SERICINI Kirby, 1837
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Morphology, Taxonomy, and Biology of Larval Scarabaeoidea
Digitized by the Internet Archive in 2011 with funding from University of Illinois Urbana-Champaign http://www.archive.org/details/morphologytaxono12haye ' / ILLINOIS BIOLOGICAL MONOGRAPHS Volume XII PUBLISHED BY THE UNIVERSITY OF ILLINOIS *, URBANA, ILLINOIS I EDITORIAL COMMITTEE John Theodore Buchholz Fred Wilbur Tanner Charles Zeleny, Chairman S70.S~ XLL '• / IL cop TABLE OF CONTENTS Nos. Pages 1. Morphological Studies of the Genus Cercospora. By Wilhelm Gerhard Solheim 1 2. Morphology, Taxonomy, and Biology of Larval Scarabaeoidea. By William Patrick Hayes 85 3. Sawflies of the Sub-family Dolerinae of America North of Mexico. By Herbert H. Ross 205 4. A Study of Fresh-water Plankton Communities. By Samuel Eddy 321 LIBRARY OF THE UNIVERSITY OF ILLINOIS ILLINOIS BIOLOGICAL MONOGRAPHS Vol. XII April, 1929 No. 2 Editorial Committee Stephen Alfred Forbes Fred Wilbur Tanner Henry Baldwin Ward Published by the University of Illinois under the auspices of the graduate school Distributed June 18. 1930 MORPHOLOGY, TAXONOMY, AND BIOLOGY OF LARVAL SCARABAEOIDEA WITH FIFTEEN PLATES BY WILLIAM PATRICK HAYES Associate Professor of Entomology in the University of Illinois Contribution No. 137 from the Entomological Laboratories of the University of Illinois . T U .V- TABLE OF CONTENTS 7 Introduction Q Economic importance Historical review 11 Taxonomic literature 12 Biological and ecological literature Materials and methods 1%i Acknowledgments Morphology ]* 1 ' The head and its appendages Antennae. 18 Clypeus and labrum ™ 22 EpipharynxEpipharyru Mandibles. Maxillae 37 Hypopharynx <w Labium 40 Thorax and abdomen 40 Segmentation « 41 Setation Radula 41 42 Legs £ Spiracles 43 Anal orifice 44 Organs of stridulation 47 Postembryonic development and biology of the Scarabaeidae Eggs f*' Oviposition preferences 48 Description and length of egg stage 48 Egg burster and hatching Larval development Molting 50 Postembryonic changes ^4 54 Food habits 58 Relative abundance. -
An Annotated Checklist of Wisconsin Scarabaeoidea (Coleoptera)
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Center for Systematic Entomology, Gainesville, Insecta Mundi Florida March 2002 An annotated checklist of Wisconsin Scarabaeoidea (Coleoptera) Nadine A. Kriska University of Wisconsin-Madison, Madison, WI Daniel K. Young University of Wisconsin-Madison, Madison, WI Follow this and additional works at: https://digitalcommons.unl.edu/insectamundi Part of the Entomology Commons Kriska, Nadine A. and Young, Daniel K., "An annotated checklist of Wisconsin Scarabaeoidea (Coleoptera)" (2002). Insecta Mundi. 537. https://digitalcommons.unl.edu/insectamundi/537 This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. INSECTA MUNDI, Vol. 16, No. 1-3, March-September, 2002 3 1 An annotated checklist of Wisconsin Scarabaeoidea (Coleoptera) Nadine L. Kriska and Daniel K. Young Department of Entomology 445 Russell Labs University of Wisconsin-Madison Madison, WI 53706 Abstract. A survey of Wisconsin Scarabaeoidea (Coleoptera) conducted from literature searches, collection inventories, and three years of field work (1997-1999), yielded 177 species representing nine families, two of which, Ochodaeidae and Ceratocanthidae, represent new state family records. Fifty-six species (32% of the Wisconsin fauna) represent new state species records, having not previously been recorded from the state. Literature and collection distributional records suggest the potential for at least 33 additional species to occur in Wisconsin. Introduction however, most of Wisconsin's scarabaeoid species diversity, life histories, and distributions were vir- The superfamily Scarabaeoidea is a large, di- tually unknown. -
Coleoptera: Scarabaeidae)
