A Revised Checklist with Distribution Maps of the Turtles of the World
John B. Iverson Earlham College
, ; ~ . .; < e e e e e e -e
e e e e e e e e -e e- It e e e e v e e e e e e e e e e e e e e e e e e e e e e e e e e e e e~ e
e
addition to dots markulg IClCalltles, e the latter are ImlDetlIa1:ely adJ:ace:nt indicate "'~ .. ~%,~% e e t:>v<~..""'nl",, Geochelone IntJfwtJlln!1ltp,lv humans e e e e e e e e recent e e e AMG Museum, Somerset Street, Grahamstown 6140, ,"",,-,v..an.v of South Africa.
AMNH American Museum of Natural York e 10024, e Australian Museum, Box 6-8 2000, e ANSP ':'Clem;es, 19th and the lY"r:!runnl PhlJladelphJla, ~J"nn"\lhJJ'!tl"IH 19103, U.S.A, e Cromwell London SW7
California Ac.aOemv of ~Clem;es, Golden San Francisco, California 941 U.S,A
Charleston MlllSel.lln, Ln,arle:ston, South Carolina U,S,A. Estaci6n l:HC'lOl1;lca e Museum of Natural and Lake Shore 60605, e e e e e Kepm)l1C of China. e
e Expos:itlC)fi Blvd., Los .... lIltCllt::S. California e Gallardo 470, 1405 4t e Camb1ridge, Massachusetts Mllseum d'Histoire Naltun~:Ue. Jardin des Plantes, Allee Jules Guesde, 3100 VUIV\j,,,,v. Haute e France e Museum National dHistoire Naturelle, 43 Rue Cuvier, 75231 Paris V, France. MNHN e COJmIJlar~i.tl,re anatomy collection is indicated the suffix MPEG Museu Paraense "Emilio Goeldi", Caixa Postal 399, 66.000 Belem, Para, Brasil. e MRAC Musee de Centrale, B-1980 Tervuren, tlelglilm. e Museo Civico di Storia Naturale de Genova 'Giacoma Doria,' Via 9, 16121 ".H"BUV"'. e MSNG e MTKD Staaliches Museum fUr Tierkunde, AUlgustusstJras:ie 2, 801 Dresden, hPrm:flnv e MVZ Museum of Vertebrate lnj"en:jtv of California, tleJrlCeleV U.s.A. MZf L,G1o!<)glcaJ Museum, UI. Vanemuizhe 43, Tartu, Estonian S.S.R., \..1 ••".':; •.1."\.. e MZUP Museo ZO'O!OglCO, Universita di Palermo, Via Archirafi 18, 90123 !Ja!."'''''''''''
MZUS Musee de LJUUl\.J1t,l''', Universite de Strasl:lOUrg Strastxmrg, France.
MZUSP Museu L.VVn/~"v, Universidade de Sao Paulo, Caixa Postal 7172, 04263 Sao Paulo, Sao Paulo, BrasiL fit NMP Natal Museum, 237 Street, Pi€~teImalritlbl.Jlrg, .1."\"t::IJUIJU\v of South Africa. e NMW Naturhistorisches MUlsellm, Postfach 417, U"rar,,"a A-1014 1, Austria. e NRM Naturhistoriska Riksmuseet, Box 50007, S-104 05 Stockholm, Sweden. e NUH LaboJ~at()rv of Natural Science Collection, National Hanoi, Socialist K.epulJl1C Vietnam. e E'>.lJ[I~1.1Ull[l. OUM Oxford In"""''''olh' Museum, Parks Oxford, Jxtorctsh:lre, United e UUleell1s1,md Museum, Terrace, Fortitude Hnsbcine, OUleellsllmd 4006, Australia. e RCSM of Museum, Lincoln's Inn Fields, London WC2, United E'>.lJHl;I.1UIJH. e RMNH Rijksrnm;eUJm van Histoire, Postbus 9517, 2300 Leiden, Holland. e RSM Scottish Museum, Chambers Street, Edinblun!h EHI UF, Scotland, United l"A,U1t,'lVl,U. e sa Shanxi Institute of Shanxi Province, l''''VU'JU'- of China. e e e e e e e e e e e e e e e - - -e e e e e e e e e e e e e 1 1
were e e e e e e e e e e e e e e e e e e e e e e e e e e e e e e xiii
3
6 7
11
BMNH 1947.3.4.21 and nof''''''I.,,,''· ...."n...... Af'th, T~."tTlf't~.t1 to tlnordliches t-\uSUijum
14
)
J
27
• )
34
42 43
09'
Podocnemis.
rhodesianus 67
70 71 ..~
) 73
75
77
83
85
87 Chelonia olivacea Es(;hs(;hollz
Reevesii soullhe:astern and eastern Asia See overview.
109
Testudo amboinensis Daudin DI~litriblJlti(m: SOtlmlern Asia to Sulawesi Bour
117
'.
