb. 1971; 2 S 5 ? Waldheim, Cradle Mtn. Nat. Park, 1 the base of segment 10 short, 7 Feb. 1971; 1 J Fisher River, Pencil Pine Grove be- two-branched, the low Lake Mackenzie dam, 15 Dec. 1974; 1 £ Pencil lower process longer than superior appendage, Pine River 6 km N of Cradle Mtn., 13 Dec 1974* apex abruptly turned downward. Inferior apend- 2 9 abdomen terminates into a slender, up- 29 Moruya tasmanica (Jacquemart) curved oviscapt; the outline of inner structure comb. nov. near the base differs from that of charadra as Figure 146 shown in fig. 145. Fork 5 in anterior wing Psytlobetina tasmanica Jacquemart, 1965b:45. rather abruptly widened to full width. Length of anterior wing: 6 7 5-9 mm; 9 The type $ which is dissected and mounted 8-9 5 mm. on three separate microscope slides, has been re-examined and figured. The wing venation as well as the basic structure of male genitalia, in- dicate that this species belong to the genus Moruya. Anterior wing with forks I, 2, 3, 4 and 5 present, discoidal cell open. Posterior wing with forks I, 2, 3 and 5 present, fork 1 long, with very short footstalk. Due to the nature of mounting, some details of the wings are ob- scure, but it appears that there are no short, up- right setae on the posterior wing. Lateral fila- ment on stcrnite 5 stout, tapered apically and terminating with laterally directed hook. S genitalia with segment 10 and superior appendages not exceeding the length of in- ferior appendages. The short upper process at the base of segment 10 simple, turned down and outward; the lower process long, but with slightly shorter superior appendage, gradually turned upward at the apex. Inferior appendage Type material: Holotype 9 Ouse River nr. Great Lake, Tas., 2 Apr. 1960, E. T. Smith (ANIC). Type seen. Material examined: Tasmania— I <$ Ouse River 8 km W of Miena, 28 Feb. 1967 (description and figures of male prepared from this specimen); 1 $ Franklin River 20 km SW or Derwcnt Bridge, 11 $ Feb. 1971; 1 <$ 1 ? Derwent River 2 km NW of Der- 1 went Bridge, 12 Feb. 1971; 3 d 6 2 Huon-Picton River junction, 18 Feb. 1967; 1 Blakes Opening, 9 Feb. 1966; 1 o* West Arthur Plains, 6 Feb. 1965; 1 2 Huon River nr. Scotts Peak, 8 Feb. 1965; 1 c? Dove River, Cradle Mtn. Nat. Park, 14 Dec. 1974; 2 9 Collingwood River bridge, Lyell h-way, 9 Dec. 1974. All specimens collected by A. Neboiss (NMV). 4 $ Derwent Bridge, 12 Feb. 1967, E. F. Riek (ANIC); 3 $ Huon-Picton junction, 17 Feb. 1967, E. F. Riek (ANIC); 1 (? 4 ? Franklin River, 10 Feb. 1967, E. F. Riek (ANIC); 3 $ Ouse River 8 km W Miena, 28 Feb. 1967, E. F. Riek (ANIC). Distribution: Tasmania—NW, C and SW pro- vinces. 36 ARTURS NEBOISS robust, coxopoditc with an elongated group of 3, 4 and 5 present, the first two sessile; discoi- strong spines on the inner surface near the dal cell closed, rather short. Small, lateral, base; harp-ago short. Phallus long and slender. blister-like protuberance on the dorsal margin 9 unknown. of sternite 5 in both sexes. Female abdomen ter- Length of anterior wing: $ 9-10 mm. minates with a pair of slender, two-segmented Type material: Holotype $ Cradle Mtn. Tas., cerci. 27 Jan. 1923, A. Tonnoir. Type dissected and KEY TO TASMANIAN SPECIES mounted on three microscope slides (IRScNB). Type seen. 1. Males with ventral process on Material sternite 6 2 examined: Tasmania— 1 c? Snake Creek, Fisher River Road, 15 Dec. 1974, A. Neboiss — . Males without ventral process on (NMV). sternite 6 laparus Distribution: Tasmania—NW and C provinces. 2. Anterior margin of segment 9 angulate in male; female without ventral process on 2 Family GLOSSOSOMATIDAE Ulmer segment 8 cralus (1903) — . Anterior margin of segment 9 broadly curved in male; female with ventral pro- The family contains mostly small, dull cess on segment 8 tasmanicus coloured caddis flies, inhabiting cool, more or less swiftly flowing streams. The larvae con- 30 A gapetus ( Synagapetus ) tasmanicus struct elongate, dome-shaped cases of small (Mosely) sand grains. For many years these insects were Figures 147-152 considered as a subfamily of the Rhyacophili- dac, but Ross (1956 and 1967) demonstrated Synagapetus tasmanicus Moscly in Mosely and Kim- mins, 1953:498; Jacquemart, 1965b:39. that family rank is warranted; he also considered Agapetus (Synagapetus) tasmanicus, Ross, 1956:161. Synagapetus as subgenus of Agapetus, this be- Small, brown to dark grey-brown species. ing the only genus found in Australia, to which According to Ross this all three Tasmanian species now belong. (1956), species, to- gether with monticolus, belong to a group of Family diagnosis: Ocelli present; maxillary palpi species in which the anterior margin of segment iivc-segmented in both sexes, the first two seg- 9 in the male is broadly curved. ments short, subequal; third the longest. An- 8 genitalia: segment 10 short, tip depressed; tennae stout, shorter than anterior wings, seg- superior appendage broad at base, lying some- ment 1 enlarged, about as long as broad, seg- what parallel to the upper posterior ment 2 short, rounded; the following ones elon- margin of segment 9, triangularly pointed at apex; in- gate. Anterior wings with forks 1, 2, 3, 4 and ferior appendage in lateral 5 present, discoidal cell closed. Posterior wing aspect broadly oval, terminal tooth slightly curved, pointing with forks I, 2, 3 and 5, or 2, 3 and 5 present, up and inward. discoidal cell either closed or open. The mid- 9 genitalia: segment 8 strongly tibia and tarsus of the females are generally chitinized, laterally considerably dilated. elongate oval, with somewhat flattened Spurs 2:4:4. ventral keel-like process, a slender, anteriorly directed, internal rod arising from anterior Genus Agapetus Curtis margin; segments 9 and 10 membraneous; seg- Agapetw Curtis, 1834:217; Ross, 1956:158. ment 9 with long, slender, internal, anteriorly Subgenus Synagapetus McLachlan directed rods; segment 10 terminates with a pair of slender, Synagapetus McLachlan, 1879:484; Moscly and Kim- two-segmented cerci. Small ventral mins, 1953:498. process on sternite 6 and two patches of dark, Agapetus subgenus Synagapetus, Ross, 1956: 159. stiff bristles on sternite 7; mid-tibia and tarsus Type species: Synagapetus duhitans McLach- strongly dilated. lan, 1879. Length of anterior wing: $ 3-5-4-75 mm; $ Anterior wings densely pubescent; forks 1, 2, 4-4-75 mm. TASMANIAN CADDIS-FLIES 37 Creek 12 km SW of Roger River, 29 Nov. 1974; 2 ? Meander River 3 km N of Westburv, 16 Dec 1974; 4 1974: 1 c? 3 ? Arthur River bridge 15 km SW of Roger River, 29 Nov. 1974; 2 J Dip River Falls, 1 Dec 1974; 7 c? 1 9 Duck River 6 km SW of Roger River, 29 Nov. 1974; 1 c? 1 9 Wilson Creek nr. Hell- ver, 29 Nov. 1974; 4 with the 'Holotypc & from Burnie 26.x. 1922'. Type material: Type s New Norfolk, Tas., In the same publication on p. 3 and in the Mar. 1938, J. W. Evans (BMNH). Type not legend for fig. 32 he has quoted Kimmins as seen. the author of the species. From the text, figures Material examined: Tasmania— 1 (? 4 ? Russell and the examination of the specimen, which is Falls, Nat. Park, 23 Feb. 1967; 2 ? same loc, 5 Dec. 1972, P. Zwick; 1 o* 3 $ Ellendale, 4 Dec. 1972, P. mounted on three microscope slides, it is quite Zwick; Buxton River nr. Mayfield, 13 Nov. 2rf 25 obvious that it must be referred to Mosely's spe- 1972; 12 c? 2 $ Apsley River, Bicheno, 9 Nov. 1972; cies (Neboiss, 1974c). 4 Blessington, 1 Mar. 1967; 8 r? 9 $ Minnow River, Synagapetus crala Mosely in Mosely and Kimmins, Paradise, 17 Nov. 1972; 2 1972, P. Zwick; 1 Tyenna River, National Park, 6 rounded, inner surface with curved ridge ter- c? Dec. 1972, P. Zwick; 1 Nov. 1972; 1 2 Henty River 12 km NW of Queens- process is present; segments 9 and 10 mem- town. 12 Feb. 1971; 7 S Eckberg Creek 12 km SW braneous; segment 9 with a pair of very long, of Roger River, 29 Nov. 1974; 3 6 Duck River 6 km anteriorly directed internal rods; segment 10 SW of Roger River, 29 Nov. 1974; 1 df Snake Creek area, Fisher River Road, 15 Dec 1974; 1 £ Dove small, terminating with a pair of slender, two- River, Cradle Mtn. Nat. Park, 14 Dec 1974; 1 c? segmented cerci. Ventral process on sternite 6 Rubicon River 8 km SE of Sassafras, 2 Dec 1974; 6 J Caves. 1974; 8 1 Quamby small; mid-tibia and tarsus strongly dilated. nr. Marakoopa 15 Dec o* 2 Brook 1 km E of Goulburn Valley, 16 Dec. 1974; 3 c? Length anterior wing: $ 4-5 4 5- of mm; ? 3 2 Lilydale, creek 2 km N, 16 Dec. 1974; 32 c? 4 2 5-25 mm. Liffey River 5 km W of Liffey. 2 Dec. 1974; 2 c? Bulls Creek, Cradle Mtn. Road, 13 Dec 1974. All speci- mens collected by A. Neboiss unless stated otherwise (NMV). 1 o* Fossey River 10 mis S Hellver Gorge, 5 Feb. 1967, E. F. Riek (ANIC); 1 32 Agapetus (Synagapetus) laparus sp. n. Figures 160-165 This small, brownish species appears to be closely related to A. cralus, but it is somewhat paler in colour and differs mainly in the male genitalia as shown in the figures. This species so far is known only from the type locality. $ genitalia: The anterior margin of seg- ment 9 is produced at the midlateral line to a blunt lobe; tergite 10 rather short; superior appendages stout, short and in lateral aspect spatulate with a group of long setae; inferior appendages short, tapering to a rounded apex in lateral aspect, but somewhat more slender Type material: Synagapetus crala Mosely. Type than in cralus in ventral aspect, the inner ridge & Cradle Mtn. Tas. 21 Jan. 1917, R. J. Till- terminating in a strong, triangular tooth near TASMANIAN CADDIS-FLIES 39 apex. There is no ventral process present on quite extensively. In this publication only a sternite 6. small number of species is described, although $ unknown. several more exist among the collected material, Length of anterior wing: $ 4-4-25 mm. but the small number of specimens in each of these species, have been considered insufficient for their description. Family diagnosis: Ocelli either present or ab- sent, according to genus. Maxillary palpi 5-seg- mented in both sexes, first two segments very short, terminal segment simple. Antennae shorter than anterior wings; shorter in females than in males; basal segment not as long as the head. Wings elongate, slender, usually acute at tips, densely covered with fine pubescence, fringe on both wings longer than the width of the wing, particularly so, in posterior wings. Mesoscutum without warts; scutellum forming a flat, triangular area with steep sides. Spurs 0:2:4; 0:3:4. The first four larval instars are slender, free living and of short duration. In the fifth instar the larvae radically change their shape and construct purse-shaped cases. KEY TO THE TASMANIAN GENERA 1. Spurs 0:2:4 2 — . Spurs 0:3:4 3 Type material: Holotype $ (T5220), 2 8 2. Ocelli present; head of male without scent paratypes (T5221-T5222) Condominion Creek organ caps Orphninotrichia nr. Mt Anne, Tas., 9 Feb. 1965, A. Neboiss — . Ocelli absent; head of male with scent (NMV);3 $ paratypes (T5223-T5225) Con- organ caps Hydroptila dominion Creek, Tas., 15 Feb. 1971, A. Ne- boiss (NMV). 3. Ocelli present 4 — . Ocelli absent Targatrichia Distribution: Tasmania—SW province. 4. Anterior wings without fork 3, abdominal 3 Family HYDROPTILIDAE Stephens segment 9 in female short 5 (1836) — . Anterior wings with fork 3 present; ab- The species of this family are among the dominal segment 9 in female extended, smallest of all Trichoptera, usually measuring slender Maydenoptila only 3 to 4 in length. They are known to mm 5. Anterior wings fork 1 present; cross-vein occur in all parts of the world and are present r-m at base of fork on M; posterior wings waters, with particular pre- in most unpolluted with fork 2 present Hellyethira ference for clean, running streams. — . Anterior wings without fork 1; cross-vein are rarely Because of their small size, they r-m basad of the base of fork on M; pos- collected by anyone except a specialist. Only terior wings without fork 2 . . Trichoglene eight species have been recorded in Australia so far, of which only one is from Tasmania. The Genus Orphninotrichia Mosely is still not satisfac- classification of the family Orphninotrichia Mosely, 1934:138; Mosely and Kim- tory, even in regions where they are known mins, 1953:510. 40 ARTURS NEBOISS Type species: Orphninotrichia maculataMosdly, 1934. Ocelli present. Anterior wings with forks 1 (?) and 2 present, fork 3 absent; cross-vein r-m at the base of fork on M. Posterior wings with forks 2 and 5 present. Antennae stout, consisting of about 30 segments; basal seg- ments barely longer than the subsequent ones. Spurs 0:2:4. At present only one species is recorded from Tasmania, which differs from the New South Wales species maculata by details of male geni- talia. 33 Orphninotrichia acta sp. n. Figures 166-173 Wings densely covered with long, black pu- bescence; the anterior wings with small patch of white hairs at the middle of posterior margin. The pubescence dorsally and the fringe on wings in closed position under some light con- ditions show metallic greenish or bluish sheen. Antennae with 29-31 segments. 3 genitalia with tergite 9 deeply excised dor- Ocelli absent. Head dorsally with a pair of sally, inner apical angles obtuse. Segment 10 lobes, which in some species in the males cover membraneous, broad at base, tapered distally, filaments, lined apex laterally compressed; lower apical angles membraneous or they are with membrane, is produced into downcurved finger-like processes. which covered with minute andro- conia. Antennae stout, always in Phallus long, slender, apex dilated, with small shorter fe- males, segments short, about equal in length, median excision. Apical margin of sternite 8 individual light rounded, slightly produced at middle. segments covered with dark or coloured hairs. Sternite 3 with longitudinal median ridge. Ventral process on sternite 7 absent. Anterior wings lanceolate, subacute, densely $ abdomen with segments 9 and 10 of about pubescent; cross-vein r-m at or slightly distad of base the same length, the latter dorso-ventrally com- of fork on M; apical forks 2 and 3 pressed; cerci short. present (interpreted by McLachlan 1880:510), and according to the interpretation Length oj anterior wing: 3 3-3 5 mm; 9 3 5 of Mosely mm. and Kimmins (1953), fork 1 also is present. Posterior wings with fringe twice Type material: Holotype $ (T4997), allotype more than the greatest 9 (T4998), 12 $ paratypes (T4999-T5010) width of the wing, fork 2 present, small. Ulverstone, 4 km NW waterfalls, Tas., 18 Nov. Spurs 0:2.4. 1972, A. Neboiss (NMV). So far two species are recorded from Tas- Other materia! mania. They are separated by the details of examined: Tasmania—7 <$ 1 ? Guide River Falls nr. Ridgley, Tas., 18 Nov. 1972, male genitalia. A. Neboiss (NMV). Distribution: Tasmania—N province. 34 Hydroptila tasmanica Mosely Genus Hydroptila Dalman Hydroptila tasmanica Mosely, 1934:147; Mosely and Hydroptila Dalman 1819:125; Ulmer, 1907:222; Kimmins, 1953:509. Moseley and Kimmins, 1953:505. Anterior wings light fuscous, fringe fuscous Type species: Hydroptila tineoides Dalman, with a white patch towards the apex. Antennae 1819. 29 segmented in male, pale fuscous. The wing . . TASMANIAN CADDIS-FLIES 41 venation and male genitalia have been illustra- row of a few small teeth on the inner margin, ted in the original description. 2 unknown. £ genitalia with long, bifurcate dorsal plate, Length of anterior wing: $ 2-5 mm. in dorsal view each branch dilated and obliquely truncate at apex. Phallus very long, slender, without the usual wrapped-around process near the apex. Small ventral process on sternite 7. 5 unknown. Length of anterior wing: £ 3 mm. Hydroptila tasmanica :;U^ M "^p V NW( ^W \ \ p j ' V. ' ne^J 7 -a J /v I XjLj ^x f . x ^^\ f \*\fpMIX. - \e^ * j£( •"" w»\ .„.. \ Jb \ ( / J ( C5 - / An. /^ h/se \ p \swA \ "%^i / Type material: Holotype $ (T5011), 1 S paratype (T5012) Scamander River, Upper Scamander, Tas., 9 Nov. 1972, A. Neboiss (NMV). « 70 « Htn « \^J> 34 ' ' Distribution: Tasmania—E province. material: Type 5 Wilmot, Tas., 9 Jan. Type Genus Targatrichia gen. n. 1917 (BMNH). Type not seen. Type species: Targatrichia zonata gen. et sp. n. No new material has been available for study. Ocelli absent. Antennae 28 to 32 segmented, Distribution: Tasmania—N province. apical segments with dark and light colour pu- 35 Hydroptila scamandra sp. n. bescence, producing zoned appearance, Anterior wings without anal lobe, apical forks 1, 2 and 3 Figures 174-176 present, fork 1 with long footstalk. Posterior densely pubescent, mottled, Anterior wings wings with fork 2 only. 28-segmented, 17 greyish-brown. Antennae Spurs 0:3:4. dark, followed by two whitish basal segments Only one species included in this genus. segments and 9 dark terminal segments, the last preceding ones. being much smaller than the 36 Targatrichia zonata sp. n. genitalia with tergite 9 short, segment 10 $ Figures 177-181 elongate, triangular. Phallus long, slender, with wrapped around, strap-like process near the Anterior wings densely covered with long erect setae, general colour apex. Inferior appendages in lateral view cla- hairs and brownish subdued mottling; cross-vein r-m distad vate; a small protuberance near the apex and a with 42 ARTURS NEBOISS of the base of fork on M. Antennae usually 32 North Esk River, Blessington, 1 Mar. 1967; 17 o* 5 $ Henty River 12 km NW Quecnstown, 10 Feb. 1971; segmented in males, 28 segmented in females, 105 c? 45 2 Huon-Picton River junction, 18 Feb. basal 18-22 segments dark, the 11 apical seg- 1967; 9 2 Hellyer River Gorge, 9 Feb. 1971; 4 2 ments always show the same colour pattern Huon River nr. Blakes Opening, 9 Feb. 1966. All specimens collected by A. Neboiss (NMV). 71 r? 34 2 with two pale, four dark, two pale and terminat- Huon-Picton junction, 17 Feb. 1967; 1 9 abdomen with terminal segments short, Ocelli present. Anterior wings with forks 1 sternite 8 with deep median cleft, at the base and 2 present, fork 3 absent; cross-vein r-m at of which arises a small, posteriorly directed tu- the base of fork on M. Posterior wings with fork bercle; segment 10 bearing a pair of slender 2 present, but very small. This genus is very cerci. close to the New Zealand genus Paroxyethira Length of anterior wing: & 3-5-3-75 mm; 9 from which it is separated by presence of foot- 3 5-4 mm. stalk on fork 1 and distinguished from genus Orthotrichia by absence of fork 3 in the an- terior wings. Antennae long, consisting of about 40 segments in males; basal segment slightly enlarged, rounded. Spurs 0:3:4. 37 Hellyethira vallecula sp. n. Figures 182-187 Anterior wings mottled, without anal lobe, forks as described in generic diagnosis. An- tennae in male 42 segmented, basal 21 segments pale, followed by 16 dark and terminal 5 pale again; usually two segments occupy the trans- gression zone from pale to dark segments; in females there is a total of about 34 segments, basal 19 segments pale, 11 dark and terminal 4 pale, although the numbers could vary slightly. 3 genitalia very short and compact, segment 9 laterally extended to a triangular, slightly curved lobe, ventral margin apically widely con- cave. Segment 10 membraneous, broad at base, truncate apically. Inferior appendages short, Type material: Holotype s (T50 13), allotype curved inwards. Phallus long, tubular, basally 2 (T5014), 6 g 12 $ paratypes (T5015- constricted near apex, spiral process wrapped T5032) St. Patricks River, Targa, Tas., 22 around the central tube. Small ventral process Feb. 1971, A. Neboiss (NMV). on sternite 7. 2 abdomen with apical margin of segment Other material examined: Tasmania— 1 <$ 1 $ Na- ventro-laterally tional Park, Tycnna River, 20 Feb. 1967; 1 38 Trichoglene columba sp. n. Figures 188-193 Wings covered with long pubescence, an- terior wings brownish, with indistinct trans- versal, dark and pale bands; posterior wings with fringe more than twice the greatest width of the wing, apical forks absent. Antennae 30- 33 segmented in males, 28-30 segmented in females. $ genitalia with segment 9 deeply and widely excised dorsally, forming a pair of con- cave dorsal lobes; a pair of shorter, apically truncate lobes formed by ventral margin; lower section of anterior margin produced forward nearly to the segment 7; between the dorsal and ventral lobes arises membraneous, finger-like process. 9 abdomen terminates with short segments 90 *0 8.0km 9 and 10, the latter with a pair of slender cerci. Type material: Holotype <} (T5033), allotype $ (T5034), 2 1971, A. Neboiss, (NMV); 1 $ paratype (T5040) Wedge River 30 mis W Maydena, Tas., 26 Feb. 1967, A. Neboiss (NMV); 1 & paratype (T5041) Huon River Crossing, Tas., 16 Feb. 1971, A. Neboiss (NMV); 1 $ para- type Hellyer River Gorge, Tas., 4 Feb. 1967, E. F. Riek (ANIC). Other material examined: Tasmania—2 c? 1 ? Sca- mander River, Upper Scamander, 9 Nov. 1972, A. Neboiss (NMV); 1 $ Flowerdale River, Meunna, 4 Nov. 1972, A. Neboiss (NMV); 2 <$ 2 ? Forth Falls, 28 Jan. 1967, E. F. Riek (ANIC). Distribution: Tasmania—E, N, NW and SW provinces. Genus Trichoglene gen. n. Type species: Trichoglene columba gen. et. sp. n. Ocelli present. Eyes covered with short setae. Antennae slender, 28 to 33 segmented, with- 44 ARTURS NEBOISS in male, terminal Length of anterior wing: £ 2-5-2-75 mm; 9 unicolorous, 42-segmcnted 2-5 mm. segment conical, slightly longer than the pre- ceding segment; in females about 33-segmentcd. Type material: Holotype $ (T5042), allotype genitalia with segment 9 deeply and widely 2 (T5043), 20 & 3 $ paratypes (T5044- $ excised dorsally, small protuberances at the T5066) Dove River, Cradle Mtn Nat. Park, upper distal angle apparently represent superior Tas., 14 Dec. 1974, A. Neboiss (NMV). appendages; basal margin mid-ventrally ex- Other material examined: Tasmania—29 c? 6 $ Huon-Picton River junction, 15 Nov. 1972, A. Ne- tended into apically rounded lobes. Segment 10 boiss (NMV); 3 c? Hellyer River Gorge. 9 Feb. long, broad at base, bluntly triangular at apex. 1971, A. Neboiss (NMV); 31 <$ 9 ? Fiowerdale River, Phallus parallel-sided, slightly curved down- Meunna, 4 Nov. 1971, A. Neboiss (NMV); 2 sp. n. segment 1 small, conical, with a pair of slender Ocelli present. Wings rather broad, lanceo- cerci. late; anterior wings with small, rounded anal Length of anterior wing: 6 3-5-4 mm; 9 4 lobes, forks 1, 2 and 3 present; posterior wings mm. with forks 2, 3 and 5 present; fringe moderately long, less than twice the greatest width of the wing. Antennae slender, 38-42 segmented in males; 28-33 segmented in females, the two basal segments enlarged, terminal segment longer than the preceding one. In females the abdomen gradually tapered, segment 9 slender, segment 10 small, conical. Spurs 0:3:4. Two Tasmanian species are included in this genus. KEY TO THE SPECIES OF MAYDENOPTILA 1. Anterior wing, fork 2 short, about as long as fork 3 rupina — . Anterior wing, fork 2 long and narrow, about twice as long as fork 3 . . cuneola 39 Maydenoptila cuneola sp. n. Figures 194-197; 203 Anterior wings brownish, with more or less distinct pale transversal line at about the middle; fork 2 very long and narrow, about twice as long as fork 3, reaching basally beyond Type material: Holotype $ (T5067), allotype the base of forks 1 and 3. Posterior wings with 2 (T5068), 9 £ 3 2 paratypes (T5069- forks 2, 3 and 5 present; fork 2 very long and T5080) Wedge River, Tas., 17 Feb. 1971, A. narrow, about twice as long as fork 3. Antennae Neboiss (NMV); 1^2$ paratypes, Wedge — TASMANIAN CADDIS-FLIES 45 River 30 mis W Maydena, Tas., 25 Feb. $ abdomen gradually tapers, segment 8 1967, E.F. Riek (ANIC). short, segment 9 slender, segment 10 short, somewhat conical, with a pair of small cerci. Other material examined: Tasmania— 1 o* Cra- croft River, 8 Feb. 1966, A. Neboiss (NMV); 1 apical forks 1, 2, 3, 4 and 5, or 1, 2, 3 and 5 present; posterior wings with forks 1, 2, 3 and 5, or 2, 3 and 5 present; discoidal cell always present in both wings. Two genera has so far been recorded from Australia, of which only one Hydrobiosella Tillyard—occurs in Tasmania. Ross (1956) regarded this as one of the seven subgenera of Stfrtova Navas (— Dolophiloides Ulmer, 1909) each one of which was apparently restricted to a single zoogeographic region. These groups are morphologically distinct and separable, and as more species have been discovered, there is sufficient evidence to retain the previously ap- plied name Hydrobiosella at the generic level for the Australian and New Zealand group of species. Genus Hydrobiosella Tillyard HvdrobioseUa Tillyard, 1924:288; Mosely and Kim- mins. 1953:387; Rick, 1970:757 Sortosa subgenus Hydrobiosella\ Ross, 1956:30. Dolophiloides subgenus HvdrobioseUa, Wise, 1973: 172. 46 ARTURS NEBOISS Type species: Hydrohhsella stenocerca Till- circular structure on sternite 4 in females yard, 1924. present. The genus is confined to Australia and New To this group belong corinna, orba, cerula Zealand with one species in New Caledonia. and anasina. The Tasmanian species are all of medium size B. Four species with abdominal segment 9 with wing span between 14 and 20 mm, dark entire, either joined or narrowly separated dor- grey to light-brownish, mottled with dull, grey- sally; posterior margin with lateral excision; ish to bright yellow-brown. superior appendages present in form of more or less Anterior wings with forks 1, 2, 3, 4 and 5 widened lobes; valvular structure present present; forks 1 and 2 sessile; cross vein be- on sternite 5 in both sexes; circular structure tween costa and subcosta at about middle of on sternite 4 present in females. costal area; two cross veins between Sc and R, To this group belong tasmanica and armata first near the R-Rs junction, the second near the with well developed circular structure on ster- distal end of discoidal cell. Posterior wings with nite 4 and sagitta and cognata with poorly de- veloped forks 1, 2, 3 and 5 present; fork 1 usually ses- circular structure. sile; three anal veins. C. One species (waddama) with dorsal part of abdominal segment 9 membraneous; su- Maxillary palp with first segment short, sec- perior appendages absent; valcular structures ond slightly longer, with a group of stiff bristles absent, circular on the inner distal angle, third as long or longer structure on sternite 4 in fe- males present. than first two together, fourth shorter than third, fifth long and flexible. It is interesting to note that of the nine Tas- manian species no less Male genitalia distinctly elongate, with two- than eight are endemic; the only segmented inferior appendages. Female abdo- exception being waddama, which is also widely distributed in Victoria men gradually tapers to a simple apex and ter- and New South Wales. minates with a pair of small but distinct cerci. Segments 8 and 9 internally with slender apo- KEY TO TASMANIAN HYDROBIOSELLA demes. SPECIES (Males only) Spurs 2:4:4. 1. Tcrgum of segment 9 membra- A total of nine species are now recognized neous waddama from Tasmania; five are described here as new, — . Tergum of segment 9 entirely or partly two of which (corinna and orba) show close sclerotized 2 affinities with the New Caledonian species Hy- 2. Posterior margin of segment 9 without drobiosella uncinata Kimmins. Ross (1956) lateral excision 3 suggested that the primitive forms would have —. Posterior margin of segment 9 with lateral possessed short superior appendages (= cerci excision . 6 of Ross, 1956) and a complete set of anal veins 3. Segment 10 very long and slender, ex- in posterior wing. This situation is found in H. tending at least to base of harpago . . 4 corinna as well as in H. orba, and these could Segment — . 10 at most extending as far as therefore be regarded as primitive forms being the middle of coxopodite 5 closest to the Hydrobiosella ancestor type. 4. Apex of segment 10 barely reaching base The Tasmanian species fall into three quite of harpago, widened at the lower distinctive groups. ang'e anasina A. Four species with abdominal — Apex of segment 9 . segment 10 reaching beyond base entire, either joined or narrowly separated dor- of harpago, curved upward, without form- sally; posterior margin without lateral excision, ing distinct angle cerula superior appendages present in a form of small, 5. Segment 9 dorsally bridged by semi-mem- simple processes; small, but distinct valvular braneous section; phallus short structure present on sternite 5 in both sexes; and stout orba , TASMANIAN CADDIS-FLIES 47 Segment 9 dorsally joined with narrow sclerotized bridge; phallus slender corinna Superior appendage broad, angular, with lower posterior angle extended into a long, upcurved spine 7 Superior appendage not angular, extended distally g 7. Apex of segment 10 in dorsal view short, triangular armata Segment 10 with small, triangular lateral flanges, after which the sides gradually curve to a narrow rounded frp tasmanica 8. Superior appendage extended distally to a finger-like, rounded process . . cognata Superior appendage extended distally to a strap-like, twisted process . . . . sagitta 41 Hydrobiosella corinna sp. n. Figures 204-205 Anterior wings yellowish to greyish-brown, irregularly mottled, with pale and dark pubes- King River, Tas., 5 Nov. 1961, N. Dobrotwor- cence. Valvular structure on sternite 5 present sky (NMV). in both sexes, and a circular structure on ster- nite 4 in females only. Other material examined; 37 <$ 6 ? 10 mis E Stra- han, 6 Feb. 1967, E. F. Riek (ANIC); 1 Murchison $ genitalia with segment tf 9 dorsally joined River, 5 Feb. 1967, E. F. Riek (ANIC). with a narrow, sclerotized bridge; posterior Distribution: Tasmania NW province. margin without lateral excision; segment 10 in — dorsal view somewhat elongate triangular, 42 Hydrobiosella orba sp. n. gradually tapering to rather pointed, laterally Figures 206-207 compressed apex, posteriorly not extending be- yond the middle of coxopodite. Superior ap- Anterior wings light yellowish-brown pendages small, somewhat pointed. Phallus mottled, pterostigma more distinct than in other long, slender, cylindrical, a pair of 'V shaped species. Valvular structure on sternite 5 present. chitinous structures internally on ventral side. $ genitalia with segment 9 rather short, Inferior appendages slender, coxopodite almost bridged dorsally with narrow, semi-membrane- twice the length of harpago, which is slightly ous section, posterior margin without lateral ex- curved inward, with a group of short, stout cision. Segment 10 shorter than in corinna, spines at the apex. elongate triangular, apex rounded and dorsally 2 abdomen gradually tapers towards die elevated. Superior appendages very short. Phal- apex, which bears a pair of small cerci; lateral lus short and stout, internally with a pair of V- margin of sternite 7 without protuberances. shaped chitinous structures. Inferior appen- Length of anterior wing: $ 7 5-8-5 mm; 9 dages slender, coxopodite slightly less than 7-5-9 mm. twice the length of harpago, the latter curved Type material: Holotype $ (T4480), allotype inwards with a group of short, stout spines at 9 (T4481), 2 8 paratypes (T4482-T4483) the apex. Corinna, Tas., 5 Nov. 1972, A. Neboiss 2 unknown. (NMV); 1 £ 2 $ paratypes (T4484-T4486) Length of anterior wing: $ 6 mm. 48 ARTURS NEBOISS Type material: Holotypc $ (T4479) Wedge Length of anterior wing: s 7-8 mm. River, Tas., 17 Feb. 1971, A. Neboiss (NMV). Type material: Holotype $ (T4478) Lake Distribution: Tasmania—SW province. Pedder, Tas., 10 Mar. 1972, A. Neboiss (NMV); 1 8 paratype Gordon River, Strath- 43 Hydrobiosella cerula sp. n. gordon 50 mis W Maydena, Tas., 26 Feb. Figures 208-209 1967, E. F. Riek (AN1C). Anterior wings light yellowish-brown, irregu- Distribution: Tasmania—SW province. larly mottled. Wing venation typical of the 44 Hydrobiosella anasina sp. genus, details of the cross-vein position shown n. in the figure. Valvular structure on sternite 5 Figures 210-211 present. Anterior wings yellowish-brown, irregularly S genitalia with segment 9 in lateral view mottled. Valvular structure on sternite 5 present somewhat obliquely elongate; dorsally bridged in both sexes, but circular structure on sternite by short, semi-membraneous section, posterior 4 in females only. margin without lateral excision; segment 10 i! genitalia with segment 9 dorsally joined by very long and slender, in dorsal view gradually a short, narrow chitinous ridge; posterior mar- tapering apically, extending beyond the base of gin without lateral excision; segment 10 long harpago, the apex curved upwards. Superior and slender, but barely reaching the base of appendages small, elongate lying close and harpago, apex distinctly widened at the base of parallel to the margin of segment 9. Phallus sharply upturned tip. Superior appendages cylindrical, slightly curved downward, apex small, lying close and parallel to the margin of widened laterally. Inferior appendages long, segment 9, shorter than in H. cerula. Phallus rather stout; coxopodite about 2 5 times longer cylindrical. Inferior appendages with coxopo- than harpago, the latter stout, armed with four dite about twice as long as harpago, which is to five strong, inwardly directed spines and a somewhat triangular in lateral view, a few group of short, stiff bristles on inner angle. stout, inwardly directed spines at apex and a 9 unknown. group of stiff bristles on inner angle. TASMANIAN CADDIS-FLIES 49 $ abdomen gradually tapering towards the S genitalia—segment 9 with tergal section apex, which terminates with a pair of small short, separated from sternal section by lateral cerci. At the middle of the lateral margin of excision; segment 10 in dorsal view with small, sternite 7 there is a small, ridge-like protuber- triangular lateral flanges at about the middle of ance. the segment, from there on the sides gradually Length of anterior wing: £ 7-9 mm; 5 7-8*5 curve to a narrow, rounded apex; dorsally at the mm. base there is a pair of minute, flat, triangular protuberances. Superior appendages ( = basal ridge or process of tenth tergite by Ross, 1956) broadly angular, upper posterior angle rounded, triangular, the lower posterior angle extended distally to a long, upcurved spine. Phallus slen- der, slightly curved downwards, with a pair of strong, ventrally directed spines at the apex, other internal, distally directed spines not al- ways visible. Inferior appendages long, coxopo- dite about 1-5 times longer than harpago, the latter slightly curved inwards with groups of short, stout bristles at the apex. $ abdomen gradually tapers apically and terminates with a pair of small cerci; tergite 8 narrow, elongate triangular. Length of anterior wing: $ 1 -5-9*5 mm; 9 8- 9-5 mm. Type material: Holotype $ (T5226), allotype 9 (T5227), 14 S 14$ paratypes (T5228- T5255) Duck River 6 km SW of Roger River, Tas., 29 Nov. 1974, A. Neboiss (NMV). Other material examined: Tasmania— 1 c? Dip River Falls 10 km S of Mawbanna, 1 Dec. 1974, A. Neboiss (NMY); 1 45 Hydrobiosella tasmanica Mosely Figures 212-217 Hydrobiosella tasmanica Mosely in Mosely and Kimmins, 1953: 390. Anterior wings yellowish to greyish-brown, irregularly mottled. Valvular structure on ster- nite 5 present in both sexes, in females circular Type material: Holotype & Tasmania, J. W. structure on sternite 4 present and well de- Evans (BMNH), without definite locality. Type veloped. seen. 50 ARTURS NEBOISS Material examined: Tasmania— 11 Feb. J 965 and 15 Feb. 1971; 12 tf 2 ? Hot Springs Creek nr. Hastings Caves, 14 Nov. 1972. All speci- mens collected by A. Neboiss unless stated otherwise (NMV). 