Systematic Entomology (2005), 31, 113–144 DOI: 10.1111/j.1365-3113.2005.00307.x The phylogeny of Sericini and their position within the Scarabaeidae based on morphological characters (Coleoptera: Scarabaeidae) DIRK AHRENS Deutsches Entomologisches Institut im Zentrum fu¨r Agrarlandschafts- und Landnutzungsforschung Mu¨ncheberg, Germany Abstract. To reconstruct the phylogeny of the Sericini and their systematic position among the scarabaeid beetles, cladistic analyses were performed using 107 morphological characters from the adults and larvae of forty-nine extant scarabaeid genera. Taxa represent most ‘traditional’ subfamilies of coprophagous and phytophagous Scarabaeidae, with emphasis on the Sericini and other melo- lonthine lineages. Several poorly studied exoskeletal features have been examined, including the elytral base, posterior wing venation, mouth parts, endosternites, coxal articulation, and genitalia. The results of the analysis strongly support the monophyly of the ‘orphnine group’ þ ‘melolonthine group’ including phytopha- gous scarabs such as Dynastinae, Hopliinae, Melolonthinae, Rutelinae, and Cetoniinae. This clade was identified as the sister group to the ‘dung beetle line’ represented by Aphodius þ Copris. The ‘melolonthine group’ is comprised in the strict consensus tree by two major clades and two minor lineages, with the included taxa of Euchirinae, Rutelinae, and Dynastinae nested together in one of the major clades (‘melolonthine group I’). Melolonthini, Cetoniinae, and Rutelinae are strongly supported, whereas Melolonthinae and Pachydemini appear to be paraphyletic. Sericini þ Ablaberini were identified to be sister taxa nested within the second major melolonthine clade (‘melolonthine group II’). As this clade is distributed primarily in the southern continents, one could assume that Sericini þ Ablaberini are derived from a southern lineage. -
Quick Guide for the Identification Of
Quick Guide for the Identification of Maryland Scarabaeoidea Mallory Hagadorn Dr. Dana L. Price Department of Biological Sciences Salisbury University This document is a pictorial reference of Maryland Scarabaeoidea genera (and sometimes species) that was created to expedite the identification of Maryland Scarabs. Our current understanding of Maryland Scarabs comes from “An Annotated Checklist of the Scarabaeoidea (Coleoptera) of Maryland” (Staines 1984). Staines reported 266 species and subspecies using literature and review of several Maryland Museums. Dr. Price and her research students are currently conducting a bioinventory of Maryland Scarabs that will be used to create a “Taxonomic Guide to the Scarabaeoidea of Maryland”. This will include dichotomous keys to family and species based on historical reports and collections from all 23 counties in Maryland. This document should be cited as: Hagadorn, M.A. and D.L. Price. 2012. Quick Guide for the Identification of Maryland Scarabaeoidea. Salisbury University. Pp. 54. Questions regarding this document should be sent to: Dr. Dana L. Price - [email protected] **All pictures within are linked to their copyright holder. Table of Contents Families of Scarabaeoidea of Maryland……………………………………... 6 Geotrupidae……………………………………………………………………. 7 Subfamily Bolboceratinae……………………………………………… 7 Genus Bolbocerosoma………………………………………… 7 Genus Eucanthus………………………………………………. 7 Subfamily Geotrupinae………………………………………………… 8 Genus Geotrupes………………………………………………. 8 Genus Odonteus...……………………………………………… 9 Glaphyridae.............................................................................................. -
3.2.6 the Phylogeny of Xenoserica Gen. N
120 D. Ahrens 3.2.6 The phylogeny of Xenoserica gen. n. The Himalayan Sericine chafers have been revised extensively using almost all collection material available to the author (Ahrens 1995a – 2004c; Ahrens and Sabatinelli 1996). These studies, recently concluded and summarized by Ahrens (2004b), revealed that flight reduction did apparently not occur among Himalayan Sericini. This peculiarity, in contrast to many other Himalayan Coleoptera (Carabidae, Chrysomelidae) with numerous apterous or brachypterous forms (Mani 1968), was explained by the absence of Sericini in the alpine and nival zone. Recently, however, brachypterous individuals were collected in the alpine zone of the slopes of the Kali Gandaki Valley (Fig. 83). Material and methods Taxon sampling and characters Nineteen species belonging to eight genera of Sericini were included in the cladistic analysis, with Comaserica bergrothi chosen as the outgroup taxon due to their rather close relationship to the ingroup taxa, but with a high probability of not being part of the ingroup (chapter 3.1). Character coding was based on 22 species belonging to nine genera (see appendix A 3.2.6). The choice of taxa to be included into the ingroup was mainly based on present and historical classification of the species and genera of Sericini (e.g. Nomura 1976, Yu et al. 1998). Representatives of the Central Asian endemic genus Trochaloschema were also included because they represent the only previously known Asian sericine chafers having reduced wings. Thirty-six adult characters were scored. The character states are illustrated in Figs 84-86. Characters resulting directly from brachyptery were not considered in the analyses to exclude the possibility that environmentally driven adaptations (wing reduction, with a high probability of convergence could influence the results of the analysis. -