137
149
error. UI}SD~eCles: None Reviewed Smith
BMNH "Borneo" Dis1tril)llltion: ,LVJl 157 (jeloer.~yara !Jum~tw!afjra diademata Maracaibo Venezuela None LOlmnlent: Reviewed and elevated to Pritchard and considered it a sut)SVlXIC~S Nicoria nasuta spe<~lml~nS) BMNH 1947 .3.5.54~57 northwestern Ecuador and Carr Almendariz 165 I 170 173 COlllSl(len:~ \:!\.rOfUlvmnn(l with the genus 189 197 Testudo carolina Lmnaelus 204 a " 207 208 213 I 217 «;-Ul 1"""._, 7.2 km north of None l.:OInmlent: Reviewed as Sternotherus /'/U'1 >11'1 """ 221 Ha(;leI110a, Chichen pelllmS11J1a, Mexico Cinosternum leucostomum 9087 232 ..,.. ,.-,..-.1 .."", and Seidel 237 ong,mally C(]ltal()~ut~<1 as USNM 6518 and • • 240 Thailand see 1 UUI'an20. Mexico" 261 "''''nth'''',..... KeplJOllC of South 268 Testudo "nn<"",,,,,,,,,,, Indonesia 269 278 285 • ~ Triorrvdlifl:le account. 301 as at the Islamic as as steindachneri northeastern Testudo TYll,nlOln f"').!:Cca1>c.lL. L. 1857. Contributions to the natural of the United States of America. Vol. L and Co., Boston. Ahl, E. 1932. neuen Schildkrote aus Australien. Ges. Naturf. "'n'Ilr1,np 129. Ahsan, and C. M. a serlrU-(iOfileslllcated eastern tlangladesll. Ahsan, F. and M. A. Saeed. 1989. distribution, historical iJac:kgroumd Alarcon Pardo, H. Anderson, UescrlptJlOn of some new Asiatic mammals and Chelonia. Ann. Natur. Hist London Anderson, J. 1878-79. and researches: Comp,nsmg the two eXloodhtlc)ns to western Yunnan in 1868 and 1 - Andersson, L. G. 1900. Museum in Stockholm. Annandale, N. 1913. The tortoises of Chota Rec. Indian Ammnclale, N. 1915. Rec. Indian Mus. \",.;uvun';AJ 11 Aoki, R. 1977. The occurrence of a short-necked chelid in the Palau Islands [in J(t~mnt:::;CI. J. l-l..rn~.t£'\1 7(2):32- 33. Arnold, E. N. 1979. Indian Ocean tortoises: Their sy~;[ernallCS and Philos. Soc. London 286B: Arnold, E. N. and J. A. Burton. 1978. A field to the and amphlol;ans of Britain and Collins, referred to {ieocllel(me (Shaw). Misc. Note 11, J. Checklist of fossil land tortoises Bull. Florida State Mus. Pt. 1. UsteoJo~;y and relatll)m>nll)S of extant and R. Franz. 1978a. and Franz. Cat. Amer. r\1l1IJIIlV. 1-2. and Franz. AuHeniJe:rg, W. and R. Franz. 1978d. Cat. 214: 1-2. and 1978e. Cat. Amer. f>lIllJ',UV. 1982. The status and distribution of the tortoise 12:95-126. and conservation status of the tortoise, rSI'lmmC'lJ(l'!(!S Doctoral Dissertation, Univ. of ~te~Uenb()scjll, Baard. E. H. W. 1991. A review of the taxonomic ge()metIJlC tortoise, Psammobates (H>/H'lLJ't"',~lH' \1...JllUl."",U<>, Babcock, H. L. 1937. Dec. Boston Soc. Baird, S. F. and C. Girard. 1852. lJe:SCIllptl,ons of new of collected under the command of Charles Wilkes, Pmc. Acad. Natur. Barbour, T. 1914a. On some Australasian l"'""'IJUj''''''. Proc. BioI. Soc. Barbour, T. 1914b. Contribution to the zoclgeC)gr;apl1:y the West Indies. Mem. Mus. Harvard. Barbour, T. 1935. Bahamas. Occ. 8:205-206. Barbour, T. and F. Carr. 1938. Another Bahamian fresh-water tortoise. Proc. New En1~larld Zooi. Club 17:75-76. Barbour, T. and A. F. Carr. 1940. Antillean tenrapins. Mem. Mus. Zoot, Harvard 54:381-415. Barbour. T. and F. Carr. 1941. from Prec. New 1...JHi5Hl1,IU Zoot 1929. and amphl,bHllns. Bull. Mus. Harvard Batsch, J. C. 1788. Versuch einer Atlileltun:g zur Kenntniss und Geschichte und Mineralien. Vol.l. Akademische lena. p. G. 1888a. Notes on the American Tnon'vchldae. Amer. Natur. 121-22. Baur, G. 1888b. ZooL Baur, G. 1888c. Usteolo~~lS(;ne Notizen tiber l'~IJUljl"H. Baur, G. 1888d. Baur, G. syslenrmtlsclle VI'''''U'''U~ von 180-191. Baur, G. ....,.,~u ••_ land tortnl<;:,p1;: of the ualap'ag()s Islands. Amer. Natur. 23:1039-1057. Baur, G. 1889c. The rel;aUC)flSJrup of the genus Amer. 23: 1099-1100. Baur, G. 1890a. Two new of tortoises from the south. Science Baur, G. 1890b. The genera of Amer. Natur. 24:486-487. Baur, G. 189Oc. On the of the Testudinata. Amer. Natur. k ..... JJ"rJJv. BauI. 189Od. An apparelltly of Amer. 24:967-968. Baur, 189la. American box-tortoises. Science 17:190-191. Baur, G. 1891b. Notes on the rru)o't'chHII1 genus P'P/fu'/'lP/',,(' Ann. Natur. Hist Baur, G. 1892. On the taxonomy of the genus Dumeril. Pmc. Amer. Philos. Soc. Jv.-rv--.-.-. G. 1893a. Two new of North American Testudinata. Amer. Natur. 321 G. 1893b. Notes on the classification and taxonomy of the Testudinata. Proc. Amer. Philos. Soc. 31:210 225. Bell, T. mc,nogn:lpn of the tortoises a moveable stemurn, with remarks on their arrangement and London 2:299-310. Bell, T. 1827. On two new genera of land tortoises. Trans. Linn. Soc. London 15:392-401. Bell, 1828a. of land tortoises. Zoot J., London 3:419-421. Bell, T. 1828b. Characters of the Order, Families, and Genera of the Testudinata. Zoot J., London T. 1834. Characters of a new genus of freshwater tortoise Proc. Zoot Soc. London 1834:17. Bennett, T. 1836. IN: G. White. The Natural Antlql.l1WeS of Selbome. sv~;tefnal:tcs in the Kinosternon .