1 c? 4 $ Russell Falls, 23 Feb. 1967 E F Riek (ANIC). Three $ paratypes were examined: it was found that the paratype 3 from Cradle Mtn., 16 Jan. 1917, R. j. Tillyard (BMNH), is Hy- drobiosella cognata Kimmins, and 2 $ para- types Hobart, Dec. 1937, J. W. Evans, are Hy- drobiosella armata Jacquemart. It is likely that the female from Hobart, Dec. 1937 and Mt Wellington 3700 ft., 7 Feb. 1942, J. W. Evans, are also H. armata, but the female from Wara- tah, collected by Lea, belongs to one of the three North West Tasmanian species. Distribution: Tasmania—SW and SE provinces. 46 Hydrobiosella armata Jacquemart Figures 218-219 Length of anterior wing: $ 5.5-7 mm; $ 6-7 Hydrobiosella armata Jacquemart, 1965b:34. mm. Anterior wings pale yellowish to greyish- Type material: Holotype $ Mt Wellington, brown with irregular mottling. Closely related Tas., 20 Nov. 1922, A. Tonnoir (IRScNB). to H. tasmanica, but smaller in size and can Mounted on three microscope slides; a label on be separated by the different shapes of the su- one of these slides reads "Nouvelle Zelande, perior appendages and of segment 10. Valvular Mont Wellington 20.11.1922", not in Tonnoir's structure of sternite 5 present in both sexes; in handwriting. females the circular structure on Material sternite 4 pre- examined: Tasmania— 1 15 Dec. 1 1 River, Pencil Pine differences were found in the holotype, which is 1974; times longer than harpago, which bear a group mi ns as author, as it appears correctly in the of small spines at the apex. same publication on p. 31 fig. 24. It is also 9 abdomen typical of the genus, segments noted that the wing venation in this figure is in- gradually tapering apically and terminating with complete. The locality 'Otrokan' should read a pair of distinct cerci. 'Strahan', a place where Tonnoir collected other Length of anterior wing: $ 7-5-8-5 mm; $ 8- insects on the same date (Neboiss 1974c: 15). 8 5 mm. Distribution: Tasmania—N, NW and C pro- vinces. 48 Hydrohiosella sagitta sp. n Figures 225-226 Anterior wings pale yellowish-brown, lightly and irregularly mottled. Valvular structure on sternite 5 present in both sexes, the circular structure on sternite 4 in females indistinct. S genitalia with segment 9 short, lateral ex- cision present but wider than in other Tasma- nian species. Segment 10 in dorsal view broadly ovoid at the base, and distinct triangular apex; a pair of parallel dorsal ridges basally widened to small, triangular, outward turned lobes. Su- perior appendages extended downward and dis- tally to a twisted, strap-like process. Phallus straight, apically expanded, clavate. Inferior ap- pendages stout, coxopodite about twice as long as harpago which bears a group of short spines at the apex. 9 abdomen similar in shape to that in H. cognata. Type material: Holotype S Lake St. Clair, Feb. 1941, J. W. Evans (BMNH). Type not Length of anterior wing: S 7*5-8-5 mm; 9 seen. 7-75 mm. 52 ARTURS NEBOISS which bears a pair of short ccrci. Stcrnite 7 with ventral keel; segment 8 longer than seg- ments 9 and 10 together. Length of anterior wing: i 6 5-8 mm; 9 7-9 mm. Type material: Holotype & (T4489), allotype 9 (T4490), 5 8 paratypes (T4491-T4495) St. Columba Falls or. Pyengana, Tas., 21 Feb. 1971, A. Neboiss (NMV). Distribution: Tasmania—NE province. Type material: Holotype $ Waddamana, R. 49 Hydrobiosella waddama Mosely Ouse, Tas., Critchley Parker (BMNH). Type Figures 227-233 not seen. Mater tat examined: Tasmania—7 <$ 18 $ St. Co- Hydrobiosella waddama Mosely in Mosely and Ktm- lumba Falls, Pyengana, 21 Feb. 1971; 4 cM2 2 St. tains, 1953:393. Patricks River, Targa, 22 Feb. 1971; 4 <$ 1 9 Ulver- Anterior wings greyish-brown, irregularly stonc, 4 km NW waterfalls on coastal cliffs, 18 Nov. mottled. Valvular structures on sternite 5 not Nov. 1972; 1 $ Flowerdale River, Meunna, 4 1972; 21 e? 7 2 Hellyer River Gorge, 9 Feb. 1971; present in cither sex, but circular structure on 1 $ Waldheim, Cradle Mtn. Nat. Park, 7 Feb. 1971; sternite 4 present in females. 80 oMO 2 Franklin River 20 km SW of Derwent 11 1 Ouse River 8 of A genitalia with segment 9 membraneous Bridge, Feb. 1971; c? km W Miena, 28 Feb. 1967; 1 S Bradys Lake, 27 Feb. dorsally, anterior margin produced into a long 1967; 26 rf 50 2 Lake Dobson, 20 Feb. 1967; 31 c? lobe. Segment 10 long, extending almost to the 12 2 Huon-Picton River junction, 18 Feb. 1967; 29 <•? 16 2 Huon River nr. Blakes Opening, 9 Feb. 1966; apex of harpago, broad at base, from about 1 r? Cracroft River, 8 Feb. 1966. All specimens col- middle tapering to narrow, rounded apex. Su- lected by A. Neboiss (NMV). 72 <$ 2 2 Franklin perior appendages entirely absent. Phallus long, River, 10 Feb. 1967, E. F. Riek (ANIC); 33 c? 17 2 Hellyer Gorge, 4 Feb. 1967, E. F. Riek (ANIC); slender, slightly curved, a pair of long processes 15 c? 19 2 Lake Dobson, 20 Feb. 1967, E. F. Riek arise from the base and extend to the apex of (ANIC); 2 2 Ouse River 5 mis W Miena, 28 Feb. 1967, segment 10. Inferior appendages two-seg- E. F. Riek (ANIC); 1 2 Russell Falls, 23 Feb. 1967, E. F. Riek (ANIC); 22 5 Family STENOPSYCHIDAE Martynov Genus Stenopsychodes Ulmer (1926) Stenopsychodes Ulmer, 1916:5; Mosely and Kim- In Australia the genus Stenopsychodes Ul- mins,' 1953:363; Schmid, 1969:197; Neboiss, 1974a:81. mer is the only known representative of the Type species: Stenopsychodes mjobergi Ulmer, family Stenopsychidae. Mosely and Kimmins 1916. (1953) placed the genus in the family Polycen- In addition to the family description, the tropodidae, but Ross (1967) analysing the genus is characterized by rather squarish head, family relationships, regarded it as a member which bears one small anterior and two larger of the family Stenopsychidae. He placed it posterior warts; eyes do not occupy the entire alongside the family Philopotamidae as belong- side of the head; a pair of minute elevations in- ing to the most primitive branch of Trichoptera. dicate the possible location of the lateral ocelli. Riek (1970) included all Australian species of The scutellum is elongate triangular, with Polycentropodidae and Stenopsychidae in the anterior quarter produced and narrow, on either one family Psychomyidae. side of it rounded mesoscutal warts. Anterior In the most recent revision only two genera wing with an oblique costal cross-vein at about were included in the family Stenopsychidae the middle of the wing; forks 1-5 all present. (Schmid 1969). The genus Stenopsyche Mc- $ genitalia elongate, with long, slender su- Lachlan, is mainly found in southern and perior appendages; inferior appendages large, eastern Asia, from India to Japan. This genus robust, single-segmented; phallus stout. is characterized by having large ocelli and slen- 9 abdomen tapers to a narrow, dorso-ven- der antennae. The entirely Australian genus trally flattened, bilobed, apical segment, which Stenopsychodes Ulmer, is the only other genus bears a pair of minute cerci. Posterior angle of included in the family and differs from the tergite 8 extended into divergent, rounded, ear- former by the absence of ocelli and rather ro- like flaps; sternite 9 internally with long, an- bust antennae. Schmid (1969) recognized six teriorly produced apodeme on lateral angles. species in this genus, to which two further spe- Only one species in Tasmania. cies were added by Neboiss (1974a). The dis- tribution of the genus was limited to the moun- 50 Stenopsychodes lineata sp. n. tain areas along the Australian east coast, from Figures 234-242 North Queensland to Victoria, within 250 km of the coast. It is therefore most interesting to Head, thorax and abdomen, as well as the record the genus and family for the first time antennae, palpi and legs black; wings golden- from Tasmania. yellow, with black hairs along the veins and Family diagnosis: Ocelli present or absent, de- with a few small, more or less distinct groups pending on genus. Antennae about as long, or at the distal section of costal margin. Mouth only slightly longer than anterior wings, slen- parts not elongate. der to robust, basal segment short. Maxillary & genitalia basically the same as in Steno- palpi 5-segmented, first two segments short, psychodes montana Tillyard, but differs by somewhat thickened, segment 3 longer than the havine a small, inwardlv directed lobe on the first two together, segment 4 shorter than seg- lower margin of lateral angle of segment 9. Su- ment 3, segment 5 long, flexible. Mesoscutum perior appendages long, slender, slightly curved, with a pair of rounded warts lateral and close to extending slightly beyond the apices of inferior scutellum, the latter somewhat elongate triangu- appendages. Tergite 10 with a pair of minute, lar with a single posteriorly situated wart. Ab- and a pair of long, slightly curved, dorsal pro- dominal sternite 5 with lateral pore, but lateral cesses; laterally a somewhat flattened, two- filament absent. Wing venation complete, regu- branched process on either side, the upper lar, without sexual dimorphism; in anterior branch very long, slender, upcurved distally, the wing the anal lobe well developed. lower branch only about half the length, acute. ventral Spurs 3:4:4. Phallus stout, with median keel. In- — 54 ARTURS NEBOISS ferior appendages long, robust, slightly dilated lished a detailed analysis of this family complex distally, rounded excision at apex, the upper and arrived at the conclusion that the larval margin of the excision terminating with two or characters strongly support differences between three minute, acute points. Polycentropodidae, Psychomyidae and Ecnomi- dae, raising the latter to family level. At about 9 genitalia sternite 8 short and broad, pos- analysed the terior margin with rounded central incision. the same time Kimmins (1957) African Psychomyidae, comparing them with Length of anterior wing: $ 9-10 mm; $ 10-11 mm. specimens from other regions in search of reli- able characters for separating Ecnominae from Stencipsychod ^s lineata Psychomyinae. The two genera found in Aus- tralia Ecnomus and Ecnomina—were placed IS in the subfamily Ecnominae. Marlier (1958) IS supported Kimmins work, and independently of Lepneva, suggested that the subfamily Ecno- minae should be raised to family level. Malicky N ^\ (1973) followed the revised family classifica- NW ' i j 7 tion proposed by Lepneva. Consequently, the G NE ; family name Psychomyidae is now removed f 1 . from the Australian fauna and replaced by family Ecnomidae. (J h \ ^ 1 \T vn S Ye^ + $L Family diagnosis: Ocelli absent. Antennae at / <3 \ V; J* "\ If] f * the most as long as the anterior wings. Maxil- $*• +- times also with setiferous punctures. Abdom- \ \ TWA inal sternite 5 without lateral filaments. In an- 1 terior wings R.! is usually forked at apex, api- o m 10 CO kit •^J]f 50 cal fork 1 present or absent, forks 2, 3, 4 and 5 * all present. In posterior wings discoidal cell present or absent, forks 2 and 5 always present, Type material: Holotype $ (T4801), allo- fork 3 either present or absent; fork 2 always type (T4802), paratype 8 (T4803), para- 2 without nygma. type 9 (T4804) Bluff Hill Creek, 12 km S of Spurs 2:4:4 or 3:4:4. Marrawah, Tas., 30 Nov. 1974, A. Neboiss (NMV). KEY TO TASMANIAN GENERA Distribution: Tasmania—NW province. 1. Anterior wings with fork 1 present; in posterior wing fork 3 and discoidal cell 6 Family ECNOMIDAE Ulmer (1903) absent; female abdomen terminates blundy, segments 9 and 10 Since the beginning of this century the short Ecnomus limitations and relationships between the vari- — . Anterior wing fork 1 absent; in posterior ous families within the polycentropodid-psy- wing fork 3 and cell present; fe- chomyid complex have been subject for detailed discoidal male abdomen gradually tapering, seg- research. Most of the recent authors (Ulmer, ments 9 and 10 long and 1951; Mosely and Kimmins, 1953; Lepneva, slender Ecnomina 1964 and others) separate Polycentropodidae from Psychomyidae, although Riek (1970) Genus Ecnomus McLachlan considered the polycentropodids to be a sub- Ecnomus McLachlan, 1864:30; Ulmer, 1907:190; family of Psychomyidae. Lepneva (1956) pub- Mosely and Kimmins, 1953: 378. TASMANIAN CADDIS-FLIES 55 Type species: Philopotamus tenellus Rambur, wardly directed tooth at the middle; terminal 1842. half straight, narrow in ventral view, slightly Maxillary palpi with segment 1 short, seg- flattened laterally. Phallus laterally compressed ment 2 slightly longer; segment 3 slightly longer at the apex. than 2; segment 4 as long as, or slightly longer The female usually slightly larger and more than 3; segment 5 about as long or slightly robust than male; abdomen terminates bluntly longer than all the others together. Mesoscutum with a pair of ventral lobes, rounded apically. and scutellum each with a pair warts. of An- Length of anterior wing: $ 6-7 mm; $ 6 5- terior wing with forked R x at apex; forks 1, 2, 85 mm. 3, 4 and 5 present; fork 1 short; discoidal, me- dian and thyridial cells all present. Posterior wing narrower than the anterior; forks 2 and 5 present; discoidal cell absent. Spurs 3:4:4; the outer pair of mid-tibial spurs shorter than the inner pair. Numerous specimens of Ecnomus continentalis Ulmer from various localities in Queensland, all had spurs 3:4:4 and not 2:4:4, as indicated by Mo- sely and Kimmins (1953:378). Female with mid-tibia and tarsus dilated. KEY TO TASMANIAN SPECIES OF ECNOMUS 1. Inferior appendage in ventral view with distal half narrow, straight (female with ventral plate of segment 8 rounded api- cally) tillyardi — . Inferior appendage in ventral view broad, curved 2 2. Inferior appendages broad at base, lower inner margin at the middle with long, finger-like process (female not known) continentalis — . Inferior appendages broad, lower inner margin with blunt central lobe (female Type material: Type $ Cradle Mtn. Tas., 22 with ventral plates of segment 8 pointed at Jan. 1917, J. W. Evans (BMNH). Type not inner apical angle) russellius seen. Material examined: Tasmania—95 c? 57 $ Leven River, Heka, 17 Nov. 1972; 15 1974; 2 J2S Meander River 3 km N of Westbury, River, Deloraine, 16 Dec. 1974; I cM ? Meander 28 Nov. 1974; 1 52 Ecnomus russellius sp. n. Figures 250-254 A greyish or dull-yellowish irrorate species, somewhat larger and more robust than E. till- Jan. E. F. Riek yardi, but the male genitalia close to that of E 9 Other material examined: Tasmania— 1 o* Mac- the lower inner margin with finger-like process quarie River 8 km Campbell Town, 9 Nov. 1972; W in the middle. Phallus compressed laterally at 1 d Lake Dobson, 20 Feb. 1967; 1 ? Russell Falls, National Park, 5 Dec. 1972, P. Zwick; 1 TASMANIAN CADDIS-FLIES 57 Ecnomus continentalis KEY TO TASMANIAN SPECIES OF ECNOMINA (Males only) 1 Inferior appendages present 2 — Inferior . appendages . absent . . . batyle 2. Inferior appendages fused to form a single elongate plate irrorata — Inferior . appendages not fused .... 3 3. Inferior appendages in lateral view as long as wide legula Inferior — . appendages in lateral view longer than wide vega 54 Ecnomina irrorata Kimmins Figures 257-260 Ecnomina irrorata Kimmins in Mosely and Kimmins, 1953:384. Small, brownish species with mottled an- terior wings. £ genitalia characterized by a pair of large, dorso-lateral processes; a strongly chitinized, downcurved, distally pointed process just below phallus. Inferior appendages fused to form a Distribution: Tasmania—N province; Queens- single, elongate apically excised plate. land; South Australia. 9 abdomen with sternite 8 developed into a pair of lateral lobes, ventrally separated by a Genus Ecnomina Kimmins deep, wide, basally rounded excision, in the Ecnomina Kimmins in Mosely and Kimmins, 1953: middle of which there is a long, narrow, sclero- 380. sed ridge. Segments 9 and 10 appear to be Type species: Ecnomina spinosa Kimmins, welded together to form a pair of slender, api- 1953. cally tapering and dorso-ventrally separated lobes; Maxillary palpi with segment 1 short, seg- a pair of small cerci at the apex. ment 2 one and half times as long again as Length of anterior wing: s 4-5-5 mm; $ 5-6 the first, segments 3 and 4 each about twice as mm. long as segment 1, segment 5 about as long as Type material: Type $ Deloraine, Tas., 26-27 the first four together, flexible. Anterior wing Dec. 1884, McLachlan Collection (BMNH). with R] forked from the cross-vein between Sc Type not seen. and Ri; apical forks 2, 3, 4 and 5 present; dis- Material examined: Tasmania—4 c? 5 2 Huon- Picton coidal, median and thyridial cells all present. River junction, 18 Feb. 1967; 2 o* 1 2 Huon Plains nr. In posterior wing Sc and R are fused near apex; Scotts Peak, 8 Feb. 1965; 2 ? Lake Ped- der, 1 Feb. 1965; 2 ? Junction Creek, West Arthur apical forks well 2, 3 and 5 as as discoidal cell Plains, 7 Feb. 1966; 10 $ Huon Crossing, Port Davey present. In female the abdomen gradually Track, 9 Feb. 1965; 2 $ Huon River, Blakes Open- ing, 9 Feb. 1966; 5 Cracroft tapers posteriorly, segments 9 and 10 long and ? River, 8 Feb. 1966 2 $ Henty River 12 km NW Queenstown, 10 Feb slender; a pair of small cerci at apex. 1971; 19 c? 2 $ Tooms Lake, 4 Dec. 1974; 2 ? Der went River Spurs 3:4:4; outer tibial spurs shorter than 3 km W of New Norfolk, 7 Dec. 1974 1 $ Apsley River, Bicheno, 9 Nov. 1972 1 $ Frank inner, particularly those of the mid-tibia; fe- lin River 20 km SW of Derwent Bridge, 11 Feb male with mid-tibia and tarsus dilated. 1971; 2 $ Derwent River 2 km NW Derwent Bridge 12 Feb. Four species in Tasmania, three of which are 1971; 1 ? Waldheim, Cradle Mtn. Nat Park, 7 Feb. 1971. All specimens collected by A here described as new. Neboiss (NMV). 58 ARTURS NEBOISS very slender, distal end slightly dilated, a pair Ecnomina irrorata '.U> of small cerci and a pair of small membraneous %^ processes at the apex. e Length of anterior wing: 8 5-5-6 mm; 9 6-7 mm. *'< + \ A ^ <^C\ < A / 1 1a ~ ^T NW — ' -•— ) r A / cV ^T\ ' ~\wL ( f M *0 Mtun o 1v-X 54 I SW * 3 ? Lake St. Clair, 13 Feb. 1967, E. F. Riek X aL- Falls, 28 Jan. 1967, E. F. Riek (ANIC); 1 c? Forth A (ANIC); 8 ? Murchison River, 5 Feb. 1967, E. F. Riek (ANIC); 1 o* Evandale, 1 Mar. 1967, E. F. (ANIC); 2 Hellyer Gorge, 4 Feb. 1967, E. F. Riek . « ? H « 10 K Riek (ANIC). L Distribution: Tasmania—N, NW, SW, C, SE and E provinces. Type material: Holotype $ (T4887), allotype $ (T4888), paratypes (T4889- 55 Ecnomina legula sp. n. 2^59 Huon River Crossing, Port Davey Figures 261-268 T4895) Track, Tas., 9 Feb. 1965, A. Neboiss (NMV); A species closely related to Ecnomina spino- River 2 Z paratypes (T4896-T4897) Huon sa Kimmins from New South Wales, but differ- Crossing, Tas., 16 Feb. 1971, A. Neboiss ing in details of the male genitalia. Anterior (NMV);1 8 1 9 paratypes (T4898-T4899) wings concolorous brownish. West Arthur Plains, Tas., 3 Feb. 1965, A. Ne- 3 genitalia tergite 9 consists of a pair of — boiss (NMV). curved, basally widened plates with a number Other material examined: 2 o* 2 $ Forth Falls, 28 of strong spines on the inner basal margin, and Jan. 1967, E. F. Riek (ANIC); 2 6* 4 ? Murchison a heavily built lateral process just below ter- River, 5 Feb. 1967, E. F. Riek (ANIC); 3 i yl 1 v' nw ( ^yj\ \ N j \ * /v/ S /\rr ^r f SE ( \ Jk A SW N. \ /V*s «J Jii7 rvA/i a, i v^s^-^r "..';> . '%\^> o w w eoim J 57 Type material: Holotype S (T4900) Mac- Type material: Holotype $ (T4861), allotype quarie River 8 km W Campbell Town, Tas., 9 9 (T4862), 12 S 12 9 paratypes (T4863- Nov. 1972, A. Neboiss (NMV) T4886) Wilmot River 10 km S of Forth, Tas., Distribution: Tasmania—E province. 13 Dec. 1974, A. Neboiss (NMV). Other material examined; Tasmania—2 o* 3 2 Me- 57 Ecnomina batyle sp. n. ander River 3 km N of Westbury, 16 Dec. 1974, A. Neboiss (NMV); 1 o* Derwent River 2 km NW of Figures 272-276 Derwent Bridge, 12 Feb. 1971, A. Neboiss (NMV); 14 10 2 Forth Falls, 28 Jan. 1967, E. F. Riek species, anterior wings with c? A small, fuscous (ANIC). dense, concolorous, brownish pubescence; the Distribution: Tasmania—N and C provinces. characteristic male genitalia separates this spe- cies from others in the genus. 7 Family POLYCENTROPODIDAE Ulmer genitalia with tergite 9 short, with a pair S (1906) of obliquely truncate processes arising from dis- Family diagnosis: Ocelli absent. Antennae tal margin. Segment 10 membraneous, with either shorter than, or as long as anterior sclerotized median ridge. Phallus long, laterally stout, wings. Maxillary palpi five-segmented in both widened at the middle, apex elevated and ex- sexes; first two segments short, terminal seg- panded dorsally; a long, bifurcate process be- 60 ARTURS NEBOISS Stephens ment long, annulated, flexible. Mesoscutum Genus Plectrocnemia with a pair of round warts, sometimes with ad- Plectrocnemia Stephens, 1836:168; Ulmer, 1907:182; 1953:353. ditional setiferous punctures. Anterior wings Mosely and Kimmins, moderately broad, densely pubescent; discoidal Type species: Hydropsyche senex Pictet, 1834. and and median cells always present; Ri simple; Ocelli absent. Maxillary palpi long, first cross-vein connecting C and Sc at about the second segment short, third long, fourth shorter centre of costal area. Posterior wings with dis- than third, fifth very long, annulated, flexible. coidal and median cells either present or ab- Antennae stout, basal segment short, bulbous. fork sent. A pair of lateral filaments arise from the Anterior wing with forks 1, 2, 3, 4 and 5, sternite 5. 2 sessile; discoidal cell long; posterior wing with forks 1, 2 and 5 present, fork 2 sessile; dis- Spurs 3:4:4. coidal cell short. Lateral filaments on sternite 5 Although Polycentropodidae was separated rather long, present in both sexes. Mid-tibia and from Psychomyidae and raised to family level tarsi of female dilated; spurs 3:4:4. as far back as 1906, occasionally it is still re- ferred to as a subfamily of the latter by some KEY TO SPECIES authors (Riek, 1970). Nevertheless, there is (Males only) sufficient evidence in the adult, as well as the 1. Anterior wing Cu 2 with thickening on the larval morphology, not only to regard it as a curve near arculus 2 family (Ross, but to split it even curve near good 1967), — . Cu without thickening on the 2 further; a group of genera, including the Aus- arculus 4 tralian Stenopsychodes, retained into family was 2. Superior appendages elongate, rounded, Stenopsychidae by Schmid (1969). without a process either on the inner sur- The only previous record—one species of face or near the base australica the family Polycentropodidae from Tasmania shorter, somewhat — . Superior appendages by Mosely and Kimmins (1953) —was based angular, with a process on the inner sur- on misidentified specimen (for details see Plec- face or near the base 3 trocnemia altera sp. n.). The family is now re- 3. Elongate, pointed, finger-like process at species corded from Tasmania with six new the base of superior appendages altera placed in three genera, one of them described trun- — . No process at the base, but instead a Poly- here as new. The third Australian genus, cate process arising from centre of inner plectropus, is not recorded from Tasmania. surface of superior appendanges . lacuna 4. Inferior appendages broadly KEY TO THE GENERA curved caudata 1. In posterior wing, forks 1, 2 and 5 pre- — . Inferior appendages double sent 3 folded manicata — . In posterior wing only forks 2 and 5 pre- sent 2 58 Plectrocnemia altera sp. n. Figures 277-286 2. In anterior wing fork 1 present, posterior Plectrocnemia australica, Kimmins non Banks, in wing with discoidal cell absent (genus not Mosely and Kimmins, 1953:354. recorded from Tasmania) Polyplectropus The examination of a series of Plectrocnemia — . In anterior wing fork 1 absent, posterior specimens from various Tasmanian localities wing with discoidal cell revealed that there is more than one species re- closed Nyctiophylax presented, and it was therefore necessary to 3. In posterior wing, fork 2 verif/ the identity by comparing the Tas- sessile Plectrocnemia manian material with specimens from Victoria including the — . In posterior wing, fork 2 with and New South Wales localities, footstalk Tasmanoplegas type of Plectrocnemia australica Banks from TASMANIAN CADDIS-FLIES 61 9-11 Mt Kosciusko, N.S.W. The type differed from Length of anterior wing: i 8-1 1 mm; 9 the published illustration of the genitalia (Fig. mm. 244) in Mosely and Kimmins (1953), which was prepared from a Tasmanian specimen, collected at Waddamana. The type of /\ austra- lica is figured here (Figs. 287-289) to illustrate the differences between the two species. The specimens from Cradle Mountain National Park which are here selected as type material of P, altera sp. n., agree with Kimmins's description (1953, p. 355) and figures. Insect greyish-brown, venter pale, anterior wings with irregular mottling. Antennae with first segment short, rounded, the subsequent ones short, smaller than the first, gradually increasing in length, pale, covered with moderately dense, dark pubescence. Mcso- scutum with a pair of round warts. In anterior wings fork 1 short; fork 2 long, sessile; fork 5 very broad; Clio with distinct thickening on the curve near arculus in both sexes. Joining point of A i to A, is separated from the joining . wings discoidal point of A- to A ;l In posterior cell short, fork 1 shorter than its footstalk; fork 2 sessile; fork 5 long and broad. Type material: Ho\olypc 3 (T4931), allotype genitalia: segment 10 membraneous, $ 9 (T4932), 6 1 5 ° St. Columba lulls, Pyengana, 21 phallus; an elongate, pointed, finger-like process Feb. 1971; r? River, Targa, 22 Feb. Feb, 1971; 1 ? St. Patricks situated at the base between superior append- River junction, 18 Feb. 1971; 3 c? I ? Huon-Pielon Plains nr. Scotts Peak, 2 Feb. 1965; ages and the hood-like cover. Inferior append- 1967; 1 S lluon 1972; 1 1 Buxton River nr. Maylieltl, 13 Nov. c? single segmented, very long, curved <£ ages Duck River 6 km SW of Roger River, 29 Nov. elephant's tusks. Phallus Strahan, 10 Dec. 1974; 3 6* upwards like a pair of 1974; 1 1 Mersey River, curved, with one dorso-ventrally flattened, Dip River Palls, 1 Dec. 1974; cJ Liena, 16 Nov. 1972. Ml specimens collected by A. apically bifurcate spine, and a pair of lateral, Neboiss (NMV). 13 Based on misidentification, this species was recorded by Mosely and'Kimmins (1953) from two Tasmanian localities. These specimens are now referred to P. altera, a new species des- cribed in this publication. The name P. aus- tralica Banks should be omitted from the Tas- manian list of species. Apart from being slightly smaller, P. aus- in tralica differs mainly in a number of points the male genitalia. The most distinct difference is the shape of the superior appendages, which are longer, more rounded and without the pointed, finger-like process at the base between the superior appendages and hood-like phallus cover. The upper spine of the phallus is shorter, and the base of inferior appendages somewhat bulbous. Holotype $ Mt Kosciusko, N.S.W. 5,000 ft., 8 Dec. 1931, Harvard Aust. Exped. P. J. Dar- lington, MCZ Type 22090. The type specimen seen. is now deposited in ANIC Canberra. Type 59 Plectrocnemia lacuna sp. n. Type material: Holotype $ (T4944), allotype paratypes (T4946-T4953) Figures 290-293 9 (T4945), 8 6 Lake Leake, Tas., 9 Nov. 1972, A. Neboiss anterior wings almost Greyish-brown species; (NMV). without mottling; fork 2 barely sessile, fork 4 Other material examined: Tasmania—3 S genitalia in general outline very similar vinces. to both P. australica and P. altera, but differing 60 Plectrocnemia manicata sp. n. in details. Segment 10 membraneous, rather slender, truncate apically; the hood-like phal- Figures 294-303 lus cover somewhat more slender; superior ap- Brownish species of moderate size, venter than in pendage more angular in lateral view yellowish; anterior wings densely covered with larger and australica, but compared with altera, yellowish and darker greyish-brown pubes- process at without the long, pointed, finger-like cence, irregularly mottled. Anterior wing with instead with a smaller truncate pro- the base, fork 1 slightly longer than its footstalk, fork 2 the centre of the inner surface. cess arising from sessile, fork 3 with very short footstalk, fork 4 does not reach the distal end The upper spine sessile, fork 5 not excessively wide. Posterior of the phallus. wing with fork 1 slightly longer than its foot- 2 genitalia very similar to that of altera, but stalk, fork 2 just reaching discoidal cell, sessile, short apex of sternite 8 broader, ventral keel extend- but it is likely that it could exist also with ing distally in form of a ridge almost to the footstalk in some specimens; fork 5 not exces- apex of the sternite; lateral lobes distally rather sively wide. First antennal segment enlarged, broad. rounded, the following ones smaller, elongate. lateral filament Length of anterior wing: $ 8-9 mm; $ 9 mm. Abdomen with well developed TASMANIAN CADDIS-FLIES 63 on sternite in 5 both sexes. Female with mid- Other material examined: Tasmania— 1 6 genitalia: segment 10 membraneous, short, in dorsal view with broad V-shaped excision apically; superior appendages in form of broad, angular plates, on the inside close to the lower margin, a number of short, peglike tubercles, each with long terminal bristle. Phallus a mem- braneous, cylindrical structure inside of which there is a long, dorso-ventrally flattened, upper lobe, protruding beyond outer casing, apex up- curved; below that, but not protruding, a pair of shorter, downturned lobes. A pair of curved, distally pointed filaments arising from the base of segment 10. Inferior appendages in form of broad, curved plates, with lower distal angle produced posteriorly. 9. unknown. Length of anterior wing: $ 8-8-5 mm. Type material: Holotype $ (T4976), 1 9 paratype (T4977) West Arthur Plains, Tas., 3 Feb. 1965, A. Neboiss (NMV); 2 3 paratypes (T4978-T4979) West Arthur Plains, Tas., 6 Feb. 1965, A. Neboiss (NMV). Other material examined: Tasmania— 1 o* Huon- Picton River junction, Type material: Holotype $ (T4954), allotype 18 Feb. 1967, A. Neboiss (NMV); 15 <$ 10 mis E Strahan, 6 Feb. 1967,'•*•*•E F 9 (T4955), 15 g 5 9 paratypes (T4956- Riek (ANIC). T4975) Hellyer River Gorge, Tas., 9 Feb. Distribution: Tasmania—NW and SW pro- 1971, A. Neboiss (NMV). vinces. — 64 ARTURS NEBOISS with forks 1, 2, 3, 4 and 5 present, fork 2 either with short footstalk or just touching dis- coidal cell; in posterior wing forks 1, 2 and 5 present, fork 2 with short footstalk. Lateral fila- ments on sternite 5 present in both sexes. Spurs 3:4:4. 62 Tasmanoplegas spilota sp. n. Figures 310-314 Head and thorax black, covered with golden hairs; antennae stout, about as long as anterior wings, yellowish-brown at base, gradually changing to dark brown distally, first segment somewhat rounded, slightly bigger than the following one. Anterior wings densely covered with black- ish-brown pubescence, interrupted by distinct white pattern—three distal spots, transverse band at distal third of the wing, broken in the middle, costal spot about midway, complete transverse band at basal third, small round spot and an angular one near the base. Wing vena- tion as given in generic diagnosis. Position of Genus Tasmanoplegas gen. n. cross-vein closing median cell in the anterior Type species: Tasmanoplegas spilota gen. et wing as well as the length of fork 3 differs in sp. n. the specimens examined, therefore in the male fork 3 and 4 both sessile, in the female both This genus is based on a single species dis- stalked. Posterior wings unicolorous, discoidal covered in South West Tasmania and captured cell short, forks 1, 2 and 5 present, fork 2 with in flight during daytime among low vegetation footstalk. on button grass plains in association with Lia- genitalia: segment 10 membraneous, short pota lavara Neb., and bears some resemblance $ and broad, superior appendages with long, to it. The wing colour pattern is unlike any downward curved, sickle shaped, upper branch, other Australian species of this family. and short, bilobed, lower branch; phallus cylin- Although possessing the main characteristics drical, distally slightly broadened, at the base of the genus Plectrocnemia, to which it appears with a pair of triangular lobes. Inferior ap- to be closely related, it may be separated by the pendages as large, curved plates. stalked fork 2 in posterior wing, different ar- 2 genitalia: lateral lobes of sternite 8 in rangement of superior appendages in male geni- form of two large, elongate, triangular plates, talia and differently formed female genitalia median lobe broad, distally rounded. the lateral lobes representing sternite 8 are Length of anterior wing: S 2 5-5-6 mm. large, elongate triangular. Type material: Holotype S (T49 80), allotype Ocelli absent; maxillary palpi with first two (T4981), 1 9 paratype (T4982) West segments short, third longer than the first two 9 Arthur Plains, Tas., 6 Feb. 1965, A. Neboiss together, fourth shorter than third, fifth long, (NMV); 1 paratype, 5 mis SW Waratah, annulated, flexible. Wing venation differing a 2000 ft., Tas. 17 Feb. 1963, I. F. B. Common slightly in sexes. An oblique cross vein between andM. S.Upton (ANIC). C and Sc at basal third of the wing; discoidal cell closed in both wings, elongate in the an- Distribution: Tasmania—SW and NW pro- terior, short in posterior wing. Anterior wing vinces. TASMANIAN CADDIS-FLIES 65 63 Nyctiophylax repandus sp. n. Figures 315-321 Anterior wings greyish-brown, abdomen yel- lowish-brown, ventral surface paler. Antennae with first segment short, bulbous, the following ones more or less elongate, with encircling me- dian ring of short, dark hairs. Lateral filament on sternite 5 slender, present in both sexes. In female mid-tibia and tarsi compressed, dilated. S genitalia: sternite 9 somewhat triangular in lateral view; segment 10 membraneous, broad, rounded distally; superior appendages in form of slightly curved plates; phallus membraneous, broad at base, cylindrical and flexible distally, encased by an upper and lower pair of pro- cesses, the upper pair pointed apically, lower ones excised ventrally at apical half, rounded at apex. Inferior appendages curved S-shaped, bluntly pointed at apex. 9 genitalia: with lateral plates rather angu- lar, widely separated by the broad sternite 8 which at the middle has a transverse edge; small, pointed lateral protuberances on seg- Genus Nyctiophylax Brauer ment 9. Nyctiophylax Brauer, 1865:419; Ulmer, 1907:186; Mosely and Kimmins, 1953:357. Type species: Nyctiophylax sinensis Brauer, 1865. Ocelli absent. Maxillary palpi with first two segments short, third about as long or longer than the first and second together, fourth slightly shorter than third, fifth shorter than the first four together. Antennae stout, about as long as anterior wings. Mesoscutum with a pair of round warts. Anterior wings with apical forks 2, 3, 4 and 5 present; fork 2 sessile; junc- tion of Ai with A 2 , and A 2 with A3 very close together. Posterior wing with forks 2 and 5 only; fork 2 sessile; discoidal cell closed in both wings. Lateral filaments on sternite 5 present in both sexes. Mid-tibia and tarsi of female dila- ted. Spurs 3:4:4. Only two specimens of this genus were pre- viously known from Australia, one male from the Northern Territory, representing N. parvus Mosely, 1953, and an undescribed female from Queensland. The genus is now represented in Length of anterior wing: $ 6-5-7-5 mm; 9 8 Tasmania by one new, endemic species. mm. 66 ARTURS NEBOISS Type material: Holotype $ (T4983), allotype Tribe Cheumatopsyche 9 (T4984), 6 2 1 9 paratypes (T4985- Hydropsychini Genus T4991) Scamander River, Upper Scamander, Subfamily Diplectrona Tas., 9 Nov. 1972, A. Neboiss (NMV); 5 $ Diplectroninae Genus paratypes (T4992-T4996) St. Columba Falls, KEY FOR SEPARATING Pyengana, Tas., 21 Feb. 1971, A. Neboiss TASMANIAN GENERA (NMV). 