Journal of the Marine Biological Association of the United Kingdom, 2013, 93(1), 143–182. # Marine Biological Association of the United Kingdom, 2012 doi:10.1017/S0025315412000847 A checklist of turtle and whale barnacles (Cirripedia: Thoracica: Coronuloidea) ryota hayashi1,2 1International Coastal Research Center, Atmosphere and Ocean Research Institute, The University of Tokyo, 5-1-5, Kashiwanoha, Kashiwa-shi, Chiba 277-8564 Japan, 2Marine Biology and Ecology Research Program, Extremobiosphere Research Center, Japan Agency for Marine–Earth Science and Technology

A checklist of published records of coronuloid barnacles (Cirripedia: Thoracica: Coronuloidea) attached to marine vertebrates is presented, with 44 species (including 15 fossil species) belonging to 14 genera (including 3 fossil genera) and 3 families recorded. Also included is information on their geographical distribution and the hosts with which they occur.

Keywords: checklist, turtle barnacles, whale barnacles, Chelonibiidae, Emersoniidae, Coronulidae, Platylepadidae, host and distribution

Submitted 10 May 2012; accepted 16 May 2012; first published online 10 August 2012

INTRODUCTION Superorder THORACICA Darwin, 1854 Order SESSILIA Lamarck, 1818 In this paper, a checklist of barnacles of the superfamily Suborder BALANOMORPHA Pilsbry, 1916 Coronuloidea occurring on marine animals is presented. Superfamily CORONULOIDEA Newman & Ross, 1976 The systematic arrangement used herein follows Newman Family CHELONIBIIDAE Pilsbry, 1916 (1996) rather than Ross & Frick (2011) for reasons taken up in Hayashi (2012) in some detail. The present author Genus Chelonibia Leach, 1817 deems the subfamilies of the Cheonibiidae (Chelonibiinae, Chelonibia caretta (Spengler, 1790) Emersoniinae and Protochelonibiinae) proposed by Harzhauser et al. (2011), as well as those included of Ross & Lepas caretta Spengler, 1790: 185, plate 6, figure 5. Frick (2007, 2011), to be unnecessary because the family Astrolepas testudinaria Gray, 1825: 105. Chelonibiidae has only three genera (Chelonibia, Emersonius Balanus chelytrypetes Hinks, 1840: 333. and Protochelonibia), and they are ignored herein. Species Chelonobia caretta Darwin, 1854: 394, plate 14, figure 2. within each genus and the references within each species are Chelonibia caretta Pilsbry, 1916: 267, plate 63, figure 5, arranged alphabetically. The original naming combinations 5a.—Hayashi, 2012: 109, figures 2 & 15b; plate 1b. and known synonyms are listed and each species is entered under the current binomen, followed by the author’s name, date of publication and page number. The geographical distri- references butions are recorded for the hosts and their general geographi- Anderson, 1994: 297; Bacon, 1976: 34; Barnard, 1924: 93; cal occurence where possible (Table 1). The fossil species are Borradaile, 1903: 443; Broch, 1924a: 16; Bugoni et al., 2001: 7; indicated by † and fossil records of Coronuloidea (including Caine, 1986: 19; Daniel, 1955: 32; Darwin, 1854: 394; extant species) are listed by geological age and location Dawydoff, 1952: 129; Dobbs & Landry, 2004: 674; Farrapeira, (Table 2). When uncertainty regarding a species record 2010: 212; Frick & Ross, 2002: 10; Frick et al., 1998: 213; exists, the species has been recorded with a query and a note. 2003a: 9; 2003b: 19; 2011: 638; Fuller et al., 2010: 1743; The literature list at the end of this paper includes the com- Gittings et al., 1986: 3; Gonza´lez et al., 2012: 132; Gramentz, plete field of scientific papers regarding the superfamily 1988: 33; Gray, 1825: 105; Gruvel, 1905a: 266; Hayashi, 2012: Coronuloidea (including taxonomy, phylogeny, ecology, 109; Hincks, 1840: 333; Hiro, 1936a: 61; 1937a: 69; Holthius, biology, palaeontology and natural history) and some preteri- 1952: 77; Innocenti, 2006: 6; Jones, 2003: 483; 2004: 122; tions in the catalogue of species by Newman & Ross (1976) are 2010: 208; Jones et al., 1990: 11; 2000: 260; Kobayashi, 2000: also compensated for in this list. 91; Kolosva´ry, 1943: 99; Korschelt, 1933: 13; Limpus et al., 1983a: 192; Loop et al., 1995: 246; Matsuura & Nakamura, 1993: 1803; Monroe, 1981: 237; Monroe & Limpus, 1979: CHECKLIST 199; Mo¨rch, 1852: 67; Nilsson-Cantell, 1938: 14; Pfaller et al., 2006: 3; 2008: 1096; Pilsbry, 1916: 267; Rawson et al., 2003: Class MAXILLOPODA Dahl, 1956 2699; Ritchie, 1924: 166; Ross, 1963b: 223; Ross & Frick, Subclass CIRRIPEDIA Burmeister, 1834 2011: 61; Scha¨rer, 2003: 88; Spengler, 1790: 185; Stubbings, 1967: 297; Utinomi, 1969: 92; Wells, 1966: 84; Weltner, 1897:

Corresponding author: 254; 1910: 528; Wirtz et al., 2006: 209; Withers, 1928a: 391; R. Hayashi 1929: 567; Young, 1991: 191; 1998: 271; Zullo, 1963: 13; 1982: Email: [email protected] 2; Zullo & Bleakney, 1966: 164.

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Table 1. Modern species of Coronuloidea indicating the host species and its collection locality. Locality records are arranged from Asia, Oceania, Indies, Mediterranean, Europe, Middle East, Africa, Atlantic, North America, South America to Pacific area.

Species Host Locality References

Chelonibia caretta (Spengler, 1790) Loggerhead turtle, Caretta caretta Yomitan, Okinawa, Japan Hayashi, 2012 (Linnaeus, 1758) Queensland, Australia (Heron Island; Mon Repos Beach; Monroe & Limpus, 1979; Jones, 2010 North Reef; Wistari Reef) Ouddorp, The Netherlands Holthuis, 1952 Alagadi Beach, Turkish Republic of Northern Cyprus Fuller et al., 2010 North-east Atlantic, Madeira, Portugal Wirtz et al., 2006 Scotland Ritchie, 1924 Greece (Zacharo; Zakynthos); Italy (Lampedusa) Gramentz, 1988 Cape of Good Hope, South Africa Pilsbry, 1916 Around Azores Frick et al., 2003b USA (Delaware Bay, New Jersey; Wassaw Island, Georgia; Pilsbry, 1916; Zullo & Bleakney, 1966; Wells, 1966; Caine, Cape Cod, Massachusetts; off the south-west pass of the 1986; Gittings et al., 1986; Frick et al., 1998; 2000; Frick Mississippi River Delta; North Padre Island, Texas; & Ross, 2002; Rawson et al., 2003; Pfaller et al., 2006; Canaveral National Sea Shore, Florida; Daytona Beach, Pfaller et al., 2008 Florida; St Teresa, Florida) Toco Depot, Trinidad and Tobago Bacon, 1976 Brazil Pilsbry, 1916; Bugoni et al., 2001 No locality records Ernst & Barbour, 1972 Green turtle, Chelonia mydas Off Madras Coast, India Daniel, 1955 (Linnaeus, 1758) Alagadi Beach, Turkish Republic of Northern Cyprus Fuller et al., 2010 South of Kharg, Iran Utinomi, 1969 Ile Europa, Mozambique, Mozambique Weltner, 1910 Table Bay, South Africa Barnard, 1924 Mauretania Broch, 1924b; Stubbings, 1967 USA (off the south-west pass of the Mississippi River Gittings et al., 1986 Delta; North Padre Island, Texas; St Teresa, Florida) Hawksbill turtle, Eretmochelys Yomitan, Okinawa, Japan Hayashi, 2012 imbricata (Linnaeus, 1766) Ream, Cambodia Dawydoff, 1952 Australia (Boydong Island; Wistari Reef, Queensland; Monroe & Limpus, 1979; Limpus et al., 1983; Loop et al., Campbell Island; Milman Island, Great Barrier Reef; 1995; Dobbs & Landry, 2004; Jones, 2004; Jones, 2010 Dampier Archipelago) Republic of Palau Hiro, 1937a Minikoi Island, India Borradaire, 1903 Cuba Kobayashi, 2000 (as Chelonibia sp.) Mona Island, Puerto Rico Scha¨rer, 2003 Antigua Frick et al., 2003a C. mydas or E. imbricata Tropical Atlantic; Indo-West Pacific Jones et al., 2000 Unidentified sea turtle, Chelonia sp. Saigon, Viet Nam Pilsbry, 1916 No host records Torres Strait Weltner, 1897; Pilsbry, 1916 Massana, New Guinea Weltner, 1897 Northern and Western Australia Darwin, 1854; Jones, 2003 Maldives Nilsson-Cantell, 1938 Mediterranean Kolosvary, 1943 Massaua, Hagere Ertra Pilsbry, 1916 West coast of Africa Darwin, 1854 Cape of Good Hope, South Africa Pilsbry, 1916 St Thomas, United States Virgin Islands Pilsbry, 1916 Venezuela Pilsbry, 1916 Chelonibia manati Gruvel, 1903 African manatee, Trichechus Joal, Senegal Stubbings, 1965 senegalensis Link, 1795 Congo coast Gruvel, 1903; Pislbry, 1916 Caribbean manatee, Trichechus Caribbean Sea, Mexico Morales-Vera et al.,2008 manatus Linnaeus, 1758 Puerto Rico Mianucci-Giannoni et al., 1999 Green turtle, Chelonia mydas Atlantic Korschelt, 1933 (as Chelonibia ramosa) (Linnaeus, 1758) Diamondback terrapin, Malaclemys Merritt Island, Florida, USA Seigel, 1983 terrapin (Schoepf, 1793) No host records Texas, USA Gittings et al., 1986 Chelonibia manati crenatibasis Pilsbry, 1916 Probably from loggerhead turtle, No locality record Pilsbry, 1916 Caretta caretta (Linnaeus, 1758) Chelonibia manati lobatobasis Pilsbry, 1916 Loggerhead turtle, Caretta caretta Florida, USA (Osprey; Bald Point) Pilsbry, 1916; Wells, 1966 (Linnaeus, 1758) Diamondback terrapin, Malaclemys Merritt Island, Florida, USA Seigel, 1983 terrapin (Schoepf, 1793) Chelonibia patula (Ranzani, 1818) Blue crab, Callinectes sapidus Turkey, Mediterranean Bakır et al.,2010 Rathbun, 1896 Cameroon Pilsbry, 1916 Gulf of Mexico Gittings et al., 1986; Spears et al., 1994

USA (Beaufort, North Carolina; Delaware Bay, Delaware; Pilsbry, 1916; McDougall, 1943; Pearse, 1947; William & checklistofturtleandwhalebarnacles 145 Virginia; Tarpon; Alligator Harbor; Seahorse Key) Porter, 1964; Wells, 1966; Crisp, 1983; Gittings et al., 1986; Gannon, 1990; Spears et al., 1994; Key et al., 1997 Sharptooth swim crab, Callinectis Off Landing Point, Gambia Stubbings, 1967 marginatus (Milne-Edwards, 1861) Blue crab, Callinectes danae Smith, Ubatsuba Bay, Sa˜o Paulo, Brazil Mantelatto et al.,2003 1869 Blue crab, Callinectes ornatus Ordway, Ubatsuba Bay, Sa˜o Paulo, Brazil Mantelatto et al.,2003 1863 Blue crab, Callinectes amnicola Off Lagos Coast, Nigeria Lawel-Are & Daramola, 2010 (Rochebrune, 1883) Speckeld swimming crab, Arenaeus Ubatsuba region, Sa˜o Paulo, Brazil Costa et al., 2010 cribrarius (Lamarck, 1818) Portunid crab, Portunus pelagicus Australia (Moreton Bay; North Reefs, Queensland; Phillips & Cannon, 1978; Shields, 1992; Gaddes & (Linnaeus, 1758) Dampier Archipelago) Sumpton, 2004; Jones, 2004; Jones, 2010 Karachi, Pakistan Javed & Mustaquim, 1994 Mediterranean coast of Israel Pasternak et al., 2002 Smooth swim crab, Portunus validus Off Lagos coast, Nigeria Lawel-Are & Daramola, 2010 (Herklots, 1851)