<,:r,r>rn,n1J7P'i! and K. leucostomum cOfnpJlex~~s KUlos1erlrndae) of Mexico and Doctoral Dissertation. Utah, J. F. and C. M. sv~;tefnal:tcs in the mud turtle, Kinosternon tlaves'cel1S \l~,1;;.:t1>1).lLJ. J. F. and J. Iverson. 1980a. Kinosternon herrerai. Cat. Amer. ,",,"'lI'"". 239:1-2. J. F. and J. B. Iverson. 1980b. new of mud turtle, genus Kinosternon, from Oaxaca, Mexico. J. 14:313-320. from Sonora, Mexico. Contrib. Sci. Natur. de Saint-Vincent. 1832. the Fr. Sci. Bickham, J. W. 1980. J...... "<'''',,...... ''' decorata. Cat. Amer. fiUlIIJIUU. 235:1-2. 1976a. Chromosome hOlll01og;y and evolution of turtles. Chromosoma Bickham, J.W. and R. J. Baker. 1976b. !'InlnhlnPl< of some necltrop!cal turtles. 1976:703~708. lhclKhatll, J. W., J. J. Bull, and 1. M. 1983. !'I.....,"'h,nPC and eV()lutl0nary relatllJm;hl1)s of Tn.onvciud turtles. VLUIU~:li:t 48:177-183. Bickham, J. W., and J. L. Carr. 1983. C!"V'pt()dllran turtles based on a cladistic of chf'OrnIOS()m,al 1978. Notes on sexual dintlorphl:sm and in 1. Natur. Hist.Soc. M. D. de. 1816. Prodrome d'une nouvelle distribution sys;ternatlqlle Sci. Soc. Philom. Paris this put'l1catum aC(~Or,r:lm2 to Smith and Smith 1853. Notices and de~;cnptl()ns or little known. J. Asiat. Soc. Calcutta 1856. Notabilia contained in the collections presenlted Berdmore and Mr. Theobald. 1. Asiat. Soc. Calcutta 24:713-723. 1859. ProceedmJ~s of Calcutta 1863. collections presenlted Berdmore and Theobald. Soc. Calcutta qW~lqLles cheloniens nouveaux appartenant la fauna Mexicaine. Ann. Natur. Testudine cartllatglflea. Kornelis van .~.~~~,_.~ ~ellck:enberglscll~~n Naturforschenden KllyniclU)Cepllalen, Schildkroten, Krokodile, Eidechsen, turtle from sOlltl1ieas:tern Sonora, Cheloniorum Tabula An;'lIVI~lCJ'l en>et()10~le ou d'histoire naturel1e SOC. London of pleurodmm the c:hl::lvOOiIO chelonians of Ann. I)p<:crinl,{\11 of new land-tortoise from South Africa, from m Proc. Zool. Soc. London ~HJ \....dtdl'J)1,tll;; of the Chelonians, Rl1yn icl1c)cepl1 131-176. IN: N. Annandale, and H. asclcuJlll Mlala:verlselS. all1thlropolo~lcal and of an eXI>edlhcm Annandale and Herbert from Yunnan. in Nat!. Natur. Pol""1''>'' williamsi J. Herpet()l. J. 1988b. Tortues 1988c. 48: 324 Bour, R. and A. Dubois. 1984c. Nomenclature ordinale et familiale des tortues \-I."";:;jJL.IU 1873, et des especes affines {K4;~PUl1a- Bourret, R. 1939. Notes sur l'Indochine XVIII. des Science de au cours de l'annee 1939. variete nouvelles. Bull. Gen. Instr. Publ. Hanoi J.7~'7\·"'I..J-J7. Bourret, R. 1941. Les tortues de rIndochine. Inst Uc!eaflIO~Jrapjh. l'Indochine. 38:1-235. (S(~hoeptl) with notes on related e. and O. 1988. The South African Tortoise Book: A Guide to South African Te:rra1pins, and Turtles. Southern Book, JOlmn,ne:solllrg. nu'v{;uu. R.e. and N. H. G. Jacobsen. 1988. On the distribution habitat and identification of Hewitt 1935 in southern Africa. Durban Mus. Novitates tlramOJle, D. 1974. Hl(>m€~ch,aml::S and eV1()!utlon. Bramble, D. M. 1982. ;'camtoctte,(vs: Generic revision evolution of M. and J. H. Hutchison. rf"PV~ln:ltl()n of kinesis in African Pelusios. 37:205-212. Bramble, D. J. H. HutchlSOltl, and 1. M. 1984. Kinosternid shell kinesis: Structure, furlction, and evolution. 1984:456-475. South African Red Data nOOK-'-j\1epl:ues and Arriohibi(lllls. South Afr. Natur. 20~21. Branch, R. 1989. The status of t1omoJPUs meN/SSC Newsletter 1981a. review of the polPulati()lls lestuclmJlda,e) OJ::curnru~ in south-eastern Africa. Provo Mus. Natur. Hr<>adlev D. G. 1981b. A review of the in southern Africa (Plem'od:ira: Occ. Natl. Mus. B, Datlern--A nl~~l€~ct€:~d taxonomic character in the G. J. Rhodin and Zool. Harvard, L:amlJ~ndl~e, Hn)adlev D. G. 1989. How many savanna mCN/sse Newsletter 10-11. 1992. review of the southern African of Ann. Transvaal Mus. Hrol1£!;erSmla, L. D. 1961. some sea Zool. Verhand. 1972. Eu:ropean Atlantic turtles. J. 1800. Essai d'une classification natureUe Bull. Sci. Soc. Phllonlatll., Paris 2:81-82, 89- 91. 1805. d'une classification naturelle Baudoouin, Paris. Brown, A. 1908. Generic types of Nearctic Pmc. Acad. Natur. Sci. Phliladlelp'hlfl 60: 112- ... "',...... "", of the \ 'VI,'''''',-"ua, Emvdld.ae) in northern Mexico. Mus. Bull, llTVnfvnPl: of Sloe-n.eCiceo jjuJrOldh?:e. A. of fiUSlnu! R. B. V'~""""Y" marmorata. North American tortoises: U. Ernst. 1977. 203:1-2. MoraU~a and C. J. l)ls;tnlmtl0n, abundance and status of tortoise. 