1. Abdominal sternite 5 with lateral process Other material examined: Tasmania—5 c? Prosser River 2 km W of Buckland, 7 Dec. 1974; 3 <$ Huon present 2 River nr. Blakes Opening, 9 Feb. 1966; 16 o* Huon- 5 without lateral pro- — . Abdominal sternite Picton River junction, 18 Feb. 1967 (specimens from cess Cheumatopsyche the last two localities are slightly smaller—5-5-6-5 mm). All specimens collected by A. Neboiss (NMV). 2. Male abdominal segments 6 and 7 with 5 Huon-Picton junction, 17 Feb. 1967, E. F. Riek TASMANLAN CADDIS-FLIES 67 genitalia: S posterior margin of tergite 9 de- 2 ? Lake River 5 km SW Delmont, 9 Nov. 1972; 45 9 abdomen terminates bluntly; posterior Genus Smicrophylax gen. n. margin of tergite 8 with broad, shallow V- Type species: Smicrophylax creektona gen. et shaped median excision; posterior margin of sp. n. sternal plate slightly concave. In the study of Smicridea species from North Length of anterior wing: $ 7-5-8-5 mm; ? and Central America, Flint (1974) reduced 8-10 mm. Rhyacophylax Muller to the status of a sub- genus of Smicridea McLachlan on the basis Cheu matopsyche rnodica ,U 5- that he was unable to separate the larvae and pupae of the two groups. Nevertheless, in the adult stage the two taxa are very distinct. .'. \**^ 1 Comparing these two American groups with \ J the Australian species included in the genus V\N /<"( Smicridea by authors, it became quite evident " " A l Lv NW \ that there are considerable morphological dif- \ V Y- \ ^r /~^ 1 j NE --'A? ferences which warrant the segregation of the n I i J ) Australian group at the generic level. The dif- ferences of the three taxa are shown in the fol- £ lowing table. ',: % vC v^f \ Y E* ^ Ocelli absent, maxillary palpi five-segmen- ..—-'-l "V?*^ flexible, J ^ - ted, terminal segment long, annulated, XfsE\ longer than the basal four segments together. Antennae shorter than the anterior wings, seg- 1 A \ \» \ )9: ,sw\ ment 3 and the subsequent ones with dark, - * V more or less distinct, oblique line. Head dor- sally with anterolateral warts present, broad, sometimes divided longitudinally, often indis- tinct; anteromesal wart absent, posterolateral '.* « » 40 HU 64 Iw warts large, well developed; dorsal sutures dis- tinct. Posterior wings broad, rounded. Abdom- Type material: Type 5 Australia, Victoria inal sternite 5 with slender lateral filament in (Edwards), McLachlan collection (BMNH). both sexes. The male abdominal segments 6 Type not seen. and 7 each with a pair of internal mem- braneous organs. Tibiae and tarsi of the inter- Material examined: Tasmania—26 c? 64 $ Derwent River 2 km NW Derwent Bridge, 12 Feb. 1971; 4 <$ mediate legs dilated and flattened in female. 14 ? Hellyer River Gorge, 9 Feb. 1971; 11 o* 2 $ Male genitalia resembling closely that of the Black Bobs Creek 15 km NW of Ouse, 9 Dec. 1974; genus Potamya from North America. 1 $ Hogarth Falls, Strahan, 10 Dec. 1974; 1 ^~^~—^^^ Genus Smicridea Rhyacophylax Smicrophylax Structure ^~~~-~~-~~^^ (sensu stricto) Anteromesal Well developed Well developed Absent wart of head Head—dorsal Absent Absent Well developed sutures Posterior Not distinctly Not distinctly Distinctly wing widened widened widened, rounded Ra+s and R^+5 R.i+5 separates Ra+a and R^+s parallel for a from R&« at a parallel for considerable distance sharp angle some distance R-M system R-M system R-M system parallel well separated approximate to but not very close from Cui to CUx M joins R some M joins R some M joins R close distance from Rs distance from Rs to Rs Lateral Absent Present Present filaments on sternite 5 Spurs 1:4:4 1:4:2 ( All the Australian species previously grouped tenor wings; the shape of sternite 8 separates in genus Smicridea should now be included in this species from Smicrophylax simplex. this new genus. Only two of the species are Length of anterior wing: s 6-7 mm; $ 65- known to occur in Tasmania. 7 5 mm. KEY TO TASMANIAN SPECIES (Males only) 1. Anterior wing white with blackish costal margin creektona Anterior — . wing with more or less pro- nounced blackish pattern on the entire wing simplex 65 Smicrophylax creektona sp. n. Figures 328-338 Head and thorax blackish, covered with long, white pubescence, eyes large, reddish- brown; anterior wings delicate, white, densely covered with white pubescence, only costal margin blackish; posterior wings white through- out. It is separated from a similar Victorian species S. parvula Mosely, by the absence of an internal, chitinous, median spine near the apex of phallus. S genitalia with tergite 9 strongly produced posteriorly, deeply cleft, apices upturned; phal- lus slender, dilated apically; inferior appen- dages long, two-segmented, harpago small. Type material: Holotype $ (T5114) Creek- $ differs from the male by its dull, uni- ton River nr. Dover, Tas., 14 Nov. 1972, A formly or slightly mottled, greyish brown an- Neboiss (NMV); allotype 9 (T5115) Huon- TASMANIAN CADDIS-FLIES 69 Picton River junction, Tas., 15 Nov. 1972, A. Neboiss (NMV); 15 $ paratypes (T5116- T5130) Creekton River nr. Dover, Tas., 14 Nov. 1972, A. Neboiss (NMV); 2 £ 1 $ paratypes (T5131-T5133) Huon-Picton River junction, Tas., 15 Nov. 1972, A. Neboiss (NMV). Other material examined: Tasmania—2 c? 1 9 Huon Plains nr. Scotts Peak, 2 Feb. 1965, A. Neboiss (NMV); 2 ? Wedge River 5 Dec. 1972, P. Zwick (NMV); 1 c? Arve River 10 km W of Geeveston, 15 Nov. 1972, A. Neboiss (NMV); 2 c? 1 S Hcnty River 12 km NW of Queenstown, 10 Feb. 1971, A. Neboiss (NMV); 1 o* Murchison River 4 km S of Tullah, 12 Dec. 1974, A. Neboiss (NMV); 1 J 10 ? Collingwood River bridge, Lyell h-way, 9 Dec. 1974, A. Neboiss (NMV). 53 c? 5 ? Murchison River, 5 Feb. 1967, E. F. Riek (ANIC). Distribution; Tasmania—SE, SW and NW pro- vinces. 66 Smicrophylax simplex (Jacquemart) comb. nov. Figures 339-345 Diplectrona simplex Jacquemart, 1965b:29. Type species: Smicridea grisea Mosely, 1933. The examination of the holotype male, Head with anterolateral warts present, nar- which is mounted as a microscope preparation row elongate, not divided longitudinally, an- on three glass slides, showed that it belongs to teromesa] wart absent, posterolateral warts the Smicrophylax group of species. It is sepa- large, well developed; dorsal sutures distinct. rated from the other Tasmanian species by the Antennae shorter than anterior wings; seg- blackish, patterned, anterior wings and the ment 3 and the subsequent ones each with a shape of the inferior appendages in the male, more or less distinct oblique line. On posterior and by the broader shape of sternite 8 in the wing fork 1 absent. Lateral filament on ab- female. dominal sternite 5 long and slender, present Length of anterior wing: $ 4-5-6 mm; $ 6-8 in both sexes. Tibiae and tarsi of the inter- mm. mediate legs dilated and laterally flattened in Cradle Mtn., Tas- Type material: Holotype $ female. Fringe of long, whitish hair on pos- Tonnoir (TRScNB). mania, 27 Jan. 1923, A. terior legs in both sexes, although more pro- Type seen. nounced in male. Male abdominal segments 6 Material examined: Tasmania—7 rT 68 ? Waldheim, and 7 without internal membraneous organs. Cradle Mtn. Nat. Park. 7 Feb. 1971, A. Neboiss Mtn. Nat. Park, Spurs 2:4:4. (NMV); 3 c? 10 $ Dove Lake, Cradle Lake Lilla, Cradle 9 Dec. 1972, P. Zwick (NMV); 1 70 ARTURS NEBOISS KEY TO THE SPECIES OF THE GENUS 7>/w? material: $ Melbourne, Australia (Ed- ASMICRIDEA wards), McLachlan collection (BMNH) 1. Male posterior wing twice as long as Type not seen. broad grisea Material examined: Tasmania 37 :-:' small creek, 8 Nov. 1972; 8 o* 43 ? Franklin River 20 km Derwent Bridge, 1# SW 11 Feb. 1971; 20 Distribution: Tasmania—all provinces; Vic- Genus Diplectrona Westwood toria; New South Wales; Queensland. Diplectrona Westwood, 1839:49; Ulmer, 1907:176; Mosely and Kimmins, 1933:336. Type species: Hydropsyche flavomaculata Ste- 68 Asmicridea grisea (Mosely) phens (1836), nee. Pictet (= Diplectrona je~ Figure 354 lix McLachlan, 1878). Smicridea grisea Mosely, 1933:217. Antennae slender, about as long or slightly Asmicridea grisea, Mosely and Kimmins, 1953:336. shorter than anterior wings. Maxillary palpi This species appears to be very closely re- 5-segmented; segment 1 very short, segment lated to edwardsi, the only reliable distin- 2 longer than segments 3 or 4; segment 5 long, guishing character is the much narrower pos- slender, flexible, barely as long as the first four terior wing in the male. More extensive ma- combined. Head dorsally with large antero- terial of grisea is required to investigate more lateral warts; anteromesal wart present; pos- fully the limits of variation in the anterior terolateral warts very large; dorsal sutures pattern, wing as well as to establish the mor- present. Anterior wing rather broad, forks 1, phological differences between the two species. 2, 3, 4 and 5 present; discoidal cell short. c5 genitalia of similar appearance to that Slender lateral filament on abdominal sternite of edwardsi. 5 present in both sexes. The male abdominal 9 unknown. segments 6 and 7 each with a pair of internal membraneous organs. Tibiae and tarsi of inter- mediate legs not dilated in female. Asmicridea grisea Spurs 2:4:4. Only a small number of specimens of this §*P genus are known from Tasmanian localities, K ..... IV'I ;.;...... j-Sj* and to them a total of four species names have J \ been applied. Diplectrona simplex Jacquemart v proved to be a member of the genus Smicro- NW * ( V ] \ phylax; D. bispinosa Jacquemart is more likely -^ f v K to be a Victorian species if the date and lo- " x • i i > i cality on the label is correct (Neboiss, 1974c). (V. X ~J X." >—^ { species lyella is v3 \<' (-- \ One new D. now added from « C ( i \ <>* | T^C\ Yt* 4 2( Western Tasmania. A large female from this "!* ti I % region (Arrowsmith Creek, Lyell h-way, 9 Dec. 1974, A. Neboiss) probably belongs to VlSE \ J another yet undescribed species. The species can be distinguished by the :\SW Y \ form of male genitalia and the differences are * : '^\ \ f" best appreciated by comparison of the draw- ings. KEY TO TASMANIAN SPECIES f%^::: ' ;: (Males only) JO »*>m .'.":."."> " \ « gaf 68 1. Coxopodite stout, widened apically . . 2 — . Coxopodite slender, not widened Type material: Type 6 Miena, Great Lake, apically 3 Tas., 3300 ft., Dec. 1929, H. M. Stephens, 2. Phallus with a pair of strongly chitinized Mosely collection (BMNH). Type not seen. claws near apex bispinosa — . Phallus with an irregular group of stout Material examined: Tasmania—2 c? Western Lakes, Jan. 1962, R. Cooper (NMV). spines at apex tasmanica Distribution: Tasmania—C province. 3. Anterior wing with cross-vein r-m close 72 ARTURS NEBOISS to cross-vein closing median cell; below Length of anterior wing; $ 10 mm. phallus a narrow, apically pointed and Type material: Type 3 National Park, Tas., downturned mesal plate lyella 3500 ft., 26 Dec. 1936, J. W. Evans (BMNH). — . Anterior wing with cross-vein r-m distad Type not seen. of median cell; without mesal plate below No new material has been available for phallus castanea study. Distribution: Tasmania SE province. 69 Diplectrona castanea Kimmins — Diplectrona castanea Kimmins in Mosely and Kim- 70 Diplectrona lyella sp. n. mins, 1953:342. Anterior wing with median cell shorter than Figures 355-362 discoidal cell; cross vein r-m distad of median Anterior wing with median cell as long or cell. The internal membraneous organs of seg- slightly longer than discoidal cell; cross vein ments 6 and 7 small, globular, not extending r-m very close or touching median cell. The beyond the width of one segment. Wings and internal membraneous organs of segments 6 male genitalia have been figured in the original and 7 small, globular, not extending beyond description. the width of one segment. £ genitalia with tergite 10 formed by two $ genitalia with a sharp, upcurved spine narrow, fringed lobes, separated dorsally by a arising from the lateral margin of segment 9, wide excision; phallus with one central lobe the lower inner margin turned inward below and two pairs of spines arising from the base; phallus. These probably are the 'pair of triangu- one pair short, upcurved, the second pair long, lar plates, set on edge' described by Kimmins slender, acute, extending to the apex of me- (1953) in D. castanea. Tergite 10 formed by dian lobe. Inferior appendages two-segmented, two narrow, fringed lobes, separated dorsally coxopodite stout, wider at apex; harpago short, by a wide excision. Phallus with a central lobe, apex acute and abruptly bent inwards. cleft apically and with minute protuberances on 9 unknown. either side at about the middle; two pairs of Diplectrona castanea 3US fl J \^>A- * 1 % Ni», *- 'jjj***^^-^ T"— J-' \ J rV 1 ^JMCTne 'A \ N 4* & ) It :; '"""^ ^^* w\ — ^L \ f I r* Jj ^ \ \L r^xjy-^-^ in fnx xTse \ : 5p : sw . \ \ \ ^vfN ^*. * 16 K »0Hn ." \ - ]}\ 69 TASMANIAN CADDIS-FLIES 73 spines arising near the base, one pair short, Further specimens of this species are required upcurved, the second pair long, slender, acute, to clarify the question of distribution, but so far extending to the apex of the median lobe; be- no specimens have been available to the present low phallus a narrow, triangular, apically author either from Victoria or Tasmania. pointed and downturned plate. Inferior appen- 8 genitalia: lateral margin of segment 9 ex- dages two-segmented, coxopodite stout, tending inwardly to a rounded lobe; phallus widened apically; harpago short, tapering and slightly curved downwards, with a pair of curved inwardly at apex. strongly chitinous spurs on the ventral side, 9 unknown. close to the apex; inferior appendages slender, two-segmented; Length of anterior wing: $ 9-5-10 mm. coxopodite with a spur near apex, harpago curved inwardly and apically Type material: Holotype $ (T5143) King pointed. The internal membraneous organs River, Tas., 5 Nov. 1961, N. Dobrotworsky of segments 6 and 7 large, oval, extending an- (NMV); paratype a (T5135) Hellyer River teriorly beyond the width of one segment. Gorge, Tas., 2 Dec, 1972, P. Zwick (NMV). 9 unknown. Distribution: Tasmania—NW province. Length of anterior wing: $ 12 mm. 7 1 Diplectrona bispinosa Jacquemart Diplectrona bi spinosa Figure 363 Mf Diplectrona bispinosa Jacquemart, 1965b:25; Neboiss, 1974c: 14. 11 The holotype male is dissected and mounted on three separate microscope slides with the t- +/ J-\ ) original label removed from the pin and glued 1 V onto the slide with the wings. The examination NW n Yp\ V if* j Ni / of this specimen shows that the length of the txr M \ V ij y anterior wing is slightly less than 12 mm and -J.- " not 21 mm as given in the description. The lo- ! ""TV l^ *'> cality label *• & + 'Sassafras ? 20.x. A. Tonnoir' was \ E^ fSr ( not apparently written by the collector himself, i w\ ""S. Jt nor does it include information regarding the ^ 4 State or year of collection, although '1923' was stated by Jacquemart in the original description, i )P . in which the locality also was misspelled as sw V 'Sassapras'. *- V The sequence of places visited by Tonnoir at M . the end of 1922 and early 1923 was recon- structed by Neboiss (1974c), who showed that 90 40 80 km ^ 71 the holotype of D. bispinosa was most likely collected at Sassafras in the Dandenong Ranges, Victoria, assuming that the collecting date was Type material: Holotype $ Sassafras ? 20.x. A. in fact 20 Oct. 1922, when Tonnoir was in Tonnoir (IRScNB), mounted on three micro- slides. Victoria. If, on the other hand, the date on the scope Type seen. original label has been 29 October (1922), in Distribution: Tasmania ? (Victoria ?) which the digit 4 9' has been mistaken for '0', 72 Diplectrona tasmanica Jacquemart then this corresponds to the time when Tonnoir was on the way from Burnie to Launceston and Figure 364 that created opportunity for him to collect in Diplectrona tasmanica Jacquemart, 1965b: 27. the vicinity of Sassafras, Tasmania, located This is the smallest of the Tasmanian Diplec- 1 5 km SE of Devonport. trona species. Anterior wings brownish; median 74 ARTURS NEBOISS cell as long or slightly longer than discoidal cell; SUPERFAMILY LIMNEPHILOIDEA cross-vein r-m close or touching median cell. Maxillary palpi with terminal segment not The internal membraneous organs of segments annulatcd. Larvae sub-eruciform or cruciform; 6 and small, globular, 7 not extending beyond head hypognathous; constructing portable cases. the width of one segment. $ genitalia with segment 9 short, hood- 9 Family PLECTROTARSIDAE Mosely shaped, lateral margin produced into an upward (1953) directed, pointed lobe; apical margin of tergite This endemic Australian family includes un- 9 with a short and wide central excavation, pos- usual moth-like, day-flying caddis flies. tero-lateral angles slightly produced, blunt. The main centre of distribution is in the Southeast Phallus slender, terminating with ventrally of Australia and Tasmania, with only one rare situated cluster of spines at the apex. Inferior species known from Southwestern Australia. appendages two-segmented, slender; coxopo- The adults can be observed flying around small dite long, somewhat cylindrical, harpago short, to medium size shrubs in curved inwards, apically pointed. damp areas in the vicinity of water during the middle of the day in 9 unknown. bright sunshine. So far all attempts to locate the Length of anterior wing: $ 6-6 5 mm. larvae have failed, although the first instar lar- vae were hatched from eggs by E. F. Riek (per- sonal communication). Family diagnosis: Ocelli present; antennae not exceeding the length of anterior wing, moder- ately stout to stout, basal segment more or less thickened; mouth parts either normal or elon- gate to form a proboscis; maxillary palpi cylin- drical, three-segmented in male, five -segmented in female. Mesoscutum with two pairs of indis- tinct warts, the anterior pair elongate, posterior pair close to scutellum, small, rounded. The warts on scutellum either rounded or elongate, somehow connected in the middle. Wings covered with dense pubescence; an- terior wing with forks 1, 2, 3 and 5 or 1, 2, 3, 4 and 5 present; posterior wing with Rj strongly bent downward where it joins Ra or ends blindly before reaching wing margin; discoidal cell small; strong frenular bristles at humeral angle. Male genitalia with inferior appendages two- segmented. Spurs 1:2:4; 1:4:4 or 2:2:4; tibiae and tarsi armed with strong spines. Type material: Holotype s Cradle Mtn., Tas., KEY TO GENERA OF PLECTROTARSIDAE 12 Jan. 1923, A. Tonnoir (IRScNB), specimen 1. Mouth parts distinctly elongate to form a dissected and mounted on three microscope proboscis; spurs 1:4:4 slides. Type seen. .... Plectrotarsus Mouth parts not elongate; mid-tibia with Material examined: Tasmania— 1 £ 10 km SW 2 spurs only Queenstown, 11 Feb. 2 1971, A. Neboiss (NMV). Spurs 2:2:4; posterior wing with anal field Distribution: Tasmania—NW province. broad with six anal veins .... Liapota I TASMANIAN CADDIS-FLIES 75 — . Spurs 1 :2:4; posterior wing with anal field separated by deep excision; superior append- narrow, with five anal ages short, slightly curved inward, apically veins Nanoplectrus rounded. Segment 10 hood-shaped, triangular in dorsal view, apex excised. Inferior append- Genus Plectrotarsus Kolenati ages two-segmented; coxopodite large, con- slender, digitiform, curved in- Plectrotarsus Kolenati, 1848:94; Ulmer, 1907:99; vex, harpago Cummings, 1914:25; Tillyard, 1918:647; Mosely, ward, arising from the apically truncate, upper 1936a:396; Mosely and Kimmins, 1953:21; Ne- angle of the coxopodite. Phallus widened at the boiss, 1959:91. middle, distal end dorso-ventrally compressed, Type species: Plectrotarsus gravenhorsti Kole- apex rounded. nati, 1848. $ abdomen terminates bluntly with large, Wings densely covered with pubescence; an- hood-shaped plate dorsally and a narrower, terior wing with forks 1, 3 and 5 or 1, 4 2, 2, 3, trough-shaped plate ventrally. and 5 present; discoidal cell long and narrow, Length anterior wing: $ 6-5-8-5 mm; 9 thyridial cell about the same length; posterior of 7-5-10 mm. wing with forks 1, 2 and 5 present in both sexes, discoidal cell short, anal field broad. ^—^— Mouth parts elongate in shape of a proboscis; Plectrotarsus gravenhorsti antennae about as long as anterior wing, stout; M segment 1 large, segment 2 small. i %> * ' Spurs 1:4:4. (. *T*tfC K pK' species in Tasmania, one of which P. Two =:} + tasmanicus is endemic. J~<"h 1 mV i KEY FOR SEPARATING TASMANIAN j A/ ) ^\£Tne y f - 3 t ' PLECTROTARSUS SPECIES 1. Anterior wing covered with orange-brown and dark, often iridescent blue pubes- 'i ' cence gravenhorsti -L n ) «\ Jf>' v. concolorous — . Anterior wing covered with dark brown pubescence, with narrow white transverse line across the wing at anasto- fe mosis tasmanicus V 73 Plectrotarsus gravenhorsti Kolenati SLH%Mm Figures 365-366 9 M Plectrotarsus gravenhorsti {gravenhorstii by many 73 authors) Kolenati, 1848:94; Walker, 1852:97; Ul- I , mer, 1904:56; 1905a:20; 1907:100; Mosely, 1936a: Neboiss, 399; Mosely and Kimmins, 1953:25; Type material: Type 'Australia occidental^, lo- 1959:94. cation unknown. Type not seen. This is one of the most colourful and well Material examined: Tasmania—2 o* 1 $ Bridge- known caddis-fly species. The orange-brown water, Derwent River, 21 Feb. 1964, A. L. Dyce and dark brownish-black, often iridescent blue and M. D. Murray (ANIC, NMV); 1 ? Rupert Pt. 3 mis N of Pieman R, 29 Dec. 1953, T. G. Camp- pubescence, often even with some small areas bell (ANIC); 3 c? 2 2 Waterhouse Estate 25 mis NE of white on anterior wings easily identifies this of Scottsdale, 17 Jan. 1948, Key, Carne, Kerr (ANIC, species. The pattern is variable. NMV). Distribution: Tasmania and pro- $ genitalia with posterior margin of sternite —N, NW SE 9 at the centre produced in two rounded lobes, vinces; Victoria; New South Wales. 76 ARTURS NEBOISS 74 Plectrotarsus tasmanicus Mosely Figures 367-373 Plectrotarsus tasmanicus Mosely, 1936a: 396; Mosely and Kimmins, 1953: 21; Neboiss, 1959:94; Jacque- mart, 1965b: 3. Plectrotarsus gravenhorsti var. Walker, 1852: 97. Plectrotarsus gravenhorsti, Banks nee. Kolenati, 1913: 234. This species is easily distinguished from P. gravenhorsti by the differently coloured anterior wing, which is dark brownish-black with a more or less conspicuous transverse line at the anas- tomosis extending across the entire wing, as well as two white spots, one near the base of fork 2, the other at the middle of the thyridial cell; a few whits hairs at several other places are less conspicuous. S genitalia very similar to that of P. gra- venhorsti, but superior appendages elongate tri- angular, apex blunt and the space separating in- ferior appendages in the ventral view is a nar- row V shape, nearly pointed at the base. 9 abdomen terminates bluntly with large dorsal hood and a ventral trough-shaped lip. Specimens from Southwest Tasmania (Lake Distribution: Tasmania—C, NW and SW pro- vinces. Pedder and Huon Plains) and those from King Island, differ slightly from the typical form as Genus Liapota Neboiss described from the vicinity of the Great Lake. Liapota Neboiss, 1959:95. The specimens from King Island on the average Tasmania Jacquemart, 1965a:2 syn. nov. are larger in size and in the male genitalia seg- Type species: Liapota lavara Neboiss, 1959. ment 10 is somewhat shorter, whereas the speci- Ocelli present; antennae stout, basal segment mens from the Southwestern localities are smal- large, segment 2 short, segment 3 and subse- ler, generally darker, with entirely black meso- quent ones slightly longer than second. Mouth thorax and slightly different shape of the phal- parts not produced into proboscis. Anterior lus. These differences are not considered to be wing with forks 1, 2, 3 and 5 present in both sufficient to warrant specific separation, but ap- sexes, all sessile; discoidal cell long and narrow, parently represent geographically separated thyridial cell about as long or slightly longer populations. than discoidal cell; posterior wing with forks 1, Length of anterior wing: g 8-9-5 mm; $ 8-11 2 and 5 present; four frenular bristles at hu- mm. meral angle. material: Type Type s Miena, Tas., Dec. Spurs 2:2:4. 1930, C. Parker (BMNH). Type not seen. Material examined; Tasmania —47 <$ 16 $ Penstock 75 Liapota lavara Neboiss Lagoon, 8 Jan. 1964, E. F. Riek (ANIC; NMV)- 1 <$ Little Pine Lagoon, 16 Feb. 1967, E. F Riek Figures 374-378 (ANIC); 17 c? 10 $ Derwent River 2 km NW Der- Liapota went Bridge, lavara Neboiss, 1959:95. 12 Feb. 1971, A. Neboiss (NMV)- 1 t? Tasmania caparti Jacquemart, 2 ? Lake St. Clair, 13 Feb. 1967, E. F. Riek (ANIC); 1965a:2 syn. nov. Fossey River 10 mis 1(?2? S of Hellyer Gorge 5 A small, very distinctive day-flying caddis-fly, Feb. 1967, E. F. Riek (ANIC); 3 c? 7 King ? Island, usually found in low Jan. J. scrubland, flying among 1907, A. Kershaw (NMV); 1 <$ Lake Pedder 1 the vegetation 31 Jan. 1965, A. Neboiss (NMV); 1 J Huon Plains during the middle of the day in nr. Scotts Peak, Feb. 8 1965, A. Neboiss (NMV). bright sunshine. The colourful, golden-yellow TASMANIAN CADDIS-FLIES 77 pubescence with dark-brown and white pattern mon and M. S. Upton (ANIC); 1 <$ same loc, 5 Feb. 1967, E. F. Riek (ANIC). is very characteristic and readily identifies the species. Distribution: Tasmania—NW and SW pro- 8 genitalia with inferior appendages two- vinces. segmented, proximal segment broad, curved Genus Nanoplectrus gen. nov. downward, distal segment small, turned down and inward; phallus with upper distal part Type species: Nanoplectrus truchanasi gen. et widened laterally. sp. n. $ abdomen terminates bluntly with a hood- Ocelli present; maxillary palp 3-segmented shaped dorsal plate with a pair of triangular in male; segment 1 short, segments 2 and 3 ventral plates. long, slender; mouth parts not produced into elliptic with forks 1, Length of anterior wing: $ 5-5-6 mm; 2 5 proboscis; anterior wing mm. 2, 3 and 5 present, discoidal cell closed, rather broad; posterior wing elongate oval, Ri bent the Liapota lavara downward and joined to R2 ; anal area of Ms wing not expanded, with five anal veins; three frenular bristles at the humeral angle. Male § genitalia with two-segmented inferior appen- dages. Cn Spurs 1:2:4; tibial spurs yellow, spines dark, brownish-black. ! ^f The wing venation and different spur formula -^ l V \ rvC_.Tne -' V separate this from the other genera. / / j 76 Nanoplectrus truchanasi sp. n. E^ : La "f\ ) «x Figures 379-389 . $ y~"~i, / J ; $*\ ^ J* J) Q: This small species was captured flying among /se\ low vegetation at the same time and locality \ : where numerous Liapota lavara specimens were v m collected. w/+ : similar plan to , r$ genitalia is basically of ' ^lkr^/^^"/ v\ j. that in L. lavara but differs in details; sternite w 8 midventrally produced into a broad, oval ' Mf lobe; lateral lobe of segment 9 narrow, truncate " . o - D M M 60 km ', \ ^ 75 apically; inferior appendages two-segmented, J 1 downturned. Phallus with lateral, slightly pos- Tas., Type material: Holotype t Cradle Mtn., teriorly directed projections. allotype Wilmot, Tas., Car- Carter and Lea; $ $ unknown. ter and Lea (SAM). Type seen. Length of anterior wing: S 4 mm. Holotype 6 of Tasmania caparti Jacq. Type material: Holotype & (T5205) Junction Cradle Mtn., Tas., 27 Jan. 1923, A. Tonnoir Creek, West Arthur Plains, Tas., 6 Feb. 1966, (IRScNB), dissected and mounted on five A. Neboiss (NMV). microscope slides. Type seen. Distribution: Tasmania—SW province. Material examined: Tasmania—25 c? 1 $ Junction Creek, West Arthur Plains, 6 Feb. 1966, A. Neboiss This species is dedicated to Olegas Truchan- (NMV); 7 2 $ West Arthur Plains, 7 Feb. 1965, c? as, a Lithuanian-born conservationist, explorer A. Neboiss (NMV); 5 c? Spring River, 4 Feb. 1966, naturalist, lost his life on 6 January A. Neboiss (NMV); 1 $ Wedge River 30 mis W of and who Maydena, 26 Feb. 1967, A. Neboiss (NMV); 1 $ 40 1972, while exploring the Gordon River Gorge mis of Maydena, 25 Feb. 1967, A. Neboiss W in Southwest Tasmania. (NMV); 1 o* 10 mis E Strahan, 20 Feb. 1963, Com- NEBOISS 78 ARTURS Genus Archaeophylax Kimmins Kimmins, Archaeophylax Kimmins in Mosely and 1953:27: Sehmid, 1955a:92. „i Mosely,w rt w Type species: Archaeophylax ochreus 1953. . roundedA A Spurs 1:2:2; anterior wings broad, wings long and at apices, discoidal cell in both narrow, longer than the footstalk. Forks 1, 2, 3 slightly pro- and 5 present in both wings. Face duced. The larvae without dorsal tubercle on the first abdominal segment, gills well developed, multi- branched; lateral line extends from third to eighth segment. The cases are straight or slightly to bent, constructed from moderately small coarse sand grains or pieces of vegetation. There are only two closely related species in Tasmania and they can be distinguished by the colour of the wings and details of the male geni- talia. 77 Archaeophylax ochreus Mosely Figures 386-389 Archaeophylax ochreus Mosely in Mosely and Kim- mins, 1953:27; Neboiss, 1958:163 (larva). of Australian cad- 10 Family LIMNEPHILIDAE Kolenati This is one of the largest dis-flies. Wings broad, rounded apically, colour (1859) variable from yellowish to greyish-brown, dark The family is widely distributed in a great specimens usually also having the tibiae and hemisphere, diversity of habitats in the northern tarsi blackish. small number of species but known from only a 3 genitalia: distal margin of tergite 9 pro- south of the equator. All of the known Aus- duced into a rounded lobe, below and distad of species occur in clear, rapidly flowing tralian which is the semi-membraneous tergite 10 as a mountain streams. short, medially deeply clefted, hood-shaped Family diagnosis: Ocelli always present; an- plate; superior appendages short, broader at tennae as long as, or slightly shorter than the base, apex truncate in lateral aspect; a pair of anterior wings, moderately stout, basal segment prominent, curved, apically downturned and enlarged; maxillary palpi in male three-seg- pointed processes arise on either side of phal- mented, in female five-segmented, slightly pu- lus; inferior appendages small, in the form of a bescent; labial palpi small. short rod; ventral margin of segment 9 ex- Legs with variable number of tibial spurs, ac- tended distally to a broad, apically truncate cording to the respective genera, the anterior plate, slightly excised in the middle. Phallus ones never with more than one spur; tibiae and consists of distally pointed median lobe with tarsi usually armed with strong spines. lateral flaps at the middle, and a pair of long Anterior wings with discoidal cell always parameres which ventrally carry elongate group closed, median cell absent; posterior wings al- of short bristles. ways much broader and less pubescent than the $ genitalia: abdomen terminates bluntly anterior ones, often iridescent, discoidal cell with broad, slightly notched dorsal plate. closed (except in the non-Australian subfamily Length of anterior wing: $ 5 15-18 mm. Apataniinae). Larvae of this widely distributed species are There is only one genus in Australia. usually found in calm water pools under stones TASMANIAN CADDIS-FL1FS 79 a small ad- in otherwise rapidly flowing mountain streams. only; anal claws short, each with Case—cylindrical, straight or slightly bent, ditional hook near the base. constructed of various materials of plant or Type material: Type $ Cuvier River, Tas., mineral origin. Plant material includes small Feb. 1941 (BMNH). Type not seen. bits of bark and '- sticks, pieces of dead leaves, Materia! examined: Tasmania— 21 cf 1 Ltik& other vegetable debris, arranged in criss-cross Dobson, 20 Feb. 1967, E. F. Riek and A. Ncboiss 1967, (ANIC; NMV); 3 78 Archaeophylax vernalis sp. n. Figures 390-393 Insect dark, smoky-brown, closely related to ochreus, but can be separated by its smaller genitalia. size, darker colour and details of male pro- $ genitalia: distal margin of tergite 9 duced into a rounded lobe; semi-membraneous hood-shaped tergite 10 with small median ex- cision; superior appendages short, triangular; a pair of prominent processes on either side of *o yiw K) phallus, slender, apically pointed without the . 80 ARTURS NEBOISS distinctive rounded excision near the base as in unexplored locality, which produced two of the ochreus; inferior appendages small, in form of three Tasmanian kokirid species. They repre- a short, apically truncate rod; ventral margin of sented two quite distinct genera, one of them segment 9 extended distally and forms a broadly having only a single spur on the anterior tibia, truncate plate; phallus with slenderly pointed although all other diagnostic features clearly median lobe, parameres with elongate group indicate its affinities with this family. Therefore of short bristles near the apex. the reduced number of anterior spurs are added 9 unknown. to the family diagnosis. The family was charac- Length of anterior wing: $ 14 mm. terized and recorded for the first time from Australia by Neboiss (1974b). Family diagnosis: Ocelli absent; antennae about as long as anterior wing, basal segment longer than broad, not bulbous; mouth parts elongate in form of proboscis; maxillary palpi cylindri- drical, 3-segmented in male, 5-segmented in female. Mesoscutum and scutellum each with a pair of warts. Anterior wing with R x curved forward, either joining Sc for a short distance or briefly connected to it basad of pterostigma before diverging with an even concave curve to the wing margin. Posterior wing with discoi- dal cell open; Rx ending blindly, becoming un- traceable distally, or joining Sc shortly before wing margin; 3-5 strong frenular bristles at humeral angle. Male genitalia with inferior ap- pendages single segmented. Spurs 1:4:4 or 2:4:4, covered with fine pu- bescence. KEY FOR SEPARATING AUSTRALIAN GENERA FROM THE TYPE GENUS (Males only) 1. Warts on scutellum short, rounded; dis- Type material; Holotype $ (T5206) Lake coidal cell in anterior wing Pedder, Tas., 12 Mar. 1972, A. Neboiss short (New Zealand) Kokiria (NMV). — . Warts on scutellum elongate, discoidal cell Distribution: Tasmania—SW province (known in anterior wing long 2 from the type locality only) 2. Tibial spurs short, 1 1 Family KOKIRIIDAE McFarlane 1:4:4 . . . , (Tasmania) Taskiropsyche (1964) — . Tibial spurs long, 2:4:4 3 3. Anterior wing with thyridial cell about as Specimens resembling the New Zealand spe- long as discoidal cies Kokiria miharo McFarlane, were found cell (Victoria) Tanjistomella among material collected in 1965 at Lake Ped- — -. Anterior wing with thyridial cell distinctly der, Southwest Tasmania. Several attempts longer than discoidal were made to obtain more material, but it was cell (Tasmania) Taskiria not until early 1972, when a single Kokiria like larva was found in the lake. Unfortunately, the Genus Taskiria gen. n. lake, together with part of the surrounding Type species: Taskiria austera gen. et sp. n. National Park, was flooded by the Hydro-elec- Ocelli absent. Mouth parts extended to a tric scheme, thus destroying this unique and slender proboscis; maxillary palp in male TASMANIAN CADDIS-FLIES 81 3-segmented, individual segments cylindrical, all ventrally open semi-tubular piece; sternite 10 approximately the same length; in female 5-seg- from the side lip-like, with V-shaped central in- mented; antennae stout, about as long as the cision and down-turned apices. anterior wing; the head dorsally with two pairs Length of anterior wing: $ 7-8 mm; 9. 