Continued 4 yt hayashi ryota 146

Table 1. Continued

Species Host Locality References

Siera Leone Stubbings, 1967 Swimming crab, Portunus spinimanus Ubatsuba region, Sa˜o Paulo, Brazil Santos, 2002 Latreille, 1819 Portunid crab, Charybdis hellerii (A. Karachi, Pakistan Javed & Mustaquim, 1994 Milne-Edwards, 1867) Atlantic horseshoe crab, Limulus USA (south-eastern and Gulf coasts; Beaufort, North McDougall, 1943; Wells, 1966; Ross & Jackson, 1972; polyphemus (Linnaeus, 1758) Carolina; Alligator Harbor, Florida) Gittings et al., 1986 Mud crab, Scylla serrata (Forska˚l, Moreton Bay, Queensland, Australia Hudson & Lester, 1994 1775) Madras, Lake Pulicat, India Nilsson-Cantell, 1938 Asian shore crab, Hemigrapsus New Jersey, USA McDermott, 1998, 1999 sanguineus (De Haan, 1835) Spider crab, Libinia dubia H. Milne Attached to the carapace of Caretta caretta, Georgia, USA Frick et al., 2004 Edwards, 1834 Long-eyed swimming crab, Hepingdao, Keelung City, Taiwan Chan et al., 2009 Podophthalmus vigil (Fabricius, 1798) Spotted box crab, Calappa Ghana Stubbings, 1967 rubroguttata Herklots, 1851 Unidentified sand crab Stradbroke, Australia Jones, 2010 On the antenna of crawfish Off Ghana Stubbings, 1961 Sea snail, Cymbium sp. Ghana Stubbings, 1967 Diamondback terrapin, Malaclemys Shired Island, California Swamp, Dixie County, Florida, Ross & Jackson, 1970 terrapin macrospilota (Hay, 1904) USA Loggerhead turtle, Caretta caretta Aegean Sea, Greece Kitsos et al., 2003, 2005 (Linnaeus, 1758) Unidentified sea turtles Tropical Atlantic; Indo-West Pacific Jones et al., 2000 A human femur Moreton Bay, Moreton Island, Australia Monroe, 1981 Some artificial materials West Peloponnesus, Greece Frazier & Margaritoulis, 1990 Nigeria (Lagos; Bonney River) Stubbings, 1967 Blue crab, Callinectis sp. Brazil (Sa˜o Paulo; Paraiba) Young, 1991 Rio Bayamon above Palo Seco, Puerto Rico Pilsbry, 1916 No host records Australia Darwin, 1854; Jones, 2003 Mediterranean Darwin, 1854 Gambia, west coast of Africa Darwin, 1854; Stubbings, 1967 USA (Charleston, South Carolina; Texas; Cedar Keys, Darwin, 1854; Pilsbry, 1916; Johnson & Bonner, 1960 Florida; North Carolina) Tampico, Tamaulipas, Mexico Celis et al., 2007 Cape Cajon, Cuba Pilsbry, 1916 Jamaica; Honduras; Brazil Darwin, 1854 Chelonibia patula dentata Henry, 1943 Warrior swimming crab, Callinectes Estero de Agiabampo, Sonora, Mexico Henry, 1943 bellicosus Stimpson, 1859 No host records China Jones et al., 2000 Chelonibia testudinaria (Linnaeus, 1758) Loggerhead turtle, Caretta caretta Japan (Otsuchi, Iwate; Tateyama, Chiba; Seto, Wakayama; Hiro, 1937b; Utinomi, 1949; Matsuura & Nakamura, (Linnaeus, 1758) Senri Beach, Wakayama; Hakata Bay, Fukuoka; Fukiage 1993; Rawson et al., 2003; Hayashi, 2009, 2012 Beach, Kagoshima; Yakushima Island, Kagoshima; Yomitan, Okinawa; Kunigami, Okinawa; Ginoza, Okinawa) Queensland, Australia (Mon Repos; Heron Island) Monroe & Limpus, 1979; Jones, 2010 Koycegiz-Dalyankoy, Turkey Geldiay et al., 1995 Malta Graments, 1988 Rovinj, Croatia Zavodnik, 1997 Adriatic, Ionian, Sicily Channel and western Basin, Italy Casale et al., 2004 Greece (Zakynthos; Kyparissia; Peloponnesus Island) Margaritoulis, 1985; Gramentz, 1988; Rawson et al., 2003 Aegean Sea Koukouras & Matsa, 1998; Kitsos et al., 2003, 2005 Aldabra Atoll Frazier, 1971 Hann, Saloum River, Senegal Stubbings, 1965 Gran Canaria, Atlantic Gonza´lez et al., 2012 Around Azores Frick et al., 2003b Northern Gulf of Mexico Gittings et al., 1986 USA (Chesapeake Bay, Virginia; New Jersey; Wassaw Pilsbry, 1916; Richards, 1930; Zullo, 1963; Wells, 1966; Island, Georgia; St Teresa, Bald Point, Florida; St Mary’s Guess, 1982; Killingley & Lutcavage, 1983; Lutcavage & River Entrance Channel, Florida; Keewaydin, Florida; Musik, 1985; Plotkin, 1996; Frick et al., 1998; Frick & Canaveral National Sea Shore, Florida; Alligator Slay, 2000; Rawson et al., 2003; Frick et al., 2004; Harbor, Florida; Mustang Island, North and South Stamper et al., 2005; Pfaller et al., 2006; Pfaller et al., Padre Islands, Texas; Pamlico, North Carolina; Core 2008 Sounds, North Carolina; Newport, Rhode Island; La Jolla, California) Bahia Magdalena, Baja California, Mexico Pilsbry, 1916; Ross & Newman, 1967; Rawson et al., 2003; Point Patuca, Honduras Pilsbry, 1916 Toco Depot, Trinidad and Tobago Bacon, 1976 Brazil Bugoni et al., 2001; Pereira et al., 2006 checklistofturtleandwhalebarnacles 147 No locality records Ernst & Barbour, 1972 Green turtle, Chelonia mydas Japan (Ginoza, Okinawa; Yomitan, Okinawa; Ogasawara, Tachikawa, 1995; Hayashi & Tsuji, 2008; Hayashi, 2012 (Linnaeus, 1758) Tokyo) Poulo Condore, Viet Nam Dawydoff, 1952 Basar Island, Sarawak, Malaysia Hendrickson, 1958 Jinyindao, Xisha Islands, South China Sea Ren, 1980 Australia (Enoe Island, Aroe Archipelago; Islands in Nilsson-Cantell, 1930b; Monroe & Limpus, 1979; Torres Strait; Heron Island, Queensland; Moreton Bay, Glazebrook & Campbell, 1990; Limpus et al., 1994; Queensland; Shoalwater Bay, Queensland; Wynnum, Flint et al., 2009; 2010; Jones, 2010 Queensland) Kaisyaru, Baberudasbu Island, Republic of Palau Hiro, 1937a India (Minikoi Island; Drusadai Island; Quilon; Bet Borradaire, 1903; Daniel, 1958; Pillai, 1958; Frazier, 1989 Dwarka Island, Gulf of Kutch) Sandpit Beach, Pakistan Mustaquim & Javed, 1993 Turtuguero, Costa Rica Biasatti, 2004 South of Kharg, Iran Utinomi, 1969 Accra, Ghana Gauld, 1957 Aldabra Atoll Frazier, 1971

Continued 4 yt hayashi ryota 148

Table 1. Continued

Species Host Locality References

Senegal Stubbings, 1965 USA (Cape Canaveral, Florida; Daytona Beach, Florida; Caine, 1986; Gittings et al., 1986 northern Gulf of Mexico) Mexico (Pragreso, Yucatan; Teopa Beach, Jalisco State) Ives, 1891; Lazo-Wasem et al., 2011 Brazil Bugoni et al., 2001; Pereira et al., 2006 Peru Brown & Brown, 1995 Hawaiian Islands Aguirre et al., 1994; Balazs, 1978; Edmonson, 1946; Edmonson & Ingram, 1939; Eldridge & Miller, 1997; Zaldus & Hadfield, 2004 Hawksbill turtle, Eretmochelys Bentota, Ceylon Deraniyagala, 1939; Nilsson-Cantell, 1932 imbricata (Linnaeus, 1766) Australia (Islands in Torres Striat; Dampier Archipelago) Glazebrook & Campbell, 1990; Jones, 2004 Republic of Palau Hiro, 1937a India (Drusadai Island; Bombay) Daniel, 1958; Wagh & Bal, 1974 Rames Waren Island, Gulf of Manaar Annandale, 1906 Rovigno, d’Istria, Adriatic Kolosvary, 1939 Jumby Bay, Antigua Frick et al., 2003b Mexico (Puerto Penasco, Sonora; Rio Lagartos, Yucata´n) Young & Ross, 2000; Ca´rdenas-Palomo & Maldonado-Gasca, 2005 Olive ridley turtle, Lepidochelys Toyama Bay, Japan Hiro, 1939 olivacea (Eschscholtz, 1829) Mexico (La Gloria, Jalisco; Playa Ceuta, Sinaloa; Teopa Hernandez-Vazquez & Valadez-Gonzalez, 1998; Beach, Jalisco) Angulo-Lozano et al., 2007; Lazo-Wasem et al., 2011 Kemp’s ridley turtle, Lepidochelys Beaufort, North Carolina, USA Walker, 1978 kempi (Garman, 1880) Northern Gulf of Mexico Gittings et al., 1986 No locality records Rudloe et al., 1991 Flatback turtle, Natator depressus Queensland, Australia (Mon Repos; Curtis Island) Monroe & Limpus, 1979; Jones, 2010 (Garman, 1880) Leatherback turtle, Dermochelys Porth Neigwl Beach, Gwynedd, North Wales, United Rees & Walker, 1993 coriacea (Vandelli, 1761) Kingdom Diamondback terrapin, Malaclemys Merritt Islands, Florida, USA Seigel, 1983 terrapin (Schoepf, 1793) Saltwater crocodile, Crocodylus Cape York Peninsula, Queensland, Australia Monroe & Garrett, 1979 porosus Schneider, 1801 American crocodile, Crocodylus Naval authorities in Puerto Vallarta harbor, Mexico Cupul-Magan˜a et al., 2011 acutus Cuvier, 1807 American alligator, Alligator Guana Lake, Ponte Vedra, Florida, USA Nifong & Frick, 2011 mississippiensis (Daudin, 1802) No host records Yamagata, Japan (Ana; Kamo, Yamagata; Nezugasaki, Pilsbry, 1916; Broch, 1931; Utinomi, 1970 Yamagata; Nagasaki) Dong-ao, Yilan County, Taiwan Chan et al., 2009 Koh Mak, Gulf of Siam, South China Sea Broch, 1931 Kota Bharu, Kelantan, Malaysia Lanchester, 1902 Australia (north-east and western coast; Broom) Darwin, 1854; Broch, 1916; Jones, 2003 Low Archipelago, French Polynesia Darwin, 1854 Ile Lifou, New Caledonia Fischer, 1884 Bay of Bengal MacDonald, 1929 India (Andaman Island; Madras; Mouth of Ganges) Nilsson-Cantell, 1938 Mediterranean Darwin, 1854 West coast of Africa Darwin, 1854; Jones, 2003 Cameroons; Ghana; Senegal Nilsson-Cantell, 1921; Stubbings, 1967 USA (Delaware Bay, New Jersey; Point Pleasant, New Pilsbry, 1916 Jersey; Beaufort, North Carolina; Key West, Florida; Pensacola, Florida; Gulf of California) Progreso, Yucatan, Mexico Pilsbry, 1916 Cape Frio, Brazil Pilsbry, 1916 California; west coast of Mexico Darwin, 1854; Pilsbry, 1916 Hawaiian Islands Pilsbry, 1916 Carolina Island, Republic of Kiribati Pilsbry, 1916 Albemarle Island, Galapagos Islands, Equador Pilsbry, 1916 Chelonibia sp. Hawksbill turtle, Eretmochelys Palau Guisaan, Sabah, Malaysia Chan et al., 1999 imbricata (Linnaeus, 1766) Caribbean Sea Carr et al.,1966 Black turtle, Chelonia mydas agassizii Baja California, Mexico Nichols et al., 2001 Bocourt, 1868 Loggerhead turtle, Caretta caretta Seychelles Frazier, 1971 (Linnaeus, 1758) Greece Frazier et al., 1985 South Africa Hughes, 1974 Coronula diadema (Linnaeus, 1767) Humpback whale, Megaptera Japan (Chikura, Chiba; Tateyama, Chiba; Rumoi, Nogata & Matsumura, 2006; Hayashi, 2012 novaeangliae Borowski, 1781 Hokkaido; Tosa-Shimizu, Kochi) Fraser Island, Queensland, Australia Paterson & Van Dyck, 1991; Jones, 2010

New Zealand Borradaire, 1916 checklistofturtleandwhalebarnacles 149 Ile Lifou, New Caledonia Fischer, 1884 Ireland (Killala Bay; Galway; Enniscrone) O’Riordan, 1979 Iceland Stephansen, 1938 Katwijk, The Netherlands Holthuis & Fransen, 2004 South Shetland Island, Scotland Nilsson-Cantell, 1930c Durban, South Africa Nilsson-Cantell, 1930c Canada (Cachalot, Vancouver, British Columbia; Snooks Pilsbry, 1916; Cornwall, 1924 Arm, Newfoundland) Brazil (Rio de Janeiro; Sa˜o Paulo) Young, 1991 South Atlantic Nilsson-Cantell, 1939b South Georgia and the South Sandwich Islands Nilsson-Cantell, 1930c; 1930d Bering Strait to the Gulf of California Dall, 1872; Pilsbry, 1916 Parque Nacional Natural Gorgona; Bahı´aMa´laga, Jime´nez et al.,2011 Colombian Pacific Southern right whale, Eubalaena Southern hemisphere Barnard, 1924 australis (Desmoulins, 1822)

Continued 5 yt hayashi ryota 150

Table 1. Continued

Species Host Locality References

Blue whale, Balaenoptera musculus Southern hemisphere Nilsson-Cantell, 1978 (Linnaeus, 1758) South Atlantic Nilsson-Cantell, 1939b Fin whale, Balaenoptera physalus North-east Atlantic, Madeira Wirtz et al., 2006 (Linnaeus, 1758) Sperm whale, Physeter catodon Southern hemisphere Hiro, 1935 Linnaeus, 1758 Northern bottlenose whale, Faroe Islands Nilsson-Cantell, 1978 Hyperoodon ampullatus (Flower, 1882) No host records Guei Shan, Yilan County, Taiwan Chan et al., 2009 Western Australia Jones, 2003 New Zealand (Waikouaiti; Cavalli Island) Darwin, 1854; Hutton, 1879; Filhol, 1885; Stead, 1903; Jennings, 1918 Tonga Island Pilsbry, 1916 Arctic Seas; Great Britain; USA Darwin, 1854 Shetland Islands, Scotland Pilsbry, 1916 Godhaven, Disco Island, Greenland Pilsbry, 1916 Nova Scotia, Canada Pilsbry, 1916 Gulf Stream Darwin, 1854 Territorial collectivity of Saint Barthe´lemy, West Indies Pilsbry, 1916 USA (Neah Bay, Washington; Mouth of Umpqua River, Pilsbry, 1916 Oregon; Unalaska; Magdalena Bay, California; Monterey, California; Neat Pigeon Point, California) Bering Sea Pilsbry, 1916 Coronula reginae Darwin, 1854 Humpback whale, Megaptera Lonstrup, Denmark; Varangerfjorden, Norway Nilsson-Cantell, 1978 novaeangliae Borowski, 1781 Iceland Stephansen, 1938; Nilsson-Cantell, 1978 Durban, South Africa Nilsson-Cantell, 1930c Canada (Snooks Arm, Newfoundland; Cachalot, Pilsbry, 1916; Cornwall, 1927 Vancouver, British Columbia) Rio Grande, Brazil Young, 1991 Blue whale, Balaenoptera musculus Lonstrup, Denmark; Varangerfjorden, Norway; Iceland Nilsson-Cantell, 1978 (Linnaeus, 1758) South Africa (Saldanha Bay; Durban) Nilsson-Cantell, 1930c South Georgia and the South Sandwich Islands Nilsson-Cantell, 1930c, 1939b Antarctic Tomilin, 1957 Fin whale, Balaenoptera physalus Lonstrup, Denmark; Varangerfjorden, Norway; Iceland Nilsson-Cantell, 1978 (Linnaeus, 1758) Saldanha Bay, South Africa Nilsson-Cantell, 1930c Sei whale, Balaenoptera borealis Lonstrup, Denmark; Varangerfjorden, Norway; Iceland Nilsson-Cantell, 1978 Lesson, 1828 Minke whale, Balaenoptera Antarctic Bushuev, 1990 acutorostrata Lace´pe`de, 1804 Loggerhead turtle, Caretta caretta Gulf of California Caldwell, 1963 (Linnaeus, 1758) No locality records Ernst & Barbour, 1972 No host records USA (Mouth of Umpqua River, Oregon; Unalaska, Pilsbry, 1916 Alaska) Iquique, Peru Darwin, 1854 Chile Darwin, 1854 Coronula sp. Humpback whale, Megaptera Carribean Sea Mianucci-Giannoni et al., 1999 novaeangliae Borowski, 1781 Cetopirus complanatus (Mo¨rch, 1852) Southern right whale, Eubalaena South Africa (Simon’s Town; Table Bay) Barnard, 1924 australis (Desmoulins, 1822) No host records Western Australia Jones, 2003 Southern Ocean Darwin, 1854 Cryptolepas rhachianecti Dall, 1872 Gray whale, Eschrichtius robustus Tomiyama, Chiba, Japan Takeda & Ogino, 2005 Lilljeborg, 1861 Russia (Sakhalin Island, Sea of Okhotsk; adjacent region) Weller et al., 1999; Sokolov & Arsen’ev, 2007 USA (Monterey Bay, California; Refugio Beach State Park, Dall, 1872; Briggs & Morejohn, 1972; Wellington & Santa Barbara, California; La Jolla, California; Anderson, 1978; Achituv, 1998 San Francisco) Baja California, Mexico Pilsbry, 1916; Kasuya & Rice, 1970; Killingley, 1980; Swartz, 1981 Bering Sea to Lower California; Hawaiian Islands Pilsbry, 1916 Killer whale, Orcinus orca Linnaeus, Bolsa Chica State Beach, California Samaras, 1989 1758 White whale, Delphinapterus leucas San Diego Bay, California, USA Ridgway et al., 1997; Maldini et al., 2010 (Pallas, 1776) Gut contents of Atherinops affinis Baja California, Mexico Swartz, 1981 Ayres, 1860

