Excerptfrom: B.Maguire. 1969. TheBotany of the GuayanaHighland-Part Vlll, Mem.New York Bot. Gard.18:1-290. [Malpighiaceae on pages43-54]

MALptGHtACnarl3 Morethan a decadeago (Mem. N. Y. Bot. Gard. B: 124. 1953.),based upon generouslyample collections,I proposedthe genus Sipapoaas distinct fuom Diacidi'a. Now,with four more speciessince discovered,the separationol Sipapoa and,Di.acidia ts reconfirmed.

12By Leandro Aristeguieta,Instituto Botanico, Apartado 2156, Caracas,. '" By Bassett Masuire, The New York Botanical Garden. ( the internal divisions oI the endosperm, X 1.25 (60735); L-O, Stamens,male plant, X 12.5 (60189): L' dlstal stamen: M, N, intermediatc stamens; O, basal stamen; P, stamen of hermaphrodite ttower,X tz.s (oooas). AA MEMOIRS OF THE NEW YORK BOTANICAL GARDEN Ivor. t8

Key separating Di.acidia and Sipapoa 1. Stipules not at all or hardly connate, hence not vaginate; anthers commonll-barbellate: sepals not becoming ampliate; microphyllous shrubs to 1.5 m high ; on loiv-altitude granitic outcrops, Rios Negro and Vaupds drainages, Venezuela and Colombia. Di 1. Stipules strongly connate,hence conspicuouslyvaginate; anthers glabrous; sepalsbccom_ prominently (except ing ampliate; macrophyllous trees S. cordata) to 20 m high; on sandstone mesas at high altitudes, western Edo. Bolivar and Terr. Amazonas, \,enezuela. siraioa. Diacidia Grisebach,Fl. Bras.12(1): 120. 1858. Key to the Speciesof Diacidia

1. Leavescommonly 4-8 cm long, 2.0-4.5 cm broad; pedicelsoften 1.5 cm long; erect shrubs to I.5 m high; granite outcrops, Rios Negro and Vaup6s, Colombia and Venezuela. 2. Leaves oblong-elliptic, ca.2 cm broad, the base and apex acute or acuminate; petioles 2-4 mm lone; pctals subequal. L D. galbhimioi,t." 2. Leaves ovate,3.5-4.5 cm broad, base obtuse, apex subacute,apiculate; petiolesci 10 mm long: petals unequal, the iifth twice the length of the smaller. 2. D. duchcnn" 1. Leaves commonly 2-3 cm long. 1.2-1.8 cm broad ; pedicels filiform. I .O cm iong; de_ pressed wiry subshrub; sandstone outcrops, Rio Vaup6s drainage, Colombia. 3. D. barailotk, t. Diacidia galphimioides Grisebach,Fl. Bras. 12(1): 120. 1858,ex discr. Type. Casiquiare,Amazonas, Venezuela, Spruce 2966 (K,2 sheets). , Diacidia duckeana \,Iaguire,IIem. N. Y. Bot. Gard.B: 124. 1953.Fig. 15,A-D. Type. Cerro Cocuy, 22 Sep t935, A. Ducke 34633 (holotype US). Additional specimenshave now come to hand: COLONTBIA. Vaup6s: Cerro Mitu, Maguire et al 44091 (NY, US, F, COL), 44100 (I\Y, US, VEN, IAN); on- posite Piedra de Cocui, Schultes& Lopez 9891; San Felipe, Schultcst al tg0g6. VENEZUELA. Amazonas: Cerro Cocuy, Rio Negro, Maguire €t Wurdack 34921, 34943; Cerro Nunca, Rio Negro, Maguire €y Wurdack 34953. Diacidia duckeanamost probably representsthe common expressionof a variable specieswhich would include the types of the original D. galphimloides. Nomenclatural adjustmentwould better await the accumulationof sufficient material to interpret properly the amplitude of the species.Its pollen is 3-colporate,prolate sphaeroidal, ca 22 X 24 1t. 3. Diacidia parvifolia Cuatrecasas,Webbia 13: 632. 1958. Type. Sufnitex reptante, terrenos cuarciticos sabana, San Jos6 del Guaviare, Vaup6s,Colombia, 12 Nov 1939,I. Cuatrecasas7703 (holotypeUS). COLOIIBIA. Vaup6s: Cerro Kafienda,Rio Kubiyri,800-900 ft, 10 Nov 1952,Schultes €t Cabrera 18307,18395;Yapobod6, Rio Kuduyari,900-1000ft, Schulteset al 20049. Diacidia parvilolia seems to be adequately set off from D. galphintioides,al- though it may represent a diminutive extreme. However, since both geographically and ecologicallyit is separatedfrom the latter, D. parvilolia will probabll'be found to maintain its distinctness.

