A New Late Triassic Palynological Assemblage from Western Gondwana (Carrizal Formation, Marayes Basin, Argentina)

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A New Late Triassic Palynological Assemblage from Western Gondwana (Carrizal Formation, Marayes Basin, Argentina) comptes rendus palevol 2021 20 33 DIRECTEURS DE LA PUBLICATION / PUBLICATION DIRECTORS : Bruno David, Président du Muséum national d’Histoire naturelle Étienne Ghys, Secrétaire perpétuel de l’Académie des sciences RÉDACTEURS EN CHEF / EDITORS-IN-CHIEF : Michel Laurin (CNRS), Philippe Taquet (Académie des sciences) ASSISTANTE DE RÉDACTION / ASSISTANT EDITOR : Adenise Lopes (Académie des sciences ; [email protected]) MISE EN PAGE / PAGE LAYOUT : Fariza Sissi & Audrina Neveu (Muséum national d’Histoire naturelle; [email protected]) RÉVISIONS LINGUISTIQUES DES TEXTES ANGLAIS / ENGLISH LANGUAGE REVISIONS : Kevin Padian (University of California at Berkeley) RÉDACTEURS ASSOCIÉS / ASSOCIATE EDITORS (*, took charge of the editorial process of the article/a pris en charge le suivi éditorial de l’article) : Micropaléontologie/Micropalaeontology Maria Rose Petrizzo (Università di Milano, Milano) Paléobotanique/Palaeobotany Cyrille Prestianni (Royal Belgian Institute of Natural Sciences, Brussels) Evelyn Kustatscher* (Museum of Nature South Tyrol, Bozen/Bolzano) Métazoaires/Metazoa Annalisa Ferretti (Università di Modena e Reggio Emilia, Modena) Paléoichthyologie/Palaeoichthyology Philippe Janvier (Muséum national d’Histoire naturelle, Académie des sciences, Paris) Amniotes du Mésozoïque/Mesozoic amniotes Hans-Dieter Sues (Smithsonian National Museum of Natural History, Washington) Tortues/Turtles Juliana Sterli (CONICET, Museo Paleontológico Egidio Feruglio, Trelew) Lépidosauromorphes/Lepidosauromorphs Hussam Zaher (Universidade de São Paulo) Oiseaux/Birds Eric Buffetaut (CNRS, École Normale Supérieure, Paris) Paléomammalogie (mammifères de moyenne et grande taille)/Palaeomammalogy (large and mid-sized mammals) Lorenzo Rook (Università degli Studi di Firenze, Firenze) Paléomammalogie (petits mammifères sauf Euarchontoglires)/Palaeomammalogy (small mammals except for Euarchontoglires) Robert Asher (Cambridge University, Cambridge) Paléomammalogie (Euarchontoglires)/Palaeomammalogy (Euarchontoglires) K. Christopher Beard (University of Kansas, Lawrence) Paléoanthropologie/Palaeoanthropology Roberto Macchiarelli (Université de Poitiers, Poitiers) Archéologie préhistorique/Prehistoric archaeology Marcel Otte (Université de Liège, Liège) RÉFÉRÉS / REVIEWERS : https://sciencepress.mnhn.fr/fr/periodiques/comptes-rendus-palevol/referes-du-journal COUVERTURE / COVER : Made from the Figures of the article. Comptes Rendus Palevol est indexé dans / Comptes Rendus Palevol is indexed by: – Cambridge Scientific Abstracts – Current Contents® Physical – Chemical, and Earth Sciences® – ISI Alerting Services® – Geoabstracts, Geobase, Georef, Inspec, Pascal – Science Citation Index®, Science Citation Index Expanded® – Scopus®. Les articles ainsi que les nouveautés nomenclaturales publiés dans Comptes Rendus Palevol sont référencés par / Articles and nomenclatural novelties published in Comptes Rendus Palevol are registered on: – ZooBank® (http://zoobank.org) Comptes Rendus Palevol est une revue en flux continu publiée par les Publications scientifiques du Muséum, Paris et l’Académie des sciences, Paris Comptes Rendus Palevol is a fast track journal published by the Museum Science Press, Paris and the Académie des sciences, Paris Les Publications scientifiques du Muséum publient aussi / The Museum Science Press also publish: Adansonia, Geodiversitas, Zoosystema, Anthropozoologica, European Journal of Taxonomy, Naturae, Cryptogamie sous-sections Algologie, Bryologie, Mycologie. L’Académie des sciences publie aussi / The Académie des sciences also publishes: Comptes Rendus Mathématique, Comptes Rendus Physique, Comptes Rendus Mécanique, Comptes Rendus Chimie, Comptes Rendus Géoscience, Comptes Rendus Biologies. Diffusion – Publications scientifiques Muséum national d’Histoire naturelle CP 41 – 57 rue Cuvier F-75231 Paris cedex 05 (France) Tél. : 33 (0)1 40 79 48 05 / Fax : 33 (0)1 40 79 38 40 [email protected] / https://sciencepress.mnhn.fr Académie des sciences, Institut de France, 23 quai de Conti, 75006 Paris. © Publications scientifiques du Muséum national d’Histoire naturelle / © Académie des sciences, Paris, 2021 ISSN (imprimé / print) : 1631-0683/ ISSN (électronique / electronic) : 1777-571X A new Late Triassic palynological assemblage from western Gondwana (Carrizal Formation, Marayes Basin, Argentina) Silvia N. CÉSARI Museo Argentino de Ciencias Naturales “Bernardino Rivadavia” – CONICET, Avenida Ángel Gallardo 470, C1405DJR Ciudad Autónoma de Buenos Aires (Argentina) [email protected] (corresponding author) Juan M. DROVANDI Carina E. COLOMBI Gustavo A. CORREA Instituto y Museo de Ciencias Naturales, Universidad