Tubicinella cheloniae Monroe & Limpus, 1979 Loggerhead turtle, Caretta caretta Japan (Yomitan, Okinawa; Yakushima Island, Kagoshima) Hayashi, 2012 checklistofturtleandwhalebarnacles 151 (Linnaeus, 1758) Mon Repos, south-east Queensland, Australia Monroe & Limpus, 1979 Green turtle, Chelonia mydas Talong Talong, Besar Islands, Sarawak, Malaysia Hendrickson, 1958 (as Stephanolepas muricata) (Linnaeus, 1758) Queensland, Australia (Moreton Bay; Wynnum; Monroe & Limpus, 1979; Flint et al., 2009, 2010; Jones, Shoalwater Bay) 2010 Hawksbill turtle, Eretmochelys Bentota, Ceylon, Sri Lanka Nilsson-Cantell, 1932 (as Stephanolepas muricata) imbricata (Linnaeus, 1766) Milman Island, Great Barrier Reef, Australia Dobbs & Landry, 2004 Olive ridley turtle, Lepidochelys No locality records Zullo, 1991 olivacea (Eschscholtz, 1829) No host records Western Australia Jones, 2003 Tubicinella major Lamarck, 1802 Southern right whale, Eubalaena False Bay, South Africa Marloth, 1900 australis (Desmoulins, 1822) Walker Bay (Gansbaai), Struisbaai, De Hoop Marine Reeb et al., 2007 Reserve, St Sebastian Bay (all on the south coast of South Africa)

Continued 5 yt hayashi ryota 152

Table 1. Continued

Species Host Locality References

Unidentified whales New South Wales, Australia Darwin, 1854 Southern Pacific Ocean Darwin, 1854 Faroe Islands Zullo, 1963 South Georgia and the South Sandwich Islands Nilsson-Cantell, 1921 Cape of Good Hope, South Africa Darwin, 1854 Western South America Darwin, 1854 Xenobalanus globicipitis Steenstrup, 1852 Long-finned pilot whale, Globicephala Faroe Islands Steenstrup, 1852; Hoek, 1883; Weltner, 1897; Pilsbry, melas Traill, 1809 1916; Nilsson-Cantell, 1921, 1978; Rappe´ &Van Waerebeek, 1988 Monaco Gruvel, 1912 Near the Azores between Madeira and England Pilsbry, 1916 Gibraltar Gruvel, 1920; Richard, 1936 Massachusetts, USA (North Dennis; Woods Hole) Pilsbry, 1916 Northern Atlantic Cornwall, 1927 Short-finned pilot whale, Gran Canaria, Atlantic Gonza´lez et al., 2012 Globicephala macrorhynchus Gray, 1846 Bottlenose dolphin, Tursiops Moreton Bay, Queensland, Australia Orams & Schuetze, 1997 truncatus Montagu, 1821 India (Parangipetti coast; Bay of Bengal; Porto Nova) Rajaguru & Shantha, 1992; Karuppiah et al.,2004 Gibraltar Dollfus, 1968; Pilleri, 1970 Natal, South Africa Barnard, 1924; Ross, 1984 Namibe Province, Angola Weir, 2010 USA (Central Atlantic coast; North Carolina; New Jersey; True, 1890; Gittings et al., 1986; Mead & Potter, 1990; Galveston Ship Channel, Texas; Santa Rosa Sound, Fertl, 1994; Watson & Gee, 2005; Toth-Brown & Hohn, Pensacola, Florida; Alligator Point, Florida; Santa 2007; Rostein et al., 2009; Bearzi & Patonai, 2010; Monica Bay, California) Pugliese et al., in press Gulf of Mexico Spivey, 1980 Rio de Janeiro, Brazil Di Beneditto & Ramos, 2000 Indian bottlenose dolphin, Tursiops Mikurajima Island, Tokyo, Japan Sakai et al.,2006 aduncus (Ehrenberg, 1833) Pantropical striped dolphin, Stenella Taiji, Wakayama, Japan Greenwood et al., 1979 attenuata (Gray, 1846) Mpelane, Mozambique Ross, 1984 Striped dolphin, Stenella coeruleoalba Western coast of Mediterranean Raga & Carbonell, 1985 (Meyen, 1833) Spanish coast, Mediterranean Raga et al., 1982; Raga & Carbonell, 1985; Duignan et al., 1992; Resendes et al., 2002 Mallorca Rappe´ & Van Waerebeek, 1988 Strait of Gibraltar Pilleri, 1970 Cape Hatteras, North Carolina, USA Mead, 1975 Brazil Carvalho et al., 2010; Ribeiro et al., 2010 Spinner dolphin, Stenella longirostris Eastern Pacific Kane et al., 2008 (Gray, 1828) Eastern spinner dolphin, Stenella Entrance of California Bay, Mexico Kane et al., 2008 longirostris orientalis Perrin, 1990 Cuvier’s beaked whale, Ziphius North Carolina, USA Bane & Zullo, 1980 cavirostris G. Cuvier, 1823 Isabera Island, Galapagos Islands, Equador Palacios et al.,2004 Franciscana dolphin, Pontoporia Urguay Danielewicz et al., 2002; Pinedo et al., 1989 blainvillei Gervais & d’Orbigny, 1844 Rio de Janeiro, Brazil Di Beneditto & Ramos, 2000, 2001 Killer whale, Orcinus orca Linnaeus, Off Kii Peninula, Japan Sakai et al.,2009 1758 Mediterranean Gruvel, 1920 Monaco; Gibaltar Richard, 1936 Golfo Dulce, Costa Rica Fertl et al., 1996 Bolsa Chica State Beach, California Samaras, 1989 Gulf of California, Mexico Guerrero-Ruiz & Urba´n, 2000 False killer whale, Pseudorca Spanish coast, Mediterranean Pilleri, 1967 crassidens (Owen, 1846) Monaco Gruvel, 1912 Miguel, Azores Gruvel, 1920; Richard, 1936 California, USA Caldwell et al.,1970 Tucuxi dolphin, Sotalia fluviatilis Rio de Janeiro, Brazil Di Beneditto & Ramos, 2000, 2004 (Gervais & Deville, 1853) Rough-toothed dolphin, Steno Mauretania Addink & Smeenk, 2001 bredanensis (G. Cuvier in Lesson, 1828) Risso’s dolphin, Grampus griseus (G. Barcelona, Spain Pilleri & Gihr, 1969 Cuvier, 1812)

South Africa Ross, 1984 checklistofturtleandwhalebarnacles 153 Azores Gruvel, 1920; Richard, 1936 Pygmy killer whale, Feresa attenuata Yenn, Senegal Stubbings, 1965 Gray, 1875 Long-beaked common dolphin, California Bay, Los Mochis, Mexico Kane et al., 2008 Delphinus capensis Gray, 1828 Short-beaked common dolphin, Western Mediterranean Pilleri, 1970 Delphinus delphis Linnaeus, 1758 Belgium Rappe´,1988 South Africa Gruvel, 1905a, 1920 Gore´e, Senegal Stubbings, 1965 Oran, Algeria Richard, 1936 Atlantic Hoek, 1883 Dusky dolphin, Lagenorhynchus Peru Van Waerebeek et al., 1993 obscurus Gray, 1828 Melon-headed whale, Peponocephala Kamitsu, Ibaraki, Japan Hayashi, 2012 electra (Gray, 1846)

Continued 5 yt hayashi ryota 154

Table 1. Continued

Species Host Locality References

Sperm whale, Physeter macrocephalus No locality records Mathews, 1938a Linnaeus, 1758 Finless black porpoise, Neophocaena Japan (Okumatsushima, Higashimatsushima, Miyagi; Hayashi, 2012 phocaenoides (G. Cuvier, 1829) Shirako, Chiba) Hong Kong Parsons et al., 2001 Karwar, India Devaraj & Bennet, 1974 Harbour porpoise, Phocoena Malaga, Spain Bellido et al., 2006 phocoena Linnaeus, 1758 Hann, Senegal Stubbings, 1965 Vaquita porpoise, Phocoena sinus Gulf of California Brownell, 1988 Norris & McFarland, 1958 Burmeister’s porpoise, Phocoena Peru Reyes & Van Waerebeek, 1995 spinipinnis Burmeister, 1865 True’s beaked whale, Mesoplodon South-eastern coast of South Africa Ross, 1984 mirus True, 1913 Minke whale, Balaenoptera North-western Pacific Uchida & Araki, 2000 acutorostrata Lace´pe`de, 1804 Antarctic Bushuev, 1990 Sei whale, Balaenoptera borealis Faroe Islands; Greenland Broch, 1924a Lesson, 1828 Norway (Finnmark; Ingoy) Nilsson-Cantell, 1921; Broch, 1924a; Rappe´ &Van Waerebeek, 1988 Saldanha Bay, South Africa Mackintosh, 1942; Matthews, 1938b; Nilsson-Cantell, 1930c Vancouver, Canada Cornwall, 1927 Tunis, Tunisia Heldt, 1950 Southern hemisphere; Northern Pacific Mathews, 1938b Bryde’s whale, Balaenoptera edeni Trujillo, Peru Kane et al., 2008 Anderson, 1879 Blue whale, Balaenoptera musculus Shetland Islands, Scotland Rappe´ & Van Waerebeek, 1988 (Linnaeus, 1758) South Africa Barnard, 1924; Mackintosh & Wheeler, 1929; Nilsson-Cantell, 1930c; Mackintosh, 1942 Pacific coast, Canada Cornwall, 1955b Fin whale, Balaenoptera physalus Shetland Islands, Scotland Calman, 1920; Nilsson-Cantell, 1930c (Linnaeus, 1758) Galicia, Spain Raga & Sanpera, 1986 South Africa Mackintosh & Wheeler, 1929; Nilsson-Cantell, 1930c; Mackintosh, 1942 Northern Atlantic; Antarctic Barnard, 1924 Humpback whale, Megaptera Piura, Peru Kane et al., 2008 novaeangliae Borowski, 1781 Unidentified Globicephalus sp. Bale´ares, Spain Richard, 1936 Atlantic Barnard, 1924; Richard, 1936 Unidentified Sotalia sp. Sa˜o Paulo, Brazil Young, 1991 Unidentified delphinid Heron Island, Queensland, Australia Pope, 1958; Jones, 2010 Unidentified porpoises North Atlantic Darwin, 1854 No host records Indian Sea Subba Rao, 2005 Bay of Biscay, United Kingdom Nilsson-Cantell, 1978 Greenland Broch, 1924a Western coast, Africa Nilsson-Cantell, 1930 Platylepas coriacea Monroe & Limpus, 1979 Leatherback turtle, Dermochelys Queensland, Australia (Wreck Rock; Mon Repos; Monroe & Limpus, 1979; Jones, 2010 coriacea (Vandelli, 1761) Alexandra Headlands) Florida, USA Biasatti, 2004 Loggerhead turtle, Caretta caretta Malta; Atlantic Gramentz, 1988 (Linnaeus, 1758) Platylepas decorata Darwin, 1854 Loggerhead turtle, Caretta caretta Australia (Mon Repos, Queensland; Heron Island, Monroe & Limpus, 1979; Jones, 2010 (Linnaeus, 1758) Queensland; Stradbroke) Green turtle, Chelonia mydas Japan (Ogimi, Okinawa; Ginoza, Okinawa; Yomitan, Hayashi, 2012 (Linnaeus, 1758) Okinawa; Ishigakijima Island, Okinawa; Yakushima Island, Kagoshima; Ogasawara, Tokyo) Jinyindao, Xisha Islands, South China Sea Ren, 1980 Heron Island, Queensland, Australia Monroe & Limpus, 1979; Jones, 2010 Teopa Beach, Jalisco State, Mexico Lazo-Wasem et al., 2011 Abrolhos Archipelago, Brazil Young, 1991 Isla Santa Cruz, Galapagos Islands, Equador Zullo, 1991 Hawksbill turtle, Eretmochelys Ishigakijima Island, Okinawa, Japan Hayashi, 2012 imbricata (Linnaeus, 1766) Australia (Wisteri Reef, Queensland; Campbell Island Monroe & Limpus, 1979; Limpus et al., 1983; Jones, 2004 Rookery; Dampier Archipelago) Kaisyaru, Baberudaobu Island, Republic of Palau Hiro, 1937a Olive ridley turtle, Lepidochelys Teopa Beach, Jalisco State, Mexico Lazo-Wasem et al., 2011 olivacea (Eschscholtz, 1829)

Unidentified sea turtle, Chelonia sp. Bahia, Brazil Young, 1991 checklistofturtleandwhalebarnacles 155 Unidentified sea turtle, ’green turtle, Tokoibuea, Little Andaman Island, India Daniel, 1962 (as Platylepas multidecorata) Eretomochelys sp.’ ? No host records Western Australia Jones, 2003 Lord Hood’s Island, Low Archipelago, French Polynesia Darwin, 1854 Corsica Gray, 1825 Galapagos Archipelago, Equador Darwin, 1854 Platylepas hexastylos (Fabricius, 1798) Loggerhead turtle, Caretta caretta Japan (Tassya, Aikawa, Sado Island, Niigata; Yomitan, Utinomi, 1970; Hayashi, 2009, 2012 (Linnaeus, 1758) Okinawa; Ginoza, Okinawa; Otsuchi, Iwate; Fukui, Fukui; Yakushima Island, Kagoshima; Tateyama, Chiba; Hachijojima Island, Tokyo) Queenland, Australia (Heron Island; Mon Repos; Monroe & Limpus, 1979; Jones, 2010 Bundaberg) Netherlands Holthuis, 1952 Aegean Sea Kitsos et al., 2005; Koukouras & Matsa, 1998 Hann, Senegal Stubbings, 1965 Malta Gramentz, 1988 Algeria to Balearic Islands Chevreux & de Guerne, 1893