7 5- of warts, the anterior pair small, oval, the pos- 8-25 mm. terior pair much larger, broadly elongate. Anterior wings with discoidal cell long, thy- ridial cell about 1-5 times longer than the dis- coidal cell; forks 2, 3 and 5 present, forks 2 and 5 sessile. Posterior wings distinctly wider than the anterior wings, discoidal cell open, forks 1, 2 and 5 present; 3-5 frenular bristles at the hu- meral angle. Spurs 2:4:4, strong, tapered to pointed apex. KEY FOR SEPARATING SPECIES (Males only) 1. Central lobe of segment 10 separated from lateral lobes by deep cleft . . . austera — . No deep cleft between central and lateral lobes mccubbini 79 Taskiria austera sp. n. Figures 394-404 Dark, blackish-brown species, head and thorax fuscous, nitid, covered with coarse, golden hairs; mouth parts and maxillary palpi brownish; abdomen dark brown, legs paler; an- terior wings fuscous, densely covered with dark, Type material: Holotype s (T5139), allotype rather coarse fuscous pubescence; posterior 9 (T5140), 15 $ 3 9 paratypes (T5141- wings hyaline, slightly darker distally, smoky T5158) West Arthur Plains, Tas., 6 Feb. 1965; covered with dense fuscous pubescence. Legs 2 $ paratypes (T5159-T5160) Junction slender, with strong, tapered tibial spurs, Creek, West Arthur Plains, Tas., 6 Feb. 1966; covered with fine pubescence; tibiae with 2 S paratypes (T5161-T5162) Wedge River, several strong spines; tarsal segments with Tas., 17 Feb. 1971; 1 $ paratype (T5163) spines arranged in two, more or less parallel Condominion Creek, Tas., 15 Feb. 1971; all size or only rows. Female about the same specimens collected by A. Neboiss (NMV). slightly larger, body and wing colouring similar Other materia} examined: Tasmania— 1 r? 1 ? West to the male. Arthur Plains, 6 Feb. 1965, A. Neboiss (NMV); 1 ? & genitalia: dorsal margin of segment 10 Spring River, 4 Feb. 1966 A. Neboiss (NMV); 1 $ 10 mis E Strahan, 6 Feb. 1967, E. F. Riek (ANIC). produced apically to a slender triangular cen- Distribution: Tasmania and pro- tral lobe, truncate at apex, and a pair of curved —NW SW vinces. lateral lobes, pointed apically and separated a deep cleft. Phallus from the centre lobe by 80 Taskiria mccubbini sp. n. long, curved downward and slightly expanded Figures 405-411 dorso-ventrally near the apex. Inferior appen- dages single segmented, somewhat broader at Dark brown species of medium size, slightly base, curved inward and slightly tapering dis- larger, but distinctly paler than T. austera. Eyes several short spines, tarsal tally. black; tibia with on segments the spines are arranged in some- 9 genitalia: segment 10 in form of a short, two 82 ARTURS NEBOISS what parallel rows. Wing venation as described palp in male 3-segmented, terminal segment for the genus. curved, longer than second, covered with short, $ genitalia distinct: central lobe of segment dense setae; antennae moderately slender, 10 excised at apex, not separated from lateral slightly shorter than the anterior wings, head lobes by deep cleft; inferior appendages broad dorsally with two pairs of warts, the anterior at base, abruptly tapering dorso-ventrally, pair rounded, posterior ones elongate, slightly somewhat flattened inwardly curved apices. larger. Phallus long and slender, curved downward. Anterior wing with discoidal cell long; thyri- 9 unknown. dial cell 1 5 times longer than the discoidal cell; Length of anterior wing: $ 9-9 5 mm. forks 2, 3 and 5 present, forks 2 and 5 sessile. Posterior wing about as wide or only slightly wider than the anterior wing; discoidal cell open, forks 2 and 5 present; 4-5 frenular bris- tles at the humeral angle. Spurs 1:4:4, rather short. 81 Taskiropsyche lacustris sp. n. Figures 412-420. In addition to the generic description, the following details characterize the species. Dark brown species of moderate size. Frons covered with stout, pale setae; eyes black; an- terior wing with a narrow group of short setae along the anal margin; posterior wing with 4-5 frenular bristles at the humeral angle, and a few, rather long, stiff bristles along the costal margin. Taskiropsyche lacustris ;U> 11 . A\ -_s* material: V^«£ Type Holotype s ( T5 1 64 ) , paratype v N pAi 6 (T5165) Lake Pedder, Tas., 1 Feb. 1965, I y't NW ! i A. Neboiss (NMV). \ (f 1 V j> NE Distribution: Tasmania—SW province (known ij \ " v from the type locality only). / -'-""n K^y "~V""ls, This species has been named after Mr C. ft j\ ^>- r >Cw cS (v< McCubbin, artist, entomologist and conserva- K?) \^ tionist, whose urge and enthusiasm to explore ,-*_.-'•' \ / 1 Southwest Tasmania, led to the discovery of a\?4j: 7 ^ this and many of the other species described in SE\ \ V this paper. \ )P SW \T \ ^C^2 • Genus Taskiropsyche gen. nov. */y Type species: Taskiropsyche lacustris gen. et ii sp. n. Ocelli absent. Mouth parts extended to a M «0 10 km r^\^J 81 short and rather broad proboscis; maxillary — ' TASMANIAN CADDIS-FLIES 83 c£ genitalia with segment 10 gradually taper- very small. Maxillary palpi in male upturned in ing to a narrow, rounded apex, below which front of the face, slightly curved, single seg- there is a pair of small lobes; inferior append- mented. Head dorsally with undivided antero- ages slender, in- and upward curved; phallus mesal, a pair of small anterolateral, and larger bilobed apically. posterolateral warts. Mesoscutum with two ? unknown. elongate warts, scutellum with one large me- Length of anterior wing: $ 8-5-9-5 mm. dian, posteriorly pointed wart. Type material: Holotype $ (T5166), 3 $ Spurs 2:4:4. paratypes (T5167-T5169) Lake Pedder, Tas., 1 Feb. 1965, A. Neboiss (NMV). 82 Tascuna ignota Neboiss Distribution: Tasmania—SW province (known Figures 421-425 from the type locality only). Tascuna ignota Neboiss, 1975:82. 12 Family OECONESIDAE Tillyard Anterior wings kept flat above the body, yellowish-brown, irregularly mottled, with dark- (1921) brown irregular spots, more or less distinct Family diagnosis: Ocelli absent. Antennae dark-brown transverse band at the apical about as long as, or slightly longer than anterior third; venation aberrant in male with reduction wings, first segment enlarged, shorter than head; in M-Cu sector and anal veins absent. Posterior maxillary palpi one or two-segmented in males, wings broad, uniformly yellowish-brown, with five-segmented in females. Wings broad, vena- the exception of mottling along the costa and tion differing in sexes, often reduced or modi- wing tip. Antennae slightly longer than the an- fied in males. R x joins R2 just before wing mar- terior wing. gin in both anterior and posterior wings in both 3 genitalia with a short, rounded median sexes. Mesoscutum with two elongate warts, lobe on dorsal margin of segment 9. Segment scutellum with a single median wart. 10 long and slender, deeply excised distally. Spurs 2:4:4. Superior appendages slender, rod-like. Inferior Only one genus with one species in Tas- mania. Tascuna ignota £Jfi The tribe Oeconesini in the family Sericosto- matidae was proposed by Tillyard (1921) to include three New Zealand genera Oecone- 1 v sus McLachlan, Pseudoeconesus McLachlan ' J and Zelandopsyche Tillyard. Another two gen- "Cn r^i 7 era were added to these by McFarlane (1960). NW ( W]\ another genus from Tas- Neboiss (1975) added ^J f \{ i mania and demonstrated that this tribe is suf- ~a_ ^aJI\J. ficiently distinct to be regarded as a family. It j closely resembles the family Goeridae, but dif- - a ( * fers from the latter by the wing venation, with ^C) \y &L Ij^B 'vV^j 1 ft and one 1 r '• ^"^ f c? Rj joining R2 before the wing margin, N i-d-i ^ or two-segmented maxillary palpi in male. "•'A \i \ - \J 7 Genus Tascuna Neboiss Tascuna Neboiss, 1975:82. 'IsA \ * ^\ r x/^*5 W c*f f x!W^ Type species: Tascuna ignota Neboiss, 1975. f ^ Anterior wings broad, sparsely covered with \y? ^v*& i in stiff, bristle-like hairs; venation aberrant ID ww.-. : ;• N. male, with anal veins completely absent. Pos- t ~S, 82 terior wings with broad anal field, discoidal cell 1 — . ARTURS NEBOISS appendages short, two-segmented; coxopodite more or less pointed apically; anterior semicircular in transversal section, ventrally wings evenly and densely covered with fine with acutely produced distal angles, from the hairs Tasimia upper margin arises a short, rounded median — . Inferior appendages very short and broad, process and a longer lateral process; harpago in anterior wings main veins covered with short, digitiform, the rounded apex dorsally scale-like hairs Tasiagma covered with a group of short, strongly chitin- Genus Tasimia Mosely ized spines. Phallus cylindrical with a pair of Tasimia Mosely. 1936a:404; Mosely and Kimmins, distally pointed and divergent paramcres. 1953:121; Riek, 1968:113. 9 unknown. Molanniclla Banks, 1939: 481; Mosely and Kimmins, 1953:147; Riek, 1968:113. Length of anterior wing: £ 16-17 mm. Type species: Tasimia palpata Mosely, 1936. Type material: Holotype $ y 2 $ paratypes The genus, in addition to the description Waldhcim, Cradle Mtn. Nat. Park, 7 Feb. given in the family diagnosis, is characterized 1971, A. Neboiss (NMV). 6 $ paratypes by its wing venation, diilering in the sexes. In 10 mis E of Strahan, 20 Feb. 1963, I. F. B. the male, anterior wing with forks 2 and 5 or 2, Common and M. S. Upton (ANIC); 1 <} para- 3 and 5 present, fork 2 either sessile or with type same loc. 6 Feb. 1967, E. F. Riek short footstalk; anal margin rolled over in a (ANIC). Type seen. long, narrow fold; posterior wing with forks 2 Distribution: Tasmania—-NW province. and 5 present, excision in wing margin at the 13 Family TASIMIIDAE Riek (1968) end of Cu 2 ; a row of stiff bristles along the Riek (1968) synonymized the genus Molan- basal half of the costa. In female the anterior niella Banks ( 1 939 ) with Tasimia Mosely and posterior wings with forks 2, 3 and 5 (1936a) and erected a new family to take this present; no fold along the anal margin of an- endemic Australian genus. The move was terior wing, and no incision at the posterior based on discovery of the larvae and the analy- wing margin. Small ventral processes on sternite sis of adult characters. It is considered to be 6 and 7 in males, on sternite 7 only in females. closely related to the family Lepidostomatidae All species are moderately small to medium of which no Australian species are known. To size and their distribution ranges from SE this family two South American genera have Queensland to Tasmania. The larvae live in been added. fast flowing, clear mountain streams. Riek Family diagnosis: Ocelli absent. Eyes covered ( 1968) gives a key for separating the described with short setae; antennae slightly longer than species, listing T. palpata Mosely as the only anterior wings, first segment long, thickened, Tasmanian species, but he apparently over- second very short, rounded; maxillary palpi 4- looked T. denticidata, described by Jacquemart segmented in male, 5-segmcnted in female; me- (1965). To these one new species from South soscutum with two rounded warts, separated West Tasmania is now added, bringing the total from the median line; anterior wings with re- number of species known from this region to duced jugal lobe. three, all endemic. Spurs 2:4:4; those of hind tibia situated well KEY FOR SEPARATING beyond middle. TASMANIAN SPECIES Larvae construct elongate, dome-shaped (Males only) cases, which resemble those of the family Glos- 1 Anterior wings with forks 2 and 5 sosomatidae, apart from the partial apical only 2 closure. — . Anterior wings with forks 2, 3 Two Australian genera are placed in this and 5 drepana family, one of which is here described as new. 2. Apex of phallus in dorso-ventral view with KEY TO THE GENERA broad lateral . projections . . . palpata (Males only) — . Apex of phallus in dorso-ventral view with 1. Inferior appendages curved, horn-like, lateral projections short ... denticulata TASMANIAN CADDIS-FLIES 85 83 Tasimia palpata Mosely Materia! examined: Tasmania—-1 o* Scumander River, Upper Scamander, 9 Nov. 1972; 1 5 km Figures 426-432 $ abdomen dorsally terminates in two rounded lobes, divided in the middle by a wide excision; ventrally somewhat flattened, with three-lobed central plate, similar to that in T. palpata, but lacking dark pigmented spot at the base of each of the lateral lobe. Length of anterior wing: $ 6-7 mm; $ 7-7 5 mm. Type material: Holotype & Ada Bay 28.xi.1922, A. Tonnoir (IRScNB), dissected and mounted on three microscope slides. Type seen. The locality 'Ada Bay' most likely is abbre- viation for Adventure Bay, Bruny Island, Tas- mania, where Tonnoir had collected a number Type material: Type * 9 Launceston, Cata- of other insects between 28 and 30 December, ract Gorge, Tas., Jan. 1923, C. L. Edwards the month 'xi' (November) is uncertain. (BMNH). Type not seen. 1922; . 86 ARTURS NEBOISS Tasimia denticulata Material examined: Tasmania—5 85 Tasimia drepana sp. n. Cui b joins Cu 2 very close to the wing margin; the main veins Figures 439-443 covered with scale-like hair; posterior wing with forks 2 and 5 present. An- Anterior wings yellowish-brown, faintly mot- tennae about as long as the anterior wing, first tled, in male with forks 2, 3 and 5 present; a segment long, thickened, second segment very narrow, rolled-over fold along the anal margin. short, rounded, the following ones elongate, This species is very close to denticulata, but slender. Maxillary palpi in male 4-segmented, differs from it by the presence of fork 3 in the held upwards in front of the face, segment 3 male anterior wing and the shape of phallus. the longest, slightly curved; terminal segment $ genitalia very similar to that of denticu- short. In female 5-segmented, segment 1 short. lata, but inferior appendages slightly less curved Eyes covered with short setae. and the phallus without the extended central lobe. £ unknown. 86 Tasiagma ciliata sp. n. Length of anterior wing: $ 6 mm. Figures 444-450 Type material: Holotype $ (T5170), 1 s Insect very pale-yellowish. Characters other paratype (T5171) Huon River 2 km W of than genitalia as given in the generic descrip- Huon-Picton River junction, Tas., 11 Feb. tion. 1966, A. Neboiss (NMV). $ genitalia; segment 9 narrow, ventrally Distribution: Tasmania —SW province (known produced to a broad, triangular plate, curved from the type locality only) and roundly excised apically; segment 10 short, TASMANIAN CADDIS-FLIES 87 broad, apically truncate; superior appendages 14 Family HELICOPSYCHIDAE Ulmer short, lying close to segment inferior 9; append- (1906) ages broadly oval, curved, with groups of setae Family diagnosis: Ocelli absent. Head dorsally on inner surface; phallus downturned, triangu- with a pair of large, densely hairy warts. Maxil- lar. Small processes on sternite 6 and 7. lary palpi 2-segmented in males*, 5-segmented 2 abdomen dorsally terminates with hood- in females, segment 1 long. Mesoscutum with a shaped plate, broadly cleft apically; ventral pair of rounded warts; scutellum rounded an- plate with a pair of central lobes, lateral mar- teriorly with a pair of somewhat oval warts. gins ridged. Posterior wings with well developed hamuli on Length of anterior wing; $ 7-7-5 mm; 9 7*5- basal half of anterior margin. 8 mm, Spurs 1:2:2; 2:2:2 or 2:2:4. The family has a world-wide distribution, but is most diverse in tropical countries. The genus Helicopsyche is so far the only one recognized from Australia and Tasmania. The larvae build very distinctive snail-like cases of small sand grains, and are found at- tached to larger stones in flowing water, usually in clear mountain streams. First Tasmanian record of larvae was made by Dyer (1879). * Some family descriptions give male maxillary palpi as being 3-segmented. Genus Helicopsyche Siebold Helicopsyche Siebold, 1856:38; McLachlan, 1880:58; Tillyard, 1924:312; Mosely and Kimmins, 1953:65; Flint, 1964:69. Type species: Helicopsyche shuttleworthi Sie- bold, 1856. Antennae about as long as the anterior wings, segment 1 very long, longer than head; gradually dilated at apex. Head dorsally with a pair of large, slightly curved warts, almost touching posteriorly. Maxillary palpi 2-seg- mented in males. Anterior wings densely pubes- cent, discoidal and thyridial cells both present, Type material: Holotype S (T5 172), allotype the latter very long. Posterior wings also densely 9 (T5173), 20 g 10 2 paratypes (T5174- pubescent, discoidal cell absent. T5203) Andover, York Rivulet, Tas., 4 Dec. Spurs 2:2:4. The external spur on anterior 1974, A. Neboiss; 1 S paratype (T5204) Port tibia reduced; the subapical pair on the pos- Arthur, Tas., 15 Feb. 1971, A. Neboiss terior tibia not far from the apical pair. Mid- (NMV). tibia with a row of small spines. Other material examined: Tasmania— 1 o* Ulver- SEPARATING stone 4 km NM, coastal waterfalls, 18 Nov. 1972; 1 first unknown. — . Frons and upper surface of segment ? of maxillary palpi with broad, scale-like Length of anterior wing: 6 5-6 mm. androconia; posterior wing without Type material: Type i Hobart, Tas., Dec. fold murrumba 1937, J. W. Evans (BMNH). Type not seen. Tasmania 1 cf Strickland 87 Helicopsyche bartona Mosely Material examined: — Avenue, Hobart, 5 Dec, 1974; 2 c? Nive River 2 km W Figures 451-455 of Bronte, 5 Dec. 1974; 1 r? Ins River tributary 15 km of Cradle Mtn, 13 Dec. 1974; 2 d Dip River Helicopsyche bartona Mosely in Mosely and Kimmins, W Falls, 1 Dec. 1974; 1 Guide River Falls, Ridgley, 1953:71. d 18 Nov. 1972. All specimens collected by A. Neboiss Abdominal sternite 3 to 5 strongly reticu- (NMV). lated, the following ones with paler reticulation. Distribution: Tasmania—SE, C, NW and N Anterior wings normal; cross-veins above dis- provinces. coidal cell not always present, only one anal vein present. Posterior wings with deep longi- 88 Helicopsyche murrumba Mosely tudinal fold from the base along the anal margin Figures 456-460 as far as the anostomosis. ilelicopsvche murrumba Mosely in Mosely and Kim- & genitalia with segment 10 elongate, dis- mins, 1953:72. tally slightly tapering, and with a somewhat In both sexes abdominal sternite 3, 4 and oval, apical incision; a pair of dorsal processes often also 5 strongly reticulated, the following arise at approximately the middle of the seg- sternites with pale reticulation; ventral pro- ment and bear several upwardly and outwardly- cess on sternite 6 stout, apically truncate. An- directed, strong spines. Superior appendages terior wing in male with a fold near the base in small, directed outward. Phallus curved, apex position of the first anal vein. The first segment slightly dilated. Inferior appendages narrow at of the male maxillary palpi dorsally with longi- base, apically widened and divided into two tudinal excavation which is covered with broad lobes, curved inward; a pair of small, apically androconia; similar androconia also on the clubbed processes arise from the base and are frons and the mesal surface of the first antennal visible in ventral view. segment. 8 genitalia with segment 10 rather narrow, deeply cleft, a strong dorsal bristle on either side of cleft near apex; close to the base a narrow central ridge and on either side a dis- tally directed process. Superior appendages small. Phallus membraneous, forked distally, apices curved upward and directed either pos- teriorly or more or less laterally. Inferior appen- dages with narrow base, widened to a broadly triangular, incurved plate; a pair of small clubbed processes ventrally. $ abdomen with tergite 9 formed by two triangular, distally pointed plates, the apices pigmented dark and curved downward; sepa- rated dorsally by a broad, mesal incision. Length of anterior wing: $ 5-6 mm; 9 6-7 mm. Type material: Type $ Murrumbidgee River, F.C.T., Jan. 1933, R. J. Tillyard; 3 <$ para- types Mt Kosciusko, N.S.W. 2700 ft., 20 Jan. 1889, McLachlan Collection (BMNH). Type not seen. TASMANTAN CADDIS-FLIES 89 Helicopsyche nurru mba Calocidae and discussed in the following para- IJf graphs. lip The re-examination of the type and para- n types of Tismana saneva Mosely revealed that J \ the two terminal segments of the maxillary palpi had been broken off in *A"7 the type, thus resulting v in erroneous diagnosis. The head dorsally has NW ( ""V CTne iAJf'' expandable filaments, which, together with the Tf' r wing venation, places the genus Tismana into / synonymy with Caloca, but the species saneva remains cS valid. A Genus Caenota, although having 3-segmented "S 6 V 3f/ f ] / i : maxillary palpi in the male, has basically simi- J\ 1. In posterior wing merges with Sc for a R x 2. Segment 10 with deep and broad median short distance distad of discoidal incision saneva cell 2 — . Segment 10 very broad, median incision — . In posterior wing Sc connected with R by narrow ascita a short cross-vein Tamasia 2. Warts on scutellum narrow, separated by space approximately equal to their own 89 Caloca tenia Mosely width; males with 5-scgmented maxillary Caloca tenia Mosely in Mosely and Kimmins, 1953: palpi Caloca 156. 3 genitalia with segment 10 longer than — . Warts on scutellum broad, separated by space less than half their own width; males wide, with small V-shaped median incision at with 3-segmented (maxillary apex and a number of strong, posteriorly palpi Caenota directed spines. Phallus with sagittate apex. In- ferior appendages with incurved lateral branch. Genus Caloca Mosely Wing venation and male genitalia have been Caloca Mosely in Mosely and Kimmins, 1953:153. figured in the original description. Tismana Mosely in Mosely and Kimmins, 1953:65. syn. nov. 9 unknown. Caloca straminea Mosely, 1953. Type species: Length of anterior wing: £ 8 mm. The genus was placed in the family Odonto- Type material: Type $ Mt Wellington, Tas., ceridae, although it did not agree with several Dec. 1937, J. W. Evans (BMNH). Type not of the family characteristics. The genus Tis- seen. mana was found to be synonymous for the No new material has been available for study. reasons explained above in the discussion of the family. Distribution: Tasmania—SE province. TASMANIAN CADDIS-FLIES 91 Caloca tenia 11 * " J. \"\^y - * V NW ( ""V \ ^vXv NV F N.V.J >c^^ r £ + i XJ > \ F ^ i^t'-- ,. -Ij, i [ / V '' ..( n^v>l i ,,'r^vv..\. XJ-..< M mt- . Vse \ Jk ; * '^\ ; ' r i x/^^j n ^t r^>v W « SOkrn ';• '. .' %i£#'*** 89 90 Caloca saneva (Mosely) comb. n. Material examined: 1 Only one species in Tasmania. 92 Caenota plicata Mosely Figures 468-479 Caenota plicata Mosely in Mosely and Kimmins, 1953:61. Antennae in male with large, flattened lobe at the back, and a small, membraneous branch in front of the segment 1; in female this seg- ment is not modified. Maxillary palpi 5-seg- mented, normal in female; 3-segmented, modi- fied irregular, apically with brush of long hairs in male. £ genitalia with segment 9 narrow dorsally. Segment 10 longer than wide, tapering dis- tally, apex with small mesal incision. Superior appendages rather small, finger-like. Inferior appendages broad at base, two branched; upper branch laterally slightly flattened, lower branch twisted apically. 9 abdomen terminates with dorsal median plate, with wide V-shaped apical incision; on either side of the plate there is a finger-like pro- Type material: Holotype $ (T5207) Lake St. cess, and below that a pair of broad, apically Clair, 14 Feb. 1956, E. T. Smith (NMV). truncate plates. Distribution: Tasmania—C province. Length of anterior wing: 6 11-13 mm; 2 12- Genus Caenota Mosely 14 mm. Caenota Mosely in Mosely and Kimmins, 1953:61 Type species: Caenota plicata Mosely, 1953. The first antennal segment normal in females, but in males with large protuberances at the back, lined with androconia and a small branch in front. Head in females with a pair of an- terior warts close to the base of antennae, pos- terolateral warts large, postocular warts normal; in male the postocular warts extended dorsally, but anterior and posterolateral warts absent, replaced by other structures. Pronotum with two pairs of warts in female, but only one pair of elongate ones in male. Mesoscutum without warts, scutellum with a pair of large warts, separated in the middle by a narrow space. Maxillary palpi 3-segmcnted and irregularly shaped in male; normal, 5-segmented in female. Mid-tibia and tarsi with spines. Wing venation normal in anterior wings of both sexes, with scales along the major longi- tudinal veins in the males; the anal margin of the posterior wings in males folded and covered with scales. TASMANIAN CADDIS-FLIES 93 Type material: Type $ Diggers Creek, Mt of costal margin covered with a row of stiff, Kosciusko area, N.S.W., 29-30 Jan. 1930, R. J. long bristles. Tillyard (BMNH). Type not seen. Spurs 2:2:4. Mid-tibia and tarsi with short spines. Material examined: Tasmania—7 c? 8 5 Corinna, 5 Nov. 1972; 10 21 Feb. 1971; 2 o* Wedge River, 17 Feb. 1971; 1 d* 93 Tamasia variegata Mosely Russell Falls, Nat. Park, 5 Dec. 1972, P. Zwick; Figures 484-492 2 <$ Mersey River, Liena, 16 Nov. 1972; 2 $ Huon- Picton River junction, 18 Feb. 1967; 1 c? 6 ? Huon Tamasia variegata Mosely, 1936a: 401; Mosely and River crossing, 16 Feb. 1971; 1 ? Hellyer River Kimmins, 1953:57; Jacquemart, 1965b:5. Gorge, 9 Feb. 1971; 2 <$ 1 ? same loc., 12 Dec. 1974; This blackish species is widely distributed 1 ? St. Patricks River, Targa, 22 Feb. 1971; 4 $ Marakoopa Caves, 15 Dec. 1974; 1 o* Eckberg Creek, throughout the state, except for SW province, 12 km SW of Roger River, 29 Nov. 1974; 2 c* 2 2 from which so far there are no records. It is Lilydale, a creek 2 km N, 16 Dec. 1974; 6 11 (where present); posterior ! below discoidal cell Tamasia variegata m# wings with hamuli or sparsely placed macro- ^%v> trichia along the basal half of costal margin. 1 Pronotum with one pair of elongate median without warts, scutel- \ warts only. Mesoscutum J ....* */ lum with a pair of small warts within unpig- V N mented area. A of short NW t Spurs 2:2:4. Mid-tibia with row j NE^-'A? spines. \ 1 originally des- r\ i The genus Alloecella was cribed by Banks (1939) in the family Molan- the family Beraei- A nidae and later transferred to dae by Mosely and Kimmins (1953). In a -—_ ^.y \ wL ( f 1 paper on the females of the British species of ^7 '.Ca. , I 1 >?^X "~M~iZs Beraeidae Kimmins (1951) discussed the po- sition of scutal warts and their value as diag- ' \ sw for nostic characters. Comparison with the Austra- k lian genus Alloecella revealed that the shape of \ UK sclerites and the position of warts is entirely dif- ferent. In Alloecella there are no warts on the mesoscutum and the shape of the scutellum is so 40 60 km * 93 proportionally shorter and somewhat more an- gular. These structures agree closely with those of Ouse, 9 Dec. 1974; 2 1 . Anterior wing with discoidal cell Other recorded localities: Tasmania—Scotts- closed Helicopha dale, Burnie. New South Wales—BathursL — . Anterior wing with discoidal cell Distribution: Tasmania—all provinces except open Alloecella SW; Victoria; New South Wales. 16 Family HELICOPHIDAE Mosely Genus Helicopha Mosely (1953) Helicopha Mosely in Mosely and Kimmins, 1953:148. Family diagnosis: Ocelli absent. Antennae about Type species: Helicopha astia Mosely, 1953. the same length or slightly shorter than an- Head dorsally with large posterolateral warts terior wings, first segment long, stout; second extending anteriorly almost to the base of an- segment short, the following ones elongate. tennae; a pair of small, rounded anterior warts Maxillary palpi slender, 5-segmented in both within a triangular area between the base of sexes, length of segments not differing greatly. antennae. Head dorsally with large posterolateral warts. Anterior wings with small discoidal cell; Wing venation differing in sexes; anterior wings hamuli along basal half of costal margin of the with discoidal cell either small or entirely ab- posterior wing, the distal ones more closely to- sent; a small hyaline area at anastomosis just gether. Median vein incomplete basally in both TASMANIAN CADDIS-FLIES 95 wings and both sexes, thus creating large, vein- Helicopha astia free areas, particularly noticeable in the pos- .-. terior wings. Two species in Tasmania. JJL KEY FOR SEPARATING \ * TASMANIAN SPECIES V (Males only) A NW I >NE 1. Segment 10 triangular, with stout ventro- D lateral process astia f # i ^J— — . Segment 10 parallel-sided, broad distally ^ with small median cleft, without ventro- v *\ iE^ lateral process delamarei \ # ' „«.•——' 1 i r J / 94 Helicopha astia Mosely f u Figures 493-495 , SW Helicopha astia Mosely in Mosely and Kimmins, 1953:149. * A .- Helicopha marlieri Jacquemart, 1965b: 17 syn. nov. S genitalia with segment 10 somewhat tri- angular, distally obtuse, bearing a few stiff !0 *0 SOkr . ."^yVJB 94 bristles at the apex, ventrolateral^ extended to stout processes with apices turned outward at a more or less sharp angle and armed with several the other—two pairs of wings; the slide which peg-like spines. Superior appendages small. In- should have contained the abdomen with geni- ferior appendages single segmented, directed talia could not be located. There were also upward; about midway, bent and directed dis- three microscope slides labelled 'Helicopha tally, apices slightly widened; a strong, curved, delamarei, Cradle 23.L23'; one slide contained apically tapered branch arises from the inner a part of an anterior wing and one posterior basal half, and a pair of small, inwardly di- wing, a second slide contained the head and rected projections at about the middle. parts of the thorax, the third, the abdomen with 9 abdomen broadly rounded at apex, a pair genitalia. The examination of these slides of small, slightly curved processes at the pos- left no doubt that the abdomen and geni- terolateral angles; darkly pigmented pattern talia labelled as "delamarei, Cradle 23.L23', on sternite 6 and 7. is the one figured and described as marlieri. The Length of anterior wing: $ 6-7 mm; 9 6-7 two small inwardly directed projections at the mm. middle of the inferior appendages are both Type material: Type $ Hythe, Tas., Nov. present, although they are not figured by Jac- details 1939; 3 8 1 $ paratypes same loc. and 4 $ quemart, 1965 fig. 13A, B, C. All other paratypes Mt Wellington, Dec. 1939, J. W. of the genitalia are also so similar to those of Evans (BMNH). Type not seen. Helicopha astia, that the two are considered The locality of the holotype g of Helicopha synonymous. marlieri Jacquemart, according to publication, The mislabelled slide containing the abdo- marlieri creates a discrepancy with is Mt Wellington, 24 Nov. 1922, A. Tonnoir men of H. (IRScNB). the published locality. Helicopha astia, of which The type material in the IRScNB collection marlieri is a synonym, is found only in the was examined by the present author and was southeast (Mt Wellington-Hythe), but dela- found to contain two microscope slides labelled marei entirely in the northwest; this leaves little 'Helicopha marlieri, Mt Wellington 24.xi.22'; doubt that the locality label also has been trans- one contained the head and parts of thorax, posed. — 96 ARTURS NEBOISS terminates with an api- The names on the slides containing wings 9 abdomen dorsally finger-like process correspond to the illustrations and their pub- cally truncate plate; a small, and 8 with darkly lished localities, but some veins are not figured on either side; sternites 6, 7 with a group and some are incorrectly interpreted. pigmented pattern; sternite 6 also No additional material has been available for of long, stiff bristles in the middle. 4-5-6-5 9 this study. Length of anterior wing: S mm; Other recorded localities: New South Wales 6-7 mm. Ebor. Type material: Holotype 6 Cradle Mtn., Tas., (IRScNB). Slide Distribution: Tasmania—SE province; New 23 Jan. 1923, A. Tonnoir South Wales. with abdomen and genitalia could not be lo- cated in the collection. The comments on iden- 95 Helicopha delamarei Jacquemart tity of the type and locality discussed under Figures 496-506 H. astia. Helicopha delamarei Jacquemart, 1965b: 1 5. Material examined: Tasmania—2 6* Corinna, 5 Nov. 1972; 3 6* Bull Creek, Cradle Mtn. Road, 13 £ genitalia with segment 10 elongate, some- Dec. 1974; 4^4? Bluff Hill Creek, 12 km S of rather broad what parallel-sided, hood-shaped, Marrawah, 30 Nov. 1974. All specimens collected by distally, with small median excision. Superior A. Neboiss (NMV). appendages small. Inferior appendages single Distribution: Tasmania—NW province. segmented, basal half directed upward, but at about the middle directed distally, apices in la- Genus Alloecella Banks clavate; or three-branched pro- teral view two Alloecella Banks, 1939:481; Mosely and Kimmins, cess arises from the inner basal section, but at 1953:142. about the middle there is a small inwardly di- Type species: Alloecella grisea Banks, 1939. rected projection. Sternite 7 without ventral Antennae about as long as anterior wings; ba- process. sal segment long, about as long as the head, sec- Genitalia figures prepared from a specimen ond and third segment short, the following ones collected at Corinna. slender. Maxillary palpi long, all segments slen- der, segment 1 only slightly shorter than seg- ment 2. Head dorsally with a pair of large pos- terolateral warts. Anterior wings with discoidal cell open; in some species males with more or less extensive longitudinal fold along R-Rs; males with one, females with two anal veins, position of anal veins differing in species and sexes. Basal half of costal margin in posterior wings with macrotrichia not distinctly closer to- gether distally. KEY TO TASMANIAN SPECIES 1. Males 2 — . Females 4 2. Anterior wing with longitudinal fold along R 3 . . . . pilosa — . Anterior wing without fold 3. Fold not extending beyond the middle of anterior wing; Ai terminating on wing margin grisea the — . Fold extending almost to the apex of wing; Ai terminating into Cu 2 longispina TASMANIAN CADDIS-FLIES 97 4. Ventral plates forming sternite 9 apically Alloecella grisea truncate grisea kM — Ventral plates . forming sternite 9 apically rounded 5 Si V 5. Distal end of plates forming sternite 9 smooth longispina ^^^ — Distal I >. \ ^5 . end of plates forming sternite 9 -r-r-V^V N r*£<( i I ridged pnosa ) r ^^i i 96 A lloecella grisea B anks ^-~^" ' x \ y. J> i \ A,v,/y v/ Figures 507-511 v *\ V3 v^i^ Alloecella grisea Banks, 1939:481. 3| 4 + &( Alloecella wameria Mosely in Mosely and Kimmins, _..—^..-'"' K *• ^"^ w\ Jl, ( ( i i 1953:144; Jacquemart, 1965b: 13, Neboiss, 1974c: J CS. 14. syn. nov. The type of grisea agrees well with the illus- trations of wameria in Mosely and Kimmins \ \ V ^W^^.^ ] swX j?x C (1953) and both are regarded as being con- * ' ^V f- specific. "V ^^7Av (j ih \j Anterior wings in the male with a fold along R not extending beyond the middle of the wing; Ai terminates at the wing margin in both sexes. JO «0 M»m 96 Head dorsally with a pair of large posterolateral warts, a pair of pale, rounded spots between Type of A. wameria: Holotype $ , Fern- them. shaw, Victoria, 21 Dec. 1884 from McLach- genitalia with segment 10 long, narrow, S lan's collection (BMNH). Type not seen. deeply clefted in the middle; lateral horns long, Jacquemart (1965b) erroneously gave Kim- slender, distally pointed, lying close paral- and mins as the author of wameria and the holotype lel to segment in lateral triangular, api- 10; view locality—'Burnie 27.x. 1922'. cally rounded protuberances represent superior Material examined: Tasmania-— 12 o* 4 ? Duck appendages. Inferior appendages terminate in River, 6 km SW of Roger River, 29 Nov. 1974; 10 o* acute inward directed apices, a small spur aris- 2 9 Hellyer River Gorge, 12 Dec, 1974; 1 Alloecella 98 Alloecella pilosa sp. n. Figures 523-528 Similar to, but slightly smaller than the two preceding species, distinguished by characteris- tic genitalia. In male anterior wing, Ai termin- ates in Cu 2 shortly before wing margin, in fe- male A 2 terminates at the wing margin. 