©Wiener Coleopterologenverein (WCV), download unter www.biologiezentrum.at

Koleopterologische Rundschau 78 264 Wien, Juli 2008 Koleopterologische Rundschau 78 265–274 Wien, Juli 2008

Buchbesprechung Pelthydrus ORCHYMONT, 1919 from () with LÖBL, I. & SMETANA, A. (Hrsg.) 2007: Catalogue of Palaearctic Coleoptera, Band 4. – Stenstrup: Apollo Books, 935 pp. Preis: DKK (Dänische Kronen) 1080.- (excl. Versand) descriptions of three new species LÖBL, I. & SMETANA, A. (Hrsg.) 2008: Catalogue of Palaearctic Coleoptera, Band 5. – Stenstrup: Apollo Books, 670 pp. Preis: DKK (Dänische Kronen) 780.- (excl. Versand) (Coleoptera: Hydrophilidae)

Die beiden neuesten Bände dieser Serie umfassen die Elateroidea, Derodontoidea, Bostrichoidea, Lyme- D. BIAN, H. SCHÖNMANN & L. JI xyloidea, Cleroidea, Cucujoidea (Band 4) sowie die Tenebrionoidea (Band 5). Am vorderen Einband und im Vorspann findet sich je eine Zeichnung des Endomychiden Eucteanus coelestinus (GERSTAECKER, 1857) – im Abbildungstext jedoch irrtümlich als “Eutectanus” bezeichnet (Bd. 4) und des Ripiphoriden Ptilophorus fallax IABLOKOFF-KHNZORIAN, 1973 (Bd. 5). Abstract Die beiden Bände sind berühmten europäischen Koleopterologen gewidmet: Karel Majer (1949–2000) bzw. Gustav Adolf Lohse (1910–1994). Six species of the genus Pelthydrus ORCHYMONT, 1919 (Coleoptera: Hydrophilidae) are recorded from Hainan Island (China). Three of these species (Pelthydrus angulatus sp.n., P. tongi sp.n., and P. In Band 4 werden 53 Familien mit fast 29.000 Taxa-Namen und 6.905 Primärzitate aufgelistet, welche waltraudae sp.n.) are described herein. The remaining three species (P. incognitus SCHÖNMANN, vor dem Stichtag (1. Jänner 2005) publiziert wurden. Insgesamt 35 Autoren sind für den Inhalt 1995, P. insularis SCHÖNMANN, 1995, and P. minutus ORCHYMONT, 1919) are recorded from Hainan verantwortlich. ĝlipiĔski ist im Autorenverzeichnis irrtümlich zwei mal angeführt. Nur einer der Autoren- for the first time. namen, nämlich F. Steinheimer ist falsch geschrieben, allerdings nicht nur an einer Stelle, sondern im gesamten Band; der Autor der Nosodendridae wird nämlich auf p. 19 als „Steinhammer“ und auf p. 21 als Key words: Coleoptera, Hydrophilidae, Pelthydrus, faunistics, taxonomy, new species, new records, „Steinhamer“ zitiert, seinen richtigen Namen sucht man in diesem Katalog vergeblich. Hainan, China. Band 5 enthält 23 Familien mit mehr als 21.000 Taxa-Namen und 4.880 Primärzitate, welche vor dem Stichtag (1. Jänner 2007 – nicht 2005, wie im Katalog irrtümlich auf Seite 10 angegeben) publiziert wurden. Die Liste der Autoren enthält 27 Namen. Der Autor bei den Tenebrionidae heißt jedoch nicht „Masomuto“ (sowohl in der Autorenliste als auch in der Adressenliste falsch geschrieben) sondern Introduction Masumoto (mit mehr als 200 Publikationen seit 1955 einer der bekanntesten Koleopterologen Japans). Die bei Rudi Schuh angegebene email Adresse (rudolf.schuh@at) wird sehr wahrscheinlich nicht Until today 55 species of the predominantly Oriental genus Pelthydrus ORCHYMONT, 1919 are funktionieren. known. Thirteen of these species occur in China, however, not a single species has so far been Das Artenregister zu den beiden Bänden ist nicht im Buch abgedruckt, sondern nur in der von den recorded from Hainan (GENTILI et al. 1995; HANSEN 1999, 2004; KOMAREK 2003; SCHÖNMANN Herausgebern im Zusammenhang mit taxonomischen Katalogen als höchst ungeeignet empfundenen 1994, 1995). elektronischen Form verfügbar (http://www.apollobooks.com/palaearcticindex.htm und http://www.ville- ge.ch/musinfo/mhng/divers/palstaph-index04.pdf [bzw. „index05.pdf“]). In the present paper we are dealing with the rich collections of Pelthydrus sampled in the course Den „Errata“ zu den Bänden 1–3 (Bd. 4) bzw. 1–4 (Bd. 5) wird breiter Raum gewidmet, insgesamt 17 of the China Water Beetle Survey (see JÄCH 2004, and http://www.nhm-wien.ac.at/Content.Node Seiten. Wobei nach dem Motto: „doppelt hält besser“ manche Korrektur gleich in beiden Bänden zu /forschung/2zoo/china-water-beetle-survey.html) in 1996, and additional specimens collected in finden ist (z.B. der Hinweis, dass Hygrobia hermanni auch für die Schweiz gemeldet wurde). Andere 2007. Three of the six species found on Hainan are new to science. They are described below. Korrekturen sucht man hingegen vergeblich, z.B. dass der Name Kodada (Band 3, p. 60, New replacement names, Stenelmis zhangi) falsch geschrieben wurde. The high number of new species (50 %) corroborates the fact, that Hainan is remarkably rich in Recht umfangreich sind auch wieder die „New nomenclatorial and taxonomic acts“, wobei man sich die endemic species (see also JÄCH & JI 1998a). Frage stellen sollte, wie sinnvoll die Verfügbarmachung von phylogenetisch nicht abgesicherten Gat- tungsnamen (z.B. Paulusiella Mandl) ist. Meiner Meinung nach hätte man in solchen Fällen die jeweiligen Spezialisten mit einbeziehen sollen. Eine Verfügbarmachung aus rein formalen Gründen ist Material and methods taxonomisch nicht vertretbar und nicht im Sinne der Nomenklaturregeln. Band 5 enthält gegenüber den früheren Bänden eine geographische Neueinführung. Der politischen Ei- Specimens were examined with an Olympus SZX16 and a LEICA–WILD M10 compound micro- genständigkeit von Montenegro Rechnung tragend wurde das Kürzel ME bei den Verbreitungsangaben scope. The body length was measured from the front margin of the labrum to the elytral apex. hinzugefügt. Male genitalia were placed in concentrated lactic acid several hours before they were examined. Obwohl schon mehrfach darauf hingewiesen, findet sich in den beiden neuen Bänden immer noch eine Aedeagi were drawn with the aid of a drawing tube. Habitus images have been captured with a unrichtige geographische Bezeichnung, nämlich Guandong statt . „Guandong“ bedeutet in Leica DFC 490 attached to a Leica MZ16 stereoscopic microscope using Automontage for chinesischer Sprache „östlich des Passes“ und bezeichnet die früher als Mandschurei bekannte Region mit stacking. Final post-processing was done in Adobe Photoshop CS. den Provinzen , und . Der Name „Guangdong“ hingegen heißt übersetzt soviel wie „Weiter Osten“. Es ist die Nachbarprovinz von (= „Weiter Westen“). Guangdong ist die bevölkerungsreichste Provinz Chinas und befindet sich im Gegensatz zu Guandong nicht im äußersten Abbreviations: Nordosten, sondern im Südosten des Landes. CWBS China Water Beetle Survey IAECAS Institute of Applied Ecology, Chinese Academy of Sciences, M.A. Jäch NMW Naturhistorisches Museum Wien ©Wiener Coleopterologenverein (WCV), download unter www.biologiezentrum.at

