J. Jpn. Bot. 84: 237–254 (2009)

Lectotypes and Original Material for Blume’s Species and Varieties of from Japan

a b c Hideaki Ohba , Shinobu Akiyama and Gerard Thijsse

aDepartment of Botany, the University Museum, the University of Tokyo, 7-3-1, Hongo, Bunkyo-ku, Tokyo, 113-0033 Japan; E-mail: h-ohba@sd. dcns. ne. jp bDepartment of Botany, National Museum of Nature and Science, 4-1-1, Amakubo, Tsukuba, 305-0005 Japan; cNationaal Herbarium Nederland, Leiden University branch, P. O. Box 9514, 2300 RA Leiden, the Netherlands

(Received on March 5, 2009)

In 1851, Carl Lodewijk von Blume, who studied extensively the specimens collected in Japan by Siebold, Bürger, their successors, and their Japanese correspondents, described a great number of new species and varieties of Quercus and other genera of Fagaceae. His treatment, especially of Quercus, influenced later workers on the family, and also studies of the flora and vegetation of Japan. To stabilize the taxonomy of the family, lectotypes for taxa described by Blume are herein designated for Quercus (13 species and 13 varieties), Lithocarpus (1 and 2), Castanea (1 and 10), and Fagus (1 and 2).

Key words: Fagaceae, Japanese flora, lectotypification, Siebold collections.

Carl Lodewijk von Blume (Karl Ludwig not designate lectotypes. The type concept was von Blume) studied extensively the specimens not as well developed at those times as it is collected in Japan by Siebold, Bürger, their now. successors, and their Japanese correspondents. Siebold, Bürger, their successors, and Blume (1851) described a great number of their Japanese correspondents gathered the new species and varieties of Quercus and specimens from various sources. The majority other genera of Fagaceae, and his treatment, of the specimens appear to be from trees especially of Quercus, influenced later workers in spontaneous situations, but some were on the family and also studies of the flora and collected from cultivated trees in various vegetations of Japan. gardens and nurseries. Although several Japanese botanists During preparation of manuscripts on studied the specimens used by Blume, they Fagaceae for recent publications on the did not designate any lectotypes. Gen-ichi Japanese flora (Ohba 1989, 2006), one of us Koidzumi in 1925 and Takenoshin Nakai in (Ohba) awakened to the need to lectotypify 1926 visited Leiden, but did not designate the names of all specific and infraspecific lectotypes for any of Blume’s taxa. In 1958 taxa described by Blume from Japan. Such Hiroshi Hara studied several specimens of an undertaking is necessary for the sake of Quercus described by Blume, but he also did stability in the application of scientific names