Continued 5 yt hayashi ryota 156

Table 1. Continued

Species Host Locality References

USA (Delaware Bay, New Jersey; Virginia; Cape Cod, Pilsbry, 1916; Wells, 1966; Zullo & Bleakney, 1966; Massachusetts; Bald Point, Florida) Lutcavage & Musik, 1985; Zardus & Balazs, 2007 Brazil Bugoni et al., 2001 Levantine Basin Koukouras & Matsa, 1998 No locality records Ernst & Barbour, 1972 Green turtle, Chelonia mydas Japan (Ginoza, Okinawa; Ogimi, Okinawa; Yomitan, Hayashi, 2012; Chan & Hayashi (in press) (Linnaeus, 1758) Okinawa; Ishigakijima Island, Okinawa; Ogasawara, Tokyo; Motojima Beach, Wakayama; Manazuru, Kanagawa) Jinyindao, Xisha Islands, South China Sea Ren, 1980 Queensland, Australia (Heron Island; Moreton Bay; Zann & Harker, 1978; Monroe & Limpus, 1979; Jones, Magnetic Island; Wilson Island) 2010 Gore´e, Senegal Stubbings, 1965 USA (Sewalls Point, Cape May, New Jersey; Chincoteague Richards, 1930; Schwartz, 1960 Bay, Marylan) Teopa Beach, Jalisco State, Mexico Lazo-Wasem et al., 2011 Brazil (Rio Grande; Paraiba) Young, 1991; Bugoni et al., 2001 Urguay Alonso et al., 2010 Hawaiian Islands. Losey et al., 1994; Eldridge & Miller, 1997 Black turtle, Chelonia mydas agassizii Otsuchi, Iwate, Japan Hayashi et al., 2011 Bocourt, 1868 Hawksbill turtle, Eretmochelys Japan (Tateyama, Chiba; Fukui, Fukui; Minabe, Hayashi, 2012 imbricata (Linnaeus, 1766) Wakayama; Uruma, Okinawa; Ishigakijima Island, Okinawa) Wistari Reef, Queensland, Australia Monroe & Limpus, 1979; Jones, 2010 Republic of Palau Hiro, 1937a Olive ridley turtle, Lepidochelys Michoacan and Oaxaca, Mexico Vivaldo et al., 2006 (as C. testudinaria) olivacea (Eschscholtz, 1829) La Gloria, Jalisco, Mexico Hernandez-Vazquez & Valadez-Gonzalez, 1998 Flatback turtle, Natator depressus Queenland, Australia (Curtis Island; Mon Repos) Monroe & Limpus, 1979; Jones, 2010 (Garman, 1880) Leatherback turtle, Dermochelys Japan (Tassya, Aikawa, Sado Island, Niigata; Hakui, westrern Utinomi, 1970 coriacea (Vandelli, 1761) coast of Noto Peninsula, Ishikawa; Wakasa Bay, Fukui) Sandy Point, Virgin Island, USA Alexander et al.,2004 Unidentified sea turtle Mediterranean Darwin, 1854 Dugong, Dugong dugong (Mu¨ller, Australia (Moreton Bay, Queensland; Magnetic Island, Darwin, 1854; Zann & Harker, 1978; Jones, 2004 1776) Queensland; Dampier Archipelago) New Caledonia Fischer, 1884 African manatee, Trichechus Joal, Senegal Stubbings, 1965 senegalensis Link, 1795 Sherbro Island, Sierra Leone Stubbings, 1965 Unidentified manatee, Trichechus sp. River Gambia Darwin, 1854 Hondulas Darwin, 1854 Joao Pessoa, Brazil Young, 1991 No host records Dong-ao, Yilan County, Taiwan Chan et al., 2009 Western Australia Jones, 2003 Mauretania Broch, 1924b Morocco Broch, 1927a Platylepas hexastylos ichtyophila Pilsbry, 1916 Green turtle, Chelonia mydas Paraiba, Brazil Young, 1991 (Linnaeus, 1758) Gar fish, Lepisosteus sp. Hernando Country, Florida, USA, in brackish water Ryder, 1879; Pilsbry, 1916 Platylepas indicus Daniel, 1958 Sea snakes Madras, India Daniel, 1958 Platylepas ophiophilus Lanchester, 1902: 371. Shaw’s sea snake, Lapemis curtus Malay Peninsula Lanchester, 1902 (Shaw, 1802) Spine-belled sea snake, Lapemis McCluer Gulf, West Iran, Indonesia Zann & Harker, 1978 hardwickii Gray, 1834 Between Sumatra and Borneo Islands Broch, 1931 (as Platylepas kru¨geri) Olive sea snake, Aipysurus laevis North Queensland, Australia Zann, 1975 Lace´pe`de, 1804 Dubois’ sea snake, Aipysurus duboisii North Queensland, Australia Zann, 1975 (Bava, 1869) Spine-tailed sea snake, Aipysurus North Queensland, Australia Zann, 1975 eydouxii (Gray, 1849) Stokes’ sea snake, Astrotia stokesii North Queensland, Australia Zann, 1975 (Gray, 1846) Elegant sea snake, Hydrophis elegans North Queensland, Australia Zann, 1975 (Gray, 1842) Blue-banded sea snake, Hydrophis Mano Bay, Sado Island, Niigata, Japan Utinomi, 1970 cyanocinctus Daudin, 1803 Greater sea snake, Disteira major North Queensland, Australia Zann, 1975 (Shaw, 1802) Hook-nosed sea snake, Enhydrina Karachi, Pakistan Nilsson-Cantell, 1938 (as Platylepas ophiopholis) schistosa (Daudin, 1803) Ganjam Coast, Madras, India Nilsson-Cantell, 1938 checklistofturtleandwhalebarnacles 157 Slender sea snake, Microcephalophis Gwador, Baluchistan, Arabian Sea Kru¨ger, 1912 (as Cryptolepas ophiophilus) gracilis (Shaw, 1802) Unidentified sea snake, Disteira sp. Siam, Thailand Pilsbry, 1916 (as Platylepas kru¨geri) Unidentified sea snake, Hydrophis sp. Siam, Thailand Pilsbry, 1916 (as Platylepas kru¨geri) Unidentified sea snake Off Borneo Darwin, 1854 (as Platylepas sp.) East coast of Australia Nilsson-Cantell, 1921 (as Platylepas decorata) No host records Western Australia Jones, 2003 Thailand Jones et al., 2000 (as Platylepas kru¨geri) Platylepas sp. Loggerhead turtle, Caretta caretta Greece Frazier et al., 1985 (Linnaeus, 1758) South Africa Hughes, 1974 Brazil Bugoni et al., 2001 Green turtle, Chelonia mydas Hawaiian Islands Greenblatt et al.,2004 (Linnaeus, 1758) Hawksbill turtle, Eretmochelys Milman Island, Northern Great Barrier Reef, Australia Dobbs & Landry, 2004 imbricata (Linnaeus, 1766) More Island, Puerto Rico Scha¨rer, 2003 Rio Lagartos, Yucata´n, Mexico Ca´rdenas-Palomo & Maldonado-Gasca, 2005

Continued 5 yt hayashi ryota 158

Table 1. Continued

Species Host Locality References

Leatherback turtle, Dermochelys Trinidad and Tobago Bacon, 1976 coriacea (Vandelli, 1761) No host records Sa˜o Paulo, Brazil Young, 1991 Cylindrolepas darwiniana Pilsbry, 1916 Loggerhead turtle, Caretta caretta Yomitan, Okinawa, Japan Hayashi, 2012 (Linnaeus, 1758) USA (Florida; Georgia) Frick & Zardus, 2010 Mon Lepos, Queensland, Australia Frick & Zardus, 2010 Hawksbill turtle, Eretmochelys Panama Frick & Zardus, 2010 imbricata (Linnaeus, 1766) Florida, USA Frick & Zardus, 2010 Olive ridley turtle, Lepidochelys La Jolla, California, USA Hubbs, 1977 olivacea (Eschscholtz, 1829) Caretta caretta or Chelonbia mydas West Indies Pilsbry, 1916 Host unknown Helena Island, Moreton Bay, Queensland, Australia Jones, 2010 Cylindrolepas sinica Ren, 1980 Loggerhead turtle, Caretta caretta Yomitan, Okinawa, Japan Hayashi, 2009 (Linnaeus, 1758) Green turtle, Chelonia mydas Japan(Ogimi,Okinawa;Yomitan,Okinawa;Ginoza,Okinawa; Hayashi, 2012 (Linnaeus, 1758) Ishigakijima Island, Okinawa; Ogasawara, Tokyo) Jinyindao, Xisha Islands, South China Sea Ren, 1980 Hawaiian Islands Zardus & Balazs, 2007 (as Platylepas decorata) Hawksbill turtle, Eretmochelys Okinawa, Japan (Yomitan; Ishigakijima Island) Hayashi, 2012 imbricata (Linnaeus, 1766) Stomatolepas dermochelys Monroe & Limpus, 1979 Leatherback turtle, Dermochelys Kuala Lumpur, Malaysia Smaldon & Lyster, 1976 coriacea (Vandelli, 1761) Wreck Rock, Bundaberg, Queensland, Australia Monroe & Limpus, 1979; Jones, 2010 King Island, Tasmania, Australia Jones, 2010 North Island, New Zealand Monroe & Limpus, 1979 Whareama River, Castle Point, North Island, New Zealand McCann, 1969 Ameland, Friesland, The Netherlands Monroe & Limpus, 1979 Belgium (De Haan; Oostende) Haetlers & Kelckhof, 1999; Haetlers et al., 2001 Skarvoy, Norway Smaldon & Lyster, 1976 Scotland (Kirkcudbrightshire; Crail, Fife) Smaldon & Lyster, 1976 Cornwall, England Smaldon & Lyster, 1976; Monroe & Limpus, 1979 Nova Scotia, Canada (Sambro; Seawall, Digby County) Zullo & Bleakney, 1966 Sandy Point, Virgin Islands, USA Alexander et al.,2004 Loggerhead turtle, Caretta caretta Florida, USA (Bald Point; Dry Tortugas) Gittings et al., 1986 (Linnaeus, 1758) Stomatolepas elegans (Costa, 1838) Loggerhead turtle, Caretta caretta Seto, Wakayama, Japan Hiro, 1937b (Linnaeus, 1758) Aegean Sea, Greece Kitsos et al., 2003; 2005 Hann, Senegal Stubbings, 1965 Green turtle, Chelonia mydas Teopa Beach, Jalisco State, Mexico Lazo-Wasem et al., 2011 (Linnaeus, 1758) Paraiba and Bahia, Brazil Young, 1991 Hawaiian Islands Zardus & Balazs, 2007 Leatherback turtle, Dermochelys Tassya, Aikawa, Sado Island, Niigata, Japan Utinomi, 1970 coriacea (Vandelli, 1761) Skreifjorden, Seiland, Finnmark, Norway Carriol & Vader, 2002 North-east Atlantic, Madeira Wirtz et al., 2006 Olive ridley turtle, Lepidochelys Teopa Beach, Jalisco State, Mexico Lazo-Wasem et al., 2011 olivacea (Eschscholtz, 1829) Unidentified sea turtle, Chelonia sp. Bahia, Abrolhos Archipelago Young, 1991 No host records Western Australia Jones, 2003 Taranto, Italy Costa, 1838; Pilsbry, 1916 Nova Scotia, Canada Frick et al., 2010a Wassaw Island, Georgia, USA Frick et al., 2010a Stomatolepas pilsbryi Frick, Zardus and Leatherback turtle, Dermochelys Gabon, Africa Frick et al., 2010a Laso-Wasem, 2010 coriacea (Vandelli, 1761) Stomatolepas praegustator Pilsbry, 1910 Loggerhead turtle, Caretta caretta Japan (Yakushima Island, Kagoshima; Otsuchi, Iwate; Hiro, 1936b; Hayashi, 2012 (Linnaeus, 1758) Ooamishirosato-machi, Chiba) Queensland, Australia (Heron Island; Mon Repos) Monroe & Limpus, 1979; Jones, 2010 USA (Tortugas, Florida; Bald Point, Florida; Wassaw Pilsbry, 1910; Wells, 1966 Island, Georgia) Green turtle, Chelonia mydas Yomitan, Okinawa, Japan Hayashi, 2012 (Linnaeus, 1758) Black turtle, Chelonia mydas agassizii Jalisco Coast, western Mexico Lazo-Wasem et al., 2007 Bocourt, 1868 Olive ridley turtle, Lepidochelys Jalisco Coast, western Mexico Lazo-Wasem et al., 2007 olivacea (Eschscholtz, 1829) Leatherback turtle, Dermochelys Wassaw Island, Georgia, USA Frick et al., 2010a coriacea (Vandelli, 1761) No host records Western Australia Jones, 2003 Stomatolepas transversa Nilsson-Cantell, 1930 Green turtle, Chelonia mydas Japan (Ginoza, Okinawa; Yomitan, Okinawa; Ishigakijima Hayashi & Tsuji, 2008; Hayashi, 2012

(Linnaeus, 1758) Island, Okinawa; Ogasawara Islands, Tokyo) checklistofturtleandwhalebarnacles 159 Jinyindao, Xisha Islands, South China Sea Ren, 1980; Liu & Ren, 2007 (as Stomatolepas pulchra) Australia (Enoe Island., Aroe Archipelago; Wistari Reef; Nilsson-Cantell, 1930b; Monroe & Limpus, 1979; Jones, Heron Island) 2010 Jao Pessoa, Brazil Young, 1991 Stephanolepas muricata Fischer, 1886 Loggerhead turtle, Caretta caretta Japan (Yomitan, Okinawa; Ginoza, Okinawa; Yakushima Hayashi, 2009; 2012 (Linnaeus, 1758) Island, Kagoshima; Otsuchi, Iwate) Queensland, Australia (Heron Island; Mon Repos) Monroe & Limpus, 1979; Jones, 2010 Marathon Bay, Florida, USA Frick et al., 2011 Green turtle, Chelonia mydas Japan (Ogimi, Okinawa; Yomitan, Okinawa; Ginoza, Hayashi, 2012 (Linnaeus, 1758) Okinawa; Ogasawara, Tokyo) Queensland, Australia (Helon Island; Moreton Bay) Monroe & Limpus, 1979; Jones, 2010 Hawaiian Islands Balazs, 1978; Eldridge & Miller, 1997 Hawksbill turtle, Eretmochelys Yomitan, Okinawa, Japan Hayashi, 2012 imbricata (Linnaeus, 1766) Pulo Condor, Condor Island, Vietnam Fischer, 1886 Boydong Island, Queensland, Australia Monroe & Limpus, 1979 Olive ridley turtle, Lepidochelys USA (North Carolina; Georgia) Frick et al., 2011 olivacea (Eschscholtz, 1829) Mexico (Baja California; Playa Ceuta, Sinaloa) Frick et al., 2011 Calyptolepas bjorndalae Frick, Zardus and Green turtle, Chelonia mydas Kennedy Space Center, Florida, USA Frick et al., 2010b Lazo-Wasem, 2010 (Linnaeus, 1758) 160 ryota hayashi

Table 2. Fossil records of Coronuloidea (including extant species) indicating their geological ages and their collection locality.