266 Koleopt. Rdsch. 78 (2008) BIAN et al.: Pelthydrus from Hainan (China) (HYDROPHILIDAE) 267

List of CWBS localities (leg. M.A. Jäch, L. Ji & M. Wang 1996) CWBS loc. 207: Hainan Province; Ledong County; Jianfeng Mountains; Jianfeng Forest Reserve, ca. 500 m NE Tian Chi Village; small streamlet, ca. 1 m wide, flowing through degraded primary CWBS loc. 182: Hainan Province; City Region; Baishiling Scenic Spot, ca. 10 km SW of forest, and deep, man-made pool, ca. 800 m a.s.l.; 22.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. Qionghai City; small stream, ca. 0.5 m wide, slowly flowing through rubber plantation, granite, sand and gravel, shaded, 20 m a.s.l.; 14.I.1996; leg. L. Ji & M. Wang. CWBS loc. 208: Hainan Province; Ledong County; Jianfeng Mountains; ca. 5 km E Tian Chi Village; two small streams, ca. 3 m wide, flowing through degraded primary forest and shrubs, ca. 800 m CWBS loc. 184: Hainan Province; Qionghai City Region; Baishiling Scenic Spot, ca. 20 km SW of a.s.l.; 23.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. Qionghai City; very small stream, slowly flowing to CWBS loc. 182; 0.3 m wide, granite, stones and gravel, shaded, 20 m a.s.l.; 14.I.1996; leg. L. Ji & M. Wang. CWBS loc. 213: Hainan Province; Ledong County; Jianfeng Town, Botanical Garden; residual pools of dried up stream, ca. 1 m wide, ca. 60 m a.s.l.; 24.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. CWBS loc. 188: Hainan Province; Qiongzhong County; Baihua Ling (= Hundred Flowers Mountain), ca. 7 km W Qiongzhong City (= Yinggen); river, ca. 5 m wide, flowing through degraded forest, CWBS loc. 214: Hainan Province; County; ca. 15 km SW Dongxing Town, ca. 1 km W large boulders, sand, cataracts, ca. 300 m a.s.l.; 16.I.1996; leg. M.A. Jäch, L. Ji & M. Wang (see Jianfeng Village; small stream, ca. 1–2 m wide, probably a tributary of CWBS loc. 215, meandering, water turbid, flowing through cultivated land (rubber plantations, bamboo groves, JÄCH & JI 1998b: Fig. 2). villages, etc.), ca. 70 m a.s.l.; 25.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. CWBS loc. 189: Hainan Province; Qiongzhong County; small tributary to CWBS loc. 188, ca. 1 m wide, densely shaded; 16.I.1996; leg. L. Ji. CWBS loc. 215: Hainan Province; Wanning County; ca. 500 m W CWBS loc. 214; river, ca. 5 m wide, with sand and gravel, flowing through cultivated land, ca. 70 m a.s.l.; 25.I.1996; leg. M.A. Jäch, CWBS loc. 191: Hainan Province; Qiongzhong County; 4 km W Qiongzhong City; ca. 150 m a.s.l., L. Ji & M. Wang. small stream ca. 0.5 m wide, partly shaded by shrubs; 16.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. CWBS loc. 216: Hainan Province; Wanning County; ca. 8–10 km W Dongxing Town; meandering river, crossing the Dongxing – Jianfeng road three times, ca. 3–6 m wide, slightly turbid, with sand and CWBS loc. 192: Hainan Province; Qiongzhong County; Maoyang Town; Changhua River, ca. 40 m gravel, flowing through cultivated land (rubber plantations, village gardens, rice fields, bamboo wide (river bed ca. 80 m wide), potamal, banks with gravel and sand, ca. 100 m a.s.l.; 17.I.1996; groves), probably being the lower course of CWBS loc. 215; ca. 70 m a.s.l.; 26.I.1996; leg. M.A. leg. M.A. Jäch, L. Ji & M. Wang (see JÄCH & JI 1998b: Fig. 4). Jäch, L. Ji & M. Wang (see JÄCH & JI 1998b: Fig. 11). CWBS loc. 193: Hainan Province; Qiongzhong County; Wuzhi Shan (= Five Finger Mountain) Resort, ca. 2 km from Wuzhi Shan Village, ca. 30 km E Maoyang Town; small stream, ca. 2–3 m wide, Check list of Pelthydrus species from Hainan shaded, flowing through degraded primary forest, below Wuzhi Shan Resort, ca. 600 m a.s.l.; 17./18.I.1996; leg. M.A. Jäch, L. Ji & M. Wang (see JÄCH & JI 1998b: Fig. 3). Pelthydrus (Globipelthydrus) jendeki group CWBS loc. 194: Hainan Province; Qiongzhong County; stream, ca. 5 m wide, densely shaded, flowing 1. Pelthydrus (Globipelthydrus) waltraudae sp.n. partly through deep gorge, through primary forest, above Wuzhi Shan Resort; ca. 700–800 m a.s.l.; 18.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. Pelthydrus (s.str.) inaspectus group CWBS loc. 197: Hainan Province; Qiongzhong County; ca. 3 km E Maoyang; Wuzhi Shan River, ca. 20 2. Pelthydrus (s.str.) incognitus SCHÖNMANN, 1995 m wide, fast flowing, with big boulders, flowing through secondary forest and pastures, ca. 200 m Pelthydrus (s.str.) minutus group a.s.l.; 19.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. 3. Pelthydrus (s.str.) angulatus sp.n. CWBS loc. 198: Hainan Province; Qiongzhong County; 3 km NE Maoyang; Wuzhi Shan River, ca. 30 m wide, slowly flowing through wide, deforested valley, only few riffles, banks with sand, grass 4. Pelthydrus (s.str.) insularis SCHÖNMANN, 1995 and mud, ca. 100 m a.s.l.; 19.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. 5. Pelthydrus (s.str.) minutus ORCHYMONT, 1919 CWBS loc. 199: Hainan Province; Tongza City Region; ca. 2 km S Mao’an; river, ca. 5 m wide, banks Pelthydrus (s.str.) sculpturatus group with shrubs, grass, sand, mud, rock pools and seepage water, ca. 250 m a.s.l.; 20.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. 6. Pelthydrus (s.str.) tongi sp.n. CWBS loc. 202: Hainan Province; Ledong County; E Jianfeng Town; several streams (including residual pools), 1–5 m wide, slowly flowing through rice fields and shrubs, only partly shaded, Pelthydrus (Globipelthydrus) waltraudae sp.n. ca. 60 m a.s.l.; 21.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. CWBS loc. 204: Hainan Province; Ledong County; foot of Jianfeng (= Sharp Peak) Mountain, ca. 4 km TYPE LOCALITY: CHINA: Hainan Province; ; Maoyang Town, 18°55'N, E Jianfeng Town; river, ca. 5–10 m wide, fast flowing, through secondary forest, with big 109°28'E, 255 m a.s.l.; small stream, ca. 0.5 m wide, unpolluted, sand, shatters, bank with sand, boulders, margins with rock pools, springfed pools and some seepage water, ca. 150 m a.s.l.; shrubs and grass. 22./24.I.1996; leg. M.A. Jäch, L. Ji & M. Wang (see JI & JÄCH 1998b: Fig. 11). TYPE MATERIAL: Holotype (IAECAS): “CHINA: Hainan Province, Wuzhishan City Maoyang Town, 18°55'N, CWBS loc. 205: Hainan Province; Ledong County; Jianfeng Mountains; Jianfeng Forest Reserve, ca. 5 109°28'E 255 m, 25.11.2007, leg. D. Bian & Y. Tong”. Paratypes: 1 , 2 (IAECAS): same data as holotype; 5 exs. km NE Tian Chi Village; river, ca. 5 m wide, flowing through very dense primary forest, banks (IAECAS): “CHINA: Hainan Province, Lingshui County Diaoluo National Natural Reserve NE [of] Diaoluo with stones and sand, ca. 800 m a.s.l.; 22.I.1996; leg. M.A. Jäch, L. Ji & M. Wang (see JÄCH & JI Holiday Village, 18°44'N 109°52'E 920 m, 16.11.2007, leg. D. Bian, Y. Tong & M. Wang”; 3 exs. (NMW): 1998b: Figs. 7, 9). “CHINA: Hainan (188) 7 km W Qiongzhong Baihua Ling, 300 m 16.1.1996, Ji & Wang”; 1 ex. (NMW): “CHINA: Hainan (188) 7 km W Qiongzhong Baihua Ling, 300 m 16.1.1996, leg. Jäch”; 1 ex. (NMW): “CHINA: Hainan (215) 1.5 km SW Dongxing 1.5 km W Jianfeng, 70 m 25.1.1996, leg. Jäch”. ©Wiener Coleopterologenverein (WCV), download unter www.biologiezentrum.at