'VQ.U''''F,'V Mus. Natur. Hist. Busack, S. D. and C. H. Ernst. Testudines, .hnwdll Buskirk, J. 1988. The Indochinese box in and out of ~JUlh"rttv 30-31. Buskirk, J. R. 1992. lieoclteltme L1UH::n.H.1. in Rio ;'YI1'nt,,'nn)(' h,r...n,nur, Carr and M8lfcfland., Mus. Univ. Ml,chl:gan 1953b. The status of the turtle zoologllca 38:137-144. Two new Cann, J. 1978. Tortoises of fiUSl.n.1lUI;!. Cantor, T. remarks on the flora and island. Ann. Natue. J. M. and E. O. MolL nalat'Ur c01np,lex of rl.U"U':I.,-'. Annual MeetIrl~ J.Jl.J..,,/,:J>JL-u.'I.. Carr, F. 1937a. Occ. Mus. ZooL Carr, F. 1937b. The status of p",,t;.un,anl1,'''' and PSt~UdiemJVs trooslii. A. 1938a. A new sul)SpeClies of P('I"un,?m,r'i: //,nr"rU1Y"n with notes 1938:105-109. Carr, F. 1938b. Notes on the r.w'1Jf1,pm1~S 1938c. a new turtle from Florida. Dec. Boston Soc. Natur. 1942a. new PS,PIHfpFJ1V." from Sonora, Mexico. Amer. Mus. Novitates 1181:1-4. Carr, 1942b. Notes on sea turtles. Proc. New J..jU15HU.IU Zoo!' Club Carr, F. 1942c. The status with notes on par'an(~hslm in v<;:,?1Jl1rp1't1V(' Club 21:69-76. Carr, F. 1952. Handbook of turtles. The Turtles of the United States. Canada, and Press, New York. Carr, 1942. A new turtle from the Florida. Proc. New hn1~larld Zoo!' Club Carr, J. L. 1990. of the turtle genus KhmG1clt?l1unvs Abstract. Tulane 'J~~" ....,,:>. Louisiana. Carr, J. L. Almendanz. 1990 Contribucion al conocimiento de la distribuci6n ~e()~rat1(~a de 2) 75-103. Inu'unlrunlt' variation in J. the Central 1981. Estudio cOInp~rrativo del cornpc.rta:mle:nto de dos espeCI~es &e(')ch,elOi1e d,entj'cUlata Y ""'V,"""'-'R' Cl)mlpar;l1bb~s de su mortojlo~ila Arl>:rentma: Heq)Ctc)falJna de las zonas y J. 1988. Geochelone carbonaria \J.'\.~0I-lUHa• Florida 14. ...... v ...... , M. 1981. et Tortue d'Hermann Testudo 11pr'm/l'nYJl ContrIbutIon de a 1£1 connaissance des dimats la France. MOlntpeHler Ecol. Prat. Haul. Etud. Chkhikvadze, V. of middle Asia and Khazachstan. Acad. 1989£1. Neocene turtles of Cocteau, J. T. and G. Bibron. 1843. ""'''''iJUHa. p. 17. IN: Ramon de la Histolre ptnrSlQlue. OO][lWlUe naturelle de rile de Cuba. Paris. :IIV~'M H. G. 1979...... ""If'~H'''''' and Arrrphiblans of Australia. Curtis Books, H .. ..('\f1 Florida. H. E. E. Cameron, and H. ZOIDlo~lcal CataJlo~tle of Australia. VoLl. Attlphllbla and Australian Govt. Collins, taxonomic arrangement for some North American amphibiians Collins, 1. J. E. 1978. Standard Common and ~clentltlc Names for North n,UJI...... '"au f\filpnml,am; Soc. Stud. Misc. Publ. H"'r1"\P,tnl Circ. 7:1- 36. Conant, 1975. field to and amiphlblillnS of eastern and central North America. H0111~hlon Mifflin, Boston. Conant, R. and J. T. Collins. 1991. Field Guide to and Arrlphlbu:ms: Eastern and Central North America. HO'U~lllton Mifflin, Boston. 450 Conant, and C. J. Goin. 1948. A new sut)Sp;eClies soft-shelled turtle from the Central United States, with comments on the of the name Occ. Mus. Zool. Univ. Ml!chl,gan 1863. Uescript:lon of Xerobates aglzsslZU. Proc. California Acad. Sci. 2:118-123. and D. G. 1990. new :rvlr>todtra: fi~sttldl'mdae) from South Africa. Studia VeOlOl~l,;a Salmanticensia 1864. On the limits and relations of the Raniformes. Proc. Acad. Natur. Sci. UWI,OellJma 16:181-183. D. 1865. Third contribution the herp;etolclgy America. Proc. Acad. Sci. 17: 185-198. 1868a. An examination of the and the upper Amazon, notes on other D. 1868b. On the of genera. Proc. Acad. Pu1b111;hed in E. D. 1869. Seventh contribution to the helpel:o!()~V of America. Proc. Amer. Philos. Soc. 11:147- Pulbhl;hed in E. 1870. Extinct Batrachia, of North America. Philos. Soc. 1-105 in 1869; In D. 1875a. Checklist of North American Batrachia D. 1875b. On the Batrachia and lChth'\i'01()2V of NIcaragua and contribution to the hel:peltol()~y of Pmc. 1887. batrachians and America and "-v....."".."""v. 1-98. C. of Geochelone 1. Herpet()L C. 1. Her'peu)L 16:317-320. 328 C. R. 1983. An annotated checklist of the fossil tortoises of China and MC)figolla. Proc. Soc. 96:567-580. C. R. 1984a. The cranial mOirptloIlleb:y of ua.lap,agc3s tortoises. Proc. California Doctoral C. R. 1984c. A nypotnieSlS for the rel,ltlonsnlp Uec)loll:tea ~allmarltlC(mSla, Vol. .hSY:leClal C. R. 1986. The of Testudo gl~tanIea ~ctlWf:HH!er. 1812: Another mt(~rplretaitlo:n. 42:237-241. C. R. 1988. nomenclatural Smithsonian He.rpeto1. Servo 75:1-17. tortoises: Evidence for Abstract. Cn,lIleston, South Carolina. C. R. and L. L. Grismer. 1990. Xerobates tep'Ul(;;cej'JaJ:ltus extensIon of Abstract. Annual HA"'''''~U'F, Carolina. Cuvier, G. L. C. F. D. 1812. Recherches sur les ossemens fossiles de QU:ldruPl~es, ou l'on retablit les caracteres du d'animaux que les revolutions Deterville, Paris. ed. 1, Cuvier, G. L. C. D. 1817. Le servir de Base al'Histoire naturelle des Animaux et d'llltrclduictlcln J..,a'uvlU"-', Paris. :17. Cuvier, G. L. C. F. D. 1825. Recherches sur les ossemens fossiles de Qu;adropl~de:s, ou ron retablit les caracteres du ....,,,,.,,,,"u...,. Paris. ed. 3, 5 vols. Le animal distribue servir de Base .a l'Histoire naturelle des &nlm,lUX et d'Introduction al'Anatomie cOlnparee. NOll'velle revue et par P. A. J..,a.u"~u". Paris. Vol 2. da Cunha, O. 1970. Una nOva sutJesl:>ecle q0<1101110, Kinosternon SCl;YlJ,Wlt1eS caro,l{JiSel.1SIS da Serra dos '-'"" ...... "". Para. Bol. Mus. Paraense 73:1-11. 