Geological age/species Locality References

Late Eocene †Emersonius cybosyrinx Ross & Newman, Florida, USA Ross & Newman, 1967 1967 Oligocene †Chelonibia melleni Zullo, 1982 Rankin and Smith Counties, Mississippi, USA Zullo, 1982 Early Miocene †Protochelonibia submersa Harzhauser & North Alpine Foreland Basin, Austria Harzhauser et al., 2011 Newman, 2011 Chelonibia caretta (Spengler, 1790) Ras Kingoje, Pemba Island, Zanzibar, United Withers, 1928 Republic of Tanzania †Chelonibia testudinaria solida Withers, Lorient, near Pessac (Gironde), France Withers, 1929 1929 Middle-Miocene †Protochelonibia capellini (de Alessandri, Italy de Alessandri, 1895 1895) Late Miocene Chelonibia patula (Ranzani, 1818) Kalksburg, Vienna, Austria Withers, 1929 Chelonibia patula (Ranzani, 1818) De Soto County, Florida, USA Ross, 1963b Coronula diadema (Linnaeus, 1767) Minami-Kanbaragun, Niigata, Japan Hatai, 1939 Miocene Chelonibia testudinaria (Linnaeus, 1758) Cuba Withers, 1953 Mio-Pliocene †Chelonibia testudinaria solida Withers, France Withers, 1929 1929 Early Pliocene Coronula diadema (Linnaeus, 1767) Masuda, Nikko-mura, Akumi-gun, Japan Hatai, 1938 Upper Pliocene †Coronula aotea Fleming, 1959 Waihua Valley and Mangaotoro River, New Fleming, 1959 Zealand †Coronula barbara Darwin, 1854 Bawdsey and Sutton, England Darwin, 1854 †Coronula barbara Darwin, 1854 San Diego and Orange County, California, USA Zullo, 1969 Pliocene †Chelonibia depressa Seguenza, 1876 Sicily, Italy Seguenza, 1876 †Chelonibia hemisphaerica Rothpletz & Grand Canary Island, Spain Rothpletz & Simonelli, 1890 Simonelli, 1890 †Coronula barbara Darwin, 1854 Orciano Pisano and Otranto, Italy de Alessandri, 1906; Menesini, 1968 †Coronula bifida Bronn, 1831 Punta Gorda, Ecuador Pilsbry & Olson, 1951 †Cetolepas hertleini Zullo, 1969 San Diego, California, USA Zullo, 1969 Late Pliocene †Coronula intermedia Buckeridge, 1983 Castlepoint Formation, Castlepoint, New Zealand Beu, 1995 Pliocene to Pleistocene Coronula diadema (Linnaeus, 1767) Punta Canoa-San Jose, Ecuador Bianucci et al., 2006 Lower Pleistocene †Coronula intermedia Buckeridge, 1983 Makara Stream, New Zealand Buckeridge, 1983 Coronula diadema (Linnaeus, 1767) Four distinct localities, New Zealand Fleming, 1959; Beu, 1971; Buckeridge, 1983 Coronula diadema (Linnaeus, 1767) Ostrakodes, Cyprus Simonelli, 1893; de Alessandri, 1906 Pleistocene Chelonibia testudinaria (Linnaeus, 1758) Punta Gorda, Charlotte County, Florida, USA Ross, 1963b †Chelonibia testudinaria solida Withers, Flagler County, Florida, USA Ross, 1963b 1929 †Coronula ficarazzensis Gregorio, 1895 Ficarazzi, near Palermo, Italy de Gregorio, 1895; de Alessandri, 1906 †Cryptolepas murata Zullo, 1961 San Quintin Bay, Baja California, Mexico Zullo, 1961 †Platylepas wilsoni Ross, 1963 South-east of Rim Ditch Canal, Florida, USA Ross, 1963a Coronula diadema (Linnaeus, 1767) Kere river mouth, Vanuatu Bianucci et al., 2006 Coronula diadema (Linnaeus, 1767) Puerto Villamil, Galapagos Islands, Ecuador Zullo, 1986 Coronula diadema (Linnaeus, 1767) Zorritos, Peru Pilsbry & Olson, 1951 Coronula diadema (Linnaeus, 1767) Quebrada, Mare Abajo, Venezuela Weisbord, 1971 †Coronula bifida Bronn, 1831 Valle del Lamato, Italy de Alessandri, 1905 Coronula diadema (Linnaeus, 1767) Riparbella, Tuscany, Italy Domocimi et al., 2011 Pleistocene to recent Coronula diadema (Linnaeus, 1767) Arauco, Chile Pilsbry & Olson, 1951 Chelonibia testudinaria (Linnaeus, 1758) Galapagos Islands, Ecuador Zullo, 1986 Holocene Cetopirus complanatus (Mo¨rch, 1852) Buenos Aires, Argentina Pastorino & Griffin, 1996

Continued checklistofturtleandwhalebarnacles 161

Table 2. Continued

Geological age/species Locality References

Coronula diadema (Linnaeus, 1767) Geelbek Dunes, South Africa Jerardino & Parkington, 1993; Kandel & Conard, 2003; Conard & Kandel, 2006 Coronula diadema (Linnaeus, 1767) Pinnacle Point near Mossel Bay, South Africa Marean et al., 2007 Coronula diadema (Linnaeus, 1767) Cantabria, Spain A´ lvarez-Ferna´ndez, 2011 Coronula diadema (Linnaeus, 1767) Dunefield Midden, west coast of South Africa Stewart, 2011 Coronula diadema (Linnaeus, 1767) Abu Dhabi, United Arab Emirates Stewart et al., 2011 distribution distribution Indo-Pacific; Atlantic; Mediterranean. Atlantic (Florida). †Chelonibia depressa Seguenza, 1876 †Chelonibia melleni Zullo, 1982 †Chelonibia depressa Seguenza, 1876: 411. †Chelonibia melleni Zullo, 1982: 1, figures 1–13. references references Ross & Frick, 2011: 61; Seguenza, 1876: 411; Withers, 1928a: Ross & Frick, 2011: 61; Zullo, 1982: 1. 390; 1929: 567; Zullo, 1982: 2. Chelonibia patula (Ranzani, 1818) †Chelonibia hemisphaerica Rothpletz & Simonelli, 1890 Coronula patula Ranzani, 1818: 86, figures 25–28. †Chelonibia hemisphaerica Rothpletz & Simonelli, 1890: 724. Coronula dentulata Say, 1822: 325. references Coronula denticula Gray, 1825: 105. Astrolepas laevis Gray, 1825: 105. Harzhauser et al., 2011: 477; Ross, 1963b: 223; Ross & Frick, Chelonobia patula Darwin, 1854: 396, plate 14, 2011: 61; Rothpletz & Simonelli, 1890: 724; Withers, 1928a: figures 3a, 3b & 4. 390; 1929: 567; Zullo, 1982: 2. references Chelonibia manati Gruvel, 1903 Achituv, 1998: 1204; Anderson, 1994: 31; Bakır et al., 2010: 1; Chelonibia manati Gruvel, 1903: 116. Bland & Amerson, 1974: 232; Broch, 1924b: 203; 1927b: 136; Chelonibia ramosa Korschelt, 1933: 2, figures 1–6. 1935: 2; 1947: 4; Celis et al., 2007: 327; Chan et al., 2009: 168; references Condrey & Gelpi, 2010: 6; Costa et al., 2010: 387; Crisp, 1974: Anderson, 1994: 297; Beck & Forrester, 1988: 633; Broch, 230; 1983: 281; Crisp & Costlow, 1963: 22; Daniel, 1955: 32; 1924b: 203; Fertl & Newman, 2009: 91; Frazier, 1986: 472; Darwin, 1854: 396; Dawydoff, 1952: 129; Edmondson, 1933: Frick et al., 2011: 637; Gruvel, 1903: 116; 1905a: 267; 231; Ernst & Barbour, 1972: 100; Farrapeira, 2010: 212; Harzhauser et al., 2011: 478; Hiro, 1936a: 61; Killingley & Frazier, 1986: 472; Frazier & Margaritoulis, 1990: 213; Frick Lutzcavage, 1983: 349; Korschelt, 1933: 17; Mignucci- et al., 2004: 615; Gaddes & Sumpton, 2004: 241; Gannon, Giannoni et al., 1999: 67; Morales-Vela et al., 2008: 594; 1990: 56; Gauld, 1957: 10; Gittings et al., 1986: 3; Gordon, Pilsbry, 1916: 265; Ross, 1963b: 223; Ross & Frick, 2011: 61; 1970: 90; Gray, 1825: 105; Gruvel, 1905a: 268; 1907: 8; 1912: Seigel, 1983: 34; Stubbings, 1965: 874; 1967: 297; Utinomi, 346; Harris et al., 2000: 395; Harzhauser et al., 2011: 475; 1950: 62; Zullo, 1982: 2. Heath, 1976: 59; Henry, 1943: 367; 1954: 444; Hiro, 1936a: 60; Hudson & Lester, 1994: 183; Inatsuchi et al., 2010: 904; distribution Innocenti, 2006: 6; Jackson et al., 1973: 495; Javed & Mustaquim, 1994: 124; Johnson & Bonner, 1960: 147; Jones, Atlantic Ocean. 2003: 483; 2004: 126; 2010: 208; Jones et al., 1990: 11; Kelly Chelonibia manati crenatibasis Pilsbry, 1916 & Sanford, 2010: 39; Key Jr et al., 1997: 426; Kitsos et al., Chelonibia manati crenatibasis Pilsbry, 1916: 266, 2003: 406; 2005: 259; Kolosva´ry, 1943: 98; Korschelt, 1933: plate 62, figure 5 & 5a. 17; Kru¨ger, 1911a: 4; Lacombe, 1977: 11; Lacombe & Rangel, references 1978: 5; Lang, 1976: 219; Lawal-Are & Daramola, 2010: 520; Lemaitre, 1995: 646; Liu & Ren, 2007: 308; Mantelatto et al., Pilsbry, 1916: 266; Ross & Frick, 2011: 61; Stubbings, 1965: 2003: 211; McDermott, 1998: 293; 1999: 427; 2007: 2380; 895; 1967: 297; Zullo, 1982: 2. McDougall, 1943: 343; Monroe, 1981: 241; Nakamura et al., 1994: 99; Negreiros-Fransozo et al., 1995: 258; Newman & distribution Ross, 1976: 18; Nilsson-Cantell, 1934: 61; 1938: 77; Norse & No locality records. Estevez, 1977: 371; Pasternak et al., 2002: 583; Pearse, 1947: 327; 1952: 7; Pe´rez-Losada et al., 2004: 247; 2008: 330; Chelonibia manati lobatobasis Pilsbry, 1916 Phillips & Cannon, 1978: 142; Pilsbry, 1916: 268; Ranzani, Chelonibia manati lobatobasis Pilsbry, 1916: 266, 1818: 86; Relini, 1969: 169; Ross, 1963b: 223; Ross & Frick, plate 62, figure 7 & 7a. 2011: 61; Ross & Jackson, 1972: 203; Ross & Newman, 1967: references 16; Sandeen & Costlow, 1961: 19; Say, 1822: 325; Seigel, Frazier & Margaritoulis, 1990: 216; Gittings et al., 1986: 3; 1983: 35; Shields, 1992: 94; Southward & Crisp, 1963: 26; Henry, 1954: 444; Kolosva´ry, 1942b: 146; Pilsbry, 1916: 266; Spears et al., 1994: 645; Stubbings, 1961: 39; 1965: 876; Ross & Frick, 2011: 61; Seigel, 1983: 35; Stubbings, 1965: 1967: 297; Tavares & Mendonc¸a Jr, 2011: 49; Utinomi, 1950: 895; 1967: 297; Wells, 1966: 86; Zullo, 1982: 2. 62; 1958: 309; Visscher, 1928: 329; Walker, 1978: 209; 1983: 162 ryota hayashi

