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Adec Preview Generated PDF File A new spider wasp from Western Australia, with a description of the first known male of the genus Eremocllrglls (Hymenoptera: Pompilidae) 1 2 1 L. Krogmann • , M.C. Day' and A.D. Austin I f\ustralian Centre for Evolutionary Biology and Biodiversity, The University of Adelaide, South Australia 5005, Australi,l. 'State Museum of Natural History Stuttgart, Rosenstein I, Stuttgart. D-70191 Germany (present address). Email: [email protected] 'National Museum Cardiff, Cathays Park, Cardiff, C1'I0 3NI', Wales, United Kingdom. Abstract - En'lllocllrglls lil/l/ilCi sI'. novo is described from Western Australia. The female of this new species is brachypterous, a unique feature within Ercl/lOClIrglls Haupt and rare within the Australian pompilid fauna. The fullv­ winged male is the first recorded for the genus. The diversity of ErCI/IOCllrgll" its distribution and brachyptery among the Pompilidae are discussed. INTRODUCTION female and the first male of the genus. At the same The Australian pompilid fauna is particularly time, we present an overview of the diversity and diverse (Austin et al. 2004) and displays a distribution of the genus, and discuss the occurrence high level of endemism. However, although of brachyptery within the Australian Pompilidae. the first Pompilidae for the continent were described by Fabricius in 1775, the group is TERMINOLOGY AND METHODS generally poorly known for Australia, and Terms for morphological structures follow Day it is likely that significantly less than half (1988) and Coulet and Huber (1993). Specimens the fauna has been described. Further, the were borrowed from and/or are deposited in the group is taxonomically difficult because of the following collections (acronyms used throughout morphological conservatism among numerous the text): Australian Museum, Sydney, Australia genera, in addition to the often extreme sexual (AM); Australian National Insect Collection, dimorphism and complex mimicry associations CSIRO, Canberra, Australia (ANIC); California seen in many species (e.g. Evans 1968, 1982; Harris Academy of Sciences, San Fransisco, U.S.A. 1987; Day 1988). Elliott (2007) lists 258 pompilid (CAS); The Natural History Museum, London, species for the Australian region of which the England (NHM); Museum Victoria, Melbourne, majority have been described from the eastern Australia (NMV); State Museum of Natural parts of the continent. Despite comprising one­ I-listory Stuttgart, Cermany (SMNS); Western third of the continent's landmass, Western Australian Museum, Perth, Australia (WAM); Australian Pompilidae have been little studied. Waite Insect and Nematode Collection, University Only 18'/" of all pompilid species (47 of 258 of Adelaide, Australia (WINC). species) are described from Western Australia, Images of the specimens were taken with a while for the subfamily Pepsinae only 12% (11 Leica lJXM 1200 digital camera attached to a Leica of 91 species) have type localities in that region MZ 16 APO microscope and processed using (data from Elliott 2007). Of the four recognised Auto-Montage (Syncroscopy) software. subfamilies (pitts et al. 2006; Elliott 2(07), the Pepsinae comprise the least known Australian genera, many of which are monotypic or contain SYSTEMATICS only few species. Severa I of these genera have Family Pompilidae LatreiIIe 1805 not been revised since their original description (Elliott 2(07), as is the case for Haupt ErclIlocurglls Haupt 1937 which is currently known from two species from Haupt 1937: 127,133. south-eastern Australia. Type species 11ere we describe a remarkable new species of the Prioclli'lIlis Smith 1868, bv original Australian endemic genus from VVestern designation. Australia, and thus record the first brachypterous 438 L. Krogrnann, M.C. Day, A.D. Austin Diagnosis Wedge Hill, 34°23'17"S, 118°1O'18"E, 2 May-17 Both sexes with fore-wings spotted or bifasciate, Dec 1996 (1 specimen SMNS, 5 specimens propodeum with relatively weakly defined WAM Entomology Reg. No. 68633-68637); 1 declivity. Female with tooth-like scales (=Haupt's 9, same data as holotype, but: Mt Magog, 'Schuppendornen') developed at bases of hind 34°23'59"S, 117°56'35"E, 25 April-3 September 1996 tibial spines. Male subgenital plate medially with (NHM); 1 9, same collection data as holotype, distinct posteriorly-directed spine. but: The Cascades, 34°22'29"S, 118°14'17"E, 4 September-17 December 1996 (WAM Entomology Remarks Reg. No. 69771); 19 9, same data as holotype, but: In his work on Australian Pepsinae, Haupt Toolbrunup Peak track, 34°23'32"S, 118°03'32"E, 23 (1937) described nine new genera (including April-3 September 1996, J.M. Waldock, BY.Main six genera with males unknown to him) in (WAM Entomology Reg. No. 69744-69762); 3 9, which he accommodated just 18 species. Two same data as holotype, but: Talyuberlup picnic of these species were placed by Haupt (1937) in site, 34°24'56"S, 117°57'18"E, 25 April-3 September Eremocllrglls: E. ephippiatlls (Smith 1868) and E. 