An Electronic Checklist of the New World Chafers (Coleoptera: Scarabaeidae: Melolonthinae)
AN ELECTRONIC CHECKLIST OF THE NEW WORLD CHAFERS (COLEOPTERA: SCARABAEIDAE: MELOLONTHINAE) Version 3 ARTHUR V. EVANS Research Associate, Department of Entomology, National Museum of Natural History, Smithsonian Institution, Washington, DC; Department of Recent Invertebrates, Virginia Museum of Natural History, Martinsville, VA; Department of Biology, Virginia Commonwealth University, Richmond, VA; c/o1600 Nottoway Ave., Richmond, VA 23227, USA; [email protected] and ANDREW B. T. SMITH Canadian Museum of Nature, P.O. Box 3443, Station D, Ottawa, ON, K1P 6P4, Canada; [email protected] INTRODUCTION The following is a checklist of all Melolonthinae (Coleoptera: Scarabaeidae) found in the New World. It has been modified from Evans (2003), Evans and Smith (2005), and Smith and Evans (2005) and has been updated to 13 March 2009. Included in this checklist are all of the available names given for New World Melolonthinae (both valid and invalid). Tribes are listed in traditional order (pseudo-phylogenetically) with genera, species, and subspecies listed alphabetically within. Under each valid generic name the subgenera and synonymies are listed as are type species and, in some cases, citations for keys, checklists, and bibliographies. Listed under each valid species are synonymies, distributional data by country, and citations for new combinations and spellings. A complete bibliography is included in the “References” section of all papers mentioned in the checklist. The purpose of this checklist is to present accurate and complete information for all the names of Melolonthinae in the New World. The taxonomy herein is based on the current literature (even if we have unpublished data contradicting what has been published) and the nomenclature carefully follows the International Code of Zoological Nomenclature. -
A New Record of the Genus Pachyserica
Short communication KOREAN JOURNAL OF APPLIED ENTOMOLOGY 한국응용곤충학회지 ⓒ The Korean Society of Applied Entomology Korean J. Appl. Entomol. 59(1) 37-40 (2020) pISSN 1225-0171, eISSN 2287-545X DOI: https://doi.org/10.5656/KSAE.2020.02.1.043 A new record of the Genus Pachyserica Brenske, 1897 (Coleoptera, Melolonthidae) in Korea A Young Kim* and Sang Wook Park1 National Institute of Biological Resources, Incheon 22689, Korea 1Research Institute of Forest Insect Diversity, Namyangju 12113, Korea 한국산 Pachyserica 속(딱정벌레목: 검정풍뎅이과)의 미기록종에 대한 보고 김아영*ㆍ박상욱1 국립생물자원관, 1산림곤충다양성연구소 ABSTRACT: We recognized Pachyserica yanoi Nomura which belongs to the genus Pachyserica Brenske, 1897 (Coleoptera, Melolonthidae) in Korea for the first time. A key to the genera of Korean Sericini, diagnosis and photographs of this species are provided. Key words: Pachyserica yanoi, Pachyserica, Melolonthidae, New record, Korea 초 록: 한국산 미기록속 비늘우단풍뎅이속(신칭)의 미기록종인 제주비늘우단풍뎅이(Pachyserica yanoi)를 보고한다. 검색어: 제주비늘우단풍뎅이, 비늘우단풍뎅이속, 검정풍뎅이과, 미기록, 한국 Tribe Sericini (Coleoptera, Melolonthidae) is a large tribe in over Asia (Ahrens, 2006; Löbl and Smetana, 2006). It is the family Melolonthidae including over 700 species in the seemed that the species in this genus are monophyly by their parleartic region (Löbl and Smetana, 2006). There were 5 phylogenetic analysis (Ahrens and Fabrizi, 2016). genera of Sericini distributed in Korea (Kim, 2011); The first Pachyserica species ‘P. yanoi Nomura, 1959’ Gastroserica, Maladera, Nipponoserica, Serica, Sericania. which was collected in Jeju island is reported in Korea. We Adding them, we report one more genus Pachyserica new to provide a key to the genera, diagnosis of the genus and species, Korea. -
Merrimac Farm WMA Insect List As of September 2014 Order Family