266 Koleopt. Rdsch. 78 (2008) BIAN et al.: Pelthydrus from Hainan (China) (HYDROPHILIDAE) 267

List of CWBS localities (leg. M.A. Jäch, L. Ji & M. Wang 1996) CWBS loc. 207: Hainan Province; Ledong County; Jianfeng Mountains; Jianfeng Forest Reserve, ca. 500 m NE Tian Chi Village; small streamlet, ca. 1 m wide, flowing through degraded primary CWBS loc. 182: Hainan Province; Qionghai City Region; Baishiling Scenic Spot, ca. 10 km SW of forest, and deep, man-made pool, ca. 800 m a.s.l.; 22.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. Qionghai City; small stream, ca. 0.5 m wide, slowly flowing through rubber plantation, granite, sand and gravel, shaded, 20 m a.s.l.; 14.I.1996; leg. L. Ji & M. Wang. CWBS loc. 208: Hainan Province; Ledong County; Jianfeng Mountains; ca. 5 km E Tian Chi Village; two small streams, ca. 3 m wide, flowing through degraded primary forest and shrubs, ca. 800 m CWBS loc. 184: Hainan Province; Qionghai City Region; Baishiling Scenic Spot, ca. 20 km SW of a.s.l.; 23.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. Qionghai City; very small stream, slowly flowing to CWBS loc. 182; 0.3 m wide, granite, stones and gravel, shaded, 20 m a.s.l.; 14.I.1996; leg. L. Ji & M. Wang. CWBS loc. 213: Hainan Province; Ledong County; Jianfeng Town, Botanical Garden; residual pools of dried up stream, ca. 1 m wide, ca. 60 m a.s.l.; 24.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. CWBS loc. 188: Hainan Province; Qiongzhong County; Baihua Ling (= Hundred Flowers Mountain), ca. 7 km W Qiongzhong City (= Yinggen); river, ca. 5 m wide, flowing through degraded forest, CWBS loc. 214: Hainan Province; Wanning County; ca. 15 km SW Dongxing Town, ca. 1 km W large boulders, sand, cataracts, ca. 300 m a.s.l.; 16.I.1996; leg. M.A. Jäch, L. Ji & M. Wang (see Jianfeng Village; small stream, ca. 1–2 m wide, probably a tributary of CWBS loc. 215, meandering, water turbid, flowing through cultivated land (rubber plantations, bamboo groves, JÄCH & JI 1998b: Fig. 2). villages, etc.), ca. 70 m a.s.l.; 25.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. CWBS loc. 189: Hainan Province; Qiongzhong County; small tributary to CWBS loc. 188, ca. 1 m wide, densely shaded; 16.I.1996; leg. L. Ji. CWBS loc. 215: Hainan Province; Wanning County; ca. 500 m W CWBS loc. 214; river, ca. 5 m wide, with sand and gravel, flowing through cultivated land, ca. 70 m a.s.l.; 25.I.1996; leg. M.A. Jäch, CWBS loc. 191: Hainan Province; Qiongzhong County; 4 km W Qiongzhong City; ca. 150 m a.s.l., L. Ji & M. Wang. small stream ca. 0.5 m wide, partly shaded by shrubs; 16.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. CWBS loc. 216: Hainan Province; Wanning County; ca. 8–10 km W Dongxing Town; meandering river, crossing the Dongxing – Jianfeng road three times, ca. 3–6 m wide, slightly turbid, with sand and CWBS loc. 192: Hainan Province; Qiongzhong County; Maoyang Town; Changhua River, ca. 40 m gravel, flowing through cultivated land (rubber plantations, village gardens, rice fields, bamboo wide (river bed ca. 80 m wide), potamal, banks with gravel and sand, ca. 100 m a.s.l.; 17.I.1996; groves), probably being the lower course of CWBS loc. 215; ca. 70 m a.s.l.; 26.I.1996; leg. M.A. leg. M.A. Jäch, L. Ji & M. Wang (see JÄCH & JI 1998b: Fig. 4). Jäch, L. Ji & M. Wang (see JÄCH & JI 1998b: Fig. 11). CWBS loc. 193: Hainan Province; Qiongzhong County; Wuzhi Shan (= Five Finger Mountain) Resort, ca. 2 km from Wuzhi Shan Village, ca. 30 km E Maoyang Town; small stream, ca. 2–3 m wide, Check list of Pelthydrus species from Hainan shaded, flowing through degraded primary forest, below Wuzhi Shan Resort, ca. 600 m a.s.l.; 17./18.I.1996; leg. M.A. Jäch, L. Ji & M. Wang (see JÄCH & JI 1998b: Fig. 3). Pelthydrus (Globipelthydrus) jendeki group CWBS loc. 194: Hainan Province; Qiongzhong County; stream, ca. 5 m wide, densely shaded, flowing 1. Pelthydrus (Globipelthydrus) waltraudae sp.n. partly through deep gorge, through primary forest, above Wuzhi Shan Resort; ca. 700–800 m a.s.l.; 18.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. Pelthydrus (s.str.) inaspectus group CWBS loc. 197: Hainan Province; Qiongzhong County; ca. 3 km E Maoyang; Wuzhi Shan River, ca. 20 2. Pelthydrus (s.str.) incognitus SCHÖNMANN, 1995 m wide, fast flowing, with big boulders, flowing through secondary forest and pastures, ca. 200 m Pelthydrus (s.str.) minutus group a.s.l.; 19.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. 3. Pelthydrus (s.str.) angulatus sp.n. CWBS loc. 198: Hainan Province; Qiongzhong County; 3 km NE Maoyang; Wuzhi Shan River, ca. 30 m wide, slowly flowing through wide, deforested valley, only few riffles, banks with sand, grass 4. Pelthydrus (s.str.) insularis SCHÖNMANN, 1995 and mud, ca. 100 m a.s.l.; 19.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. 5. Pelthydrus (s.str.) minutus ORCHYMONT, 1919 CWBS loc. 199: Hainan Province; Tongza City Region; ca. 2 km S Mao’an; river, ca. 5 m wide, banks Pelthydrus (s.str.) sculpturatus group with shrubs, grass, sand, mud, rock pools and seepage water, ca. 250 m a.s.l.; 20.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. 6. Pelthydrus (s.str.) tongi sp.n. CWBS loc. 202: Hainan Province; Ledong County; E Jianfeng Town; several streams (including residual pools), 1–5 m wide, slowly flowing through rice fields and shrubs, only partly shaded, Pelthydrus (Globipelthydrus) waltraudae sp.n. ca. 60 m a.s.l.; 21.I.1996; leg. M.A. Jäch, L. Ji & M. Wang. CWBS loc. 204: Hainan Province; Ledong County; foot of Jianfeng (= Sharp Peak) Mountain, ca. 4 km TYPE LOCALITY: CHINA: Hainan Province; Wuzhishan City; Maoyang Town, 18°55'N, E Jianfeng Town; river, ca. 5–10 m wide, fast flowing, through secondary forest, with big 109°28'E, 255 m a.s.l.; small stream, ca. 0.5 m wide, unpolluted, sand, shatters, bank with sand, boulders, margins with rock pools, springfed pools and some seepage water, ca. 150 m a.s.l.; shrubs and grass. 22./24.I.1996; leg. M.A. Jäch, L. Ji & M. Wang (see JI & JÄCH 1998b: Fig. 11). TYPE MATERIAL: Holotype (IAECAS): “CHINA: Hainan Province, Wuzhishan City Maoyang Town, 18°55'N, CWBS loc. 205: Hainan Province; Ledong County; Jianfeng Mountains; Jianfeng Forest Reserve, ca. 5 109°28'E 255 m, 25.11.2007, leg. D. Bian & Y. Tong”. Paratypes: 1 , 2 (IAECAS): same data as holotype; 5 exs. km NE Tian Chi Village; river, ca. 5 m wide, flowing through very dense primary forest, banks (IAECAS): “CHINA: Hainan Province, Lingshui County Diaoluo National Natural Reserve NE [of] Diaoluo with stones and sand, ca. 800 m a.s.l.; 22.I.1996; leg. M.A. Jäch, L. Ji & M. Wang (see JÄCH & JI Holiday Village, 18°44'N 109°52'E 920 m, 16.11.2007, leg. D. Bian, Y. Tong & M. Wang”; 3 exs. (NMW): 1998b: Figs. 7, 9). “CHINA: Hainan (188) 7 km W Qiongzhong Baihua Ling, 300 m 16.1.1996, Ji & Wang”; 1 ex. (NMW): “CHINA: Hainan (188) 7 km W Qiongzhong Baihua Ling, 300 m 16.1.1996, leg. Jäch”; 1 ex. (NMW): “CHINA: Hainan (215) 1.5 km SW Dongxing 1.5 km W Jianfeng, 70 m 25.1.1996, leg. Jäch”. ©Wiener Coleopterologenverein (WCV), download unter www.biologiezentrum.at

