PART 3 Polychaeta Errantia
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National Monitoring Program for Biodiversity and Non-Indigenous Species in Egypt
UNITED NATIONS ENVIRONMENT PROGRAM MEDITERRANEAN ACTION PLAN REGIONAL ACTIVITY CENTRE FOR SPECIALLY PROTECTED AREAS National monitoring program for biodiversity and non-indigenous species in Egypt PROF. MOUSTAFA M. FOUDA April 2017 1 Study required and financed by: Regional Activity Centre for Specially Protected Areas Boulevard du Leader Yasser Arafat BP 337 1080 Tunis Cedex – Tunisie Responsible of the study: Mehdi Aissi, EcApMEDII Programme officer In charge of the study: Prof. Moustafa M. Fouda Mr. Mohamed Said Abdelwarith Mr. Mahmoud Fawzy Kamel Ministry of Environment, Egyptian Environmental Affairs Agency (EEAA) With the participation of: Name, qualification and original institution of all the participants in the study (field mission or participation of national institutions) 2 TABLE OF CONTENTS page Acknowledgements 4 Preamble 5 Chapter 1: Introduction 9 Chapter 2: Institutional and regulatory aspects 40 Chapter 3: Scientific Aspects 49 Chapter 4: Development of monitoring program 59 Chapter 5: Existing Monitoring Program in Egypt 91 1. Monitoring program for habitat mapping 103 2. Marine MAMMALS monitoring program 109 3. Marine Turtles Monitoring Program 115 4. Monitoring Program for Seabirds 118 5. Non-Indigenous Species Monitoring Program 123 Chapter 6: Implementation / Operational Plan 131 Selected References 133 Annexes 143 3 AKNOWLEGEMENTS We would like to thank RAC/ SPA and EU for providing financial and technical assistances to prepare this monitoring programme. The preparation of this programme was the result of several contacts and interviews with many stakeholders from Government, research institutions, NGOs and fishermen. The author would like to express thanks to all for their support. In addition; we would like to acknowledge all participants who attended the workshop and represented the following institutions: 1. -
Annelida, Phyllodocida)
A peer-reviewed open-access journal ZooKeys 488: 1–29Guide (2015) and keys for the identification of Syllidae( Annelida, Phyllodocida)... 1 doi: 10.3897/zookeys.488.9061 RESEARCH ARTICLE http://zookeys.pensoft.net Launched to accelerate biodiversity research Guide and keys for the identification of Syllidae (Annelida, Phyllodocida) from the British Isles (reported and expected species) Guillermo San Martín1, Tim M. Worsfold2 1 Departamento de Biología (Zoología), Laboratorio de Biología Marina e Invertebrados, Facultad de Ciencias, Universidad Autónoma de Madrid, Canto Blanco, 28049 Madrid, Spain 2 APEM Limited, Diamond Centre, Unit 7, Works Road, Letchworth Garden City, Hertfordshire SG6 1LW, UK Corresponding author: Guillermo San Martín ([email protected]) Academic editor: Chris Glasby | Received 3 December 2014 | Accepted 1 February 2015 | Published 19 March 2015 http://zoobank.org/E9FCFEEA-7C9C-44BF-AB4A-CEBECCBC2C17 Citation: San Martín G, Worsfold TM (2015) Guide and keys for the identification of Syllidae (Annelida, Phyllodocida) from the British Isles (reported and expected species). ZooKeys 488: 1–29. doi: 10.3897/zookeys.488.9061 Abstract In November 2012, a workshop was carried out on the taxonomy and systematics of the family Syllidae (Annelida: Phyllodocida) at the Dove Marine Laboratory, Cullercoats, Tynemouth, UK for the National Marine Biological Analytical Quality Control (NMBAQC) Scheme. Illustrated keys for subfamilies, genera and species found in British and Irish waters were provided for participants from the major national agencies and consultancies involved in benthic sample processing. After the workshop, we prepared updates to these keys, to include some additional species provided by participants, and some species reported from nearby areas. -
