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The Mesosomal Anatomy of Myrmecia Nigrocincta Workers and Evolutionary Transformations in Formicidae (Hymeno- Ptera)
7719 (1): – 1 2019 © Senckenberg Gesellschaft für Naturforschung, 2019. The mesosomal anatomy of Myrmecia nigrocincta workers and evolutionary transformations in Formicidae (Hymeno- ptera) Si-Pei Liu, Adrian Richter, Alexander Stoessel & Rolf Georg Beutel* Institut für Zoologie und Evolutionsforschung, Friedrich-Schiller-Universität Jena, 07743 Jena, Germany; Si-Pei Liu [[email protected]]; Adrian Richter [[email protected]]; Alexander Stößel [[email protected]]; Rolf Georg Beutel [[email protected]] — * Corresponding author Accepted on December 07, 2018. Published online at www.senckenberg.de/arthropod-systematics on May 17, 2019. Published in print on June 03, 2019. Editors in charge: Andy Sombke & Klaus-Dieter Klass. Abstract. The mesosomal skeletomuscular system of workers of Myrmecia nigrocincta was examined. A broad spectrum of methods was used, including micro-computed tomography combined with computer-based 3D reconstruction. An optimized combination of advanced techniques not only accelerates the acquisition of high quality anatomical data, but also facilitates a very detailed documentation and vi- sualization. This includes fne surface details, complex confgurations of sclerites, and also internal soft parts, for instance muscles with their precise insertion sites. Myrmeciinae have arguably retained a number of plesiomorphic mesosomal features, even though recent mo- lecular phylogenies do not place them close to the root of ants. Our mapping analyses based on previous morphological studies and recent phylogenies revealed few mesosomal apomorphies linking formicid subgroups. Only fve apomorphies were retrieved for the family, and interestingly three of them are missing in Myrmeciinae. Nevertheless, it is apparent that profound mesosomal transformations took place in the early evolution of ants, especially in the fightless workers. -
Phylogeny of the North American Vaejovid Scorpion Subfamily Syntropinae Kraepelin, 1905, Based on Morphology, Mitochondrial and Nuclear DNA
Cladistics Cladistics 31 (2015) 341–405 10.1111/cla.12091 Phylogeny of the North American vaejovid scorpion subfamily Syntropinae Kraepelin, 1905, based on morphology, mitochondrial and nuclear DNA Edmundo Gonzalez-Santill an a,b,*,†,‡ and Lorenzo Prendinib aThe Graduate Center, City University of New York, CUNY, 365 Fifth Avenue, New York, NY, 10016, USA; bScorpion Systematics Research Group, Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79th Street, New York, NY, 10024-5192, USA; †Present address: Laboratorio Nacional de Genomica para la Biodiversidad, Centro de Investigacion y de Estudios Avanzados del Instituto Politecnico Nacional, Km 9.6 Libramiento Norte Carretera Leon, C.P. 36821, Irapuato, Guanajuato, Mexico; ‡Present address: Laboratorio de Aracnologıa, Departamento de Biologıa Comparada, Facultad de Ciencias, Universidad Nacional Autonoma de Mexico, Coyoacan, C.P. 04510, Mexico D.F., Mexico Accepted 25 June 2014 Abstract The first rigorous analysis of the phylogeny of the North American vaejovid scorpion subfamily Syntropinae is presented. The analysis is based on 250 morphological characters and 4221 aligned DNA nucleotides from three mitochondrial and two nuclear gene markers, for 145 terminal taxa, representing 47 species in 11 ingroup genera, and 15 species in eight outgroup genera. The monophyly and composition of Syntropinae and its component genera, as proposed by Soleglad and Fet, are tested. The follow- ing taxa are demonstrated to be para- or polyphyletic: Smeringurinae; Syntropinae; Vaejovinae; Stahnkeini; Syntropini; Syntrop- ina; Thorelliina; Hoffmannius; Kochius; and Thorellius. The spinose (hooked or toothed) margin of the distal barb of the sclerotized hemi-mating plug is demonstrated to be a unique, unambiguous synapomorphy for Syntropinae, uniting taxa previ- ously assigned to different subfamilies. -