433; Wass, 1963: 36; Weeks et al., 2006: 452; Wells, 1966: 86; Lanchester, 1902: 371; Lazo-Wasem et al., 2011: 222; Weltner, 1897: 254; Williams & Porter, 1964: 150; Withers, Limpus et al., 1983a: 192; 1983b: 180; 1994: 145; Linnaeus, 1929: 567; Young, 1991: 191; 1995: 230; 1998: 271; 2000: 85; 1758: 668; 1767: 1108; Liu & Ren, 2007: 307; Lucas, 1968: Yusa et al., 2010: 260; Zevina & Litvinova, 1970: 174; Zullo, 136; Lutcavage & Musik, 1985: 453; MacDonald, 1929: 537; 1963: 14; 1982: 1; Zullo & Lang, 1978: 160. Margaritoulis, 1985: 453; Matsuura & Nakamura, 1993: 1803; Meischner, 2001: 1; Monod, 1931: 404; Monroe, 1981: distribution 237; Monroe & Garrett, 1979: 108; Monroe & Limpus, 1979: Cosmopolitan. 199; Mo¨rch, 1852: 67; Moriarty et al., 2008: 1; Mustaquim & Javed, 1993: 73; Newman & Abbott, 1980: 516; Newman Chelonibia patula dentata Henry, 1943 et al., 1969: 289; Nifong & Frick, 2011: 557; Nilsson-Cantell, Chelonibia patula dentata Henry, 1943: 370, plate 1921: 369; 1930b: 19; 1931: 116; 1932: 258; 1937: 95; 1938: 31 figures 4–13. references 77; 1939a: 5; 1957: 7; Pasternak et al., 2002: 583; Pereira et al., 2006: 17; Pfaller et al., 2006: 1; 2008: 1096; Pillai, Henry, 1943: 370; 1960: 147; Liu & Ren, 2007: 310; Ross, 1962: 1958: 126; Pilsbry, 1916: 264; 1928: 316; Plotkin, 1996: 79; 6; 1963b: 223; Ross & Frick, 2011: 61; Zullo, 1982: 2; 1991: 175. Ranzani, 1818: 85; Rawson et al., 2003: 2697; Rees & distribution Walker, 1993: 189; Relini, 1969: 169; 1980: 88; Ren, 1980: 187; Richards, 1930: 143; Riedl, 1963: 258; Ross, 1962: 6; Mexico; China. 1963a: 156; 1963b: 223; 1981: 102; Ross & Frick, 2011: 61; Chelonibia testudinaria (Linnaeus, 1758) Ross & Newman, 1967: 15; Rudloe et al., 1991: 50; Santoro Lepas testudinaria Linnaeus, 1758: 668. & Mattiucci, 2009: 509; Scha¨rer, 2003: 88; Seigel, 1983: 34; Verruca testudinaria Ellis, 1758: 852, plate 34, figure 12. Spivey, 1981: 155; Stamper et al., 2005: 637; Stubbings, 1965: Astrolepas rotundarius Gray, 1825: 105. 876; 1967: 296; Tachikawa, 1995: 67; Utinomi, 1949: 24; Chelonobia testudinaria Darwin, 1854: 392, plate 14, 1950: 53; 1958: 209; 1969: 92; 1970: 359; Utinomi & figure 1a–d. Kikuchi, 1966: 8; Visscher, 1928: 329; Vivaldo et al., 2006: Chelonibia testudinaria Pilsbry, 1916: 264, plate 62, figures 434; Wagh & Bal, 1974: 121; Walker, 1978: 205; 1983: 433; 1–4.—Hayashi, 2012: 107, plate 1a, figures 1 & 15a. Wass, 1963: 36; Weeks et al., 2006: 452; Wells, 1966: 83; Weltner, 1895: 298; 1897: 254; 1899a: 443; 1910: 528; references Withers, 1928a: 390; 1929: 567; Yamaguchi, 1987: 337; Aguirre et al., 1994: 9; Anderson, 1994: 297; Angulo-Lozano Young, 1991: 191; 1998: 271; Young & Ross, 2000: 224; et al., 2007: 13; Annandale, 1906: 138; Aznar et al., 2010: Yusa et al., 2010: 260; Zann, 1975: 268; Zardus & Hadfield, 306; Bacon, 1976: 34; Balazs, 1978: 128; Barnard, 1924: 92; 2004: 409; Zavodnik, 1997: 116; Zullo, 1963: 14; 1982: 2; Biasatti, 2004: 206; Borradaile, 1903: 443; Broch, 1916: 14; 1986: 55; 1991: 175; Zullo & Lang, 1978: 160. 1924b: 204; 1931: 122; 1947: 7; Brown & Brown, 1995: 238; Bugoni et al., 2001: 7; Caine, 1986: 19; Ca´rdenas-Palomo & distribution Maldonado-Gasca, 2005: 32; Casale et al., 2004: 1005; Cosmopolitan. Caziot, 1921: 51; Chan et al., 2009: 168; Cupul-Magan˜a et al., 2011: 213; Daniel, 1955: 31; Darwin, 1854: 392; †Chelonibia testudinaria solida Withers, 1929 Davenport, 1994: 735; Dawydoff, 1952: 129; de Alessandri, †Chelonibia testudinaria solida Withers, 1929: 568, 1895: 391; 1906: 314; Deraniyagala, 1939: 204; Edmondson, plate 17, figures 5–7. 1946: 231; Edmondson & Ingram, 1939: 255; Eldredge & references Miller, 1997: 11; Ellis, 1758: 852; Farrapeira, 2010: 212; Ross, 1963b: 230; Ross & Frick, 2011: 61; Ross & Newman, Fischer, 1884: 359; Flint et al., 2009: 741; 2010: 299; Frazier, 1967: 19; Withers, 1929: 568; Zullo, 1982: 2. 1971: 394; 1986: 472; 1989: 250; Frazier & Margaritoulis, 1990: 216; Frazier et al., 1985: 129; Frick & Ross, 2001: 16; Genus †Emersonius Ross & Newman, 1967 2002: 10; Frick & Slay, 2000: 103; Frick et al., 1998: 213; †Emersonius cybosyrinx Ross & Newman, 1967 2000: 3; 2003b: 19; 2004: 615; 2010b: 169; 2011: 629; Fuller †Emersonius cybosyrinx Ross & Newman, 1967: 8, figures 1–4. et al., 2010: 1743; Gauld 1957: 10; Geldiay et al., 1995: 428; references Gittings et al., 1986: 3; Glazebrook & Campbell, 1990: 102; Gonza´lez et al., 2012: 132; Gordon, 1970: 94; Gramentz, Monroe, 1981: 244; Newman et al., 1969: 290; Ross & Frick, 1988: 35; Gray, 1825: 105; Gruvel, 1903: 115; 1905a: 266; 2011: 61; Ross & Newman, 1967:7; Zullo, 1982: 2. 1907: 8; 1931: 404; Guess, 1982: 122; Gunter, 1981: 90; Genus †Protochelonibia Harzhauser & Newman, 2011 Harding et al., 2011: 236; Harzhauser et al., 2011: 478; †Protochelonibia capellini (de Alessandri, 1895) Hayashi, 2009: 2; 2012: 107; Hayashi & Tsuji, 2008: 121; †Chelonibia capellini de Alessandri, 1895: 300, plate 3, figure Heller, 1866: 759; Hendrickson, 1958: 523; Henry, 1941: 5a, b.—1906: 313, plate 18, figures 3–5. 105; 1943: 368; 1954: 444; 1960: 147; Hernandez-Vasquez & †Protochelonibia capellini Harzhauser et al., 2011: 473. Valadez-Gonzalez, 1998: 121; Hiro, 1936a: 61; 1936b: 312; references 1937a: 69; 1937b: 470; 1939: 214; Innocenti, 2006: 6; Ives, de Alessandri, 1895: 300; 1906: 313; Harzhauser et al., 2011: 1891: 189; Jones, 2003: 484; 2004: 126; 2010: 209; Jones 473; Ross, 1963b: 223; Ross & Frick, 2011: 61; Withers, et al., 1990: 11; 2000: 260; Kelly & Sanford, 2010: 39; 1928a: 390; 1929: 567; Zullo, 1982: 2. Killingley & Lutzcavage, 1983: 345; Kitsos et al., 2003: 407; 2005: 259; Kolosva´ry, 1939: 38; 1942b: 149; 1943: 99; 1951: †Protochelonibia submersa Harzhauser & Newman, 2011 411; Korschelt, 1933: 16; Koukouras & Matsa, 1998: 136; †Protochelonibia submersa Harzhauser & Newman, 2011: Kru¨ger, 1911a: 57; 1911b: 461; Laguna, 1990: 409; 473, figures 2–5b. checklistofturtleandwhalebarnacles 163 reference 1872: 300; Darwin, 1854: 417; 1855: 39; Dominici et al., Harzhauser et al., 2011: 473. 2011: 95; Dyer, 1989: 1088; Ellis, 1758: 851; Fe´lix et al., 2006: 979; Fertl & Newman, 2009: 90; Filhol, 1885: 489; Family CORONULIDAE Leach, 1817 Fischer, 1872: 433; 1884: 359; Fleming, 1959: 246; Ford & Genus Coronula Lamarck, 1802 Reeves, 2008: 63; Foster, 1978: 115; Geraci & Aubin, 1987: †Coronula aotea Fleming, 1959 407; Gonza´lez et al., 2012: 132; Gray, 1825: 105; 1843: 269; †Coronula aotea Fleming, 1959: 243, figures 1–5, 7, 9 &10.— Gruvel, 1903: 152; 1905a: 273; 1905b: 308; 1911: 292; Guiler, Beu, 1971: 899, figures 9 & 10. 1956: 3; Haelters et al., 2010: 96; Hatai, 1938: 98; 1939: 262; references Hayasaka, 1933: 49; 1935: 1; Hayashi, 2012: 110; Heath, 1976: 59; Henry, 1943: 368; Hiro, 1935: 226; 1936a: 62; Beu, 1971: 899; Bianucci et al., 2006a: 331; 2006b: 118; 1936b: 318; 1937b: 471; 1939: 214; Hoek, 1883: 163; 1909: Buckeridge, 1983: 81; Fleming, 1959: 243; Ross & Frick, 271; Holmes et al., 1997: 153; Holthuis & Fransen, 2004: 12; 2011: 64. Holthuis et al., 1998: 352; Hutton, 1879: 329; Innocenti, †Coronula barbara Darwin, 1854 2006: 6; Jennings, 1918: 62; Jerardino & Parkington, 1993: 6; †Coronula barbara Darwin, 1854: 421, plate 15, figure 6. Jime´nez et al., 2011: 178; Jones, 2003: 484; 2010: 209; Jones †Coronula bifida barbara Menesini, 1968a: 395, plate 3, et al., 1990: 12; Kandel & Conard, 2003: 91; Kolosva´ry, figures 3a–5c; plat 4, figures 2, 4 &6. 1942a: 138; 1942b: 149; 1943: 99; Korschelt, 1933: 18; references Linnaeus, 1767: 1108; Liu & Ren, 2007: 325; Marean et al., Beu, 1971: 900; Bianucci et al., 2006a: 331; 2006b: 116; 2007: 906; Marloth, 1900: 1; Menesini, 1968a: 389; Monroe, Buckeridge, 1983: 82; Darwin, 1854: 421; 1855: 38; de 1981: 241; Mo¨rch, 1852: 66; 1911: 667; Moss & Erlandson, Alessandri, 1895: 303; 1906: 317; Dominici et al., 2011: 98; 2010: 3367; Mu¨ller, 1776: 250; Newman & Ross, 1971: 178; Menesini, 1968a: 387; Ross & Frick, 2011: 64; Stewart et al., Newman et al., 1969: 289; Nilsson-Cantell, 1921: 371; 1930c: 2011: 3685; Weisbord, 1971: 91; Withers, 1928b: 63; Zullo, 256; 1930d: 212; 1931: 116; 1938: 14; 1939b: 237; 1957: 7; 1969: 21; 1986: 58. 1978: 75; Nogata & Matsumura, 2006: 92; O’Riordan, 1967: 294; Pasternak et al., 2002: 583; Paterson & Van Dyck, 1991: †Coronula bifida Bronn, 1831 345; Pilsbry, 1916: 273; Pilsbry & Olson, 1951: 203; Ranzani, †Coronula barbara Bronn, 1831: 126. 1818: 88; Rees, 1953: 33; Rice, 1963: 185; Ross, 1962: 6; Ross †Coronula dormitor Pilsbry & Olson, 1951: & Frick, 2011: 64; Sars, 1880: 15; Scarff, 1986: 129; Scheffer, 202—Weisbord, 1971: 94. 1939: 67; Schumacher, 1817: 91; Seilacher, 2005: 29; Slijper, †Coronula bifida bifida Menesini, 1968a: 387, 1962: 90; Stebbing, 1910: 571; Stephensen, 1938: 6; Stewart, plates 1–3, figures 1a–2b; plate 4, figures 1, 3 2011: 163; Stewart et al., 2011: 3685; Stubbings, 1961: 15; &5. references Tachikawa, 1995: 67; Tarasov & Zevina, 1957: 241; Van Beneden, 1870: 349; Visscher, 1928: 328; Weisbord, 1971: Bianucci et al., 2006a: 327; 2006b: 116; Bronn, 1831: 126; 91; Weltner, 1895: 290; 1897: 254; 1898: 8; 1899b: 102; 1900: Darwin, 1854: 423; 1855: 40; de Alessandri, 1895: 302; 1906: 302; 1922: 86; Wirtz, et al., 2006: 209; Wolff, 1960: 8; 315; Dominici et al., 2011: 96; Menesini, 1968a: 387; 1968b: Young, 1991: 193; 1998: 273; 2000: 85; 2001: 746; Zullo, 584; Pilsbry & Olson, 1951: 202; Ross & Frick, 2011: 64; 1963: 14; 1986: 55; 1991: 175. Seguenza, 1876: 324; Stewart et al., 2011: 3685; Weisbord, 1971: 94; Withers, 1928b: 63. distribution Coronula diadema (Linnaeus, 1767) Cosmopolitan (without Mediterranean). Pediculus ceti Ellis, 1758: 851, figure 7, 7a. †Coronula ficarazzensis Gregorio, 1895 †Coronula ficarazzensis Gregorio, 1895: 5. Lepas diadema Linnaeus, 1767: 1108. references Lepas balaenaris Mu¨ller, 1776: 250. Diadema vulgaris Schumacher, 1817: 91.—Fischer, 1884: Bianucci et al., 2006b: 118; de Alessandri, 1906: 315; Gregorio, 359. 1895: 5. Diadema candidum Ranzani, 1818: 88. †Coronula intermedia Buckeridge, 1983 Coronula diadema Lamarck, 1818: 387.—Darwin, 1854: †Coronula intermedia Buckeridge, 1983: 81, figure 417, plate 15, figsure 3, 3a, 3b; plate 16, figures 1, 2 & 7. 60. Polylepas (Diadema) kleinii Gray, 1825: 105. references Coronula biscayensis Van Beneden, 1870: 349. Beu, 1995: 19; Bianucci et al., 2006a: 331; 2006b: 116; Diadema japonica Van Beneden, 1870: 354. Buckeridge, 1983: 81. Coronula macsotayi Weisbord 1971: 91, plate 20, figures 1–4. Coronula reginae Darwin, 1854 references Coronula reginae Darwin, 1854: 419, plate 15, Achituv, 1998: 1204; A´ lvarez-Ferna´ndez, 2011: 336; figure 5; plate 16, figure 4.—Broch, 1924a: 93, Anderson, 1994: 297; Barnard, 1924: 94; Barnes & Klepal, plate 1, figure 11.—Nilsson-Cantell, 1930c: 1971: 88; Bassindale, 1964: 43; Best, 1991: 585; Beu, 1971: 257, plate 1, figures 5–10. 902; Bianucci et al., 2006a: 326; 2006b: 115; Borradaile, Coronula regina MacKintosh & Wheeler, 1929: 1916: 135; Broch, 1924a: 91; Buckeridge, 1983: 81; Chan 373. et al., 2009: 180; Clarke, 1966: 155; Conard & Kandel, 2006: references 335; Cornwall, 1924: 421; 1927: 503; 1928: 9; 1953: 83; Anderson, 1994: 81; Barnard, 1924: 94; Barnes & Klepal, 1971: 1955a: 51; 1955b: 40; Crisp & Stubbings, 1957: 179; Dall, 88; Best, 1991: 585; Broch, 1924a: 93; Buckeridge, 1983: 82; 164 ryota hayashi

