Great Basin Naturalist

Volume 47 Number 1 Article 6

1-31-1987

Robber of (Diptera: )

C. Riley Nelson Utah State University

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Recommended Citation Nelson, C. Riley (1987) "Robber flies of Utah (Diptera: Asilidae)," Great Basin Naturalist: Vol. 47 : No. 1 , Article 6. Available at: https://scholarsarchive.byu.edu/gbn/vol47/iss1/6

This Article is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Great Basin Naturalist by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. ROBBER FLIES OF UTAH (DIPTERA: ASILIDAE)

C. Riley Nelson'

Abstract —Reported are 158 of Asilidae (Diptera) in 50 genera from Utah. Keys to subfamilies genera and species are given, along with information on seasonal and distributional occurrence in Utah. Seventy-six maps and 56 line drawings show the Utah distribution of each species and illustrate important characters used in the keys. A table summarizes the current status of names used in earlier state lists.

The Asilidae (Diptera) have long attracted lections of asihds from the state revealed 158 the attention of collectors. As a result, numer- species in 50 genera. ous specimens have been deposited in collec- tions in the state of Utah. The systematics of Review of Literature the have been studied rather inten- sively so that the taxonomic status of most Brown (1929) pubhshed the first paper groups is known. The purpose of this study is dealing with Asilidae of Utah. He listed 22 to assemble the numerous records and to species of robber flies as occurring in Utah, identify the species of asilids caught in Utah. one of which was not determined to species. The catalog of Diptera (Stone et al. 1965) He included drawings of the genitalia of most shows that about 50 species of asilids are ei- of these species. Table 1 summarizes the cur- ther listed as occurring in Utah or are in- rent status of the species listed by Brown cluded in the broad geographical ranges de- (1929). lineated under a given species. A preliminary Knowlton and Harmston (1938) published a examination of the specimens from the Utah list of Asilidae found in Utah. A summary of State University Entomological Museum re- the current status of these species is given in vealed that identifications made by prominent Table 2. workers of the family existed for over 150 A further list of asilids captured in Utah was different species. A careful examination of col- given by Knowlton, Harmston, and Stains

Table 1. Current status of species listed in Brown (1929).

1. abdominalia (Say) = O. abdominalis 2. modestiis Loew = S. inqiiinattis 3. Stenopogon consanguineus Loew Specimens are neglectus 4. Stenopogon californiae (Walker) Specimens are S. rufibarbis 5. Stenopogon sp. female Not enough information 6. ludius (Coquillett) Wilcox lists as H. senilis 7. Dasijllisfernaldi Back = Laphriafernaldi 8. Deromijia bigoti (Bellardi) = grossus 9. Proctacanthiis arno Townsend = P. nearno 10. Erax harhatus (Fabricius) = albibarbis 11. Erax interniptus (Macquart) = interrupta 12. Erax stamineus WiUiston = Efferia staminea 13. Erax sidjpilosns SchaefFer Neither Wilcox (1966) nor I saw specimens of this species. 14. fulviventris Macquart Unrecognizable to Cole and Pritchard (1964). 15. Mallophora guildiana Williston = Megaphonis guildiana 16. bastardii (Macquart) = P. dimidiattis 17. Promachus nigripes Hine Specimens are P. uldrichii. 18. Asihi.s lecythiis Walker Utah specimens are occidentalis. 19. Asihis tenehrosits Williston = Machimus ariseus

'Department of Biology, Utah State University, Logan, Utah 84322. Present address: M. L. Bean Museum. Brigham Young University, Provo, UtahS4602.

38 39 January 1987 Nelson: Utah Robber Flies

and ILirmston (1938). Table 2. Changes in status of species listed in Knowlton

Utah specimen is Omninablauttis nigronotu)n L Ablaiitus trifariiis Loevv = Cerotainiops abdominalis 2. abdominalis (Brown) = MacJiimus callidus 3. Asihis callidus = Polacantha coniposita 4. A. compositus Hine Specimens are Machimus paropus 5. A. erijthrocnemius Hine Specimens are Polacantha coniposita 6. A. grad/wsWeidemann = Negasilus mesae 7. A. mesae Tucker = Machimus occidentalis 8. A. occidentalis Hine = Machimus griseus 9. A. f(?nc/jrosH.s- Williston = fernaldi 10. Bomboinima fernaldi (Back) specimens seen; this is a species; Utah speci- 11. callipedihis Loew No mens were probably C. montanus Diogmites grossus 12. Deromyia bigoti Bellardi Probably D. henshawi 13. parvula Coquillett Specimen is E. benedicti 14. Efferia Candida Coquillett spp. 15. Erax spp. Efferia Utah specimens are Efferia benedicti 16. Erax argyrosoma Hine = albibarbis 17. Erax barbatus Fabricius Efferia specimen seen, specimen was probably Efferia benedicti 18. E. californicus No Nomen nudum 19. E. (fw/ni/.s Williston = Triorla interrupta 20. E. interruptiis = benedicti 21. E. knotvltoni Bromley Efferia seen in collections 22. £. pallididus Hine Not seen in collections 23. E. texana Banks Not specimen seen 24. Eucyrtopogon macidosus Coquillett No specimen is H. maculinervis 25. Heteropogon lautiis Loew Utah = guildiana 26. Mallophora gitildiana Williston = Mallophora fautrix 27. Mallophora faiitricoides Curran = auriannulatus 28. Neoitamushardyi Bromley = N. utahensis 29. Nicodes punctipennis Melander = O. abdominalis 30. Osprioceriis ventralis (Say) = nearno 31. arno Townsend P. = cacopiloga 32. P. cacopiloga Hine specimens are P. aldrichii 33. Promachus nigripes Hine Utah Eastern species; Utah specimen is P. dimidiatus 34. P. quadratus Weidemann = Ospriocerus longulus 35. Scleropogon longultis Loew is Scleropogon indistinctus 36. Stenopogon helvolus Loew Specimen = Scleropogon neglectus 37. Stenopogon neglectiis Bromley Utah specimens are S. martini 38. Stenopogon obscurivenths Loew = Scleropogon picticornis 39. S. picticornishoew

Methods (1939). A review of the species from that pub- hcation and their current status is given in Utah collections of Asilidae were examined Table 3. State University, Brigham Young Johnson's (1936) unpubhshed master's the- at Utah University, the University of Utah, Southern sis is an important contribution to the study of State College, Dixie College, and the the feuna of the state. He hsted 73 species as Utah of Eastern Utah. Additionally, a occurring in Utah. Table 4 summarizes the College through literature dealing with Asili- changes in the status of the species hsted in search iTiade, and records of specimens col- Johnson (1936). Johnson's specimens are in dae was Utah were included in the synopsis the Brigham Young University collection and lected in were examined during this study. of each species. throughout the state were Numerous other records of asilids caught in Numerous areas and extensive collections of Asilidae Utah are scattered throughout the literature. visited, made during this study. Areas which These are listed unmarked under the discus- were particular attention were: Washing- sions of individual species. received .

40 Great Basin Naturalist Vol. 47, No. 1

Table 3. Changes in status of species listed in Knowlton, Harmston, and Stains (19,39).

1. lauttis Loew Probably Heteropogon senilis 2. Asilus avidus Van der Wulp Specimens are Machimus adustus 3. A. belli Curran = Negasilus belli 4. A. delicatulus Hine Specimen unidentifiable 5. A. enjthrocnemhis Hine Specimens are Machimus paropus 6. A. occidentalis Hine = Machimus occidentalis 7. A. paropw,s Walker = Machimus paropus 8. Bomhomima fernaldi Back = Laphria fernaldi 9. Dioctria pleiiralis Banks Specimens are Dioctria vera 10. Efferia Candida Coquillett Specimens are E. davisi 1 1 Erax harhatus Fabricius = Efferia albibarbis 12. E. bicaudatus Specimens are Efferia frewingi 13. E. dubiiis Williston Nomen nudum 14. E. zonatus Hine = Efferia zonata 15. phaeonotiis Loew Specimens are H. albipilosus 16. H. seniculus Loew Specimens are H. albipilosus 17. sexfasciatus Say Specimen is L. tolandi 18. Mallophora bromleiji Curran = M. faittrix 19. M. purdens Pritchard = Megaphorus willistoni 20. M. pulchra Pritchard = Megaphorus pulcher 21. Neoitamti.s hardiji Bromley = Asilus auriannulatus 22. piinctipennis Melander = N. utahensis 23. Ospriocerus ventralis Coquillett = O. abdominalis (Say) 24. Proctacanthus arno Townsend = P. nearno Martin 25. Promachus nigripes Hine Specimens are P. aldrichii 26. Stenopogon helvohis Loew Specimens are Scleropogon coyote 27. Stenopogon neglectiis Bromley = Scleropogon neglectus 28. Stenopogon obscuriventris Loew Specimens are S. mar-tini 29. S. picticornis Loew = S. picticornis

ton County, the only area of the state where 1. R2+3 ending in costa (Figs. 40, 46) 2 species of Mohave origins could be — Ro+3 joining Rj, closing cell rj (Fig. 43) 3 collected; the Raft River Mountains; Cache 2(1). Abdominal segment 2 elongate, five or more County; Box Elder County; Juab County; times as long as wide; resembling, somewhat, Emery County; Grand County; and San Juan small County. — Abdominal segment 2 shorter, not more than five times as long as wide; not particularly The taxonomic synopsis presented in this re- sembling damselflies paper consists of information regarding origi- 3(2). Antennae without a terminal style, ending nal citation and author, type locality, type l^luiit'y repository, all of which were obtained from — Antennae bearing a terminal style the literature. The county and seasonal distri- butions of each species were taken from speci- Key to the Genera of Utah Leptogastrinae and men labels. The seasonal distributional data Dasypogoninae are expressed as a range of earliest and latest (Modified from Wood known collection dates for Utah specimens. 1981) Keys to subfamilies, genera, and species 1. Abdominal segment 2 elongate, five or more times as long as wide; abdomen very were constructed using published informa- thin and elongate Meigen" tion and new insights. A map was produced — Abdominal segment 2 shorter, not more than showing the distribution of each species five times as long as wide; abdomen generally listed, and 56 figures were drawn using a cam- shorter and stout 9 era lucida on a Wild M8 stereomicroscope. 2(1). Foretibia bearing a terminal, clawlike spur on The figures illustrate important characters ventral surface which is thicker and sigmoid (Figs. 54-55) used in the keys. : 3

Much Keys difficulty was met using the key given by Martin (1957). Although several species of Leptogaster are present in Utah collections, reliable spe- cies determinations were not made. The distribution Key to the of this in Utah is Subfamilies of Utah Asilidae given in Fig. 100. January 1987 Nelson: Utah Robber Flies 41

Table 4. Changes in status of species listed in Johnson (1936).

L Ahlautus squamipes Cole Specimens are A. mimiis = 2. Andrenosoma similis (Brown) Cerotainiops abdoinitialis

3. Asilus avidus Van der Wulp Specimens are Machimus adusfiis

4. A. brevicomtis Hine = bnwicumu.s

5. A. mesae (Tucker) = Negasihis mesae 6. A. occidentalis Hine = Machimus occidentalis

7. A. paropus Walker = Machimus paropus 8. A. feneirosws Williston Specimens are Machimus griseus -— 9. Bombomima fernaldi (Back) Laphria fernaldi 10. Chnjsoceria pictitarsis (Bigot) = pictitarsis 11. texana Bromley No specimens seen 12. Cyrtopogon dasijllis Williston Specimens are C. dasylloides 13. C. leucozona Loew = C. montanus 14. C. nwgafor Osten Sacken Specimens are C. banksi 15. Diogmites pulchra (Back) Specimens are Diogmites grossus 16. Ecthopoda sp. No label; spelling Ecthodopa 17. Erax argyrosoma (Hine) Specimens are Efferia benedicti 18. EraxbarbatusFahricius = Efferia albibarbis (Macquart) 19. Erax bicaudatus Hine Probably Efferia fre wingi 20. Erax Candidas (Coquillett) Specimens are Efferia davisi 21. Erax caneUiis Bromley Specimens are Efferia benedicti 22. Erax cingulatus Bellardi No specimens seen; not Nearctic 23. Erax dubius (Williston) Nomen nudum 24. Erax interruptus (Macquart) = Triorla interrupta 25. Erax pilosus Hine No specimens seen 26. Erax stramineiis Williston Specimens are Efferia benedicti 27. Holopogon senicuhis Loew Specimens are Holopogon albipilosus 28. Laphystia limatuhis Coquillett Specimen is Laphystia rubra 29. L. rufiventris Curran Specimens are Laphystia tolandi 30. Lestomyia fraudigera Williston Specimen is Lestomyia sabulona 31. Mallophora bromleyi Curran = Mallophora fautrix 32. Proctacanthella leucopogon (Will. No specimens seen 33. Proctacanthiis arno Townsend = Proctacanthiis nearno 34. Promachus bastardii (Macquart) Damaged; probably P. dimidiatus 35. P. nigripes Hine Specimens are P. aldrichii 36. Stenopogon californiae (Walker) Specimens are S. rufibarbis neglectus 37. S. helvohis (Loew) Probably Scleropogon

38. S. obscuriventris Loew Specimens are S. rufibarbis

vein; costa; CuAi, first anterior branch of cubitus; Fig. 1. Proctacanthus nearno Martin, right wing: A,, first anal C, h, humeral crossvein; M„ first branch cua,, anterior cubital cell; CuA,, second anterior branch of cubitus; d, discal cell; medial cell; M3, third branch of media; m,, third of media; m,, first medial cell; M,, second branch of media; m., second of second and third branches of radius; r,, third medial cell; R,, first branch of radius; r,, first radial cell; R,.,, fusion of radius; r,, fifth radial cell; Sc, subcosta. radial cell; R4, fourth branch of radius; r„ fourth radial cell; R5, fifth branch 42 Great Basin Naturalist Vol. 47, No. 1

Gibbosity

Gibbosity

Figs. 2-5. Heads of Utah Asilidae; 2, Sfenopogon rufiharbis Bromley, lateral aspect; 3, Stenopo{i,on inquinatus Loew, lateral aspect; 4, monticola Melander, cephalic aspect; 5, Neoitamus brevicomiis(Hine) lateral aspect.

— Foretibia without a terminal differentiated spur — Cell m, closed; one llagellomere 6 on ventral surface 12 6(5). Scutellar bristles present Diogmites Loew 3(2). Differentiated spur of foretibia meeting a — Scutellar bristles absent raised denticulate area on basitarsus (Fig. 54) Blepharepium sonorensis Papavero & Bernardi 4 7(3). Pulvilli present 8 — Differentiated spur of foretibia not meeting a — Pulvilli absent H differentiated denticulate area on basitarsus 8(7). Face inflated on lower (Fig. 55) 7 two-thirds or more; ante- rior mesonotum bearing a manelike crest of 4(3). Face inflated and haired on lower two- hairs on midline Curran thirds Lestotmjia sabulona Osten Sacken — Face flat or slightly rounded; mesonotal crest — Face flat, hair confined to lower one-third ... 5 absent 9 5(4). Cell m3 open; two flagellomeres present, the 9(8). Style of anteapical, in a notch at about second being small, bearing a pit with apical midlength of dorsal surface of flagellomere 1; ab- bristle mohaivki Wilcox . . domen pitted . ruficaudus Curran January 1987 Nelson: Utah Robber Flies 43

— Style apical; abdomen not pitted 10 — Hind femur and tibia not club-shaped, at most slightly expanded near apex 21 10(9). Wings with distinct spots on most crossveins and furcations, otherwise hyaline; male with 21(20). Flagellomeres 1 and 2 lobed at apex only six visible tergites, last two visible ter- Mijelaphus lohicornis Osten Sacken gites widened and covered with dense silvery — All flagellomeres simple, not lobed Nicocle.s Jaennicke Dicolomis sparsipilustim Back — Wings hyaline more washed with brown, 22(19). Pulvilli absent; flattened, scalelike hairs spots, if present, indistinct present on thorax, legs, and wing base Cophiira Osten Sacken Loew 11(7). Presutural dorsocentral bristles present; — Pulvilli present; scalelike hairs absent 23 length less than 5 mm Omninahlaiitus (Hgronofum Wilcox 23(22). Midtibia bearing a pair of stout spines at apex which are directed outward at an angle near — Presutural dorsocentral bristles absent; 90 degrees Callinicus Loew length greater than 7 mm Hodoplujlax hasinncri Pritchard — Midtibia with all apical spines directed dis- tally 24 12(2). Sides of frons greatly divergent toward vertex (Fig. 4) ' 13 24(23). Wing with dark spot on r-m, furcation of R4 at apex of cell d, other spots may — Sides of frons snbparallel or converging to- and R5 and 25 ward vertex I'l also be present pattern not 13(12). Face strongly inflated and haired on lower — Wings hyaline or darkened; spot three-fourths Lasiopogon Loew as above 26

— Face but slightly convex; mystax confined to 25(24). Presutural dorsocentral bristles stout; meso- lower margin Loew notum strongly arched carinatus Wilcox 14(12). Face and frons narrow; head about as wide as high and appearing small in proportion to — Presutural dorsocentral bristles absent; thorax 15 mesonotum not strongly arched Eucijrtopogon Curran — Face and frons wider; head wider than high . 17

15(14). Hypopleuron (katatergite) lacking bristles or 26(24). Face strongly inflated, extending greater than hairs (Fig. 31) Stenopogon Loew length of scape past eye margin when viewed from the side 27 — Hypopleuron bearing bristles or hairs (Fig. 30) 16 — Face flat or but slightly convex 29

16(15). Flagellomere 1 at most one and three-fourths 27(26). Dorsocentrals weak, hairlike length of antennal segments 1 and 2 com- Cyrtopogon Loew

bined; long, slender terminal style present . — Dorsocentrals bristlelike, prominent 28 Scleropogon Loew 28(27). Mystax composed of bristles; male with wing — Flagellomere 1 at least two times length of tip darkened and basal half of wing covered antennal segments 1 and 2 combined; termi- with white bloom . . . Coleomyia alticola James nal style, if present, short . . . Ospriocerus Loew — Mystax composed ofweak hairs; wings ofboth 17(14). R2+3 bent sharply forward near apex, making sexes hyaline nearly a 90 degree angle with wing margin Nannocyrtopogon aristatus ]ames (Fig. 45) Laphijstia Loew Hind tibia club-shaped, broadest near apex, — R2+3 not bent sharply forward to make such an 29(26). as or thicker than hind femur angle with wing margin (Fig. 40) 18 as thick Holopogon Loew 18(17). Basal half of wing darkened; eyes set off from club-shaped but spindle- face along lower margin by grooves running — Hind tibia not broadest near middle, along vertical margin of eyes; CuAi and A, shaped or straight, 30 fused before wing margin thinner than hind femur ittahensis Wilcox Haplopogon 30(29). Hairs of thorax branched .. Heteropogon Loew although may — Basal half ofwing not darkened, — Hairs of thorax straight or crinkled, not be spotted; eyes not set off by vertical branched 31 grooves; CuAo and A; not joining before wing 31(30). Legs and tergites with greenish metallic re- margin (except in Myelaphiis) 19 flections Sintoria cazieri Wilcox 19(18). Mystax limited to lower one-third of face or — Legs and tergites without metallic reflections with only a few hairs near antennae in addi- painteri Wilcox tion to lower mystax 20 — Mvstax extending more than halfway up face 22 Key to the Species of Utah Ablautus Loew