268 Koleopt. Rdsch. 78 (2008) BIAN et al.: Pelthydrus from Hainan (China) (HYDROPHILIDAE) 269

DIAGNOSIS: Habitus as in Fig. 1. Body length 3.0–3.5 mm; body width 1.9–2.2 mm. Hainan Province, Lingshui County Diaoluo National Natural Reserve NE Diaoluo Holiday Village, 18°44'N Externally, very similar to P. dudgeoni SCHÖNMANN, 1995. Body strongly convex, broadly oval 109°52'E, 920 m, 16.11.2007, leg. D. Bian, Y. Tong & M. Wang”; 3 exs. (NMW): “CHINA: Hainan (182) 10 km and acute at elytral apex, upper surface shining, darkish brown to black, lateral margins of pro- SW Qionghai, 20 m Baishiling, 14.1.1996 leg. Ji & Wang”; 2 exs. (NMW): “CHINA: Hainan (191) 4 km W Qiongzhong 150 m, 16.1.1996 leg. Ji”; 2 exs. (NMW): “CHINA: Hainan (191) 4 km W Qiongzhong 150 m, notum and elytra as well as legs reddish brown, antennae and maxillary palpi yellowish brown. 16.1.1996 leg. Jäch”; 1 ex. (NMW): “CHINA: Hainan (198) 3 km NE Maoyang Wuzhi Shan Riv., 100 m 19.1.1996, Head trapezoidal, about 1.8 times as wide as long, distinctly narrower than pronotum; covered Ji & Wang”; 1 ex. (NMW): “CHINA: Hainan (205) Jianfeng Mts., 800 m 5 km NE Tian Chi 22.1.1996, leg. Jäch”; 1 with deeply impressed punctures of two different sizes, appearing wrinkled along inner margins ex. (NMW): “CHINA: Hainan (215) 15 km SW Dongxing 1.5 km W Jianfeng, 70 m 25.1.1996, Ji & Wang”. of eyes, clypeal suture distinctly impressed; anterior margin of clypeus concave, anterior corners DIAGNOSIS: Habitus as in Fig. 2. Body length 1.9–2.2 mm; body width 0.9–1.1 mm. External- angulate; maxillary palpi distinctly longer than antennae, as long as width of head. ly, very similar to P. minutus. Body sub-ovoid, moderately convex, somewhat acute posteriorly, Shape of pronotum trapezoidal, about 2.4 times as wide as long; lateral sides narrowed distally, elytral apices separately rounded; surface shining dark brown to black; lateral margins of head, ridged, anterior part obscurely dentate; punctures of two different sizes, with denser arrangement pronotum, elytra and legs reddish brown; antennae, labial and maxillary palpi yellowish brown. than on head; posterior corners broadly rounded and shining transparent. Head rectangular, about 1.6 times as wide as long; densely punctate; lateral sides of head and Elytra broadest at basal fifth, slightly narrowed posteriorly towards acute apex; ten distinct striae inner margins of eyes appearing wrinkled; clypeal suture slightly impressed; anterior margin of consisting of bigger punctures; intervals not raised, densely covered with small punctures. clypeus concave, anterior corners distinctly angulate. Underside reddish brown, covered with fine pubescence; mentum somewhat rugulose; pro- Pronotum trapezoidal, about 2.5 times as wide as long, punctation similar to that on head; lateral sternum with medio-longitudinal carina; mesoventrite strongly elevated, producing a triangular margins ridged, anterior corners angulate, posterior corners broadly rounded. plate; metaventrite highly raised, with a ́-shaped projection anteriorly; hind margin of fifth Elytra about 1.4 times as long as wide, lateral margins smoothly ridged; broadest part at two sternite somewhat truncate in males, with a distinct mesal emargination in females. basal fifths, from there slightly narrowed anteriorly and distinctly narrowed posteriorly, Aedeagus (Fig. 4): 1.2 mm long, slender, outline nearly parallel-sided; phallobasis approximately terminating in two separately rounded elytral apices; punctation similar to that of head and pro- as long as median lobe; median lobe strongly narrowed from midlength towards base, broadest notum, consisting of equally distributed, completely unordered punctures. subapically, with a sharply pointed apex; parameres gradually narrowed from base to apices, Ventral side mostly black; mentum rugulosely microreticulate, anterior tenth shining and strongly narrowed and medially arched in distal third, encircling the broadest part of median sparsely punctate; prosternum with a medio-longitudinal carina; mesoventrite elevated lobe. posteriorly, produced into a triangular plate pointing towards anterior part; metaventrite progres- DIFFERENTIAL DIAGNOSIS: This species is a member of the Pelthydrus jendeki species sively elevated towards posterior part of plate, with a cordiform, shining, and glabrous area group. It can easily be distinguished from the other members of the P. jendeki group by the posteriorly; first sternite with a medio-longitudinal carina not reaching hind margin. combination of the following characters: head and pronotum with double sized punctation; elytra Aedeagus (Fig. 5): 0.6 mm long, median lobe abruptly narrowed at base, then parallel-sided and with ten punctate striae, intervals not elevated and densely covered with small punctures; slender, corona in apical position; parameres nearly parallel-sided from base to level of apex of aedeagus longer and much more slender than in P. dudgeoni, its shape more narrow, elongated median lobe, then abruptly narrowed and sharply angled laterad extending into acute apices. and parallel-sided than in all other species of the group. DIFFERENTIAL DIAGNOSIS: Pelthydrus angulatus is a member of the P. minutus species DISTRIBUTION: Hainan. CWBS locs. 188, 215. Diaoluo National Natural Reserve (Fig. 7). group. Body shape and body surface similar to P. minutus but easily to be distinguished by DERIVATIO NOMINIS: Waltraude Neiszer-Kühnberger runs the small restaurant “Zum separately rounded elytral apices and mainly by the unique shape of the parameres. Schwarzen Mohren” not far from the Vienna Natural History Museum. She is very interested in DISTRIBUTION: Hainan. CWBS locs. 182, 191, 198, 205, 215. Diaoluo National Natural various aspects of biology and nature protection. Since more than twenty years she warmly Reserve (Fig. 7). welcomes coleopterists from all over the world, especially at lunch time. After serving “all you DERIVATIO NOMINIS: angulatus (Latin for “angled”) in reference to the typical shape of the can eat” lunch, she usually invites coleopterists to have coffee, beer or wine “on the house”. apical part of the parameres. In the name of all beetle lovers who had the opportunity to enjoy her hearty hospitality we want to express our warmest thanks by dedicating this little roundish species to her. Pelthydrus (s.str.) incognitus SCHÖNMANN Pelthydrus incognitus SCHÖNMANN 1995: 139. – GENTILI et al. 1995: 214. – HANSEN 1999: 128. – HANSEN 2004: Pelthydrus (s.str.) angulatus sp.n. 59. ADDITIONAL MATERIAL EXAMINED: 1 ex. (IAECAS): CHINA: Hainan Province, Changjiang County; TYPE LOCALITY: CHINA: Hainan Province; Wuzhishan City; Maoyang Town, 18°55'N, Bawangling National Natural Reserve, 19°05'N, 109°04'E, 127 m, 23.11.2007, leg D. Bian & Y. Tong; 7 exs. 109°28'E, 255 m a.s.l.; small stream, ca. 0.5 m wide, unpolluted, sand, shatters, bank with sand, (IAECAS): CHINA: Hainan Province, Lingshui County, Diaoluo National Natural Reserve, NE Diaoluo Holiday shrubs and grass. Village, 18°44'N, 109°52'E, 920 m, 16.11.2007, leg. D. Bian, Y. Tong & M. Wang. TYPE MATERIAL: Holotype (IAECAS): “CHINA: Hainan Province, Wuzhishan City Maoyang Town, 18°55'N, DISTRIBUTION: This species is widely distributed from northern Thailand to southern China. 109°28'E 255 m, 25.11.2007, leg D. Bian & Y. Tong”. Paratypes: 4 , 7 (IAECAS): same data as holotype; 4 exs. (IAECAS): “CHINA: Hainan Province, Ledong County Jianfeng Town, Jianfeng Mountains, Jianfeng Forest It is here recorded from Hainan for the first time. CWBS locs. 188, 193, 194, 207. Diaoluo Reserve, 18°44'N, 108°52'E 785 m, 21.11.2007, leg. D. Bian, Y. Tong & M. Wang”; 1 ex. (IAECAS): “CHINA: National Natural Reserve (Fig. 7), Bawangling National Natural Reserve. ©Wiener Coleopterologenverein (WCV), download unter www.biologiezentrum.at