Sipapoa \Iaguire,Mem. N. Y. Bot. Gard.B: 124. 1953. Key to the Speciesof. Sipapoa 1. Inflorescencepaniculate, thyrsoid. 2. Shrub to 1.5 m high; leaves elliptic, obtuse, -1-6 cm long,2 3 cm broad, glaucous beneath, cordate; stipule lobes free; known only from the rim of Cafion Grande, Cerro Neblina, Amazonas, Venezuela. 1. S. clrd'ata' 2. Trees to 20 m high; Ieaves 10 cm or more long; stipules foliar, thc lobes connate, intrapetiolar, exceeding6 cm long; Cerro Neblina, Amazonas, Venezuela and .

..: lir ,t? s i* .ffi W

p6el OF THE GUAYANA HIGHLAND_PART VIII 45

=<-=.

Frc. 11. Sipapoa cordata (Maguire et al 42537D, type): A, habit, X 1; B, detail of paired, connatestipulesi strowing iree stipule lobes, )( 5; C, detail, ultimate branch of inflorescence, brac- teoles,pediiel, opening flower bud, X 10; D, expanded flower, X 10; E, mature pistil, X 10; F' nearmedian section, maturing ovary, X 20; G, stamen, X 20; pollen, diagram polar view, X 1000.

ii ru 46 MEMOIRS OF THE NEW YORK BOTANICAL GARDEN lvor. 1g

and adjacent Serra Pirapucir, Amazonas, Brazil. 3. Leaves strongll' rufus-tomentose beneath, 2. S. rulo 3. Leaves glaucousonly beneath. 1. S. gtauciloiia,. 'I . Inflorescenceraccmosc, trces to 20 m high. 4. Inflorescence glabrous, glaucous; leaves glabrous; stipules foliar, intrapetiolar, ex- ceeding2 cm in length; maturc sepalslanceolate, entire. 4. S. rfiliiloris. 4. Inflorescencestrongly pubescent; stipules not foliar. 5. Leaves glabrous on the upper surface; stipules tardily iI at all separating. 6. Mature sepals oblong or lanceolate. 7. Leaves broadly elliptic to suborbicular, glabrous and glaucous beneath ex- cept for the sparsely pilose nerves and conspicuously ciliate margins; stipule lobes connate, intrapetiolar; sepals denticulate. 5. S. hunharrtr;; 7. Leaves elliptic, membranous, densely sericeous beneath, the nerves obscured, margins not ciliate; stipule lobes free; sepals entire. 6. S. h;,loleur.n 6. Mature sepals ovate-cordate, margins ciliolate, bractlets 4-angled, densell' rcd- hirsute; leaves coriaceous,oblong-elliptic 4-6 cm long,1-2 cm broad, stipules soon separeting,Iobes free. 7. S. Jerrugines 5. Leaves denselypubcscent on the upper surface; stipulessoon separating,lobes iree. 8. Tomentum of undcrsurface oi leaf lanate, underlfing glaucescencepulvinate- squamulose; sepalsstrongly ampliate, to 12 mm long. 8. S. t,estjla. 8. Tomentum of undersurfaceoi leaf appressed-sericcous,underlying glaucesccnce densely cretaceous, mature sepals moderately ampliate, 5-6 mm long. Q. S. steyermarkii

