JOURNAL OF PALEONTOLOGY, VOL. 22 PLATE 66

Durham and Roberts, Cretaceous Asteroids CRETACEOUS ASTEROIDS FROM CALIFORNIA 437

In paratype no. 4860 most of the aboral ogy of the University of California at integumental covering has been removed Berkeley, Calif. Unfortunately part of the and the rays apparently were not flattened mold of the ambulacral area of the holotype during fossilization. In this specimen (pre- was destroyed during the making of the served so that the aboral surface is seen) the casts. Fortunately the photographs used in ambulacral plates are arched, presumably as illustrating it, and an excellent latex impres- in life, making an angljp of about 150° with sion of it were taken before it was damaged. one another, except adjacent to the mouth Astropecten pewei Miller and Unklesbay where it is about 120°. Most specimens have (1943) from the Jurassic Sundance forma- been flattened during preservation and in tion of Wyoming does not appear to be those that show least evidence of further closely related to the present ; it has distortion (as in the holotype, and paratype fewer and differently shaped marginal plates. no. 4859) it appears that the ambulacral, Order SPINULOSA adambulacral, and marginal plates were all Family slightly inclined towards the tip of the ray, Genus Gray usually at an angle of about 70° with the axis of the ray. Henricia GRAY (1840), Ann. Mag. Nat. Hist., vol. 6, p. 184; CLARK (1946), Carnegie Inst. Some specimens, for example paratype Washington, Pub. 566, p. 148. no. 4864, have the rays much more rounded. This may have been the outline of the ani- Genotype: Henricia oculata Gray =Asterias mal during life, or it may be that the flat- sanguinolenta Mtiller tened impressions like the holotype and HENRICIA (?) VENTURANA Durham and paratype no. 4859 represent more nearly Roberts n. sp. the true shape in life. If this latter is the Plate 66, figures 1,3 ' case, then specimens like paratype no. 4865 This species is represented by both oral must represent individuals which had died and aboral impressions of a single individual. some time previous to burial and had their Unfortunately the details are not consist- rays somewhat rolled up before interment. ently well preserved, but it is believed that At first it was thought that these individuals sufficient characters are preserved to war- represented another species, but upon care- rant giving it a specific name. The disc is ful examination it was seen that they had very small in comparison to the length of the same number of marginal plates, and the arms. The arms are more or less grace- these plates are the same shape as in in- fully curved, both the oral and aboral sur- dividuals like the holotype, so it was con- faces are represented. The margins of the cluded that they are distorted individuals arms are not distinct but it is believed that of the present species. this is due to a lack of conspicuous marginal Types.—In the California Institute of plates. However on one ray of the aboral Technology Paleontological Collections, mold, the integument has been removed and holotype no. 4858; paratypes nos. 4859- the aboral surface of the ambulacral plates 4865, 4867, 4868. In addition there are the is exposed and in this particular area the fragments and poorly preserved impressions ambulacral plates appear to terminate representing the remaining 25 specimens laterally against vertical plates which might which have not been assigned type numbers. represent marginal plates. Plaster casts of all the numbered specimens Rays 5, radius of rays about 40 mm., have been deposited with the U. S. National radius of disk about 5 mm. Rays slender, Museum and in the Museum of Paleontol- width at base about 6.8 mm., tapering grace-

EXPLANATION OF PLATE 66 FIGS. 1,3—Henricia venturana Durham and Roberts, n. sp. Holotype no. 4866.1, oral surface no. 4866a, X 1.7. 3, aboral surface no. 4866b, X 1.8. ' (p. 437) 2, 4, 5—Astropecten matilijaensis Durham and Roberts, n. sp. 2, Paratype no. 4868, X1.8, aboral surface. 4, Paratype no. 4867, Xll, madreporite (retouched), 5, Paratype no. 4864, X2.45, oral surface, note contracted state of arms. (p. 435) 438 _ J. WYATT DURHAM AND WA YNE A. ROBERTS