Nacional de San Juan, Avenida España 400 (Norte), 5400 San Juan (Argentina) and CIGEOBIO, CONICET (Argentina) [email protected] [email protected] [email protected] Luis A. SPALLETTI Centro de Investigaciones Geológicas, Universidad Nacional de La Plata, Diagonal 113 no 64, 1900 La Plata, Buenos Aires (Argentina) [email protected] Submitted on 7 December 2019 | Accepted on 3 June 2020 | Published on 18 August 2021 urn:lsid:zoobank.org:pub:4B7ED75F-5A4B-490F-AD8E-1495380095AA Césari S. N., Drovandi J. M., Colombi C. E., Correa G. A. & Spallettil A. 2021. — A new Late Triassic palynological as- semblage from western Gondwana (Carrizal Formation, Marayes Basin, Argentina). Comptes Rendus Palevol 20 (33): 677-700. https://doi.org/10.5852/cr-palevol2021v20a33 ABSTRACT The Carrizal Formation, exposed in the Marayes-El Carrizal Basin (western Argentina), has been the focus of palaeobotanical studies since the late 1800s. The recent finding of well-preserved palyno logical assemblages provides the first detailed studies about its palynofloras. In this paper, the 63 taxa identified in the unit are illustrated and discussed, as well as their stratigraphic distribution in equivalent palynological assemblages of Argentina. Some spore species are revised: Uvaesporites hammenii (Herbst) Césari, comb. nov., Retusotriletes wielandii (Jain) Césari, comb. nov. and Lund- KEY WORDS Triassic, bladispora stellae (Herbst) Césari, comb. nov. The recognition ofCadargasporites baculatus de Jersey & Carnian, Paten emend. Reiser & Williams, Craterisporites rotundus de Jersey, Enzonalasporites vigens Leschik, palynology, Leptolepidites argenteaeformis (Bolkhovitina) Morbey, Protodiploxypinus americus Dunay & Fisher and systematics, Gondwana, Rugulatisporites permixtus Playford, among others, appears to be useful for local and intercontinental Argentina. correlations. A Carnian age is proposed for the palynofloras. COMPTES RENDUS PALEVOL • 2021 • 20 (33) © Publications scientifiques du Muséum et/and Académie des sciences, Paris. www.cr-palevol.fr 677 Césari S. N. et al. RÉSUMÉ Nouvel assemblage palynologique du Trias supérieur de l’Ouest du Gondwana (formation Carrizal, bassin de Marayes, Argentine). La formation de Carrizal, exposée dans le bassin de Marayes-El Carrizal (Ouest de l’Argentine), fait l’objet d’études paléobotaniques depuis la fin duxix e siècle. La découverte récente d’assemblages palynologiques bien conservés fournit les premières études détaillées sur ces palynoflores. Dans cet article, les 63 taxons identifiés dans l’unité sont illustrés et commentés, ainsi que leur répartition stratigraphique dans des assemblages palynologiques équivalents de l’Argentine. Certaines espèces de spores sont révisées : Uvaesporites hammenii (Herbst) Césari, comb. nov., Retusotriletes wielandii MOTS CLÉS Trias, (Jain) Césari, comb. nov., et Lundbladispora stellae (Herbst) Césari, comb. nov. La reconnaissance de Carnien, Cadargasporites baculatus de Jersey & Paten emend. Reiser & Williams, Craterisporites rotundus de palynologie, Jersey, Enzonalasporites vigens Leschik, Leptolepidites argenteaeformis (Bolkhovitina) Morbey, Protodi- systématiques, Gondwana, ploxypinus americus Dunay & Fisher et Rugulatisporites permixtus Playford, entre autres, semble être utile Argentine. pour les corrélations locales et intercontinentales. Un âge Carnien est proposé pour les palynoflores. INTRODUCTION GEOLOGICAL SETTING The Argentinian Triassic paleofloras are characterized mainly The Marayes-El Carrizal Basin is part of a series of extensional by 17 orders and a group of incertae sedis genera, all included basins developed in the SW of Pangea during the early Meso- in four classes (Lycopsida; Sphenopsida; Filicopsida and zoic. This depocenter and the neighbourg Ischigualasto-Villa Gymnospermopsida) known as the “Dicroidium Flora”, Unión Basin were located at around 48°S drifting toward North comprising a total of about 200 and 36 species (e.g. Artabe during the Late Triassic (Goddéris et al. 2008; Kent et al. et al. 2007). The paleofloristic assemblage of the Carrizal 2014). In this paleogeographic context the Marayes-Carrizal Formation (San Juan Province) has been studied since the Basin would be placed in the austral temperate humid belt. late 1800s (Geinitz 1876), and more recently by Lutz & This basin is in the San Juan Province, at S and SW of Arce (2013) and Morel et al. (2015). The paleoflora is char- La Huerta Hill (Fig. 1) and is characterized by a 2300 m acterized mainly by Pleuromeiales, Marattiales, Equisetales, thick sedimentary succession (Bossi 1976). The stratigraphy Osmundales, Umkomansiales (Corystospermales), Peltasper- of this basin changed through time, firstly Borrello (1946) males, Ginkgoales, Coniferales, Cycadales and incertae sedis defined four units: Esquina Colorada, Quebrada de la Mina, genera. According to its
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