ARTICLE 155 et al. 673U 5 Phthirusa Phthirusa et al !""4 2 $ `SQ* ! Key words: Basidiomycota > Pucciniaceae Pucciniomycotina Urediniomycetes IMA FUNGUS · 6(1): 155–162 (2015) 155–162 · 6(1): FUNGUS IMA 54 ` G< .5 ! et al.566!V*@Z [ <>/? Uromyces Loranthaceae !""# ? V "7 G 7 _6"7 G ! 47 G ! #5 G ! #5 G 4 [ $ ;G O !z sequencing V % $ / % . !4* M56 GV O ? [ . <>/ $ GV/+ <I<!""6;G 54%[64ZV ?<>/; ; 653U )* 4*!6 !4UUG `@U.#66!WU X U G Z % + Z+ ; [$ !6 . 54/!!6 % K $ !6\ 3* <>/K . %; NV@7. NV@4. ; NV@ >/ 5 % * F/ 566FX I9 !""6 * 6 ! U % < +; ! , , Loranthaceae U U U. , as it Phthirusa urbanianus + O G U. urbanianus U U. nilagiricus Uromyces A. goyazense , Q@ Aecidum goyazense + O ! Loranthaceae loranthi U U. socius U. loranthi, % V Aecidium goyazense , U. bahiensis stelis
UG % ! + O phthirusae U. evastigatus, U , Phthirusa et al. 5664 , V$
[ Uromyces < W N G + Z % + #6"!6."66+
Table 1.]+ Uromyces hawksworthii Species GenBank accession numbers (LSU)* Source and Country Uromyces hawksworthiiZ+55_E5 2 E5!!_" ; + O Uromyces hawksworthiiZ+55E#4 2 E5!!76 ; + O Cumminsiella mirabilissima <_474_! / 566F] ARTICLE Cumminsiella mirabilissima /W75F56F U et al566_] Puccinia coronata <_4745F / 566FZ@/ Puccinia coronata SZE4!!7! et al.56!! Puccinia graminis /W455!## + et al.!""5 Puccinia graminis 97!5F7E < et al.56!! Puccinia hemerocallidis ]Z64E656 < $et al56!6Z@/ Puccinia hemerocallidis <_474!" / 566FZ@/ Puccinia heucherae <_4"#6! 9 et al.566#Z2 Puccinia heucherae <_4"#65 9 et al.566#Z2 Puccinia hordei <_4745# / 566FZ@/ Puccinia melanocephala 2;56!E_E et al.Z 56!7G Puccinia melanocephala 2;56!E_" et alZ 56!7G Puccinia nakanishikii ]Z64E665 < $et al.56!6Z@/ Puccinia peperomiae SZE4!!7F et al.56!!G Uromyces acuminatus ]Z!6"5E5 et al.56!6S Uromyces appendiculatus 2U57"E#6 UV 56!7/ Uromyces appendiculatus /#74#67 U et al.Z 5664 Uromyces ari-triphylli <_4745" / 566FZ@/ Uromyces ixiae <"!##_E U et al.566#@ / Uromyces ixiae <"!##_" U et al.566#@ / Uromyces pisi /W75F56! U et al.566_G S Uromyces striatus 97!5F45 < et al.56!!Q Uromyces striatus 9_!#4!5 * et al.56!4 G Uromyces trifoli ]Z"_FF_7 et al.56!!G Uromyces viciae-fabae 2`#!F_7_ ; IU2O 56!7> Uromyces viciae-fabae /W75F!"" U et al.566_G S Uromyces vignae /+!!4F7" Get al.5667` Melampsora larici-populina `675546 + et al.56!5Z@/ Ê*@Z[ <>/
!V/S Pucciniaceae+ et al.!""5U et al.566_G % O ZX [ O et al.5667/ 566FU et al.566F9 et al. ;G S$@/;.NV&Z@+ 566#U et al.566#et al.56!6< $et al.56!6 ? / + /+N_!_6$U < et al56!! et al.56!!+ et al. 56!5 G Z % + UV 56!7; U2O 56!7* et al. V ? + S 56!4 / ? 9 56!5/? W/@V/ ? [ UZ@G*S&S 5667 US]/ ? > ? %F V et al.56!!/ ]+ [\\ %\ ] \V ! V V +/@S [\\ \ !#FF# + +N PHYLOGENETIC ANALYSIS U % G U G *@Z?+ G? ]+ R +N U +*/@V + +*/@V U UQ
156 IMA FUNGUS 78 9 6 8 6 on Pihthirusa (Loranthaceae)
V . % % Diagnosis[ / Uromyces Phthirusa stelis ARTICLE % Loranthaceae $ ]V N] % !5 V . GN; S@ U et al. 56!6 U + % _5 ? I 9 Type[Brazil: ] @ ; Phthirusa stelis 56!5W UGUG Loranthus~/!E"5Ule 1909 +5"74M K !6666666 V + ] N/ G ; % !666 !6666 + !4 7E)75!5P@&7#:4E)554_P % V[ 5466 . Phthirusa stelis"W 56!7J. C. DianeseZ+UG ; 55E#4Mepitype designated hereU+V56!4_4 . #466 V Melampsora larici-populina Description[ Spermogonia Aecidia 4MF % O W V 566" $ & _66M766 ) \ % 64M! RESULTS !5 K peridial cells _6M36M4# & 5!M22M_4 ) Phylogeny / [ ? *@Z NV@ <>/ Aecidiospores 57M54M5"M_4 & !