—237— 238 植物研究雑誌 第 84 巻 第 4 号 2009 年 8 月 to Japanese and their allies. Since 2005, Additional original material: Japonia. Akiyama and Thijsse have collaborated in Bürger s.n. (L 0103235). the typification of taxa described by Siebold Current name: Castanea crenata Siebold & and Zuccarini, and Miquel based on Siebold’s Zucc. collections. We have also extended our 1a) [β] var. quercina Blume, Mus. Bot. 1: interests to Blume’s taxa, of which the type 285 (1851). materials are also kept in the ‘Herbarium Type: Japonia. Siebold s.n. (L 0103229 Japonicarum Generale’ in Leiden (L), and –lectotype, designated here). with corresponding specimens in München Additional original material: Japonia. (M). All the specimens in L and M collected Siebold s.n. (L 0103236). by Siebold, Bürger, their successors, and their This specimen represents wild individuals Japanese correspondents listed in Yamaguchi with small leaves. et al. (2003). Current name: Castanea crenata Siebold & This paper aims to provide data related Zucc. to the types and original materials of all 1b) [γ] var. longispina Blume, Mus. Bot. 1: specific and infraspecific taxa described from 285 (1851). Japan by Blume in 1851. Blume (1851) did Type: Japonia. Siebold s.n. (L 0103254– not designate types nor cite the specimens lectotype, designated here). he examined. Except in a few cases, no Current name: Castanea crenata Siebold & lectotypifications of Blume’s taxa ofFagaceae Zucc. have been made. We designate here lectotypes 1c) [δ] var. kusakuri Blume, Mus. Bot. 1: for all taxa described by Blume from Japan 285 (1851). according to guidelines in the Vienna Code Type: Japonia. Bürger s.n. (L 0103252– of the International Code of Botanical lectotype, designated here). Nomenclature (McNeill et al. 2006). The Additional original material: Japonia. designations are listed in alphabetical order, Bürger s.n. (L 0103253), with Koidzumi’s Castanea (1), Fagus (2), and Quercus (3–24). identification slip. The present taxonomic status of all taxa Current name: Castanea crenata Siebold & are given in Ohba (2006). The specimens, Zucc. recorded on the CD-ROM database titled “Von This specimen, consisting of a small branch Siebold’s Botanical Treasures in Leiden” (de with many small leaves, appears to have been Kort et al. 2000), are indicated with an asterisk collected from an unusual or juvenile . after the register number of the Nationaal 1d) [ζ] var. ookasi Blume, Mus. Bot. 1: 285 Herbarium Nederland, Leiden University (1851). Branch (L) in parentheses after the collector’s Type: Japonia. Bürger s.n. (L 0103234 number of each specimen. –lectotype, designated here). Additional original material: Bürger s.n. (L Castanea 0103233). 