I. Sipapoa cordata l,Iaguire,sp nov. Fig. 11,A-G. Frutex ad 1.5 m altus. Rami teretestenues, mox glaucescentes;internodiis 2-4 cm longis. Folia opposita: laminis oblongisvel late oblongo-ovatis,vulgo 4-6 cm longis,2-3 cm latis, glabris,subtus glaucis,rubritinctis; apicibusrotundatis vel ob- tuse acutiusculis,basibus cordatis. Petioli 6 1Omm longi, 4-6 mm connato-vaginati, extus glabri, glauci, intus denselongi-fusco-pilosi; lobis stipulis jugatis, deltoideis. lanceolatis,acutis, 5-7 cm longis, liberis; petiolulo supra libero deficiente;costa prominenta,rubra; venis lateralibus 10 12-jugis,prominulis. Inflorescentiaetermi- nales angustethyrsoideae; rhachidibus 5-10 cm longis, cymosis; ramulis 1-3 cm longis,secundis, rubritinctis, 3-5-floribus; pedicellistenuibus, 8-20 mm longis,arti- culatis: bracteisbracteolisque ovatis, obtusis, glabris, glaucis, 3-7 mm longis,caducis. Floreszygomorphi; sepalis 5, Ianceolatis,acutiusculis, cordatis, integris, apicibus, re- curvatis,ca 3 mm longis,glabris, venatis, ad basimduobus rotate glandibusrecurvatis, ca 1 mm longis instructis; sepalisin maturate ampliatis 6-8 mm longis: petalis5, ca 5 mm longis,laminis ovato-orbicularibus,3.0-3.5mm diam, suberosemarginatis, venatis,unguibus 1.0-1.5 mm longis,tenuibus, trigonis, sulcatis; petalo quinto majori, ca 10 mm longo,lamina elliptico-obovata;staminibus 8-10, ca 3 mm longi-s,petalo quinto oppositolongiori; filamentisteretibus, introrsis: antherisglabris, bilocularibus, oblongis,ca 0.7-0.9 mm longis, thecis 1-2-spinuloso-aristatis,aristis 0.4-0.6 mm longis, introrse nectantibus,connectivo crasso, thecis excedente.Grana pollina 3- colporata,prolati-sphaeroidea, 30-33 X 28-30 p; sporodermaminute vermiformi- rugulosavel reticulata. Ovarium globosumglabrum biloculare, tristylare; stylis subu- latis ca 2.0-2.5 mm longis; stigmatibus minutis intrbrse subterminalibu-siovulis suspensis.Fructus non-visus. Type. Branched,several-stemmed shrub 1 m high, petalsyellow, calyx greenish, infrequent,rim of Cafion Grande east of Cumbre Camp, elev 1800 m, , Rio Yatira, Territorio de Amazonas,Venezuela, 23 Dec 7957, Bassett Magui,re,lohn l. Wurdack €t Celia K. Maguire 42537D (holotype NY). Distribution. VENtrZUELA. Amazonas:Cerro de la Neblina, wandlike shrub 0.5 1.5 m high, leavesglaucous, red-edged, occasional around exposedsandstone out' crops,south slope toward CafionGrande, alt 1500-1700m, 16 Jan 1954,Maguire et al p6el BOTANY OF THE GUAYANA HIGHLAND-PART VIII

,wi qt{F# StrFe

N'l%\/t h.i ,6' Vd

'l' '

,,] -- .

Frc. 12. Sipa\oa rttla (Maguire et al 42036,type): A, habit, y 1; B, undersideof leaf, X 10; C, ultimate branct of inilorescence, showing pedicel, bracteoles, flower bud, X 10; D, expanded 10; H, near median Ilorver,X 10; E, stamen, X 20; F, mature calyx, X 10; G, mature sepal, N long section, maturing pistil, X 10. GARDEN 48 MEMoTRS oF THE NEw YoRK BorANrcAL [vor. 18