6 genitalia with segment 10 slender, bifur- cate, deeply cleft in the middle, not extending as far as the distal end of the inferior appendages, mid-ventral margin of each process with a long, slender projection. Superior appendages short, digitiform. Inferior appendages tapering to acute, inwardly directed apices, inner basal margin extended to a flat, angular lobe. Phal- lus long, apex gradually widened and curved downward, a ventral, collarlike flap near base. Small spatulate ventral process on sternite 7. $ abdomen terminates with broad, rounded tt> *0 Wkm dorsal plate, slightly produced and cleft in the middle, ventral plates rounded, obliquely — TASMANIAN CADDIS-FLIES 99 ridged. Sternite 8 narrow, sparsely covered with Sericostomatinae by 'atrophied scutal warts'. long bristles. He did not name the genera to be included and Length of anterior wing: & 5-6 mm; $ 5-5- the genus Conoesucus is the only one which is 6-5 mm. recognized by its name: consequently, it be- comes the type genus. The absence of mesoscutal warts is also given as the only distinguishing character by Malicky (1973). He indicates that the distribution is not limited to Australia and New Zealand, but extends to Asia and Africa, but except for genus Conoesucus from Australia, does not name any other genera. The analysis of the Australian genera which do not possess mesoscutal warts, and were placed in the family Sericostomatidae by Mo- sely and Kimmins (1953), indicates that other structures of major importance also differ from typical sericostomatid species, and therefore the subfamily Conoesucinae is here raised to family level. The family diagnosis is prepared from the Australian species, hence some amend- ments might be necessary later, when more de- tails of genera found elsewhere become avail- able. This move eliminates the family Sericostoma- tidae s. str. from the Tasmanian and Australian list. Six Australian genera Coenoria, Matasia, Costora, Lingora and Conoesucus and Type material: Holotype $ (T5208), allo- Hampa, three Zealand genera Pycnocentria, Olin- type $ (T5209), 1 2 paratype (T5210) Lake New in Lilla, Cradle Mtn. Nat. Park, Tas., 14 Dec. ga and Conuxia are included this family. All other Australian genera which were incorpor- 1974, A. Neboiss (NMV); 8 $ paratypes ated by Mosely and in the Sericosto- (T5211-T5218) 4 km E of Liena, Tas., 17 Kimmins matidae are transferred to other families as Nov. 1972, A. Neboiss (NMV); 1 $ paratype (T5219) Dip River Falls 10 km S of Maw- follows: banna, Tas., 10 Dec. 1974, A. Neboiss (NMV). Antipodoecia to family Antipodoeciidae Tamasia to family Calocidae Other material examined: Tasmania— 1 c? West Arthur Plains, 3 Feb. 1965; 1 2 Murchison River 4 Caenota to family Calocidae km S of Tullah, 12 Dec, 1974; 1 <$ Condominion Tismana = Caloca to family Calocidae Creek nr. Mt Eliza, 9 Feb. 1965; 1 ? Hogarth Falls, Strahan, 10 Dec. 1974; 1 $ Russell Falls, Nat. Park, Helicopsyche to family Helicopsychidae 23 Feb. 1967. All specimens collected by A. Ne- Saetotricha to family Helicopsychidae boiss (NMV). Tasimia to family Tasimiidae Distribution: Tasmania and SE —N, NW, SW The New Zealand genera Oeconesus, Pseudo- provinces. econesus, Zelandopsyche, Tarapsyche and Zepsyche are now placed in the family Oecone- 1 7 Family CONOESUCIDAE Ross sidae, but three genera Beraeoptera, Pycnocen- stat. 1967 nov. trodes and Confluens still remain in Sericosto- In a similar fashion to the family Calocidae, matidae. Ross (1967) erected the subfamily Conoesu- Family diagnosis: Ocelli absent. Antennae cinae, stating only that it differs from subfamily moderately stout, shorter than anterior wings; 100 ARTURS NEBOISS basal segment long, segment 2 very short, seg- some of them are rather doubtful and based on ment 3 slightly longer than 2. Maxillary palpi male characteristics only. in males 1 to 3-segmented, often partly or en- KEY TO AUSTRALIAN GENERA OF THE tirely membraneous, upturned in front of the FAMILY CONOESUCIDAE face and modified; in female 5-segmented, nor- 1. Spurs 2:2:2 (not in Tasmania) Coenoria mal. Labial palpi 3-segmented, slender, segment — . Spurs 2:2:4 2 1 only slightly shorter than segment 2 or 3. 2. Anterior wing with base of fork 3 basad Head dorsally with a pair of oval warts, simi- of cross-vein r-m 3 lar in both sexes; postocular warts short. . Anterior wing with base of fork at, or Pronotal warts long, narrow. Mesoscutum with — 3 distad of cross-vein r-m deep, mesal depression anteriorly, warts absent; 4 scutellum with a pair of warts. 3. Mesonotum wider than long; male an- In females the basal margin of sternite 5 with terior wing with curved, longitudinal pigmented, transverse lines, a rounded loop groove; superior appendages in genitalia produced distally on either side. Males without absent Hampa such structure. — . Mesonotum longer than wide; male an- Anterior wing venation often modified in terior wing with one groove along Sc and another males, discoidal cell present in both sexes, al- along Cu 2 ; superior appendages in though sometimes modified in males of certain genitalia present Lingora species; hyaline line along M, widened at the 4. Posterior wing with fork 1 sessile, or at the distal end of thyridial cell and occasionally most with very short footstalk .... 5 extending along the cros-vein r-m below the — . Posterior wing with footstalk of fork 1 as discoidal cell; all three anal veins present, long as Ai the width of the fork . . Matasia ends at the wing margin some distance basad 5. Male with bilobed hinged process on the from arculus. Posterior wings with bristle-like frons Costora hairs along basal section of anterior margin; — . Male without bilobed process on discoidal cell present in both sexes. the frons Conoesucus Spurs 2:2:2 or 2:2:4; mid- and posterior tibiae densely pubescent, sometimes with small Genus Hampa Mosely spines. Hampa Mosely in Mosely and Kimmins, 1953:44. The separation from the family Sericostoma- Type species: Hampa patona Mosely, 1953. tidae sensu stricto is based on: This genus is very close to the New Zealand genus 1. absence of transverse line on sternite 5 Pycnocentria, but is separated from it by in males, having fork 2 sessile in the anterior wing. 2. absence of warts on mesoscutum, Head dorsally with pair of oval warts, similar in 3. absence of hyaline area along cross-vein both sexes; postocular warts small. Maxillary palpi closing discoidal cell, 2-segmented in males, segment 1 long, terminal 4. by Ai ending some distance basad from segment very short; in females 5-seg- arculus. mented, normal. Frons with flat, broad, bilobed process in the male. Pronotal warts long and A revision of the entire group is required to narrow; mesonotum wider than long, scutellum establish the genera to be included in this broadly triangular with pair of warts. Anterior family. The number of undescribed species wings with base of fork 3 basad of cross-vein from Australian mainland will undoubtedly in- r-m; hyaline line along M only. fluence the generic limitations, but such work is Spurs 2:2:4. outside the scope of this paper, and therefore no attempt has been made to change the generic 99 Hampa patona Mosely placings of some species at this stage. Figures 529-533 The Tasmanian species at present are placed Hampa patona Mosely in Mosely and Kimmins, in five genera, 1953: although the distinction between 44. TASMANIAN CADDIS-FLIES 101 In the male, anterior wing with a single lon- Material examined: Tasmania— 12 <$ 2 $ Henty River 12 Queenstown, gitudinal groove; maxillary palpi 2-segmented, km NW 10 Feb. 1971: 4 though Mosely expressed the belief that it was 3-segmented. Head dorsally with a pair of elon- gate warts; postocular warts small. Frons with broadly U-shaped process in the middle; mem- braneous structures on either side of it absent. Pronotum with a pair of long and narrow warts; mesoscutum with deep mesal depression; scutel- lum with a pair of short warts. Wing venation irregular in the male anterior wings; discoidal cell present in both wings and both sexes. In female anterior wings, a narrow hyaline line along M, widened at anastomosis; a small hya- line area across Qa& near arculus. Spurs 2:2:4. 1 00 Matasia satana Mosely Figures 534-539 Matasia satana Mosely, 1936a:411; Mosely and Kim- mins, 1953:42. Wings densely covered with dark, blackish pubescence; in males there are two longitudinal folds, discoidal cell distinctly Type material: Type s , 2 2 paratypes, Hamp- enlarged, an in- ton, N.S.W., Jan. 1918, R. J. Tillyard (BMNH). complete vein between R and Rs; in the centre Type not seen. a large area covered with short, upright, clavate ^ 102 ARTURS NEBOISS hairs. Sternite 7 with strong, apically rounded, Genus Costora Mosely flattened process; in female the process is short Costora Mosely, 1936a:403; Mosely and Kimmins, and triangular. 1953:45. Mosely, 1936. $ genitalia with tergite 9 as strongly chitin- Type species: Costora iena ized process; segment 10 slender, bifurcate, a Head dorsally with a pair of large warts; post- pair of small dorsal processes at about the ocular warts small; maxillary palpi 2-segmen- middle. Superior appendages absent. Inferior ted and modified in males, 5-segmented and appendages large, ear-shaped. normal in females. In the centre of frons in males a flat or rounded, bilobed process, pos- $ abdomen with posterior half of sternite 8 sibly a kind of scent organ; in most species unpigmented, sparsely covered with stiff there is also a membraneous structure located bristles; sternite 9 formed by a pair of widely between the scent organ and the eyes. Prono- separated, triangular, ventrally flattened plates. tum with a pair of elongate warts; scutellum Length of anterior wing: s 5-6 mm; 9 5-5-7 long, slightly curved warts. An- mm. with a pair of terior wings with narrow hyaline line along M, widened near anastomosis, a small hyaline Matasia satan; i spot on Cu 2 near arculus; discoidal cell in both wings and both sexes shorter than its footstalk. 1 In females a pale, broad, transverse band along v with scattered stiff bristles. 1 - . ' -+• '. Spurs 2:2:4. V . N NW i \ -^ i }~NE KEY TO THE SPECIES OF GENUS \ iJ COSTORA \1— 4 \\ '<^ (Males only) " "vrC 1 ^ 1. Anal veins in anterior wing short, fused \ E V and reduced delora .-.--.'' — . Anal veins in anterior wing normal, reach- SE \ ing close to anastomosis 2 i sV| 2. Anterior wing with short, longitudinal s, sw C fold at the base of M; line of distinct black *• scales along Cu near the base . . rotosca *%k 2 — . Anterior wing without a fold or line of i\ w dark scales 3 o n *0 SO km 3. Segment 10 in lateral view curved up- i>'»; 1 00 wards, gradually tapering; without apical processes 4 Type material: Type 3 Miena, Tas., Dec. 1929, — . Segment 10 in lateral view widened to H. M. Stephen (mounted in balsam), 1 $ large, rounded lobes on either side shortly paratype Miena, Tas., Dec. 1930, C. Parker before the apex; a pair of long apical pro- (BMNH) . Type not seen. cesses ebenina Material examined: Tasmania—47 Distribution: Tasmania—SE, C, SW and N pro- — . Mid-dorsal projection of tergite 9 minutely vinces. trilobed at apex luxata — TASMANIAN CADDIS-FLIES 103 6. Inferior appendages with ventromesal No additional material has been available projection long and slender .... ramosa for this study. Inferior — . appendages with ventro-mesal Distribution: Tasmania—C province. projection short and stout 7 7. Superior appendages slender, extending 102 Costora delora Mosely slightly beyond the apex of mid-dorsal Figures 540-545 projection of tergite 9 iena Costora delora Mosely in Mosely and Kimmins, — . Superior appendages short and stout, not 1953:49. reaching the apex of mid-dorsal projection The colour of specimens fade considerably of tergite 9 krene with time; this species is blackish-brown when 101 Costora iena Mosely alive, but fades to yellowish as noted by Mo- sely in the original description. Costora iena Mosely, 1936a:403; Mosely and Kim- mins, 1953:47. The male anterior wings have a conspicuous group of The male of this species has been described dark, upright hairs arising near the anal margin, close to the of the wing. and figured in the previous publications; the base Anal veins are female, although listed among the paratypes, fused; a shallow longitudinal groove along is densely covered with hairs. Wing has not been described. The species is distin- M venation in female guished by the long and slender superior appen- normal, with all three anal veins present. The bilobed scent organ the dages, which extend slightly beyond the apex on frons in the male rather narrow, deeply ex- of mid-dorsal projection of tergite 9. cised in the middle; on either side there is a Length of anterior wing: $ 11mm; $ 12 mm. membraneous structure usually extending up- wards beyond the base of antennae. Costora iena $ genitalia very distinct, inferior appen- dages armed with several strong spines. Ventral margin of segment 9 with two widely separated long processes. Broad, spatulate ventral process on sternite 7. ^"^* .1 \ \ v /^-r v-**^ $ abdomen with small, triangular ventral a nw ~y ( ] \ process on sternite 7; the pale transversal band with stiff bristles on sternite 8 interrupted in the middle; sternite 9 formed by two ventrally flattened plates, dorsally the abdomen ter- + minates with a pair of elongate, apically ^F xj \ \ p ^ $ t ;'. 1 JL 1 ^ / rounded lobes. Length anterior wing: 7-7-5 8-9 A \!L \ 3; X. of $ mm; $ ' \ v. / V. mm. Type material: Type $ Deloraine, Tas., 25 Dec. 1884, McLachlan Collection + ^y/^*j \>{t. (BMNH). v^\ f I w m? r Type not seen. Material examined: Tasmania— 10 c? 58 ? Scaman- der River, Upper Scamander, 9 Nov. 1972. Victoria 9 3 4 ? Yea River 7 km S of Glenburn, 1 Dec. 1972; TO «0 Mum * vJ 101 1 c? 3 $ 3 km SE of Taggerty, Little River, 6 Jan. ' 1972; 1 Costora delora Length of anterior wing: $ 6-7 5 mm; 9 7-9 mm. 103 Costora ehenina sp. n. Figures 546-552 When alive this species is entirely black, but Type material: Holotype S (T5256), allotype when preserved, gradually fades to yellowish- 9 (T5257), 4 $ paratypes (T5258-T5261) brown. Bull Creek, Cradle Mtn. Road, Tas., 13 Dec. Wing venation in both sexes normal. The 1974, A. Neboiss, 2 S paratypes (T5262- bilobed scent organ on frons in the male short T5263) Sassafras creek 4 km W of Mole and broad; membraneous structure on either Creek, Tas., 17 Nov. 1972, A. Neboiss; 15 g side small. 1 9 paratypes (T5264-T5279) Iris River $ genitalia: dorsal margin of tcrgitc 9 form- tributary 15 km W of Cradle Mtn., Tas., 13 ing a rather short, triangular process, seg- Dec. 1974, A. Neboiss (NMV). ment 10 long, median incision deep, reaching nearly to the base; on either side a ventrally Other material examined; Tasmania— 1 o* Franklin River, Frankford, 19 Nov. 1972, A. Neboiss (NMV); directed, rounded lobe near the apex; a long, 2 c? 1 9 Russell Falls, Nat. Park, 23 Feb. 1967, A. Ne- twisted process at the tip on either side. Phallus boiss (NMV); 9 normal, frons with the bilobed scent organ Other material examined: Tasmania— 1 <$ Iris River rather broad, the membraneous structures on tributary 15 km N of Cradle Mtn., 13 Dec. 1974, A. either side small. Neboiss (NMV). Broad ventral process on ster- nite 7. Distribution: Tasmania—NW and SE provin- c5 genitalia ces. with dorsal margin of tergite 9 developed into a triangular process; segment 10 105 Costora krene sp. long, n. bifurcate, curved upwards and tapering apically. Figures 554-555 Superior appendages short, stout. In- ferior small, appendages each with four, approxi- A brownish species, closely resembling mately ramosa, equal-sized, marginal projections, and but differing in details of male geni- talia. one long, slender, ventro-mesal projection. Wing venation normal. The bilobed scent organ $ unknown. on the frons broad, the membraneous structure small. Broad ventral process on ster- Length of anterior wing: s 6-7 mm. nite 7. S genitalia with dorsal margin of tergite 9 developed into a triangular process; segment 10 slender, bifurcate, curved upwards and tapering apically. Superior appendages short, stout. Inferior appendages each with five un- equal-sized marginal projections, the ventrome- sal projection short and stout, larger than the preceding marginal projection. 9 unknown. Length of anterior wing: $ 5-5 mm. Type material: Holotype $ Geeveston, Tas., 4 Feb. 1922, A. Tonnoir (IRScNB). Dissected and mounted on four microscope slides. Type examined. Date on labels, none of which has been written by Tonnoir, is incorrect, because he did not arrive in Tasmania until late October 1922 and departed early February 1923 (Ne- boiss 1974c). Other insects captured at Gee- veston and described by Tonnoir himself, were collected on 4 Dec. 1922, and it is reasonable Type material: Holotype $ (T5280) Spring to assume that this should also be the date for River, Tas., 5 Feb. 1966, A. Neboiss (NMV). the holotype of C. ramosa. Distribution: Tasmania—SW province. H 106 ARTURS NEBOISS 106 Costora seposita sp. n. The bilobed scent organ on frons very small; Figures 556-559 membraneous structure entirely absent. Broad ventral process on sternite 7. Dark, blackish-brown species. Wing venation $ genitalia with dorsal margin of tergite 9 normal. The bilobed scent organ as well as the extended distally to a moderately broad, api- membraneous structure on the frons present. cally trilobed process. Segment 10 bifurcate, Broad ventral process on sternite 7. slender, apically tapered, slightly curved up- 8 genitalia with segment 9 rather broad, wards; a pair of small, rounded dorsal projec- dorsal margin extended distally to a moderately tions near the base. Superior appendages short. broad, apically bilobed process; the segment 10 Inferior appendages curved, each ventro-me- bifurcate, rather broad, apically tapered, slightly sally extended to a flat trifid lobe. flattened dorso-ventrally and curved upwards; $ unknown. dorsally at the base a pair of short, flat pro- Length cesses, truncate at apex, lower angle slightly of anterior wing: $ 5*5 mm. produced. Superior appendages short and stout. Type material: Holotype $ (T5282), 1 s Inferior appendages broad with unequal-sized marginal projections. $ unknown. Length of anterior wing: S 7 mm. Costora seposita paratype (T5283) Waldheim, Cradle Mtn. Nat. Park, Tas., 7 Feb. 1971, A. Neboiss «Q IX.) km (NMV). X ^L£ - 106 L Distribution: Tasmania—NW province. Type material: Holotype $ (T5281) Creek 1 08 Costora rotosca Mosely near Marakoopa Caves, Tas., 15 Dec. 1974 A Figures 565-569 Neboiss (NMV). Costora rotosca Mosely in Mosely and Kimmins, Distribution: Tasmania—N province. 1953:49. Dark brown species. 107 Costora luxata sp. n. Anterior wings in male with a short fold along near the Figures 560-564 M base; a conspicuous line of dark scales along Cu2 . Wing Dark brown species. Wing venation normal. venation in both sexes normal, but differing TASMANIAN CADDIS-FLIES 107 from other species in the genus by having the Material examined: Tasmania— 1 <$ Ouse River 8 km W of Miena, 5 Dec. 1974; 1 c? Huon River nr. distal end of Cu2 connected by short veins to Blakes Opening, 9 Feb. 1966; 21 <$ 3 5 Huon-Picton Cuib as well as to the wing margin. The bilobed River junction, 18 Feb. 1967. All specimens collected scent organ on frons distinct, a pair of long by A. Neboiss (NMV), 5 <$ Huon-Picton junction, 17 Feb. 1967, E. F. Riek (AN1C); 2 o* Derwent bristles on each lobe; the membraneous struc- Bridge, 12 Feb. 1967, E. F. Riek (ANIC). tures not present. In female the anterior mar- Distribution: Tasmania—C and SW provinces. gin of sternite 5 with pigmented dark line, roundly produced on either side; sternite 7 Genus Lingora Mosely without ventral projection. In male a broad ven- Lingora Mosely, 1936a :406; Mosely and Kimmins, tral projection on sternite 7 present. 1953:93. $ genitalia with segment 9 broad, dorsal Type species: Lingora aurata Mosely, 1936. margin produced in the middle to apically trifid Head dorsally with a pair of large warts, projection. Segment 10 short, diverging apic- postocular warts small; antennae shorter than ally. Superior appendages short, dilated api- anterior wings, segment 1 about as long as the cally. Inferior appendages in lateral view nar- head, segment 2 and the subsequent segments row, small projections along outer margin, ven- short. Maxillary palpi single segmented, mem- tro-mesal projection wider, trifid apically. braneous, modified in males; 5-segmented, nor- 9 abdomen terminates with a pair of mal in females. Frons in males without bilobed rounded dorsal lobes; broad transversal band scent organ and without membraneous struc- on sternite 8 covered with stiff bristles. tures. Pronotum with a pair of long and narrow Length of anterior wing; 6 7-8 mm; 2 7-8-5 warts; mesoscutum without warts, but with mm. deep mesal depression; scutellum with a pair of elongate warts. Anterior wings in males with longitudinal grooves, venation irregular, pos- terior wings normal; in females wing venation normal, in the anterior wing a narrow hyaline line along M widened at anastomosis. Dis- coidal cell present in both wings and both sexes. In females the anterior margin of sternite 5 with curved, laterally produced darkly pig- mented line. Spurs 2:2:4. The two Tasmanian species are separated by the shape of inferior appendages. 109 Lingora aurata Mosely Figures 570-579 Lingora aurata Mosely, 1936a:407; Mosely and Kim- mins, 1953:93. Lingora caparti Jacquemart, 1965b:8 syn. nov. Anterior wings in males densely covered with yellowish-brown hairs, showing golden lustre in live specimens, two longitudinal grooves, one along Sc, the other along Cuo bent forward at anastomosis, specialized upright standing hairs on major veins; in females the anterior wings Type material: Type $ Tasmania (without de- with two patches of golden pubescence, one finite locality), J. W. Evans (BMNH). Type near the base of the wing, the second at anasto- not seen. mosis. 108 ARTURS NEBOISS genitalia S with tergite 9 produced in the Holotype 6 of Lingora caparti Jacquemart, middle to an apically pointed projection, the 4 N. Port 16 Dec. 1922', A. Tonnoir (IRScNB), lateral margins becoming membraneous and ex- dissected and mounted on three microscope tending below phallus to join segment 10 above, slides. Type examined and found that the geni- which is formed by a pair of blade-like proces- talia agree in all respects with those of aurata. ses. The membraneous plate below phallus was The two names apply to one species and cap- overlooked by Mosely (1936), who observed arti becomes an synonym of aurata. only the more sclerotized triangular apical sec- The locality 'N. Port' refers to the township tion, therefore the illustrated ventral aspect of of National Park, near Russell Falls (Neboiss the male genitalia is incomplete. Superior ap- 1974c). pendages clavate apically and situated at the 1 Material examined: Tasmania—2 J 2 $ St. Pat- base of segment 10. Inferior appendages ricks River, in Targa, 22 Feb. 1971; 2 c? 1 2 South Esk River, Evandale. 1 lateral view broad apically with slender stems. Mar. 1967; 4 c? 2 ? Mac- quarie River 8 km W of Campbell Town, 9 Nov A pair of long, slender, upcurved spines arising 1972; 5c?2? Lake River 5 km SW Delmont. 9 Nov. from the interrupted ventro-mesal 1972; 3 Leven River section of 3 nr. Heka, 17 Nov. 1972; 1 ? Mersey River. Liena, 16 Nov. segment 9. Broad, spatulate ventral process on 1972; 3 rf Dip River Falls, 1 Dec. 1974; 43 110 Lingora vesca sp. n. Figures 580-583 Anterior wings similar to those of aurata with two longitudinal grooves and specialized hairs; colour yellowish-brown, with golden lustre. 6 genitalia is characterized by diverging apices of segment 1 0, shorter and broader mem- braneous plate below phallus and rather short, narrow and (in lateral view) obliquely trun- cate apices of the inferior appendages. The middle projection of tergite 9 rounded apically; the long, slender, upcurved processes arising from the ventro-mesal margin bowed outward in the middle. Broad, spatulate ventral process on sternite 7 present. 9 unknown. Length of anterior wing: $ 5 -5 mm. Type material: Holotype <$ (T5284) North Esk River, Type material; Type 3 and paratype g Great Blessington, Tas., 1 Mar. 1967, A Lake, Miena, Tas., Dec. 1930, C. Parker Neboiss (NMV). (BMNH). Type not seen. Distribution: Tasmania—NE province. TASMANIAN CADDIS-FLIES 109 Lingora vesca TT ventral process 2 — . Abdominal sternite 7 without broad spatu- late ventral process 3 g 2. Abdominal segment 9 with lateral margin produced to somewhat triangular lobe norelus N — . Abdominal segment 9 with lateral margin NW curved jromus NE.J 3. Segment 10 in lateral view slightly curved *'**. 1 f upwards, apex broad, rounded digitijerus S cS (* H I turned upwards at almost a right angle, c"~'' v sw \ — . Posterior wing fork 1 sessile or nearly so brontensis 1 1 1 Conoesucus jromus Mosely 110 Figures 584-588 Conoesucus jromus Mosely, 1936a:409; Mosely and Genus Conoesucus Mosely Kimmins, 1953: 88. Conoesucus tnosefyi Jacquemart, 1965b: 12 syn. nov. Conocsucus Mosely, 1936a:408; Mosely and Kim- Dark, blackish species, both pairs of wings mins, 1953:87. densely pubescent; in anterior wing Cuo joins Type species: Conoesucus jromus Mosely, Cuib; in posterior wing Sc joins R shortly be- 1936. fore wing margin. In males the longitudinal fold Head dorsally with a pair of large, oval warts along R in the anterior wing reaches nearly as situated close to the eyes; postocular warts far as the distal end of the discoidal cell. Maxil- small, antennae shorter than anterior wings; lary palpi in male 3-segmented, flattened, seg- maxillary palpi in males 3-segmented, lying ments 1 and 3 short, segment 2 longer than 1 close to the face, directed upwards, segment 1 and 3 together. Broad spatulate ventral pro- short, segment 2 long, segment 3 modified and cess on sternite 7 in male, a small, pointed one differing in species; in females, maxillary palpi in female. 5-segmented and normal. Pronotum with a pair $ genitalia with dorsal margin of tergite 9 of elongate warts; mesoscutum without warts, produced distally into a pair of short, broad, but pair on scutellum. Anterior wings with a apically rounded processes. Superior append- narrow hyaline line along M, widened near an- ages very short, rounded. Segment 10 formed astomosis, a small hyaline spot near arculus; by a pair of parallel, laterally somewhat flat- usually a more or less distinct fold in males tened, apically upcurved processes. Phallus along near the base; discoidal cell present R parallel sided, rather large, slightly curved closed in both sexes, about as long or and downwards. Inferior appendages broad, con- longer than its footstalk. cave, two-branched, upper branch broad, apic- Spurs 2:2:4; mid-tibia sometimes with a ally rounded, lower branch smaller, the outer few small spines. margin between the two branches with irregu- KEY TO THE SPECIES OF GENUS lar, more or less distinct excisions which dif- CONOESUCUS fer even between the two sides in a single speci- (Males only) men; a pair of long processes with twisted 1. Abdominal sternite 7 with broad spatulate apices arise from the inner basal margin, curve ! 110 ARTURS NEBOISS Mosely 1 Conoesucus norelus up and protrude posteriorly on either side of 1 2 Figures 589-593 phallus. Kimmins, The examination of available specimens Conoesucus norelus Mosely in Mosely and 1953:90. show that the type of C. moselyi Jacquemart is Small, dark species, wings densely covered only a variant of C. fromus. with fine pubescence. Anterior wings in males 5 abdomen terminates bluntly, but can be with longitudinal fold along R; Cu2 in both other species by the shape of separated from palpi in males 3- sexes joins Cu t b. MaxiLlary ventral incision. segmented, flattened, segments 1 and 3 short, Length of anterior wing: $ 6-7*5 mm; 9 7-8 segment 2 longer than 1 and 3 together. Broad, mm. spatulate ventral process on sternite 7 in male, one in female. Conoesucus fromus a short, triangular * pro- M $ genitalia with segment 9 laterally duced into a somewhat triangular lobe; dorsal IX margin extended distally into a pair of broad, slightly curved processes. Superior appendages short, rounded. Segment 10 in the form of two y 1 >, \ *fc-_-^^^W ^V N c-A 7 slender, apically upcurved rods. Inferior ap- A NW ^v \ margin irregu- ^ J pendages curved upwards, outer larly serrate; an acute spur arising mesally near the base, directed inwards; a pair of long, slen- der processes arising from inner basal margin * {&( and situated on either side of phallus. y£% 2 abdomen terminates bluntly with end ter- y€r~^ J jfls gite concave in the middle; ventral incision ((SE \ wider near the base. \ Y_ Length of anterior wing; $ 5-6-5 mm; $ 6-7 '\swA \ r mm. Conoesucus norel us '.U: v> ':.'*" » « eOhm \ in £a , p - ^jTV^ffi 0$4 material: Type $ paratype 9 Miena, Type , ^\^> -Vkn r^< I Great Lake, Tas., Dec. 1930, C. Parker NW (A ^m \ A (BMNH). Type not seen. 1 /~^ TmlJaI Jacque- L A] ijMl Holotype 6 of Conoesucus moselyi 1 1 J mart, Burnie, Tas., 27 Oct. 1922, A. Tonnoir Specimen dissected and mounted (IRScNB). c^ + on four microscope slides. Type seen. V^ rA) ' Blackman 1 ) r \ Material examined: Tasmania—2 o* J) <=i. \ River 15 km NW Oatlands, 5 Dec. 1974; 1 1 r? North Esk River, 27 Nov. 1959, D. Scholes (AN1C). pi! Distribution: Tasmania—NE, N, C and NW M to *-• ^ 112 provinces. TASMANIAN CADDIS-FLIES 111 Type material: Type $ New Norfolk, Tas., Inferior appendages concave, broad at base, Mar. 1938, J. W. Evans (BMNH). Type not outer margin irregularly excised; a pair of long, seen. slender processes arise from the inner basal margin, curve protrude posteriorly on Material examined; Tasmania— 1 c? 1 ? Plenty up and River 6 km E of Moogara, 7 Dec. 1974; 1 tf Dip either side of phallus; a small mesal projection River Falls, 1 Dec. 1974; 5 Distribution: Tasmania—all provinces. 113 Conoesucus digitiferus Jacquemart Figures 594-599 Conoesucus digitiferus Jacquemart, 1965b :9. Dark, blackish species. The anterior wings in both sexes with basal section of R thickened and elevated, but without longitudinal fold; Cu2 ends at wing margin and a short cross-vein near the apex connects it with Cu x b. Maxillary palpi 3-segmented in male, segment 1 short, material: Holotype & 'Bradle' misspelling segment 2 about twice as long, broad; segment Type Cradle Mtn., Tas., 12 Jan. 1923, A. Ton- 3 membraneous, covered with brush of long for noir (IRScNB ) . Specimen dissected and hairs. Posterior wing in both sexes with Sc and mounted on three microscope slides. Type seen. R running separate to the wing margin. There Wald- are no ventral processes on the abdomen in Material examined: Tasmania— 18 c? 7 $ heim, Cradle Mtn. Nat. Park, 7 Feb. 1971, A. Ne- either sex, although in females the posterior boiss (NMV); 1 o* Cradle Mtn., Lake Dove, 9 Dec. margin of sternite 7 is broadly curved pos- 1972, P. Zwick (NMV); 4 o* 6 $ Lake Lilla, Cradle Mtn. Nat. Park, 14 Dec. 1974, A. Neboiss (NMV). teriorly and thickened. Distribution: Tasmania—NW province. S genitalia: segment 9 with dorsally ele- vated, transverse ridge, produced posteriorly in 114 Conoesucus nepotulus sp. n. the middle; distal margin extended to a pair Figures 600-603 of long, broad, slightly curved and apically rounded processes. Superior appendages short, Greyish-brown to brown species. Anterior longitudinal fold along rounded apically. Segment 10 formed by a pair wings without R; Cu2 margin, and a short cross-vein near of laterally compressed, slightly upcurved pro- ends at wing connects it with Posterior wings cesses. Phallus dilated laterally near the apex. the apex Qiib. 112 ARTURS NEBOISS with Sc and R running separately to wing mar- Other material examined: Tasmania— 14 r? 4 $ Guide River Falls nr. Ridgley, 18 Nov. 1972; 4 c? gin; fork 1 with footstalk. Maxillary palpi in Dove River, Cradle Mtn, Nat. Park, 14 Dec. 1974; male 3-segmented; segment 1 short, segment 1 c? Bull Creek, Cradle Mtn. Road, 13 Dec. 1974; 1 J River trib. 4 km Liena, 15 Dec. 1974; 1 g 2 about twice as long as segment 1, broad; seg- Mersey W Iris River trib. 15 km N of Cradle Mtn., 13 Dec. ment 3 membraneous, long, covered with long, 1974; 1 o* Arrowsmith creek 18 km SW Derwent dark hairs. Abdominal sternites without ven- Bridge, 9 Dec. 1974; 4 <$ 4 km E of Liena, small creek, 17 Nov. 1972; 1 Leven River nr. Heka, 17 tral processes in either sex. Conoesucus n epotul US the most with very short footstalk. Maxillary palpi in male 3-segmented; segment 1 short, segment 2 about twice as long as segment 1, broad; segment 3 membraneous, long, with short, sclerotized basal section, covered with long, dark hairs. Abdominal sternites without H ^"4 \ ? ventral processes in either sex. NW ( n JX£7 NE £ genitalia: segment 9 dorsally with ele- vated transversal ridge, produced posteriorly 'J ) '*~"\ at the middle; distal margin extended to a pair of short, broad, curved processes. Superior ap- + $i a pendages short, rounded. Segment 10 in form of --1a I two parallel, laterally flattened plates, distal \ NT ends curved upwards at almost a right angle, SE \ tapered apically. Phallus widened and rounded V I A apically. Inferior appendages short, basal sec- ';. :\ SW it \ tion of outer margin irregularly excised; a pair \ , • * '^\ ~{i r 7 $r^7 *v of long, slender processes arise from the inner } \ lJsi : protrude posteriorly . \5"\j. basal margin, curve up and pp on either side of phallus; a small mesal projec- 20 « >0Hm '"'-* tion on each process near the base. :. I 114 • , $ abdomen with small mesal dome, the me- dian incision in ventral plate widened near its Type material: Holotype S (T5285), allotype base. $ (T5286), 15 $ 15 $ paratypes (T5287- T5316) Dip River Falls, 10 km S of Maw- Length of anterior wing: S 7-8 mm; 9 8-5-10 banna, Tas., 1 Dec. 1974, A. Neboiss (NMV). mm. TASMANIAN CADDIS-FLIES 113 genus with a single species. In addition, Mosely and Kimmins (1953:165) doubtfully refer to a single ma'e form Deloraine as being close to Marilia bola Mosely. So far this species has not been confirmed. For the genus Caloca Mosely which was placed in this family (Mosely and Kimmins, 1953), a new family Calocidae was created by Ross (1967). Genus Atriplectides Mosely Atriplectides Mosely, 1936a:119; Mosely and Kim- mins, 1953:167; Jacquemart, 1965b: 17. Type species: Atriplectides dubia Mosely, 1936. Anterior wings long and narrow; discoidal cell small in both wings; posterior wings with broad anal field, a row of short bristles along costal margin. Male genitalia with two-seg- mented inferior appendages. Only one species in this genus. 116 A triplectides dubia Mosely Figures 610-614 Atriplectides dubia Mostly, 1936 i: 120: Mosely and Type material: Holotype «j (T53 17), allotype Kimmins, 1953:168; Jacquemart, 1965b:17, Ne- 1974c: 14. 