Morphological Investigation and Analysis of Ribosomal DNA Phylogeny of Two Scale-Worms (Polychaeta, Polynoidae) from the Gulf of Thailand
Songklanakarin J. Sci. Technol. 40 (5), 1158-1166, Sep. - Oct. 2018 Original Article Morphological investigation and analysis of ribosomal DNA phylogeny of two scale-worms (Polychaeta, Polynoidae) from the Gulf of Thailand Arin Ngamniyom1*, Rakchanok Koto2, Weerawich Wongroj3, Thayat Sriyapai1, Pichapack Sriyapai4, and Busaba Panyarachun5 1 Faculty of Environmental Culture and Eco-tourism, Srinakharinwirot University, Watthana, Bangkok, 10110 Thailand 2 Department of Biology, Faculty of Sciences, Srinakharinwirot University, Watthana, Bangkok, 10110 Thailand 3 Prasarnmit Elementary Demonstration School, Srinakharinwirot University, Watthana, Bangkok, 10110 Thailand 4 Department of Microbiology, Faculty of Sciences, Srinakharinwirot University, Watthana, Bangkok, 10110 Thailand 5 Department of Anatomy, Faculty of Medicine, Srinakharinwirot University, Watthana, Bangkok, 10110 Thailand Received: 14 December 2016; Revised: 7 June 2017; Accepted: 5 July 2017 Abstract Scale-worms are polychaetes of the family Polynoidae that are commonly distribute in marine environments. This study aims identify and introduce two scale-worms as Capitulatinoe cf. cupisetis and Eunoe cf. oerstedi from the western coast of the Gulf of Thailand. Using scanning electron microscopy of adult worms, the antennae, palps, prostomium, cirri, setigers, parapodia, saetae and elytra are described. In addition, the phylogenetic relationships of our specimens with other polychaete species were analyzed based on partial sequences of 28S, 18S and 16S ribosomal DNA (rDNA) genes. The rDNA sequences identified C. cf. cupisetis and E. cf. oerstedi were respectively recovered within Arctonoinae and Polynoinae in a monophyletic Polynoidae. The congruence or incongruence of the morphological and molecular data is discussed in the text. These findings increase the knowledge of polynoid polychaete worms in Thailand, although two scale-worms remain to be identified of the precise species. -
Onetouch 4.0 Sanned Documents
Vol. 82, pp. 1-30 29 May, 1969 PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON REVIEW OF SOME SPECIES REFERRED TO SCALISETOSUS MCINTOSH (POLYCHAETA, POLYNOIDAE) BY MABIAN H. PETTIBONE Smithsonian Institution, Washington, D. C. In connection with an extended review of the polynoid gen- era, based on a study of the type-species, it was foimd that Scalisetosus Mclntosh ( 1885) has been used for a heterogenous group of species. The genus has served to include species with setae as transparent as crystal and the neurosetae character- ized by the presence of a basal semilunar cusp or pocket, al- though this particular feature was not shown on the figure of the neiu-osetae of the type-species, S. ceramensis, by Mclntosh (1885, pi. lOA, fig. 14). Any species equipped with this pe- culiar type of neiu-osetae has been placed in Scalisetosus, re- gardless of other characters. Saint-Joseph (1899) proposed the new genus Adyte for three species {Polynoe pellucida Ehlers, Hermadion assimile Mclntosh, and H. echini Giard) having the peculiar type of neurosetae, separating them from S. cera- mensis, which lacks the basal semilunar cusps. Mclntosh (1900) was responsible for changing the diagnosis of Scali- setosus to include the species referred to Adyte by Saint- Joseph, Subsequently, Adyte was abandoned and was synony- mized with Scalisetosus by Fauvel (1914, p. 47). During a visit to the British Museum of Natural History in May 1967, I was able to examine the unique type of Scalise- tosus ceramensis and to verify that the neurosetae indeed do lack the basal semilunar cusps and that the species therefore lacks one of the key characters that has been attributed to the genus. -