268 Koleopt. Rdsch. 78 (2008) BIAN et al.: Pelthydrus from Hainan (China) (HYDROPHILIDAE) 269

DIAGNOSIS: Habitus as in Fig. 1. Body length 3.0–3.5 mm; body width 1.9–2.2 mm. Hainan Province, Lingshui County Diaoluo National Natural Reserve NE Diaoluo Holiday Village, 18°44'N Externally, very similar to P. dudgeoni SCHÖNMANN, 1995. Body strongly convex, broadly oval 109°52'E, 920 m, 16.11.2007, leg. D. Bian, Y. Tong & M. Wang”; 3 exs. (NMW): “CHINA: Hainan (182) 10 km and acute at elytral apex, upper surface shining, darkish brown to black, lateral margins of pro- SW Qionghai, 20 m Baishiling, 14.1.1996 leg. Ji & Wang”; 2 exs. (NMW): “CHINA: Hainan (191) 4 km W Qiongzhong 150 m, 16.1.1996 leg. Ji”; 2 exs. (NMW): “CHINA: Hainan (191) 4 km W Qiongzhong 150 m, notum and elytra as well as legs reddish brown, antennae and maxillary palpi yellowish brown. 16.1.1996 leg. Jäch”; 1 ex. (NMW): “CHINA: Hainan (198) 3 km NE Maoyang Wuzhi Shan Riv., 100 m 19.1.1996, Head trapezoidal, about 1.8 times as wide as long, distinctly narrower than pronotum; covered Ji & Wang”; 1 ex. (NMW): “CHINA: Hainan (205) Jianfeng Mts., 800 m 5 km NE Tian Chi 22.1.1996, leg. Jäch”; 1 with deeply impressed punctures of two different sizes, appearing wrinkled along inner margins ex. (NMW): “CHINA: Hainan (215) 15 km SW Dongxing 1.5 km W Jianfeng, 70 m 25.1.1996, Ji & Wang”. of eyes, clypeal suture distinctly impressed; anterior margin of clypeus concave, anterior corners DIAGNOSIS: Habitus as in Fig. 2. Body length 1.9–2.2 mm; body width 0.9–1.1 mm. External- angulate; maxillary palpi distinctly longer than antennae, as long as width of head. ly, very similar to P. minutus. Body sub-ovoid, moderately convex, somewhat acute posteriorly, Shape of pronotum trapezoidal, about 2.4 times as wide as long; lateral sides narrowed distally, elytral apices separately rounded; surface shining dark brown to black; lateral margins of head, ridged, anterior part obscurely dentate; punctures of two different sizes, with denser arrangement pronotum, elytra and legs reddish brown; antennae, labial and maxillary palpi yellowish brown. than on head; posterior corners broadly rounded and shining transparent. Head rectangular, about 1.6 times as wide as long; densely punctate; lateral sides of head and Elytra broadest at basal fifth, slightly narrowed posteriorly towards acute apex; ten distinct striae inner margins of eyes appearing wrinkled; clypeal suture slightly impressed; anterior margin of consisting of bigger punctures; intervals not raised, densely covered with small punctures. clypeus concave, anterior corners distinctly angulate. Underside reddish brown, covered with fine pubescence; mentum somewhat rugulose; pro- Pronotum trapezoidal, about 2.5 times as wide as long, punctation similar to that on head; lateral sternum with medio-longitudinal carina; mesoventrite strongly elevated, producing a triangular margins ridged, anterior corners angulate, posterior corners broadly rounded. plate; metaventrite highly raised, with a ́-shaped projection anteriorly; hind margin of fifth Elytra about 1.4 times as long as wide, lateral margins smoothly ridged; broadest part at two sternite somewhat truncate in males, with a distinct mesal emargination in females. basal fifths, from there slightly narrowed anteriorly and distinctly narrowed posteriorly, Aedeagus (Fig. 4): 1.2 mm long, slender, outline nearly parallel-sided; phallobasis approximately terminating in two separately rounded elytral apices; punctation similar to that of head and pro- as long as median lobe; median lobe strongly narrowed from midlength towards base, broadest notum, consisting of equally distributed, completely unordered punctures. subapically, with a sharply pointed apex; parameres gradually narrowed from base to apices, Ventral side mostly black; mentum rugulosely microreticulate, anterior tenth shining and strongly narrowed and medially arched in distal third, encircling the broadest part of median sparsely punctate; prosternum with a medio-longitudinal carina; mesoventrite elevated lobe. posteriorly, produced into a triangular plate pointing towards anterior part; metaventrite progres- DIFFERENTIAL DIAGNOSIS: This species is a member of the Pelthydrus jendeki species sively elevated towards posterior part of plate, with a cordiform, shining, and glabrous area group. It can easily be distinguished from the other members of the P. jendeki group by the posteriorly; first sternite with a medio-longitudinal carina not reaching hind margin. combination of the following characters: head and pronotum with double sized punctation; elytra Aedeagus (Fig. 5): 0.6 mm long, median lobe abruptly narrowed at base, then parallel-sided and with ten punctate striae, intervals not elevated and densely covered with small punctures; slender, corona in apical position; parameres nearly parallel-sided from base to level of apex of aedeagus longer and much more slender than in P. dudgeoni, its shape more narrow, elongated median lobe, then abruptly narrowed and sharply angled laterad extending into acute apices. and parallel-sided than in all other species of the group. DIFFERENTIAL DIAGNOSIS: Pelthydrus angulatus is a member of the P. minutus species DISTRIBUTION: Hainan. CWBS locs. 188, 215. Diaoluo National Natural Reserve (Fig. 7). group. Body shape and body surface similar to P. minutus but easily to be distinguished by DERIVATIO NOMINIS: Waltraude Neiszer-Kühnberger runs the small restaurant “Zum separately rounded elytral apices and mainly by the unique shape of the parameres. Schwarzen Mohren” not far from the Vienna Natural History Museum. She is very interested in DISTRIBUTION: Hainan. CWBS locs. 182, 191, 198, 205, 215. Diaoluo National Natural various aspects of biology and nature protection. Since more than twenty years she warmly Reserve (Fig. 7). welcomes coleopterists from all over the world, especially at lunch time. After serving “all you DERIVATIO NOMINIS: angulatus (Latin for “angled”) in reference to the typical shape of the can eat” lunch, she usually invites coleopterists to have coffee, beer or wine “on the house”. apical part of the parameres. In the name of all beetle lovers who had the opportunity to enjoy her hearty hospitality we want to express our warmest thanks by dedicating this little roundish species to her. Pelthydrus (s.str.) incognitus SCHÖNMANN Pelthydrus incognitus SCHÖNMANN 1995: 139. – GENTILI et al. 1995: 214. – HANSEN 1999: 128. – HANSEN 2004: Pelthydrus (s.str.) angulatus sp.n. 59. ADDITIONAL MATERIAL EXAMINED: 1 ex. (IAECAS): CHINA: Hainan Province, Changjiang County; TYPE LOCALITY: CHINA: Hainan Province; Wuzhishan City; Maoyang Town, 18°55'N, Bawangling National Natural Reserve, 19°05'N, 109°04'E, 127 m, 23.11.2007, leg D. Bian & Y. Tong; 7 exs. 109°28'E, 255 m a.s.l.; small stream, ca. 0.5 m wide, unpolluted, sand, shatters, bank with sand, (IAECAS): CHINA: Hainan Province, Lingshui County, Diaoluo National Natural Reserve, NE Diaoluo Holiday shrubs and grass. Village, 18°44'N, 109°52'E, 920 m, 16.11.2007, leg. D. Bian, Y. Tong & M. Wang. TYPE MATERIAL: Holotype (IAECAS): “CHINA: Hainan Province, Wuzhishan City Maoyang Town, 18°55'N, DISTRIBUTION: This species is widely distributed from northern Thailand to southern China. 109°28'E 255 m, 25.11.2007, leg D. Bian & Y. Tong”. Paratypes: 4 , 7 (IAECAS): same data as holotype; 4 exs. (IAECAS): “CHINA: Hainan Province, Ledong County Jianfeng Town, Jianfeng Mountains, Jianfeng Forest It is here recorded from Hainan for the first time. CWBS locs. 188, 193, 194, 207. Diaoluo Reserve, 18°44'N, 108°52'E 785 m, 21.11.2007, leg. D. Bian, Y. Tong & M. Wang”; 1 ex. (IAECAS): “CHINA: National Natural Reserve (Fig. 7), Bawangling National Natural Reserve. ©Wiener Coleopterologenverein (WCV), download unter www.biologiezentrum.at