9 (T5318), 11 $ 10 2 paratypes (T5319- boiss, T5339) 5 km W Bronte, small creek, Tas., 8 Wings dark fuscous, with dark, indistinct, Nov. 1972, A. Neboiss (NMV). irregular mottling. Basal half of the costal mar- gin of posterior wing with a row of macrotri- Other materia! examined: Tasmania—3 6" Fisher River, Pencil Pine Grove below Lake McKenzie dam, chia, distal ones slightly longer. Antennae 1 Collingwood River bridge, 15 Dec. 1974; 15 o* S slender, segment 1 somewhat bulbous, segment Lyell h-way, 9 Dec. 1974; 1 o* Mersey River nr. Liena, short, segment 3 and the sub- 16 Nov. 1972. All specimens collected by A. Neboiss 2 very rounded, (NMV). sequent ones long, cylindrical. Maxillary palpi Distribution: Tasmania—C, N and NW pro- with segment 1 short, segments 2 and 3 long vinces. and stout, segments 4 and 5 slender. S genitalia with distal margin of segment 9 18 Family ODONTOCERIDAE Wallengren in lateral view produced into a broad, blunt, (1891) triangular projection; segment 10 short and Family diagnosis: Ocelli absent. Antennae broad, with apical angles extended laterally. Su- longer than the anterior wings, basal segment perior appendages short, broad, with sub-acute two-segmented; large, second segment small. Maxillary palpi apices. Inferior appendages curved, wider pos- long and stout, five-segmented. Wing venation coxopodite long, slightly sometimes irregular in males, often differing teriorly; harpago short, somewhat squarish, ridge. Phallus stout, from that in the females. Discoidal cell present apex truncate with serrate and closed in both wings; median cell absent. curved downwards. Mesoscutum with mesal line only faintly in- $ abdomen terminates bluntly with a pair dicated; scutellum round and distinctly dome- of distally downturned, apically rounded dor- shaped, the wart appears to occupy most of the sal plates, separated in the middle by deep cleft; either side. sclerite. a small outer lobe on 10-12 Spurs 2:4:4. Length of anterior wing: S 5 mm; $ In Tasmania there is only one confirmed 12-13 mm. 114 ARTURS NEBOISS 1 4 2 Hellyer Gorge, 2 Feb. 1967; 1 d Lake Dove, Cradle Mtn., 30 Jan. 1967. All specimens of the above group were collected by E. F. Riek (ANIC). Victoria—from numerous southern and eastern localities. Other recorded localities: Penstock, Cradle Mtn. Lake Lilla, Hobart, Dcloraine. Distribution: Tasmania—all provinces except NE; Victoria. ] 9 Family CALAMOCERATIDAE Ulmer (1906) Family diagnosis: Ocelli absent. Antennae longer than the anterior wings, first segment thick, shorter than head. Maxillary palpi 5- or 6-segmented, hairy, terminal segment simple. Mesoscutum with two longitudinal lines of setiferous punctures, scutellum small, squar- ish. Anterior wing usually broad, somewhat triangular. Discoidal, median and thiyridial cells always present and closed; venation alike in both sexes; forks 1, 2, 3, 4 and 5 present. Posterior wings with discoidal cell open or Type material: Type $ Great Lake, Miena, closed, median cell open. Tas., Jan. 1931, C. Parker (BMNH). Type not Spurs 2:4:2, 2:4:3 or 2:4:4, those on the an- seen. terior tibia short. Material examined: Tasmania— 1 c? 3 $ Miena, Only one genus 3400 ft, 24 Jan. 1961, L. Couchman; 1 £ Lagoon of Anisocentropus in Australia, Islands, 5 Dec. 1974; Hi? Bradys Lake, 9 Dec. which is found also in Tasmania. 1974; 12 c? Derwent River 2 km NW of Derwent Bridge, 12 Feb. 1971; 1 ? Franklin River 20 km of Genus Anisocentropus SW Derwent Bridge, 11 Feb. 1971; 1 TASMANIAN CADDIS-FLIES 115 coloratus have been included to illustrate the differences. 117 Anisocentropus latijascia (Walker) Figures 615-621 Notidobia latijascia Walker, 1852:90. Goera elegans Walker, 1852:95. Anisocentropus latijascia, McLachlan, 1863:495; Ul- mer, 1906:54; Ulmer, 1929:168; Betten and Mo- sely, 1940:39: Mosely and Kimmins, 1953:172; Kimmins, 1958:167; Fisher, 1965:5; 1972:45. Anterior wings densely covered with tawny or yellowish pubescence, either with more or less distinct oblique dark brown band across the middle of the wing, or bicolorous with yellowish basal and dark brown apical half. In latter case the wing membrane at the basal section also is yellow. S genitalia with segment 10 broad, bent downwards, apical margin bearing a row of short spines, excised in the middle, apical angles extended to a rounded lateral lobe with a ven- tral claw-like process. Superior appendages stout, slightly clavate. Inferior appendages very short, either truncate or somewhat rounded South Esk River nr. Evandale, 1 Mar. 1967; 1 $ Tooms Lake, 4 Dec. 1974; 1 ? Sorell River 3 km N apically in lateral view, inner surface covered of Sorell, 8 Dec. 1974; 2 <$ Prosser river 2 km W of with a group of short, stout spines. Buckland, 7 Dec. 1974; 1 o* 3 ? Rubicon River 8 km SE of Sassafras, 2 Dec. 1974; 13 S 1 ? Derwent River 9 abdomen terminates bluntly with a pair of 3 km W of New Norfolk, 7 Dec. 1974. All specimens broad, rounded dorsal lobes; apical margin of collected by A. Neboiss (NMV). 6 ? Evandale, 1 (ANIC). Victoria— 12 <$ 5 tergite 9 more or less evenly rounded, not pro- Mar. 1967, E. F. Riek ? Moorabool River, Meredith, 12 Feb. 1959, A. Neboiss duced into a median projection; sternite 9 with (NMV); 7 o* 7 2 Lake Barracoota, 28 Jan. 1975, A. narrow, triangular apical lobes; dorsal pig- Neboiss (NMV); 1 c? 1 2 Murray River, Irymple, 25 Nov. 1964. A. Neboiss (NMV). New South Wales line broadly V-shaped. Sternite 8 with mented 1 1 d 4 ? Batemans Bay, 18 Feb. 1958, E. F. Riek broad. mid-ventral incision short and Other (ANIC); 1 c? Canberra, A.C.T., 10 Mar. 1958, E. F. details as figured. Riek (ANIC). pro- Length of anterior wing: $ 9-10-5 mm; 9 Distribution: Tasmania—N, E and SE 10-12 mm. vinces; Victoria; New South Wales. Type material: Type $ (BMNH) without lo- cality label, but erroneously described as com- Anisocentropus bicoloratus (Martynov) ing from 'North America'. Type seen. Figures 622-626 Type s of Goera elegans Walker (BMNH) Ganonema bicolorata Martynov, 1914:132. without locality, but also described as being Anisocentropus latijascia, Mosely and Kimmins, 1953: 172; Fisher, 1972:45. '? from North America'. Type seen. Anisocentropus bicoloratus, Fisher, 1965:3. history of the two types and their iden- The Mosely and Kimmins (1953), without ex- been discussed in detail by Kimmins tity have amining the type specimen, considered that In Victoria and New South Wales (1958). 'Martynov's Ganonema bicolorata is almost latijascia occasionally has been recorded from certainly a synonym' of Anisocentropus latijas- the same localities as bicoloratus. cia Walker. Later Kimmins (1958) analysed Material examined: Tasmania— 1 <$ 1 ? Macquarie the types of latijascia and elegans, and recorded River nr. Campbell Town, 9 Nov. 1972; 1 o* 1 9 Sca- the intention by Riek to describe a new species mander River, Upper Scamander, 9 Nov. 1972; 3 $ 1,6 ARTURS NHROISS oi Anisocentropus from New South Wales, but large, thickened, second segment small, usually this has not been done to date. about as long as wide. Maxillary palpi usually The examination of the type female of 5-segmented in both sexes, but segmentation Ganonema bicolorata Martynov by the present reduced in males of some genera; first segment author, clearly indicated that this species is not with a nodule on the mesal surface near apex, synonymous with latijascia, but agrees with A pair of slender, upturned, single segmented, specimens found in Victoria and New South finger-like processes or pilifers on the frons of Wa]es - males in most genera. In anterior wings Rj All specimens so far examined and identified joins R 2 shortly before the wing margin; an as bicoloratus, have distinctly bicolorous wings, oval sclerotized area on the anal margin, which the divisional line between the two colours be- sometimes is extended into a more or less dis- mg almost straight. The bright yellowish colour tinct lobe. Discoidal cell present and closed in of the basal half of the wing includes the pu- both pairs of wings; it can also be seen in wings bescence and part of the wing membrane, ex- with a longitudinal fold (males of genus Aust- tending to all thoracic segments. The intensity rheitbrus). of dark brown colouring of the apical half of Spurs?~ "4-4 .. the wing varies '. considerably. J! ' ' t *""* ,S kno n m Austraha a genitalia of the same general plan as that ,?° J *? ^"f *** fc 1 m ^ata^Wto 1973) in b&mia, but differing mainly in the shap T "? £™ f ' md ( &hm d 955* ) The mam of ' the inferior appendages, which are short, tri- *"$ *£"?* a ea ° .distribution nevertheless lies eastern angular in lateral view; inner margin produced m / f Australia where it is well represented in areas distally to a short digitiform process abdomen g construct 9 terminates with 'a pair of Vl« ' TT" ?* ^^ S ' lghtly T rounded dorsal cylmdncal CaSes niade from lobes; apical margin of tergite . coarse sand°Tg^ns. 9 extends to an elevated median projection; dorsal pigmented line curved; apical lobes of KEY TO TASMANIAN GENERA n e U ^*^P> 1- Scutel.um with pair of distinct warts 3 menmented.ted «££ Sternite TS*8 with • mid-ventral incision c™,^n.,~ -*u * i long, tapering anteriorly. - * ^ ^^terous J^S.Pictures 2 Length 2 of anterior wing; $ 9-105 mm; 9 , 2 *f 9 5-11 5 mm. * Mesoscutum with two rows of setiferous punctures (diverging posteriorly); prono- Type material: Type ? (Leningrad Museum) tUm Wlth two airs of warts Tasmanthrus 'Nov. Holl.' without exact locality. Type seen. P — MesoscutUtm with Or Material ' without warts, pro- examined: Victoria—30 rf 14 ? Yarra River, Warrandyte, 27 Feb. 19^4; 1 $ Kangaroo notum with a pair of warts Ground, Dec. 27 1954: 4 Buffalo River . 4. Anterior wing with sclerotized area n ., . of anal Distribution: Victoria, New South Wales. margin produced to a distinct lobe. Males 20 Family PHILORHEITHRIDAE Mosely without pilifers, antennae with segments 3 to 16 pectinate ,j Ramiheithrus 1 936 ' — • Anterior wing with sclerotized area of Family diagnosis: Ocelli absent. Antennae about anal margin not produced to a distinct as long or longer than the anterior wings in lobe; males with pilifers, antennae not males, slightly shorter in females; first segment pectinate Aphilorheithrus — TASMANIAN CADDIS-FLIES 117 Several very interesting morphological struc- pairs of warts, the anterior pair small, between tures are found in this family. The males in or slightly basad of the bases of the antennae, some genera have a pair of single segmented, close to the median suture; posterolateral warts slightly curved cylindrical processes or pilifers large, more or less elongate oval. Pronotum in front of the face. It is believed that they with lateral warts only; mesoscutum with or function as scent-organs. Of the five Tasmanian without warts according to species; scutellum genera, two Austrheithrus and Ramiheith- without warts. rus—do not possess the pilifers. Anterior wings in male with a longitudinal Antennal segments, particularly the first 10 fold along the middle of the wing; Rj joins R 2 or 15 in males, are often either modified or shortly before the wing margin in females, but bear special hairs, such as groups of conspicu- in males it varies according to species; discoi- ous long hairs, setae or papillae; modifications dal cell present in both wings and both sexes, of the shape of the first segment are also found. although often obscured in the anterior wings Some of the subsequent segments may be pec- of males by the fold. Sclerotized area on the tinate—as in genus Ramiheithrus, or with ex- anal margin extended to a narrow lobe. In cisions as in Austrheithrus and Kosrheithrus. posterior wings R x ends separately from R 2 at The position and shape of waits found on the the wing margin. head, pronotum, mesoscutum and scutellum are Two species recorded from Tasmania. varied and are important characters in taxo- nomic analysis. Usually those of the mesoscu- KEY FOR SEPARATING TASMANIAN SPECIES tum and scutellum have been used in family level, but within Philorheithridae they differ not 1. Mesoscutum without warts; male anten- only between genera but even between species. nal segments 4 to 7, each with mesal In the anterior wings, some variation occurs excavation glymma in the position of anal veins, and comparative — . Mesoscutum with a pair of distinct warts; male antennal segments 4 to 7 without ex- work shows that A 2 is always present, whereas cavations Ai is often either absent, or present in part ronewa only, while A3 is usually very short. Between the anal veins and wing margin in males of 1 1 8 Austrheithrus ronewa Mosely some species is a longitudinal blind vein, which Figures 627-632 starts in the vicinity of the sclerotized area and ends shortly before the arculus; sometimes it Austrheithrus ronewa Moselv in Mosely and Kimmins, 1953:192. forms a groove which is covered with long, dense hairs (Tasmanthrus). This species is distinguished from Austrhei- The tracheal openings on sternites 2 to 7 are thrus glymma by having a pair of distinct meso- each connected to the anterior and posterior scutal warts. margins of the sternites by a more or less dis- The first antennal segment long, longer than tinct dark line. head, upper mesal margin widened in male, normal in female; segment 2 very short, seg- Genus Austrheithrus Mosely ment 3 more than twice the length of second; the subsequent ones cylindrical. dorsally Austrheithrus Mosely in Mosely and Kimmins, 1953: Head 190. with distinct median suture; a pair of small an- Type species; Austrheithrus dubitans Mosely, terior warts situated between bases of antennae; 1953. the large posterolateral warts elongate oval, Maxillary palpi in male 3-segmented; seg- slightly curved; basal margin of the head with ment 1 very short, segments 2 and 3 long and small, pale spot on either side of median suture. slender; labial palpi 3-segmented, slightly Pronotum with a pair of rounded lateral warts longer than maxillary palpi; pilifers absent. In only; the mesoscutal warts elongate oval, very female, maxillary palpi normal, 5-segmented, close together and located anteriorly of the longer than labial palpi. Head dorsally with two middle. 118 ARTURS NEBOISS Anterior wings apparently with A t absent; A2 Mar. 1935; 1 9 paratype no locality, Nov. and A.t separated for a very short distance near 1938, J. W. Evans (BMNH). Types not seen. the base only; an oblique cross-vein between Cu Material examined: Tasmania— 10 Mitta Mitta River 8 km NE Ben- § genitalia with segment 10 short, formed ambra, 5 Feb. 1974, A. Neboiss (NMV). by a pair of strongly downcurved plates, fused Distribution: Tasmania NW, N, and dorsally. Superior appendages moderately long, — E SE provinces; Victoria; New South Wales. parallel, not clavatc at the apices. Inferior ap- pendages short, three branched, upper branch 1 1 Austrheithrus curved and tapered apically; centre branch the 9 glymma sp. n. longest, broader at base, gradually tapers to Figures 633-640 rounded apex. This species differs from ronewa by the ab- 9 abdomen terminates dorsally with a pair sence of mesoscutal warts and mesally exca- of subquadrate plates separated in the middle vated antennal segments in male. by narrow incision; sternite 8 with broad, U- Head dorsally with anterior pair of warts shaped, darkly pigmented area. round, and situated between the bases of an- Length of anterior wing; £ 10-12 mm; $ tennae; posterolateral warts short, rounded. 12-14 mm. Mesoscutal warts usually absent, although there are some specimens known which have a pair of small, rounded warts in the middle. In males antennal segment 1 expanded dorsomesally, covered with long hairs on mesal surface, seg- ment 2 short, segment 3 slightly longer, seg- ments 4 to 7 excavated mesally. The position of anal veins in anterior wings of both sexes similar to that in ronewa. In male R] does not join R3a but terminates at the wing margin. S genitalia of similar pattern to that in ronewa, but the superior appendages are cla- vate. Inferior appendages short, three-branched, upper branch curved and broadly spatulate api- cally, centre branch in lateral view narrow at base, broader apically. 9 abdomen terminates with a pair of angu- lar dorsal plates, separated by broad central incision. Length of anterior wing: s 10-12 mm; 9 1 1-13 mm. Type material: Uolotypt $ (T5340), allotype Type material: Type $ Mt Kosciusko, N.S.W., 9 (T5341) 5 £ 5 9 paratypes (T5342- 2700 ft., 22 Jan. 1885, McLachlan Collection T535 1 ) St. Patricks River, Targa, Tas., 22 Feb. (BMNH); 1 $ paratype New Norfolk, Tas., 1971, A. Neboiss (NMV). . . TASMANIAN CADDIS-FLIES 119 vated mesally. Head dorsally with a pair of Austrheithrus glymma small, rounded anterior warts, situated between the bases of antennae; posterolateral warts Ik large, oval. "x Anterior wings apically rounded in male, ) somewhat elongate in female. Ri joins R 2 in C N both sexes and both wings. Sclerotized area on \ A anal margin of anterior wing produced into a i \ distinct lobe. 1 K". i v \ \ - r J e '••.._ \ Only one species known from Tasmania. j ,.-• 120 Kosrheithrus remulus sp. n. +\ ?4 + ... fi^ ( \p\\ \E^ Figures 641-646 '"''" i .''^fV^-'--'/ t / j4 .^ ' greyish-brown species, but smaller p Dark, l i ^^ T' ji (iSE \ than tillyardi from New South Wales and Vic- toria. \ SW N A \ J§ $ genitalia with segment 9 hood-shaped, ter- minating with a pair of broad plates which are separated in the middle by a deep excision. Segment 10 is situated below segment 9, has distal angles produced into apically rounded V « Mkm VV 119 divergent lobes, the middle dorsal surface of raised to a high triangular crest. Phallus Other material examined: Tasmania 3 ' produced into a more or less distinct, apically W *0 to kin ". , . \ _^ 120 pointed plate; segment 2 small, segment 3 exca- _ 120 ARTURS NEBOISS Type material: Holotypc 3 (T5352), 1 & circular, shorter than lateral lobes. Phallus paratype (T5353), National Park, Russell broad, apex rounded. Inferior appendages fused Falls, Tas., 23 Feb. 1967, A. Neboiss (NMV); ventrally and appear as broad, bilobed plate. 1 6 paratype (T5354) loc, same 19 Feb. 9 unknown. 1971, A. Neboiss (NMV); 2 <$ paratypes Length of anterior wing: 6 9 mm. (T5355-T5356) same loc, 5 Dec. 1972, P. Zwick (NMV); 4 $ paratypes same loc, 16 Ramiheithrus kocinus Jan. 1965, E. F. Riek (ANIC). Other material examined: Tasmania—2 c? Hellyer River Gorge, 9 Feb. 1971, A. Neboiss (NMV); 1 . (NMV); 4 <$ Murchison River, 5 Feb. 1967, E. F. 1 Riek (ANIC); 1 J Hellyer Gorge, 4 Feb. 1967, E. F. -"_. j \ * /"" N Riek (ANIC). V ^f *pv H Distribution: Tasmania—NE, SE and NW pro- NE vinces. vj— S. tL-^^j^ f J ""-v Genus Ramiheithrus Neboiss Ramiheithrus Neboiss, 1974d:322. Type species: Ramiheithrus virgatus Neboiss, 1974. Antennae about as long as anterior wing, SE basal )P segments 3 to 16 pectinate in male, SW branches becoming shorter distally; the seg- ments of apical half simple. Maxillary palpi 5- segmented in male; pilifers absent. Mesoscutum and scutcllum each with a pair of rounded warts. Anterior wing without longitudinal fold, 121 apex broad, obliquely truncate; sclerotized area on the anal margin produced to a rounded lobe. Type material: Holotype g and 1 $ paratype Posterior wing with Sc fused with R : for a short Corinna, Tas. (small creek in forest), 5 Nov. distance, then separating; Sc terminates into 1972; A. Neboiss and G. Kocins (NMV). wing margin, Rj joins R 2 shortly before wing Distribution: Tasmania—NW province. margin. Only one species in Tasmania. Genus Aphilorheithrus Mosely Aphilorheithrus 1 Mosely, 1936a:417; Mosely and Kim- 2 1 Ramiheithrus kocinus Neboiss mins, 1953:186. Figures 647-652 Type species: Aphilorheithrus stepheni Mosely, Ramiheithrus kocinus Neboiss, 1974d:323. 1936. Blackish species with pale, irregular mott- Maxillary palpi 5-segmented in both sexes, ling on anterior wings. Head dorsally with a pair first segment short with well developed apical of rounded anterior warts, situated posteriorly nodule, other segments slender; in male the at the bases of antennae; posterolateral warts pilifers present. Head dorsally with two pairs of somewhat reniform; pronotum with two pairs warts; the anterior pair round, situated at about of warts. the middle of the head; posterolateral warts & genitalia with segment 9 dorsally produced very large and occupying most of the posterior into a large, deeply cleft hood, the inner surface portion of the head. Pronotum with two pairs of each side covered with short spines. Seg- of warts. Mesoscutum with a pair of slightly ment 10 in form of trilobed, hood-like plate oval warts which are only slightly larger than above phallus; the median lobe broad, semi- those of the scutellum. . TASMANIAN CADDIS-FLIES 121 Anterior wings without longitudinal fold in ventral part appears as a pair of squarish male; R x joins R2 shortly before wing margin plates. Sternite 9 in form of a short, broad and in both wings and both sexes. The first anal apically rounded plate. vein (Ai) usually not developed; long, A2 well Length of anterior wing: S 12-14 mm; 2 developed; A3 very short; an additional veinlet 13-17 mm. between the anal vein and wing margin. Sclero- tized area well developed but not produced into Aphilorheithrus stepheni a distinct lobe. KEY TO TASMANIAN SPECIES (Males only) 1 Head with dorsal median sulcus narrow 2 — . Head with dorsal median sulcus broad and deep luteolus 2. First anal vein in anterior wing present decoratus — . First anal vein in anterior wing absent 3 3. Tergite 9 in lateral view terminating with rounded apex pauxillus — . Tergite 9 in lateral view with apex pro- duced downward into triangular keel stepheni 122 Aphilorheithrus stepheni Mosely Figures 653-657 Aphilorheithrus stepheni Mosely, 1936a:418; Mosely and Kimmins, 1953:188. Detailed descriptions of this dark-brownish Type material: Type $ Miena, 3300 ft., Tas., species are to be found in the previous publica- Dec. 1929, H. M. Stephen (BMNH). Type not tions cited and are not repeated here. It should seen. only be added that warts on scutellum are situ- Material examined: Tasmania— 15 <$ 10 Crossing, Port Davey track, 8 Feb. 1965; 1 <$ same loc, 16 Feb. 1971; 10 rf 16 ? Mersey River, Liena, 16 Nov. 1972; 1 J nr. Marakoopa Caves, 15 Dec. 1974; 4 c? 2 $ Leven River nr. Heka, 17 Nov. 1972. All g specimens collected by A. Neboiss unless stated otherwise (NMV). 3 c? 3 ? Murchison River, 5 Feb. 1967, E. F. Riek (AN1C); 19 <$ 6 ? Forth Falls, 28 Jan. 1967, E. F. Riek (AN1C); 1 ? Iris River, Cradle Mtn. ft., 2250 31 Jan. 1967, E. F. Riek (AN1C); 1 ? North Esk River, Waisloca, 12 Dec. 1959, D. Scholes (ANIC). Victoria—7 c? 4 $ Gibbo River-Morass Creek junc- tion, 1 Feb. 1974; 16 Thomson River 1 km up- stream. Cowwarr weir, 26 Oct. 1973. All specimens collected by A. Neboiss (NMV). Other recorded localities; Tasmania—Delo- raine, Waratah; New South Wales—Kiandra, Mt Kosciusko, Upper Murrumbidgee River. Distribution: Tasmania—all provinces; Vic- toria; New South Wales. 123 A philorheithrus pauxillus sp. n. Figures 658-661 Smaller and darker species than A, stepheni, but differs in some details. Anterior wings with A x joined to Cu for a short distance near the base, but distally it be- 124 A philorheithrus decor atus sp. n. comes untraceable soon after separating from Figures 622-666 Cu. In posterior wings Rs joins R 2 before the cross-vein Sc-R. The arrangement of warts on This species has the most distinct, although the head, mesoscutum and scutellum is similar irregular colour pattern on anterior wings of all to that in stepheni. the Tasmanian Philorheithridae. Particularly S genitalia with tergite '9 narrowed to a noticeable is the pale lunular area bordered deeply cleft apex, which is not produced ven- with white and dark-brown at the apex of the trally. Segment 10 hood-shaped, apically trun- wing. cate and cleft in the middle. Phallus slightly The arrangement of warts on head, mesoscu- curved downward. Inferior appendages 2- tum and scutellum is similar to that in stepheni. segmented coxopodite long, distally with slightly Anterior wings with Ax present except for a expanded, inner apical angle, but not produced short distance between the cross-vein Cu-A and into a lobe; harpago small, clavate, apically the base, where it is indistinct. Posterior wing covered with group of stout teeth. Small ven- with Rs joining R2 at the cross-vein Sc-R. tral process on sternite 6 and 7. There are no ventral processes on sternites 6 or $ unknown. 7 in either sex. Length anterior 7-5-8 of wing: $ mm. $ genitalia with distal section of tergite 9 Type material: Holotype £ (T5357), 3 $ triangular, narrowly and deeply cleft, apices paratypes (T5358-T5360) St. Columba Falls, only slightly produced ventrally to rounded Pyengana, Tas., 21 Feb. 1971, A. Neboiss ridges. Segment 10 hood-shaped, triangular in (NMV). ventral view, cleft apically. Phallus curved Other material examined: Tasmania— 1 r? Hellyer downward, dorso-ventrally flattened, apex trun- River Gorge, 12 Dec. 1974, A. Neboiss (NMV). cate or slightly concave. Inferior appendages Distribution: Tasmania—NE and NW pro- two-segmented, coxopodite slightly curved, vinces. widened in the middle, truncate apically; har- TASMANIAN CADDIS-FLIES 123 pago round, curved inwards, apex truncate and Other material examined: Tasmania— 1 able before joining A . Lunular area at the .-^ 2 apex of wing similar to that in A. decoratus. 1 >. v ^* \N -A* / Posterior joining at the cross- wing with R x R2 ^J / v .:\ NW ( ""V vein Sc-R. nK. v NE \ -^ f V j> -c' / - Head with broad and deep dorsal median jT'\ J Tl 1 sulcus; anterior warts rounded, situated at about W^Tv - - CS . + large, curved, anterior end tapered to a narrow \^ $ ( >" point. Pilifers present. :^l --_, —\_, ^r^vv-.l J\ 0$i- S genitalia with posterolateral angles of ter- Vse\ gite 9 produced to acute points. Segment 10 Y \ i \ )p appears as a broad, apically rounded hood, with ;.\ SW \. ?*L "« a pair of elevated, triangular ridges dorsally *~ —\ r "V near the apex. Phallus curved downwards, apex '.-;. nv W M « «0 Km : >^ 124 Type material: Holotype & (T5361), allotype 9 (T5362), 1 S 1 9 paratypes (T5363- T5364) Corinna, Tas., 5 Nov. 1972; 2 S para- types (T5365-T5366) Lake Pedder, Tas. 1 Feb. 1965; 2 S paratypes (T5367-T5368) Con- dominion Creek, Tas., 15 Feb. 1971; 1 $ paratype (T5369) Farm Creek 4 km N of Tullah, Tas., 6 Nov. 1972; 1 s paratype (T5370) Damper Inn, Port Davey track Mt Mueller area, Tas., 12 Feb. 1965; 1 $ para- type (T5371) Pencil Pine River 6 km N of Cradle Mtn., Tas., 13 Dec. 1974; 2 & para- types (T5372-T5373) Waldheim, Cradle Mtn. Nat. Park, Tas., 7 Feb. 1971. All specimens collected by A. Neboiss (NMV). 4 S paratypes 10 mis E Strahan, 6 Feb. 1967, E. F. Riek (ANIC); 1 & paratype Mur- chison River, 5 Feb. 1967, E. F. Riek (ANIC). I 124 ARTURS NEBOISS depressed dorsally. Inferior appendages 2- The details described in the generic diagnosis segmented; coxopodite long, apex truncate; har- refer entirely to this species as the only member pago small, broad at base, narrowed distally of the genus. Ventral process on sternite 6 in and curved inwards, apex truncate and minutely both sexes very small, in males often entirely dentate. Sternites 6 and 7 without ventral absent. processes. 6 genitalia with tergite 9 short, slightly pro- ? slightly larger than male; anterior wings duced in the middle; superior appendages large, with similar colour pattern; genitalia not dis- broad at base, gradually tapering to rounded sected. apices. Segment 10 formed by two plates joined Length of anterior wing: £ 13-13 5 mm; $ in the middle. Phallus curved downward, vis- 15 5 mm. ible from below only. Inferior appendages single Type material: Holotype (T5374), allotype $ segmented, two-branched; the upper branch 9 (T5375), 2 t paratypes (T5641-T5642) directed upward and variable in shape, from Cracroft River Tas., 8 Feb. 1966, A. Neboiss a simple, (NMV). upright, digitiform process to one with laterally dilated base; the lower branch Distribution: Tasmania SW province. — dentate at the inner surface. Genus Tasmanthrus Mosely 9 abdomen terminates bluntly with apical Tasmanthrus Mosely, 1936a:414; Mosely and Kim- margin of segment 9 excised at the middle to mins, 1953:183. form small, somewhat triangular plates on either Type species: Tasmanthrus angustipennis Mo- side. sely, 1936. Length of anterior wing: Maxillary 6 9-11 mm; $ 12-14 palpi in male 4-segmented, all seg- mm. ments modified, short, partly membraneous; pilifers present. In female maxillary palpi nor- Tasmanthrus angustipennis \ \ mal, 5-segmented. Antennae about as long as the anterior wing, segment 1 longer than head, stout; in male with longitudinal mesal lobe. Head dorsally with obliquely situated, some- what rectangular posterolateral warts; anterior warts present only in female, sometimes partly or fully divided; absent in male, but instead there is a pair of membraneous papillae. Pro- notum with two pairs of warts, the middle pair distinctly elevated. Mesoscutum with two rows of posteriorly diverging setiferous punctures in the middle; scutellum with a pair of setiferous punctures near the anterior margin; there are no warts on mesoscutum or scutellum. Anterior wings with A x present, although sometimes absent or indistinct basally of cross- vein Cu-A in male; sclerotized area on the anal margin produced into a distinct, rounded lobe. R x joins R2 shortly before wing mar- gin in both wings and both sexes. 126 Tasmanthrus angustipennis Mosely Figures 673-682 Type material: Type s Miena, Great Lake, Tasmanthrus angustipennis Mosely, 1936a:414; Mo- Tas., Dec. 1930, C. Parker; paratype, sely and Kimmins, 1953:183. $ same loc, Dec. Tasmanthrus galbinomaculatus Jacquemart, 1965b: 19 1929, H. M. Stephen (BMNH). syn. nov. Type not seen. TASMANIAN CADDIS-FLIES 125 Holotype $ of Tasmanthrus galbinomacula- lected by A. Neboiss unless stated otherwise (NMV). 19 42 2 Forth Falls, 28 Jan. 1967, E. F. Riek tus Jacquemart, Cradle Mtn. Tas., 12 Jan. c? (ANIC); 5 o* 11 2 Franklin River, 10 Feb. 1967, 1923, A. Tonnoir, dissected and mounted on E. F. Riek (ANIC); 32 mart, 1965b fig. 15D and E) show numerous 2 1 Family LEPTOCERIDAE Leach presumed errors. The male genitalia is similar (1815) to that of angustipennis, except that the upper branch of the inferior appendage has a distinct Family diagnosis: Ocelli absent. Anterior wings triangular lateral widening, whereas from other long, narrow, venation sometimes irregular in cell always localities it is obtuse, rounded or almost absent. males; discoidal cell closed; median either or wide, Due to this variability it is not considered to be absent. Posterior wings narrow sufficient for erection of a separate species and depending on species; discoidal cell closed in therefore galbinomaculatus is considered as subfamily Triplectidinae, open in subfamily synonym of angustipennis. Leptocerinae; one or two short rows of curved macrotrichia or hamuli at about the middle of slender. Material examined: Tasmania-—26 c? 68 $? Sca- the costal margin. Antennae long and mander River, Upper Scamander, 9 Nov. 1972; 14 $ Maxillary palpi slender, 5-segmented in both 21 $ Derwent River 2 km NW Derwent Bridge, 12 sexes, covered with dense hairs, segment 5 Feb. 1971; 11 KEY FOR SEPARATING vein closing discoidal cell slightly concave; thy- dis- TASMANIAN GENERA ridial cell in males only slightly longer than than in Triplectides. In 1. In anterior wing cross-vein closing discoi- coidal cell, but broader absent. Male genitalia dal cell concave; lower apical angle at posterior wings fork 1 superior ap- cross-vein r-m more or less produced with segment 10 short and broad, downwards towards the thyridial pendages short, dorso-ventrally flattened; in cell 2 female dorsal lobes short. closing dis- — . In anterior wing the cross-vein Spurs 2:2:4. coidal cell straight; lower apical angle at cross-vein r-m not produced down- 127 Westriplectes pedderensis sp. n. wards 3 Figures 683-689 2. Posterior wing — fork 1 Apex of the anterior wings rounded in males, present Triplectides more acute in females; posterior wings broader, fork 1 — . Posterior wing — with wider anal fan in males. absent Westriplectes $ genitalia with dorsal margin of tergite 9 3. Spurs 2:2:2 Symphitoneuria produced into a moderately long, narrow, — . Spurs 2:2:4 4 membraneous projection, which has minute me- 4. Cross-vein r-m in both pairs of wings dian excision at apex. Segment 10 short, broad, either directly, or nearly directly below apex widely and deeply excised. Superior ap- the cross-vein closing discoidal cell; vena- pendages short, oval. Phallus short, partly tion in the male anterior wing membraneous, with sclerotized, lateral, ven- normal 5 trally directed lobes near the middle. Inferior its length branch long, — . Cross-vein r-m at least own appendages four-branched; main basad or cross-vein closing discoidal cell; short bristles on the inner surface; median venation in male anterior wing modi- branch short, flat, curved inward, apical margin fied 6 covered with a row of minute chitinous teeth; 5. In posterior wing fork 1 absent, or if present, then very narrow, indistinct; in- ferior appendages in male genitalia with- out the large, strong slightly curved middle branch Notalina 1 distinct; inferior — . In posterior wing fork appendages with large, curved middle branch Notoperata 6. In anterior wing in males fork 1 with footstalk, thickening of veins below dis- coidal cell short; in females thyridial cell longer than discoidal cell . . Triplectidina with fork 1 sessile, — . Anterior wing in males thickening of veins below discoidal cell long; in females thyridial cell shorter than discoidal cell Lectrides Genus Westriplectes gen. n. Type species Westriplectes pedderensis gen. et sp. n. Anterior wings long, narrow; discoidal cell somewhat similar to that in Triplectides; cross- TASMANIAN CADDIS-FLIES 127 mesal branch short, truncate; lower branch 4. In anterior wing of males, part of lower broad at base. margin of discoidal cell and a short sec- 9 abdomen terminates with acutely pointed, tion of M both thickened forming a short triangular, dorsal projection on segment 9; fold truncatus margin of lower angles acute; dorsal plates short, truncate — . In anterior wing of males, lower apically; lateral plates oval. discoidal cell not so thickened ... 5 Length of anterior wing: $ 14-15 mm; 2 5. Inner marginal plates at the base of in- 10-11-5 mm. ferior appendages broadly Type material; Holotype $ (T5376), allo- triangular elongatus plates at the base of in- type $ (T5377),3 S 6 $ paratypes (T5378- — . Inner marginal T5386) Lake Pedder, Tas., 1 Feb. 1965, A. ferior appendages bilobed apically . 6 Neboiss (NMV). 6. Inferior appendages with basal branch Distribution: Tasmania—SW province. short, broad at base, tapered apically bilobus Genus Triplectides Kolenati basal branch — . Inferior appendages with Triplectides Kolenati, 1859:247; Mosely, 1936b:92; long, cylindrical proximus Mosely and Kimmins, 1953:199 (list of references and synonymy); Fisher, 1965:59. Triplectides ciuskus Mosely Pseudonema McLachlan, 1862:305. 128 Notanatolka McLachlan, 1866:256; Ulmer, 1906:31; Figures 690-693 Ulmer, 1907:130; Mosely, 1936b:93. Triplectides ciuska Mosely in Mosely and Kimmins, gracilis Burmeister, Type species: Mystaeides 1953:209. 1839. Triplectides ciuskus, Fisher, 1972:62. The anterior wings slender, discoidal cell Anterior wings brown to blackish-brown; with lower distal angle more or less produced mesoscutum elongate with two narrow rows of downwards; cross-vein closing discoidal cell setiferous punctures. concave, often less distinctly so in females; Spurs 2:2:2. forks 1 and 5 present in males, forks 1, 3 and S genitalia in dorsal view with distal margin 5 present in females; thyridial cell long and of tergite 9 slightly produced, a small, mem- narrow. Posterior wings with more or less braneous median projection immediately below. widened anal area; fork 1 present with short Superior appendages of medium size, tapering footstalk; fork of M does not reach cross-vein apically. Segment 10 apically truncate with r-m. slight depression in the middle, sides produced Spurs 2:2:2; 2:2:4. downwards into small lateral lobes. Phallus The genus is widely distributed throughout slightly dilated apically, with median excision. the SE Asian and Australian regions, and it is Inferior appendages three-branched, stout at also known from South America. A more de- base; upper branch the longest, rounded apic- tailed study of tropical forms might call for ally; second branch shorter, strongly chitinized, further taxonomic changes. curved inwards, apex acute; third branch arises KEY FOR SEPARATING from the base, reaches only half the length of TASMANIAN SPECIES upper branches, rounded apically. Inner mar- (Males only) ginal plates hooked apically. pairs of 1. Spurs 2:2:2 2 $ abdomen terminates with two plates. — . Spurs 2:2:4 4 2. Mesoscutum with two widely separated Length of anterior wing: $ 9 14-15 mm. Kosciusko, ft., rows of setiferous punctures . . ciuskus Type material: Type 8 Mt 3000 setiferous punctures ar- N.S.W., 20 Jan. 1914 (BMNH); & 9 para- — . Mesoscutum with ranged in two bands 3 types Dunwich, Qld., Sept. 1926, Mackerras 3. Eyes normal in males magnus (ANIC). Type not seen. exceptionally large in —. Eyes Material examined: Tasmania— 12 c? 4 $ Scamander males simOis River, Upper Scamander, 9 Nov. 1972; 1 § Buxton 128 ARTURS NEBOISS This species, originally described from Tas- mania, has since been recorded from numer- ous localities as far away as India, China and Japan. It is distinguished from the other three Tasmanian Triplectides species with spurs 2:2:2, by the normal sized eyes, gradually widening rows of setiferous punctures on the mesoscutum and a pair of double punctures on the scutellum. 