1) Castanea japonica Blume, Mus. Bot. 1: Current name: Castanea crenata Siebold & 284 (1851). Zucc. Type: Japonia. Siebold s.n., as Castanea This is an unusual form with relatively vesca Gaert. (L 0103250, 0103251 [two small, narrowly oblong leaves with 14 to 16 register numbers for one specimen] pairs of lateral veins, truncately or rounded –lectotype, designated here). [Fig. 1] cuneate equal base and acuminate apex. The August 2009 Journal of Japanese Botany Vol. 84 No.4 239 procera Blume [β] var. Fagus crenata Blume. Bürger s.n. (L 0103268). s.n. (L Blume. Bürger Fig. 2. Lectotype of Blume. Siebold s.n. Castanea japonica (L 0103250, 0103251 [two register numbers for one specimen]). (L Fig. 1. Lectotype of 240 植物研究雑誌 第 84 巻 第 4 号 2009 年 8 月 spines on the margins are not extruding, and Bürger s.n. (L 0103255); Bürger s.n. (L most of them are short and inconspicuous, and 0103256). some of them are ascending. Current name: Castanea crenata Siebold & 1e) [η] var. spontanea Blume, Mus. Bot. 1: Zucc. 285 (1851). This specimen represents the rather Type: Siebold s.n., with name “Chiuguri” common wild individuals with narrowly (in Japanese) (L 0103264–lectotype, oblong leaves and small cupules. Pierot 647, designated here). collected “in sylvis prope urbem Yedo ins. Additional original material: Japonia. Nippon,” apears to be a duplicate of Bürger Siebold s.n. (L 0103263). s.n. (L 0103256). Current name: Castanea crenata Siebold & 1i) [λ] var. elongata Blume, Mus. Bot. Zucc. 1: 286 (1851). 1f) [θ] var. vulgaris Blume, Mus. Bot. 1: Type: Japonia. Bürger s.n. (L 0103247 285 (1851). –lectotype, designated here). Type: Japonia. Siebold s.n., as Castanea Additional original material: Bürger s.n. (L vesca α.vulgaris (L 0103265–lectotype, 0103248). designated here). Current name: Castanea crenata Siebold & Current name: Castanea crenata Siebold & Zucc. Zucc. 1j) [µ] var. canescens Blume, Mus. Bot. 1: 1g) [ι] var. siba-kuri Blume, Mus. Bot. 1: 286 (1851). 286 (1851). Type: Japonia. Siebold s.n. (L 0420819 Type: Japonia. Pierot s.n. (L 0103228– –lectotype, designated here). lectotype, designated here). Additional original material: Japonia. Additional original material: Japonia. In Siebold s.n. (L 0420820). collibus sylvosis juxta pagum Kawasaki ins. Current name: Castanea crenata Siebold & Nippon. Pierot 420 (L 0103237); Bürger s.n. Zucc. (L 0103231). The leaves are conspicuously pale whitish Current name: Castanea crenata Siebold & with dense soft hairs on the lower surface and Zucc. has 16 to 18 conspicuous lateral veins on each Pierot s.n. (L 0103228) has rather narrow side. leaves with unequal rounded base, long acuminate apex, serrulate-cuspidate margins, Fagus and sparse glandular hairs when young, and 2) Fagus crenata Blume, Mus. Bot. agrees with the original description. Pierot 420 1: 307 (1851). (L 0103237) consists only of a branch with Type: Japonia. Keiske Ito (in Herb. Siebold) many leaves. Some leaves of the basal part 14, with names “Sobagurumi” and “Kurobuna” are widely lanceolate, 21 x 6.7 cm, others in (in Japanese) and “Fagus Kurobuna” (L the apical part are narrowly oblong. This also 0040617*–lectotype, designated here). agrees with the original description. Additional original material: Japonia. 1h) [κ] var. oblexta Blume, Mus. Bot. Siebold s.n., with names “Bunanoki” 1: 286 (1851). (in Japanese) and “Fagus ferruginea” Type: Japonia. Bürger s.n. (L 0103232 (L 0175953). –lectotype, designated here). Current name: Fagus crenata Blume. Additional original material: Japonia. 2a) [β] var. procera Blume, Mus. Bot. August 2009 Journal of Japanese Botany Vol. 84 No.4 241