DEEP SEA LEBANON RESULTS of the 2016 EXPEDITION EXPLORING SUBMARINE CANYONS Towards Deep-Sea Conservation in Lebanon Project
DEEP SEA LEBANON RESULTS OF THE 2016 EXPEDITION EXPLORING SUBMARINE CANYONS Towards Deep-Sea Conservation in Lebanon Project March 2018 DEEP SEA LEBANON RESULTS OF THE 2016 EXPEDITION EXPLORING SUBMARINE CANYONS Towards Deep-Sea Conservation in Lebanon Project Citation: Aguilar, R., García, S., Perry, A.L., Alvarez, H., Blanco, J., Bitar, G. 2018. 2016 Deep-sea Lebanon Expedition: Exploring Submarine Canyons. Oceana, Madrid. 94 p. DOI: 10.31230/osf.io/34cb9 Based on an official request from Lebanon’s Ministry of Environment back in 2013, Oceana has planned and carried out an expedition to survey Lebanese deep-sea canyons and escarpments. Cover: Cerianthus membranaceus © OCEANA All photos are © OCEANA Index 06 Introduction 11 Methods 16 Results 44 Areas 12 Rov surveys 16 Habitat types 44 Tarablus/Batroun 14 Infaunal surveys 16 Coralligenous habitat 44 Jounieh 14 Oceanographic and rhodolith/maërl 45 St. George beds measurements 46 Beirut 19 Sandy bottoms 15 Data analyses 46 Sayniq 15 Collaborations 20 Sandy-muddy bottoms 20 Rocky bottoms 22 Canyon heads 22 Bathyal muds 24 Species 27 Fishes 29 Crustaceans 30 Echinoderms 31 Cnidarians 36 Sponges 38 Molluscs 40 Bryozoans 40 Brachiopods 42 Tunicates 42 Annelids 42 Foraminifera 42 Algae | Deep sea Lebanon OCEANA 47 Human 50 Discussion and 68 Annex 1 85 Annex 2 impacts conclusions 68 Table A1. List of 85 Methodology for 47 Marine litter 51 Main expedition species identified assesing relative 49 Fisheries findings 84 Table A2. List conservation interest of 49 Other observations 52 Key community of threatened types and their species identified survey areas ecological importanc 84 Figure A1. -
Annelida, Amphinomidae) in the Mediterranean Sea with an Updated Revision of the Alien Mediterranean Amphinomids
A peer-reviewed open-access journal ZooKeys 337: 19–33 (2013)On the occurrence of the firewormEurythoe complanata complex... 19 doi: 10.3897/zookeys.337.5811 RESEARCH ARTICLE www.zookeys.org Launched to accelerate biodiversity research On the occurrence of the fireworm Eurythoe complanata complex (Annelida, Amphinomidae) in the Mediterranean Sea with an updated revision of the alien Mediterranean amphinomids Andrés Arias1, Rômulo Barroso2,3, Nuria Anadón1, Paulo C. Paiva4 1 Departamento de Biología de Organismos y Sistemas (Zoología), Universidad de Oviedo, Oviedo 33071, Spain 2 Pontifícia Universidade Católica do Rio de Janeiro , Rio de Janeiro, Brazil 3 Museu de Zoologia da Unicamp, Campinas, SP, Brazil 4 Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro (UFRJ) , Rio de Janeiro, RJ, Brasil Corresponding author: Andrés Arias ([email protected]) Academic editor: C. Glasby | Received 17 June 2013 | Accepted 19 September 2013 | Published 30 September 2013 Citation: Arias A, Barroso R, Anadón N, Paiva PC (2013) On the occurrence of the fireworm Eurythoe complanata complex (Annelida, Amphinomidae) in the Mediterranean Sea with an updated revision of the alien Mediterranean amphinomids. ZooKeys 337: 19–33. doi: 10.3897/zookeys.337.5811 Abstract The presence of two species within the Eurythoe complanata complex in the Mediterranean Sea is reported, as well as their geographical distributions. One species, Eurythoe laevisetis, occurs in the eastern and cen- tral Mediterranean, likely constituting the first historical introduction to the Mediterranean Sea and the other, Eurythoe complanata, in both eastern and Levantine basins. Brief notes on their taxonomy are also provided and their potential pathways for introduction to the Mediterranean are discussed. -