6 genitalia of similar plan to that in ciuskus but stouter. The second branch of inferior ap- pendages distinctly bidentate. 9 abdomen with distal margin of end tergite truncate; the lower pair of terminal plates striate on inner surface. Length of anterior wing: $ 15-18 mm; 9 17-18 mm. River, Mayfield, 13 Nov. 1972; 1 J South Esk River, Evandale, 1 Mar. 1967; 3 o* Huon River nr. Scotts Peak, 8 Feb. 1965; 1 8 km SE of Sassafras, 2 Dec. 1974; 1 o* Meander River 3 km N of Westbury, 16 Dec. 1974; 1 ? Lily- dale, creek 2 km N, 16 Dec. 1974; 1 j Wilmot River 10 km S of Forth, 13 Dec. 1974; 1 ? Sorell River 3 km N of Sorell, 8 Dec. 1974; 3 J Dee River 8 km NW of Ouse, 9 Dec. 1974; 1 $ Derwent River 3 km W of New Norfolk, 7 Dec. 1974; 1 £ Dip River Falls, 1 Dec. 1974; 1 129 Triplectides magnus (Walker) Type material: Type $ 'Van Dieman's Figures 694-696 Land. From Dr Hooker's Collection' (BMNH). Type Leptocerus magnus Walker, 1852:73; McLachlan not seen. 1862:307. Notanatolica magna, McLachlan, 1866:257; Ulmer Material examined: Tasmania—3 c? 3 $ Lake Ped- 1907:130. der, 31 Jan. 1965, A. Neboiss (NMV); 1 130 Triplectides similis Mosely Material examined: Tasmania— 1 (J Sundown Creek 25 km S of Marrawah, 1 Huon- Figures 697-698 30 Nov. 1974; o* Pieton River junction. 18 Feb. 1967; 3 ? St. Columba TriplcaiJcs similis Moselv in Moselv and Kimmins, Falls. Pyengana, 21 Feb. 1971; 1 $ Lilydale, creek 2 1953:210. km N. 16 Dec. 1974. All specimens collected bv A. Neboiss (NMV). 2 M S Huon-Picton junction. 17 Anterior wing venation similar to that in Feb. 1967. E. F. Riek (ANIC); 12 ? He'llver Gorge, ciuskus, but lower distal angle not as much 4 Feb. 1967, E. F. Riek (AN1C). produced downward. In posterior wings fork 1 Distribution: Tasmania—NE, N, NW and SW with very short footstalk; fork 5 extended ba- provinces. sally well beyond cross-vein r-m, discoidal cell long; the eyes are very large; mesosternum 131 Triplectides truncatus sp. n. laterally in males slightly constricted near the Figures 699-702 base of the wings; setiferous punctures ar- Anterior wings with discoidal cell short and ranged in two rather broad lines; scutelluni with broad, in males part of the vein forming the setiferous punctures forming a pair of small lower margin of the cell and a short section of warts. M at anastomosis thickened and developed into $ genitalia with upper branches of inferior a short fold. appendages more slender than in magnus, the 3 genitalia of the same pattern as in elonga- second branch only half the length of the upper tus, but distinguished by inner marginal plates and terminates with mesally directed claw. of inferior appendages, truncate at apex and 9 abdomen with end tergite truncate distally, outer apical angles slightly produced. the lower pairs of terminal plates striate on 9 abdomen terminating bluntly, the ventral inner surface. side of dorsal lobes with small projections near Length of anterior wing: $ 17-18 mm; 9 18 mm. the base; the lower pair of plates vertically striate on the inner surface. Triplectides similis xi~ Length of anterior wing: 3 10-12 mm; 9 12 mm. Triplectides truncatus 130 Type material: Type g Deloraine, Tas., 27 Dec. 1884, McLachlan collection (BMNH). TO *0 M Type not seen. 130 ARTURS NEBOISS Type material: Holotype $ (T53 87), allotype Triplectides bilobus ;• 9 (T5388), 16 $ 1 9 paratypes (T5389- .U T5405) Bluff Hill creek 12 km S of Marrawah, Tas., 30 Nov. 1974, A. Neboiss (NMV). jjjL . w t ^^ ^-^_v Other material examined: Tasmania— 1 c? Sundown 1 v Creek 25 km S of Marrawah, 30 Nov. 1974; 2 £ ^y~v s, y'""," s *v Duck River 6 km SW Roger River, 29 Nov. 1974. All T"* *^^s_ <— . ^^"^s W N J-<(- specimens collected by A. Neboiss V (NMV). " V'7 ( 1 \ NW ( ^ Distribution: Tasmania—NW province. pNE \ J / ' s 1 I . r 132 Triplectides bilobus sp. n. "\\^W~*/ Figures 703-706 A r*4 E^ + j>/ . The species in colour and general appearance U '"""^a] /£* "wLV——— i)\ / i is very close to the others in the group of spe- '"^vV-i- \ cies with spurs 2:2:4, but differs in details of the male and female genitalia. 1 is a "v- S genitalia with distal margin of tergite 9 \ SW \j K\ ! rounded, closely below the margin in the middle * "'•* \ r "v- a pair of small, semi-membraneous triangular jC^ protuberances. Superior appendages slender, M rod-like, apices rounded. Segment 10 short, tri- 'sTV*—-A. ' K W t0t.m 132 angular, wide at base, a small incision at apex. P Inferior appendages three-branched, basal part enlarged, upper branch slender, small spines on lected by A. Neboiss unless stated otherwise (NMV). inner apical surface; second branch half the Distribution: Tasmania—C, NW and SW pro- length of upper branch, lower margin near the vinces. apex minutely dentate; basal branch short, broader at base, tapering curved outward and 133 Triplectides proximus sp. n. distally, inner marginal plate bilobed, mesal Figures 707-708 lobe smaller and somewhat pointed, outer lobe larger and rounded apically. Very similar to bilobus, but general colour- 9 abdomen terminates with truncate dorsal ing blackish-brown; otherwise only genitalic plate which has a pair of small, triangular structures separate these two closely related projections at the centre of distal margin. species. Main area of distribution of proximus Length of anterior wing: $ 16-18 mm; 9- 15- lies North and East of that of bilobus. 17 mm. $ genitalia with distal margin of tergite 9 Type material: Holotype (T5406), allotype g rounded, a small, rounded semi-membraneous 5 (T5407), 3 $ 6 9 paratypes (T5408- protuberance in the middle just below the mar- T5416) Franklin River 20 km SW of Derwent gin. Inferior appendages with basal branch Bridge, Tas., 11 Feb. 1971, A. Neboiss long, cylindrical, apices rounded; the inner (NMV). marginal plate bilobed, mesal lobe short, broad, Other material examined: Tasmania—5 6* 1 9 rounded, outer lobe about as wide or narrower. Henty River 12 km NW Queenstown, 10 Feb. 1971; 1 ? Derwent River 2 km NW Derwent Bridge, 12 $ abdomen similar to that in bilobus, but Feb. 1971; 5 <$ 15 $ Huon River Crossing, 16 Feb. dorsal plate with distal margin slightly concave; 1971; 10 The Tasmanian specimens from several lo- calities, including one male from Cradle Moun- tain, were identified as Triplectides dubius as described by Mosely and Kimmins 1953. These were compared with the type of Triplectides elongatus Banks and found that they did not show sufficient differences to regard them as separate taxonomic units and therefore both these forms are now considered conspecific. In the absence of female specimens from Tasmania, the figure and description of this sex is given from the type female collected at Barrington Tops, N.S.W. £ genitalia more slender than in truncatus, the inner marginal plate at the base of inferior appendages triangular. $ abdomen terminates with a pair of stout, dorsal, finger-like projections; below them a pair of curved lobes, concave at lower margin near apex. Length of anterior wings: $ 12-14 mm; $ 14 mm. material: , Type Holotype £ (T5417) Leven River nr. Heka, Tas., 17 Nov. 1972; allotype 2 (T5418) St. Patricks River, Targa, Tas., 22 Feb. 1971; 10 $ paratypes (T5419-T5428) Leven River nr. Heka, Tas., 17 Nov. 1972; 5 3 5? paratypes (T5429-T5438) St. Pat- ricks River, Targa, Tas., 22 Feb. 1971. All specimens collected by A. Neboiss (NMV). Other material examined: Tasmania— 1 <$ Guide River Falls nr. Ridgley, 18 Nov. 1972; 2 134 Triplectides elongatus Banks Figures 709-711 Triplectides elongatus Banks, 1939:486; Fisher, 1965: 62; 1972:62. Triplectides dubius Mosely in Mosely and Kimmins, 1953:219; Fisher, 1972:62. syn, nov. Dark, blackish-brown species. Anterior wings with discoidal cell short and broad; in Type material: Type $ Blackheath, Blue Mtns. males the vein forming lower margin of the 3000 ft., N.S.W., 21 Jan. 1932, Harvard Ex- cell is not thickened and there is no fold on M pedition, Darlington; MCZ Type 22079 at the anastomosis. (ANIC); type $> Barrington Tops, 5000 ft., — 132 ARTURS NEBOISS N.S.W., 9 Feb. 1932, Harvard Expedition, 135 Notoperata sparsa (Kimmins) comb. n. Darlington (ANIC). Both types examined. Figures 712-716 Triplectides dubius Mosely, Type $ Cradle Hudsonema sparsa Kimmins in Mosely and Kimmins, Mtn., Tas., 18 Jan. 1917, R. J. Tillyard 1953:246. (BMNH). Type not seen. Anterior wings dark, blackish-brown, nygma Material round. In posterior wings the distance between examined: Tasmania— 1 Posterior wings with fork 1 present, footstalk longer than the width of discoidal cell. Male genitalia typical of those found in Triplectides, except that the basal branch of the three- branched inferior appendages very small. Spurs 2:2:4, slender. Two species known from mainland Australia and both are also found in Tasmania. KEY FOR SEPARATING SPECIES 1. Superior appendages short, rounded apic- ally; anterior wings less than 10 mm long sparsa — . Superior appendages elongate oval; an- Type material: Type $ Cradle Mtn., Tas., 18 terior wings more than Jan. 1917, R. J. Tillyard (BMNH). Type not 12 mm long maculata seen. TASMANIAN CADDIS-FLIES 133 Material examined: Tasmania—44 1 c? 2 ? same loc, 15 Feb. 1971; 1 c? Huon-Picton River junction, 18 Feb. 1967; 1 Distribution: Tasmania—NW and SW pro- vinces. 136 Notoperata maculata (Mosely) comb. n. Figures 717-720 Hudsonema maculata Mosely in Mosely and Kim- mins, 1953:245. Triplexina lobata Jacquemart, 1965 : 24; Neboiss, 1974c:15; syn. nov. Anterior wings dark brown, nygma distinctly elongate. In posterior wings the distance be- tween cross-vein r-m and form of M very short, shorter than the length of the cross-vein. Material examined; Tasmania— 1 ? National Park, $ genitalia similar to sparsa, but somewhat 1 20 Feb. 1967; 1 J Derwent River 2 km NW Derwent more elongate; superior appendages elongate Bridge, 12 Feb. 1971; 9^4? Waldheim, Cradle Mtn. Nat. Park, Feb. 10 Huon-Picton oval. The basal branches of inferior appendages 1971; <$ River junc- tion, 18 Feb. 1967; 1 °. Huon Crossing, Port Davey short, finger-like; inner marginal plates hooked Track, 8 Feb. 1965; 1 c? 1 ? Huon Plains nr. Scotts and somewhat truncate apically. Peak, 2 Feb. 1965; 1 c? 2 2 same loc, 8 Feb. 1965; 1 o* Condominion Creek nr. Mt Eliza, 10 Feb. 1965; 2 genitalia with tergite distally produced 9 12 c? 3 2 same loc, 15 Feb. 1971; 2 2 Henty River 12 into a more or less distinct, bipointed median km NW Queenstown, 10 Feb. 1971; 4 <$ 8 2 Wedge River, 17 Feb. 1971; 12 134 ARTURS NEBOISS Posterior wings similar in both sexes, wider than the anterior wings; fork 1 absent; fork of M reaches basad beyond the cross-vein below the discoidal cell. Male genitalia pattern close to that found in the genus Triplectides. Spurs 2:2:2. Only one species recorded from Tasmania. 137 Symphitoneuria opposita (Walker) Figures 721-723 Leptocerus oppositus Walker, 1852:73. Notanatolica opposita, McLachlan, 1866:73; Ulmer, 1906:31; 1907:131. Loticana opposita, Betten and Mosely, 1940: 13 1 Ulmer, 1951:404. Symphitoneuria opposita, Mosely and Kimmins, 1953: 264 (also further references). This dark-greyish species, although being one of the first described from Tasmania, was not found to be at all common during this sur- vey. Early records from Victoria are substanti- ated by new material from several localities, but no new material is available from Western Australia. ? Canal at Interlaken, 2 Feb. 1966, G. F. Edmunds $ genitalia with tergite 9 broadly triangular. (ANIC). Victoria— 1 <$ Tyres River, 26 Oct. 1973; 1 Superior appendages moderately long, gradu- S Tanjil River, Walhalla Rd bridge, 22 Oct. 1973; 1 c? ally dilated apically. Segment 10 long and Latrobe River, Yallourn, No 3 lake, 24 Oct. 1973; all collected by broad, apex deeply and narrowly excised; la- C. McCubbin during Latrobe River sur- vey (NMV). teral margins with small, triangular projections Distribution: Tasmania C, E and SE provin- about midway, where there is also a transversal — ces; Victoria; Western Australia; Celebes. chitinous ridge across the segment. Phallus short, with a short, truncate plate beneath. In- Genus Triplectidina ferior appendages three-branched; the upper Mosely branch slightly shorter than the second, which Triplectidina Mosely, 1936b: 107; Mosely and Kim- mins, 1953:233. terminates with claw-shaped apex; a small, an- Type species: gular, inward and downward directed mesal Triplectides oreolimnetes Till- yard, 1924. plate, bearing several strong spines, arises from the inner margin of the upper branch. From Anterior wing venation differing in sexes; in the base of inferior appendages arise inwardly males, part of lower margin of discoidal cell directed, subangular plates and ventrally a and part of M at the same distance thickened; moderately long third branch. thyridial cell absent; in females venation nor- 9 abdomen terminates bluntly with a pair mal; distal end of discoidal cell only a short dis- of dorsal and a pair of lateral plates. tance distad of anastomosis, cross-vein closing the Length of anterior wing: $ 11-12 mm; 9 cell straight; thyridial cell present. Posterior 11 mm. wings with fork 1 present, although often rather material: indistinct Type Type $ 'Tasmania', no other and sometimes absent. R 2 very short, data forming almost right (BMNH). Type not seen. angle with R3 . Material examined: Tasmania— 1 <$ Andover York Spurs 2:2:4. Rivulet, 4 Dec. 1974, A. Neboiss (NMV); 1 138 Triplectidina nigricornis Mosely Material examined: Tasmania—8 o* 2 ? Waldheim, Cradle Mtn. Nat. Park, 7 Feb. 1971; 1 S Huon River Figures 724-726 Crossing, 16 Feb. 1971; 1 o* 1 ? Huon-Picton River junction, 18 Feb. 1967; 1 1 River nr. Triplectidina nigricornis Mosely, 1936b: 109; Mosely o* ? Huon 1 and Kimmins, 1953:235. Blakes Opening, 9 Feb. 1966; 1 J Wedge River, 17 Feb. 1971; 1 J Franklin River 20 km SW of Derwent Anterior wings dark brown with greyish and Bridge, 11 Feb. 1971; 2 ? Derwent River 2 km NW fuscous pubescence, venation as described in Derwent Bridge, 12 Feb. 1971; 1 o* Fossey River 8 km E of Waratah, 12 Dec. 1974. All specimens col- generic diagnosis. lected by A. Neboiss (NMV). 1 o* Huon-Picton junc- $ genitalia with distal margin of tergite 9 tion, 17 Feb. 1967, E. F. Riek (ANIC). developed into a short, triangular projection. Distribution: Tasmania—C, NW and SW pro- Superior appendages short, situated close to- vinces. gether, dorso-ventrally flattened. Segment 10 long, narrowly and deeply cleft at apex, distal Genus Lectrides Mosely half of lateral margins covered with short, stiff Lectrides Mosely in Mosely and Kimmins, 1953:270. bristles. Phallus short. Inferior appendages Type species: Lectrides varians Mosely, 1953. three-branched, the main upper and outer Anterior wing venation differing in sexes; in branch the longest, apex with few short, dark males a long fold in the centre of the wing, peg-like spines inner on surface; the inner merging the lower margin of discoidal cell with branch slender, curved mesally; lower branch M; venation normal in females, discoidal cell shorter, finger-like. long, extending distad of cross-vein r-m; nar- 9 abdomen with terminal segment dorsally row hyaline line above thyridial cell widened slightly produced at its centre. at the cross-veins; thyridial cell shorter than Length of anterior wing: $ 9-5-12 mm; $ discoidal cell. Posterior wings similar in both 10-12-5 mm. sexes, fork 1 absent; fork of the media usually does not reach the cross-vein below the discoi- dal cell. The male genitalia diverge considerably from the normal Triplectides pattern; inferior ap- pendages two-branched. Spurs 2:2:4. So far only one species has been included in this genus. 139 Lectrides varians Mosely Figures 727-730 Lectrides varians Mosely in Mosely and Kimmins, 1953:270. Anterior wings brown with indistinct dark area near the base and a pair of dark patches at pterostigma. The females are similar in appearance and size to the males; posterior wings wider than the anterior pair, but without distinctly ex- panded anal fan; a short row of hamuli along costa extend from about the middle to distal end of discoidal cell. S genitalia with distal margin of tergite 9 Type material: Type $ and para-types Miena, broad, bluntly triangular, below which is a Tas., 3300 ft, Dec. 1929, H. M. Stephen, in forked process, considered by Mosely and Kim- Mosely's collection (BMNH). Type not seen. mins as the superior appendages fused at the 136 ARTURS NEBOISS base. Segment 10 broad at base, tapering apic- Distribution: Tasmania- -N, NW, SW and SE ally, excised in the middle. Phallus slender, ter- provinces; Victoria. minating with a pair of rounded, lateral lobes, separated by deep mesal cleft. Inferior appen- Genus Notalina Mosely dages two-branched, the upper branches slightly Notalina Mosely, 1936b: 114; Mosely and Kimmins, 1953:248. shorter, the lower ones longer and curved in- Triplexina Mosely in Mosely and Kimmins, 1953:231 wards apically. syn. nov. 9 abdomen terminates with broadly triangu- Type species: Notalina parked Mosely, 1936. lar tergite; dorsal plate short, lateral plates The genus was erected by Mosely (1936) to slightly longer and apically truncate if viewed include three species, Notalina parked from from the side. Tasmania, which was designated as the type Length of anterior wing: 10*5-12 S 5 mm; 9 species, and two other species from Southwest 10-12 mm. Australia, described by Ulmer in the genus Triplectides. The main distinguishing character was the absence of fork 1 in the posterior wings. Further species were added in 1953, all of which shared distinctive male genitalia. The discovery of a Tasmanian species, in which the male genitalia and wing venation is similar to that in other Notalina species (except that in posterior wings a very narrow fork 1 is usually present), required widening of the generic diagnosis. Analysis of characters found in the genus Triplexina, with its sole species nigra, clearly indicates the close relationship with No- talina. In longer series of specimens of nigra, it was observed that the fork 1 in the posterior wing sometimes is indistinct or occasionally absent. The male genitalia is typical of that found in Notalina. Therefore genus Triplexina is now placed in synonymy with Notalina, but the species nigra retains its specific status. Anterior wings in male with discoidal and thyridial cells of approximately equal length, but the thyridial cell is longer in females; Rs slightly curved towards M between the base and Type material: Type $ Sheffield, Tas., 8 Jan. discoidal cell. Posterior wings with discoidal 1917, R. J. Tillyard (BMNH). Type not seen. cell rather long and slender, closed distally Material examined: Tasmania 3 cf 1 9 Huon-Pic- — by more or less oblique cross-vein; fork 1 ton River junction, 18 Feb. 1967, A. Neboiss (NMV); 1 absent, or if present, very narrow. ? same loc, 15 Nov. 1972, A. Neboiss (NMV); 2 o* River, Wedge 17 Feb. 1971, A. Neboiss (NMV); 1 o* Mesoscutum elongate, dorsally flattened, an- Bushy Park, 23 Feb. 1967, A. Neboiss (NMV); 2 o* terior margin somewhat produced anteriorly; Murchison River, 5 Feb. 1967, E. F. Riek (ANIC). Victoria—8 J1 Myrtleford, 23 Jan. 1973, A. Ne- scutellum as a small sloping plate, raised boiss (NMV); 1 $ Koornalla, Traralgon Creek, 10 slightly above mesoscutum. Dec. 1973, C. McCubbin (NMV); 1 2. Segment 10 in lateral view obliquely trun- Notalina parkeri cate, apex directed upwards .... nigra — . Segment 10 in lateral view tapered to rounded apex parkeri 3. Lower margin of inferior appendages with angular or rounded sub-basal lobe, carry- ing a small, acute spine 4 NW ^V \ { ] \ — . Lower margin of inferior appendages with two acute spurs bifaria 4. Inferior appendages in lateral view with sub-basal lobe appearing as a curved hook fulva — . Inferior appendages in lateral view with only a minute spur on lower Vse\ J^ .."' margin tillyardi KM 140 Notalirta parkeri Mosely : hs j £_ \Z^*) N ** i^-^&' ' Figures 731-736 Notalina parkeri Mosely, 1936b: 1 14; Mosely and Kimmins, 1953:249. - C » m Mkm '.»• ?N ^ 140 General colour black; anterior wings in males with thyridial cell long dis- about as as Type material: Type £ Great Lake, Miena, coidal cell, females the thyridial cell is dis- in Tas., Jan. 1930, C. Parker (BMNH). Type not tinctly longer. In posterior wings discoidal cell seen. long, closed with oblique cross-vein; fork 1 Material examined: Tasmania—3 6* 1 ? Miena, absent. 3400 ft, 24 Jan. 1961, L. Couchman (NMV); 10 d* 8? Shanon Lagoon, 27 Feb. 1967, A. Neboiss (NMV); 12 $ genitalia short and broad; distal margin of c? 4 ? Penstock Lagoon, 27 Feb. 1967, A. Neboiss small pro- tergite 9 excised in the middle, two (NMV); 1 J Ouse River 8 km W of Miena, 28 Feb. tuberances on either side of the excision. Su- 1967, A. Neboiss (NMV); 8 oM ? Bradys Lake, 9 Dec, 1974, A. Neboiss (NMV); 1 o* Interlaken, Lake perior appendages spindle shaped, slightly Sorell, 5 Dec. 1974, A. Neboiss (NMV); 21 9 abdomen terminates with a pair of tri- Figures 737-7'40 are angular dorsal plates, at the base of which Notalina fulva Kimmins in Mosely and Kimmins, a pair of small protuberances; lateral plates 1953:257. large subquadrate. Anterior wings yellowish to brownish, the Length of anterior wing: $ 11-13 mm; 9 fringe fuscous at arculus; posterior wings pale, 9-11 mm. yellowish, fork 1 absent. Wing venation and . 138 ARTURS NEBOISS genitalia similar to N. parked. The species is Distribution: Tasmania—all provinces; Vic- widely distributed and has been recorded from toria; New South Wales. New South Wales, Victoria and Tasmania. 142 Notalina tillyardi Kimmins $ genitalia, compared with that of parked, has longer and more slender superior append- Notalina tillyardi Kimmins in Mosely and Kimmins, 1953:258. ages, the inner margin of sub-basal lobe more Although a large number of specimens of rounded and the branch at the base in ventral the genus Notalina were examined view is somewhat irregular in outline. from nu- merous Tasmanian localities, none $ genitalia very similar but separated from appeared to match the figures given parked by the yellowish colour. by Mosely and Kim- mins (Fig. 179, p. 259). The description notes Length of anterior wing: 6 9-16 mm; $ 7-11 mm. that it can be distinguished from julva by paler colouring and that the male genitalia in lateral view shows only a small spur on the ventral margin of inferior appendages. The female, although listed among the orig- inal material, is neither described nor figured. Length of anterior wing: s 12-13 mm; $ 10-11 mm. Type material: Type $ Gisborne, Victoria, 7 Nov. 1915 (BMNH). Type not seen. Material examined: Tasmania—4 c? Sorell River 3 km N of Sorell, 8 Dec. 1974; 4 lar spots on wing membrane; anal margin from SW of Derwent Bridge, 11 Feb. 1971; 2 rf Grt. Forester River 5 km Forester, 11 Nov. 1972. All base to arculus and almost as far as Cu vein NW specimens collected by A. Neboiss (NMV). pale yellow; fringe 1 distally from arculus fuscous. 2 d Forth Falls, 28 Jan. 1967, E. F. Riek (ANIC). Posterior wings pale-yellowish, fork 1 usually Distribution: Tasmania—C, N, NW, SW and present, although sometimes indistinct or en- SE provinces. tirely absent. $ genitalia of the same pattern as in other 144 Notalina nigra (Mosely) comb. n. species of the genus, but lower margin of in- Figures 744-746 ferior appendages sub-basally with a pair of strong, inwardly directed spurs, welded together Triplexina nigra Mosely in Mosely and Kimmins, 1953:232. at their base. The reason for the present placing of this 2 unknown. species are explained in the description of the Length of anterior wing: S 12-13 mm. genus. Anterior wings covered with dense, black Notalina bifaria AM pubescence; head, thorax and abdomen also black. Anterior wings in males with thyridial II cell slightly longer, in females distinctly longer than discoidal cell. Posterior wings with fork 1 although sometimes indistinct A ^A7 usually present, '^Cn r or missing. ,S C r ' \ ;\ NW^ V \ J $ genitalia with segment 9 narrow dorsally, A J / VfLj) lateral margins broadly rounded. Superior ap- \ '1 t ^ pendages short and stout, tapered distally. Seg- A^-^TiT^ ment 10 broad at base, bifurcate distally, api- + - Phallus short, - truncate. oT Vj \ V f^ $ I ces obliquely mem- L A% 5 $ braneous, curved downwards, on either side with down-turned, acutely pointed parameres. Inferior appendages stout, small acute mesal processes on lower margin towards the base; " V \ SW\* A, inwardly directed branch at the base; upper mesal margin developed into a plate with curved, distally directed hooks. 9 abdomen ends with a pair of small dorsal processes and a pair of rounded lateral lobes. M «0 Wltm ":"*' "\-/ 143 Length of anterior wing: 8 7-5-9 mm; 9 7-8 mm. Type material: Holotype $ (T5439), 5 S Type material: Type (NMV); 6 c? 4 ? Little Pine Lagoon, 27 Feb. 1960, Other materia! examined: Tasmania—3 <$ Huon D. Scholes (ANIC); 1 ? Navarre River, 12 Feb. Scotts Peak, 8 Feb. 1965; 4 140 ARTURS NEBOISS coidal cell closed with sloping cross-vein; forks 1 and 5 present in male, forks 1, 3 and 5 present in female; thyridial cell about twice as long as discoidal cell; apical cellule 4 with foot- stalk. Posterior wing broader than the anterior wing, discoidal cell open, forks 1, 3 and 5 present in both sexes. Spurs 2:2:4. The combination of characters separate Con- docerus from the cosmopolitan genus Athrip- sodes and members of the Australian mainland Leptocerus. 145 Condocerus paludosus sp. n. Figures 747-755 Anterior wings densely covered with fine pubescence which is darker along costal and anal margins, thus showing a pale, longitudinal central band which disappears in rubbed speci- mens. The first two pairs of legs covered with short, dark pubescence and appear much darker than the posterior pair. Subfamily LEPTOCERINAE Ulmer $ genitalia—distal margin of segment 9 with a pair of small, elongate protuberances in the The subfamily is distinguished by the open middle of the base of segment 10 and larger, discoidal cell in the posterior wings; other apically tapered one on either side. Segment characters as described for the family. 10 downturned, gradually tapered towards the KEY EOR SEPARATING upcurved apex. Phallus slightly clavate; in- TASMANIAN GENERA ferior appendages single segmented, robust, Subfamily LEPTOCERINAE Ulmer curved inwards, with short, broad median, in- 1 In anterior wing thyridial wardly-directed lobe. cell absent Triaenodes 9 abdomen terminates bluntly with a down- — . In anterior wing thyridial curved, oval, dorsal plate, at the base of which cell present 2 is a pair of small, flat protuberances and 2. Spurs 2:2:4 3 further laterally a large, horizontally situated, Spurs — . 0:2:2, 1:2:2 or 2:2:2 .... Oecetis rounded lobe, below that a vertically orien- 3. Posterior wing narrow; fork 3 tated, lateral lobe. absent Leptorussa Length of anterior wing: $ 10-11 mm; $ 9 5- — . Posterior wing with broad anal fan, fork 3 11 mm. present Condocerus Type material: Holotype <$ (T5450), allotype 9 (T5451) Genus Condocerus gen. n. 6 £ 6 $ paratypes (T5452- T5463) Condominion Creek, Tas., 15 Feb. Type species: Condocerus paludosus gen. et 1971, A. Neboiss (NMV);2 ^ 3 5 paratypes sp. n. (T5464-T5468) Lake Pedder, Tas., 1 Feb. Antennae at least twice as long as anterior 1965, A. Neboiss (NMV). wing, basal segment enlarged, about as long as Other material examined: Tasmania 1 Hellyer the diameter of the eye, segment 2 very short, — <$ River Gorge, 9 Feb. 1971; 3 ? Wedge River, 17 Feb. rounded, segment 3 long slender. $> and Maxillary 1971; 4 Huon River Crossing, 16 Feb. 1971; 3 ? St, Patricks River, Targa, palpi long, segment 2 the longest, about twice as 22 Feb. 1971; 1 $ Huon- Picton River junction, 18 Feb. 1967; 1 2 West Arthur long as segment 1 . Anterior wings slender, dis- Plains, 6 Feb. 1965. All specimens collected by A. TASMANIAN CADDIS-FLIES 141 146 Leptorussa darlingtoni (Banks) Condocerus paludosus m Figures 756-758 Leptocerus darlingtoni Banks, 1939:487. ^ Leptorussa darlingtoni, Mosely and Kimmins, 1953: n 275. Leptorussa russata, Mosely in Mosely and Kimmins, 1953:274; Jacquemart, 1965b:21. syn. nov. J-<( A slender, blackish-brown species. The type ) i V \ nw 7^ "- v I \ \ ^- M 1 *7 y of darlingtoni agrees well with the illustrations J NE - f \ / \6o ,^-oMo of russata in Mosely and Kimmins (1953) and both are regarded as being conspecific. 3 genitalia with upper distal margin of seg- ment 9 produced into a pair of long, down- curved, apically clavate processes; segment 10 in form of a broad, deeply cleft plate, each branch AH /^ mTse \ apically tapered and terminating with outwardly directed triangular apex. Phallus appears as a C narrow central spine. Inferior appendages with |-^\ i. +- (^ vj \_/ i n ^f r ^»- ^V" inner upper angle extended into a ventro- laterally curved process. $ abdomen terminates in a bluntly triangu- lar apex, formed a large, hood-shaped upper M «0 flOkm • \^v^ 145 by 1 plate. Length of anterior wing: $ 8-9 mm; 9 8-9 Neboiss (NMV). 1 ? 10 mis E of Strahan 20 Feb. mm. 1963, I. F. B. Common and M. S. Upton (ANIC). Victoria—3 <$ 2 $ Merrijig, 25 Mar. 1958, A. Ne- boiss (NMV); 6 Genus Leptorussa Mosely Leptorussa Mosely in Mosely and Kimmins, 1953: 272. Type species: Leptorussa russata Mosely. Antennae long and slender, from about one and a half times the length of anterior wing in female, to more than twice the length in male; basal segment enlarged, bulbous, segment 2 short, rounded; segment 3 long, slender. An- terior wing with forks 1 and 5 in male, 1, 3 and 5 in female; discoidal cell closed, thyridial cell almost twice the length of discoidal cell. Pos- terior wing narrow, forks 1 and 5 present. Spurs 2:2:4. The genus is close to the cosmopolitan genus Athripsodes, but differs from it by having a narrowed posterior wing, different structure of male genitalia and different spur formula. Type material: Holotype s Aldgate, Mt Lofty Only one species recorded from Tasmania. Range, S.A., 29 Nov. 1931, Harvard Aust. Ex- 142 ARTURS NEBOISS ped. P. J. Darlington (MCZ Type 22081), 9 abdomen terminates with short, triangular paratypc $ same date. The type specimens are dorsal plate and a pair of large, rounded lateral now deposited in ANIC Canberra. Type seen. lobes. Type of $ Leptorussa russata Mosely, Lee's Length of anterior wing; 3 7-8 mm; 9- 7-5-8 Spring, ACT., 1932, R. J. Tillyard (BMNH). mm. Type not seen. Material °_ examined: Tasmania—6 J 1 9 Grt. Forester River 5 km NW Forester, II Nov. 1972, A. Neboiss (NMV); 1 rf Russell Falls, National Park. 5 Dec. 1972, P. Zwick (NMV). Other localities recorded: Tasmania: Deloraine, Hobart, Orford, St. Patricks River. Distribution; Tasmania—N, SE and E pro- vinces; Victoria; New South Wales. Genus Triaenodes McLachlan Triaena McLachlan, 1865a:34 not Triaena Leconte, 1847:365. Triaenodes McLachlan, 1865b: 110; 1877:319; Ulmer, 1907:140; Ross, 1944:244; Mosely and Rimmins, 1953:276. Type species: Leptocerus hicolor Curtis. Antennae about twice as long as the an- terior wing, basal segment longer than the diameter of eye, stout; segment 2 short, rounded; segment 3 and the subsequent ones long, slender. Maxillary palpi long, segments slender, thickly covered with long pubescence. Anterior wings long, narrow, apical forks 1 and 5 present, discoidal cell long, thyridial cell ab- sent; posterior wing narrow, fork 1 only present. Spurs 1:2:2. Type material: Holotype g (T5469), allotype 2 (T5470), 5 S 10 $ paratypes (T5471- 147 Triaenodes intricate sp. n. T5485) South Esk River, Evandale, Tas., 1 Figures 759-763 Mar. 1967, A. Neboiss (NMV); 1 <$ 1 $ A slender, dark-brown species. In male, the paratypes (T5486-T5487) Grt. Forester River upper surface of basal antennal segment v/ith a 5 km NW Forester, Tas., 11 Nov. 1972, A. long, flat, somewhat spatulate, scale-like pro- Neboiss (NMV); 1 s paratype (T5488) cess, near the base of which arises a group of Huon-Picton River junction, Tas., 18 Feb. long hairs; in female, the basal antennal seg- 1967, A. Neboiss (NMV); 1 $ 5 9 paratypes, ment is simple but of similar shape and length. Evandale, Tas., 1 Mar. 1967, E. F. Riek $ genitalia with upper margin of segment 9 (ANIC). produced into a pair of small, flattened Oilier material pro- examined: Tasmania— 1 ? Scaman- cesses and a long central rod-like projection; der River, Upper Scamander, 9 Nov. 1972, A. Ne- boiss (NMV); 3 ? Derwent River 2 km Derwent segment 10 formed by a pair of NW asymmetric, Bridge, 12 Feb. 1971, A. Neboiss (NMV); 4^1? gradually tapered processes, the one on right Tooms Lake, 4 Dec. 1974, A. Neboiss (NMV). side curved, tusk-like. Phallus narrow at base, Distribution: Tasmania—SW, C, N and E widened distally. Inferior appendages complex provinces. with a pair of long, slender, curved processes Genus Oecetis arising from the base; the distal portion with McLachlan broad, spiny, upper lobe and a curved, pointed, Oeeetis McLachlan, 1877:329; Mosely and Kimmins 1953:281; Fisher, 1966:106 latero-distal angle. (list of references and synonymies). — TASMANIAN CADDIS-FLIES 143 Type species: Leptocerus ochraceus Curtis, margin divided, mid-ventral angle pro- 1825. duced into a slender finger-like Antennae long, slender, slightly longer than process australis the anterior wings in females, to more than — . Inferior appendages with upper margin twice their length in males of some species; forming rounded lobe; lower margin segment 1 about as long or slightly longer than not divided 3 the diameter of eye; segment 2 short, segment 3. Central area of anterior wing with patch of 3 and the subsequent ones long and slender. scale-like hairs 4 Anterior wings long and narrow; Sc connected — . Central area of anterior wing without to Ri by thickened cross-vein; thickening of R x patch of scale-like hairs gilva from cross-vein gradually decreasing towards the wing margin; discoidal cell closed; forks 1 4. Inferior appendages in lateral view with and 5 present. Posterior wings narrow, forks 1 lower margin obtuse; parameres and 5 present. long pechana Spurs0:2:2, 1:2:2 or 2:2:2. — . Inferior appendages in lateral view with distally; parameres This world-wide genus in Australia is re- lower margin produced presented by two very distinct species groups short umbra the minasata group which has the fork 1 in 5. Cross-vein r-m at approximately the same anterior wing with footstalk and the australis level as the cross-vein closing discoidal group, with fork 1 sessile in anterior wing. To cell 6 the australis group belong the Australian spe- — . Cross-vein about its own length distad cies australis (Banks), pechana (Mosely), from cross-vein closing discoidal species gilva lurida (Kimmins) and two new cell arcada and umbra, described in this paper; also the New Zealand species unicolor (McLachlan), 6. Spurs 0:2:2 7 iti (McFarlane), chatamensis (Tillyard) and —. Spurs 1:2:2 or 2:2:2 8 the Papuan species squamosa (Kimmins). 7. In male, segment 10 about twice as long A number of generic names have been pro- as superior appendage asmanista posed for various groups of species, but they — . In male, segment 10 about as long or have all been placed into synonymy with Oece- only slightly longer than superior append- tis. attempt is made here to relate Tas- No . . age . . minasata manian species to any other than those on the 8. Spurs 1:2:2 9 Australian mainland; as there is a number of . Spurs 2:2:2 scirpicula undescribed species, other grouping might later — be recognized. 