fully tb tip. Interbrachial arcs acute. Margin , 1935, A Pliocene Ophiuran from Trini- of rays uncertain, probably rounded, margi- dad: Jour. Paleontology, vol. 9, pp. 430-433. , 1937, An Ophiuran from the Byram Marl nal plates not conspicuous. The ambulacral (Oligocene) of Mississippi: Jour. Paleontol- plates occupy over one-half the width of oral ogy, vol. 11, pp. 235-240. surface of rays. It is suspected that this is , 1938, Ophiurans from the Upper Senonian due to flattening during fossilization. Aboral of South Limburg, Holland: Jour. Paleontol- ogy; vol. 12, pp. 61-71, pis. 14-16. surface covered by a coarse granular net- , 1939, More Complete Remains of Ophiura work, possibly a few traces of short spines marylandica: Proc. Am. Phil. Soc., vol. 80, near the margins of the rays. Apparently no. 1, pp. 87-94, 1 pi., 1 fig. about 35 ambulacral plates to a ray. Tube , 1941a, Cretaceous Ophiurans from Texas: Jour. Paleontology, vol. 15, pp. 61-67, feet pores large, about 0.8 mm. in longitudi- pis. 9-11. nal diameter, adambulacral plates appar- , 1941b, Tertiary Ophiurans from Vene- ently corresponding to ambulacrals, not zuela: Jour. Paleontology, vol. 15, pp. 68-70, spinose, apparently about as long as wide. pi. 11. , 1942, A New Ophiuran from the Eocene Madreporite dorsal, on margin of disk, of New Jersey: Jour. Paleontology, vol. 16, about 1.5 mm. in diameter, surface marked pp. 393-396, pi. 60. by numerous vermiform radiating ridges. CLARK, B. L., 1915, Fauna of the San Pablo Holotype, California Institute of Tech- Group of Middle California: Univ. Calif. Publ., Bull. Dept. Geol. Sci., vol. 8, pp. 385- nology Paleontological Collections, nos. 572, pis. 42-71. 4866A (oral surface), 4866B (aboral sur- , 1918, The San Lorenza Series of Middle face). Plaster casts have been deposited California: Univ. Calif. Publ., Bull. Dept. "with the U. S. National Museum, and in the Geol. Sci., vol. 11, pp. 45-234, pis. 3-24. CLARK, H. L., 1913, Asterozoa in Eastman Museum of Paleontology of the University Zittel: pp. 244-257, figs. 347-364 (Macmillan of California at Berkeley, Calif. and Co., Limited, London). The reference of this species to the genus , 1946, The Fauna of Austra- lia: Carnegie Inst. Washington, publ. 566, pp. Henricia is very uncertain because of the 1-567. apparent wide ambulacral grooves and lack CLARK, W. B., 1893, The Mesozoic Echinoder- of adambulacral spines, but it is possible mata of the United States: U. S. Geol. Surv., that this is an apparent condition produced Bull. 97, pp. 1-207, pis. 1-50. by flattening during fossilization. If, as sug- , 1904, Maryland Geological Survey, Mio- cene Report: pp. 430-433. gested on the arm showing the aboral sur- , and TWITCHELL, M. W., 1915, The face of the ambulacral plates, there are Mesozoic and Cenozoic Echinodermata of the conspicuous marginals bounding the ray, United States, U. S. Geol. Surv. Mon. 54, pp. the species might better be assigned to 1-134, pis. 1-108. COOKE, C. W., GARDNER, J., and WOODRING, Linckia which may also have granular W. P., 1943, Correlation of the Cenozoic For- adambulacral plates. mations of the Atlantic and Gulf Coastal Plain and the Caribbean Region: Bull. Geol. Soc. BIBLIOGRAPHY Am., vol. 5*4, pp. 1713-1733. ADKINS, W. S., 1920, The Weno and Pawpaw for- DOUGLAS, E., 1903, Astropecten ? mantanus, a mations of the Texas Comanchean: Univ. new from the Fort Benton, and some Texas Bull. no. 1856, pp. 1-174, pis. 1-11. geological notes, Carnegie Mus., Ann., vol. 2, , 1928, Handbook of Texas Cretaceous fos- pp. 5-8. sils: Univ. Texas Bull. no. 2838, pp. 1-385, pis. DURHAM, J. W., 1944, Megafaunal zones of the 1-37. Oligocene of northwestern Washington, Univ. , and WINTON, W. M., 1920, Paleontological Calif. Publ., Bull. Dept. Geol. Sci., vol. 27, pp. Correlation of the Fredericksburg and Washita 101-212, pis. 13-18. formations in North Texas, Univ. Texas Bull, GABB, W. M., 1869, Paleontology of California, no. 1945, pp. 1-128, pis. 1-21. vol. 2, pp. 1-299, pis. 1-36. ALEXANDER, C. I., 1931, A new Lower Creta- , 1876, Note on the discovery of representa- ceous ophiuroid: Jour. Paleontology, vol. 5, tives of three orders new to the Cretaceous for- pp. 152-153, pi. 20, figs. 19-20. mation of North America, Proc. Acad. Nat. ARNOLD, R., 1908, Description of a new brittle Sci. Philadelphia, vol. 28, pp. 178-179. star from the upper Miocene of the Santa Cruz HOWE, H. V., 1942, Neglected Gulf Cbast Ter- Mountains, Calif., U. S. Nat. Mus., Proc., tiary microfossils, Bull. Am. Assoc. Petr. Geol., vol. 34, pp. 403-406, pi. 40. vol. 26, pp. 1188-1199, figs. 1-25. BERRY, C. T., 1934, Miocene and Recent KERR, P. F., and SCHENCK, H. G., 1928, Signifi- Ophiura skeletons: Johns Hopkins Univ. cance of the Matilija Overturn, Bull. Geol. Studies Geol., no. 11, pp. 9-136. Soc. America, vol. 39, pp. 1087-1102. CRETACEOUS ASTEROIDS FROM CALIFORNIA 439