Neanthes Limnicola Class: Polychaeta, Errantia
Phylum: Annelida Neanthes limnicola Class: Polychaeta, Errantia Order: Phyllodocida, Nereidiformia A mussel worm Family: Nereididae, Nereidinae Taxonomy: Depending on the author, Ne- wider than long, with a longitudinal depression anthes is currently considered a separate or (Fig. 2b). subspecies to the genus Nereis (Hilbig Trunk: Very thick segments that are 1997). Nereis sensu stricto differs from the wider than they are long, gently tapers to pos- genus Neanthes because the latter genus terior (Fig. 1). includes species with spinigerous notosetae Posterior: Pygidium bears two, styli- only. Furthermore, N. limnicola has most form ventrolateral anal cirri that are as long as recently been included in the genus (or sub- last seven segments (Fig. 1) (Hartman 1938). genus) Hediste due to the neuropodial setal Parapodia: The first two setigers are unira- morphology (Sato 1999; Bakken and Wilson mous. All other parapodia are biramous 2005; Tusuji and Sato 2012). However, re- (Nereididae, Blake and Ruff 2007) where both production is markedly different in N. limni- notopodia and neuropodia have acicular lobes cola than other Hediste species (Sato 1999). and each lobe bears 1–3 additional, medial Thus, synonyms of Neanthes limnicola in- and triangular lobes (above and below), called clude Nereis limnicola (which was synony- ligules (Blake and Ruff 2007) (Figs. 1, 5). The mized with Neanthes lighti in 1959 (Smith)), notopodial ligule is always smaller than the Nereis (Neanthes) limnicola, Nereis neuropodial one. The parapodial lobes are (Hediste) limnicola and Hediste limnicola. conical and not leaf-like or globular as in the The predominating name in current local in- family Phyllodocidae. (A parapodium should tertidal guides (e.g. -
H1.1 Open Water
PAGE .............................................................. 392 ▼ H1.1 OPEN WATER The open-water offshore habitat covers an area of by which solar energy enters the marine ecosystem, Nova Scotia larger than the land mass, and includes similar to the layer of plants on land. The ocean H1.1 Open Water salt water in inlets, bays and estuaries. The water waters are distinctive in having fostered the origins and the organisms it supports are the primary means of life on the planet. Plate H1.1.1: Right Whale, north of Brier Island (Unit 912). Photo: BIOS Habitats Natural History of Nova Scotia, Volume I © Nova Scotia Museum of Natural History .............................................................. PAGE 393 ▼ FORMATION PLANTS Oceans are formed as part of major geological events. The plants of the open ocean are almost entirely Nova Scotia’s open-ocean habitats are part of the microscopic algae, collectively known as phyto- Atlantic Ocean, which opened during the Jurassic plankton. Many different species occur, including Period and has been in continuous existence ever representatives of the prochlorophytes (blue-green since. The quality and depth of the water column algae—evolutionary intermediates between bacteria have fluctuated in relation to post-glacial climatic and algae), diatoms, dinoflagellates, chrysomonads, conditions. cryptomonads, minute flagellates and unicellular reproductive stages of macroscopic algae. Phyto- H1.1 PHYSICAL ASPECTS plankton are often grouped in size classes: Open Water 1. Water conditions, such as salinity, temperature, macroplankton: 200–2000 micrometres, includes ice-formation, turbidity, light penetration, tides larger diatoms. and currents, are extremely variable in the microplankton: 20–200 micrometres, includes waters offshore. most diatoms. 2. Air-water interaction, surface-water turbulence nanoplankton: 2–20 micrometres, includes determines the level of wave and gas exchange. -