1: 307 (1851). 299 (1851), ut bürgeri. Type: Japonia. Bürger s.n. (L 0103268 Type: not yet found in L and M. –lectotype, designated here). [Fig. 2] 5) Quercus canescens Blume, Mus. Bot. 1: Additional original material: Japonia. 296 (1851). Siebold s.n. (L 0103267). Type: Japonia. Bürger s.n. (L l0040861* Current name: Fagus crenata Blume. –lectotype, designated here). 2b) [γ] var. undulata Blume, Mus. Bot. 1: Additional original material: Japonia. 307 (1851). Siebold s.n. (L 0040862). Fagus sieboldii Endl. var. undulata Blume Current name: Murray. ex Miq., Ann. Mus. Lugduno-Batavi 3: 194 6) Quercus crispula Blume, Mus. Bot. 1: (1867); H. Ohba & al. in Bull. Univ. Mus. 298 (1851); H. Ohba in K. Iwats. & al., Fl. Univ. Tokyo (41): 38 (2005). Jap. 2a: 47 (2006). Type: Bürger s.n. (L 0040618*–lectotype, Type: Japonia. [In saltibus montis designated here). Hakone insulae Nippon]. Bürger s.n. Additional original material: Japonia. (L 0040838–lectotype, designated here). Bürger s.n. (L 0040619). [Fig. 4] Current name: Fagus crenata Blume. Current name: Quercus crispula Blume. Bürger s.n. (L 0040618) was also Pierot 642 (L 0040837*), with a note on designated as the lectotype of Fagus sieboldii the habitat, “In saltibus montis Hakone ins. var. undulata Blume ex Miq. by Ohba et al. Nippon,” was also collected in Hakione, and (2005). is regarded as a duplicate of the lectotype, because the Pierot collection from Japan was Quercus actually collected by Bürger. The lectotype, 3) Quercus aliena Blume, Mus. Bot. which lacks detailed collection information, 1: 298 (1851). was probably collected in Hakone (central Type: Japonia. Siebold s.n. (L 0040832* Honshu) as mentioned by Pierot. –lectotype, designated here). The treatment of Quercus crispula is still Additional original material: Japonia. controversial, with two schools of thought: Bürger s.n. (L 0103196). one is that it is a distinct species endemic to Current name: Quercus aliena Blume. Japan (with some extension to the adjacent 3a) [β] var. pellucida Blume, Mus. Bot. 1: regions), and two, that it is conspecific withQ. 298 (1851). mongolica Fisch. Ohba (1989, 2006) treats it Type: Japonia. Siebold s.n. (L 0040831 as a distinct species. –lectotype, designated here). [Fig. 3] 7) Quercus gilva Blume, Mus. Bot. Additional original material: Japonia. 1: 306 (1851). Siebold s.n. (L 0103195); Bürger s.n. Type: Japonia. Bürger s.n. (L 0103348 (L 0103197). –lectotype, designated here). [Fig. 5] Current name: Quercus aliena Blume var. Additional original material: Japonia. pellucida Blume. Bürger s.n. (L, L 0103347); Bürger s.n., with The glabrous or sparsely stellate-hairy name “Quercus Konara” (L 0103346); Siebold lower surface of the leaves of var. pellucida s.n. with names “Ichiikasi” (in Japanese), distinguish it from var. aliena, which has “Quercus itsikas” (L 0103345). dense stellate hairs. Current name: Quercus gilva Blume. 4) Quercus buergeri Blume, Mus. Bot. 1: 7a) [β] var. procera Blume, Mus. Bot. 242 植物研究雑誌 第 84 巻 第 4 号 2009 年 8 月 Blume. Bürger s.n. Quercus crispula (L 0040838). (L Fig. 4. Lectotype of pellucida Blume [β] var. Quercus aliena Blume. Siebold s.n. (L 0040831). Blume. Siebold s.n. (L Fig. 3. Lectotype of August 2009 Journal of Japanese Botany Vol. 84 No.4 243