37320 (paratvpe); caflon Grande,23 Dec 1957, Maguire et ul 42537D (holotypeNY. isotypesNY, COL, F, US, S, P, L, UC, IAN, U). 2. Sipapoa rufa N{aguire,sp nov. Fig. 12, A-H; Fig. 16,H-K. Arboresparvae, ad 20 m altae. Rami teretes,7-10 mm crassi,glabri: internodiis 3-6 cm longis. Folia opposita; laminis late oblongo-obovatis,vulgo 15-20 cm longis,vulgo (8) 10 15 cm iatis, obtusis,supra glabris,subtus denserufo-appresso- tomentosis,tomento subter dense glauco; costa nervisque lateralibus densioribus tomentosis,supra prominentibus, vulgo 14-15-jugis.Petioli 15-20 mm longi,I'aginati, connati, intus densepilosi; stipulis intrapetiolaribus,connatis, conspicue foliaribui oblanceolatis,glabris, vulgo 6-8 cm longis; petiolulosupra libero ca 10-12 mm longo. Inflorescentiaeterminales thyrsoideae; rhachidibus ad 30 cm longis cymosis,ruio- tomentosis;ramulis ad 3 cm longis,3-7-fioribus; pedicellis6-10 mm longis,articu- latis; bracteisdeltoideo-ovatis, 3-5 mm longis, caducis. Flores zygomorphi;sepalis 5, angustelanceolatis, vulgo acuminatis,evidenter serrato-dentatis, ca 5 mm longis, apicibusrecurvatis, demum ampliatis 8-10 mm longis, valde reticulato-nervosis,ad bisim duobusglandibus oblongis subtendentibus; petalis 5, ca 5-6 mm longis;laminis ovato-orbicularibus,4-5 mm diam, suberosemarginatis, venatis, unguibus 1.0-1.5 mm longis, tenuibus, trigonis, sulcatis; petalo quinto marjori, ca L0-12 mm longo, lamina elliptico-obovata;staminibus 8-10, introrsis,petalo quinto oppositolongiori: filamentisleretibus, introrsis, antheris, glabris, bilocularibus, oblongis, ca 0.8-1.0mm longis,thecis l-2 spinuloso-aristatis,aristis 0.5-0.7 mm longis introrsenectantibus; connectivocrasso. Grana pollina 3-colporata,prolati-sphaeroidea, 20-22 X 19-20p; sporodermaminute reticulata. Ovarium biloculare, globosum; stylis subulatisca 2-3 mm longis, stigmatibusminutis introrse subterminalibusiovulis suspensis.Pyrene ovata, bilobata, ca 2 mm longa,valde reticulatenervata' Type. Tree 4 8 m high, flowers yellow, occasionalon escarpmentslopes above Camp 3, elev 1600-1700m, Cerro de la Neblina, Rio Yatira, Territorio Amazonas' Venezuela,17 Nov 1957, Bassett Maguire, lohn J. Wurdack €v Celia K' Llaguire 42036 (holotypeNY; isotypesF, IAN, US, VEN)' Distribution. VENEZUELA. Amazonas:Cerro de la Neblina, Rio Yatfa, tree 5-10 m high, frequenton upper escarpmentslopes, alt 1600-1700m,27 Dec 1953, Maguire et al 36917 (NY, US, VEN, F, K); tree 5-10 m high, frequenton escarp- -.nt, ult 1650-1750m,30 Dec 1953,Maguire et al 36991 (NY); smalltree 3-8 m high, leaves silky brown-pubescentbeneath, stipules connate at base, glaucous,in- flJrescencetawny, floweri yellow, frequent on low bush slopes near Cumbre Camp, VEN' IAN' alt 18OOm, 4 Jan 1954,Maguire et al 37034 (paratypesNY, B' COL, GH, F); tree 5-10 m high, locally frequenton northwestescarpment, alt 1700-1800 ., iO jun 1954,Magui'ri et at 37152 (NY, K, S' MO); tree 5-12 m high' frequent m, on lower aralt igOOm, 18 Jan 1954,Magui,le et al 3734S(NY); open tree to 10 P); escarpmentslopes at alt 1OOOi, Z+ 1^n 1954,Maguire et al 37378 (NY, L:C' tree i-8 m high, on escarpmentslopes at alt 1600-1700m a1t,17 Nov 1957,Xlaguire. of et at 42036 (lolotype NY; isotypesF, IAN, US, VEN); small tree along rim CaffonGrande ar alt 1800 ^, iS D", 1957,Magui,re et aL 42461 (Ny,IIICT]' BRAZIL. Amazonas:2 m high, 2 cm diam, in low humid catingaforest at alt 1300- (para- l7O0 m, Serrapirapucir, Rio 1iaturach,2T Jan 1966, Si,tva& Brazao 60930 typesNY, IAN, F, LTS). E-G' 3. Sipapoa glaucifolia N{aguire,sp nov. Fig' 13; Fig' 16' A S. rula Mag. affinis et similis, sed foliis glabris, subtus dealbatis,farinoso- squamulosis;inflorescentiis paucipubescentiis differt' 49 p6e) BOTANY OF THE GUAYANA HIGHLAND-PART VIII

1; B, expandedflower' Frc. 13. Sila.poagloucilolia (Maguire et ol 42057,type): A, habit, .X pistil' 10; X 10; C, stamen,k zo;O, maturecalyx, X 10; E, nearmedian iong section,maturing X F. pol)en,diagram of polar view. X 1000. 50 MEMoTRSoF THD NEw yoRK BorANrcALGARDEN I vor. 1g