20(19). Hind femur and tibia club-shaped; R4 reach- Including Omninablautus Pritchard ing wing margin anterior to wing tip Dioctria Meigen (Modified from Wilcox 1966c) 44 Great Basin Naturalist Vol. 47, No. 1

Figs. 6-17. Utah Asilidae: 6, male genitalia of Machimits occidentalis (Hine), ventral aspect; 7, male genitalia of Machimus caUidus (Williston), ventral aspect; 8, antenna ofOspriocenis abdominalis (Say); 9, antenna of Ospriocerus vallensis Martin- 10, male genitalia of Regfl.«/i/.s Wanton! Bromley, lateral aspect, E = epandrium; 11, male genitalia of lepidns (Hine), lateral aspect, E = notched epandrium; 12, male genitalia of Lasiopo^on aldrichii Melander, lateral aspect; 13, surstyli of La slop ogon aldrichii Melander from below; 14, male genitalia of Lasiopogon monticola Melander, lateral aspect; 15, surstyli of Lasiopogon monticola Melander from below; 16, surstyli of Lasiopogon cinereus Cole from below; 17, male genitalia of Asilus auriannulatus (Hine), lateral aspect,' E = epan- drium. January 1987 Nelson: Utah Robber Flies 45

sides — Scutellum clothed with long, thin, light hairs 1. Mesonotum with black, shining spots on Omninahlaiitus nig.ronotum Wilcox scitula (Williston)

— Mesonotum entirely pollinose 2 Key to the Species of Utah Cijriopogon Loew scalelike 2(1). Face, frons, thorax, and legs without from Wilcox and Martin 1936) hairs ^ (Modified

— Face, frons, thorax, and legs with scalelike hairs 1. Last segment of male foretarsus extremely 4 flattened albifacies Johnson

scale- flattened 2 3(2). Frons golden brown pollinose, male with — Last segment of male foretarsus not like hairs of foretarsi black laterally and pale 2(1). Scutellum convex, mostly shining; face usu-

. hind tibia and femur black . . brown medially, ally strongly gibbous; antennae situated at mimiis Osten Sacken about three-fourths height of head; more — Frons white pollinose, male with scalelike hairs densely pilose and less poUinose species .... 3 of hind femur and of foretarsi all brown, apex — Scutellum flattened, largely or entirely polli- rufotibialis Back middle of hind tibia reddish nose; face not usually strongly gibbose; anten- pollinose, at one-half height of head; 4(2). Frons white or slightly yellowish nae situated about scalelike hairs above antennae on frons not lim- less pilose and more pollinose species 13 of eyes flavipes Coquillett ited to margins 3(2). Antennal segment 3 yellowish red 4 — Frons golden brown pollinose; scale hairs of — Antennal segment 3 black or dark brown .... 5

frons limited to margins of eyes . coquilletti Wilcox 4(3). Abdomen with yellowish orange hairs, denser in male; pollinose fasciae of abdomen yellow-

mystax mostly golden; length 9-12 mm . Key to the Species of Utah CaUinicus Loew ish; aiiratus Cole

(Modified from Wilcox 1936a) — Abdomen with hairs light yellowish silver; pollinose fasciae of abdomen silver; mystax 1. Femora black, central stripe of mesonotum whitish yellow with much black; length 10- 15 presutural black spots ex- reaching scutellum, mm pulcher Back tending to the lateral margins .... pollenius Cole 5(3). Abdomen with dense orange-yellow hairs on — Femora yellow, central stripe of mesonotum dorsum 6 reaching scutellum, presutural black spots small — with dense hairs (may be somewhat and isolated from lateral margin by golden pol- Abdomen lateral mar- pictitarsis Bigot yellowish or black) limited to the len "^ gins

Orange hairs of abdomen limited to segments Curran 6(5). Key to the Species of Utah Comantella 1-4 curtistyhis Curran segments (Modified from James 1937b) — Orange hairs of abdomen on at least 1-5 dasyUoides Williston as antennal segment 3; 1. Style one-third as long at least 7(5). Male with disc of broad black hairs on venter black-haired, at least on anterior seg- apical two segments of midtarsus; foretarsal ments 2 segments 2-5 of male with a differentiated, — Style one-half as long as antennal segment 3; dense, narrow fringe of silvery hairs 8 with all hairs white pacifica Curran venter — Male without disc of black hairs on apical two vitta; male 2(1). Thoracic mane set on a well-defined black segments of midtarsus; foretarsus of hairs of mystax black on entire length without a differentiated fringe of silvery hairs rotgeri James

vitta; en- — Thoracic mane set on poorly defined black 8(7). Long hairs of lateral margins of abdomen hairs of mystax with tips white faUei (Back) tirely yellowish white ... p/awsor Osten Sacken Long hairs of lateral margins of abdomen in part black willistoni Curran Key to the Species of Utah apex black, apex of anal cell Cophiira Osten Sacken 9(7). Wing of male with also black; wing offemale darkened somewhat (Modified from Pritchard 1943) in same areas; pleura largely shining bimacula Walker

1. Scutellum entirely pollinose, base of tibia with Wing without black spots; pleura extensively dark yellow band poUinosa Curran pollinose ^^ of tibia without — Scutellum shining black, base at least 10(9). Tibiae and tarsi more or less reddish, dark yellow band 2 below, rarely male with only underside of ^1 2(1). Wings hyaline albosetosa Hine tarsi reddish

— Wings brownish 3 Tibiae and tarsi either both black or with tib- yellow with white bris- iae all black and foretarsi 3(2). Scutellum clothed with short, stout, dark hairs ^^ tles brevicornis (Williston) 46 Great Basin Naturalist Vol. 47, No. 1

Figs, 18-29. Utah Asilidae, terminalia; 18, male genitalia of P/i//o/hc»,s flmonen.sJs (Williston) dorsal aspect E = epandnum, P - proctiger; 19, female genitalia of P/H7o;HC«,sflmonen,s/.s(Williston), lateral aspect; 20, male genitalia of Polacanthacomposita (Hine), dorsal aspect, E = epandrium. P = proctiger; 21, female genitalia of Polacantha composita (Hme), lateral aspect; 22, male genitalia oi Proctacanthella cacopiloga (Hine), lateral aspect E = epan- drmm; 23, female genitalia of Proctacanthella cacopiloga (Hine), lateral aspect; 24, male genitalia oiMachimus griseus Hme, lateralaspect, with left epandrium removed, E = epandrium unnotched, P = proctiger without ventral process 25, male genitalia oiMachimus adustus Martin, lateral aspect, with left epandrium removed E = epandrium P = proctiger with ventral process; 26, male genitalia of Procfflca»if/j».s nearno Martin, dorsal aspect, E = epandrium' P = proctiger; 27, female terminalia of Proctacanthus nearno Martin, lateral aspect; 28, male genitalia of Proctacanthus micans Schiner, dorsal aspect, = E epandrium, P = proctiger; 29, female terminalia of Proctacanthus micans Schiner lateral aspect. January 1987 Nelson: Utah Robber Flies 47

indistinctus (Bromley) showing hairs Figs. 30-33. Utah Asilidae, lateral aspect of head and thoim; 30, Sclcropo^on (h\'popleuron); Machimiis on katatergite (hvpopleuron); 31, Stenopofion inquinatus Loew showing bare katatergite 32, Williston showing hairs occidentalis (Hine) showing hairs on both katatergite and anatergite; 33, Promachus cdbifacics on katatergite and bare anatergite.

Scutellar hairs all black, pollinose fasciae of 11(10). Mystax largely black; foretarsus of male yellowish — jemezi Wilcox & Martin abdominal segments not extending forward . banksi Wilcox & Martin — Mystax largely white, particularly above (some females with a few black hairs above); Key to the Species of Utah Dioctria Meigen foretarsus of male black montantis Loew (Modified from Adisoemarto and Wood 1975) 12(10). Foretarsus of male yellowish with white hairs; of male white above ... rufotarsus Back mystax 1. Pleuron with golden pollinose band extending — Foretarsus of male black with black hairs; uninterrupted from between base of wing and Martin humeral callus to coxa vera Back mystax all black . . . stenofrons Wilcox & discontinuous pollinose band, 13(2). Pollinose fasciae of abdominal segment 1 en- — Pleuron with anterior margin of meso- tire or nearly so l^i broadly interrupted on pleuron; interruption as wide as high 2 — PoUinosity of abdominal segment 1 limited to of mesopleuron restricted to and patches near lateral margins, not extending 2(1). PoUinosity covered by haired area near prothoracic medially across segment 16 mostly spiracle ptisio Osten Sacken 14(13). Tibiae and femora reddish below; mystax — PoUinosity of mesopleuron extending from black except for a few central, white hairs . . spiracle to base of wing profusus Osten Sacken prothoracic henshawi Johnson — Tibiae and femora black, midtibia sometimes reddish in male; mystax white above, black Key to the Species of Utah below 15 Eucyrtopogon Curran 15(14). PoUinosity of abdominal segments narrowly interrupted along dorsal midline The genus Eucyrtopogon Curran occurs in Cole thompsoni Utah with possibly as many as four or five — PoUinosity of abdominal segments entire but species represented. The Curran (1923a) key spot on several segments lacking with central easily distinguishes one species, E. comantis pollen ablautoides Melander Curran, by the presence of a medial crest of large part white; pollinose 16(13). Scutellar hairs in white hairs on the mesonotum of the males. fasciae of abdominal segments extending The other representatives of this genus in the anteriorly along lateral margins are only eight idahoensis Wilcox 6f Martin state do not key out well. There 48 Great Basin Naturalist Vol. 47, No. 1

Figs. 34-46. Utah Asilidae, right wing: 34, Efferia aestuans (Linnaeus); 35, Efferia albibarhis (Macquart)- 36 Ejjeria apache Wilcox; 37, Efferia benedicti (Bromley); 38, Efferia costalis (Williston); 39, Efferia pernicus Coquillett- 40, Cyrtopogon willistoni Curran; 41, Efferia tucsoni Wilcox; 42, Promachus albifacies Williston; 43 Machimus occidentahs (Hine); 44, Efferia freivingi Wilcox; 45, Laphijstia tolandi Wilcox; 46, Leptogaster sp. from Clear Creek Raft River Mountains, Box Elder Co. , Utah. January 1987 Nelson: Utah Robber Flies 49

lacking pollen on posterior portion of specimens before me from the collections at 2(1). Scutellum disc near margin wilcoxi Martin BYU, UU, SUSC, and CEU. These speci- — Scutellar disc pollinose (narrow margin may ap- mens seem to represent three distinct spe- pear to be lacking pollen) 3 cies. The specimens from USU have been 3(2). Scutellar disc lacking hair currani Martin sent to R. Lavigne at the University of Wyo- — Scutellar disc with a few to many hairs 4 ming, as he is at present studying the genus. Anterior mesonotal hairs sparse, erect, mostly Seven of the USU specimens are labeled E. 4(3). brown mingusae Martin nebulo as determined by Bromley; other — Anterior mesonotal hairs dense, somewhat re- are simply labeled Eucyrtopogon specimens cumbent, white alhipihsus Curran sp. Any determination of the species in Utah (except E. comantis) would be very tenuous Key to the Species of Utah Laphystia Loew on my part, due to the small number of speci- mens which I have seen; thus, determinations (Modified from Wilcox 1960) of Eucyrtopogon from Utah must await the 1. Scutellum with marginal bristles nearly as work which Lavigne is preparing. strong as lateral bristles of mesonotum; femora mostly dark brown, apices yellowish tolandi Wilcox Key to the Species of Utah Heteropogon Loew — Scutellum with marginal hairs weak and pro- from Wilcox 1965) (Modified cumbent or absent; femora usually concolorous . 2 procumbent; L Scutellum without marginal bristles; wing with 2(1). Marginal scutellar hairs weak and dark spots on crossveins and furcations abdomen not reddish 3 maculinervis James — Marginal scutellar hairs absent; abdomen red- Hull — Scutellum with marginal bristles; wings hyaline dish brown rubra 2 3-5 3(2). Abdomen yellowish, fasciae on segments of length of segments; femora of both 2(1). Scutellum polished on hind margin, disc one-third abdominal fas- scutellum pollinose; terminal abdominal seg- sexes wholly reddish; pollen of Wilcox ments red stonei Wilcox ciae white utahensis pollen of — Scutellum entirely pollinose, sometimes thinly — Abdomen brown, femora yellowish, yellow annulata Hull so, subshining black; terminal abdominal seg- abdominal fasciae ments black 3 extending greater than 3(2). Black bristles of mystax Key to the Species of Utah Lasiopogon Loew one-third distance from oral margin to antennae 4 (Modified from Cole and Wilcox 1938)

— Black bristles of mystax confined to oral margin extending less than one-third distance from oral 1. Marginal scutellar bristles white alhidus Cole & Wilcox margin to antennae ^ of scutellar bristles black 2 4(3). Male with brush of black hairs in basal third — Marginal hairs of mystax both black and white; one-third midtibia; 2(1). Surstyli of male genitalia one and senilis Bigot scutellar bristles black times as long as high in lateral view (Fig. 12); abdominal — Males without brush of black hairs on niidtibiae; male genitalia wider than adjacent (Fig. 13); sternite 9 of hairs of mystax all white; scutellar bristles black segment in dorsal view species 1 female genitalia amber colored aldrichii Melander 5(3). Bristles of thorax and scutellum white, rarely or more times as long one or two presutural black brisdes; bristles of Surstyli of male about two genitalia . lateral view (Fig. 14), male mystax black and white, white bristles above . as high in martini Wilcox narrower than abdomen in dorsal view; sternite 9 of female genitalia dark brown or black 3 — Bristle of thorax both black and white; bristles of with ash-colored pol- . covered . of male scutellum black; brisdes of mystax all black . 3(2). Surstyli arizonensis Wilcox len; surstyli without tooth on ventral margin before base (Fig. 16) (Note: male genitalia may be rotated 180 degrees) cinereus Cole Key to the Species of Utah Holopogon Loew Surstyli of male shining black or brown, surstyli on ventral margin near (Modified from Martin 1959) with prominent tooth base (Fig. 15) monticola Melander laterally 1. Mesonotum without presutural bristles caesariatus Martin Key to the Species of Utah Nicocles Jaennicke — Mesonotum with two or more presutural bris- (Modified from Wilcox 1946) tles laterally 2 50 Great Basin Naturalist Vol. 47, No. 1

Figs 47-56. Utah Asilidae, lateral aspect of male terminalia and legs: 47, male terminalia of Efferia utahensis (Bromley); male terminalia 48, oi Efferia frewingi Wilcox; 49, male terminalia otEfferia mortensoni Wilcox; 50 male termmalia of Efferia tucsoni Wilcox; 51, male terminalia of Efferia bicolor (Bellardi); 52, male terminalia of Efferia MibarbisiMacquRrt)- male 53, terminalia of Efferia zonata (Hine); 54, foretihia and basitarsus of Diogmites grossus Bromley B - basitarsus, T = tibia; foretibia 55, and basitarsus of Cophura scitiila Williston, B = basitarsus T = tibia 56, middle leg of Callinicus pollenius (Cole), T = tibia. January 1987 Nelson: Utah Robber Flies 51

L Abdominal segments 3-7 of male and 4-6 of fe- 1. Gibbosity of face extending about one-half to distance oral margin to an- male red, silver fasciae covering entire dorsnm oi three-fourths from male segments 5-6 ahdominalis Williston tennae (Fig. 3); fore coxae with numerous bris- tles 2 — All abdominal segments of male and female black; extending atjout five-sixths to seven- silver fasciae of male covering entire dorsnm of — Gibbosity oral margin to antennae segment 6 and basal portion of segment 5 eighths distance from utcihcnsis Banks (Fig. 2); fore coxae with numerous long hairs ... 3

2(1). Gibbosity weak, face not strongly inflated; shiny black triangle between mystax and antennae ab- Key to the Species of Utah Ospriocems Loew sent; small, length 9-10 mm . . utahensis Bromley

(Modified from Martin 1968) — Gibbosity strong, face strongly inflated; shiny black triangle between mystax present; large, 1. Abdominal segments entirely black length 20-37 mm inquinatus Loew minos Osten Sacken 3(1). Abdomen densely pollinose; long hairs on inner

or brown . reddish, . — Abdominal segments in part arms of hypandrium continuing across base . pollinose ^ martini Bromley

in 2(1). Abdominal segments brown pollinose, tan — Abdomen subshining black; long hairs on inner ground color lon^tihts (Loew) arms of hypandrium interrupted medially 4

pollinose . — Abdominal segments in part red, not 3 4(3). Hind tibiae with indistinct reddish area at least in- near apex; mystax yellowish 3(2). Tip of flagellum with notch containing small engelhardti Bromley conspicuous spine (Fig. 8) ahdominalis (Say) reddish or yellowish, mystax red- — Tip of flagellum produced as apical segment — Hind tibiae all dish rufibarhis Bromley (Fig. 9) vallensis Martin

Key to the Utah Stichopogon Loew Key to the Species of Utah Scleropogon Loew (Modified from Wilcox 1936b) (Modified from Wilcox 1971) 1. Scutellum with marginal hairs or bristles; pollen of abdomen golden and brown; length 3-5 mm 1. Cells rj and nij with long petioles subequal to fragilis Back length of rm crossvein picticornis Loew — Scutellum lacking marginal hairs or bristles; pol- — Cells r, and m, either open or with one or other len of abdomen silver; length greater than 6 mm having a short petiole 2 2 2(1). Antennal segment 3 one and three-fourth times 2(1). Tergite 4 mostly silvery pollinose sometimes as long as segments 1-2; abdomen slender, gray with medial portion of apex lacking pollen .... pollinose; wings tinged with brown trifasciattis Say duncani (Bromley)

— Tergite 4 with black band covering basal third . — Antennal segment 3 at most one and one-half salinus (Melander) times as long as segments 1-2; abdomen stock-

ier; wings hyaline 3 Key to the Genera of Utah Laphriinae 3(2). Hypopleuron with bristles, cell nij closed and a sclerotized arch usually short petiolate ^ 1. Postmetacoxal area forming when viewed from below; flagellum bearing ei- — Hypopleuron with hairs, cell nij broadly open . . 5 ther a short dorsal bristle or terminal style 2 Abdomen stout, apex of wings reaching to ter- 4(3). — Postmetacoxal area entirely membranous; fla- gite 6; ground color of most of abdomen tan; gellum without dorsal bristle or terminal style, length 15-19 mm coyote (Bromley) apical pit may contain a short bristle 3 — Abdomen elongate, apex of wings not reaching 2(1). Coloration and pile resembling a ; to tergite 6 but covering base of tergite 5; ground anatergite bare; length over 25 mm abdomen dark brown with bases and color of Dasylechia atrox (WiUiston) apices of segments tan; length 19-26 mm indistinctus (Bromley) Body quite bare, not resembling a bumblebee; anatergite haired or with bristles; length less 5(3). Abdomen black, narrow posterior and lateral than 20 mm mucida Osten Sacken margins of tergites tan; densely pollinose species neglectus (Bromley) 3(1). Proboscis compressed laterally; some in pile and coloration on resembling — Abdomen reddish brown, black laterally Laphria Meigen basal segments; thinly pollinose bradleyi (Bromley) or S. ^Wilcox has determined specimens as being either S. eng.elhardti with the final couplet rufibarhis from the same locahties which do not agree used in this key (adapted from Key to the Species of Utah Stenopogon Loew of this key. It may be that the characters of these two Wilcox 1971) are not dependable in separating Utah specimens (Modified from Wilcox 1971) species. 52 Great Basin Naturalist Vol. 47, No. 1

;i-:^ .