1. S. and R. Mertens. Zwei unrveroffentliche Schildkrotentafeln von Pallas. Salamandra Das, 1. 1991. to the Turtles and Tortoises of the Indian Subcontinent. Publ., gel[H~rale nllt·tlC"nh,~rp Daudin, M. 1802. Histoire naturelle, et des Dufart, Paris. 2:1-432. L--"--"-J put)hsll1ed in 1801, to Helnrnmg, Dean, H. and J. W.Bickham. 1983. \.·t/l·tl..,~h""H' salvinii. Cat. Amer. .r>.....J.UlJ. entre Ie fractionnement du Gondwana et la Cretace. Studia Salmanticensia. P. E. P. 1939. Vol. 1. Testudinates and Crocodilians. Dulau and London. de N. 1915. The the Indo~Australian An;nlt>elaigo. Vol. 1. Lacertilia, Chelonia, bmlydi()sa,una. E. J. Brill, Leiden. Derr, J. N., J. W. Bickham, 1. Greenbaum, A. G. 1. Rhodin, and R. A. Mittermeier. 1987. Biochemical sysiternallcs and evolution in the South American turtle genus PI/l,fa!i1.1\!<· 329 a new name for the mid-Albemarle de Vries. T. 1984. The man. Pl'. 145-156. IN: R. environments: ualap:ago,s. Dixon, J. R. 1987. A"""II-'U4"'" of Texas. Texas A&M Univ. Press, Station. J. L. 1981. The taxonomic reI.atlcmsJhip between Murlal:ler,nys 1844 and Zool. Bot ....,..v.,-,'v.... Dodd, SVlnor)~i~ of the bl010i:Jlcal 10ggerhe~la sea turtle Caretta caretta \"-'Juua",..." U. BioI. Dodd, C. K. 1990a. Caretta. Cat. Amer. rUUlUJUV. 482:1-2 Dodd, 1990b. Caretta caretta. Cat. Amer. r'\IIIlJl,II U. 483:1-7. the flattened musk Sci. Dowler, C. and 1. W. Bickham. 1982. Chromosomal rel,:ltlo,nstllps of the tortoises ueIlewca 58:189-197. of EnlVdme turtles. Abstract. Annual Meetuli: Soc. "..,J"u"r"•. L. S. 1989. osteol!oglcal data. Abstract. First World Duerden, J. E. 1906. South African tortoises of the genus n o'mClpus, with descnlDtIcm of a new Mus. 1:405-411. Dumeril, Andre M. C. 1806. methode naturelle de classification des animaux, rendue facile al'aide de tableaux synloplt1ql1es. Paris. Dumeril, Andre M. C. and G. Bibron. 1834. Er)JCt4:>10gie genlerale ou histoire naturelle cornpllete des ""'!-J'w...""". voL 1. Librairie de Roret, Dumedl, Andre M. C. and G. Bibron. 1835. EI1ootlologle gellierale ou histoire naturelle cornpl,ete des vol. Librairie de Roret. Dumeril, Andre M. C. and G. Bibron. EI1:>et4oIoi:le i:enlerale ou histoire naturelle cOlnpete des voL 3. Librairie de Dumeril, M. C. and LAU,,"".:>\."" H. Dumeril. 1851. CataI()gUle rrleUlodlqllle de la collection des du Museum d'Histoire de Paris. and Dumeril, H. 1852. des nouveaux ou connus de la collection du Museum d'Histoire Naturelle, et remarques sur la classification et les characteres des Mus. Hist. Natur. Paris 6:209-264. '-""',lU-"'''', l"'I.Ul;:U~L~ H. A. 1856. Note sur les du Gabon. Rev. ZooL, Paris \kl!l).jV7- Dumeril, r>,U:I::.U~"~ H. and M. F. Bocourt. 1870. sur les Mission sClentlt1clue dan's Anlerlqllie Centrale ... recherches zocllo~~lqlles. Part 3. Dundee, H. category for the ampnltmulS Dundee, The Anlpllilblans and n''"'IJu"~,, of Louisiana. Louisiana State Univ. Dunn, E. 1930. from Costa Rica. Proc. New .t:.nj;larld Club Dunn, InU~rgr,adaltlOn betwetm the C. c. HI'11nn/~/pl'nnrlJ(, annulata. 250:1-2. hmocl4emlnys nasuta. 264:1. Ernst, 1981b. KhJrnOlr;lernnrys tHJlrl1,prrlmn 276:1-2. 1981d. KhjrnO~r;lelnm~vs rubida. 1981e. 1983a. t-'ln,fPW1IJ.(' radiolala. liel:Jgr:aphlc variation in ULU"H \1.'- Cat. 461:1-2. Ernst, and W. Barbour. 1972. Turtles of the United States. Univ. Press ...",,,,,,,1\,...'.,1:>.,/ W. tlaI'OOUlf. 1989. Turtles of the World. Smithsonian 1977. 1982. 299:1-4. Ernst, and 1989. The distribution and status of the tlatleIloo musk turtle Tulane Stud. Zool. Bot. Gibbons, and S. S. Novak. 1988. H. and J. Lovich. 1990. \E\.t;;PUllj1; Testudines: bnlvdlld,ae Islands. FlOC. BioI. SOC. and J. F. McBreen. 1991. Cat. 1:1-6. H. and J. F. McBreen. prr'nn,?Y/p carolina Two new turtles southeast Asia. FlOC. BioI. SOC. R. and B. Iverson. 1990. 485:1-2. M. 1980. taxonomic concinna in the Stud. Zool. Bot. 22:49-66. Fan, 1931. Preltmmal"V report from Yaoshan, BioI. Sci. Univ. F.W. on chelonians of China. P.W. on some Cht~!Oll1aII1S Felix, J. Ziva 13:227-229. ''-'''"H;'J,''''U Feuer, C. 1971. ZOO!oglsChtm l\Ilusemn zu t'ltzmgt~r, L. J. F. J. 1835. ~CllllalcrO,ren nach den Un.mqlsalzen der BraumUller Anlpl1l11Jla, from the 'IJ,,1·tp}w,;,1lp animals. HI. Mount. 1969. Forsskat P. Uescriptl,onles Animalium, avium, Piscium, Insectorum, Orientali nh,~pnJJ:lV1t Petrus Forskal ...post mortem anetoris edidit Carsten N1f"hnl"lr Fowler, H. 1906. Some cold blooded vertebrates from the Proc. Acad. Natur. 58:77-113. Frair, 1979. Taxonomic relations among sea turtles elucidated ser~[)lo,glc.al tests. 244. Frair, W. 1980. ::>elrOI()j:?,lCal survey of ple:uf()dtran turtles. 65B:505-511. Frair, W. 1982a. Serum electr~[)ptlon~sls and sea turtle classification. Biochem. 