1: 306 (1851). Nakai. Type: Japonia. Bürger s.n. (L 0103350 Two specimens collected by Bürger s.n. –lectotype, designated here). [Fig. 6 ] (L 0103280, 0103278) have the handwritten Additional original material: Japonia. “Quercus glabra Thunb. var. micrococca,” Bürger s.n. (L 0103351); Siebold s.n. with probably by the same person on the lectotype. names “Yamabiwa” (in Japanese) and Blume characterized this variety by the larger “Quercus Yamabiwa” (L 0103349). leaves (2.5–6.5 poll., 3/4–2.5 poll. wide; Current name: Quercus gilva Blume. petioles 6–10 lin.), and the minor cupules, The leaves of the lectotype and original which have the nuts usually concave apically. material of Quercus gilva (var. gilva) are These two specimens lack cupules, and the mostly 5–7 cm long and 1.7–2 cm wide; those leaves are no larger than those of Bürger s.n. of var. procera are 7–17 cm long and 3–4 (L 0103279). cm wide. In Quercus gilva the leaves vary The characters used by Blume for this continuously from 6–14 cm long and 2–4 cm variety are considered to be taken only from wide (Ohba 2006). No other differences can Bürger L 0103279. be found between these two varieties. 9) Quercus glandulifera Blume, Mus. Bot. 8) Quercus glabra Thunb. 1: 295 (1851). 8a) [β] var. sublepidota Blume, Mus. Bot. [var. glandulifera]. 1: 289 (1851). Type: Japonia. Siebold ex an unknown Type: Japonia. Bürger s.n. (L 0103281 Japanese collector [? Keiske 14] s.n. –lectotype, designated here). [Fig. 7] (L 0040833*–lectotype, designated here). Current name: Lithocarpus edulis (Makino) Current name: Quercus serrata Murray. Nakai. 9a) [β] var. subattenuata Blume, Mus. Bot. The Bürger specimen (L 0103281) includes 1: 295 (1851). two branches, probably collected in early Type: Japonia. Bürger s.n. (L 0040863* summer with staminate inflorescence and –lectotype, designated here). nuts of the second year. The petiolate leaves Additional original material: Japonia. are 7–9 cm long and 2.8–3.3 cm wide with Bürger s.n. (L 0325670, 0103292, 0103293); cuspidate apex, long cuneate base and entire Keiske 330, with names “Sodara”, margins. These leaves, without doubt, fall “Konara”, “Ho” (L 0175762); ?Siebold s.n. within the range of variation seen in leaves of (L 0103294). Lithocarpus edulis. Pierot 203 (L 0103282, Current name: Quercus serrata Murray. 0103272), collected “in sylvis montis Kuni Mi 9b) [γ] var. inaeaqualiserrata Blume, Mus. Yama ins. Kiu Siu,” is regarded as a duplicate Bot. 1: 295 (1851). of the lectotype. Pierot retained the Japanese Type: Japonia. Siebold s.n. (L 0040836* name “Satuma Si” on the label. –lectotype, designated here). 8b) [γ] var. micrococca Blume, Mus. Bot. Additional original material: Japonia. 1: 290 (1851). Siebold s.n. (L 0103289). Type: Japonia. Bürger s.n. (L 0103279 Current name: Quercus serrata Murray. –lectotype, designated here). [Fig. 8] 9c) [δ] var. subcrenata Blume, Mus. Bot. Additional original material: Japonia. 1: 295 (1851). Bürger s.n. (L 0103280); Bürger s.n. Type: Japonia. Siebold s.n. (L 0103290 (L 0103278). –lectotype, designated here). [Fig. 9] Current name: Lithocarpus edulis (Makino) Current name: Quercus serrata Murray. 244 植物研究雑誌 第 84 巻 第 4 号 2009 年 8 月 procera Blume [β] var. Quercus gilva Blume. Bürger s.n. (L 0103350). s.n. (L Blume. Bürger Fig. 6. Lectotype of Blume. Bürger s.n. Quercus gilva (L 0103348). (L Fig. 5. Lectotype of August 2009 Journal of Japanese Botany Vol. 84 No.4 245