California (Scorpiones: Vaejovidae)
do PAN-PACIFIC ENTOMOLOGIST 62(4), 1986, pp. 359-362 A New Species of Uroctonus from the Sierra Nevada of California (Scorpiones: Vaejovidae) STANLEY C. WILLIAMS San Francisco State University, San Francisco, California 94132. Abstract. —A new species of Uroctonus is described and named Uroctonus franckei Williams. This species has only been found at elevations of over 2133 meters in the Sierra Nevada of California. The closest relative of this new species appears to be Uroctonus mordax Thorell. During 1980, a series of collecting trips was conducted along the eastern slope of the Sierra Nevada of California. Sampling at higher elevations (i.e., over 2000 meters) indicated an abundant and diverse scorpion community. Of particular interest was a large, dark, previously undescribed species which was only found at elevations above 2133 meters on slopes dominated by yellow pine (Pinus jeffreyi Grer. & Balf.). This new species is here described and named. Measurements cited are as defined by Williams (1980). I am indebted to Paul H. Arnaud, Jr. for furnishing research facilities at the California Academy of Sciences which aided this study. Much appreciation is due Vincent F. Lee, David Herlocker, and Jack T. Tomlinson who critically read this manuscript. Thanks also to Jett S. Chinn for help with illustrations. Uroctonus franckei Williams, NEW SPECIES (Fig. 1, Table 1) Diagnosis.—Total length up to 57 mm; base color of body dark reddish-brown, often appearing blackish; frontal margin of carapace bibbed, median ocelli small, ratio of carapace width to diameter of diad 6.2-6.8; pedipalps with palm swollen prolaterally in oblique plane, ratio of chela length to palm width 3.3-3.4; fixed finger of chela with trichobothrium id at finger origin, supernumerary denticles 7 on fixed finger, 8 on movable finger, primary row denticles divided into 6 subrows on fixed finger, 7 subrows on movable finger; brachium with three ventral trich- obothria; soles of telotarsi with single row of spiniform setae ventrally; pectine teeth 13-14 in males, 9-12 in females. -
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A new spider wasp from Western Australia, with a description of the first known male of the genus Eremocllrglls (Hymenoptera: Pompilidae) 1 2 1 L. Krogmann • , M.C. Day' and A.D. Austin I f\ustralian Centre for Evolutionary Biology and Biodiversity, The University of Adelaide, South Australia 5005, Australi,l. 'State Museum of Natural History Stuttgart, Rosenstein I, Stuttgart. D-70191 Germany (present address). Email: [email protected] 'National Museum Cardiff, Cathays Park, Cardiff, C1'I0 3NI', Wales, United Kingdom. Abstract - En'lllocllrglls lil/l/ilCi sI'. novo is described from Western Australia. The female of this new species is brachypterous, a unique feature within Ercl/lOClIrglls Haupt and rare within the Australian pompilid fauna. The fullv winged male is the first recorded for the genus. The diversity of ErCI/IOCllrgll" its distribution and brachyptery among the Pompilidae are discussed. INTRODUCTION female and the first male of the genus. At the same The Australian pompilid fauna is particularly time, we present an overview of the diversity and diverse (Austin et al. 2004) and displays a distribution of the genus, and discuss the occurrence high level of endemism. However, although of brachyptery within the Australian Pompilidae. the first Pompilidae for the continent were described by Fabricius in 1775, the group is TERMINOLOGY AND METHODS generally poorly known for