72B:I-4. Frair, W. ::>elro!<)j:?,lcal studies of the red HERP, Bull. New He:rpeto1. Soc. Frair, W. 1982c. ::>elroll)j:?,lcal studies of """,rUlL"O/l and some other testudinid turtles. 978. Frair, W. 1983. ::>elrol()~lCal survey of softshells with other turtles. J. Herpetol. Frair, W. 1985. The eml~rrlatllc Plateless River Turtle, ,'arett<7c;lelys, In ser~[)lo,glc.al survey. J. tlerpetoL 523. Frair, R. Mittermeier, and A. G. J. Rhodin. 1978. Blood t)]OCneml~;trv turtles Biochem. Frazier, J. 1985. Misidentifications of sea turtles in the east Pacific: Carella carella and Lejrna,OC!.lellIS oIlV£lCe~(l. J. M. A. 1936. Una neuva tortuga del norte ArJ~entin~[). 12:169-171. A. 1945. Una nueva de tortuga del genero 20:19-23. M. A. 1969. M. A. 1973. Dos nuevas tortugas terrestres de Arj~erlbna. Bol. Soc. BioI. COllcelXlcm 46:81-93. J. 1977. Les Cheloniens de Francaise. 1. Thesis, Unlverslltv J. IN: R.t Fritts, 1984. tortoises in va,lap;a~o,s. BioI. J. Fritz, mit HelrucKslchttglmg 33. Fritz, U. 1844. innerartlichen Variabilitat orbicularis Erne neue v H""'''' <>... ~ huroplalSch(~n SUilllpn;chlldJlaote aus 1\.le~InaSle]rl. Salamandra 333 Fritz, U. and H.-D. Bienert. 1981. Ubersicht tiber die neuweltliche Sump,fsc:hildkJrotc~n-Gattun~ ''';''I'lOnVlTI fUr D. B. 1915. On a new Chelodina from Australia, with a to Brisbane Fu,J. and of Sacalia Qu('uJrlOcellllla. Asiatic Herpetoi. Res. che~lvdlrid turtles: A of shared derived characters in the skull. nh'vloiZel1lV and classification of the categl)fll~S of turtles. Bull. Amer. Mus. Natur. E.S. 1977. The sidenecked turtle of rel S. 1984. Historical l:111iillVSIl> of theories of chelonian rel'iho,nstup. E. S. 1988. A ClaGOl:!:ram of the pleurodmm turtles. Acta Zool. Cracov E. S. and P. A. ~V"'~Y~"'h The Vh'lIlo'IJPrIV and Cl,lSS111c:atlcm '-'.u,,,...... E. S., P. and A. 1991. cornptlter assisted of the catc~gones of turtles. t..:l,aGlSW;S 7:in press. C. 1985. Comments on two checklists. Garden, A. 1772. IN: T. Pennant. An account of two new tortoises: in a letter to Matthew M.D. Sci. Philos. Trans. Soc. London Garman, S. 1880. On certain of Chelonioidae. Bull. Mus. Zool. Harvard 6:123-126. Garman, S. 1891. On a tortoise found in Florida and Cuba, Cinoslernum baurii. Bun. Essex lnst. Garrett, J. M. and D. G. Barker. field to and am:phlbi,ims of Texas. Texas M{)fltltHy Press, Austin, Texas. {jeottrov St. Hilaire, E. F. 1809a. Sur les tortues moUes, nouveau genre sous Ie nom et sur la formation des carapaces. Ann. Mus. Hist. Natur. Paris 14:1-20. Hilaire, E. F. 1809b. Memoire tomes moUes. Nouv. Bull. Soc. Philom. Paris 22:363-367. literature Adams. sy~;ternatlcs of Australian Chelid turtles. Abstract. First World LOlngress of Heroetoll)gy the Australian chelid turtles on aU()zvme ele~ctropJhOl·eS]$. Gibbons, J. 1990. of the Slider turtle. Novak, and fossil turtle, Ki,wSl~enlon anZ0I1er,lse, from of Glenn, J. of the 1789. C. Linnaeus. cum Characteribus Ulltterentlls, SVI110flVnlHL Gmelin, G. 1774. Reise durch KU:SSl Goode, 1. 1967. Freshwater tortoises of Australia and Melbourne. Graham, E. 1991. P. Grandidier, A. 1867. 1866, sur la Zoo1. 1869. ql,l4elqlues animaux nouveaux dec:owverlti, l'annee 1869, sur la cote ouest Paris A.lllpnlOla, with a descfllDbcln some new collection of ''''G'U' ·-'u",,,"'. in J. E. 1831a. 1-110 IN: E. Griffith and The Class J.'\.cJ.lUjua, ~rr~n(r..l1 des,cnptl()fis. Vol. 9. \A!1'l•• tt"Ir"".. and Co., LJVllUVU. pUllJlll~atllon date for parts of this J. E. 1831b. KelPWJlUm or short the Part 1. tortoises, croicmilles, enaliosaunans. Treuttel, Wurtz J. E. 1831c. Characters of new genus of freshwater tortoise from China. Proc. Zoo1. Soc. London 107. J. E. 1834a. Characters of several new from India Zool. Soc. London 1834:53-54. J. E. 1834b. Characters of two new genera Proc. Zoo1. 1834:99-100. J. Eo 184LA catallogl11e of and ampl11tna with a de:;;criptllon from Western t'\U:SLHI,U J. 1842a. IN: J. E. J. E. 1842b. of two hitherto unrecclrd(;~d British Museum Dr. Diffenbach. pp. Wurtz & Co., London. J. E. 1844. CatallJglle of the tortoises, crocodiles, and ampl1lSbaerl1aJIS in the Collection of the Museum. Natur. Hist., London. J. E. 1847. Descrlptlon of a new genus of Errlvdldae. Proc. Zoot Soc. London 1847:55-56. J. 1849. DescrIPtiOn of a new box tortoise from Mexico. Proc. Zoot Soc. 17. J. of tortoises. Zoo!' Soc. London British MUlsellm. 1859. D~scnp'hon of of tres,hw,ater 1859:478-79 of Mammalia and tortoises from CamuoJla. London Ue~)cnph<:m of in the E. 1861. of water-tortoise «(i~?OctenlmJls mtetc.tno,sleirna from Proc. Zoot Soc. London of from the Lao Mountains, Observations on the with Zoo!' Soc. London 1863b. On the npl\Jlm'~~ from Australia, with the ti..,,:rrl'nh,\n of a new Natur. Hist. London 1863c. Note on a new rejfOmteal'4Sa from Natal. Natur. Hist. London J. E. 1863d. Note on American r.rrlvdlda,e. observations on my cataloJgue of them. Natur. Hist. London J. 1863e. Notice of a new IHdH11f"'.. from North-Western India. Froc. Zoot Soc. 863:253. J. and other tortoises from Natur. 