270 Koleopt. Rdsch. 78 (2008) BIAN et al.: Pelthydrus from Hainan (China) (HYDROPHILIDAE) 271

Figs. 4–6: Aedeagus, dorsal view; 4) Pelthydrus waltraudae sp.n.; 5) P. angulatus sp.n.; 6) P. tongi sp.n.

Pelthydrus (s.str.) insularis SCHÖNMANN Pelthydrus insularis SCHÖNMANN 1995: 129–130. – GENTILI et al. 1995: 214. – HANSEN 1999: 128. – HANSEN 2004: 59. DISTRIBUTION: Described from Lantau Island (), this species also occurs in Guangxi, and is now recorded from Hainan for the first time. CWBS locs. 188, 189, 191, 193, 194, 198, 199, 204, 208, 215.

Pelthydrus (s.str.) minutus ORCHYMONT Pelthydrus minutus ORCHYMONT 1919: 153. – ORCHYMONT 1932: 695–697. – SATÔ 1960: 80. – SCHÖNMANN 1995: 128–129. – GENTILI et al. 1995: 214. – HANSEN 1999: 128. – HANSEN 2004: 59. DISTRIBUTION: This species is most common and widely distributed in the Oriental Region. It is here recorded from Hainan for the first time. CWBS locs. 188, 192–194, 197–199, 215, 216.

Pelthydrus (s.str.) tongi sp.n.

TYPE LOCALITY: CHINA: Hainan Province; Wuzhishan City; Maoyang Town, 18°55'N, Figs.1–3: Habitus, dorsal view; 1) Pelthydrus waltraudae sp.n.; 2) P. angulatus sp.n.; 3) P. tongi sp.n. 109°28'E, 255 m a.s.l.; small stream, ca. 0.5 m wide, unpolluted, sand, shatters, bank with sand, shrubs and grass. ©Wiener Coleopterologenverein (WCV), download unter www.biologiezentrum.at

270 Koleopt. Rdsch. 78 (2008) BIAN et al.: Pelthydrus from Hainan (China) (HYDROPHILIDAE) 271

Figs. 4–6: Aedeagus, dorsal view; 4) Pelthydrus waltraudae sp.n.; 5) P. angulatus sp.n.; 6) P. tongi sp.n.

Pelthydrus (s.str.) insularis SCHÖNMANN Pelthydrus insularis SCHÖNMANN 1995: 129–130. – GENTILI et al. 1995: 214. – HANSEN 1999: 128. – HANSEN 2004: 59. DISTRIBUTION: Described from Lantau Island (Hong Kong), this species also occurs in Guangxi, and is now recorded from Hainan for the first time. CWBS locs. 188, 189, 191, 193, 194, 198, 199, 204, 208, 215.

Pelthydrus (s.str.) minutus ORCHYMONT Pelthydrus minutus ORCHYMONT 1919: 153. – ORCHYMONT 1932: 695–697. – SATÔ 1960: 80. – SCHÖNMANN 1995: 128–129. – GENTILI et al. 1995: 214. – HANSEN 1999: 128. – HANSEN 2004: 59. DISTRIBUTION: This species is most common and widely distributed in the Oriental Region. It is here recorded from Hainan for the first time. CWBS locs. 188, 192–194, 197–199, 215, 216.

Pelthydrus (s.str.) tongi sp.n.

TYPE LOCALITY: CHINA: Hainan Province; Wuzhishan City; Maoyang Town, 18°55'N, Figs.1–3: Habitus, dorsal view; 1) Pelthydrus waltraudae sp.n.; 2) P. angulatus sp.n.; 3) P. tongi sp.n. 109°28'E, 255 m a.s.l.; small stream, ca. 0.5 m wide, unpolluted, sand, shatters, bank with sand, shrubs and grass. ©Wiener Coleopterologenverein (WCV), download unter www.biologiezentrum.at