Type. Tree 8 m high, adult leaveswaxy-glaucous beneath, flowers velioq,,shooto :if from lower trunk, frequentin Clusi,qtangle just south of Camp 3, lowland and st*" forests,ait 700 m, Cerro de la Neblina, Rio Yatira, Territorio de Amazonas,\'.n1" zuela, 14-16 Nov 1957, Bassett Maguire, Iohn l. Wurdack €y Celia X. Uo,-u'i)) 42057 (holotypeNY; isotypesNY, US, VEN, F, K, GH, UC). Distribution. VENEZUELA. Amazonas:Cerro de la Neblina, Rio Yatua, tree 6-15 m high, leaveshighly glaucousbeneath, in Clusi.a"moss-forest," alt 650-700m 23 Dec t953, Maguire et al 36821 (Ny, W, LIC, GH, N{); in Clusia "rno=r-for.st'il alt 650 m, 29 Dec 1953,Magai.re et al 36968 (paratypesNY, US, VEN, K, S, COi F, P, B); tree to 15 m, frequentin Clusia tangle in slopeforest, alt 650 m. 2J Tan 1954,Maguire et al 37362 (l{Y, US, VEN, LL, IAN); tree 8 m high, aclultleaves waxy-glaucousbeneath, frequent in Clusiatangle, alt 700 m, 14-16 Nov 1957,IIaguire et al 42057 (holotypeNY; isotypesNY, US, VEN, F, K, GH, UC); tree 8 m hish. adult leaveswaxy-glaucous beneath, frequent in Clusi.atangle, alt 700 m, l-i 16 \6r, 1957,Maguire et a|420571 (VEN, NY, US) ; Maguire et a|420578 (NY, LrS,VEN, F); tree 5-10 m high, leaveswaxy-glaucous beneath, along banks of Caion Grande, alt 1100m, 24 Nov 1957,Maguire et al 42216 (Ny, US, VEN, RIO). BRAZIL, Amazonas:Tree 8 m high,5 cm diam, intrapetiolar stipule large, beti,veenXlissao Salesianaand SerraPirapucri, alt 800-i000 m, Rio nfaturac6, 23 Jan 1966,Sift,a & Brazdo 60867 (paratypesNY, IAN, US, F, S, K, GH, UC, COL, RIO).

4. Sipapoa stipularis llaguire & Phelps,Mem. N. Y. Bot. Gard. B: 127. 1953. Type. Cerro Parf, Rio Ventuari,Amazonas, Venezuela,2 Feb 1951,Couan €r Wurdack 31200 (h:nlotype NY). Known only from the cumbre of Cerro Parir: Phelps€y Hitchcock471; Cowan& Wurdack31112,31168,31292.

5. Sipapoa kunhardtii lIaguire, Mem. N. Y. Bot. Gard. B: 125. 1953. Type. Cerro Sipapo,Rio Orinoco,Amazonas, Venezuela, 15 Dec 1948,Maguire & Poli,ti 27677 (holotype NY). Known only from the cumbre drainageof Cafro )iegro, Cerro Sipapo Maguire & Politi, 27622, 28105.

6. Sipapoa hypoleuca Maguire,N{em. N. Y. Bot. Gard. B: 125. 1953. Tlpe. Cerro Yapacana,Rio Negro, Amazonas,Venezuela, 3 Jan 1951,Xlaguire, Cowan €t Wurdack 30704 (holotype NY). Known only from the cumbre of Cerro Yapacana:Holt & Blake 707; Maguireet al 30672,30635,30708,30710.

7. Sipapoa ferruginea Maguire & Phelps,NIem. N. Y. Bot. Gard. B: 126. 1953. Type. Cerro Pari, Rio Ventuari, Amazonas,Venezuela,4 Feb 1951,Cowan €v Wurdack 31233 (holotype NY). Known only from the cumbre of Cerro Paru: Phelps €t Hitchcock 516; Couan €t lVurdack 31066, 31079, 31080, 31081,31150, 31171.