#M5!M54M5#4 ) % Z+55_E5 Z+55E#4 % K walls 5M_4M74 ) V [ 5E@ <>/ NV@ % Uredinia Telia!M2M_ ? !466 746 %/ > *@Z[Z+55_E52 E5!!_"Z+55E#42 E5!!76 % \ NV@[ Z+55_E52 E5!!_# Z+55E#42 E5!!_E V *@Z NV@ ? . Teliospores _7M V <>/ _"M7_M7F & !EM 55M57 M5F4 ) !. . *@Z [ K @SU %K lateral wall 54M_M7 3 % [ 4MFM#3 . Kpedicels 9et al.56!7VNV@? 7EME7M!7_M!4#&7M4MFM#3 ]+ Z>NVS> et al.56!7+ . *@Z $ \$ 9et al 56!7 _! $ Other specimens examined %Phthirusa stelis[Brazil[ *@Z % V + ] N/ G ; + !6_# "# !EU 56!7J. C. DianeseZ+U55E#"K/ > G %5!! % E!6 % Z % + Z % !#@56!5 E. S. C. de Souza Z+ U 55_E"K ; ? Q
VOLUME 6 · NO. 1 157 Souza et al. ARTICLE
Fig. 1. A–H. Uromyces hawksworthiiZ+UG55E#4[ % A. S B. N C.V D. G % E. / F.; $ G. / H.< % + [/MG5< 463S_663W563]!639!3
158 IMA FUNGUS 78 9 6 8 6 on Pihthirusa (Loranthaceae) ARTICLE
Fig. 2. A–E. Uromyces hawksworthiiZ+UG55E#4[A.G $ B.S C. / % . D.V @SUE.V @SU+ [/!+5G!63<!663S43
Key to Uromyces species on Loranthaceae
! V . _6M74&5!M_63 E3 U. nilagiricus V . 5
5! V 763 3 V 763 #
_5 V U. ornatipes V % 7
7_ V . 4 V % F
47 V U. circumscriptus V %53 U. bahiensis
F7 V [% K U. loranthi V K U. socius
#5 V . 463 E V . "6 !F63!6
VOLUME 6 · NO. 1 159 Souza et al. ARTICLE
Fig. 3. ; + *@Z ? Uromyces $ V + % %6#4 ]+ V + ? <>/? %V Melampsora larici-populina
E# V % [ K % U. phthiruzae V % % % "
"E V % . K % U. urbanianus V % U. evastigatus
!6# V .% K . U. struthanthi V .% K % U. hawksworthii
DISCUSSION ]+ + ]+ *@Z Uromyces hawksworthii ? Puccinia heucherae 9@45"F\64]+ + O Loranthaceae, in <_4"#65KN "Ez!6_F\!6F6U. acuminatus GV.X6E6F5_._ ]+ ]Z!6"5E5K N "E z _4 .[ % !6_4\!64" U. ari-triphylli ZF_# ]+ <_4745"K W U. hawksworthii N "E z !6_7\!64# Puccinia graminis Z.F#7 $ ]+ 97!5F7EKN "Ez!65_\!67E
160 IMA FUNGUS 78 9 6 8 6 on Pihthirusa (Loranthaceae)
P. hordei/WVQ*.N<!765]+ <_4745#KN / `G!"!EUredinales] ARTICLE "Ez!6!#\!67_/ <9 MNNAecidiaceae$ %Puccinia Uromyces hawksworthii $ American Journal of Botany 5[756M77F Uromyces + V< X + @U ]O O < 9 <@ et al. ari-triphylli Puccinia peperomiaeW _ !""5 S% [ 6"E >/?Molecular Phylogenetics U. hawksworthii and Evolution 1[5_!M57! $ / @ + ;S / UG > ] 56!5 W / O Populus angustifolia + <>/ ? > / Fungal Biology 116[#"5ME6! G9V VQ2 U5667U V Uromyces appendiculatus U. Aecidium goyazense9 !E"4 vignae ` Mycoscience 45[5__M577 % V Uromyces < U*/@ /U/U ? UW
VOLUME 6 · NO. 1 161 Souza et al.
U %`U 999 @!""4; / 566" FigTree X !5_ S [ N in Ganoderma S% + Z % S 54@ <>/?Mycologia 87[55_M5_E ? WVU% U ;/ <*< /et al > 992<; ` UV *et al. 56!5 U _5[ [ + 56!7 N % NV@ ? [ Systematic
ARTICLE Fungal Diversity 67[!!M!" Biology 61[4_"M475 ; UU2O S9G56!7/ @9;!"56>% Annales Mycologici 18[ > Myosotidium !47M!F6 G N Phytotaxa 174[55_M5_6 V 2 ; < ; > @ ] > U 2 @ ; .@OQ; U56!7@ Uromyces 56!!U 4[ % Puccinilaes, Basidiomycota Loranthaceae Tropical Plant $ % $ Pathology 39[!7!M!4_ Molecular Biology and Evolution 28[5#_!M ; U @ @ +/ 566! @? % 5#_" \ . X 9 U !""6 [ Uromyces pisi Euphorbia cyparissias Mycological O [ <>/ % Research 105[4#MFF Cryptococcus Journal of Bacteriology 172[75_EM757F ; <+V5667U % 9U /U< $*`G */56!6W [ % / Uromyces acuminatusHonckenya peploides + % Systematic Plant Disease 94[5#" Biology 53[#"_ME6E G + ; U U 56!! W \ V@ 2 !"46/ Uredinales O U XNNNProceedings of the Indian Academy of Science,+ /<> 5E@Revista de Biología Tropical 3[!65M!!6 59[4!#M476
162 IMA FUNGUS