LOGAN, W. N., 1900, The Stratigraphy and in- Part Third, pp. 67-90, pis. 17-26 (Paleonto- vertebrate faunas of the Jurassic formation in graphical Society, London). the Freeze-out Hills of Wyoming, Kansas , 1907, A Monograph on the British fossil Univ. Quart., ser. A, vol. 9, pp. 109-134, pis. Echinodermata from the Cretaceous forma- 25-31. tions. Volume Second. The Asteroidea and MEEK, F. B., 1873, United States geological and Ophiuroidea, Part Fourth, pp. 91-132, pis. 27- geographical survey of territories, Pal. Rept. 29 (Paleontographical Society, London). for 1872, pt. 2, pp. 429-518. VANDERPOOL, H. C., 1933, Upper Trinity micro- MERRIAM, C. W., 1931, Notes on a Brittle-star fossils from southern Oklahoma, Jour. Paleon- Limestone from the Miocene of California, Am. tology, vol. 7, pp. 406-411, pi. 49. Jour. Sci., ser. 5, vol. 21, pp. 304-310, 2 figs. WADE, BRUCE, 1926, The Fauna of the Ripley MILLER, A. K., and UNKLESBAY, A. G., 1943, A formation on Coon Creek, Tennessee, U. S. New Asteroid from the Jurassic of central Wy- Geol. Surv. Prof. Pap. 137, pp. 1-272, pis. 1-72. oming, Jour. Paleontology, vol. 17, pp. 179- WARING, C. A., 1917, Stratigraphic and Faunal 180, pi. 30, fig. 5. relations of the Martinez to the Chico and PACKARD, E. L., 1916, Faunal studies in the Cre- Tejon of southern California, Proc. Calif. taceous of the Santa Ana Mountains of south- Acad. Sci., ser. 4, vol. 7, pp. 41-124, pis. 7-16. ern California, Univ. Calif. Publ., Bull. Dept. WELLER, S., 1905, A fossil starfish from the Cre- Geol. Sci., vol. 9, pp. 137-159. taceous of Wyoming, Jour. Geol., vol. 13, pp. SLADEN, W. P., 1889, Report on the Asteroidea, 257-258. Sci. Res. Voy. H.M.S. Challenger, 1873-76, , 1907, A Report on the Cretaceous paleon- Zool., vol. 30, pp. 1-935, pis. 1-118. tology of New Jersey, Geol. Surv. New Jersey, , 1891, A Monograph on the British fossil Pal. ser., vol. 4, pp. 1-871, pis. 1-111. WHITEAVES, J. F., 1903, Mesozoic fossils (Geol. Echinodermata from the Cretaceous forma- Surv. Canada), vol. 1, pt. 5, pp. 309-416, pis. tions. Volume Second. The Asteroidea, part 40-51. first, pp. 1-28, pis. 1-8 (Paleontographical WHITFIELD, R. P., 1880, Paleontology of the Society, London). Black Hills of Dakota in Newton and Jenney's , 1893, A Monograph on the British fossil report on the geology and resources of the Echinodermata from the Cretaceous forma- Black Hills of Dakota, U. S. Geog. and Geol. tions. Volume Second. The Asteroidea, Part Survey Rocky Mountain Region, pp. 325-468, Second, pp. 29-66, pis. 9-16 (Paleontographi- pis. 1-16. cal Society, London). WHITFIELD, R. P., 1877, Preliminary report on SPENCER, W. K., 1905, A Monograph on the Brit- the paleontology of the Black Hills, U. S. ish fossil Echinodermata from the Cretaceous Geog. and Geol. Survey Rocky Mtn. region, formations. Volume Second. The Asteroidea, pp. 1-49.