January 1987 Nelson: Utah Robber Flies 53

enclose an — Proboscis flattened dorsoventrally; abdomen — Epandria of male not arching to but mostly bare and reddish orange open space when viewed from above, Cerotainiops ahdominalis (Brown) touching along most of dorsomedial margin (Fig. 20); terminalia of female lacking stout bristles (Fig. 21) H Key to the Genera and Species of 1 Length of antennal style at least one and one- Utah AsiHnae 1(10). third length of antennal segment 3; inner sur- male with patch of erect (Modified from Wood 1981) face of epandrium of spines Polacantha Martin Anatergite (metanotal callus) bare (Fig. 33) .. 2 L — Length of antennal style of antenna shorter — Anatergite (metanotal callus) with bristles or than or subequal to length of antennal seg- hairs (Fig. 32) 9 ment 3; inner surface of epandrium lacking erect spines 12 2(1). R4 with recurrent vein; recurrent branch ofmale notched near apex (Fig. 11) (stump vein) sometimes short (Fig. 35), end- 12(11). Epandria Neomochtheriis Osten Sacken ing in cell r,+3 or extending basally and joining R,,3(Fig.42) 3 — Epandria ofmale not notched near apex (Fig. 24) 13 — R4 without recurrent vein (Fig. 43) 7 13(12). Style about one-fourth length of antennal seg- 3(2). Cell rj narrow, apical half of R5 subparallel ment 3; bristles of tergites 2 and 3 short and 35-39) Effeha Coquillett with R4 (Figs. recumbent Negasilus Curran half of R5 diverging — Cell r4 broad, apical — Style longer than one-fourth length of anten- strongly from R4 (Fig. 42) 4 nal segment 3; bristles of tergites 2 and 3 longer and erect 14 4(3). Claws acute; abdomen elongate 6 Epandria of males pointed and directed up- — Claws blunt; abdomen stout 5 14(13). ward at apex (Fig. 17), apices diverging .... 5(4). Lower face inflated; hind femur club-shaped, Asilus aurianntdattis Hine with row of bristles below near apex; Utah — Epandria of males not pointed and not di- specimens have black pile on thorax . rected upward at apex, apices converging . Mallophora fautrix Osten Sacken Asilus vescus Hine, and A . . . Machimiis Loew, — Lower face not greatly inflated; hind femur formosiis Hine spindle-shaped and without bristles below near apex; pile of thorax in Utah specimens Key to the Species of Utah Efferia Coquillett Bigot mostly gray and/or yellow . . . Megaphonis (Modified from Wilcox 1966a) 6(4). Recurrent branch (stump vein) short (similar vein 1. Three submarginal cells in wings (no to Efferia), not reaching R2+:3 submarginal cell, but continuing Triorla interrupta Macquart ending in a to meet Ro), (Fig. 39)(Anomala group) 2 — Recurrent vein extending to and connecting vein with R2+3 Prornachus Loew — Two submarginal cells in wings, 'stump'

present and ending in first submarginal cell . . 3 7(2). R5 reaching wing margin anterior to apex of much longer wing Proctacanthus Macquart 2(1). Hairs on abdominal sternites than antennal segments 1-2 combined — R5 reaching wing margin posterior to apex of pernicus Coquillett wing 8 — Hairs on abdominal sternites subequal to an- elongate, longer 8(7). Epandria of male black and tennal segment 1 in length davlsi Wilcox terminalia of females lacking than tergite 6; cell (Figs 37, 3(1). R4+5 forking before apex of discal strong bristles 44) 4 Regasilus hlantoni Curran B^^5 forking at or beyond apex of discal cell red and short, subequal to — Epandria of male (Figs. 34-36, 38, 41) H length of tergite 6 (Fig. 22); terminalia of fe- bristles or hairs at 4(3). Mesonotum anteriorly with male bearing strong bristles (Fig. 23) least as long as antennal segments 1-2 com- Proctacanthella cacopiloga Bromley bined; color ofmale abdominal segments 2-5 9(1). Occipital bristles long and fine, distal third alternating black and white; lacking long strongly bent anteriad at nearly a 90 degree parted hairs; segment 6-7 white pollinose; (Hine) and split at tip when . hrevicomus angle (Fig. 5) . . Neoitamiis pointed viewed from above (Pogonias group) 5 — Occipital bristles stout, not bent forward at a sharp angle, but may bend forward gradually Mesonotum anteriorly with bristles and hairs 10 throughout length shorter than antennal segments 1-2 com- bined; color ofmale abdominal segments gray 10(9). Epandria of male arching to enclose an open often with white parted space when viewed from above (Fig. 18); ter- or white pollinose, ovipositor rounded at tip, not spht minalia offemale bearing several pairs of stout hairs; when viewed from above (Staminea group) . . 7 bristles (Fig. 19) Loew 54 Great Basin Naturalist Vol. 47, No. 1

'il:{y-p' A Atomosia mucida # Bleehare£iurii sonorenslj ' 'a Cerotainiops , j f^- # abdominal is A Ca ninicus pictitarsis

Callinicus pollenius

-'x?'^^-S 61 ym&

--trr-'*' -.—^-,-^.--r^ January 1987 Nelson: Utah Robber Flies 55

Anterior part of mesonotum compressed lat- 5(4). Lower forceps of male with apical lobes form- 12(11). ing an acute angle when viewed laterally (Fig. erally; acrostical crest of long hairs or bristles group) 13 47); scutellar bristles and hairs white (some- present (Carinata black); mysta.\ with at most four times one — Anterior part of mesonotum not compressed black bristles; females with fore and middle laterally; crest, if present, extends onto dor- black, sometimes brownish dorsally tibiae socentral row as well as acrostical row 15 near bases titahensis (Bromley) 13(12). Abdomen of male lacking long, white parted — Lower forceps of male with apical lobes meet- hairs; black of segments 2-4 extensive; ing basally as a broad U or with a flattened mesonotal crest short, subec}ual to antennal area between them at base (Figs. 48-49); segment 1; ovipositor greater than 4 mm in scutellum with black bristles and white hairs length wiUistoni (Hine) or white bristles and black hairs, bristles and hairs not all white; mystax with at least eight — Abdomen of male with long, white parted black hairs or bristles along oral margin; fe- hairs; black of segment 2-4 limited (at most) males usually with fore and middle tibiae to hind margins; mesonotal crest subequal to 1-3; ovipositor brown, if black then scutellar bristles black . . 6 length of antennal segments usually less than 4 mm in length (if longer, 6(5). Lower forceps of male with apical lobes join- then mesonotal crest long) 14 ing in a broad, evenly curved U when viewed laterally (Fig. 48); scutellar bristles usually 14(13). Abdominal segment 7 of male with mostly black (at least in part), hairs of scutellum usu- black hairs; ovipositor about 4 mm long .... ally white; common frewingi Wilcox subcuprea (Schaeffer)

— Lower forceps of male with a broad relatively — Abdominal segment 7 of male with mostly flat area between the apical lobes (Fig. 49); white hairs; ovipositor 3.5 mm or less in scutellar bristles white (at most one black), length (wing. Fig. 38) costalis (Williston) hairs of scutellum all black; rare, single speci- 15(12). Mesonotum anteriorly with numerous erect men from St. George mortensoni Wilcox hairs or bristles as long as antennal segments yellow (slightly 7(4). Mystax yellow, pollen of foce 1-3; marginal bristles of scutellum numerous; contrasting with white yellow in female but bristles of tarsi mostly white (Arida group) .. 16 hairs of postgena) staminea (Williston) — Mesonotum anteriorly with bristles and hairs and pollen efface silvery white, hairs — Mystax shorter than antennal segments 1-2; fewer mystax and postgena both white 8 of than ten marginal scutellar bristles 18

8(7). Sternite 8 of male produced medially at apex; 16(15). Mystax mostly white, 1-8 black hairs below female with R4^5 forking at or before middle of and laterally; discal hairs of scutellum mostly the distance between rm and apex of discal white (especially in female); bristles and hairs cell 9 of postalar callus mostly white (or a few of medially, — Sternite 8 of male not produced weaker bristles and hairs black) . apache Wilcox straight when viewed from below; female — Mystax with dense black hairs below and lat- with R4+5 forking beyond middle of distance erally in male, lateral black hairs sparser in between rm and apex of discal cell 10 female but more than eight present; white 9(8). Anterior mesonotal crest shorter than anten- hairs of scutellum limited to base and sides (or nal segment 1; sparse, white parted hairs on absent); bristles and hairs of postalar callus abdominal segments 2-4; abdomen grayish mostly black (with at least one strong white ^'^ pollinose henedicti (Bromley) bristle) Anterior mesonotal crest subequal to length — 17(16). Foretibia mostly black, somewhat lighter 1 and 2; long, white of antennal segments basally; scutellar bristles black in male, white hairs on abdominal segments 2-5 and parted in female arida (Williston) short parted on 6-7; abdomen silvery-white basally and black apically; pollinose basini Wilcox Foretibia reddish scutellar bristles white (only male seen; may 10(8). Fringe of hairs on surstyli of male white; . (Bromley) simply be variant oi arida ) subarida white parted abdominal hairs long and dense tubercles on on segments 1-7 of male; female indistin- 18(15). Males with prominent, ventral slender guishable from that oicana .... deserti Wilcox abdominal segments 4-7 (Fig. 50), species (Tuberculata group) 19 — Fringe of hairs on lower forceps of male black in part; white parted abdominal hairs long on Males without prominent tubercles on ab- segments 1-4, short on segments 5-7, sparse dominal segments (Fig. 52); usually larger, 20 on all; female indistinguishable from that of stouter species (Albibarbis group) deserti cana (Hine) 19(18). Male terminalia and tubercles reddish; fe- 11(3). R5 reaching costa before apex of wing (Figs. male abdominal segment 7 and base ofovipos- 35-38, 41) 12 itor reddish; abdominal tergites largely white hairs of tergite 7 black — R5 reaching costa after apex of wing (Fig. 34) haired, some Wilcox aestuans (Linnaeus) tucsoni .

56 Great Basin Naturalist Vol. 47, No. 1

# Cyrtopogon bimacula

A Cyrtopogon curti stylus

4.J f-:^. Cyrtopogon dasyHoides

66

9 Cy rtopo gon ida hoen s

A Cyrtopogon Jemezi

Figs. 65-68. Utah Asilidae, distribution: 65, Cyrtopogon ablaiifoidcs , Cyrtopogon auratus , and Cyrtopogon banksi; 66, Cyrtopogon himacula, Cyrtopogon curtistylns , and Cyrtopogon dasylloides ; 67 , Cyrtopogon idahoensis, Cyrtopogon jemezi , and Cyrtopogon albifacies ; 68, Cyrtopogon montanus .

January 1987 Nelson: Utah Robber Flies 57

— Male terminalia and tubercles black; female — Larger, greater than 12 mm in total length; male abdominal segment 7 and base of ovipositor terminalia black or dark brown 7

black; abdominal tergites broadly black 7(6). Pollen of abdomen golden; color of incisures not haired producta (Hinc) contrasting greatly with color of remainder of each segment paropus (Walker) 20(18). Male terminalia black; surstyli with ventral tooth near apex (Fig. 51); marginal scutellar — Pollen of abdomen silvery; incisures yellowish, bristles black with discal hairs long and white contrasting with the black remainder of each bicolor Bellardi segment callidus (Williston)

— Male terminalia reddish; surstyli without ventral tooth near apex (Figs. 52, 53); scutel- Key to the Species of Utah Megaphorus Bigot lum with either black bristles and some short (Modified from Cole and Pritchard 1964) black hairs or with bristles and hairs both 21 white 1. Costa ending near wing tip 2

21(20). Mystax all white; transverse black (or brown- — Costa continuing around wing tip to hind margin 4 ish) bands of abdomen interrupted by longitu- 2(1). Cell rj open pulcher (Pritchard) dinal lighter stripe; length of ovipositor sub- — Cell r, closed, usually petiolate 3 equal to length of abdominal segments 6-7; male genitalia as in Fig. 52 3(2). Wing veins brown tvillistoni (Cole)'* alhibarhis (Macquart) — Wing veins yellow guildiana (Williston)'' black and white; transverse black — Mystax 4(1). Pile of thorax and abdomen white, wings yellow- uninterrupted; length of bands of abdomen ish; length 10 mm pallidus (Johnson) ovipositor greater than length of abdominal — Pile of thorax and abdomen bright yellow, wings segments 6-7; male genitalia as in Fig. 53 . . brownish; length 12-14 mm . . fnistra (Pritchard) zonata (Hine)

Key to the Species of Utah Negasihis Curran Key to the Species of Utah Machimus Loew 1. Margin of scutellum with bristles . . . mesae (Tucker) Including Asilus formosiis Hine and — Margin of scutellum lacking bristles ... belliCunan Asilus vescus Hine

(Modified from Hine 1909 and Martin 1975) Key to the Species of Utah NeomochtJierus Osten Sacken 1. Hind femur longitudinally yellowish brown on ventral surface and black on dorsal surface 2 1. Face white, disc of scutellum lacking long, white — Hind femur with yellowish brown confined to hairs hypopygialis (Shaeffer) apex, or hind femora entirely black 4 — Face brown; disc of scutellum with long, white hairs 2 2(1). Wing with some dark coloration; especially near in apex 3 2(1). Wings of male with brown color concentrated area near middle of anterior half of wing; anal — Wing entirely hyaline Asilus fonnosits Hine lobe of male hyaline lepidus (Hine) 3(2). Wing with dark spots in medial cells; distal — Wings of male with brown color evenly dis- fourth of wing with dark coloration isolated from tributed throughout; anal lobe of male tinged veins except at extreme tips; proctiger without with brown alhicornis (Hine) ventral process near apex griseus (Hine)

— Wing without dark spots in medial cells, distal to the Species of Utah Philonicus Loew fourth of wing with faint dark wash reaching Key apex; proctiger with ventral process veins before (Williston)' 1. Wing smoky; legs dark red . . arizonensis near apex (Fig. 25) adustns Martin (Hine) — Wing clear; legs pale reddish . limpidipennis 4(1). Sternite 8 of male produced apically and bearing tuft of longer hairs on apex of produced area (Fig. 6) 5 Key to the Species of Utah Proctacanthus Macquart — Sternite 8 of male not produced apically, hairs of

posterior margin (Fig. 7) . 6 uniform length across (Modified from Hine 1911)

5(4). Proctiger with two processes near its attachment to abdomen; medial areas of tergites covered state. to M. willistnni and M. nuildiana. Cole and Pritchard pollen sestertius Martin ^With regard with brown blend with 'Some populations [oi willl'iUmi]. particularly in Utah, tend to found in the — Proctiger lacking processes; medial areas of ter- Ruildiam." After examining the specimens of these species the key lead most comfortably to tvillistoni. gites covered with dense, silvery pollen state, 1 found have been identified by Bromley and Wilcox as occidcntalis (Hine) 'Specimens of both species derived from occurring scattered throughout the state. Using the above key to I found that specimens from Utah seem 6(4). Small, less than 11 mm in total length; male Hine (19()9) and Bromley (1934), terminalia reddish brown Asilus vescus Hine run best to arizonensis .

58 Great Basin Naturalist Vol. 47, No. 1

W^ • Cyr topogon plausor # Cyrtopogon pulcher A Cyrtopogon profusus A Cyrtopogon rufotarsus

H Cyrtopogon stenofrons 69

'•y%

• Cyrtopogon wilHstoni

A Cyrtopogon thompsoni

:•#.

•^•.