Merrimac Farm WMA Insect List as of September 2014 Order Family Common Name Scientific Name Acari Ixodidae American Dog Tick Dermacentor variabilis Araneae Anyphaenidae Ghost Spider Hibana sp. Araneae Araneidae Larinia directa Larinia directa Araneae Araneidae Star-bellied Orbweaver Acanthepeira stellata Araneae Araneidae White Micrathena Micrathena mitrata Araneae Araneidae Spined Micrathena Micrathena gracilis Araneae Lycosidae Wolf Spider Hogna sp. Araneae Lycosidae Thin-legged Wolf Spider Pardosa sp. Araneae Lycosidae Rabid Wolf Spider Rabidosa rabida Araneae Oxyopidae Lynx Spider Oxyopes aglossus Araneae Salticidae Jumping Spider Pelegrina proterva? Araneae Salticidae Jumping Spider Phidippus princeps Araneae Salticidae Jumping Spider Tutellina elegans Araneae Salticidae Peppered Jumper Pelegrina galathea Araneae Thomisidae Northern Crab Spider Mecaphesa asperata Araneae Thomisidae Swift Crab Spider Mecaphesa celer Araneae Thomisidae White-banded Crab Spider Misumenoides formosipes Blattodea Cryptocercidae Brown-hooded Cockroach Cryptocercus punctulatus Coleoptera Cantharidae Margined Leatherwing Chauliognathus marginatus Coleoptera Cantharidae Soldier Beetle Podabrus rugosulus Coleoptera Carabidae Vivid Metallic Ground Beetle Chlaenius sp. Coleoptera Carabidae Vivid Metallic Ground Beetle Chlaenius emarginatus Coleoptera Carabidae Six-spotted Tiger Beetle Cicindela sexguttata Coleoptera Cerambycidae Flower Longhorn Beetle Strangalia luteicornis Coleoptera Cerambycidae Locust Borer Megacyllene robiniae Coleoptera Cerambycidae Red -
Maladera Castanea Arrow 1913
Fact Sheet Maladera castanea Arrow 1913 Taxonomy Sub family: Melolonthinae / Tribe: Sericini / Genus: Maladera Distinguishing Features Small oblong shaped beetles, body length 7-11mm. Body colouration brown, reddish brown to Maladera castanea dorsal view orange, with a iridescent sheen. Setae and scales absent on elytra. Antennae with 10 segments Photographer: and a 3-segmented club. Clypeus with apex appearing rounded to subquadrate, heavily upturned. Pia Scanlon Labrum indistinct, fused to the apical margin of the clypeus. Mesocoxae widely separated, metasternal anterior border rounded medially and not projecting forward of coxae. Metacoxae greatly enlarged. Penultimate ventrite and propygidium separated by a suture. Metatibial apical spurs separated, so that tarsi can move between them. Tarsal claws with a median tooth. Related and Similar Species Maladera castanea lateral view Photographer: M. castanea is a member of the Sericini tribe which shares the following features: Pia Scanlon Generally small sized with a convex body shape. Labrum indistinct, attached to the front margin of the clypeus. Metacoxae large, projecting forward where metasternum would usually be, and commonly covering first two abdominal sternites. Widely separated mesocoxae. Metatibial spurs widely separated, so that the tarsi can pass between them. A key to Australian Sericini can be found in Britton (1957). A key to North American genera is provided by Ratcliffe, Jameson and Smith (2002). Maladera castanea ventral view Other members of Sericini dealt within the key are the North American/Oriental/Ethiopian Photographer: occurring Serica, and the native Australian Phyllotocus. Maladera castanea can be clearly separated Pia Scanlon from these Genera by having 10 antennal segments (Serica 8-9, Phyllotocus 8). -
An Electronic Checklist of the New World Chafers (Coleoptera: Scarabaeidae: Melolonthinae)
AN ELECTRONIC CHECKLIST OF THE NEW WORLD CHAFERS (COLEOPTERA: SCARABAEIDAE: MELOLONTHINAE) ARTHUR V. EVANS Research Associate, Department of Entomology, National Museum of Natural History, Smithsonian Institution, Washington, DC; Department of Recent Invertebrates, Virginia Museum of Natural History, Martinsville, VA; c/o1600 Nottoway Ave., Richmond, VA 23227, USA; [email protected]. and ANDREW B. T. SMITH Canadian Museum of Nature, P.O. Box 3443, Station D, Ottawa, ON, K1P 6P4, Canada; asmith@mus- nature.ca INTRODUCTION The following is a checklist of all Melolonthinae (Coleoptera: Scarabaeidae) found in the New World. It has been modified from Evans (2003) and Smith and Evans (2005) and has been updated to 1 September 2005. Included in this checklist are all of the available names given for New World Melolonthinae (both valid and invalid). Tribes are listed in traditional order (pseudo-phylogenetically) with genera, species, and subspecies listed alphabetically within. Under each valid generic name the subgenera and synonymies are listed as are type species and, in some cases, citations for keys, checklists, and