micrococca Thunb. [γ] var. var. [γ] Thunb. Quercus glabra glabra Quercus Blume. Bürger s.n. (L 0103279). (L s.n. Bürger Blume. Fig. 8. Lectotype of of Lectotype 8. Fig. Blume. sublepidota Thunb. [β] var. Quercus glabra Bürger s.n. (L 0103281). s.n. (L Bürger Fig. 7. 246 植物研究雑誌 第 84 巻 第 4 号 2009 年 8 月

Mohnike s.n. (L 0040834) and Pierot 185, Ywaja circa altitudinem 800 ped [sic], cum collected in “saltibus montis Hakone ins. vere cuspida et KuroKaj. Siebold s.n., with Nippon” (L 0103291) are also representative. name “Quercus Sirokasi” (L 0103316sic); 9d) [ε] var. polymorpha Blume, Mus. Bot. Bürger s.n. (L 0103317). 1: 295 (1851). Current name: Quercus salicina Blume. Type: Japonia. Bürger s.n. (L 0103286* All of these specimens represent Quercus –lectotype, designated here). salicina Blume with lanceolate leaves. Additional original material: Japonia. 10d) [ε] var. caesia Blume, Mus. Bot. Buerger s.n. (L 0103287); Siebold s.n., with 1: 303 (1851). name “itsikas” (L 0040835). Type: Not yet found in L and M. Current name: Quercus serrata Murray. 11) Quercus grosseserrata Blume, Mus. Blume described Quercus glandulifera Bot. 1: 306 (1851). including four varieties, Q. canescens and Quercus crispula Blume var. grosseserrata Quercus urticiifolia as species different from (Blume) Miq., Ann. Mus. Lugduno-Batavi 1: Quercus serrata Murray, and considered 104 (1893). to be Quercus serrata the present Quercus Type: Japonia. Bürger s.n. (L 0040839* acutissima Carruth. The three species and four –lectotype, designated here). varieties described by Blume are conspecific Current name: Quercus crispula Blume. with Quercus serrata. The lectotype specimen consists of a branch 10) Quercus glauca Thunb. with five, almost sessile, widely oblanceolate 10a) [β] var. nudata Blume, Mus. Bot. leaves with fine rough serrations. It represents 1: 303 (1851). the typical form of tillers with leaves of Type: Japonia. Bürger s.n. (L 0103310 Quercus crispula. De Kort et al. (2000) –lectotype, designated here). [Fig. 10] errorenously reported this specimen to be Additional original material: Japonia. the type of Quercus crispula f. grosseserrata Bürger s.n. (L, L0103311); Siebold s.n. Blume. (L 0103312). 12) Quercus hirsutula Blume, Mus. Bot. Current name: Quercus glauca Thunb. 1: 298, in nota sub Q. aliena Blume (1851). Pierot 652, collected “in sylva prope Type: not found in L and M. pagum Umesima ins. Nippon” (L 0103313), 13) Quercus holosericea Blume, Mus. Bot. apears to be a duplicate of Siebold s.n. 1: 298, in nota sub Q. aliena Blume (1851). (L 0103312). Type: not found in L. and M. 10b) [γ] var. fasciata Blume, Mus. Bot. 1: 14) Quercus humosa Blume, Mus. Bot. 1: 303 (1851). 298 (1851), in nota sub Q. aliena Blume. Type: Japonia. Siebold s.n. (L 0103307 Type: Not found in L and M. –lectotype, designated here). [Fig. 11] 15) Quercus lacera Blume, Mus. Bot. Current name: Quercus glauca Thunb. 1: 306 (1851). 10c) [δ] var. stenophylla Blume, Mus. Bot. Type: Japonia. Siebold (ex Keisuke) s.n., 1: 303 (1851). with name “Hiryukasi” (L 0103319–lectotype, Type: Japonia. Siebold s.n., with name designated here). [Fig. 13] “Quercus sirokasi” (L 0103315–lectotype, Current name: Quercus glauca Thunb. designated here). [Fig. 12] This is Quercus glauca Thunb. with deeply Additional original material: Japonia. lobulate leaves. ?Sukeroku Mizutani (L 0103314); in monte 16) Quercus laevigata Blume, Mus. Bot. August 2009 Journal of Japanese Botany Vol. 84 No.4 247 nudata Blume. Bürger Quercus Quercus glauca Thunb. [β] var. s.n. (L 0103310). s.n. (L Fig. 10. Blume [δ] Quercus glandulifera Blume. Siebold s.n. (L 0103290). Blume. Siebold s.n. (L subcrenata var. Fig. 9. Lectotype of 248 植物研究雑誌 第 84 巻 第 4 号 2009 年 8 月 Thunb. [δ] Quercus glauca Blume. Siebold s.n. (L 0103315). stenophylla Blume. Siebold s.n. (L var. Fig. 12. Lectotype of fasciata Thunb. [γ] var. Quercus Quercus glauca Blume. Siebold s.n. (L 0103307). Blume. Siebold s.n. (L Fig. 11. Lectotype of August 2009 Journal of Japanese Botany Vol. 84 No.4 249