• •;

Figs. 69-72. Utah Asilidae, distribution: 69, Cyrtopogon plausor and Ci/rtopogon profusus- 70, Cyrtopogon pulcher, Cyrtopogon rufotarsus, and Cyrtopogon stenofrons; 71, Cyrtopogon ti)i//i.stont and Cyrtopogon thompsoni;

72, Uioctria vera , Dioctria henshawi , and Dioctria pusio January 1987 Nelson: Utah Robber Flies 59

1. Body with ground color reddish orange Ablautus flavipes Coquillett 1904: 178 Iiinci Bromley [Types: Three males and two females, Los — Body with ground color black and orange 2 Angeles and San Diego counties, California in

2. Wings with uniform brownish tinge USNM]. Utah distribution: Juab Co.: Topaz milberti Macquart Mtn.; Tooele Co.: Dugway Proving Ground. — Wings clear or with brown concentrated near 26April-20June. veins 3 Ablautus mimus Osten Sacken 1877: 290 outward and 3(2). Epandria of male genitalia curving [Holotype: male, Crafton near San then back to enclose a space between tips and Bernardino, California, March in MCZ]. Utah proctiger when viewed from above (Fig. 26); Woodhill; terminal keellike sternite of female lacking distribution: Carbon Co.: Emery strong spines ventrally (Fig. 27); circlet of spines Co.: Elmo; Kane Co.: Kanab; Tooele Co.: present dorsally nearno Martin Cedar Mtns.; Utah Co.: Alpine, Provo, — Epandria not enclosing a space, but straight and Spanish Fork, Vineyard (now Geneva Steel), terminalia of compact against proctiger (Fig. 28); west side Utah Lake. 6 April-4 July. female with ventral keellike projection covered Ablautus rufotibialis Back 1909: 182 [Holo- with stout spines in addition to dorsal circlet of female, Ysleta, Texas, 2 April 1902 in spines (Fig. 29) micans Schiner type: AESP]. Utah distribution: Grand Co.: Moab; Juab Co.: Topaz Mtn.; San Juan Co.: Comb Key to the Species of Utah Promachiis Loew Wash 32 mi SW Blanding; Tooele Co. : Dug- (Modified from Hine 1911) way Proving Ground. 17-22 April. Blepharepium sonorensis Papavero and 1. First submarginal cell with dark, distinct gray cloud 2 Bernardi 1973: 175 [Holotype: , Palos Uni- — First submarginal cell clear or with very faint, Verdes, viii. 1949, in Museu Zoologia da narrow gray line; black areas of abdomen with versidade de Sao Paulo]. Utah distribution: lateral and poste- few to many white hairs near Washington Co.: St. George, Coalpits Wash, margins of segments aldrichii Hine rior Zion National Park. 5-18 August. Male genitalia without silvery hairs above; tibiae 2(1). Bohartia sp. Utah distribution: Rich Co.: yellow in contrast to black femora . . sackeni Hine Dry Basin. 4 August. — Male genitalia with silvery hairs above; tibiae Callinicus pictitarsis (Bigot) 1878: 411 and femora more or less concolorous, not con- California; in Bigot Collection]. trasting greatly 3 [Holotype: Utah distribution: Washington Co.: Zion Na- 3(2). Mystax white, sometimes slightly yellow; thorax gray pollinose; male abdominal tergite 3 without tional Park. No dates on labels. lateral patch of dense black hairs; light hairs of Callinicus pollenius (Cole) in Cole and abdomen whitish gray albifacies Williston Lovett, 1919: 237 [Holotype: male. Hood — Mystax yellowish; thorax brownish yellow polli- River, Oregon, 24 September 1917; #491 in with lateral patch nose; male abdominal tergite 3 CAS]. Utah distribution: Cache Co.: Black- of dense black hairs; light hairs of abdomen yel- smith Fork Canyon; Rich Co.: Logan Canyon lowish dimidiatus Hine summit. 30 July-1 September. Coleomyia alticola James 1941: 37 [Holo- Synopsis OF Species type: male. Science Lodge near Ward, Colo- Subfamily Dasypogoninae rado, 24 July 1939, repository not listed], Utah distribution: Cache Co.: Beaver Moun- Ablautus coquilletti Wilcox 1935: 226 tain, Blacksmith Fork Canyon, Tony Grove; Angeles Co., California [Holotype: male, Los Rich Co.: Logan Canyon summit, Monte Utah distribution: Washington in USNM]. Cristo; Uintah Co.: Whiterocks. 6 June-14

: , 4 June. Co. Leeds Cyn. Santa Clara. May-2 August. Comantella fallei (Back) 1909: 278 [Lecto- type: male, Fort Collins, , 30 March including M. T. James, Wilcox, and S. W. Bromley have '^Authors J. Carbon identified these three species as occurring in Utah. I have difficulty, how- 1900; in CSU]. Utah distribution: making ever, in seeing a distinct brownish tinge of the wings of P. milberti, Co.:Sunnyside. 11-10-1936. the male specimens determined by these people run to P. micans in Hine s key (1911). Females labelled P milberti lack the spines on the terminal Comantella pacifica Curran 1926: 311 P. milberti is composed of keellike stemites. The dorsal circlet of spines of [Holotype: Penticton, B.C., 4 April 1919, several rows (2-4) of spines, while those of P. micans and P. nearno are made reduced to a single row of stouter spines. These observations were #2320 in CNC]. Utah distribution: Box Elder without examining types but by comparing specimens in the USU collection Co. : Price Airport; : Hills; Carbon which had previously been identified by the above authors. Co. Wildcat 60 Great Basin Naturalist Vol. 47, No. 1

Eucyrtopogon comanti's

Asilidae, distribution: 73, Diofimites oj-osstts and Dicolonus sparsipilosum, 74, Eucyrtopogon mantisSf nVvPr^^ir-md ^^'i h s Eucyrtopogon spprJS, Effcna apache And Effehadavisi- 76, Efferiaalbibarbis. ^ January 1987 Nelson: Utah Robber Flies 61

Daggett Co.: Sheep Creek; Millard Co.: tion: Cache Co.: Avon, Blacksmith Fork Kanosh; Tooele Co.: Cedar Mountains. 26 Canyon, Cold Spring-Mendon, Providence, March-4 May. Smithfield, Tony Grove, Twin Creek, Comantella rofgeri James 1937b: 61 [Holo- Wellsville; Daggett Co.: Ashley National type: male, Stollsteimer, Colorado, 6,500 ft, Forest, Elk Park; Grand Co.: La Sal Moun- 29 October 1935, no repository listed]. Utah tains; Juab Co.: Mount Nebo; Rich Co.: Gar- distribution: Iron Co.: Cedar City. 26 Octo- den City, Logan Canyon summit, Monte

ber. Cristo; Utah Co. : American Fork Canyon, As- Cophura albosetosa Hine 1908: 202 [Syn- pen Grove, Mt. Timpanogos, North Fork types: two males and one female, Hope Provo Canyon; Wasatch Co.: Wolf Creek Mountains, B.C.; probably in OSU]. Utah Pass. 12 June-14 August. distribution: Cache Co.: Green Canyon; We- Cyrtopogon bimacuhi (Walker) 1851: 102 August. [Syntypes: Four males and four females; ber Co. : Willard Peak. 21 July-13 Cophura brevicornis Williston 1883: 22 'North America,' in British Museum]. Utah [Syntypes: Two specimens, sex not known, distribution: Beaver Co.: Beaver; Duchesne

: Boulder from Washington, in KU]. Utah distribution: Co. : Wolf Creek Pass; Garfield Co. Washington Mountain, Aquarius Plateau; Iron Co.: Cedar Uintah Co. : Whiterocks Canyon; Co.: Zion National Park; Wayne Co.: near Breaks; San Juan Co.: La Sal; Sanpete Co.: Grover. 12 July-7 August. Ephraim, Gunnison. 10 July-24 July. Cophura pollinosa Curran 1930: 10 [Holo- Cyrtopogon curtistylus Curran 1923a: 133 type: male, Kitt Peak, Rincon, Baboquivari [Holotype: male. Cache Junction, Utah, 3 Mountains, Arizona, 1-4 August 1916, 4,050 June 1912, inC. W. Johnson collection]. Utah Elder Co.: Brigham City, ft, in AMNH]. Utah distribution: Washington distribution: Box

Co.: Santa Clara, Zion National Park. 30 Willard; Cache Co. : Cache Junction, Beaver Canyon, Sardine May-27 July. Creek, Logan, Logan Cophura scitula Williston 1883: 19 [Holo- Canyon; Davis Co.: Bountiful; Rich Co.: Lo-

: Park, type: sex not known, Washington Territory, gan Canyon summit; Utah Co. Glacier Springs, Pump- in KU]. Utah distribution: Cache Co.: Green Timpanogos, 2 mi W Cascade Canyon, Logan, Logan Canyon, Tony Grove, house Hill. 2 July-3 August. West Hodges Canyon; Grand Co.: Wilson Cyrtopogon dasylloides Williston 1883: 11 Mesa; Rich Co.: Logan Canyon summit; San- [Holotype: male, Washington Territory, in KU]. Utah distribution: Cache Co.: Black- pete Co. : Ephraim Canyon; Utah Co. : Ameri- can Fork Canyon, Aspen Grove, Emerald smith Fork Canyon, Logan. 16 May-12 June. Martin Lake. 10 July-23 September. Cyrtopogon idahoensis Wilcox and Cyrtopogon ablautoides Melander 1923a: 1936: 82 [Holotype: male, Parma, Idaho, 13 111 [Syntypes: four males and six females, May 1934, in CAS]. Utah distribution: Cache Mabton, Washington, 3 May 1911, in Me- Co.: Blacksmith Fork Canyon, Cold Sardine lander Collection]. Utah distribution: Millard Spring-Mendon, Logan, Mendon,

: Rich Co.: 15 mi N Delta. 31 May. Canyon; Garfield Co. Aquarius Plateau; Ears; Cyrtopogon albifacies Johnson 1942: 1 Co.: Monte Cristo; San Juan Co.: Bear Co: 13 mi S [Holotype: male. Glacier Lake, Mount Tim- Utah Co. : Payson Canyon; Weber panogos, Utah, 10,600 ft; August 1928, in Monte Cristo. 9 June-23 July. Martin BYU]. Utah distribution: Utah Co.: type lo- Cyrtopogon jemezi Wilcox and Valle Grande, cality. July-August. 1936: 47 [Holotype: male, Je- Cyrtopogon auratus Cole in Cole and mez Mountains, , 6 July 1930, in Lovett, 1919: 230 [Holotype: male, Mt. CAS]. Utah distribution: Grand Co.: Lake Canyon. 10 Rainier, Washington, White River Camp, 4 Oowah; Uintah Co.: Brownie September 1932, in CAS]. Utah distribution: June-22 July. 1874: 362 Cache Co.: Logan, Logan Canyon; Duchesne Cyrtopogon montanus Loew in MCZ]. Co.: Mirror Lake; Rich Co.: Monte Cristo; [Holotype: male. Sierra , August. Utah distribution: Box Elder Co.: Willard Summit Co. : Trial Lake. 5 August-14 Canyon, Cyrtopogon banksi Wilcox and Martin Peak; Cache Co. : Card Canyon, Dry Hodges 1936: 79 [Holotype: male, Puyallup, Wash- Franklin Basin, Green Canyon, West Naomi, ington; 3 June 1933, in CAS]. Utah distribu- Canyon, Hyrum, Logan, Mount J .

62 Great Basin Naturalist Vol. 47, No. 1

# Efferia aestuans 9 Efferia cana Efferia deserti A A Efferia bicolor H Efferia pernicus

{ ~'-\.y'^'^^§00M'^0^

'--,.^7,-:' -

#|f-i|'' *:^

Efferia costal is # Efferia benedicti i .Vi-V/'/ •

80

U'ns'^^

1- -.--;•#: % ' 1 .1 T ^(^Tt:^K.,» ;, •

[i:*"^"*

-'- ^. iv^rV/ ;«

<»i >* .

^^g^VrJ"?^-, ^^^^ Asilidae distribution: 77, £/jfena aestuans, Efferia deserti, and E/jfena pernicus, 78, Efferta ana Ei/ena bicolor , , and E^ena fca.sini ; 79, Efferia benedicti 80, Efferia costalis :

January 1987 Nelson: Utah Robber Flies 63

Providence, Tony Grove, Wellsville Moun- Reef. 1 September-17 October. tains; Daggett Co.: Summit Springs; Duch- Cyrtopogon thompsoni Cole in Cole and esne Co.: Duchesne, Mirror Lake, Wolf Lovett, 1921: 255 [Syntypes: male and fe- Creek Pass, Yellowstone Ranger Station; male, Burns, Oregon, May 1919, in CAS].

Garfield Co.: Aquarius Plateau; Juab Co.: Utah distribution: Box Elder Co. : Curlew Val- Mount Nebo Loop; Rich Co.: Logan Canyon ley; Daggett Co.: Clay Basin; Utah Co.: summit; San Juan Co.: Blanding; Sevier Co.: Goshen Springs. 16 June. willistoni Curran 1922: 277 Highway 4; Summit Co. : Bear River Ranger Cyrtopogon Station, Beaver Creek Ranger Station; Uintah [Holotype: male, Chilcotin, British Colum- Co.: Brownie Canyon, Brush Creek; Wash- bia, 16 June 1920, in CNC]. Utah distribu- ington Co.: Pine Valley Mountains. 12 tion: Box Elder Co. : Devils Gate, Raft River May-15 July. Mtns., Willard Peak; Cache Co.: Franklin Cyrtopogon plausor Osten Sacken 1877: Basin, Logan, Logan Canyon, Mount Naomi, 297 [Syntypes: four males and two females, no Providence, Sardine Canyon, Tony Grove, locality listed, in MCZ]. Utah distribution: Wellsville; Carbon Co.: Carbon Airport; Cache Co.: Cache Valley; Garfield Co.: Davis Co.: Bountiful; Grand Co.: Geyser Aquarius Plateau, Boulder Mountain; Grand Pass, Lake Warner, La Sal Mountains, Min-

Co.: La Sal Mountains; San Juan Co.: Bland- ers Basin, Moab; Iron Co. : Parowan Canyon; ing, Elk Ridge; Summit Co.: Bear River Juab Co.: Trout Creek; Millard Co.: Kanosh,

Ranger Station; Utah Co.: Aspen Grove; Oak Creek Canyon; Rich Co. : Bear Lake Val- ley, Garden City, Logan Canyon summit, Wasatch Co. : Strawberry Valley; Washington

Co.: Pine Valley, Zion National Park; Wayne Walton (Allen) Canyon; San Juan Co. : La Sal Co.: Bicknell, Boulder Mountain, Hanksville; Mtns., Wilson Mesa; Uintah Co.: White- rocks; Wasatch Co.: Strawberry Valley; Weber Co. : Farr West. 4 June-10 August. Cyrtopogon profusus Osten Sacken 1877: Washington Co.: Leeds, Pine Valley, Zion

305 [Syntypes: male and female, Morino Val- National Park; Weber Co. : Monte Cristo. 16 ley, New Mexico, 1 July, in MCZ]. Utah dis- May-14 August. Dicolonus sparsipilosum Back 1909: 247 tribution: Grand Co. : La Sal Mountains; Utah [Cotypes: Two males, Bozeman, Montana, in Co. : Mount Timpanogos. 21 July-17 August. Cyrtopogon pulcher Back 1909: 274 [Holo- Univ. of Mass. and Montana State Univ.].

type: male. Palmer Lake, Colorado, 10 July, Utah distribution: Cache Co. : Franklin Basin, in USNM]. Utah distribution: Box Elder Co. Tony Grove; Rich Co.: Logan Cyn. summit; Devils Gate, Willard Basin; Cache Co.: Utah Co.: Provo Cyn.; Wasatch Co.: Straw- Beaver Mountain, Blacksmith Fork Canyon, berry Valley. 1-17 July. Dry Lake, Logan Canyon, Tony Grove; Dioctria henshatvi Johnson 1918: 103 Daggett Co.: Deep Creek, Sheep Creek; [Holotype: Yakima, Washington, 2 July 1882, Emery Co.: Green River; Garfield Co.: Boul- in MCZ #10036]. Utah distribution: Cache Cyn., Spring Hollow; Rich Co.: der Mountain; Rich Co. : Logan Canyon sum- Co.: Green mit; Summit Co.: Bear River Ranger Station; Logan Cyn. summit, Walton (Allen) Cyn.; Utah Co.: Aspen Grove, Mount Timpanogos; Summit Co.: 10 mi E Kamas; Utah Co.: Oquirrh Mtns., Provo; Washington Co.: Pine Wasatch Co. : Daniels Canyon summit; Wash- ington Co.: Leeds. 22 May-14 August. Valley, St. George; Weber Co.: Ogden. 20 Cyrtopogon rufotarsus Back 1909: 275 July-5 September. [Holotype: male, Gallatin Co., Montana, Dioctria pusio Osten Sacken 1877: 288 California, 8,000-9,000 ft, 9-11 July, in University of [Holotype: Female, Sonoma Co., Massachusetts collection]. Utah distribution: 4 July, in MCZ]. Utah distribution: Utah Co.: Duchesne Co.: Mirror Lake, Uinta Moun- Castella. 22 August. tains; Grand Co.: Geyser Pass; Kane Co.: Dioctria vera Back 1909: 256 [Holotype: Long Valley; Sanpete Co.: Ephraim Canyon; Male, Monterrey Co., California, 2 July 1896,

in AMNH]. Utah distribution: Cache Co. : Lo- Summit Co. : Trial Lake. 14 June-13 August. Cyrtopogon stenofrons Wilcox and Martin gan; Duchesne Co.: Whiterocks; Garfield 1936: 52 [Holotype: male. Grant Co., New Co.: Boulder Mtn.; Grand Co.: Castleton; Mexico, 24 June 1934, in CAS]. Utah distribu- Juab Co.: Deep Creek Mtns. near Callao; Kanosh tion: Cache Co.: Logan; Wayne Co.: Capitol Kane Co.: Mt. Carmel; Millard Co.: 64 Great Basin Naturalist Vol. 47, No. 1 , :

January 1987 Nelson: Utah Robber Flies 65

Cyn., Oak Cr. Cyn.; San Juan Co.: La Sal State Exp. Sta. ; Utah Co. : Payson Cyn. , West Mtns., Mill Cr.; Washington Co.: Deep Cr., Cyn.; Wasatch Co.: Soldier Summit; Wash- Leeds, Pine Valley; Wayne Co.: Grover. L3 ington Co.: Zion National Park; Weber Co.: June-20 August. Huntsville. 6 August—4 September. Diogmites grossus Bromley 1936: 236 Heteropogon martini Wilcox 1965: 207 [Holotype: Male, Lamar, ProwerCo., Colo- [Holotvpe: Male, Montgomery Pass, Nevada, rado, 25 August 1925, in Bromley Collection]. 6 July 1958, in Wilcox Collection in CAS]. Utah distribution: Box Elder Co.: Pilot Utah distribution: Box Elder Co.: Snowville; Range, Promontory, Snowville; Cache Co.: Cache Co.: Logan, Wellsville Cyn.; Daggett

Cornish, Dry Cyn., Hyrum, Logan, Provi- Co. : Sheep Cr. ; Davis Co. : Clearfield; Emery dence; Davis Co.: Clearfield; Duchesne Co.: Co.: 4 air mi N Gilson Butte; Millard Co.:

Bluebell; Grand Co. : Moab; Juab Co. : Topaz Fillmore, warm spring near Gandi; Utah Co.: Mtn.; Kane Co.: Kanab; Millard Co.: Delta, Fairfield, west side Utah Lake, Lehi. 16 Flowell, Garrison, Holden; Salt Lake Co.: June-4 September. Magna, Midvale; Tooele Co.: Camelback, Heteropogon senilis (Bigot) 1878: 423 Little Granite Mtn., Skull Valley, Simpson [Holotype: sex not known, California, in Bigot Butte; Utah Co.: Alpine, Goshen, Hobble collection]. Utah distribution: Box Elder Co.: Cr., Lincoln Beach, Provo, Rock Cyn., Lucin, 1 mi NE Mantua; Cache Co.: Card

Spanish Fork; Washington Co.: Leeds, St. Cyn. , Green Cyn. , Logan Cyn. ; Daggett Co.