1864a. and with the des,cripti()ns of some new Froc. Zool. Soc. J. DescrIPtion of a London J. 1869a. cl1a,racters afforded of J. 1869b. Ue::;cnptlcm of M£1,Ur,pI'l1'\J,~ a new freshwater London 1869:499-500. 1. 1870a. Notice of new Chilian London 191 1870b. Notes on of tortols~~S Zool. Soc. 1870:706-708. J. E. 187Oc. Testudinata and two new of Australian freshwater tortoises. J. 1871b. Notes on Australian freshwater tortoises. Ann. Natur. Hist. London J. 1872a. A"'''''If'~U'''''' in the Collection of the British MlllSetlm. Testudinata J. E. 1872b. Lauw:>gule of Shield A"''''''IJU1'vJ in the Collection ofthe British Museum. Part n. bnlydlos:aurlal1IS, Khym;hOCel)flal1a, and and Francis, London. J. E. 1872c. Notes on India Ann. Natur. Hist.London J. E. 1873a. siX~cnnetlS of shield the British Museum. British MUISetlm, J."VJ,JUV". J. E. 1873b.Notes on tortoises. Ann. Nator. Hist. London 1873c. On a new freshwater tortoise from Borneo Natur. Hist. London 156-157. de~;cnptu)ns of new genera and Ann. Natu!. J.E. of Mexico' of MM ,",UI''''-1H and Bocourt. Ann. J. E. 1873f. Notes on Chinese mud-tortoises with the desCrI1PUCm sent to the British Museum Swinhoe, and observations on the male Natur. Hist. London 1976. Patterns in the distributions of Southern African terrestrial tortoises ZooL Africana 11:249-273. G. and Shine. 1985. An open letter to an herpetologls1ts. Rev. Data Book, Testudines, Cn)ccidyl1a, Khyn(~hoICel)h3ll1a Uf()oulbrJLdg,e, B. and R. Luxmoore. 1989. The green turtle and hawkbill (Kt~ptllla: Chelonndat:~): World status, eXI)lOlclatJLon and trade. CITES, Lausanne, Switzerland. 601 pp. E. Moll, and J. 1983. KeljlS(;Overv of a Halrdw'lcke, London. GUnther, A. 1885. and Batrachia. 1- 326. IN: Americana. and Dulau & London. was put)11slrted in 40 parts; Part 1, pages 1-56 aPlJeared Haiduk, M. W. and 1. W. Bickham. 1982. emph~lSlS on the sysltematlc pialcernlent Ualap,ag()s Islands, cmnm,onJly known as Harlan, R. 1835. Genera of North American a synopsIs of the Med. Res. 1835:84-163. G., Harless, M. and H. Morlock 1979. Turtles: Persn€\dl've!i\ and research. John Sons, New York. 1940. Some works of ...... , ... "U', iJVIUIUU, and their upon American nel~roiio~l.calli~rature. Haltwe:g, N. 1934. Description of a new kmost~ernld Univ. MIlCnlJ\l.an 277:1-2. N. 1939a. A new ~m,efli[;an rSt~ual'?m)is. Mus. ZooL, Univ. Mllchl!san 397:14. Haltwe:2. N. 1939b. Proc. Amer. alaclemn:IYS, a genus of turtles. U. S. Bur. Fish. 24:1-20. 1976. 184:1-2. D. and R. R. 1974. new the Uuad~I!UlJe River system in Texas. Tulane Stud. Zool. Heatwole, H. Letter to the Editor. Heimann, E. 1990. graeca Linnaeus. SAURIA I.C.Z.N. Direction 97 (name and International ComrrUSSlon Henderson, J. R. 1912. Pre\!munarv note on a new tortoise from South India. Indian 218. de~;cnptl()ns Hewittl J. 1927. Further of and batrachians from South Africa. Mus. Hewitt, J. 1931. lJeSCfJtptlons of some African tortoises. 6:461-506. Hewitt, J. 1935. Some new forms of batrachians and from South Africa. Rec. Mus. 4:283-357. C. 1990. Tortoises of Africa: Ta:1\0l1l0rrlY evolution with notes on field studes in Tunisia. J. Chelon. Heroetol. revision the Testudines of North Africa, and J. C. and J. Martin. 1989b. Testudo whilei Bennett 1836: New on an old carapace- Gilbert White's ~eJlOOme tortoise re-discovered. J. t)lOlO~ICal data on the green turtle, Chelonia 1758. F. 1980a. Chelonia. Amer. McllllJl.!lIJ. Hirth, 1980b. Chelonia Cat MClrptlon:letI]CS of the tortoise, Geochelone sulcata, in: the Sudan. Holbrook, J. 1836. North fiITICnCan Helrpellolclgy; the 1I111laDltltlig the 1. Dobson and Holbrook, J. 1838. North American tie·nx:tology; des:Cf11phCl!1 of United States. Vol. J. Dobson and Son, PhlIIa(leIl,hla. Holbrook, 1840. desCf11Dt!Cl!1 of Ed. Vol des:cnJptlCl!1 of the Holman, J. A. 1977. Comments on the Natuulrlijke J..U"'''.H'''' with Houseal, lieogJraphlC variation in the mud Hsu,H. pre:l1n:llnary note on a new Contr. BioI. Lab. Sci. 6(1):1-7. HutchlSOlil, 1. H. and D. M. Bramble. 1981. Klfloslterrl1dl'te and related turtles. Hutchu<;on, J. 1991. Killos·terrlin~te U'lcelJ'uu;;a; Testudlll1e:s) and their ptl'ylogelletJlC slgmtIc:mce. J. Vertebrate Paleo. Iverson, J. 1976a. KUWSiren10n sonoriense. Cat Amer. fill.llJllllJ. 176:1-2. J. B. 1976b. The genus Kinosternon in Belize Cf(;$tuidultes: KUloslterrl1dl'te Herpet()IO~~lCa 32:258-262. Iverson, J. 1977a. Kinosternon subrubrum. Cat. Amer. '"" ....ll.ll". 193:1-4. Iverson, J. B. 1977b. Sternotherus rln....,n('"".'" Cat. Amer. r\.1I1Vllll.). 194:1-2. Iverson, 1. B. 1977c. Sternotherus minor. Cat. 195:1-2. Iverson, J. B. 1977d. lieo~u:aptllC variation in the musk turtle, Sternotherus Iverson, J. mud turtles, Kinosternon baurii \J."