272 Koleopt. Rdsch. 78 (2008) BIAN et al.: Pelthydrus from Hainan (China) (HYDROPHILIDAE) 273

TYPE MATERIAL: Holotype (IAECAS): “CHINA: Hainan Province, Wuzhishan City Maoyang Town, 18°55'N, 109°28'E 255 m, 25.11.2007, leg Y. Tong”. Paratypes: 6 , 8 (IAECAS): same data as holotype; 12 exs. (IAECAS): “CHINA: Hainan Province, Qionghai City Nanfeng Town, 19°10'N, 110°12'E, 110 m 15.11.2007, leg. D. Bian, Y. Tong & M. Wang”; 3 exs. (NMW): “CHINA: Hainan (182) 10 km SW Qionghai, 20 m, Baishiling, 14.1.1996 leg. Ji & Wang”; 1 ex. (NMW): “CHINA: Hainan (184) 20 km SW Qionghai, 20 m Baishiling, 14.1.1996 leg. Ji & Wang”; 1 ex. (NMW): “CHINA: Hainan (191) 4 km W Qiongzhong 150 m, 16.1.1996 leg. Jäch”; 1 ex. (NMW): “CHINA: Hainan (198) 3 km NE Maoyang Wuzhi Shan Riv., 100 m 19.1.1996, Ji & Wang”; 2 exs. (NMW): “CHINA: Hainan (199) 2 km S Mao’an 250 m, 20.1.1996 leg. Ji & Wang”; 5 exs. (NMW): “CHINA: Hainan (202) E Jianfeng 60 m, 21.1.1996 leg. Jäch”; 1 ex. (NMW): “CHINA: Hainan (204) 4 km E Jianfeng, 150 m Jianfeng Mt., 1996 22./24.1., Ji & Wang”; 1 ex. (NMW): “CHINA: Hainan (213) Jianfeng, 700 m Bot. Garden 24.1.1996, leg. Jäch”; 38 exs. (NMW): “CHINA: Hainan (214) 15 km SW Dongxing 1 km W Jianfeng, 70 m 25.1.1996, Ji & Wang”; 21 exs. (NMW): “CHINA: Hainan (214) 15 km SW Dongxing 1 km W Jianfeng, 70 m 25.1.1996, leg. Jäch”; 4 exs. (NMW): “CHINA: Hainan (215) 15 km SW Dongxing 1.5 km W Jianfeng, 70 m 25.1.1996, Ji & Wang”; 11 exs. (NMW): CHINA: “Hainan (215) 15 km SW Dongxing 1.5 km W Jianfeng, 70 m 25.1.1996, leg. Jäch”; 4 exs. (NMW): “CHINA: Hainan (216) 8-10 km W Dongxing 70 m, 26.1.1996 leg. Jäch”. DIAGNOSIS: Habitus as in Fig. 3. Body length 2.4–2.8 mm; body width 1.3–1.5 mm. Externally, very similar to P. siamensis SCHÖNMANN, 1995. Body elongate, ovoid and strongly convex; gradually narrowed in posterior part with a broadly rounded elytral apex. Shining dark brown with yellowish brown margins; legs, maxillary palpi and anterior part of antennae yellowish brown, last three antennomeres dark brown. Head rectangular, about 1.5 times as wide as long, distinctly narrower than pronotum, surface microreticulate between sparsely distributed large punctures; inner margins of eyes wrinkled, clypeal suture slightly impressed; clypeus with distinctly angulate anterior corners. Pronotum rectangular, about 2.3 times as wide as long; lateral margins reddish brown, ridged and finely serrated, straight and nearly parallel-sided; punctation larger and denser than on head, interspaces with a distinct mesh-like microreticulation. Elytra broadest at basal three tenths, about 1.2 times as long as wide; lateral margins reddish Fig. 7: Stream in Diaoluo National Natural Reserve, Hainan, China. Habitat of Pelthydrus angulatus, brown and ridged, finely serrated in distal half and rather strongly reflexed towards broadly P. incognitus and P. waltraudae. Photograph by D. Bian. rounded apex; punctation similar to that on pronotum, but smooth and shining between punctures. Underside mostly covered with white pubescence; mentum microreticulate with anterior margin Acknowledgements shining and sparsely punctate; prosternum reddish brown with a median longitudinal We want to express our gratitude to Dr. Manfred A. Jäch and Dr. Albrecht Komarek (NMW) for progressively elevated carina; mesoventrite strongly elevated posteriorly forming a triangular helpful comments and suggestions on our manuscript. This study was supported by The plate with a median longitudinal carina; metaventrite black with a lumpish protuberance, Botanical Gardens and Biological Taxonomy Program of the Chinese Academy of Sciences medially with a ́-shaped portion projecting between the mesocoxae and the rhombic shining (KSCX2–YW–Z–037), by National Key Technology R & D Program (2006BAD03A0901), by area situated posteriorly; first sternite with a median longitudinal carina, which almost reaches The Ministry of Science and Technology of the People’s Republic of China (2006FY110500–3), the hind margin. and partly by The Open Fund of Liaoning Key Laboratory of Non–commercial Forest Manage- Aedeagus (Fig. 6): 0.5 mm long, moderately wide; phallobasis approximately as long as median ment. Additionally, our warmest thanks must be submitted to the “CWBS-Project” and its lobe and parameres; median lobe broadest basally, gradually narrowed towards broadly rounded proponents having collected most of the specimens mentioned above. Our thanks also go to Dr. apex; parameres gradually narrowed from base towards straight apices. Yanfeng Tong for providing further specimens collected in Hainan and to Harald Schillhammer DIFFERENTIAL DIAGNOSIS: This species is a member of the Pelthydrus sculpturatus species for the habitus photographs of the new species. group. Body shape and body surface similar to P. siamensis, but aedeagus resembles P. vitalisi ORCHYMONT, 1926, from which it can be distinguished by its more slender shape of the median References lobe, which is a bit longer than the parameres and by the equally narrowed parameres, which are not arched. GENTILI, E., HEBAUER, F., JÄCH, M.A., JI, L. & SCHÖDL, S. 1995: Hydrophilidae: 1. Check list of the Hydrophilinae recorded from China (Coleoptera), pp. 207–219. – In: Jäch, M.A. & Ji, L. (eds.), DISTRIBUTION: Hainan. CWBS locs. 182, 184, 191, 198, 199, 202, 204, 213–216. Nanfeng. Water Beetles of China, Vol. I. – Wien: Zoologisch–Botanische Gesellschaft in Österreich and DERIVATIO NOMINIS: This species is dedicated to Dr. Yanfeng Tong, who recently has Wiener Coleopterologenverein, 410 pp collected a large number of the type specimens. ©Wiener Coleopterologenverein (WCV), download unter www.biologiezentrum.at

272 Koleopt. Rdsch. 78 (2008) BIAN et al.: Pelthydrus from Hainan (China) (HYDROPHILIDAE) 273

TYPE MATERIAL: Holotype (IAECAS): “CHINA: Hainan Province, Wuzhishan City Maoyang Town, 18°55'N, 109°28'E 255 m, 25.11.2007, leg Y. Tong”. Paratypes: 6 , 8 (IAECAS): same data as holotype; 12 exs. (IAECAS): “CHINA: Hainan Province, Qionghai City Nanfeng Town, 19°10'N, 110°12'E, 110 m 15.11.2007, leg. D. Bian, Y. Tong & M. Wang”; 3 exs. (NMW): “CHINA: Hainan (182) 10 km SW Qionghai, 20 m, Baishiling, 14.1.1996 leg. Ji & Wang”; 1 ex. (NMW): “CHINA: Hainan (184) 20 km SW Qionghai, 20 m Baishiling, 14.1.1996 leg. Ji & Wang”; 1 ex. (NMW): “CHINA: Hainan (191) 4 km W Qiongzhong 150 m, 16.1.1996 leg. Jäch”; 1 ex. (NMW): “CHINA: Hainan (198) 3 km NE Maoyang Wuzhi Shan Riv., 100 m 19.1.1996, Ji & Wang”; 2 exs. (NMW): “CHINA: Hainan (199) 2 km S Mao’an 250 m, 20.1.1996 leg. Ji & Wang”; 5 exs. (NMW): “CHINA: Hainan (202) E Jianfeng 60 m, 21.1.1996 leg. Jäch”; 1 ex. (NMW): “CHINA: Hainan (204) 4 km E Jianfeng, 150 m Jianfeng Mt., 1996 22./24.1., Ji & Wang”; 1 ex. (NMW): “CHINA: Hainan (213) Jianfeng, 700 m Bot. Garden 24.1.1996, leg. Jäch”; 38 exs. (NMW): “CHINA: Hainan (214) 15 km SW Dongxing 1 km W Jianfeng, 70 m 25.1.1996, Ji & Wang”; 21 exs. (NMW): “CHINA: Hainan (214) 15 km SW Dongxing 1 km W Jianfeng, 70 m 25.1.1996, leg. Jäch”; 4 exs. (NMW): “CHINA: Hainan (215) 15 km SW Dongxing 1.5 km W Jianfeng, 70 m 25.1.1996, Ji & Wang”; 11 exs. (NMW): CHINA: “Hainan (215) 15 km SW Dongxing 1.5 km W Jianfeng, 70 m 25.1.1996, leg. Jäch”; 4 exs. (NMW): “CHINA: Hainan (216) 8-10 km W Dongxing 70 m, 26.1.1996 leg. Jäch”. DIAGNOSIS: Habitus as in Fig. 3. Body length 2.4–2.8 mm; body width 1.3–1.5 mm. Externally, very similar to P. siamensis SCHÖNMANN, 1995. Body elongate, ovoid and strongly convex; gradually narrowed in posterior part with a broadly rounded elytral apex. Shining dark brown with yellowish brown margins; legs, maxillary palpi and anterior part of antennae yellowish brown, last three antennomeres dark brown. Head rectangular, about 1.5 times as wide as long, distinctly narrower than pronotum, surface microreticulate between sparsely distributed large punctures; inner margins of eyes wrinkled, clypeal suture slightly impressed; clypeus with distinctly angulate anterior corners. Pronotum rectangular, about 2.3 times as wide as long; lateral margins reddish brown, ridged and finely serrated, straight and nearly parallel-sided; punctation larger and denser than on head, interspaces with a distinct mesh-like microreticulation. Elytra broadest at basal three tenths, about 1.2 times as long as wide; lateral margins reddish Fig. 7: Stream in Diaoluo National Natural Reserve, Hainan, China. Habitat of Pelthydrus angulatus, brown and ridged, finely serrated in distal half and rather strongly reflexed towards broadly P. incognitus and P. waltraudae. Photograph by D. Bian. rounded apex; punctation similar to that on pronotum, but smooth and shining between punctures. Underside mostly covered with white pubescence; mentum microreticulate with anterior margin Acknowledgements shining and sparsely punctate; prosternum reddish brown with a median longitudinal We want to express our gratitude to Dr. Manfred A. Jäch and Dr. Albrecht Komarek (NMW) for progressively elevated carina; mesoventrite strongly elevated posteriorly forming a triangular helpful comments and suggestions on our manuscript. This study was supported by The plate with a median longitudinal carina; metaventrite black with a lumpish protuberance, Botanical Gardens and Biological Taxonomy Program of the Chinese Academy of Sciences medially with a ́-shaped portion projecting between the mesocoxae and the rhombic shining (KSCX2–YW–Z–037), by National Key Technology R & D Program (2006BAD03A0901), by area situated posteriorly; first sternite with a median longitudinal carina, which almost reaches The Ministry of Science and Technology of the People’s Republic of China (2006FY110500–3), the hind margin. and partly by The Open Fund of Liaoning Key Laboratory of Non–commercial Forest Manage- Aedeagus (Fig. 6): 0.5 mm long, moderately wide; phallobasis approximately as long as median ment. Additionally, our warmest thanks must be submitted to the “CWBS-Project” and its lobe and parameres; median lobe broadest basally, gradually narrowed towards broadly rounded proponents having collected most of the specimens mentioned above. Our thanks also go to Dr. apex; parameres gradually narrowed from base towards straight apices. Yanfeng Tong for providing further specimens collected in Hainan and to Harald Schillhammer DIFFERENTIAL DIAGNOSIS: This species is a member of the Pelthydrus sculpturatus species for the habitus photographs of the new species. group. Body shape and body surface similar to P. siamensis, but aedeagus resembles P. vitalisi ORCHYMONT, 1926, from which it can be distinguished by its more slender shape of the median References lobe, which is a bit longer than the parameres and by the equally narrowed parameres, which are not arched. GENTILI, E., HEBAUER, F., JÄCH, M.A., JI, L. & SCHÖDL, S. 1995: Hydrophilidae: 1. Check list of the Hydrophilinae recorded from China (Coleoptera), pp. 207–219. – In: Jäch, M.A. & Ji, L. (eds.), DISTRIBUTION: Hainan. CWBS locs. 182, 184, 191, 198, 199, 202, 204, 213–216. Nanfeng. Water Beetles of China, Vol. I. – Wien: Zoologisch–Botanische Gesellschaft in Österreich and DERIVATIO NOMINIS: This species is dedicated to Dr. Yanfeng Tong, who recently has Wiener Coleopterologenverein, 410 pp collected a large number of the type specimens. ©Wiener Coleopterologenverein (WCV), download unter www.biologiezentrum.at