bibliographies. Listed under each valid species are synonymies, distributional data by country, and citations for new combinations and spellings. A complete bibliography is included in the “References” section of all papers mentioned in the checklist. The purpose of this checklist is to present accurate and complete information for all the names of Melolonthinae in the New World. The taxonomy herein is based on the current literature (even if we have unpublished data contradicting what has been published) and the nomenclature carefully follows the International Code of Zoological Nomenclature. It is our intent to update this checklist periodically as errors are corrected and new papers are published requiring changes. -
Report of Astaena Pygidialis Kirsch (Coleoptera: Scarabaeidae)
Report of Astaena pygidialis Kirsch (Coleoptera: Scarabaeidae), the main chafer beetle causing damage to avocado fruit and young leaves in Antioquia Department, Colombia Claudia M. Holguin1,*, and Rosa H. Mira1 Abstract Scarab beetles (Coleoptera: Scarabaeidae) are one of the most limiting pests of avocado in Antioquia Department, Colombia. The establishment of management strategies to control these beetles has been difficult because the species causing damage to avocado fruit and young leaves have not been identified. Selected commercial avocado fields were surveyed at different times of d for scarab beetle adults affecting small fruit and young leaves, as well as other plant structures such as flowers, tender buds, stems, and mature leaves. Additionally, ultraviolet light traps were placed at each field to quantify the species captured. After direct scouting of avocado trees, the only scarab beetle species detected causing damage to fruit and young leaves at all sampling sites was Astaena pygidialis Kirsch (Coleoptera: Melolonthinae). Adults were observed chewing the green skin of small fruit causing striations on the fruit and skeletonization of young leaves by chewing the leaf tissue between veins on the upper surface. Other species observed include Anomala cincta Say (Coleoptera: Rutelinae), Charioderma xylina Blanchard, and Strigoderma sp. (all Coleoptera: Rutelinae) associated with flowers, Isonychus sp. (Coleoptera: Melolonthinae) with flower buds, Cyclocephala fulgurata Burmeister (Coleoptera: Dynastinae) with tender buds, stems and flowers, andAstaena valida Burmeister and Plectris pavida (Burmeister) (both Coleoptera: Melolonthinae) with mature leaves. The species detected through scouting also were captured with light traps. The dominant species captured with light traps was Phyllophaga obsoleta (Blanchard) (Coleoptera: Melolonthinae), which was not observed causing damage to avocado fruit or young leaves. -
New Species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae)
European Journal of Taxonomy 621: 1–20 ISSN 2118-9773 https://doi.org/10.5852/ejt.2020.621 www.europeanjournaloftaxonomy.eu 2020 · Ranasinghe S. et al. This work is licensed under a Creative Commons Attribution License (CC BY 4.0). Research article urn:lsid:zoobank.org:pub:CF9060EF-84DA-4195-81AE-25B06B69D0DF New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae) Sasanka RANASINGHE 1, Jonas EBERLE 2, Suresh P. BENJAMIN 3 & Dirk AHRENS 4, * 1, 2, 3, 4 Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 160, D-53113 Bonn, Germany. 3 National Institute of Fundamental Studies, Hantana Road, 20000 Kandy, Sri Lanka. * Corresponding author: [email protected]; [email protected] 1 Email: [email protected] 2 Email: [email protected] 3 Email: [email protected] 1 urn:lsid:zoobank.org:author:E3B80CA3-910F-4DC0-ACB1-7BBF1B89D923 2 urn:lsid:zoobank.org:author:3A048DFB-D6E1-4F17-9705-9B2EB2753B94 3 urn:lsid:zoobank.org:author:986CED51-9425-4CB6-84A4-A9464BB9207E 4 urn:lsid:zoobank.org:author:DEDCE5CF-AA11-4BBF-A2C6-D7C815019714 Abstract. In a recent project, extensive fieldwork was carried out in several parts of Sri Lanka to investigate scarab biodiversity. Here we present the first results of this survey and describe four new Sericini species: Selaserica athukoralai sp. nov., Neoserica dharmapriyai sp. nov., Maladera cervicornis sp. nov., M. galdaththana sp. nov. Further, new locality records for 20 already known species are given. The genitalia and the habitus of all new species are illustrated and photos of the habitats of the new species are given. Keywords. Beetles, chafers, Sri Lanka, taxonomy.