OREGON ESTUARINE INVERTEBRATES an Illustrated Guide to the Common and Important Invertebrate Animals
OREGON ESTUARINE INVERTEBRATES An Illustrated Guide to the Common and Important Invertebrate Animals By Paul Rudy, Jr. Lynn Hay Rudy Oregon Institute of Marine Biology University of Oregon Charleston, Oregon 97420 Contract No. 79-111 Project Officer Jay F. Watson U.S. Fish and Wildlife Service 500 N.E. Multnomah Street Portland, Oregon 97232 Performed for National Coastal Ecosystems Team Office of Biological Services Fish and Wildlife Service U.S. Department of Interior Washington, D.C. 20240 Table of Contents Introduction CNIDARIA Hydrozoa Aequorea aequorea ................................................................ 6 Obelia longissima .................................................................. 8 Polyorchis penicillatus 10 Tubularia crocea ................................................................. 12 Anthozoa Anthopleura artemisia ................................. 14 Anthopleura elegantissima .................................................. 16 Haliplanella luciae .................................................................. 18 Nematostella vectensis ......................................................... 20 Metridium senile .................................................................... 22 NEMERTEA Amphiporus imparispinosus ................................................ 24 Carinoma mutabilis ................................................................ 26 Cerebratulus californiensis .................................................. 28 Lineus ruber ......................................................................... -
(Polychaeta) Borings in Paraspirifer Bownockeri (Brachiopoda: Devonian)1
114 A. E. ANNALA AND L. A. KAPUSTKA Vol. 83 Copyright © 1983 Ohio Acad. Sci. 003O-O95O/83/0003-O114 $2.00/0 VERMIFORICHNUS (POLYCHAETA) BORINGS IN PARASPIRIFER BOWNOCKERI (BRACHIOPODA: DEVONIAN)1 R. D. HOARE and R. L. WALDEN, Department of Geology, Bowling Green State University, Bowling Green, OH 43403 ABSTRACT. Shells of Paraspirifer bownockeri (Stewart) from the Silica Formation, Middle Devonian of northwestern Ohio, commonly contain numerous borings of a polychaete worm forming the endolithic trace fossil Vermiforichnus clarki Cameron (1969a) which can be exposed by acidizing the specimens. The borings are most abundant on the brachial valve, and their surface openings tend to be concentrated along major growth lines thence extending dominantly in the general direction of the beaks of the valves. In- festations of the polychaete occurred at 2 different time intervals as indicated by the spac- ing of the borings on 2 major growth lines with renewed shell growth between them. Growth of the host was severely reduced immediately following the infestation and in some areas damage to the mantle caused deformation in the shell of the host. OHIO J. SCI. 83 (3): 114-119, 1983 INTRODUCTION (1932) by Hoare and Steller (1967) (fig. 1), Previous interpretations of the larger as boring sponges by Kesling and Chilman borings commonly seen in the brachiopod (1975) and as "Clionoides" sp. by Steller Paraspirifer bownockeri (Stewart) from the (1965), Kesling et al. (1980) and Sparks Silica Formation in northwestern Ohio et al. (1980). These interpretations were have been alluded to as sponge borings, based on the external configuration of the Clionoides thomasi Fenton and Fenton surface opening of the boring only. -
Bacterial Diversity in the Gorgonian Coral Eunicella Labiata and How