1: 304 (1851). crenata Siebold & Zucc., but the veinlets Type: not yet found in L and M. always end at the apex of marginal teeth. 17) Quercus marginata Blume, Mus. Bot. 20b) [γ] var. attenuata Blume, Mus. Bot. 1: 1: 304 (1851). 297 (1851). Type: Japonia. Bürger s.n. (L 0040823* Type: Japonia. Bürger s.n. (L 0040824* –lectotype, designated here). –lectotype, designated here). Current name: Quercus acuta Thunb. Additional original material: Japonia. Quercus marginata falls within the Bürger s.n. (L 0040825). synonymy of Quercus acuta Thunb. The Current name: Carruth. margins of the leaves of Quercus acuta are Blume separated var. attenuata from usually entire, but rarely have small teeth Quercus acutissima var. acutissima by the apically or are shallowly undulate as in the narrow leaves and often attenuate leaf base. lectotype of Quercus marginata. This is only an extreme example of the 18) Quercus myrsinifolia Blume, Mus. variation in the leaves of Quercus acutissima. Bot. 1: 305 (1851). 20c) [δ] var. nana Blume, Mus. Bot. Type: Siebold (ex Keisuke 321), with name 1: 297 (1851). “Hosobagasi” and “tyo” (in Chinese letter) s.n. Type: Japonia. Bürger s.n. (L 0040827* (L 0175956*–lectotype, designated here). –lectotype, designated here). Current name: Quercus myrsinifolia Blume. Additional original material: Japonia. 19) Quercus salicina Blume, Mus. Bot. 1: Siebold s.n. (L 0040828). 305 (1851). Current name: Quercus acutissima Carruth. Type: Japonia. Bürger (? ex Keisuke) s.n. 20d) [ε] var. obtusata Blume, Mus. Bot. 1: (L 0040842–lectotype, designated here). 297 ([1850], actually 1851). [Fig. 14] Type: Japonia. Bürger s.n. (L 0040830 Additional original material: Japonia. –lectotype, designated here). [Fig. 15] Siebold (actually Bürger) s.n. (L 0040840*); Additional original material: Japonia. ?Bürger s.n. (L 0040843); ?Bürger Siebold s.n. (L 0040829*). (L 0040841). Current name: Quercus acutissima Carruth. Current name: Quercus salicina Blume. Siebold s.n. (L 0040829) is more similar 20) Quercus serrata Thunb. to var. nana than var. obtusata, and probably Blume considered Thunberg’s Quercus Blume mistakenly wrote that name. serrata to be Quercus acutissima Carruth. All 21) Quercus sessilifolia Blume, Mus. Bot. varieties described under Quercus serrata are 1: 305 (1851). H. Hara in J. Jap. Bot. 33: 146 synonyms of Quercus acutissima. (1958). 20a) [β] var. tanbakuri Blume, Mus. Bot. Quercus paucidentata Franch. ex Nakai in 1: 297 (1851). Bot. Mag. (Tokyo) 40: 583 (1926). Type: Japonia. Bürger s.n. (L 0040826* Cyclobalanopsis sessilifolia (Blume) –lectotype, designated here). Schottky in Bot. Jahrb. Syst. 47: 652 (1912). Current name: Quercus acutissima Carruth. Nakai in J. Jap. Bot. 15: 202 (1939), as C. Blume cited a Japanese name “Tanbakuri” sessilifolia (Blume) Nakai. as “fide Herb. Bürger,” but this specimen lacks Type: Japonia. Siebold (ex Keiske 84 indication of the Japanese name. Additional (deleted) ) s.n. (L 0040844*–lectotype, specimens might still be found. The shape designated here). of the leaves is similar to those of Castanea Current name: Quercus sessiliflora Blume. 250 植物研究雑誌 第 84 巻 第 4 号 2009 年 8 月 Blume. Bürger (? ex Keisuke) s.n. (L 0040842). (? ex Keisuke) s.n. (L Blume. Bürger salicina Fig. 14. Quercus Blume. Siebold (ex Keisuke) s.n. Quercus lacera (L 0103319). (L Fig. 13. August 2009 Journal of Japanese Botany Vol. 84 No.4 251 Blume. Siebold s.n. (L 0103327). sieboldiana Blume. Siebold s.n. (L Fig. 16. Quercus Blume. obtusata Thunb. [ε] var. Quercus serrata Bürger s.n. (L 0040830). s.n. (L Bürger Fig. 15. 252 植物研究雑誌 第 84 巻 第 4 号 2009 年 8 月 Blume. Siebold (ex Sukeroku