B. Sipapoa vestita (Bentham) Maguire, N{em. N. Y. Bot. Gard. B: 127. 1953. ('Mountains Tlpe. of British Guiana,Schomburgh s z" (holotype K). Sipapoavestita has not sincebeen collectedin "British Guiana." It is probable that the original was collected by Schomburgk in VenezuelanAmazonas ancl on the ('British' label erroneouslyascribed to Guiana." VENEZUELA. Amazonas:, Tate 563; ,Maguire et al 29806,29858,29869,30091, 30111.30221. 30252. p6el BOTANY OF THE GUAYANA HIGHLAND-PART VIII 5l

,,,1 l' \ il t,,'r

Sn-noon

Frc.14. Sipapoa steyermarkii (Steyermark 98016,holotype): A, habit, X 1; B, detail showing separatedstipules r'vith free lobes at the summit, v.2; C, flower and petioles lvith paired bracteoles at base,X 10; D, flower, X 10; E, stamen, X 20.

54 MDMoTRS oF THE NEw YoRK BorANrcAL GARDEN lvol. 18

9. Sipapoa steyermarkii I'{aguire,sp nov. Fig. 14, A-E; Fig' 16. A-D. A S. vestitaeBentham valde affinis, sed laminis cordatiset pubescentiisciensiori- bus, nervis magis prominentibus,differt. Arbor parva 4-5 m alta. Rami teretesdense cinereo-piloso-hirsuti. mox glabres- centes; iniernodiis 1.5 2,0 cm apartis. Folia opposita; laminis ellipticis vel lanci- ellipticis, 6,7 cm longis, 3.0 4.5 cm latis, cum 7-9-jugis nervorum primariorum lateralium, supra tenuiter puberulis, nervis impressis,subtus densissimiscinereo- piloso-hirsutis,pilis valde stellato-pustulatis; nervis prominentibus; apicibus s6- iusiusculis,caespituiis; basibus cordatis; marginibus aliquantum involutis' Petioli 2.0-3.5 cm longi, partibus liberis 7-10 mm longis, ad basim ca l-2 cm connatis; vaginis aen.. piioii. intus; stipulis iobis liberis, ovatis.vel lanceolatis,acutis vel acuminatis,12-15 cm longis, glabris intus. Inflorescentiae10-15 cm iongae,r4qg- mosae,dense piloso-hirsutae; pedicellis tenuibus, 10-12 cm_.longis,bibracteolatis, bracteolisca 1.5mm longis,cadr"rcis; floribus flavis 12-14 mm diam: sepalis5. lanceo- latis, acutis, ca 4 mm longis, valde hirsutis, glandibusoblongis ca 12 m.mlongis; petaiis 5, flavis, subaequalibus,6-8mm longis,unguibus ca 1'5 mm longis,laminis trbicularlovatis, 4-5 mm longis ca 4 mm iatis, irregulariter erosecrenulatis. acl basim cordatis,petalo quinto 7-8 mm longo,lamina ovata; staminibus8-9, ca 3.5 4'3 mm Iongis, posterioribuslongioribus, filamentis teretibus, ad basim intus pauci-longi- pilolii, connectivoprominenti obtusoO.I-0.2 mm projecto,antheris 2-theci-s oblongis ca 0.i mm longis, introrsis, apice 2-spinuloso-aristato,aristis nectantibus. Grana pollina 3-colporata,prolati-sphaeroidea, 22-23 X 2O-22 p, sporodermagranulari. buariu- globosum 2-loculare,glabrum, 3-stylare; loculis 1-ovrrlatis,ovulis dependen- tibus; stylis subulatis,divergentibus, 2.5 mm longis,acutis. Fructus non-vistts. Type. Tree 4-5 m with erect branches, leaves subcoria-ceous,rich green with impresied nerves abover sericeous-buffbelow, alt 1900-2100 m, Cerro Jaua, Rio Kanarakuni,Rio caura, Eclo.Bolivar, Venezuela,22-27 Mar 1967,lulian A' Steyer' mak 98016(holotype NY; isotypevEN). Known only by the type collection. This handsometreer clearly allied to Si,papoauestita, is representedonll' b-vthe type collection. It is readily distinguisheclfrom s. vestita by its cordateleave-", denser pub.scence,and more prominent nerves,which are impressedin the upper suriace is is characteristicof its immediatecongener, the connatestipules soon separate'

Pollen general Pollen grains ol Di,acidia and Sipap\a are similar, and conform to the (20-33 x pattern of malpighiaceouspollens, viz, 3-colporate,prolate sphaeroidal 19-31 r.), with finely reticulatesporoderms.