1996, J.M. Waldock, BY. Main (1 specimen SMNS, pictipennis (Smith, 1855). In recent years it has 2 specimens WINC); 3 9, same data as holotype, become apparent that Eremocllrglls and other but: Porongurup National Park, south end of pepsine genera putatively related to it require Millinup Pass, 34°41'43"S, 117°53'51"E, 28 April-2 extensive reinterpretation to recognise more September 1996 (l specimen ANIC, 2 specimens natural groups, and this will likely involve WAM Entomology Reg. no. 68631-68632); 1 9, synonymy of some genera and an expansion of the Porongurup National Park, Yate Flats, malaise, 13 limits of Eremocllrglls. Such a study is currently November 1987, M.E. Irwin, E.!. Schlinger (CAS); being undertaken by the authors as part of a 1 9, Stirling Range National Park, Bluff Knoll at larger generic revision of Australian Pomplilidae, Cascade Track, malaise, 17-20 November 1987, and the description of the present species makes a M.E. Irwin, E.I. Schlinger (CAS); 2 0, 7 km N. of small contribution in this respect. The diagnosis Kent River Bridge, 34°54'S, 117°02'E, 19 April 1983, of Eremocllrglls sensu Haupt presented above has E.5. Nielsen, ED. Edwards (ANIC). been modified to include both the brachypterous Diagnosis female and first recorded male for E. linnaei. The female of E. linnaei differs from the Based on this species, Eremocllrglls exhibit a previously described species of Eremocurglls by striking level of sexual dimorphism, which led the following characters: head with red-golden to the males of E. limzaei remaining unrecognised pubescence (Figure le), fore- and hind-wing in the ANIC collection. Our postulated sex brachypterous (Figure la, c), propodeum elongate, association is supported by the close geographic with transverse striae, that are medially weakly occurrence of males and females. indicated (Figure If), toothed scales on hind tibia Eremocllrglls linnaei is only known from Western only weakly developed (Figure 19). The male is the Australia, while E. ephippiatlls has a broad first described for the genus and characterized by distribution covering Victoria, South Australia the following diagnostic features: head black, face and Western Australia (collection data), whereas with orange area extending laterodorsally from E. pictipennis is apparently restricted to Victoria. level of toruli (Figure 1h). Male with metapost­ notum well developed, reaching three quarters of Eremocurgus linnaei Krogmann, the median length of the metascutellum (Figure Day and Austin, sp. novo 1i). Figures la-j Description Material examined Holotype Female (Figure la, c, e-g) Australia: Western Australia: 9, Stirling Range Body length: 13.0-17.1 mm, head plus mesosoma: National Park, S. face of Pyungoorup Peak, 330m, 6.5-8.3 mm. Colour: Body and legs brownish­ 34°22'17"S, 118°19'20"E, 27 April-4 September 1996, purple. Antennae brown. Conspicuous red­ wet pitfall traps ex spirit collection, MS Harvey, golden pubescence present on head, tibiae, tarsi J.M. Waldock, BY. Main (WAM Entomology Reg. and dorsal surfaces of meso- and metasoma. No. 68761). Two patches of silvery pubescence present on anterolateral portions of metasomal tergites 2 and Pa ra types 3. Australia: Western Australia: 1 9, same Head (Figure le): Inner eye margin only very collection data as holotype (WAM Entomology slightly incised, eye 1.9-2.1 times as long as wide. Reg. No. 68760); 6 9, same data as holotype, but: Clypeus 3.1-3.3 times as long as wide and as A new species of Eremocurgus 439 a lmm e h Figure 1 ErellloCllrgus lillllaei sp. nov., paratypes - a, habitus female, lateral view; b, habitus male, lateral view; c, forewing female; d, forewing male; e, head female, frontal view; f, mesosoma female, dorsal view; g, hind tibia female, dorsolateral view; h, head male, frontal view; i, posterior mesosoma male, dorsal view; j, metasoma with subgenital plate male, lateral view. Abbreviations: elc=extended lateral corner; nol=pronotum; no3=metanotum; mpn=metapostnotum; ms=metascutellum; pd=propodeum; sc=mesoscutum; scl=mesoscutellum; SMC=submarginal cell. 440 L. Krogmann, M.C. Day, A.D. Austin wide as distance between bottoms of orbits. First Mesosoma: Pronotu m short, with long flagellomere 3.7-4.2 times as long as wide and pubescence. Mesonotum with fine punctuation, 1.1-1.2 times as long as second. Ocell%cular­ parapsidal sulcus present, reaching close to distance 1.7-1.8 times as wide as interocellular­ anterior mesoscutal margin, notaulus absent. distance, ocelli not enlarged. Metapost-notum well developed, posterior margin Mesosoma (Figure If): Pronotum transversely only slightly incised medially (Figure li). Median divided by distinct groove (=streptaulus), length of metapostnotum reaching three quarters anteriorly with extended lateral corners. of the median length of the metascutellum (Figure Mesonotum narrower than pronotum, paraspidal 1i). Propodeal surface without transverse striae, sulcus present, reaching half the length of but two strong dorsolateral carinae reaching mesoscutum.
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