Australia, and Terms for morphological structures follow Day it is likely that significantly less than half (1988) and Coulet and Huber (1993). Specimens the fauna has been described. Further, the were borrowed from and/or are deposited in the group is taxonomically difficult because of the following collections (acronyms used throughout morphological conservatism among numerous the text): Australian Museum, Sydney, Australia genera, in addition to the often extreme sexual (AM); Australian National Insect Collection, dimorphism and complex mimicry associations CSIRO, Canberra, Australia (ANIC); California seen in many species (e.g. -
THE TRUE ARMY ANTS of the INDO-AUSTRALIAN AREA (Hymenoptera: Formicidae: Dorylinae)
Pacific Insects 6 (3) : 427483 November 10, 1964 THE TRUE ARMY ANTS OF THE INDO-AUSTRALIAN AREA (Hymenoptera: Formicidae: Dorylinae) By Edward O. Wilson BIOLOGICAL LABORATORIES, HARVARD UNIVERSITY, CAMBRIDGE, MASS., U. S. A. Abstract: All of the known Indo-Australian species of Dorylinae, 4 in Dorylus and 34 in Aenictus, are included in this revision. Eight of the Aenictus species are described as new: artipus, chapmani, doryloides, exilis, huonicus, nganduensis, philiporum and schneirlai. Phylo genetic and numerical analyses resulted in the discarding of two extant subgenera of Aenictus (Typhlatta and Paraenictus) and the loose clustering of the species into 5 informal " groups" within the unified genus Aenictus. A consistency test for phylogenetic characters is discussed. The African and Indo-Australian doryline species are compared, and available information in the biology of the Indo-Australian species is summarized. The " true " army ants are defined here as equivalent to the subfamily Dorylinae. Not included are species of Ponerinae which have developed legionary behavior independently (see Wilson, E. O., 1958, Evolution 12: 24-31) or the subfamily Leptanillinae, which is very distinct and may be independent in origin. The Dorylinae are not as well developed in the Indo-Australian area as in Africa and the New World tropics. Dorylus itself, which includes the famous driver ants, is centered in Africa and sends only four species into tropical Asia. Of these, the most widespread reaches only to Java and the Celebes. Aenictus, on the other hand, is at least as strongly developed in tropical Asia and New Guinea as it is in Africa, with 34 species being known from the former regions and only about 15 from Africa. -
Scorpiones, Euscorpiidae) from Turkey 63 Doi: 10.3897/Zookeys.219.3597 Research Article Launched to Accelerate Biodiversity Research
A peer-reviewed open-access journal ZooKeys 219:A 63–80 new (2012) species of Euscorpius Thorell, 1876( Scorpiones, Euscorpiidae) from Turkey 63 doi: 10.3897/zookeys.219.3597 RESEARCH artICLE www.zookeys.org Launched to accelerate biodiversity research A new species of Euscorpius Thorell, 1876 (Scorpiones, Euscorpiidae) from Turkey Gioele Tropea1,†, Ersen Aydın Yağmur2,‡, Halil Koç3,§, Fatih Yeşilyurt4,|, Andrea Rossi5,¶ 1 Società Romana di Scienze Naturali, Rome, Italy 2 Alaşehir Vocational School, Celal Bayar University, Manisa, Turkey 3 Sinop University, Science and Art Faculty, Biology Department, Sinop, Turkey 4 Kırıkkale University, Science and Art Faculty, Biology Department, Zoology Section, Kırıkkale, Turkey 5 Aracnofilia, Centro Studi sugli Aracnidi, Massa, Italy † urn:lsid:zoobank.org:author:92001B12-00FF-4472-A60D-3B262CEF5E20 ‡ urn:lsid:zoobank.org:author:8DB0B243-5B2F-4428-B457-035A8274500C § urn:lsid:zoobank.org:author:77C76C8B-3F8F-4617-8A97-1E55C9F366F7 | urn:lsid:zoobank.org:author:FDF24845-E9F2-4742-A600-2FC817B750A7 ¶ urn:lsid:zoobank.org:author:D48ACE18-1E9B-4D68-8D59-DDC883F06E55 Corresponding author: Ersen Aydın Yağmur ([email protected]) Academic editor: W. Lourenço | Received 