Caldwell, 1963: 569; Clarke, 1966: 157; Cornwall, 1927: 507; distribution 1928: 9; 1955a: 53; 1955b: 43; Darwin, 1854: 419; 1855: 39; Southern Ocean. Dominici et al., 2011: 97; Fe´lix et al., 2006: 979; Fertl & Newman, 2009: 90; Ford & Reeves, 2008: 63; Gruvel, 1903: Genus Cryptolepas Dall, 1872 152; 1905a: 272; Haelters et al., 2010: 96; Hiro, 1936a: 62; †Cryptolepas murata Zullo, 1961 1936b: 318; Holmes et al., 1997: 153; Holthuis & Fransen, †Cryptolepas murata Zullo, 1961: 14, plate 3, figures 1–5. 2004: 12; Innocenti, 2006: 7; Jerardino & Parkington, 1993: 6; Kolosva´ry, 1942a: 140; 1942b: 141; 1943: 99; Kru¨ger, references 1927a: 15; 1927b: 5; MacKintosh & Wheeler, 1929: 373; Ross, 1962: 4; Zullo, 1961: 14; 1969: 17. Matthews, 1938a: 126; Monroe, 1981: 241; Moss & Erlandson, 2010: 3367; Newman & Ross, 1971: 178; Cryptolepas rhachianecti Dall, 1872 Nilsson-Cantell, 1926: 15; 1930c: 256; 1939b: 238; 1957: 8; Cryptolepas rhachianecti Dall, 1872: 300.— 1978: 75; Petriconi, 1969: 539; Pilsbry, 1916: 275; Rees, Scammon, 1874: 22, plate 10, figure 6.— 1953: 34; Rice, 1963: 185; Scarff, 1986: 129; Slijper, 1962: 90; Hayashi, 2012: 110, plate 2b, figures 4 & 15d. Stephensen, 1938: 7; Stewart et al., 2011: 3685; Tarasov & Cryptolepas rhachianectis Gruvel, 1903: 153, Zevina, 1957: 244; Tomilin, 1957: 62 (as Coronula sp.); plate 2, figures 4–10; plate 3, figures 201 & 2011.—1905: 274, figure 299. Weltner, 1897: 254; 1898: 11; Wolff, 1960: 8; Young, 1991: references 194; 1998: 273; Zullo, 1963: 14. Achituv, 1998: 1204; Briggs & Morejohn, 1972: 287; Clarke, 1966: 157; Cornwall, 1955a: 49; 1955b: 44; Dall, 1872: 300; distribution: Fertl & Newman, 2009: 90; Ford & Reeves, 2008: 64; Frick Cosmopolitan (without Mediterranean). et al., 2011: 637; Genzer & Marmur, 2008: 745; Geraci & Aubin, 1987: 407; Grutter, 2002: 71; Gruvel, 1903: 153; Genus †Cetolepas Zullo, 1969 1905a: 274; Hayashi, 2012: 110; Hiro, 1935: 227; 1936a: 62; †Cetolepas hertleini Zullo, 1969 1936b: 318; Hoek, 1883: 7; Jones et al., 1984: 34; Kasuya & †Cetolepas hertleini Zullo, 1969: 17, figures Rice, 1970: 42; Killingley, 1980: 759; Kolosva´ry, 1943: 101; 47–72. Korschelt, 1933: 21; Maldini et al., 2010: 460; Monroe, 1981: references 241; Moss & Erlandson, 2010: 3367; Pasternak et al., 2002: Ross & Frick, 2011: 64; Zullo, 1969: 17. 583; Pilsbry, 1916: 279; Rice & Wolman, 1971: 100; Rice, 1963: 186; Ridgway et al., 1997: 377; Roest, 1970: 411; Ross Genus Cetopirus Ranzani, 1818 & Frick, 2011: 64; Samaras, 1989: 85; Samaras & Durham, Cetopirus complanatus (Mo¨rch, 1852) 1985: 113; Scammon, 1874: 22; Scarff, 1986: 130; Seilacher, Lepas balaenaris Spengler, 1790: 187. 2005: 33; Sokolov & Arsen’ev, 2006: 56; Swartz, 1981: 360; Coronula balaenaris Dufresne, 1802: 473, plate 30, figures Swartz et al., 2006: 78; Takeda & Ogino, 2005: 151; Tarasov 2–4.—Lamarck, 1818: 387.—Darwin, 1854: 415, plate 15, & Zevina, 1957: 246; Tomilin, 1957: 323; Visscher, 1928: figure 2, 2b; plate 16, figures 1–4. 328; Walker, 1983: 433; Weller et al., 1999: 1213; Cetopirus balaenaris Ranzani, 1818: 87. Wellington & Anderson, 1978: 292; Weltner, 1897: 278; Lepas quinquevalvis Mawe, 1823: 5, plate 2, figure 3. Zullo, 1961: 13; 1969: 17. Coronula balanarum Blainville, 1824: 380. Polylepas (Cetopirus) vulgaris Gray, 1825: 105. distribution Coronula darwini Stebbing, 1910: 572. Pacific Ocean. Coronula complanata Pilsbry, 1916: 276, plate 63, figures 1–3a.–Broch, 1924b: 204.—Nilsson-Cantell, 1930c: 257.— Genus Tubicinella Lamarck, 1802 Cornwall, 1953: 83, figure 6.—Stubbings, 1967: 300.— Tubicinella cheloniae Monroe & Limpus, 1979 Seilacher, 2005: 29, figure 2. Stephanolepas muricata Nillson-Cantell, 1932: 258, Coronula complanatus Hiro, 1936a: 62. plates 1–3.—Hendrickson, 1958: 524, plate 10, Cetopirus complanatus Scarff, 1986: 137, figure 7. figure c.—Liu & Ren, 2007: 323, figure 144. references Tubicinella cheloniae Monroe & Limpus, 1979: 199, Barnard, 1924: 95; Best, 1991: 585; Blainville, 1824: 380; figure 6; plate 1, figures 5 &6.—Monroe, 1981: Broch, 1924b: 204; Cornwall, 1953: 83; Darwin, 1854: 415; 241.—Hayashi, 2012: 112, plate 2c, figures 5 & 15e. 1855: 39; Dominici et al., 2011: 97; Dufresne, 1802: 473; Chelolepas cheloniae Ross & Frick, 2007: 3, figures 1d, Fertl & Newman, 2009: 90; Gray, 1825: 105; Gruvel, 1903: e & 2a. 152; 1905a: 271; Guiler, 1956: 3; Haelters et al., 2010: 96; references Hiro, 1936a: 62; 1936b: 318; Holthuis et al., 1998: 349; Anderson, 1994: 91; Dobbs & Landry, 2004: 674; Flint et al., Jones, 2003: 484; Jones et al., 1990: 12; Kolosva´ry, 1942a: 2009: 741; 2010: 301; Hayashi, 2012: 112; Hendrickson, 141; 1943: 100; Mawe, 1823: 5; Monroe, 1981: 241; Mo¨rch, 1958: 524; Jones, 2003: 484; 2010: 200; Jones et al., 1990: 13; 1852: 67; Murray, 1895: 449; Newman et al., 1969: 289; Kirkwood & Hooper, 2004: 1; Limpus et al., 1994: 147; Liu Nilsson-Cantell, 1930c: 257; 1931: 116; 1938: 14; 1978: 79; & Ren, 2007: 323; Loop et al., 1995: 246; Monroe, 1981: 241; Pastorino & Griffin, 1996: 769; Pilsbry, 1916: 276; Ranzani, Monroe & Limpus, 1979: 197; Nilsson-Cantell, 1932: 258; 1818: 87; Ross & Frick, 2011: 64; Scarff, 1986: 130; Seilacher, Ross & Frick, 2007: 2; 2011: 63. 2005: 29; Stebbing, 1910: 572; Stubbings, 1967: 300; Tarasov & Zevina, 1957: 245; Weisbord, 1971: 94; Weltner, 1897: distribution 254; 1898: 8; 1900: 307; Zullo, 1961: 13; 1969: 18. Indo-Western Pacific. checklistofturtleandwhalebarnacles 165 remarks 407; Gonza´lez et al., 2012: 132; Greenwood et al., 1979: 24; Ross & Frick (2007) revised the taxonomic position of T. che- Gruvel, 1903: 159; 1905a: 280; 1920: 55; Guerrero-Ruiz & loniae and erected a new genus Chelolepas for this species Urba´n, 2000: 148; Hayashi, 2012: 113; Heldt, 1950: 25; (Ross & Frick, 2007). However this nomenclatural act was Heyning, 1989: 303; Heyning & Dahlheim, 1988: 4; Hiro, published in Marine Turtle Newsletter. The journal is open 1936a: 62; 1936b: 318; Hoek, 1909: 271; Holmes et al., 1997: to the public via a website while this is not available work 153; Jones, 2010: 210; Jones et al., 1990: 13; Kane et al., under the International Code of Zoological Nomenclature 2008: 395; Karuppiah et al., 2004: 879; Kolosva´ry, 1943: 100; 4th edition (Article 9). This nomenclatural act cannot be vali- Kru¨ger, 1911a: 59; MacKintosh & Wheeler, 1929: 373; dated and this was mentioned in Hayashi (2012), too. So, Maisey, 1978: 660; Matthews, 1938b: 238; Mead, 1975: 1157; Chelolepas is the void nomenclature and this species should Mead & Potter, 1990: 177; Monroe, 1981: 241; Newman & be listed as Tubicinella cheloniae. Ross, 1971: 180; Nilsson-Cantell, 1921: 375; 1930c: 258; 1978: 81; Orams & Schuetze, 1998; O’Riordan, 1967: 295; Tubicinella major Lamarck, 1802 Palacios et al., 2004: 129; Parsons et al., 2001: 776; Perrin, Tubicinella (major) Lamarck, 1802: 463, plate 30, figure 2001: 4; Pilleri 1970: 248; Pilleri & Knuckey, 1969: 66; 1.–Pilsbry, 1916: 281, plate 65, figure 1. Pilsbry, 1916: 283; Pinedo et al., 1989: 48; Pope, 1958: 159; Tubicinella (minor) Lamarck, 1802: 463, plate 30, figure 1. Pugliese et al., 2012; Rajaguru & Shantha, 1992: 197; Raga & Lepas trachealis Shaw, 1806: 726. Carbonell, 1985: 210; Raga & Sanpera, 1986: 489; Raga Lepas tracheaeformis Wood, 1815: 31, plate 4, figures 1–3. et al., 1982: 1; Rappe´, 1988: 100; Resendes et al., 2002: 206; Tubicinella anulata Ranzani, 1818: 89. Reyes & Van Waerebeek, 1995: 349; Ribeiro et al., 2010: 1; Tubicinella balaenarum Lamarck, 1818: 385. Rice, 1963: 184; Richard, 1936: 55; Richard & Neuville, Tubicinella balaenae Oken, 1821: 659. 1897: 108; Ross & Frick, 2011: 64; Rotstein et al., 2009: 589; Tubicinella trachealis Darwin, 1854: 431, plate 17, figure Sakai M. et al., 2006: 975; Sakai Y. et al., 2009: 81; Samaras, 3a–3e.—Hutton, 1879: 330.—Weltner, 1897: 253.— 1989: 85; Sandison, 1950: 312; Scarff, 1986: 130; Seilacher, Marloch, 1900: 1.—Stebbing, 1900: 62.—Gruvel, 1903: 2005: 32; Spivey, 1977: 8; 1980: 292; 1981: 155; Stebbing, 148.1905a: 278. 1923: 12; Steenstrup, 1852: 64; Stubbings, 1965: 902; 1967: Tubicinella striata Stebbing, 1910: 573.—Barnard, 1924: 95. references 301; Subba Rao, 2005: 121; Tarasov & Zevina, 1957: 250; Tomilin, 1957: 107; Toth-Brown & Hohn, 2007: 1271; Barnard, 1924: 95; Darwin, 1854: 431; Fertl & Newman, 2009: True, 1891; Uchida & Araki, 2000: 85; Van Bressem et al., 90; Ford & Reeves, 2008: 64; Gruvel, 1903: 148; 1905a: 278; 1999: 54; Van Waerebeek et al., 1993: 159; Watson & Gee, Hiro, 1936a: 62; 1936b: 318; Huttons, 1879: 330; Innocenti, 2005: 448; Weir, 2010: 4; Weltner, 1897: 253; 1898: 11; 2006: 7; Kenney 2002: 808; 2009: 964; Kolosva´ry, 1943: 100; Wilson et al., 1999: 1079; Young, 1991: 193; 1998: 273; Lamarck, 1802: 463; Marloth, 1900: 1; Monroe, 1981: 239; Zullo, 1963: 15. Monroe & Limpus, 1979: 201; Marloch, 1900: 1; Mo¨rch, 1852: 66; Nilsson-Cantell, 1921: 373; 1931: 116; 1957: 8; Oken, 1821: 659; Pilsbry, 1916: 281; Ranzani, 1818: 89; Reeb distribution et al., 2007: 600 (Tubicinella sp.); Reeves & Kenney, 2003: Cosmopolitan. 439; Ross & Frick, 2011: 64; Scarff, 1986: 129; Seilacher, Family PLATYLEPADIDAE Newman & Ross, 1976 2005: 31; Shaw, 1806: 726; Stebbing, 1900: 62; 1910: 573; Weltner, 1897: 253; 1898: 7; 1900: 307; Wood, 1815: 31; Zullo, 1963: 15. Genus Platylepas Gray, 1825 Platylepas coriacea Monroe & Limpus, 1979 distribution Platylepas coriacea Monroe & Limpus, 1979: North and South Atlantic Ocean; Southern Ocean. 208, figure 12; plate 5, figures 1–8. references Genus Xenobalanus Steenstrup, 1852 Anderson, 1994: 298; Biasatti, 2004: 206; Gramentz, 1988: 33; Xenobalanus globicipitis Steenstrup, 1852 Jones, 2010: 200; Jones et al., 1990: 12; Kirkwood & Hooper, Xenobalanus globicipitis Steenstrup, 1852: 64, plate 17, figure 2004: 1; Monroe & Limpus, 1979: 197; Ross & Frick, 2011: 62. 4a–c.—Hayashi, 2012: 113, plate 2d, figure 6. Xenobalanus natalensis Stebbing, 1923: 12, plate 16. distribution references Western Pacific; Atlantic. Addink & Smeenk, 2001: 42; Aguilar & Raja, 1993: 527; Anderson, 1994: 298; Aznar et al., 1994: 159; 2005: 239; Platylepas decorata Darwin, 1854 Baird et al., 1993: 461; Bane & Zullo, 1980: 2; Barnard, Platylepas decorata Darwin, 1854: 429, plate 17, 1924: 96; Barnes & Klepal, 1971: 88; Bassindale, 1964: 43; figure 2a, b.—Hayashi, 2012: 114, figure 8; plate Bearzi & Patonai, 2010: 37; Bellido et al., 2006: 38; Broch, 3b. 1924a: 95; Brownell, 1988: 7; Calman, 1920: 165; Carvalho Platylepas hexastylos ichtyophila Pilsbry, 1916: et al., 2010: 116; Clarke, 1966: 155; Cornwall, 1927: 510; 287, plate 67, figure 2.—Henry, 1954: 444. 1928: 9; 1955a: 56; 1955b: 46; Danilewicz et al., 2002: 31; Platylepas multidecorata Daniel, 1962: 641, Darwin, 1854: 440; Devaraj & Bennet, 1974: 579; Dhermain figures 1 & 2. et al., 2002: 5; Di Beneditto & Ramos, 2000: 159; 2001: 185; references 2004: 1245; Dollfus, 1968: 55; Duignan et al., 1992: 243; Anderson, 1994: 297; Bugoni et al., 2001: 7; Daniel, 1962: 641; Fertl, 1994: 308; Fertl et al., 1996: 606; Foote et al., 2011: Darwin, 1854: 429; Farrapeira, 2010: 212; Fertl & Newman, 631; Gannon & Waples, 2004: 528; Geraci & Aubin, 1987: 2009: 91; Frick & Zardus, 2010: 292; Frick et al., 2010b: 169; 166 ryota hayashi