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HANSEN, M. 1999: Hydrophiloidea (s.str.) (Coleoptera). – In: World Catalogue of Insects, Volume 2. – Stenstrup: Apollo Books, 416 pp. HANSEN, M. 2004: Hydrophilidae, pp. 44–68. – In: Löbl, I. & Smetana, A. (eds.): Catalogue of Palae- Zelenkaesalus subgen.n. for three Southeast Asian species of arctic Coleoptera, Vol. 2. – Stenstrup: Apollo Books, 942 pp. Echinoaesalus ZELENKA, including E. javanus sp.n. JÄCH, M.A. & JI, L. 1998a: Foreword, p. I. – In: Jäch, M.A. & Ji, L. (eds.), Water Beetles of China, Vol. II. – Wien: Zoologisch–Botanische Gesellschaft in Österreich and Wiener Coleopterologenverein, (Coleoptera: Lucanidae: Aesalinae) II + pp. 371. JÄCH, M.A. & JI, L. 1998b: China Water Beetle Survey (1995–1998), pp. 1–23. – In: Jäch, M.A. & Ji, L. J. KRIKKEN (eds.): Water Beetles of China, Vol. II. – Wien: Zoologisch-Botanische Gesellschaft in Österreich and Wiener Coleopterologenverein, II + pp. 371. JÄCH, M.A. 2004: The China Water Beetle Survey (CWBS) – a biodiversity project of the superlative th celebrates its 10 anniversary. – Procupine! 30: 5–7. Abstract KOMAREK, A. 2003: Hydrophilidae: I. Check list and key to Palearctic and Oriental genera of aquatic Aesaline stag beetles of the genus Echinoaesalus ZELENKA, 1993 (Coleoptera: Lucanidae) were Hydrophilidae (Coleoptera), pp. 383–395. – In: Jäch, M.A. & Ji, L. (eds.), Water Beetles of hitherto recorded from Peninsular Malaysia/Thailand, Borneo, Sumatra, and Sulawesi. In this paper a China, Vol. III. – Wien: Zoologisch–Botanische Gesellschaft in Österreich and Wiener Coleo- new species from Java, Echinoaesalus javanus, is described and illustrated. It is closely related with pterologenverein, VI + 572 pp. two named Southeast Asian species with deep metapectoral and abdominal grooves accommodating the middle and hind legs. A new subgenus, Zelenkaesalus (type-species Aesalus timidus KRIKKEN, ORCHYMONT, A. d’ 1919: Contribution à l’étude des sous-familles des Sphaeridiinae et des Hydrophil- 1975), is established for these three species. The broader taxonomic and biogeographic context of the inae (Coleoptera: Hydrophilidae). – Annales de la Société Entomologique de France LXXXVIII: Aesalini is briefly discussed. 105–168. Key words: Coleoptera, Lucanidae, Aesalini, Echinoaesalus, Zelenkaesalus, new subgenus, key, ORCHYMONT, A. d’ 1932: Zur Kenntnis der Kolbenwasserkäfer (Palpicornia) von Sumatra, Java und Bali. species list, new species, Southeast Asia. – Archiv für Hydrobiologie, Supplement-Band IX (Tropische Binnengewässer II): 623–714, pl. XIV–XVII. SATÔ, M. 1960: One new genus and two new species of the subtribe Helocharae from Japan (Coleoptera: Hydrophilidae). – Transactions of the Shikoku Entomological Society VI: 76–80. Introduction SCHÖNMANN, H. 1994: Revision der Gattung Pelthydrus Orchymont, 1. Teil: Globipelthydrus subgen.n. During the past few decades two new genera and several new species of tropical Aesalini have (Coleoptera: Hydrophilidae). – Koleopterologische Rundschau 64: 189–222. been proposed, following their initial discovery in both Southeast Asia and South America in the SCHÖNMANN, H. 1995: Revision der Gattung Pelthydrus Orchymont, 2. Teil: Pelthydrus s.str. (Coleo- early seventies. The subfamily Aesalinae apparently comprises odd, ancient lineages within the ptera: Hydrophilidae). – Koleopterologische Rundschau 65: 105–144. Lucanidae (HOSOYA & ARAYA 2005, NIKOLAJEV 1993), and the tribe Aesalini was for a long time thought to have a monogeneric Palearctic-Mesoamerican distribution. The first aesaline Dongju BIAN from the Oriental tropics was described from North Sumatra by KRIKKEN (1975), who already Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang, 110016, China ([email protected]) emphasized its differences from the “normal” species of Aesalus FABRICIUS, 1801 then known. Graduate University of Chinese Academy of Sciences, , 100039, China At present the total number of species known from the area between Thailand and Sulawesi,

Dr. Heinrich SCHÖNMANN meanwhile all referred to Echinoaesalus ZELENKA, 1993, has risen to nine (see check list below), Naturhistorisches Museum, Burgring 7, A – 1010 Wien, Austria ([email protected]) including records from most major islands, except Java. There are no records from the Philippines and the area east of Sulawesi. Further new Aesalini from mainland Eurasia have been Prof. Dr. Lanzhu JI placed in Aesalus (see, for instance, BARTOLOZZI 1991, ARAYA et al. 1998, ARAYA & Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang, 110016, China ([email protected]) YOSHITOMI 2003), and more aesaline novelties are anticipated. Recently, after having been beyond reach for two decades, a small, diverse lot of Asian aesalines came back to Leiden, including a specimen from West Java collected there about 75 years ago – to my knowledge the first aesaline to be recorded from the island. This specimen, from the well- known volcano Tangkubanperahu, north of the city of Bandung, certainly represents a new species of the group characterized by deep, sharply defined ventral sulci, which includes two more Sundaland species, Echinoaesalus timidus (KRIKKEN, 1975) and E. sabahensis ZELENKA, 1994. The metapectoral and abdominal sulci, apparently adapted to receive and protect the meso- and metatibiae and tarsi, evidently constitute a synapomorphy, and, consequently, a new subgenus is proposed for this conspicuous set of three species. In this paper the new subgenus is briefly characterized, the three included species are keyed, and the new species from Java is described; the genus is also briefly characterized. For more synoptic work on Southeast Asian