Quercus variabilis Mizutani) s.n. (L 0103333, lower). Mizutani) s.n. (L Fig. 18. Blume. Siebold s.n. (L 0040845). Blume. Siebold s.n. (L urticifolia Fig. 17. Quercus August 2009 Journal of Japanese Botany Vol. 84 No.4 253

Nakai (1926, 1939) wrote that the type Type: Japonia. Siebold (ex Sukeroku specimen of Quercus sessilifolia is in the Mizutani) s.n. (L 0103333 lower–lectotype, Rijksherbarium at Leiden, but did not designated here). [Fig. 18] designate a lectotype. Hara (1958) later wrote Additional original material: Siebold (ex that he examined the type at Leiden in 1954 ?Keiske 271) (L 0103333, upper). and annotated the sheet indicating that it was Current name: Quercus variabilis Blume. the type of Quercus sessilifolia, but he also did not designate a lectotype. We greatly appreciate Dr. Hans- 22) Quercus sieboldiana Blume, Mus. Bot. Joachim Esser, Botanisohe Staatssammlung 1: 290 (1851). München. Thanks also due to Dr. David E. Type: Japonia. Siebold s.n., with name Bouford, the Harvard University Herbaria, for ‘Kashi’ (in Japanese) (L 0103327–lectotype, critically reading the manuscript. This study designated here). [Fig. 16] was partly supported by Grants-in-Aid for Additional original material: Japonia. Scientific Research of the Japan Society for Siebold s.n. (L 0103326); Bürger s.n. the Promotion of Science (nos. 16401036; (L 0103328); Bürger s.n. (L 0103325). 20300286 to Dr. Tokuhei Tagai) and the Japan Current name:Lithocarpus glabra (Thunb.) Foundation (Japanese Studies and Intellectual Nakai. Exchange in 2006 to H.O.). 23）Quercus urticifolia Blume, Mus. Bot. 1: 296 (1851), ut urticaefolia. References Type: Japonia. Siebold s.n. (L 0040845 Blume C. L. (1850) 1851. Ord. Cupulifereae. In: Blume, –lectotype, designated here). [Fig. 17] Museum Botanicum Lugduno-Batavi 1: 282–310, E. J. Hara annotated this specimen as the Brill, Lugduni-Batavorum (now Leiden). de Kort I., Ridder-Numan J, Thijsse G., Kato N. and lectotype, but did not publish his choice. Yamaguchi T. 2000. Von Siebold’s Botanical Treasures Additional original material: Siebold, in Leiden, CD-ROM with a Booklet (18 pp.), ETI/NHN, with “No. 3, Quercus konara Variet. ß,” Amsterdam. (L 0103331, 0103330 [duplicate of 0103331]); Hara H. 1958. Critical notes on some type specimens of Siebold s.n. (L 0103329). East-Asiatic in foreign herbaria. J. Jpn. Bot. 33: 146–152. Current name: Quercus serrata Murray. McNeil J. (Chairman). 2006. International code of Botanical In Siebold No. 3 the scales of the cupule Nomenclature (Vienna Code). A. R. G. Gautner Verlog are narrowly subulate and similar to those KG, Ruggell (Liechtenstein). of Quercus dentata and Quercus acutissima. Nakai T. 1926. Notulae ad plantas Japoniae & Koreae Such cupules with narrowly subulate scales XXXIII. Bot. Mag. (Tokyo) 40: 1–627. Nakai T. 1939. Castanopsis, Pasania and their allies in the are rarely found in Quercus serrata or Quercus Japanese Empire. I. J. Jap. Bot. 15: 185–204 (in Japanese crispula, they are thought to be the result of with Latin diagnosis). teratological atavism. Ohba H. 1989. Fagaceae. In: Satake Y., Hara H., Watari S. 23a) [β] var.: foliis brevius petiolatis basi and Tominari T. (eds.), Wild Flowers of Japan, Woody acutis v. obtusis. Plants. 1: 66–78, Heibonsha, Tokyo (in Japanese with This “variety” is based on Siebold s.n. scientific names). Ohba H. 2006. Fagaceae. In: Iwatsuki K., Boufford D. E. and with “Kumaside” and “Quercus Konara” (L Ohba H. (eds.), Flora of Japan. 2a: 42–60. Kodansha, 0103332). Tokyo. Current name: Quercus serrata Murray. Ohba H., Akiyama S. and Thijsse G. 2005. Miquel's new taxa 24) Quercus variabilis Blume, Mus. Bot. of the vascular plants described from Japan in Prolusio 1: 297 (1851). Florae Japonicae and some other works. Univ. Mus. 254 植物研究雑誌 第 84 巻 第 4 号 2009 年 8 月

univ. Tokyo Bull. 41: 31–140. Bürger and von Siebold’s collaborators, and now housed Yamaguchi T, Fujii S, Tamura M. N., Nagamasu H., Kato in the National Herbarium of the Netherlands and the N., Wada H. and Yamamoto A. 2003. The specimens Botanische Staatssammlung München. Calanus Special of Japanese plants collected by Ph. F. von Siebold, H. No, V, 191–576.

大場秀章 a，秋山 忍 b，G. タイセ c：Blume が 1851 年に日本から記載したブナ科の種及び変 種のレクトタイプと原資料 シーボルト・コレクションから Blume が記載 Blume は， 日 本 か ら の Siebold，Bürger， に用いた標本（原資料）を特定した．16 種 27 Pierot，Textor，Mohnike，Keiske（伊藤圭介） 変種については，記載に最もよく合致する標本 らの採集品からなるシーボルト・コレクション を選定し，レクトタイプに指定した．しかし， 中のブナ科を研究し，その結果を 1851 年に論 記載された分類群のうち，5 種 1 変種について 文にまとめた．そのなかで多数の新種や新変種 は該当する標本を見出すことができなかった． が発表されたが，記載ではまったくタイプの指 （a 東京大学総合研究博物館， 定がなされなかった．学名の安定化などの目的 b 国立科学博物館植物研究部， で，ライデンならびにミュンヘンに収蔵される c オランダ・国立植物標本館ライデン大学分館）