2011: 638; Hayashi, 2012: 114; Gruvel, 1905a: 276; 1912: 350; distribution Henry, 1954: 444; Hiro, 1936a: 61; 1936b: 319; 1937a: 70; Indo-Pacific; Mediterranean; Atlantic Ocean. Jones, 2003: 484; 2004: 140; 2010: 211; Jones et al., 1990: 12; 2000: 260; Korschelt, 1933: 22; Lazo-Wasem et al., 2011:222; Platylepas hexastylos ichtyophila Pilsbry, 1916 Limpus et al., 1983a: 192; 1994: 147; Liu & Ren, 2007: 314; Platylepas hexastylos ichtyophila Pilsbry, 1916: 287, Monroe, 1981: 237; Monroe & Limpus, 1979: 206; plate 67, figure 2.—Young, 1991: 197, figure 2g, h. Nilsson-Cantell, 1921: 376; Pilsbry, 1916: 285; Ren, 1980: references 190; Ross, 1963a: 153; Ross & Frick, 2011: 62; Ryder, 1879: Farrapeira, 2010: 212; Henry, 1954: 444; Hiro, 1936a: 61; 453; Utinomi, 1970: 363; Wells, 1966: 87; Weltner, 1897: Pilsbry, 1916: 287; Ross, 1963a: 156; Wells, 1966: 87; Young, 253; Young, 1991: 194; 1998: 272; Zullo, 1968: 225; 1991: 175. 1991: 195; 1998: 272; Zullo, 1968: 225. distribution distribution Paraiba, Brazil; Hernando County, Florida, USA. Indo-Pacific; Atlantic Ocean. Platylepas indicus Daniel, 1958 Platylepas hexastylos (Fabricius, 1798) Platylepas indicus Daniel, 1958: 755. Lepas hexastylos Fabricius, 1798: 35, plate 10, references figures 1 & 2. Daniel, 1958: 755; Ross & Frick, 2011: 62. Coronula bisexlobata Blainville, 1824: 379, plate 117, figure 1. distribution Platylepas pulchra Gray 1825: 105 Indian Ocean. Platylepas bissexlobata Darwin, 1854: 428, plate 17, figure 1a–d.—Gruvel, 1903: 151, plate 3, Platylepas ophiophilus Lanchester, 1902 figure 13. Platylepas -? Darwin, 1854: 430. Platylepas hexastylos Pilsbry, 1916: 285, plate 67, Platylepas ophiophilus Lanchester, 1902: 371, plate 35, figures 1, 1c & 3.—Hayashi, 2012: 114, figure 7; figure 5, 5b. plate 3a. Cryptolepas ophiophilus Kru¨ger, 1912: 12. references Platylepas kru¨geri Broch, 1931: 122. Platylepas ophiopholis Nilsson-Cantell, 1938: 77. Aguirre et al., 1994: 12; Alexander et al., 2004: 14; Alonso Platylepas ophiophila Utinomi, 1970: 360, plate 18, figure 12. et al., 2010: 167; Anderson, 1994: 297; Angulo-Lozano et al., Platylepas ophiophia Ren, 1980: 191.—Liu & Ren, 2007: 2007: 14; Bacon, 1976: 34; Balazs, 1985: 17; Barnes & Klepal, 316, figure 140. 1971: 89; Beck & Forrester, 1988: 633; Broch, 1924a: 18; references 1924b: 203; 1927a: 30; Bugoni et al., 2001: 7; Chan et al., Anderson, 1994: 175; Broch, 1931: 122; Darwin, 1854: 430; 2009: 176; Chan & Hayashi, in press; Chevreux & de Frick et al., 2011: 638; Gruvel, 1905a: 277; Hiro, 1936a: 61; Guerne, 1893: 443; Crisp, 1967: 136; Daniel, 1955: 33; 1936b: 319; Jones, 2003: 484; 2010: 213; Jones et al., 1990: Darwin, 1854: 428; Eckert & Eckert, 1987: 684; 1988: 401; 12; Korschelt, 1933: 22; Kru¨ger, 1912: 12; Lanchester, 1902: Eldredge & Miller, 1997: 11; Fabricius, 1798: 35; Farrapeira, 371; Liu & Ren, 2007: 316; Nilsson-Cantell, 1921: 376; 1938: 2010: 212; Fertl & Newman, 2009: 91; Fischer 1884: 359; 77; Pilsbry, 1916: 285; Ren, 1980: 191; Ross, 1963a: 153; Foster, 1978: 117; Frazier, 1971: 394; 1989: 250; Frick et al., Ross & Frick, 2011: 62; Utinomi, 1970: 360; Zann, 1975: 2011: 629; Glazebrook & Campbell, 1990: 102; Gonza´lez 267; Zann & Harker, 1978: 206. et al., 2012: 132; Gramentz, 1988: 35; Gruvel, 1903: 151; 1905a: 276; Harding et al., 2011: 236; Hayashi, 2009: 2; distribution 2012: 114; Hayashi et al., 2011; Henry, 1954: 444; Indo-West Pacific. Hernandez-Vazquez & Valadez-Gonzalez, 1998: 121; Hiro, 1936a: 61; 1936b: 312; 1937a: 70; 1937b: 472; Holmes et al., †Platylepas wilsoni Ross, 1963 1997: 153; Holthuis, 1952: 72; Innocenti, 2006: 7; Jones, †Platylepas wilsoni Ross, 1963: 153, figures 2 & 3. 2003: 484; 2004: 140; 2010: 199; Jones et al., 1990: 12; Kitsos et al., 2005: 259; Kolosva´ry, 1943: 101; 1951: 412; Korschelt, references 1933: 22; Koukouras & Matsa, 1998: 136; Kru¨ger, 1912: 13; Ross, 1963a: 153; 1963b: 229 (as Platylepas sp.); 1964: 278; Limpus et al., 1983a: 192; Liu & Ren, 2007: 312; Losey et al., Ross & Frick, 2011: 62; Utinomi, 1970: 363. 1994: 686; Lucas, 1968: 132; Lutcavage & Musick, 1985: 453; Mead & Potter, 1990: 177; Monroe, 1981: 240; Monroe & Genus Cylindrolepas Pilsbry, 1916 Limpus, 1979: 205; O’Riordan, 1979: 356; Pilsbry, 1916: 285; Relini, 1969: 169; Ren, 1980: 188; Richards, 1930: 143; Ross, Cylindrolepas darwiniana Pilsbry, 1916 1963a: 156; Ross & Frick, 2011: 62; Ryder, 1879: 453; Cylindrolepas darwiniana Pilsbry, 1916: 288, plate Sandison, 1950: 312; Schwartz, 1960: 116; Spivey, 1981: 155; 68, figure 3 & 3b.—Hayashi, 2012: 115, figures 9 & Stubbings, 1965: 876; 1967: 300; Tachikawa, 1995: 67; 15f; plate 3c. Utinomi, 1950: 62; 1959: 384; 1970: 360; Vivaldo et al., references 2006: 431; Walker, 1983: 433; Wass, 1963: 36; Wells, 1966: Frick & Zardus, 2010: 292; Frick et al., 2010b: 169; Hayashi, 87; Weltner, 1897: 253; Young, 1991: 195; 1998: 272; Zann, 2012: 115; Hiro, 1936a: 61; 1936b: 319; Hubbs, 1977: 265; 1975: 268; Zann & Harker, 1978: 206; Zavodnik, 1997: 116; Jones et al., 1990: 12; Monroe, 1981: 237; Pilsbry, 1916: 288; Zullo, 1963: 15; Zullo & Bleakney, 1966: 164; Zullo & Lang, Ren, 1980: 195; Ross & Frick, 2011: 62; Zullo, 1968: 223; 1978: 160. 1991: 175. checklistofturtleandwhalebarnacles 167 distribution Stomatolepas dermochelys Monroe & Limpus, 1979 Pacific Ocean; Atlantic Ocean. Stomatolepas elegans Zullo & Bleakney, 1966: 162, figure 2.—McCann, 1969: 152, figures 3—5.—Smaldon & Cylindrolepas sinica Ren, 1980 Lyster, 1976: 317. Cylindrolepas sinica Ren, 1980: 194, figure 6; plate 2, figures Stomatolepas dermochelys Monroe & Limpus, 1979: 203, 12–20.—Hayashi, 2012: 118, figures 10 & 15g; plate 3d. figure 9; plate 3, figures 1–5. Platylepas decorata Zardus & Balazs, 2007: 1303, figures Stomatolepas eretomochelys Jones 2010: 200. 7–9. references references Alexander et al., 2004: 14; Anderson, 1994: 297; Brongersma, Frick & Zardus, 2010: 294; Hayashi, 2009: 1; 2012: 118; Liu & 1972: 60; Carriol & Vader, 2002: 82; Eckert & Eckert, 1987: Ren, 2007: 321; Ren, 1980: 194; Ross & Frick, 2011: 62; Zardus 684; 1988: 401; Gitting et al., 1986: 33; Haelters & Kerckhof, & Balazs, 2007: 1303. 1999: 30; Haelters et al., 2001: 14; Hayashi, 2012: 124; Holthuis, 1969: 44; Jones, 2010: 200; Jones et al., 1990: 12; distribution McCann, 1969: 152; Monroe & Limpus, 1979: 197; Ross & Pacific Ocean. Frick, 2011: 63; Smaldon & Lyster, 1976: 317; Zullo & Bleakney, 1966: 162. Genus Stomatolepas Pilsbry, 1910 distribution remarks Western Pacific Ocean; Atlantic Ocean. Genus Stomatolepas was described by Pilsbry (1910) for a new species S. praegustator, collected from the gullet of a logger- Stomatolepas elegans (Costa, 1838) head sea turtle Caretta caretta, at Tortugas, Florida. Prior to Coronula elegans Costa, 1838: 17, plate 1, figures this work, another species S. elegans had been described as 1–3. Coronula elegans by Costa (1838) from Taranto, Italy. Chelonobia elegans Costa, 1939: 14. Pilsbry (1916) regarded this species also to be included in Stomatolepas elegans Pilsbry, 1916: 289, plate 68, the genus Stomatolepas. He suspected that S. praegustator figure 2, 2a. might be identical with Costa’s S. elegans, but hesitated to references make that determination, because Costa’s description of S. Bugoni et al., 2001: 7; Carriol & Vader, 2002: 82; Costa, 1838: elegans was imperfect. Afterwards, Nilsson-Cantell (1930b) 17; 1839: 14; Farrapeira 2010: 212; Frick et al., 2010a: 123; described a new species with transversely elongated imbricat- Gitting et al., 1986: 33; Gonza´lez et al., 2012: 132; Hiro, ing scales, named as S. transversa. Stomatolepas transversa is 1936a: 61; 1936b: 312; 1937b: 473; Holmes et al., 1997: 153; easily distinguished by the shell and the scales Jones, 2003: 484; Kitsos et al., 2003: 407; 2005: 259; (Nilsson-Cantell, 1930b; Monroe & Limpus, 1979; Young, Lanfranco, 1979: 24; Lazo-Wasem et al., 2011: 222; Liu & 1991; Hayashi, 2012). While, Hiro (1936b) regarded S. trans- Ren, 2007: 317; Nilsson-Cantell, 1930b: 20; O’Riordan & versa as a synonym of S. elegans, Ren (1980) described other Holmes, 1978: 152; Pilsbry, 1916: 289; Quigley & Flannery, similar looking specimens as S. pulchra. Among these barna- 1993: 153; Ren, 1980: 193; Ross & Frick, 2011: 63; Spivey, cles, Stomatolepas transversa and S. pulchra have been well- 1981: 155; Stubbings, 1965: 876; 1967: 300; Utinomi, 1970: described in shell structure and soft-part morphology by 363; Walker, 1983: 433; Wirtz et al., 2006: 209; Young, 1991: many authors (Monroe & Limpus, 1979; Ren, 1980; Young, 193; 1998: 272; Zardus & Balazs, 2007: 1303; Zullo, 1991; Hayashi, 2012); it is clear that S. pulchra is a junior 1968:227; Zullo & Lang, 1978: 160. synonym of S. transversa. Monroe & Limpus (1979) reviewed the complicated taxonomy of Stomatolepas and indicated the distribution missing types and inadequate descriptions of S. elegans by Costa (1838). However, they hesitated to select a neotype for Pacific Ocean; Mediterranean; Atlantic Ocean. S. elegans and described S. dermochelys as new, while regarding Stomatolepas pilsbryi Frick, Zardus and Laso-Wasem, 2010 S. elegans as a nomen dubium. According to Winston (1999), Stomatolepas pilsbryi Frick, Zardus and Laso-Wasem, 2010: anyone working on a species that resembles one in a question- 127, figsures 7–19. able illustration or an unclear old description would do better to references describe the species as new, making a note of the early name as Frick et al., 2010a: 127; Ross & Frick, 2011: 63. a possibility. Winston believed that should his original speci- mens someday be discovered, restudy might result in its distribution name becoming a junior synonym, but at least the history of Gabon, Africa. the name and the decision would be clear. Also, Quicke remarks (1993) stated that a neotype specimen should come from the same locality as the original as much as possible. After a few Frick et al. (2010a) described this species using soft-part mor- decades, Frick et al. (2010a) redescribed S. elegans with a phology, however comparison with other species was made neotype; however, this neotype was collected from Nova only in terms of the parietal wall and attachment habitat. Scotia whereas Costa’s (1838) original S. elegans was from Stomatolepas praegustator Pilsbry, 1910 Taranto, Italy. A revision of Stomatolepas species is certainly needed, but in this checklist I rely on the decisions of previous Stomatolepas praegustator Pilsbry, 1910: 304, figure authors, particularly Monroe & Limpus (1979) in honour of 1.—1916: 289, plate 68, figure 1, 1a & 1b.— their detailed and exhaustive observations including shell and Hayashi, 2012: 118, figures 11 & 15; plate 4a. soft-part morphology. Stomatolepas elegans Hiro, 1936b: 312, figures 1–5. 168 ryota hayashi

references references Anderson, 1994: 297; Carriol & Vader, 2002: 82; Frick et al., Frick et al., 2010b: 169; Ross & Frick, 2011: 63. 2010a: 123; Hayashi, 2012: 118; Henry, 1954: 444; Hiro, 1936b: 312; Holthuis, 1969: 44; Jones, 2003: 484; 2010: 213; distribution Jones et al., 1990: 12; Lazo-Wasem et al., 2007: 153; Atlantic Ocean. Monroe, 1981: 241; Monroe & Limpus, 1979: 202; Nilsson-Cantell, 1930a: 2; 1932: 257; Pilsbry, 1910: 304; remarks 1916: 289; Relini, 1968: 223; 1969: 169; Ren, 1980: 193; Ross Frick et al. (2010b) described this species as new and simul- & Frick, 2011: 63; Vivaldo et al., 2006: 431; Wells, 1966: 87; taneously placed it in a new genus and subfamily, however, Zullo, 1968: 227; Zullo & Bleakney, 1966: 163. no description of the soft parts was provided. distribution Pacific Ocean; Atlantic Ocean. ACKNOWLEDGEMENTS Stomatolepas transversa Nilsson-Cantell, 1930 Stomatolepas transversa Nilsson-Cantell, 1930: 20.— The comments and suggestions of Dr Benny K.K. Chan Monroe & Limpus, 1979: 202, plate 6, figures 1–6.— (editor), Professor John Buckeridge, Dr Diana Jones and an Young, 1991: 202, figure 4a–k.—Hayashi, 2012: 119, anonymous scientist (referees) improved the quality of the figure 12; plate 4b. manuscript. This study was supported by the programme Stomatolepas pulchra Ren, 1980: 191, figures 4 & 5; ‘Bio-Logging Science of the University of Tokyo (UTBLS)’. plate 2, figures 5—11.—Liu & Ren, 2007: 318, figure 142. references CORONULOIDEA LITERATURE CITED Anderson, 1994: 297; Bugoni et al., 2001: 7; Carriol & Vader, 2002: 82; Farrapeira, 2010: 212; Hayashi, 2012: 119; Hayashi & Achituv Y. (1998) Cirral activity in the whale barnacle Cryptolepas rha- Tsuji, 2008: 122; Jones, 2010: 214; Jones et al., 1990: 12; chianechi. Journal of the Marine Biological Association of the United Limpus et al., 1994: 147; Monroe & Limpus, 1979: 202; Liu Kingdom 78, 1203–1213. & Ren, 2007: 318; Nilsson-Cantell, 1930b: 20; Ren, 1980: 191; Ross & Frick, 2011: 63; Stubbings, 1967: 301; Young, Addink M.J. and Smeenk C. (2001) Opportunistic feeding behaviour of rough-toothed dolphins Steno bredanensis off Mauritania. 1991: 210; 1998: 272. Zoologische Verhandelingen 334, 37–48. distribution Aguilar A. and Raja J.A. (1993) The striped dolphin epizootic in the Mediterranean Sea. AMBIO 22, 524–528. Pacific Ocean; Atlantic Ocean. Aguirre A.A., Balazs G.H., Zimmerman B. and Spraker T.R. (1994) Genus Stephanolepas Fischer, 1886 Evaluation of Hawaiian green turtles (Chelonia mydas) for potential pathogens associated with fibropapillomas. Journal of Wildlife Stephanolepas muricata Fischer, 1886 Diseases 30, 8–15. Alexander J., Garrett K., Garner S.A., Conrad J. and Coles W. (2004) Stephanolepas muricata Fischer, 1886: 193, plate Tagging and nesting research on leatherback sea turtle (Dermochelys 4, figures 9–11.—Frick et al., 2011: 629, figures coriacea) on Sandy Point, St Croix, US Virgin Islands, 2004. Annual 1–3.—Hayashi, 2012: 121, figures 13 & 14; plate Report to Fish and Wildlife Service, 40 pp. 4c, 4d. Alonso L., Estradesm A., Scarabino F. and Calcagno J. (2010) references Conchoderma virgatum (Spengler, 1790) (Cirripedia: Pedunculata) Aguirre et al., 1994: 12; Anderson, 1994: 297; Balazs, 1978: associated with sea turtles in Uruguayan shallow coastal waters. 128; Broch, 1947: 7; Bustard, 1976: 347; Dawydoff, 1952: Pan-American Journal of Aquatic Sciences 5, 166–168. 128; Deraniyagala, 1939: 204; Eldredge & Miller, 1997: 11; A´ lvarez-Ferna´ndez E. 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