27 July 2012 | Accepted 15 August 2012 | Published 4 September 2012 urn:lsid:zoobank.org:pub:CE885AF1-B074-4839-AD1D-0FB9D1F476C3 Citation: Tropea G, Yağmur EA, Koç H, Yeşilyurt F, Rossi A (2012) A new species of Euscorpius Thorell, 1876 (Scorpiones, Euscorpiidae) from Turkey. ZooKeys 219: 63–80. doi: 10.3897/zookeys.219.3597 Abstract A new species of the genus Euscorpius Thorell, 1876 is described based on specimens collected from Dilek Peninsula (Davutlar, Aydın) in Turkey. It is characterized by an oligotrichous trichobothrial pat- tern (Pv= 7, et= 5/6, eb= 4) and small size. -
The Scorpion Fauna of Mona Island, Puerto Rico (Scorpiones: Buthidae, Scorpionidae)
The Scorpion Fauna of Mona Island, Puerto Rico (Scorpiones: Buthidae, Scorpionidae) Rolando Teruel, Mel J. Rivera & Alejandro J. Sánchez August 2017 – No. 250 Euscorpius Occasional Publications in Scorpiology EDITOR: Victor Fet, Marshall University, ‘[email protected]’ ASSOCIATE EDITOR: Michael E. Soleglad, ‘[email protected]’ Euscorpius is the first research publication completely devoted to scorpions (Arachnida: Scorpiones). Euscorpius takes advantage of the rapidly evolving medium of quick online publication, at the same time maintaining high research standards for the burgeoning field of scorpion science (scorpiology). Euscorpius is an expedient and viable medium for the publication of serious papers in scorpiology, including (but not limited to): systematics, evolution, ecology, biogeography, and general biology of scorpions. Review papers, descriptions of new taxa, faunistic surveys, lists of museum collections, and book reviews are welcome. Derivatio Nominis The name Euscorpius Thorell, 1876 refers to the most common genus of scorpions in the Mediterranean region and southern Europe (family Euscorpiidae). Euscorpius is located at: http://www.science.marshall.edu/fet/Euscorpius (Marshall University, Huntington, West Virginia 25755-2510, USA) ICZN COMPLIANCE OF ELECTRONIC PUBLICATIONS: Electronic (“e-only”) publications are fully compliant with ICZN (International Code of Zoological Nomenclature) (i.e. for the purposes of new names and new nomenclatural acts) when properly archived and registered. All Euscorpius issues starting from No. 156 (2013) are archived in two electronic archives: • Biotaxa, http://biotaxa.org/Euscorpius (ICZN-approved and ZooBank-enabled) • Marshall Digital Scholar, http://mds.marshall.edu/euscorpius/. (This website also archives all Euscorpius issues previously published on CD-ROMs.) Between 2000 and 2013, ICZN did not accept online texts as "published work" (Article 9.8). -
Taxonomical Updates for Buthidae
Taxonomical updates in The Scorpion Files for Chactidae (2008 →) Taxa Status Distribution Comments Reference Auyantepuia (Gonzalez- Unclear The genus Auyantepuia has been Sponga, 1978) synonymized with other genera. Lourenço & Qi (2007) have chosen not to accept this synonymization, and described the new species in Auyantepuia. The taxonomy of The Scorpion Files follows Soleglad & Fet (2005), but it is impossible for me to know where to put the new species. I have chosen to reinstate Auyantepuia in The Scorpion Files for this species until a new revision on the family Chactidae is published. Auyantepuia is not counted in the number of genera for the family, but the species is included. Auyantepuia aluku Ythier, New sp. French Guiana Ythier E. A synopsis of the 2018 scorpion fauna of French Guiana, with description of four new species. ZooKeys. 2018(764):27-90. Auyantepuia aurum Ythier, New sp. French Guiana Ythier E. A synopsis of the 2018 scorpion fauna of French Guiana, with description of four new species. ZooKeys. 2018(764):27-90. 2020 © Jan Ove Rein, The Scorpion Files Auyantepuia royi Ythier, New sp. Brazil Ythier E. A new species of 2018 Auyantepuia GonzálezSponga, 1978 (Scorpiones, Chactidae) from Brazil. Arachnida - Rivista Aracnologica Italiana. 2018;4(20):13-22. Auyantepuia surinamensis New sp. Surinam Lourenco WR, Duhem B. A new Lourenco, 2010 species of Auyantepuia Gonzalez-Sponga, 1978 (Scorpiones, Chactidae) from Suriname. Entomol Mitt Zool Mus Hamburg. 2010 Jun;15(182):137-45. Belisarius New placement Transferred back to Troglotayosicidae. See family website for more info. Broteochactas cauaburi New sp. Brazil Lourenco WR, Araujo J, Lourenco, Araujo & Franklin E. -