Jamie M. Maxwell Gorgoniapolynoe caeciliae (Annelida: Phyllodocida) revisited: previously unknown characters and undescribed species from the Equatorial North Atlantic UNIVERSIDADE DO ALGARVE Faculdade de Ciências e Tecnologia 2019 Jamie M. Maxwell Gorgoniapolynoe caeciliae (Annelida: Phyllodocida) revisited: previously unknown characters and undescribed species from the Equatorial North Atlantic Mestrado em Biologia Marinha Supervisor Dr Regina Cunha Co-supervisor Dr Michelle Taylor UNIVERSIDADE DO ALGARVE Faculdade de Ciências e Tecnologia 2019 Declaração de autoria de trabalho Gorgoniapolynoe caeciliae (Annelida: Phyllodocida) revisited: previously unknown characters and undescribed species from the Equatorial North Atlantic. Declaro ser o autor deste trabalho, que é original e inédito. Autores e trabalhos consultados estão devidamente citados no texto e constam da listagem de referências incluída. _______________________________________ Jamie M. Maxwell A Universidade do Algarve reserva para si o direito, em conformidade com o disposto no Código do Direito de Autor e dos Direitos Conexos, de arquivar, reproduzir e publicar a obra, independentemente do meio utilizado, bem como de a divulgar através de repositórios científicos e de admitir a sua cópia e distribuição para fins meramente educacionais ou de investigação e não comerciais, conquanto seja dado o devido crédito ao autor e editor respetivos. Acknowledgments I would like thank Dr Regina Cunha for accepting me as a student and for her help and guidance throughout the project. Thank you to Dr Michelle Taylor, for inviting me to Essex University and suppling me with the specimens needed to complete this project. She welcomed me into her lab and provided much appreciated support, guidance, and some amazing BBQs. Next, I would like to thank Dr Sergio Taboada and Dr Ana Riesgo who hosted me in the Natural History Museum in London. -
Polychaete Worms Definitions and Keys to the Orders, Families and Genera
THE POLYCHAETE WORMS DEFINITIONS AND KEYS TO THE ORDERS, FAMILIES AND GENERA THE POLYCHAETE WORMS Definitions and Keys to the Orders, Families and Genera By Kristian Fauchald NATURAL HISTORY MUSEUM OF LOS ANGELES COUNTY In Conjunction With THE ALLAN HANCOCK FOUNDATION UNIVERSITY OF SOUTHERN CALIFORNIA Science Series 28 February 3, 1977 TABLE OF CONTENTS PREFACE vii ACKNOWLEDGMENTS ix INTRODUCTION 1 CHARACTERS USED TO DEFINE HIGHER TAXA 2 CLASSIFICATION OF POLYCHAETES 7 ORDERS OF POLYCHAETES 9 KEY TO FAMILIES 9 ORDER ORBINIIDA 14 ORDER CTENODRILIDA 19 ORDER PSAMMODRILIDA 20 ORDER COSSURIDA 21 ORDER SPIONIDA 21 ORDER CAPITELLIDA 31 ORDER OPHELIIDA 41 ORDER PHYLLODOCIDA 45 ORDER AMPHINOMIDA 100 ORDER SPINTHERIDA 103 ORDER EUNICIDA 104 ORDER STERNASPIDA 114 ORDER OWENIIDA 114 ORDER FLABELLIGERIDA 115 ORDER FAUVELIOPSIDA 117 ORDER TEREBELLIDA 118 ORDER SABELLIDA 135 FIVE "ARCHIANNELIDAN" FAMILIES 152 GLOSSARY 156 LITERATURE CITED 161 INDEX 180 Preface THE STUDY of polychaetes used to be a leisurely I apologize to my fellow polychaete workers for occupation, practised calmly and slowly, and introducing a complex superstructure in a group which the presence of these worms hardly ever pene- so far has been remarkably innocent of such frills. A trated the consciousness of any but the small group great number of very sound partial schemes have been of invertebrate zoologists and phylogenetlcists inter- suggested from time to time. These have been only ested in annulated creatures. This is hardly the case partially considered. The discussion is complex enough any longer. without the inclusion of speculations as to how each Studies of marine benthos have demonstrated that author would have completed his or her scheme, pro- these animals may be wholly dominant both in num- vided that he or she had had the evidence and inclina- bers of species and in numbers of specimens.