George, Zion National Park; Weber Co. : Og- Palisade Park Camp; Iron Co.: Kanarraville, den. South Weber. 30July-ll September. near Parowan; Millard Co.: Oak Cr. Cyn.;

Eucyrtopogon comantis Curran 1923a: 116 Utah Co. : American Fork. 16 June^ Septem- [Holotype: Male, Chilcotin, B.C., 29 April ber. 1920, in CNC #565]. Utah distribution: Utah Heteropogon stonei Wilcox 1965: 207-222 Co.: Aspen Grove, Emerald Lake, Glacier [Holotype: Male, Hualapai Mtns., Arizona, Lake, Mount Timpanogos. No dates with la- 6,000 ft, 4 June 1962, in CAS]. Utah distribu- bels. tion: Beaver Co. : Beaver Cyn. No date given. Eucyrtopogon spp. Utah distribution: Heteropogon species 1. Two specimens, Cache Co.: Dry Cyn., Logan Cyn., Logan undescribed. Utah distribution: Box Elder Mtns.; Tooele Peak, Twin Cr. ; Duchesne Co. : Forest Camp, Co.: Tecoma Range—Copper

Uintah Cyn., Uinta Mtns., Wolf Cr.; Iron Co. : Lofgreen. No dates on labels. Co.: Cedar City; Rich Co.: Logan Cyn. sum- basingeri Pritchard 1938: 130 Spring, San mit; Tooele Co. : Vernon Cr. ; Utah Co. : Aspen [Holotype: Female, Quail Grove; Wasatch Co.: Soldier Summit; Weber Bernardino Co., California, 5 October 1934, Co.: Eden. in Basinger collection]. Utah distribution:

Haplopogon utahensis Wilcox 1966d: Washington Co. : St. George. No date on la- 99-106 [Holotype: Male, 7 mi N St. George, bel. Utah, Hwy 91, 1 June 1963, in CAS]. Utah Holopogon albipilosus Curran 1923b: 207 distribution: Washington Co.: Washington. [Holotype: Male, Vernon, B.C., 5 August 1-8 June. 1920, in CNC #569]. Utah distribution: Heteropogon arizonensis Wilcox 1941: 55 Beaver Co.: Beaver, Greenville, 9 mi E [Holotype: Male, White Mtns., Arizona, Sep- Beaver; Box Elder Co.: Park Valley, Portage; tember, in CAS]. Utah distribution: Millard Cache Co.: Valley, Blacksmith Fork Co.: Oak City. 6-25 June. Cyn., Logan, Mendon; Duchesne Co.: Duch- Heteropogon maculinervis James 1937a: 12 esne, Fort Duchesne, Tabiona, Whiterocks; [Holotype: Female, Masonville, Colorado, 4 Garfield Co.: Panguitch; Grand Co.: Dead Sep 1934, in CSU]. Utah distribution: Beaver Horse Point, Green River, 378 river mile; Paragonah, Parowan; Juab Co.: Co. : 9 mi E Beaver; Box Elder Co. : Clear Cr. Iron Co.: Fielding; Cache Co.: Avon Cyn., Green Nephi, Trout Cr.; Kane Co.: Kanab; Millard Cyn., Providence, Twin Cr.; Daggett Co.: Co.: Oak City, Pahvant, Scipio; Morgan Co.: Pipe Cr.; Grand Co.: La Sal Mtns.; Iron Co.: Devils Slide; Piute Co.: Circleville; Rich Co.: Cedar City; Kane Co.: Zion—Chamberlain Laketown, North Eden, Walton (Allen) Cyn.; Ranch; Tooele Co.: Granite Mtn., Johnsons Salt Lake Co.: Bluffdale, Magna, Salt Lake

Pass; Uintah Co.: Little Mtn., Summit, Utah City; Sanpete Co. : Indianola, Manti, Moroni; 66 Great Basin Naturalist Vol. 47, No. 1

'^ ~ • ' ' . i • ' i?' January 1987 Nelson: Utah Robber Flies 67

Sevier Co. : Elsinore, Glenwood, Richfield; Laphystia utahensis Wilcox 1960: 345 Summit Co.: Red Rock Cyn.; Tooele Co.: [Holotype: Male, St. George, Utah, 12 mi NE Camelback, Dugway Proving Grounds, Little Hwy 17, 23 May 1959, in CAS]. Utah distribu- Granite Mtns.; Uintah Co.: Hayden, La tion: Washington Co.: 12 mi NE St. George Point, Naples, Tridell, Vernal, Whiterocks; (near Hurricane). 22-23 May. Utah Co.: Alpine Loop, Aspen Grove, Lehi, Lasiopogon alhidus Cole and Wilcox 1938: Lindon, North Fork Provo Cyn., Payson, 25 [Holotype: Male, 8 mi E Kiona, Washing- Provo, Provo Cyn., Spring Lake, Springville; ton, 23 April 1933, in CAS]. Utah distribu- Wasatch Co.: Heber, Strawberry Valley; tion: Emery Co.: Green River (Gunnison Washington Co.: Pine Valley; Wayne Co.: Rutte); Grand Co. : Moab. 7 May. Cainville, Grover. 1 June-5 August. Lasiopogon aldrichii Melander 1923b: 139 1-40 Holopogon caesariatus Martin 1959: [Syntypes: Male and female, Moscow Mtn., [Holotvpe: Male, Alpha, Long Valley, Idaho, Idaho, 29 June 1918, repository not listed]. distribution: 6 July 1934, in AMNH]. Utah Utah distribution: Cache Co.: Logan Cyn.;

: Rlacksmith Cyn. 7 August. Cache Co. Fork Duchesne Co.: Roosevelt; Grand Co.: La Sal 1959: 17 [Holo- Holopogon currani Martin Mtns.; Rich Co.: Monte Cristo; Uintah Co.: 1949, in type: Male, Winona, Arizona, 21 July Rrush Cr., Whiterocks; Utah Co.: Mt. Tim- Co.: AMNH]. Utah distribution: Daggett panogos; Washington Co.: Leeds, Pinto. 24 Manila; Washington Co.: Leeds Cyn., Zion June-21 July. National Park; Wavne Co.: Grover. 11-28 Lasiopogon cinereus Cole, in Cole and July. Lovett 1919: 229 [Holotype: Male, Hood Holopogon mingusae Martin 1959: 21 River, Oregon, 24 September 1918, in CAS [Holotype: Male, Mingus Mtn., Arizona, 3 #476]. Utah distribution: Daggett Co.: July 1949, in AMNH]. Utah distribution: Manila; Duchesne Co. ; Uintah Co. : Merkeley Garfield Co.: Aquarius Plateau, The Pass; Park (near Vernal), Whiterocks; Utah Co.: Washington Co.: Zion National Park; Wayne Hobble Cr. 6 July-16 August. Co.: Grover. 11 July-21 August. Lasiopogon monticola Melander 1923b: Holopogon wilcoxi Martin 1959: 33 [Holo- 142 [Syntypes: Males and females, Mt. type: Male, San Carlos Lake, Arizona, May, Adams, Washington, 24 July 1921, no reposi- in CAS]. Utah distribution: Reaver Co.: tory listed]. Utah distribution: Cache Co.: Reaver; Tooele Co.: South Camel Mtn. 13 Green Cyn., Logan Cyn., Mt. Logan; Grand July. Co.: Moab, Wilson Mesa-La Sal Mtns.; Rich Laphystia annulata Hull 1957: 72 [Holo- Co.: Logan Cyn. summit, Monte Cristo; Uin- type: Male, near Navajo, Arizona, 11 July tah Co.: Whiterocks; Weber Co.: Ogden. 26 1954, in Hull collection]. Utah distribution: May-9 July. Kane Co.: Coral Pink Sand Dunes, Mount Lestomyia sabulona Osten Sacken 1877: Carmeljct. 14 July. female, Grafton, Laphystia rubra Hull 1957: 74 [Holotype: 292 [Syntypes: Male and near San Rernardino, California, March, on Female, near Navajo, Arizona, 11 July 1954, dry gravelly soil, in MCZ]. Utah distribution in Hull collection]. Utah distribution: Emery

Rox Elder Co. : Promontory Point; Cache Co. Co.: Goblin Valley, 4 air mi N Gilson Rutte, Logan; Juab Co.: Topaz Mtn.; Salt Lake Co. Wild Horse Creek; Grand Co. : Green River,

Salt Lake Citv; Tooele Co. : Dugway Proving 366 river mile; San Juan Co. : Rluff, Lime Cr. 7 Grounds, Skull Valley. 24 May-9 June. July-26 August. Laphystia tolandi Wilcox 1960: 344 [Holo- Metapogon carinatus Wilcox 1964: 193 type: Male, Lahontan Reservoir, Churchill [Holotype: Male, Whitewater, California, 13 Co., Nevada, 13 June 1949, in CAS]. Utah January 1948, in CAS]. Utah distribution: distribution: Rox Elder Co.: Corinne, Loco- Uintah Co.: Split Mtn. Gorge. Date uncer- "55-28" motive Springs, Tremonton; Cache Co.: Cor- tain, (2-8-1955?). nish; Juab Co.: Fish Springs, Topaz Mtn.; Myelaphus lobicornis Osten Sacken 1877: Millard Co.: Delta, Fillmore; Sevier Co.: 287 [Holotype: Male, Snake River, Idaho, in Richfield; Tooele Co.: Camelback, Granite MCZ]. Utah distribution: Cache Co.: Rlack- Mtn.; Utah Co.: Spanish Fork. 14 June-14 smith Fork Cyn., Green Cyn., Hyrum Dam, August. Logan. 17-22 June. .

68 Great Basin Naturalist Vol. 47, No. 1

9 Heteropoqon macul inervis Holopoqon albipilosus A Heteropogon senilis

B Heteropoqon stonei

*" ; /^i.

)'^'"l • S'^fe

9 Holopogon currani 'Wl^ Holopogon mingusae ?t f A Holopogon caesariatus Holopogon wilcoxi

.->''• "Sv. 91 92

'^M$l --,/'',i^^.'''^*$C'-'

Figs. 89-92. Utah Asilidae, distribution: 89, Heteropogon 7naculinervis , Heteropogon senilis, and Heteropogon stonei; 90, Holopogon albipilosus; 91, Holopogon currani and Holopogon caesariatus ; 92, Holopogon mingusae and Holopogon wilcoxi January 1987 Nelson: Utah Robber Flies 69

Nannocyrtopogon aristatus James 1942: Ospriocerus minos Osten Sacken 1877: 291 126 [Holotype: Male, Arboles, Colorado, [Holotype: Male, Golden City, Colorado, 3

6,700 ft, 17 May 1939, inCSU]. Utah distribu- July, in MCZ]. Utah distribution: Emery Co.: tion: Iron Co.: 5 mi E New Harmony; Wash- 3 mi SSE Temple Mtn., San Rafael Desert; ington Co.: Leeds Cyn., Oak Grove. 23 Grand Co.: Arches National Monument;

May-6 June. Tooele Co. : Dugway Proving Grounds; Uin-

Nicocles abdominalis Williston 1883: 17 tah Go. : 3 mi SW Jensen, Vernal; Utah Co. : 1

[Holotype: Male, California, in KU]. Utah mi W Elberta; Wayne Co. : Notom. 25 July-31 distribution: Washington Co.: Zion National July. Park. No date on label. Ospriocerus vallensis Martin 1968: 401 Nicocles utahensis Banks 1920: 66 [Holo- [Holotype: Male, Grand View, Owyhee Co., type: Female, Eureka, Utah, 31 May, in MCZ]. Idaho, 9 July 1958, in CAS]. Utah distribu- Utah distribution: Cache Co.: Green Cyn., tion: Box Elder Co.: Cedar Hills, Steamboat Hyrum, Logan, Logan Cyn., Sardine Cyn., Springs, Thiokol; Cache Co.: Providence;

USU; Emery Co.: Green River; Grand Co.: Rich Co. : Sage Cr. Jet. ; Sevier Co. : Richfield. Moab; Juab Co.: Eureka; Salt Lake Co.: Little 4 June-U July. Saropogon mohawki Wilcox 1966b: 134 Mtn.; Utah Co. : Provo. 5 March-5 June. Omninablautus nigronotum (Wilcox) 1935: [Holotype: Male, Mohawk, Arizona, 16 July 1 [Holotype: Male, Prairie Hill, Grant Co., 1962, in CAS]. Utah distribution: Washington Oregon, in CAS]. Utah distribution: Box El- Co.: Ivins. 16 July. bradleyi (Bromley) 1937a: 309 der Co. : Etna. 12 August. Scleropogon Ospriocerus abdominalis (Say) 1824: 375 [Holotype: Male (female on same pin). Grant [Holotype: Lost]. Utah distribution: Beaver Forest, California, 9-13 August 1937, in Co.: Minersville; Box Elder Co.: Blue Cr., USNM]. Utah distribution: Wayne Co.: 6 mi Collinston, Mantua, Promontory, Snowville; WTorrey. 15-20 July. Cache Co.: Beaver Mtn., Cornish, Clarkston, Scleropogon coyote (Bromley) 1931: 429 Hyrum, Lewiston, Logan, Mendon, Provi- [Syntypes: Male and female, near Lander, dence, Smithfield; Daggett Co.: Douglas Di- , July, in OSU]. Utah distribution:

nosaur Quarry, Flaming Gorge; Davis Co.: Emery Go. : 4 air mi N Gilson Butte; Grand Farmington; Duchesne Co.: Monarch, Co.: La Sal Mtns.; Utah Co.: Colton. 20-23 Mountain Home, Myton; Emery Co.: 2 mi E July. Gilson Butte, 9 air mi E Castledale, Sinbad Scleropogon duncani (Bromley) 1937a: 307 Country, Wild Horse Cr. north of Goblin Val- [Holotype: Male, Silver City, New Mexico, ley, Woodside; Grand Co.: Castleton, Moab; 24 June 1933, in CAS]. Utah distribution: Iron Co.: Cedar City; Juab Co.: Deep Cr. Washington Co.: Rockville, Santa Clara, Zion Mtns., Fish Springs, Jericho, Levan, Nephi, National Park. 22-24 June. Trout Cr.; Kane Co.: Kanab; Millard Co.: Scleropogon indistinctus (Bromley) 1937a: Delta, Fillmore, Holden, Oasis, Pahvant, Sci- 308 [Syntypes: Male and female. White pio; Rich Co.: Bear Lake Valley, Dry Basin; Mtns., Arizona, August 1930, in CAS]. Utah Provi- Sevier Co.: Fremont Jet., Monroe Cyn., distribution: Cache Co.: Logan Cyn., Richfield; Tooele Co.: Cedar Mtns., Dugw^ay dence; Daggett Co.: Hideout Cyn., Pipe Cr.; Proving Grounds, James Ranch, Lakeside Juab Co.: Nephi; Millard Co.: Antelope Mtns., Little Granite Mtn., Ophir; Uintah Springs; Salt Lake Co.: Salt Lake City; Sevier Co.: Fort Duchesne, Gusher, 3 mi SW Co.: Richfield. 26 May-16 August. Jensen, Vernal; Utah Co.: Cedar Valley, Scleropogon neglectus (Bromley) 1931: 430 Lehi, Provo, West Utah Lake; Wasatch Co.: [Syntypes: Male and Female, near Lander, Deer Cr. Reservoir; Washington Co.: Pine Wyoming, August, in OSU]. Utah distribu- Valley, St. George, Zion National Park; tion: Beaver Co.: Beaver; Box Elder Co.: Wayne Co.: Notom; Weber Co.: Ogden. 9 Clear Cr., Yost; Cache Co.: Beaver Mtn., June-6 September. Hyrum, Logan, Providence; Daggett Co.: Ospriocerus longulus (Loew) 1866: 28 Hideout Cyn., Manila; Emery Co.: Buckskin

[Holotype: in MCZ]. Utah distribution: Springs, Goblin Valley; Garfield Go. : Boulder Grand Co.: Castleton; Washington Co.: Santa Mtn., 15 mi N Boulder; Grand Co.: La Sal Millard Clara. 8-29 July. Mtns.; Juab Co.: Callao, Topaz Mtn.; 70 Great Basin Naturalist Vol. 47, No. 1

Laphria fernaldi

'AC 93

,v-.'*-

"R January 1987 Nelson: Utah Robber Flies 71

Co.: Fillmore, Sutherland; Piute Co.: Utah Cyn. , Salt Lake City; San Juan Co. : Bear Ears; Agricultural Research Station; Rich Co.: Sanpete Co.: Gunnison, Indianola; Sevier Woodruff; Tooele Co.: Camelback, Dugway Co.: Gooseberry, USDA (center of county); Proving Grounds, Lakeside Mtns.; Washing- Tooele Co.: Dugway Proving Grounds, Co.: Douglas, Fort Duch- ton Co. : Toquerville. 29 May-16 August. Timpie; Uintah Scleropogon picticornis Loew 1866: 26 esne, Vernal; Utah Co.: Aspen Grove, Provo [Holotype: Female, California, in MCZ]. Cyn.; Washington Co.: Dixie Park (Snow National Park; Co.: Boul- Utah distribution: Kane Co. : Kanab; San Juan Cyn.), Zion Wayne der Mtn., Hanksville. 8 May-29 Co. : Bluff. 20 August-12 September. July. Sintoria cazieri Wilcox 1972a: 51 [Holo- Stenopogon martini Bromley 1937a: 303 type: Male, Holden, Utah, 18 September [Syntvpes: Male and female, Parma, Idaho, 1959, in AMNH]. Utah distribution: Emery 13 May 1934, in CAS]. Utah distribution: Box Co.: Goblin Valley; Iron Co.: Beryl; Millard Elder Co.: Collinston, Fielding, Snowville; Co.: Holden; Washington Co.: Beaver Dam Cache Co.: Blacksmith Fork Cyn., Cornish, Slope on Joshua Trees, Red Cliffs Cmpgd. at Logan; Daggett Co.: Manila; Davis Co.: An- lights; Weber Co.: Five Points. 16-27 Sep- telope Island, Bountiful, Farmington; Duch- tember. esne Co.: Bluebell, Roosevelt; Garfield Co.: Stenopogon engelhardti Bromley 1937a: Panguitch, Panguitch Lake; Grand Co.:

301 [Syntvpes: Male and female, Jacumba, Moab; Iron Co. : Miners Peak, Parowan Cyn.; California,' 26 April 1935, in CAS]. Utah dis- Juab Co.: Eureka, Topaz Mtn.; Millard Co.: tribution: Box Elder Co.: Raft River Mtns.; Delta, Hatton, McCormick; Rich Co.: Gar- Cache Co.: Blacksmith Fork Cyn., Logan den City, Laketown, Walton (Allen) Cyn.; Cyn., Tony Grove, Wellsville Cyn.; Daggett Salt Lake Co.: Taylorsville; Sanpete Co.: Co.: Elk Park; Duchesne Co.:' 30 mi SW Fountain Green, Indianola, Seely Cr.; Sum- Duchesne; Garfield Co.: Boulder Mtn., mit Co.: Elk Park Cmpgd.; Tooele Co.: Bryce Cyn.; Iron Co.: Cedar City; Juab Co.: Grantsville, Stansbury Island, Timpie, Ver- Eureka; Rich Co.: Logan Cyn. summit; Salt non Cyn.; Uintah Co.: Fort Duchesne, Ver-

Lake Co.: Dry Cyn., Little Mtn., Salt Lake nal; Utah Co. : Aspen Grove, Provo, Spanish City; Sanpete Co.: Indianola; Sevier Co.: Fork; Wasatch Co.: Strawberry Valley; Richfield; Tooele Co.: Lookout Mtn., Stans- Wayne Co.: Fremont, Torrey; Weber Co.: Cyn., Riverdale. 15 May-6 bury Island; Utah Co. : Alpine, Aspen Grove, Ogden, Ogden Payson Cyn., Provo; Washington Co.: Zion September. National Park; Wayne Co.: Hanksville. 8 Stenopogon mexicanus Cole 1923: 463 June-14 August. [Holotvpe: Male, Guaymas, , Mexico, Stenopogon inquinatus Loew 1866: 27 10 April 1921, in CAS '#1341]. Utah distribu- [Syntypes: Male and female, Nebraska, in tion: Cache Co.: Logan; Grand Co.: Castle- MCZ]. Utah distribution: Beaver Co.: Mil- ton. 24 July-1 September. ford; Box Elder Co.: Cedar Cr., Clear Cr., Stenopogon rufiharhis Bromley 1931: 431 Cutler Dam, Devils Gate, Mantua, [Holotype: Male, Lassen Co., California, 20 Washakie; Cache Co.: Green Cyn., Logan, July 1911, in OSU]. Utah distribution: Beaver

; Co. : Devils Gate, Logan Cyn., Providence, Tony Grove, Co. : Beaver Cr. Box Elder Wellsville Cyn.; Carbon Co.: Dragerton, Raft River Mtns., Willard Peak; Cache Co.: Price, Woodhill; Daggett Co.: Flaming Blacksmith Fork Cyn., Card Cyn., Logan, Gorge, Hideout Cyn., Manila, Spirit Lake Newton, Sardine Cyn., Wellsville; Daggett Road, Summit Ranger Station; Duchesne Co.: Manila; Duchesne Co.: Duchesne; Co.: Roosevelt, Yellowstone Ranger Station; Garfield Co.: Aquarius Plateau, The Pass; Garfield Co.: Boulder Mtn., Bryce Cyn., Es- Grand Co.: Castle Valley, Moab (La Sal calante, Panguitch Lake, Tropic; Grand Co.: Mtns.); Iron Co.: Cedar City; Juab Co.: Eu- Val- La Sal Mtns., Westwater; Iron Co.: Parowan reka, Mt. Nebo; Kane Co. : Alton, Long Salt Rich Co. : Garden City; Salt Lake Co. : : Trout Cr. ; Mil- ley; Cyn. ; Juab Co. Fish Springs, lard Co.: Fillmore, Oak Cr. Cyn.; Piute Co.: Lake City; San Juan Co.: Geyser Pass; San- Co.: Fountain Green; Summit Co.: Park Marysvale; Rich Co. : Bear Lake Valley, Lake- pete Vernal; town, Logan Cyn. summit. Woodruff; Salt City; Tooele Co. : Tooele; Uintah Co. : Lake Co.: Big Cottonwood Cyn., Parleys Utah Co.: Alpine, Aspen Grove, Spanish 72 Great Basin Naturalist Vol. 47, No. 1

Wp] % Laphystia rubra '"^ A Laphystia utahensi '"n

97

. : : ;

January 1987 Nelson: Utah Robber Flies 73

Val- Subfamily Laphriinae Fork; Washington Co. : Leeds Cyn. , Pine Hanksville; Weber Co.: Og- ley; Wayne Co.: Atomosia mucida Osten Sacken 1887: 184 den. 20 May-25 July. [Holotype: Sex not known, Presidio, Mexico]. 8 Stenopogon utahensis Bromley 1951: Utah distribution: Washington Co.: Leeds Utah, 20 [Syntypes: Male and female, Leeds, Cyn., Rockville, Santa Clara. 18 July-20 Au- distribution: June 1929, in USNM]. Utah gust. 1-20 Washington Co.: Leeds, St. George. Cerotainiops abdominalis (Brown) 1897: June. 103 [Holotype: information not found]. Utah Stichopogonfragilis Back 1909: 334 [Holo- distribution: Beaver Co.: Milford; Garfield Alamagordo, New Mexico, 24 type: Female, Co. : Escalante Desert, The Hall; Millard Co. April 1902, in AESP]. Utah distribution: Hatton, Hinckley, Kanosh; San Juan Co.: SSE Tem- : Rafael Desert 3 mi Emery Co. San Bluff; Tooele Co. : Dugway Proving Grounds.

Co. : Delta; Tooele ple Mtn. , 5,300 ft; Millard 5 June-12 July. Grounds; Washington Co. : Dugwav Proving Dasylechia atrox (Williston) 1883: 28 Co.: Santa Clara; 3 mi NW St. George. 24 [Holotype: Sex not known, Pennsylvania].

May-4 July. Utah distribution: Cache Co. : Logan. No date Stichopogon salinus (Melander) 1923c: 216 on label, Utah specimen in AMNH. [Holotype: Male, Great Salt Lake, Utah, 31 Laphria felis (Osten Sacken) 1877: 286 July 1908, in Aldrich Collection]. Utah distri- [Holotype: Webber Lake, Sierra Nevada, bution: Beaver Co.: Minersville; Cache Co.: California, no sex or repository listed in Osten Cornish, Logan; Juab Co.: Topaz Mtn.; Mil- Sacken (1878) or McAtee (1919)]. Utah distri- Salt Lake lard Co. : Delta, Hatton, Pahvant; bution: Box Elder Co.: Willard Peak; Kane

: Aspen Grove, Co. : Great Salt Lake; Utah Co. Co.: Duck Lake, ; San Juan Co.: Goshen, Provo, Spanish Fork, west Utah Bear Ears; Summit Co.: Trial Lake; Weber Lake. 20 June-3 August. Co.: Ogden, Ogden Peak, Willard Peak. 17 (Say) 1823: 51 June-3 August. [Holotype: Lost]. Utah distribution: Duch- Laphria fernaldi (Back) 1904: 290 [Syn- Kane Co. in either esne Co. : Indian Creek, Roosevelt; types: Males and females, Colorado, Kanab, Zion National Park near east entrance; UM or AESP]. Utah distribution: Cache Co.: Uintah Co.: Naples; Washington Co.: Santa Green Cyn., Logan, Logan Cyn., Twin Au- Co. Clara, Zion National Park. 25 June-19 Creek; Daggett Co. : Elk Park; Duchesne gust. Garfield Co.: Bryce Cyn.; Iron Co.: Deer Taradieus ruficaudus Curran 1930: 4 Haven Cmpgd.; Rich Co.: Logan Cyn. sum- [Holotype: Female, Mud Springs, Santa mit; San Juan Co.: Bear Ears; Summit Co.: Catalina Mtns., Arizona, 17-20 July 1916, in Trial Lake; Uintah Co.: Whiterocks Cyn., AMNH]. Utah distribution: Alpine. 14 July. Utah Co.: Aspen Grove, Hobble Cr. Cyn., Na- : Pine Valley, Zion Wilcoxia painteri Wilcox 1972b: 43-47 Provo; Washington Co. Co.: Hanksville. [Holotype: Male, Datil, Continental Divide, tional Park; Wayne Catron Co., New Mexico, 17 July 1930, in June-September. [Holo- Painter Collection?]. Utah distribution: Box (Linnaeus) 1758: 605 Lo- Emery type: Lost]. Utah distribution: Cache Co. : Elder Co. : Howell, Nafton, Showell;

Co.: east of Block Mtn., Sinbad Country; gan; Garfield Co. : Bryce Cyn. 5 June. janus McAtee 1919: 153 [Holo- : mi Garfield Co. : 4 mi N Boulder; Kane Co. 40 Laphria E Kanab, 16 mi W Glen Cyn., 3 mi W Wah- type: Male, near summit of Mt. Washington, weep, Zion National Park near east entrance; New Hampshire, in USNM]. Utah distribu- Co.: Uinta Mtns.; Garfield : 16 mi tion: Duchesne Millard Co. : 23 mi W Delta: Uintah Co. dates on labels. SW Vernal; Wayne Co.: east edge Capitol Co. : Aquarius Plateau. No Reef. 19 May-16 September. Laphria sadales Walker 1849: 378 [Holo- British Willistonina bilineata (Williston) 1883: 11 type: Sex not known. New York, in Co.: [Holotype: Female, Northern California, in Museum]. Utah distribution: Cache Mirror Lake, KU]. Utah distribution: Wasatch Co.: Trout Green Cyn.; Duchesne Co.: Cyn. summit, Cr. Spring (near Strawberry Reservoir). 20 Uinta Mtns.; Rich Co.: Logan in Rock August. Monte Cristo; Summit Co.: Hole 74 Great Basin Naturalist Vol. 47, No. 1

% Machimus adustus J^Wrl r, K, >. i-

101

•^^ V

-•"•i

9 Machimus griseus

103

'^l^r(-^ :; :

January 1987 Nelson: Utah Robber Flies 75

Cyn.; Uintah Co.; Utah Co.: A.spen Grove, Efferia apache Wilcox 1966a: 141 [Holo-

Provo Cyn. ; Weber Co. : Pineview. 21 July-16 type: Male, Chambers, Arizona, 18 May August. 1965, in CAS]. Utah distribution: Emery Co.: Laphria vivax Williston 18(S3: 30 [Holo- 4 mi N Gilson Butte, Goblin Valley, San type: Sex not known, Washington Territory, Rafael Desert, 3 mi SSE Temple Mtn.', 5,300 no repository hsted in McAtee (1918)]. Utah ft; San Juan Co. : Bluff. 16 April-29 May. distribution: Cache Co.: Logan Cyn., Tony Efferia arida (Williston) 1893: 254 [Holo- Grove; Summit Co.: 3 mi SE Bear River type: Male, Death Valley Expedition, April Ranger Station; Wasatch Co.: Silver Lake. 5 1891, in USNM]. Utah distribution: Washing- August-11 September. ton Co.: Leeds. 4-23 April. E/jTerifl [;asmi Wilcox 1966a: 190 [Holotype: Subfamily Asilinae Male, 28.5 mi W Eureka, Eureka Co., Ne- Asilus auriannulatus Hine 1906 [Syntypes: vada, 6 Jime 1960, in CIS]. Utah distribution: Male and female, Hope Mtns., B.C., 1 July, Box Elder Co.: Cedar Cr., Snowville; Grand no repository listed]. Utah distribution: Co.: 12 mi NW Moab; Wayne Co.: Horse Cache Co.: Blacksmith Fork Cyn., Green Valley. 28 May-19 June. Cyn., Little Bear Cr., Logan, Logan Cyn., Efferia benedicti (Bromley) 1940: 15 [Holo- Twin Creek, Wellsville; Rich Co.: Logan type: Male, Winslow, Arizona, 13 June 1937, Cyn. summit. 1-26 July. in KU]. Utah distribution: Beaver Co.: Mil- Asilus formosus Hine 1918: 321 [Holotype: ford, Minersville Reservoir; Box Elder Co.: Male, Clary Co., Kansas, 29 August 1911, in Clear Cr., Kelton, Lampo Jet., Mantua, the Hine Collection (OSU?)]. Utah distribu- Promontory, Raft River Mtns., Rattlesnake tion: Box Elder Co.: 25 mi SW Snowville; Pass, Snowville, Tecoma Range; Cache Co.: Cache Co.: Petersboro; Millard Co.: Fill- Beaver Mtn., Blacksmith Fork Cyn., Dry more; Tooele Co.: Lakeside Mtns.; Weber Cyn., Green Cyn., Logan, Logan Cyn., Prov-

Co.: Huntsville. 10 July-13 August. idence; Carbon Co. : Carbon Airport, Drager- Asilus vescus Hine 1918: 320 [Holotype: ton, Kennilworth, Price, Woodhill; Davis Male, Monterrey Co., California, 2 July 1896, Co.: Antelope Island, Farmington; Daggett in AMNH]. Utah distribution: Box Elder Co. Co.: Bridgeport; Duchesne Co.: Duchesne; Willard Basin; Cache Co.: Blacksmith Fork Emery Co.: Buckskin Springs, Block Mtn., 4 Cyn.; Daggett Co.: Manila; Millard Co.: mi N Gilson Butte, Sinbad Country; Garfield Delta; Rich Co.: Randolph. 15 June-1 Sep- Co.: Boulder, 11 mi E Escalante, Escalante tember. Desert, Halls Cr. , Long Hollow, Shitamaring Efferia aestuans (Linnaeus) 1763: 413 Cyn., Ten Mile, Willow Tanks; Grand Co.:

[Holotype: Lost]. Utah distribution: Daggett Castle Valley, Moab; Iron Co. : Cedar City, Springs, Callao, Co. : Manila; Garfield Co. : Bryce Cyn. ; Wash- Parowan; Juab Co.: Antelope Range, To- ington Co. : Snow Cyn. 19 May-11 August. Deep Cr. Mtns., Levan, Thomas Efferia albibarbis (Macquart) 1838: 118 paz Mtn., Trout Cr., Utah Agricultural Ex- [Holotype: Sex not known. North America,' periment Station north of Levan; Kane Co. repository not found]. Utah distribution: Glendale, Kanab, Kodachrome Basin, Zion Cache Co.: Cornish, Logan, Paradise, National Park; Millard Co.: Confusion Pass, Wellsville; Duchesne Co.: Myton; Emery Delta, Fillmore, Flowell, Garrison, Hatton, Co.: Green River, Woodside; Garfield Co.: Oak City, Pahvant, Scipio Lake; Piute Co.: Henrieville, Ten Mile (Escalante Desert); Marysvale; Salt Lake Co.: Dry Cyn., Fort

Grand Co.: Arches, Castleton, Castle Valley, Douglas, Salt Lake City; San Juan Co. : Bland- La Sal Mtns., Moab; Juab Co.: Trout Cr. ing, Hite, Kane Springs, Lime Cr., Monti- Millard Co.: Delta, Fillmore; San Juan Co.: cello, Bluff, Monument Valley, Natural Bluff; Tooele Co.: Little Granite Mtns.; Uin- Bridges National Monument, Navajo Mtn. tah Co.: Fort Duchesne, Maeser; Utah Co.: Trading Post; Sanpete Co.: 8 mi NE Fountain east side Utah Lake, Provo; Washington Co.: Green; Sevier Co.: Fish Lake, Fremont Jet., Leeds, Pinto, Rockville, Santa Clara, Snow Gooseberry, Richfield; Tooele Co.: Cedar Cyn., St. George, Zion National Park—Coal- Mtns., Clifton, Delle, Dugway Mtns., be- passes. Lookout pits Wash, Narrows; Wayne Co. : Notom; We- tween Johnson and Douglas Valley, Simpson Springs, ber Co. : Hooper, Riverdale. 7 May-1 August. Mountain, Skull .