Arachnides 76
Arachnides, 2015, n°76 ARACHNIDES BULLETIN DE TERRARIOPHILIE ET DE RECHERCHES DE L’A.P.C.I. (Association Pour la Connaissance des Invertébrés) 76 2015 0 Arachnides, 2015, n°76 LES PREDATEURS DES SCORPIONS (ARACHNIDA : SCORPIONES) G. DUPRE Dans leur revue sur les prédateurs de scorpions, Polis, Sissom & Mac Cormick (1981) relèvent 150 espèces dont essentiellement des espèces adaptées au comportement nocturne de leur proie (chouettes, rongeurs, carnivores nocturnes) mais également des espèces diurnes (lézards, rongeurs, carnivores....) qui débusquent les scorpions sous les pierres ou dans leurs terriers. Dans une précédente note (Dupré, 2008) nous avions effectué un relevé afin d'actualiser cette étude de 1981. Sept ans après, de nouvelles données sont présentées dans cette synthèse. Voici un nouveau relevé des espèces prédatrices. Nous ne faisons pas mention des scorpions qui feront l'objet d'un futur article traité avec le cannibalisme. Explication des tableaux: La première colonne correspond aux prédateurs, la seconde aux régions concernées et la troisième aux références. Dans la mesure du possible, les noms scientifiques ont été rectifiés en fonction des synonymies ou des nouvelles combinaisons appliquées depuis les dates de publication d'origine. ARTHROPODA ARACHNIDA SOLIFUGAE Solifugae Afrique du Nord Millot & Vachon, 1949; Punzo, 1998; Cloudsley-Thompson, 1977 Eremobates sp. USA Bradley, 1983 ARACHNIDA ARANEAE Acanthoscurria atrox Brésil Lourenço, 1981 Aphonopelma sp. et autres Amérique centrale Mazzotti, 1964 Teraphosidae Phormictopus auratus Cuba Teruel & De Armas, 2012 Brachypelma vagans Mexique Dor et al., 2011 Epicadus heterogaster Brésil Lourenço et al. 2006 Latrodectus sp. USA Baerg, 1961 L. hesperus USA Polis et al., 1981 L. mactans Cuba Teruel, 1996; Teruel & De Armas, 2012 L. -
Scorpiones: Vaejovidae)
A New Species of Vaejovis from Chaparral Habitat Near Yarnell, Arizona (Scorpiones: Vaejovidae) Richard F. Ayrey July 2014 — No. 188 Euscorpius Occasional Publications in Scorpiology EDITOR: Victor Fet, Marshall University, ‘[email protected]’ ASSOCIATE EDITOR: Michael E. Soleglad, ‘[email protected]’ Euscorpius is the first research publication completely devoted to scorpions (Arachnida: Scorpiones). Euscorpius takes advantage of the rapidly evolving medium of quick online publication, at the same time maintaining high research standards for the burgeoning field of scorpion science (scorpiology). Euscorpius is an expedient and viable medium for the publication of serious papers in scorpiology, including (but not limited to): systematics, evolution, ecology, biogeography, and general biology of scorpions. Review papers, descriptions of new taxa, faunistic surveys, lists of museum collections, and book reviews are welcome. Derivatio Nominis The name Euscorpius Thorell, 1876 refers to the most common genus of scorpions in the Mediterranean region and southern Europe (family Euscorpiidae). Euscorpius is located at: http://www.science.marshall.edu/fet/Euscorpius (Marshall University, Huntington, West Virginia 25755-2510, USA) ICZN COMPLIANCE OF ELECTRONIC PUBLICATIONS: Electronic (“e-only”) publications are fully compliant with ICZN (International Code of Zoological Nomenclature) (i.e. for the purposes of new names and new nomenclatural acts) when properly archived and registered. All Euscorpius issues starting from No. 156 (2013) are archived in two electronic archives: Biotaxa, http://biotaxa.org/Euscorpius (ICZN-approved and ZooBank-enabled) Marshall Digital Scholar, http://mds.marshall.edu/euscorpius/. (This website also archives all Euscorpius issues previously published on CD-ROMs.) Between 2000 and 2013, ICZN did not accept online texts as "published work" (Article 9.8). -