9. Anterior wing mottled; cross-vein r-m and are The spurs on anterior legs sometimes m-cu at the same level laustra very small. The differences between chatamen- — . Anterior wing with dark, longitudinal in the sis, unicolor and umbra is found mainly lines; cross-vein m-cu basad of cross-vein shape of the inferior appendages. r-m inscripta KEY FOR SEPARATING TASMANIAN SPECIES 148 Oecetis pechana Mosely (Males only) Figures 764-769 1. Anterior wing with fork 1 Oecetis pechana Mosely in Mosely and Kimmins, sessile {australis group) 2 1953:302. 1 with — . Anterior wing — fork Anterior wings brown, with dark brown spots footstalk {minasata group) 5 at vein junctions, the central area of the wing 2. Inferior appendages with upper margin in males covered with brown scales. Posterior produced into a distinct process; lower wings narrow, apically acute. Spurs 1:2:2. 144 ARTURS NEBOISS * genitalia with segment 9 slightly pro- 149 Oecetis umbra sp. n. duced mid-dorsally, distal margin developed Figures 770-773 into a pair of rounded lobes. Segment 10 in shape of elongate, membraneous plate, triangu- Anterior wings greyish-brown, small groups lar apically with a short, vertical ridge. Phallus of dark hairs on vein junctions, but less dis- membraneous, parameres very long, slightly tinct than those in pechana. In male the cen- curved, pointed apically. Inferior appendage tral area of anterior wing just below discoidal short and broad, upper lobe rounded and cell covered with patch of dark scales. Spurs curved inward. 1:2:2. 9 abdomen terminates bluntly, lateral lobes S genitalia very similar to unicolor and rounded apically; central tubular piece with chatamensis, but differs by having a more pro- dorsal margin apically excised. nounced, longer, finger-like process on the inner, upper margin of the inferior appendage. Length of anterior wing: $ 9-10 5 mm; $ 8-11 mm. Phallus short, apex truncate with downturned ventral ridge; parameres asymmetric, short, acute apically. $ genitalia similar to that of pechana but upper margin of tubular piece not distinctly ex- cised. Length of anterior wing: $ 12 mm; $ 10 5 mm. Type material: Type S Yanchep, W.A., 20-31 Dec. 1935, R. E. Turner (BMNH). Type not seen. Material examined: Tasmania—2 c? 3 $ South Esk River, Evandale, 1 Mar. 1967; 1 TASMANIAN CADDIS-FLIES 145 150 Oecetis gilva sp. n. Distribution: Tasmania—N province. Figures 774-776 151 Oecetis australis (Banks) The species is of overall light, reddish-brown Figures 777-783 colour. Anterior wings rounded apically; both Oecetina australis Banks, 1920:350. cross-veins below discoidal cell are equally far Oecetis australis, Mosely and Kimmins, 1953:305. basad from the cross-vein closing discoidal cell. The anterior wing pattern is similar to that The wing membrane around vein junctions and in pechana, but differs in some minute details, cross-veins pigmented dark; in males there are overall paler colour and absence of scale-like no scale-like hairs at the centre of anterior hairs in the male anterior wings. Tasmanian wings. Spurs 1:2:2. specimens, compared with the holotype & of £ genitalia with middle of dorsal margin of australis, show some difference in the shape of segment 9 broadly rounded, a pair of small the upper lobe of the inferior appendages. protuberances at the base just above dorsal Variations of this structure were observed also lobes. Segment 10 in form of gradually tapering among the Victorian specimens. The darkened plate. Phallus short, lower margin produced area at the base of the discoidal cell is flanked into a downward directed acute projection, by pale, elongate spots basad and distal. parameres short and pointed. Inferior append- S genitalia with gradually tapering segment ages with upper margin broadly rounded. 10 at and above the base of which is a pair of 9 genitalia similar to that of pechana but oval lobes. Superior appendages dorso-ven- central tubular piece less protruding distally. trally flattened, rather long, in dorsal view Length of anterior wing: s 10-10-5 mm; $ apically clavate. Phallus membraneous, para- 9 5 mm. mere long, slender, apically bent at right angle. Inferior appendages ventrally separated by a Oecetis gilva V-shaped incision, central margin extended 'Mi into a long, narrow projection, but lateral margin extended into two lobes, variable in H shape. Oecetis australis ;y •- "\ N F I ' \ ** f V \ fl j] % ^"\ 1 + $ t .4 NW (A ~ Jtk^^^ 1 r ' ^ in \fr 4J-< 4 • |p*' VSE \ + \ ( C\ iff &?\ m ^ \^\ + &l :.\sw \. "\ v^ \ „ _,.** * ^\ r *&' \ Kypy J\ ^. ip >p- ''•'**' ' A - sw M *0 »0*r> . > 150 t ^Y a Type material: Holotype $ (T5491), allotype Uvhfi 9 (T5492), 7 £ 1 $ paratypes (T5493- T5500) South Esk River nr. Evandale, Tas., 1 » «0 50 k... ^4P 151 Mar. 1967, A. Neboiss (NMV). 146 ARTURS NEBOJSS 9 abdomen terminates bluntly, lateral plate in side view with lower angle produced to a rounded lobe. Length of anterior wing: £ 7-8 mm; 9 7- 8-5 mm. Type material: Holotype $ New Holland, Mel- bourne (Victoria), Thorey (ANIC). Type seen. Material examined: Tasmania—] c? St. Columba Falls, Pyengana, 21 Feb. 1971; 1 ? National Park, 20 Feb. 1967; 2 $ Huon River Crossing 16 Feb. 1971; 1 South ? Esk River, Evandale, 1 Mar. 1967; 1 $ Great Forester River 5 km NW Forester, 11 Nov 1972; 1 c? Bradys Lake, 27 Feb. 1967; 1 $ St. Patricks River, Targa, 22 Feb. 1971; 1 c? Wedge River, 17 Feb. 1971; 3 9 Hellyer River Gorge, 9 Feb. 1971. All specimens collected by A. Neboiss (NMV). Distribution: Tasmania—NE, N, C, NW, SW and SE provinces; Victoria. 152 Oecetis minasata Mosely Figures 784-787 Oecetis minasata Mosely in Mosely and Kimmins, 1953:282. Anterior wings brownish, slightly darker, ob- 153 Oecetis laustra Mosely liquely transversal line in the middle of wing at anastomosis, thyridial cell slightly longer than Figures 788-791 discoidal cell. Spurs 0:2:2. Oecetis laustra Mosely in Mosely and Kimmins, 1953: 295. & genitalia with segment 10 about as long Anterior wings brownish with some mottling. or only slightly longer than superior append- The specimens from Tasmania differ slightly ages. Phallus directed downward, apex rounded. from those of Western Australia, but are still Inferior appendages broad at base, upper mar- regarded as belonging to the same species. gin extended upward and terminating with in- Spurs 1:2:2. ward directed hook. $ genitalia with distal margin of tergite 9 9 abdomen in lateral view with triangular produced in the middle. Superior appendages projection at the base of lateral plate, which short, broadly triangular in lateral view. Seg- has the lower margin produced downward into ment 10 membraneous, transparent, apex ex- a rounded lobe. cised. Phallus stout, apical section abruptly Length of anterior wing: $ 6 5-7-5 mm; 9 bent downward. Inferior appendages broad, up- 7 mm. per apical angle extended to a rounded, up- Type material: Type $ Tasmania', J. W. Evans wardly and inwardly curved lobe; the inner, (BMNH). Type not seen. lower margin smooth with a curved tooth on Material examined: Tasmania — 1 ;: the Oecetis laustra detected in the drawings of wing venation; ;Uv lateral view of genitalia is shown with ventral side up. The differences between Jacquemart's B species and asmanista are not substantiated and both species are regarded as synonymous. cf genitalia characterized by long and narrow ^N segment is almost the length V r^<( 10, which twice NW '~V ]&' \ { Y\ \ p^r ' of superior appendages. Inferior appendages t / •^ \ / ) broad at base, distal section abruptly narrowed, turned inwards at apex and terminating with \^Tfp^ short few spines. *• ' ^ lateral plates somewhat \ <^v I ^ $ abdomen with ''"""^ rectangular, lower angle slightly produced t£\ -•—— >3l ( J ) r J ^ ^ t? -L-yS ^ ]M downward. 20 « SO Km V -/ 153 Type material: Type a Yanchep, 32 mis N of Perth, W.A., 13-23 Nov. 1935, R. E. Turner (BMNH). Type not seen. River nr. Evandale, 1 Mar. 1967, A. Neboiss (NMV); 3 <$ same loc, 1 Mar. 1967, E. F. Riek (ANIC). Material examined: Tasmania—3 154 Oecetis asmanista Mosely Figures 792-795 Oecetis asmanista Mosely in Mosely and Kimmins, 1953:282. Oecetis ochracea Jacquemart, 1965b:23 non Curtis, 1825. Oecetis geevestonia Neboiss, 1974c: 15 syn. nov. This species is similar to minasata but slightly larger in size; anterior wings brownish with Type material: Type $ Tasmania, J. W. Evans darker oblique line at anastomosis, thyridial cell (without definite locality) (BMNH). Type not about the same length as discoidal cell. Spurs 0:2:2. seen. Holotype of Oecetis ochracea Jacquemart, The species described by Jacquemart s Geeveston, Tas., 7 Dec. 1922, A. Tonnoir (1965b) as ochracea, was renamed geevestonia (IRScNB); dissected and mounted on three by Neboiss (1974c) because the name is pre- microscope slides. Type seen. occupied by the type species ochracea Curtis 1825. Material examined: Tasmania—5 6* 7 $ Prosser River nr. Orford, 13 Nov. 1972; 1 6* Prosser River 2 Comparing Jacquemart's type with the fig. 18 km W of Buckland, 7 Dec. 1974; 5 c? 10 $ Huon- (Jacquemart, 1965b) several omissions were Picton River junction, 18 Feb. 1967; 1 6* Huon River 148 ARTURS NEBOISS nr. Blakes Opening, 9 Feb. 1966; 1 155 Oecetis arcada Mosely Figures 796-800 Oecetis arcada Mosely in Mosely and Kimmins, 1953: 290. Oecetis atbodecorata Jacquemart, 1965b:23 syn. nov. This brownish species is the only one in Tas- mania with the cross-vein r-m situated its own length distad of the cross-vein closing discoidal cell. Small groups of darker hairs on cross- veins at anastomosis and on vein junctions on anterior wings, the intensity varies considerably from distinct to almost non existent. Spurs 1:2:2. The types of both species arcada and albo- decorata came from Cradle Mountain area. The Material examined: 5 o* 19 ? Waldheim, Cradle comparison of descriptions and figures of both, Mtn. Nat. Park, 7 Feb. 1971; 12 o* 45 $ Huon-Pic- ton River junction, 18 Feb. 1967 and 15 Nov. with specimens from the locality, leaves no 1972; 1 ? Lake Dobson, 20 Feb. 1967; 2 o* Cracroft River, 8 doubt of their synonymy. Feb. 1966; 4 1 ""Tv""t^ Type material: Holotype g (T5501), allotype &*( \ \ + tf^t • $ (T5502) Penstock Lagoon, Tas., 27 Feb. 1967, A. Neboiss (NMV). '* •7 Distribution: Tasmania—C province. fSE \ / \i ft- DISCUSSION \ sw V In their work on Australian and New Zea- * ; ^\ land Trichoptera Mosely and Kimmins (1953), listed 17 families of which only 13 were re- % M*u corded from Tasmania. Since then a consider- 156 o n 4C 60 km able number of changes has taken place in ^ > 157 higher classification (Lepneva 1956, 1964; Ross 1967). Some families have been sub- 9 abdomen terminates bluntly, lateral lobes divided, new families described, and others re- small. placed. The family classification adapted in this Length of anterior wing: S 8-11 mm; $ 9-10 paper is that used by Malicky (1973), which mm. is an amended version of that proposed by Type material; Type & Bathurst, N.S.W., 2400 Ross (1967). ft., 12 Nov. 1884, McLachlan Collection In the present study 21 families are recog- Type not seen. (BMNH). nized as occurring in Tasmania, although the South Esk Material examined: Tasmania—3 o* 8 $ position of three families—Calocidae, Helico- Evandale, 1 Mar. 1967, A. Neboiss (NMV); River, require fur- 12 phidae and Conoesucidae—might 1 myidae s. str. is now omitted from the Aus- have strong northern relationships, 13 of the 30 tralian fauna. Tasmanian species arc widespread on mainland Kokiriidac —the family had recently been Australia, and some have also been reported recognized in the Australian fauna (Neboiss from SE Asian localities. 1974b), and is now known to be represented The family Rhyacophilidae is represented by in Tasmania by three endemic species. The first two subfamilies—Apsilochoreminae and Hy- apparent record by Bayly et al ( 1972), of the drobiosinae. Apsilochoreminae, with only two genus similar to Kokiria from Lake Pedder, was species in Tasmania, both of which are also based on a preliminary identification of material widely distributed on the Australian mainland, by the present author, which is fully described represent a group with strong, northern relation- in this paper. ships. This subfamily is not known from New Tasimiidae—genus Tasimia, segregated from Zealand. the Sericostomatidae by Rick (1968) to form The second subfamily—Hydrobiosinae—is a separate family. dominated by trans-antarctic distribution pat- Calocidae —genus Caloca, segregated from the tern and at present contains 27 Tasmanian Odontoceridae by Ross (1967) to form a new species, of which only seven are shared with family; genus Caenota and Tamasia is now mainland Australia. There are no species com- added to it. The New Zealand family Pycnocen- mon to both Tasmania and New Zealand. The trellidae is reduced to synonymy. distribution of Australian species was discussed Oeconesidae group —a of New Zealand and illustrated by Neboiss (1962:522), de- genera previously regarded as a tribe of the monstrating that the largest number of species Sericostom atidae by Ti 1 lyard (1921), was was concentrated in the southeast of the con- raised to family rank by Neboiss (1975), when tinent, with rapidly decreasing numbers in the a new genus from Tasmania was added. This northerly direction. The present investigation is the only family found in Tasmania, but not transfers the centre of concentration to Tas- recorded from the Australian mainland. mania, emphasizing the southern origin of the Beraeidae the only Australian genus — placed group. The subfamily is represented with more in the family was Alloecella Banks, which is than 50 species in New Zealand. now transferred to the family Helicophidae; Several of the smaller families, most of them consequently, the family name Beraeidae is re- with very limited distribution, have typical moved from the list of Australian and Tas- trans-antarctic distribution patterns. To this manian Trichoptera. group belong families Philopotamidae (genus Conoesucidae erected — as a subfamily of Hydrobiosella only), Kokiriidae, Oeconesidae, the Sericostomatidae by Ross (1967), is raised Tasimiidae, Calocidae, Helicophidae, Conoesu- in this publication to family rank. The main cidae and Philorheithridae, all of them are clas- diagnostic features of the family are described sified as cool adapted forms (sensu Ross 1956). and the names of all general included are given. The available information on distribution A number of other Australian genera, pre- shows that 116 species or 74% are endemic to viously placed in the family Sericostomatidae, Tasmania. Future, more extensive collecting in had already been transferred to other families the southeastern part of the Australian main- (see p. 99), and as all the remaining ones be- land, particularly in Victoria, quite likely will long to Conoesucidae, the family name Sericos- reveal that some of them have wider range of tomatidae had to be removed from the list of distribution, but nevertheless, Tasmania is far Tasmanian and Australian Trichoptera. from being fully explored, and without doubt, In Tasmania, there are now recorded a total a number of yet unknown species could be dis- of 66 genera with 157 species, which are placed covered. It is therefore reasonable to assume in 21 families. Only two of these families con- that the proportion between endemic and wide- tain more than 20 species each Rhyacophili- — spread species will remain quite close to the dae, with 29 species and Leptoceridae with 30 present level. The proportion of endemic spe- species. Both subfamilies of the Leptoceridae cies is similar to that found in other aquatic TASMANIAN CADDIS-FLIES 151 insect groups in Tasmania, e.g. Plecoptera (as out of a total of 103 species, 75 are endemic, high as 82%—Hynes, 1976). The above- of which 14 are found only in this province. mentioned figures disagree with the statement The lowest proportion of endemic species is in by Guiler (1965) that \ . . Tasmania, although the E province, where, out of 42 recorded spe- once connected to the Australian continent, has cies, 21 or 50% are endemic to Tasmania, and very few endemics, both in its vertebrate and only two of them are restricted to the province. invertebrate fauna'. A difference of more than 20% in endemic Analysing individual provinces, it can be content between the two provinces seems large seen that not all of them have the same pro- enough to be of some significance. portion of endemic species. Calculated from the During the Pleistocene glacial phases, the total number of species recorded from each sea level was substantially lower, which al- province, the highest proportion of Tasmanian lowed formation of land bridge across the rela- endemics—as much as 73%—are found in the tively shallow Bass Strait, connecting Victoria, NW and SW provinces. In the NW province, from Wilsons Promontory over the Furneaux Map 3 —Endemism level of Tasmanian Trichoptera. 152 ARTURS NEBOISS Group of islands, with NE Tasmania (Jennings, spread, warm adapted species, common to the 1971). Recently accumulated evidence indicate Australian mainland and Tasmania, are found that there has been more than one glacial phase mainly NE of a line between Hobart and in Tasmania (Davies, 1974), and the last such Burnie, which coincides with the dominance of connection probably existed for a considerable sclerophyll forests (Vegetation map, Davies length of time between 32 000 and 10 000 1965). By contrast, the cooler and wetter con- years before present. ditions SW of the line, allowed uninterrupted persistence of It also has been established from the avail- the cool adapted, endemic spe- cies, able geological evidence, that during the maxi- coinciding with dominance of rainforests. mum of the last glaciation, Tasmania had a wet The total number of species, the number of endemic species western region and a much dryer region in the and the percentage of endemics from east, essentially as at present (Davies, 1974). the total, calculated from each province separately, To a large extent this explains why the wide- are shown on Map 3. CHECK LIST OF TASMANIAN TRICHOPTERA SUPERFAMILY RHY OPHI AC LOWEA N NE E C NW SW SE 1. Fam. Rhyacophilidae Stephens Apsilochorema Ulmer (1) obliqitum (Mosely) x X X X X X (2) gisbum (Mosely) X X X X Allochorema Mosely X X (3) tasmanicum Mosely Austrochorema Mosely X X X (4) pegidion Neboiss X X X X (5) evansi (Mosely) X (6) crinitum sp. n. X X (7) lepnevae Jacquemart X (8) complexa Jacquemart X X Ipsebiosis gen. n. (9) spicula sp. n. X X X Ulmer ochorema Mosely X X (10) breve (Mosely) X X (11) seona (Mosely) X X X X X X (12) lentum Neboiss X X (13) ony chion sp. n. X (14) tasmanicum (Mosely) X X X X X X (15) rubiconum Neboiss x X X X X X Ethochorema gen. n. X (16) secutum sp. n. X (17) nesydrion (Neboiss) X X X (18) kelion sp. n. X Taschorema Mosely X X (19) asmanum Mosely X X X X X X (20) apobamum sp. n. X X X X X X (21) ferulum sp. n. X X X X X (22) viridarium Neboiss X X X (23) pedunculatum Jacquemart X (24) evansi Mosely X X Ptychobiosis XXX X X X gen. n. (25) nigrita (Banks) X X Koetonga Neboiss X X X (26) clivicola Neboiss X Moruya Neboiss X (27) charadra Neboiss X X X X X (28) opora Neboiss X X X (29) tasmanica (Jacquemart) X X 2. Fam. Glossosomatidae Ulmer Agapetus Curtis (30) tasmanicus (Mosely) X cralus XXX X X X (31) (Mosely) X X X X X (32) laparus sp. n. X TASMANIAN CADDIS-FLIES 153 N NE NW sw SE 3. Fam. Hydroptilidae Stephens Orphninotrichia Mosely (33) acta sp. n. X Hydroptila Dalman (34) tasmanica Mosely X (35) scamandra sp. n. X Targatrichia gen. n. (36) zonata sp. n. X X X X Hellyethira gen. n. (37) vallecula sp. n. X X X X Trichoglene gen. n. (38) columba sp. n. X X X Maydenoptila gen. n. (39) cuneola sp. n. X X X (40) rupina sp. n. X SUPERFAMILY HYDROPSYCHOIDEA 4. Fam. Philopotamidae Wallengren Hydrobiosella Tillyard (41) corinna sp. n. X (42) orba sp. n. X (43) cerula sp. n. X (44) anasina sp. n. X (45) tasmanica Mosely X (46) armata Jacquemart X (47) cognata Kimmins X X X (48) sagitta sp. n. X (49) waddama Mosely X X X X 5. Fam. Stenopsychidae Martynov Stenopsychodes Ulmer (50) lineata sp. n. 6. Fam. Ecnomidae Ulmer Ecnomus McLachlan (51) tillyardi Mosely X X X X X X (52) russellius sp. n. X X X X (53) continentalls Ulmer X Ecnomina Kimmins (54) irrorata Kimmins X X X X X X (55) legula sp. n. X X X (56) vega sp. n. (57) batyle sp. n. 7. Fam. Polycentropodidae Ulmer Plectrocnemia Stephens X X X (58) altera sp. n. X (59) lacuna sp. n. X X (60) manicata sp. n. X X (61) caudata sp. n- X X Tasmanoplegas gen. n. X (62) spilota sp. n. X Nyctiophylax Brauer X (63) repandus sp. n. 8. Fam. Hydropsychidae Curtis Cheumatopsyche Wallengren X X (64) modica (McLachlan) X Smicrophylix gen. n. X X X (65) creektona sp. n. X (66) simplex (Jacquemart) Asmicridea Mosely X X X X (67) edwardsi (McLachlan) X (68) grisea (Mosely) Diplectrona Westwood X (69) castanea Kimmins (70) lyella sp. n. (71) bispinosa Jacquemart (72) tasmanica Jacquemart SUPERFAMILY LIMNEPHILOIDEA 9. Fam. Plectrotarsidae Mosely 1 154 ARTURS NEBOISS N NE C NW sw Plectrotarsus Kolenati (73) gravenhorsti Kolenati X X (74) tasmanicus Mosely X X X Liapota Neboiss (75) lavara Neboiss X Nanoplectrus gen. n. (76) truchanasi sp. n. X 10. Fam. Limnephilidae Kolenati Archaeophylax Kimmins (77) ochreus Mosely X (78) vernalis sp. n. X 1 . Fam. Kokiriidae McFarlane Taskiria gen. n. (79) austera sp. n. X (80) mccubbini sp. n. X Taskiropsyche gen. n. (81) lacustris sp. n. 12. Fam. Oeconesidae Tillyard Tascuna Neboiss (82) ignota Neboiss 13. Fam. Tasimiidae Riek Tasimia Mosely (83) palpata Mosely X XX (84) denticulata Jacquemart X (85) drepana sp. n. Tasiagma gen. n. (86) ciliata sp. n. 14. Fam. Helicopsychidae Ulmer Helicopsyche Siebold (87) bartona Mosely X X X X (88) murrumba Mosely X X 15. Fam. Calocidae Ross Caloca Mosely (89) tertia Mosely X (90) saneva (Mosely) X X X (91) ascita sp. n. Caenota Mosely (92) plicata Mosely X X X X X Tamasia Mosely (93 ) variegata Mosely X X X X X 16. Fam. Helicophidae Mosely Helicopha Mosely (94) astia Mosely (95) de lamarei Jacquemart X Alloecella Banks (96) grisea Banks X X X X (97) longispina Jacquemart X X X X (98) pilosa sp. n. X X X X 17. Fam. Conoesucidae Ross Hampa Mosely (99) patona Mosely X X X X Matasia Mosely (100) satana Mosely X X X X Costora Mosely (101) iena Mosely X (102) delora Mosely X X (103) ebenina sp. n. X X X X (104) ramosa Jacquemart X X (105) krene sp. n. X (106) seposita sp. n. (107) luxata sp. n. X (108) rotosca Mosely X X Lingora Mosely (109) aurata Mosely X X X (110) vesca sp. n. Conoesucus Mosely (111) from us Mosely X X X X (112) norelus Mosely X X X X X TASMANIAN CADDIS-FLIES 155 N NE E c NW sw SE (113) digitiferus Jacquemart X (114) nepotulussp.n. X X X X (115) brontensis sp. n. X X X Fam. Odontoceridae Wallengren Atriplectides Mosely (116) dubia Mosely X X X X Fam. Calomoseriatidae Ulmer Anisocentropus McLachlan (117) latifascia (Walker) X Fam. Philorheithridae Mosely Austrheithrus Mosely (118) rone wa Mosely X X X (119) glymma sp. n. X X X X Kosrheithrus Mosely (120) remulus sp. n. X X X Ramiheithrus Neboiss ( 121 ) kocinus Neboiss X Aphilorheithrus Mosely (122) stepheni Mosely XXX X X X X (123) pauxillus sp. n. X X (124) decoratus sp. tl. X X (125) luteolus sp. n. X Tasmanthrus Mosely (126) angustipennis Mosely X Fam. Leptoceridae Leach Westriplectes gen. n. (127) pedderensis sp. n. X Triplectides Kolenati (128) ciuskus Mosely X X X (129) magnus (Walker) X X (130) similis Mosely X X X (131) truncatus sp. n. X (132) bilobus sp. n. X X (133) proximus sp. n. X (134) elongatus Banks X X Notoperata gen. n. (135) sparsa (Kimmins) X X (136) maculata (Mosely) X X X X Symphitoneuria Ulmer (137) opposita (Walker) X X Triplectidina Mosely (138) nigricornis Mosely X X X Lectrides Mosely (139) varians Mosely X X X Notalina Mosely (140) parked Mosely X X (141) fulva Kimmins X X X X X X (142) tillyardi Kimmins X (143) bifaria sp. n. X X X X X (144) nigra (Mosely) X X X Condocerus gen. n. (145) paludosus sp. n. X X Leptorussa Mosely (146) darlingtoni (Banks) X X Triaenodes McLachlan (147) intricata sp. n. X X Oecetis McLachlan (148) pechana Mosely X X X X (149) umbra sp. n. X (150) gilva sp. n. X (151) australis (Banks) X X X X (152) minasata Mosely X (153) laustra Mosely X (154) asmanista Mosely X (155) arcada Mosely X X X (156) inscripta Kimmins X X X (157) scirpiculasp.n. X IS6 ARTURS NFBOISS REFERENCES Kimmins, D. F. 1951. A note on the females of the British species of Beraeidae (Trichoptera). Ef> Hanks, N. 1913. Synopses and descriptions of exolic (omologist. 84: 19-21. Neuroptera. Tratis. Am. ent. Soc, 39: 201-242. 1957* Notes on the Psychomyidac from the - 1920, New neuropterotd insects, Htdl. Mus. African Trans. Sac. Land. 109: nun mainland. R. ent, ik ZooL llurv. 64: 299-362. 259-273. 1939, New genera and species of Neurop- — 1958. types of Anisocentropus latifas- Keroid insects. Hull. Mus. comp. ZooL Harv* 8S: The 440 soj ria (Walker) and elegans (Walker). Entomo- logist. 91: 167-168. IUyiv, I. A. I ., LAKE, P. S., Swain, R. and I aykor, I*. 1960. to 'The Trichoptera (Cad- A. 1972. Lake Pcddei : its importance lo bio- Corrections logical science. 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Diplectrona 71 asmanum, Taschorema 27, 28, 29, Condocerus 140 ferulum, Taschorema 27, 29, 164 163, 164 Confluens 99 flavomaculata, Hydropsyche 71 Asmicridea 66, 69, 71 70, Conoesucidae 10, 99, 100, 149, fromus, Conoesucus 109, 110, 190 astia, Helicopha 94, 95, 96, 185 150 fulva, Notalina 137, 138, 201 Athripsodes 140, 141 Conoesucinae 99 Atriplectides 113 Conoesucus 99, 100, 109, 110, 111, aurata, Lingora 107, 108, 190 112 austera, galbinomaculatus, Tasmanthrus Taskiria 80, 81, 179, 180 continentalis, Ecnomus 55, 56, 777 124, 125 australica, Plectrocnemia 9, 60, 61, corinna, Hydrobiosella 46, 47, 169 Ganonema 115, 116 62, 173 Conuxia 99 australis, Oecetina 145 geevestonia, Oecetis 147 Costora 99, 100, 102, 103, 104, australis, gilva, Oecetis 143, 145, 203 Oecetis 143, 145, 203 105, 106 Austrheithrus gisba, Apsilochorema 13 116, 117, 118 crala, Synagapetus 37 Austrochorema gisba, Bachorema 13 11, 14, 15, 16, 17, cralus, Agapetus 36, 37, 38, 167 18 gisbum, Apsilochorema 12, 13. creektona. Smicrophylax 67, 68, 158 775, 776 Glossosomatidae 9, 36, 84, 149 crinitum, Austrochorema 15, 16, B glymma, Austrheithrus 117, 118, 17, 160 Bachorema 12 794 cuneola, Maydenoptila bartona, Helicopsyche 44, 168, Goera 115 87, 88, 182 169 batyle, Ecnomina 57, 59, 172 Goeridae 83 Beraeidae 94, 150 D gracilis, Mystacides 127 Beraeoptera 99 darlingtoni, Leptocerus 141 gravenhorsti, Plectrotarsus 75, 76, bicolor, 777 Leptocerus 142 darlingtoni, Leptorussa 141, 202 bicolorata, grisea, Alloecella Ganonema 115, 116 decoratus, Aphilorheithrus 121, 96, 97, 186 bicoloratus, Anisocentropus 9, 114, 122, 123, 196 grisea, Asmicridea 70, 71, 776 grisea, 115, 116,795 delamarei, Helicopha 96, 185, 186 Smicridea 69, 71 bifaria, Notalina 137, 138, 139, delora, Costora 102, 103, 188 201 denticulata, Tasimia 84, 85, 87, H bilobus, Triplectides 127, 130, 181 Hampa 99, 100 198, 199 digitiferus, Conoesucus 109, 111, Helicopha 94, 95, 96 bispinosa, Diplectrona 71, 73, 777 797 Helicophidae 10, 94, 149, 150 bola, Marilia 113 Diplectrona 66, 69, 71, 72, 73 Helicopsyche 87, 88, 99 206 TASMANIAN CADDIS-FLIES 207 Helicopsychidae 10, 87, 99 Loticana 133, 134 obliquum, Apsilochorema 12, 13, Hellyethira 39, 42 lurida, Oecetis 143 158 hesperium, Ethochorema 24 luteolus, Aphilorheithrus 121, 123, ochracea (Curtis), Oecetis 147 Hudsonema 132, 133 196 ochracea (Jacquemart), Oecetis 147 Hydrobiosella 45, 46, 47, 48, 49, luxata, Costora 102, 106, 189 ochraceum, Ethochorema 24 50,51,52, 150 lyella, Diplectrona 71, 72, 177 ochraceus, Leptocerus 143 Hydrobiosinae 10, 11, 14, 150 ochraceus (Curtis), Oecetis 143 Hydrobiosini 10, 14 M ochreus, Archaeophylax 78, 179 Hydrobiosis 19 maculata, Orphninotrichia 40 Odontoceridae 9, 89, 90, 113 Hydropsyche 60, 66, 70, 71 maculata, Hudsonema 133 Oecetina 145 Hydropsychidae 9, 66 maculata, Notoperata 132, 133, Oecetis 133, 140, 142, 143, 144, Hydropsychinae 66 199 145, 146, 147, 148, 149 Hydropsychini 66 Macronematinae 66 Oeconesidae 9, 83, 99, 150 Hydropsychodes 66 magna, Notanatolica 128 Oeconesini 83 Hydropsychoidea 45 magna, Triplectides 128 Oeconesus 83, 99 Hydroptila 39, 40, 41 magnus, Leptocerus 128 Olinga 99 Hydroptilidae 9, 39 magnus, Triplectides 127, 128, 129, onychion, Ulmerochorema 19, 21, 162 I 198 manicata, Plectrocnemia 60, 62, opora, Moruya 34, 166 iena, Costora 102, 103 173 opposita, Loticana 134 ignota, Tascuna 83, 181 Marilia 113 opposita, Notanatolica 134 illustris, Anisocentropus 114 marlieri, Helicopha 95 opposita, Symphitoneuria 134, 200 indicum, Psilochorema 12 Matasia 99, 100, 101 oppositus, Leptocerus 1, 134 inscripta, Oecetis 143, 148, 205 Maydenoptila 39, 44, 45 orba, Hydrobiosella 46, 47, 169 intricata, Triaenodes 142, 202 mccubbini, Taskiria 81, 180 oreolimnetes, Triplectides 134 Ipsebiosis 11, 18 Megogata 11 Orphninotrichia 39, 40 irrorata, Ecnomina 57, 171 miharo, Kokiria 80 Orthotrichia42 iti, Oecetis 143 minasata, Oecetis 143, 146, 204 P K mjobergi, Stenopsychodes 53 pallescens, Taschorema 27 kelion, Ethochorema 24, 26, 163 modica, Cheumatopsyche 66, 175 palpata, Tasimia 84, 85, 181 kimminsi, Taschorema 27 modica, Hydropsyche 66 paludosus, Condocerus 140, 201, kocinus, Ramiheithrus 120, 195 modica, Hydropsychodes 66 202 Koetonga 11, 33 Molannidae 94 parkeri, Notalina 136, 137, 138, Kokiria 80, 150 Molanniella 84 200 Kokiriidae9, 80, 150 montana, Stenopsychodes 53 Paroxyethira 42 Kosrheithrus 116, 117, 119 monticolus, Agapetus 36 parvula, Smicrophylax 68 Nyctiophylax 65 krene, Costora 103, 105, 189 Moruya 10, 11, 33, 35 parvus, moselyi, Conoesucus 109, 110 patona, Hampa 100, 187 L murrumba, Helicopsyche 88, 182, pauxillus, Aphilorheithrus 121, lacuna, Plectrocnemia 60, 62, 173 183 122, 196 lacustris, Taskiropsyche 82, 180, Mystacides 127 pechana, Oecetis 143, 144, 145, 181 N 202, 203 laparus, Agapetus 36, 38, 167 pedderensis, Westriplectes 126, latifascia, Anisocentropus 114, 115, Nanoplectrus 75, 77 197 116, 193 nepotulus, Conoesucus 109, 111, pedunculata, Taschorema 30 latifascia, Notidobia 115 191, 192 pedunculatum, Taschorema 27, laustra, Oecetis 143, 146, 204 nesydrion, Ethochorema 24, 25, 30, 165 lavara, Liapota 64, 76, 77, 178 163 pegidion, Austrochorema 14, 15, Lectrides 126, 135 nesydrion, Taschorema 24, 25 159 legula, Ecnomina 57, 58, 172 nigra, Notalina 137, 139, 201 Philopotamidae 9, 45, 53, 150 lentum, Ulmerochorema 19, 21, 22, nigra, Taschorema 32 Philopotamus 55 162 nigra, Triplexina 136, 139 Philorheithridae 9, 116, 117, 122, lepida, Hydropsyche 66 nigricornis, Triplectidina 135, 200 150 Lepidostomatidae 84 nigrita, Notiobiosis 32 pilosa, Alloecella 96, 97, 98, 187 lepnevae, Austrochorema 15, 17, nigrita, Ptychobiosis 32, 165 Plectrocnemia 60, 61, 62, 63, 64 160 nigrita, Taschorema 32 Plectrotarsidae 9, 74 Leptoceridae 9, 125, 150 nivea, Smicridea 70 Plectrotarsus 74, 75, 76 Leptocerinae 125, 140 norelus, Conoesucus 109, 110, 191 plicata, Caenota 92, 184 Leptocerus 128, 133, 134, 140, 142 Notalina 126, 136, 137, 138, 139 Polycentropodidae 9, 53, 54, 59, Leptorussa 140, 141, 142 Notanatolica 127, 128, 134 60 Liapota 64, 74, 76, 77 Notidobia 115 Polyplectropus 60 Limnephilidae 9, 78 Notiobiosis 26, 32 Potamyia 67 Limnephiloidea 74 Notoperata 126, 132, 133 proximus, Triplectides 127, 130, lineata, Stenopsychodes 53, 170 Nyctiophylax 60, 65 799 Pseudoeconesus 99 Lingora 99, 100, 107, 108 O 83, lobata, Triplexina 133 Pseudonema 127 longispina, Alloecella 96, 97, 186, obliqua, Apsilochorema 12 Psilochorema 12 187 obliqua, Bachorema 12 Psychomyidae 53, 54, 60, 149 1 208 ARTURS NEBOISS Psychomyinae 54 Smicridea 67, 68, 69, 70 tasmanicus, Synagapetus 36, 37 Psyllobetina 10, 11, 35 Smicrideini 66 Tasmanoplegas 60, 64 Psyllobetini 10, 33 Smicrophylax 66, 67, 68, 69, 70, 71 Tasmanthrus 116, 117, 124, 125 Ptychobiosis 11, 32 soena, Anachorema 20 tenellus, Philopotamus 55 Pycnocentrella 89 Sortosa 45 Pycnocentrellidae tertia, Caloca 90 89, 150 sparsa, Hudsonema 132 Pycnocentria tillyardi, Notalina 137, 138 99, 100 sparsa, Notoperata 132, 133, 199 tillyardi, Ecnomus 55, 56, 171 Pycnocentrodes 99 spicula, Ipsebiosis 18, 161 tillyardi, Kosrheithrus 119 spilota, Tasmanoplegas 64, 174 tineoides, Hydroptila 40 spinosa, Ecnomina 57, 58 Tiphobiosis 10 Ramiheithrus 116, 117, 120 spinosa, Hydrobiosella 50, 51 Tismana 89, 90, 91,99 ramosa, Costora 103, 104, 105, squamosa, Oecetis 143 Triaena 142 stenocerca, Hydrobiosella 46 Triaenodes remulus, Kosrheithrus 140, 142 119, 195 Stenoosyche 53 Trichoglene repandus, Nyctiophylax 39, 43 65, 174 Stenopsychidae 9, 53, 60, 149 175 Triplectides 126, 127, 128, 129, Stenopsychodes 53, 60, 149 130, 131,132, 134, 135, 136 Rhyacophilidae 9, 10, 150 stepheni, Aphilorheithrus 120, 121, Triplectidina 126, 134, 135 Rhyacophilinae 10 122, 195 Triplectidinae 125 Rhyacophiloidea 10 stigma, Hydrobiosis 19 Triplexina 133, 136, 139 Rhyacophylax 67, 68 straminea, Caloca 90 truchanasi, rieki, Ptychobiosis 32 Nanoplectrus 77, 178, styliferus, Synagapetus 37, 38 179 rieki, Taschorema 32 Symphitoneuria 126, 133, 134 truncatus, ronewa, Austrheithrus Triplectides 127, 129, 117, 118 Synagapetus 36, 37, 38 194 131, 198 turbidum, Ethochorema rotosca, Costora 102, 106, 189 24 rubiconum, Ulmerochorema 19 23, 162 U Tamasia 89, 90, 93, rugulum, 99, 150 Ulmerochorema 10, 11, 18, 19, Taschorema 27 Tanjilana rupina, Maydenoptila 1 21,22,23 44, 45, 168, Tanjistomella 169 80 umbra, Oecetis 143, 144, 203 Tarapsyche 99 uncinata, russata, Leptorussa 141, 142 Hydrobiosella 46 Targatrichia 39, 41 unicolor, russellius, Ecnomus 55, 56, 171 Oecetis 143, 144 Taschorema 10, 11, 24, 26, 27, 32, S 158 Tascuna 83 vallecula, Hellyethira Saetotricha 99 Tasiagma 42, 168 84, 86 varians, Lectrides sagitta, Hydrobiosella 46, 47, 51 135, 200 ' Tasimia 84, 85, 86. 99, 150 170 variegata, Tamasia 89, 93, 184, 185 Tasimiidae 10, 84, 99, 150 saneva, Caloca vega, Ecnomina 57, 58, 172 89, 91, 183 Taskiria80, 81 saneva, vernalis, Archaeophylas Tismana 89, 90, 91 Taskiropsyche 79, 179 80, 82 vesca, Lingora satana, Matasia 101, 187, 188 Tasmania 108, 190 76, 77 virgatus, scamandra, Hydroptila 41, Ramiheithrus 120 168 tasmanica, Allochorema 13 scirpicula, viridarium, Taschorema Oecetis 143, 149, 205 tasmanica, 27, 29, 30, Anachorema 22 165 secutum, Ethochorema 24, 162 tasmanica, Diplectrona 71, 73, 177 senex, Hydropsyche 60 tasmanica, Hydrobiosella W seona, Anachorema 46, 47 20 49, 50, 169 waddama, seona, Hydrobiosella 46, 52, Ulmerochorema 19, 20 22 tasmanica, Hydroptila 40 161 170 tasmanica, Moruya 33, 35, 166 warneria, Alloecella 97 seposita, Costora 102, 106, 189 tasmanica, Psyllobetina 35 wenta, Austrochorema 14 Sericostomatidae 83, 99, 100, 150 tasmanicum, Allochorema 13, 159 Westriplectes 126 shuttleworthi, Helicopsyche 87 tasmanicum, Ulmerochorema 19, similis, Triplectides 127, 129, 198 22, 23, 162 simplex. Diplectrona 69, 71 tasmanicus, Agapetus 36, 37, 167 Zelandopsyche 83, 99 simplex, Smicrophylax 68, 69, 176 tasmanicus. Plectrotarsus 1, 75, sinensis, Zepsyche 99 Nyctiophylax 65 76, 777, 178 zonata, Targatrichia 41, 168 Plates 1-3 1. Huon Plains looking west from Mt. Eliza, showing Scotts Peak Dam access road, Con- dominion Creek crossing indicated by the widening in the road. Forested line across the plain indicates Huon River. Lake Pedder in the background. 2. Condominion Creek—in the foreground disturbed creek bed near the road. 3. Russell Falls, National Park. MEM. NAT. MUS. VICT. 38 PLATE 1 MEM. NAT. MUS. VICT. 38 PLATE 2 MEM. NAT. MUS. VICT. 38 PLATE 3 ^rt«2-a>o5 £! > & ' ^.j^h5= « 7? O