76 Great Basin Naturalist Vol. 47, No. 1

Machimus paropus

105

i,/';:\-v: , V-. W&i

j(^M- : —

January 1987 Nelson: Utah Robber Flies 77

Stansbury Island, Tinipie, Wendover; Uintah tvpe: Male, Antelope Mtn., Harney Co., Ore- Co.: Dinosaur National Monument, Evacua- gon, 6,500 ft, 9 August 1931, in CAS]. Utah tion Cr., Vernal; Utah Co.: Alpine, Aspen distribution: Box Elder Co. : Clear Cr. , Kelton Grove, Cedar Valley, Goshen, Oquirrh Pass, Promontory, Yost; Cache Co.: Logan; Mtns., Payson, Provo, Rock Cyn., Spanish Daggett Co.: Dutch John, Hideout Cyn., Fork, West Cyn., west Utah Lake, Y Mtn.; Manila; Duchesne Co.: Duchesne, Monarch, Washington Co.: Beaver Dam Mtns., Dixie My ton, Roosevelt, 10 mi N Neola; Emery National Forest, Hurricane, Ivins, Leeds Co.: Buckhorn Flats, fossil beds. Green Co.: Cyn., St. George, Summit, Toquerville, River, Gilson Butte; Garfield Hen- rieville, Henry Mtns.; Juab Co.: Eureka, To- Veyo, Zion National Park; Wayne Co. : Exper- iment Station, Fruita, Grover, Hanksville, paz; Kane Co.: Zion National Park—Cham- Notom, Torrey, 6 mi W Torrey; Weber Co.: berlain Ranch; Millard Co.: Fillmore, Desert FarrWest, North Ogden, Ogden. 23April-28 Experiment Station; Sanpete Co.: 8 mi NE August. Fountain Green; Sevier Co.: Richfield; Mountain; Efferia bicolor (Bellardi) 186L 47 [Holo- Tooele Co.: Clifton, Little Granite Na- type: Male, 'Mexico,' repository not hsted in Uintah Co.: Diamond Mtn., Dinosaur Vernal; Wilcox (1966)]. Utah distribution: Box Elder tional Monument, Jensen, Red Wash, Co.: Kelton; Cache Co.: Logan; Daggett Co.: Utah Co.: Alpine, Goshen, Provo, west Utah

Bridgeport, Hideout Cyn.; Iron Co.: Cedar Lake; Wasatch Co. : Strawberry Valley; Wash- Leeds, City; Juab Co.: Trout Cr.; Washington Co.: ington Co.: Crystal Cr., Hurricane, Zion National Park. 19 July-24 July. Pintura. 26 July-8 September. 1937b: 104 Efferia cana (Hine) 1916: 22 [Syntypes: Efferia knowltoni (Bromley) Males and females, Claremont, California, in [Syntypes: Male and female. Trout Creek, not seen OSU]. Utah distribution: Juab Co.: Fish Utah, 5-6 1934, in USU or USNM ( Trout Springs; Kane Co.: Wahweep Va.; Millard at USU)]. Remarks: A specimen labeled Creek, Utah, 5-6 1934 but without a type : Co. Co. : Delta; Utah Co. Provo; Washington examined, 15 mi SW Shivwits, Hurricane, La Verkin, label in the USU collection was Wilcox Leeds, Pintura, Santa Clara, St. George, 3 mi and it ran to Efferia benedicti in the specimens examined N St. George; Wayne Co.: Hanksville. 23 (1966a) key. No male speci- April-1 June. during this study, including numerous species in Efferia costalis (Williston) 1885: 64 [Holo- mens from Trout Creek, ran to this run to this type: None hsted]. Utah distribution: Box El- the key. Some females do indeed of the der Co.: Curlew Valley; Cache Co.: Franklin species based on the position of the fork closer Basin; Carbon Co.: Price; Daggett Co.: Elk third vein (in the Wilcox key). Upon Sta- observation this character was found variable Park; Duchesne Co. : Yellowstone Ranger in those from the same tion; Garfield Co.: Bryce Cyn., Daves Hol- in the specimens, even species low, Panguitch Lake; Iron Co.: Kanarraville; locality. Therefore, status of this

£ . bene- Kane Co.: Coral Pink Sand Dunes; Millard should be that of synonomy under this species Co.: Antelope Mtn.; Rich Co.: Sage Cr. Jet.; dicti until characters separating Sah Lake Co.: Parleys Cyn.; San Juan Co.: can be found. 173 Blanding; Tooele Co.: James Ranch—Gov- Efferia mortensoni Wilcox 1966a: [Holotype: Male, 2 mi NE Portal, Arizona, 23 ernment Spring; Washington Co. : New Har- mony. 9 June-7 August. October 1962, in CAS]. Utah distribution: 7 October. Efferia davisi Wilcox 1966a: 132 [Holotype: Snow Cyn., St. George. [Syn- Male, 12 mi N Sasabe, Arizona, 5 August Efferia pernicus Coquillett 1893: 175 Los Angeles and San 1962, in CAS]. Utah distribution: Washington types: Male and female, in USNM]. Utah Co.: Ivins, Leeds, Rockville, Santa Clara, Diego counties, California, Zion National Park Zion National Park. 15-19 July. distribution: Kane Co.:

Washington Co. : Boilers Efferia deserti Wilcox 1966a: 199 [Holo- Chamberlain Ranch; Flats. 29 type: Male, 10 mi E Desert Center, Califor- in Washington, Rockville—Duncan nia, 13 April 1941, in CAS]. Utah distribution: July-5 September. 1919: 136 [Syn- Washington Co.: Santa Clara, St. George, Efferia producta (Hine) Springs, Zion National Park. 4-15 May. tvpes: Male and female, Flinn 9 August 1917, in OSU]. Efferia frewingi Wilcox 1966a: 169 [Holo- Lakeside, California, 78 Great Basin Naturalist Vol. 47, No. 1

Wetapoqon carinatus

Myelaphus lobicornis

hlannocyrtopoqon aristatus

W W- 9 niw Negasilus mesae

111

5"^-^;*'-^ii:*-2i:;L...::.„.: January 1987 Nelson: Utah Robber Flies 79

Utah distribution: Kane Co. : Kanab; San Juan Lake City; San Juan Co. : Grand Gulch, Natu- Co.: Navajo Mtn.; Washington Co.: Zion Na- ral Bridges National Monument; Washington tional Park. 17 July-28 August. Co. : Crystal Cr. , Deep Cr. , Leeds Cyn. , Zion Efferia staminea (Williston) 1885: 68 [Holo- National Park—Court of the Patriarchs, Oak type: Male, Montana, in KU]. Utah distribu- Cr. 6 May-6 August. tion: Daggett Co. : Pipe Creek. 6 August. Efferia zonata (Hine) 1919: 112 [Syntypes: Efferia subarida (Bromley) 1940: 14 [Holo- Male and female, southern Arizona, August type: Types—male and female, Tucson, Ari- 1902, in OSU]. Utah distribution: Box Elder zona, 8 March 1937, in KU]. Utah distribu- Co. : Clear Cr. , 5 mi N Kelton, Mantua; Cache tion: Garfield Co.: Shitamaring Cyn. 18 May. Co.: Blacksmith Fork Cyn., Dry Cyn., Green Efferia siibcuprea (Schaeffer) 1916: 66 Cyn., Hatties Grove, Logan, Providence [Holotype: Male, Prescott, Arizona, in Cyn.; Daggett Co.: 14 mi S Manila; Emery USNM]. Utah distribution: Beaver Co.: Co.: San Rafael Reef, S. Temple Wash;

Beaver: Box Elder Co.: Willard Basin; Garfield Co. : Henrieville; Juab Co. : Antelope Garfield Co.: Aquarius Plateau; Grand Co.: Springs; Millard Co.: Delta, Fillmore, Moab; Iron Co.: Parowan; Kane Co.: Paria Gaudy; San Juan Co.: Hite; Sanpete Co.: River; San Juan Co.: Kane Springs; Tooele Ephraim; Tooele Co.: Deep Cr. Mtns., Hick- Co.: Little Granite Mtn.; Utah Co.: Provo man Cyn., Johnson Pass, Mercur, Whiskey

Cyn. ; Washington Co. : Zion National Park. 29 Springs; Utah Co.: Payson, Provo; Washing- June-29 August. ton Co.: Leeds, Snow Cyn., Zion National Efferia siibpilosa (Schaeffer) 1916: 67 Park. 30 June-28 August. [Holotype: Male, Beaver Creek Hills, Beaver Machimus adustus Martin 1975: 27 [Holo- Co., Utah, in USNM]. Remarks: Wilcox type: Male, White Mtns., Hive Springs, (1966a) sensed problems with respect to the Arizona, 18 July 1949, in CAS]. Utah distribu- identity of this species. Apparently no speci- tion: Beaver Co.: Milford; Box Elder Co.: mens which he studied could confidently be Clear Cr., Mantua; Cache Co.: Green Cyn., placed in this species. I have not seen any Logan, Logan Cyn., Tony Grove, Twin Cr., specimens which would fit the descriptions West Hodges Cyn.; Daggett Co.: 4 mi E outlined by the Wilcox (1966a) key. The type Deep Cr., Pipe Creek Summit Ranger Sta- should be examined to determine the status of tion; Garfield Co.: Boulder, Bryce Cyn.; Mil-

this species. lard Co. : Delta; Rich Co. : Garden City, Logan Efferia tucsoni Wilcox 1966a: 231 [Holo- Cyn. summit; San Juan Co.: Kane Springs, type: Male, Portal, Arizona, 23 July 1963, in Navajo Mtn.; Sanpete Co.: Centerfield; Sum-

CAS]. Utah distribution: Garfield Co.: Shita- mit Co. : 16 mi "P Kamas; Tooele Co. : Tooele; maring Cyn.; San Juan Co.: Natural Bridges Uintah Co.: Whiterocks Cyn.; Utah Co.: As- National Monument; Washington Co.: Leeds pen Grove; Wasatch Co.: Keetley, Straw- Cyn., Paradise Cyn. (nearlvins), Santa Clara, berry Reservoir; Washington Co.: Kolob, Zion National Park. 14-27 July. Pine Valley, Zion National Park—East Rim, Efferia utahensis (Bromley) 1937b: 103 Lava Point; Weber Co.: Huntsville, Uintah. [Holotype: Male, Price, Utah, 26 August 20 June-18 September. 1935, in USU or USNM (not seen at USU)]. Machimus callidus Williston 1893: 75 [Syn- Utah distribution: Carbon Co.: 1 mi S Price; types: Male and female, Mt. Hood, in KU]. Emery Co.: 2 air mi W Little Gilson Butte, Utah distribution: Box Elder Co.: Clear Cr., Goblin Valley (in wash), 4 air mi N Gilson Willard Basin, Yost; Cache Co.: Blacksmith Butte; Garfield Co.: Henry Mtns., Wickiup Fork Cyn., Franklin Basin, Green Cyn., Lo- Pass; Grand Co.: Westwater; San Juan Co.: 8 gan, Logan Cyn., Providence, Tony Grove, mi NE Mexican Hat. 15 August-17 Septem- White Pine Lake; Daggett Co.: Hideout ber. Cyn., Manila; Grand Co.: Castle Valley; Juab Efferia willistoni (Hine) 1919: 110 [Syn- Co.: Trout Cr.; Rich Co.: Logan Cyn. sum- types: Male and female, Williams, Arizona, 21 mit, Monte Cristo, Walton (Allen) Cyn.; July, in USNM]. Utah distribution: Cache Tooele Co.: Vernon; Uintah Co.: Jensen; Co.: Green Cyn., Logan Cyn.; Grand Co.: Washington Co.: Zion National Park. 6 Castleton, La Sal Mtns.; Millard Co.: Cove May-10 September. Fort, Fillmore; Salt Lake Co.: Magna, Salt Machimus griseus Hine 1906: 29 [Syntypes: 80 Great Basin Naturalist Vol. 47, No. 1

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January 1987 Nelson: Utah Robber Flies 81

Male and female, 'Southwestern Colorado," tory listed]. Utah distribution: Uintah Co.: no repository listed]. Utah distribution: Randlett (?); Washington Co.: La Verkin, St. Cache Co.: Blacksmith Fork Cyn.; Daggett George, Washington, Zion National Park. 4 Co.: Manila; Grand Co.: Castleton; Juab Co.: July-15 August.

Callao; Kane Co. : Kanab; San Juan Co. : Natu- Megaphorus frustra (Pritchard) 1935: 11 ral Bridges National Monument; Uintah Co.: [Holotvpe: Male, Tem (south of Whitewater), Fort Duchesne, Whiterocks; Utah Co.: Hob- Arizona, 19 August 1934, in AMNH]. Utah ble Cr. Cyn.; Washington Co.: Boilers distribution: Garfield Co.: Cass Cr. Reser- Washington, Crystal Cr., Deep Cr., Leeds, voir; Iron Co.: Cedar City, Iron Springs, Zion National Park—Court of the Patriarchs; Parowan; Kane Co.: Kanab; Millard Co.:

, : Wayne Co. : Grover. 10 July-12 September. Chalk Cr. Leamington; San Juan Co. Bland- Machimus occidentalis (Hine) 1909: 147 ing, Monticello; Tooele Co.: Cedar Mtns.; [Type: None designated, several specimens, Washington Co.: Anderson Jet., Ash Cr., noted by Hine (1909) from B.C., Nevada, Gunlock, Leeds Cyn., Little Pinto, Pintura, California, Oregon, and Washington]. Utah Utah State Line U 59, Zion National Park. 14 distribution: Box Elder Co.: Clear Cr., June-26 August. Curlew Valley, Hansel Valley, Mantua, Raft Megaphorus guildiana (Williston) 1885: 60 River Mtns.; Cache Co.: Ant Valley, Black- [Types: western Kansas, Montana, North smith Fork Cyn., Green Cyn., Logan Cyn., Carolina, in KU]. Utah distribution: Box El- Providence, Tony Grove, Twin Creek; Grand der Co.: Snowville; Cache Co.: Green Cyn., Co.: La Sal Mtns.; Juab Co.: Trout Cr.; Mil- Logan; Daggett Co.: Forest Camp, Hideout lard Co.: Delta, Holden; Rich Co.: Logan Cyn.; Juab Co.: Nephi; Kane Co.: Kanab;

Cyn. summit; San Juan Co.: 25 mi S Moab; Sanpete Co. : Fairview; Uintah Co. : La Point,

: 5-23 Uintah Co. : Jensen; Utah Co. South Fork of Tridell. August. Provo Cyn.; Washington Co.: Leeds, Pine Megaphorus paUidus (Johnson) 1958: 41

Valley, Zion National Park; Weber Co. : Og- [Holotype: Male, west side of Little Granite den. 3 June-25 July. Mtn., Tooele Co., Utah, 21 August 1955; in Machimus paropus (Walker) 1849: 455 Johnson collection (paratypes seen at BYU)]. [Holotype: Not listed in Hine (1909)]. Utah Utah distribution: Millard Co.: 9 mi E Delta; distribution: Beaver Co.: Milford; Box Elder Tooele Co.: Cedar Mtns., Hickman Cyn., Co.: Grouse Cr., 25 mi SW Snowville; Cache Simpson Mtns., west side of Little Granite Co.: Clarkston, Cornish, Mendon, Peters- Mtn. 23 July-21 August. boro, Richmond, Smithfield Cyn.; Carbon Megaphorus pulcher (Pritchard) 1935: 11 Co.: Helper; Daggett Co.: Manila; Juab Co.: [Holotype: Carlsbad, New Mexico, 29 August Chicken Cr. Reservoir; Millard Co.: Delta, 1934, in AMNH]. Utah distribution: Wash- Val- Fillmore; Rich Co. : Laketown, Randolph; Salt ington Co.: Enterprise, Middleton, Pine St. National Lake Co. : Midvale; Sanpete Co. : Centerfield, ley, Rockville, George, Zion Ephraim, Fayette, Sanpete; Tooele Co.: Mills Park. 10 August-14 October.

Jet.; Uintah Co.: Fort Duchesne; Utah Co.: Megaphorus willistoni (Cole,) in Cole and American Fork, Provo, South Fork of Provo Pritchard 1964: 74 [Holotype: Male, Hallelu- Cyn., Spanish Fork; Washington Co.: St. jah Jet., Lassen Co., California, 26 July, in George; Weber Co.: Huntsville. 6 May-2 CAS]. Utah distribution: Beaver Co.: Beaver; September. Box Elder Co.: Clear Cr.; Cache Co.: Black- Machimus sestertius Martin 1975: 29 smith Fork Cyn., Green Cyn., Logan; Emery

, Co. [Holotype: Male, 4.5 mi E Moenkopi, Co- Co. : Block Mtn. Sinbad Country; Grand conino Co., Arizona, 14 July 1966, in CAS]. Moab; Millard Co.: Fillmore; Salt Lake Co.:

Utah distribution: Kane Co.: Coral Pink Sand Salt Lake City; Sanpete Co. : 8 mi NE Foun- Dunes, Kanab, 10 mi N Kanab; Tooele Co. tain Green; Utah Co.: BYU, Provo, Rock Dugway Proving Grounds; Washington Co.: Cyn., Spanish Fork, Spring Lake; Washing- Crystal Cr., Deep Cr., Santa Clara, St. ton Co.: Zion National Park. 8-24 August. George, Zion National Park. 13 May-21 Sep- Negasilus belli Curran 1934: 184 [Holo- tember. type: None listed]. Utah distribution: Box El- MallophorafautrixOsten Sacken 1877: 191 der Co.: Portage; Cache Co.: Logan, Men- [Holotype: Presidio, Mexico, no sex or reposi- don; Duchesne Co.: Duchesne; Garfield Co.: 82 Great Basin Naturalist Vol. 47, No. 1

\A .im ;

January 1987 Nelson: Utah Robber Flies 83

Boulder; Juab Co.: Chicken Cr., Nephi; Mil- Co.: Crystal Cr., Deep Cr. , Leeds Cyn., lard Co.: Delta; Rich Co.: Laketown, Ran- Rockville, Zion National Park—Birch Cr., dolph, Walton (Allen) Cyn., Woodruff; San- Coalpits Wash, Court of the Patriarch, Nar- pete Co.: Fairview; Sevier Co.: Koosharem rows, Taylor Cr. 17 June-28 August. Reservoir; Uintah Co.: Vernal; Utah Co.: Philonicus limpidipennis (Hine) 1909: 167 Provo; Washington Co.: Santa Clara, Virgin, [Syntypes: Male and female, southwestern Zion National Park. 13 June-23 July. Colorado, 14 July 1899, repository not listed]. Negasilus mesae (Tucker) 1907: 51 [Holo- Utah distribution: Grand Co.: Moab; Juab type: Colorado Springs, Colorado, in KU]. Co. : Callao; Salt Lake Co. : Salt Lake City; San Utah distribution: Box Elder Co.: Bird Juan Co.: Indian Cr.; Sevier Co.: Richfield;

Refuge, Blue Cr., Brigham City, Corinne, Uintah Co. : Vernal Refuge; Utah Co. : Spanish

Howell, Kelton, Locomotive Springs, Nafton, Fork; Washington Co. : Rockville. 18 June-30 Snowville; Cache Co.: Beaver Mtn., Benson, August. Cornish, Logan Cyn. —Twin Cr.; Daggett Polacontha composita (Hine) 1918: 321 Co.: Manila; Duchesne Co.: Myton; Emery [Holotype: Male, San Diego, California, 30 Co.: 9 air mi E Castledale, Sinbad Country June 1913, in Hine Collection]. Utah distribu-

east of Block Mtn.; Crand Co.: Moab; Kane tion: Cache Co. : Logan; Iron Co. : Cedar City; Co.: Navajo Lake; Millard Co.: Delta, Pah- Juab Co.: Trout Cr.; Utah Co.: Saratoga; vant; Rich Co.: Sage Cr. Jet., Woodruff; San Washington Co.: Leeds, Leeds Cyn., Par- Juan Co.: Kane Springs; Sanpete Co.: Indi- adise Cyn., St. George, Virgin, Zion National

anola; Sevier Co. : Fish Lake, Sevier; Summit Park. 27 June-28 August. Co.: Park City; Tooele Co.: Delle, losepa, Proctacanthella cacopiloga (Hine) 1909: 65 Timpie; Uintah Co.: Jensen; Utah Co.: El- [Syntypes: Several specimens from many berta, Fairfield; Wasatch Co.: Heber, Provo states, see Hine (1909), in KU]. Utah distribu- Cyn. 18 May-12 August. tion: Daggett Co.: Manila; Duchesne Co.: Neoitamus brevicomus (Hine) 1909: 155 Roosevelt; Emery Co.: Green River, Wild [Syntypes: Male and female, Kalso, B.C., Horse Cr. north of Goblin Valley; Grand Co.: repository not listed]. Utah distribution: 12 mi NW Moab; Uintah Co.: Vernal. 6

Garfield Co. : Long Hollow; Rich Co. : Monte June-16 August.