Reanalysis of the Genus Scorpio Linnaeus 1758 in Sub-Saharan Africa and Description of One New Species from Cameroon
ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg Jahr/Year: 2011 Band/Volume: 15 Autor(en)/Author(s): Lourenco Wilson R. Artikel/Article: Reanalysis of the genus Scorpio Linnaeus 1758 in sub-Saharan Africa and description of one new species from Cameroon (Scorpiones, Scorpionidae) 99-113 ©Zoologisches Museum Hamburg, www.zobodat.at Entomol. Mitt. zool. Mus. Hamburg15(181): 99-113Hamburg, 15. November 2009 ISSN 0044-5223 Reanalysis of the genus Scorpio Linnaeus 1758 in sub-Saharan Africa and description of one new species from Cameroon (Scorpiones, Scorpionidae) W ilson R. Lourenço (with 32 figures) Abstract For almost a century, Scorpio maurus L., 1758 (Scorpiones, Scorpionidae) has been considered to be no more than a widespread and presumably highly polymorphic species. Past classifications by Birula and Vachon have restricted the status of different populations to subspecific level. In the present paper, and in the light of new evidence, several African populations are now raised to the rank of species. One of these, Scorpio occidentalis Werner, 1936, is redescribed and a neotype proposed to stabilise the taxonomy of the group. A new species is also described from the savannah areas of Cameroon. This is the second to be recorded from regions outside the Sahara desert zone. Keywords: Scorpiones, Scorpionidae, Scorpio, new rank, new species, Africa, Cameroon. Introduction The genus Scorpio was created by Linnaeus in 1758 (in part), and has Scorpio maurus Linnaeus, 1758 as its type species, defined by subsequent designation (Karsch 1879; see also Fet 2000). -
Geological History and Phylogeny of Chelicerata
Arthropod Structure & Development 39 (2010) 124–142 Contents lists available at ScienceDirect Arthropod Structure & Development journal homepage: www.elsevier.com/locate/asd Review Article Geological history and phylogeny of Chelicerata Jason A. Dunlop* Museum fu¨r Naturkunde, Leibniz Institute for Research on Evolution and Biodiversity at the Humboldt University Berlin, Invalidenstraße 43, D-10115 Berlin, Germany article info abstract Article history: Chelicerata probably appeared during the Cambrian period. Their precise origins remain unclear, but may Received 1 December 2009 lie among the so-called great appendage arthropods. By the late Cambrian there is evidence for both Accepted 13 January 2010 Pycnogonida and Euchelicerata. Relationships between the principal euchelicerate lineages are unre- solved, but Xiphosura, Eurypterida and Chasmataspidida (the last two extinct), are all known as body Keywords: fossils from the Ordovician. The fourth group, Arachnida, was found monophyletic in most recent studies. Arachnida Arachnids are known unequivocally from the Silurian (a putative Ordovician mite remains controversial), Fossil record and the balance of evidence favours a common, terrestrial ancestor. Recent work recognises four prin- Phylogeny Evolutionary tree cipal arachnid clades: Stethostomata, Haplocnemata, Acaromorpha and Pantetrapulmonata, of which the pantetrapulmonates (spiders and their relatives) are probably the most robust grouping. Stethostomata includes Scorpiones (Silurian–Recent) and Opiliones (Devonian–Recent), while