Cristo; Salt Lake Co. : Mill Creek Ranger Sta- Proctacanthus hinei Bromley 1928: 13 tion; Utah Co.: Aspen Grove. 12 July-11 Au- [Holotype: male, Albuquerque, New Mexico, gust. 3 August 1925, S. W. Bromley., in Bromley Neomochtherus albicomus {Hine) 1909: 136 Collection]. Utah distribution: San Juan Co.: [Syntvpes: Male and female, Montana, in Sand Island Cmpgd. 2 mi S Bluff. 22-23 Au- USNM]. Utah distribution: Box Elder Co.: gust.

Clear Cr. ; Cache Co. : Green Cyn. , Twin Cr. Proctacanthus micans Schiner 1867: 397 Daggett Co.: Hideout Cyn., Manila. 19 [Holotype: None listed in Hine (1911)]. Utah July—4 September. distribution: Duchesne Co.: Boneta; Kane Neomochtherus hypopygialis (Schaeffer) Co.: Kanab, Zion National Park; Millard Co.: 1916: 68 [Syntypes: Two males, Beaver Cyn., Delta; San Juan Co.: Navajo Mtn.; Uintah Utah, in AMNH]. Remarks: No specimens of Co.: Bonanza, Fort Duchesne, Jensen, Ver-

this species were seen in collections during nal; Washington Co. : Leeds Cyn. , Pine Valley this study. Cmpg, Zion National Park; Wayne Co.; We- Neomochtherus lepidus (Hine) 1909: 136 ber Co.: Farr West, Ogden. 15 July-19 Au- [Holotype: Male, Colorado, also female. gust.

White Mtns. of New Mexico about 6,000 ft, 23 Proctacanthus milberti Macquart 1838: July, in USNM]. Utah distribution: Box Elder 124 [Holotype: None listed in Hine (1911)]. Co.: Clear Cr.; Uintah Co.: Bonanza; Wash- Utah distribution: Cache Co.: Cornish; ington Co.: Pine Valley, Zion National Park. Daggett Co.: Manila; Duchesne Co.: Roo-

24 August. sevelt; Uintah Co. : Vernal; Utah Co. : Goshen; Philonicus arizonensis (Williston) 1893: 76 Washington Co.: Zion National Park. 27 [Holotype: Female, Arizona, in KU]. Utah June-11 August. distribution: Kane Co.: Long \'alley; Millard Proctacanthus nearno Martin 1962: 187 Co.: Fillmore, Kanosh Cyn.; Washington [Holotype: Male, Baboquivari Mtns., Ari- 84 Great Basin Naturalist Vol. 47, No. 1

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January 1987 Nelson: Utah Robber Flies 85 zona, 19 July 1950, in KU], Utah distribution: Dam Mtns., Leeds, old airport. Paradise Cache Co.: Cornish; Carbon Co.: Price; Cyn., Rockville, St. George, Washington, Daggett Co.: Bridgeport; Duchesne Co.: Zion National Park; Wayne Co.: Teasdale. 8 Bluebell, Uintah Cyn.; Emery Co.: Wild- June-17 September. horse Cr. north of Goblin Valley, 4 mi N Promachus dimidiatus Curran 1927: 87 Gilson Butte, 2 mi E Little Gilson Butte; [Holotype: Male, Aweme, Manitoba, July Garfield Co.: Boulder, Ten Mile, The Halls; 1920, in CNC]. Utah distribution: Cache Co.: Grand Co.: Castleton, Moab; Juab Co.: Logan. 4-20 July. Callao; Kane Co.: Coral Pink Sand Dunes; Promachus sackeni Hine 1911: 166 [Syn- Millard Co.: Delta, Fillmore; San Juan Co.: types: Male and female, southern Arizona, Goulding; Sevier Co.: Richfield; Tooele Co.: July and August, no repository listed]. Utah

Skull Valley; Utah Co. : Provo, Spanish Fork; distribution: Washington Co.: Leeds Cyn., Washington Co.: Duncan Flats near Zion National Park. 4-20 July. Rockville, Leeds Cyn., Middleton, Rockville, Regasilus blantoni Bromley 1951: 35 [Holo- Santa Clara, Shunesburg, Zion National type: Male, Wills (Wells?), Nevada, 19 Au- Park—Coalpits Wash; Wayne Co.: Hanks- gust 1939, in Bromley Collection]. Utah dis- Pass; ville; Weber Co. : Ogden Cyn. 19 July-5 Sep- tribution: Millard Co.: Skull Rock tember. Washington Co. : Beaver Dam slope near Ari- Promachus albifacies Williston 1885: 63 zona border. 10 September-8 October. [Holotype: Sex not known, Arizona, in KU]. Triorhi interrupta (Macquart) 1834: 310 Utah distribution: Box Elder.: Clear Cr. [Holotype: Georgia, location unknown]. Utah Cyn.; Cache Co.: Logan; Daggett Co.: Pipe distribution: Cache Co.: Cornish, Logan;

Cr., Manila; Davis Co.: Wasatch Forest Garfield Co. : Bryce Cyn. ; Uintah Co. : White- boundary near Bountiful; Garfield Co.: rocks; Washington Co.: Rockville—Duncan Aquarius Plateau, Escalante, Panguitch; Mil- Flats, confluence East and North forks Virgin lard Co.: Kanosh Cyn.; Piute Co.: Junction; River, St. George. 10 July-11 August.

Tooele Co. : Lake Point; Uintah Co. : La Point; Utah Co.: American Fork, Lehi, Provo; Acknowledgments

Washington Co. : Ash Cr. bridge, Leeds Cyn.

Pinto, Zion National Park; Wayne Co.: Thanks are given to W. J. Hanson, G. F. Hanksville. 10 June-11 August. Knowlton, and G. E. Bohart of Utah State Promachus aldrichii Hine 1911: 171 [Syn- University for help given during the prepara- types: Male and female, Utah, repository not tion of the thesis from which this publication is listed, probably in Hine Collection or OSU]. derived. Additional thanks are given to the Utah distribution: Beaver Co.: Beaver, Mil- following curators and institutions which al- ford, Minersville Reservoir; Box Elder Co.: lowed use of their specimens: S. L. Wood and Collinston; Cache Co.: Logan, Petersboro; R. W. Baumann, BYU; M. Fors, UU; A. Bar- Daggett Co.: Bridgeport, Manila; Duchesne num, Dixie College; R. D. Anderson, SUSC;

Co.: 10 mi N Neola; Emery Co.: Paige Flat, andL. King, CEU. I thank J. K. Nelson and J. Sinbad Country; Grand Co.: Castle Valley, A. Stanger for help with the illustrations, the to use the Moab; Iron Co.: Cedar City; Juab Co.: Eu- J. Wilcox family for permission reka, Nephi, Topaz Mtn., Trout Cr.; Kane drawings of Efferia terminalia, and A. Co.: Escalante Desert, Willow Tank; Millard Provonsha for the map. Many thanks are ex- Co.: Confusion Pass, Desert Research Sta- tended to the D Elden Beck family for provid- tion, Pahvant; Salt Lake Co.: Hunter; San ing funds to publish this work.

Juan Co. : Piute Pass near Grand Gulch; San- pete Co. : 8 mi NE Fountain Green, Fairview; List OF Museum Abbreviations Tooele Co.: Cedar Mtns., Delle, Dugway Proving Grounds, Flux, Hickman Cyn., In- AESP American Entomological Society, Philadelphia dian Springs, Mercur, Skull Valley, Whiskey AMNH American Museum of Natural History Young University Springs; Uintah Co.: Bonanza, Dinosaur Na- BYU Brigham CAS California Academy of Sciences tional Monument, Jensen, Vernal, White- CEU College of Eastern Utah rocks; Utah Co.: Lehi, Orem, Rock Cyn., CIS California Survey west Utah Lake; Washington Co.: Beaver CNC Canadian National Collection 86 Great Basin Naturalist Vol. 47, No. 1

# Stenopogon engelhardti

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Figs. 12.5-128. Utah Asilidae, distribution: 12.5, Scleropo^on nefilectiis, Sclcropo^on picticonm , and Scleropogon bradleyi, 126, Stenopogon engelhardti- 127. Stenopogon inqiiinatus 128, Stenopogon martini. January 1987 Nelson: Utah Robber Flies 87

CSU Colorado State University 1921. An annotated list of the diptera (flies) of KU UniversitN' of Kansas Oregon. Proc. California Acad. Sci., Ser. 4, 11: MCZ Museum of Comparative Zoology 197-344. OSU Ohio State University Cole, F. R., and A. E. Pkitchard 1964. The genus Mal- SUSC Southern Utah State College lophora and related asilid genera in North Amer- UM L^niversity of Massachusetts ica (Diptera: Asilidae). Univ. California Publ. En- USNM National Museum tomol. 36:43-100.

USU Utah State University Cole, F. R, and J. Wilco.x. 1938. The genera Lasiopogon UU University of Utah Loew and Alexiopogon Curran in North America (Diptera: Asilidae). Ent. Americana 18; 1-90. CoQUlLLETT, D. 1893. A new asilid genus related to Literature Cited W Erax. Canadian Ent. 25; 175-177. 1904. New North American Diptera. Proc. Ent. Nearctic Adisoemarto. S . AND D M Wood 1975. The Soc. Washington 6: 166-192. species oWioctria and six related genera (Diptera: Curran, C H 1922. New Diptera in the Canadian Na- Asihdae). Quaest. Ent. 11: 505-576. tional Collection. Canadian Ent. 54: 277-287. Back, E. A. 1904. New species of North American Asili- 1923a. Studies in Canadian Diptera. I. Revision of dae. Canadian Ent. 36: 289-293. the asilid genus Cijrtopogon and allied genera. II. 1909. Therobberfliesof America north of Mexico, The genera of the family Blepharoceridae. Cana- belonging to the subfamilies Leptogastrinae and dian Ent. 55; 92-95, 116-125, 132-142, 169-174, Dasypogoninae. Trans. Amer. Ent. Soc. 35; 185-190, 266-269. 137-400. 1923b. Apparently undescribed Canadian Asilidae Banks. N 1920. Descriptions ofa few new Diptera. Cana- and Dolichopodidae (Diptera). Canadian Ent. 55: Ent. 52: 65-67. dian 207-211. Bellardi, L. 1861. Saggio di ditterologia messicana. 1926. A new species of Comantclla (Asilidae, Parte II. Torino. Diptera). Canadian Ent. 58; 310-312. Bigot, M. F. 1878. Dipteres nouveaux ou pen connus. J. 1927. Descriptions of Nearctic Diptera. Canadian Tribu des Asilidi. Ann. Soc. Ent. France Ser. 5, 8: Ent. 59: 79-92. 31-48, 40-48, 213-240, 401-446. 1930. New Diptera from North and Central Amer- Bromley, S. W 1928. Notes on the genus Proctacanthus ica. Amer. Mus. Novitates 415; 1-16. with descriptions of two new species (Diptera: 1934. The families and genera of North American Asilidae). Psyche 35: 12-15. Diptera. New York. 512 pp. 1931. New Asilidae, with a revised key to the belonging to HiNE. J S 1906. Two new species of Diptera genus Stenopogon Loew: (Diptera). Ann. Ent. Asilinae. Ohio Nat. 7; 29-30. Soc. Amer. 24: 427-435. 1908. Two new species of Asilidae from British 19.34. The robber flies of Texas (Diptera, Asilidae) Columbia. Canadian Ent. 40; 202-204. Ann. Ent. Soc. Amer. 27; 74-111. 1909. Robber flies of the genus A.St/us . Ann. Ent. 1936. The genus Diogmites in the United States of Soc. Amer. 2; 136-172 (= contributions from the with descriptions of new species America Ohio State Univ. Dept. Zool. and Ent. No. 32). (Diptera; Asilidae). New York Ent. Soc. 44; J. 1911. Robber flies of the genera Promachus and 225-237. Proctacanthus . Ann. Ent. Soc. Amer. 4: 153-172 1937a. The genus Stenopogon Loew in the United (= contributions from the Ohio State Univ. Dept. Diptera). York States of America (Asilidae; J. New of Zool. and Ent. No. 33). Ent. Soc. 45; 291-309. 1916. Descriptions of robber flies of the genus 1937b. New and little-known Utah Diptera with Erax. Ohio Jour. Sci. 17: 21-22. notes on the of the Diptera. Proc. Utah 1918. Descriptions of seven species ofA.si/H.s (fam-

Acad. Sci. 14:99-109. ily Asilidae). Ohio Jour. Sci. 18; 319-322. 1940. New U.S.A. robber flies (Diptera; Asilidae). 1919. Robber flies of the genus Erax. Ann. Ent. Bull. Brooklyn Ent. Soc. 35; 13-21. Soc. Amer. 12; 103-154 (= contributions from the 1951. Asilid notes (Diptera) with descriptions of 32 Ohio State Univ. Dept. of Zool. and Ent. No. 57). new species, Amer. Mus. Novitates 1532: 1-36. Hull, F M. 1957. Some new species of robber flies Brown, B. 1897. Two new species of asilids from New (Diptera: Asilidae). Psyche 64; 70-75. Mexico. Kansas Univ. Quart. Ser. A, 6: 103. James, M. T. 1937a. New Colorado Asilidae (Diptera). systematical Brown. C. J D. 1929. A morphological and Ent. News 48; 12-15. study of Utah Asilidae (Diptera). Trans. Amer. 1937b. The genus Comantella Curran (Diptera; Ent.' Soc. 54; 295-320. Asilidae). Pan-Pac. Ent. 13: 61-63. Cole, F R 1923. Expedition of the California Academy of 1941. The robber flies of Colorado (Diptera; Asili- in 1921. 14; 27-53. Sciences to the The dae). J. Kansas Ent. Soc. (bee flies); Diptera from the islands 1942. Additions to the "Robber Flies of Colorado."

II. 15; 124-126. and adjacent shores of the Gulf of California. J. Kansas Ent. Soc. General report. Proc. California Acad. Sci., Ser. Johnson, C W 1918. Notes on the species of the genus 4, 12: 289-314. Dioctria. Psyche 25; 102-103. Cole, F. R., and A. L. Lovett. 1919. New Oregon Johnson. D E 19.36. A further study of Utah Asilidae. Diptera. Proc. California Acad. Sci., Ser. 4, 9; Unpublished thesis, Brigham Young University, 221-255. Provo, Utah. Great Basin Naturalist Vol. 47, No. 1

•i .

January 1987 Nelson: Utah Robber Flies 89

1942. A new Cijrtopogon (Asilidae, Diptera) from Pritcmard, a E 1935. New Asilidae from the southwest- Utah. Great Basin Nat. 3: 1-4. ern U.S. (Diptera). Amer. Mus. Novitates 813: 1958. A new species oiMallophora from the Great 1-13. Salt Lake Desert (Diptera: AsiHdae). Great Basin 1938. The genus Hodoplnjlax James, with a de-

Nat. 18: 41-42. scription oi hasingeri , new species (Diptera: Asili- Knowlton, G. F., and F C Harmston 1938. Utah Asih- dae). Pan-Pac. Ent. 14: 129-131. dae. Proc. Utah Acad. Sci. 15: 123-125. 1943. Revision of the genus Cophura Osten Knowlton, G. F.. F. C. Harmston, and G. S. Stains. Sacken (Diptera: Asilidae). Ann. Ent. Soc. Amer. 1939. Utah Diptera. Mimeo Ser. 200 (Tech.) 5: 36: 281-309. 4-5. Say.T 1823. Descriptions of dipterous of the U.S. Linnaeus, G. 1758. Systema naturae per regna tria natu- 3: 73-104. J. Acad. Nat. Sci. Phila. 9-54, rae. Ed. 10. Stockhohn. 824 pp. 1824. Appendix, Part 1. Natural History. 1. Zool- Academicae, sen disserta- 1763. Amoenitates ogy. E. Class Insecta. Pages 268-278 in W. H. physicae, medicae, botanicae, ante- tiones variae Keating, Major Long's second expedition [q.v.]. hac seorsim editae, nunc collectae et auctae cum Vol. 2. Philadelphia. 459 pp. tabuUs aeneis. Vol. 6. Stockhohn. 486 pp. SCHAEFFER, C. 1916. New Diptera of the family Asilidae LOEW, H. 1866. Diptera Americae septentrionahs indi- with notes on known species. J. New York Ent. gena. Genturia septima. BerHner Ent. Zeitschr. Soc. 24: 65-69. 10: 1-54. SCHINER, I. R. 1867. Neue oder weniger bekannte Asili- 1874. Neue nordamerikanshe Dasypogonina. Ber- den der K. zoologischen Hofcabinetes in Wien. hner Ent. Zeitschr. 18: 353-377. K.;K. Zool.-Bot. Gesell. Wien, Verhandl. 17 (Ab- MacQUART, 1834. Histoire naturelle des insectes. J. handl): 355-412. Diptera. Vol. 1. Paris. 578 pp. (Gollection des Stone, A., C. W. Sabrosky, W W Wirth, R H Foote, suites a Buffon) [q.v.]. and R. CouLSON 1965. A catalog of the Diptera 1838. Diptereexotiques nouveau.xou peuconnus. J, ofAmerica north of Mexico. U.S. Dept. Agr., Agr. Soc. Rov. des Sci., de I'Agr. et des Arts, Lille, Res. Serv., Washington, D.C. 1696 pp. (Asilidae, Mem. 1838 (2): 9-225. pp. 360-401, 1113-1116). Martin, G. H. 1957. A revision of the Leptogastrinae in desultory collecting of the United States (Diptera: Asilidae). Bull. Amer. Tucker, E. S. 1907. Some results of Colorado. Kansas Univ. Sci. Mus. Nat. Hist. 111:347-385. insects in Kansas and = ser., 14): 51-112 = Kansas Univ. 1959. The Holopogon complex of North America, Bull. 4 ( whole [ excluding Mexico, with the descriptions of a genus Bull. 7(5)]. and a new subgenus (Diptera: Asilidae). Amer. Walker, F 1849. List of the specimens of dipterous Mus. Novitates 1980: 1-40. insects in the collection of the British Museum. 1962. The mistaken identity of Proctacanthus Four parts and three supplements. London. Pt. 2: arno Townsend and a new species (Diptera: Asili- 231-484. 35: 185-188. saundersiana; or characters of unde- dae). J. Kansas Ent. Soc. 1851. Insecta 1968. A revision of Ospriocerus (Diptera: Asili- scribed insects in the collection of William Wilson dae). Proc. Galifornia Acad. Sci. 35: 401-424. Saunders, Esq. in 5 parts. 474 pp.

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