\C!J~l.U", Testudines, Kinosternidae). J. Iverson, J. B. 1978b. Distributional prclDle:ms of the genus Kinosternon in the cu.,,,,....,,,, southwest. Iverson, J. B. 1979b. Sternotherus carinatus. 1980a. KU1OsterJ'10n UL~HU1H. Iverson, 1. and W. P. McCord. 1992a. of the Proc. BioI. Soc. Iverson, J. B. and W. P. McCord. 1992b. A new sut)SpleClles Hainan Island, China. Proc. BioI. Iverson, J. B. and W. P. McCord. Proc. BioL Soc. Iverson, J. and Mittermeier. 1980. Vel'matemydlliae, lJermol'em'lIs D. Cat. 237:1-4. Jackson, R. 1978a. Amer. 210:1-2. Jackson, R. 1978b. Evolution and fossil record the chicken turtle with a re-evaluation of the genus. Tulane Stud. ZooL Jerdon, T. 1870. Notes on Indian Helrnel:ol()l?V Soc. "-c<;U,","HHl 1870:66-85. Juvik, J. O. 1975. The radiated Juvik, J. J. Andrianarivo, and C. Blanc. "''''....''15'''''''-''''' tortoise in northwestern Mada}~as(~ar. Khan, M. 1982. Chelonians of tlangJ.a<1leSn and their conservation. J. tlOllTIO;;lV Natur. Hist. Soc. 10- 116. RrlOnl'!m:vs - nouveau genre de tortue terrestres (Tl;~stuldirl1d;;le BulL W. and R. L. Burke. 1989. Crocodilian, Tuatara, and Turtle laJi.onllmJIC and Ue,ogl:aphlC Reference. Assoc. CoIl., M. and J. Miller. 1985. Letter to the Editor. Ba:staI"die:rurlg von Testudo fU"llc(CnpUl'lI und Testudo h. ner'rtlfJ'nfll Kizirian, 1990. reassessment of the pn~ylogelletllc of soft-shelled turtles Abstract. Annual Tulane, Univ. New Orleans. A. 1984. in the Misc. PubL Mus. ZooL ~CIIOe]Dn. 1792 auf den Nordlichen ~pc)raClen, of the threatened sawback turtle, and a new record to China Krefft, Austr,ah2ln animals in New Guinea with of Ann. Mus. Civ. Stor. Natur. CriltlC2111y en Kuhl, H. am Li:t'\';t':I-X::Ut':, B. G. E. 1788. Histoire naturalle des serpens. VoL 1. ImlJfUlnerle du Paris. AJ""''''''''''. P. N. 1986. Cat. Amer. rl.HIUIJLlU. 396:1-2 Lamar. W. W. and F. Medem. 1984. Notes on the chelid turtle PHlr"n,nnt' in Colornblla V"'~IJ'Ul1i1; TestudJlnes: Salamandra T.• J. C. Avise. and J. W. Gibbons. tortoise Evolution T.and J. E. Lovich. 1990. mud turtle (AtflO,sle,rnlm in the Carolinas and Lambert, M. R. K. 1983. Some factors intl!uetlciIlg Moroccan distribution of the western Me:Olterrammn tortoise, Testudo graeca graeca and those its survival in NW Soc. 79:149-179. J. L. 1979. and olstribruti()n). Smithson. Lataste, F. 1886. UescrtptlOn d'une tortue nouvelle do .t1aUh::>eIlellal a101n017US Le Naturaliste 287. Latreille, P. A. 1801. IN: C. N. S. Sonnini de Marloncourt and P. Latreille. Dessinnees Nature. Deterville, Paris. Vol. 1, 01s ,-"un""••" R. F. 1956. Contribution a rH<~IDeto,1021e I. Generalites, n. CMloniens, '-"UU<;oII" R. F. 1965. A contribution to the Ict1l1Wlp£h71." of the Afr. Cent. Zoo1. 1-33. LeConte, J. 1829. Americarl tortoises. AJV"'...... Natur. Hist. New York 3:91-131. LeConte, J. 1854. UescrlpUlon of four of Kinosternum. Proc. Acad. Natur. Sci. PhIladelphia 7:180-190. ;rm,Ho,..ut' from north-central Mexico. Univ. Kansas Pub1. Mus. Natur. J. M. 1960a. Natural of the ornate box turtle, Univ. Kansas Pub1. Mus. Natur. Hist. 1 turtle from Coahuila, Mexico. Univ. Kansas J. 1965. of turtle, genus KinosteYnon from Central America. Univ. Pub!. Mus. Natur. Hist. J. M. 1966. Notes on the natural of a rare Central American turtle, KinosteYnon anflu,f;tllJOl:IS 118-122. 1981. Pleurodira: '-',,"'..'...... J. and distribution of the slider turtle. Smithsonian Insbtllhcln and 484: UL"',,-,"'UVVU, A. R. 1908. A new genus and of turtle from North Australia. Rec. Mus. 128. McDiarmid, R. W. and M. S. Foster. 1987. Additions to the Stud. l'!Jp,..trr.., McDowell, S. B. 1964. Partition of the pro1blems in the taxonomy of the Testudinidae. Froc. Zooi. Soc. McDowell, S. B. 1983. The genus in New Guinea with notes on the mo,rptlo!<)gy of Pleurodira, pp. 169-189. IN: A. G. J. Rhodin andK. (eds.). Au'vances heI1)eltol()~V and Mus. Zool. Harvard, Mass. McKeown, S. and G. Webb. 1982. Softshell turtles in Hawaii. J. tierpetol. 11. McMorris, J. R. 1976. The rea,Sslgmnel1t of tteoefnVC;fa t(~he.tJOJu:n!$lS Bourret. Chelonia Medem, 1958. Informe sobre Colombianos. (II). EI conocimiento actual los Testudinata en Colombia. Bol. Mus. Cienc. Medem, F. 1977. Contribuci6n al conocimiento sobre eeologla de la "Bache" '~"'-'f-' Melville, R. V. 1985. and Iverson, Melville, R. V. 1989. 1825 and Pelusios Conserved. Merkle, D. Merrem, B. 1820. Versuch eines ~vs;tenrlS der Anlptllbl.en. Tentamen ~Yl;ternatlc A,mr.hltlOnlm. HelmeJrkungt~n tiber Rassen von Pelomedusa Anz. 117:139-142. Mertens, R. 1946. Mediterrane SclliklkriJtelo -~~as:)en. SenckelnbeJrgiana 27:111-118. Mertens, 1954a. Zur Kenntnis der Schildkrotenfauna Senck. BioI. Mertens, R. aus Sud- Zentralamerika. Mertens, R. 1967a. Bemerkenswerte Su:ssV\rasl;en;chlldJuoten Brasilien. Senck. BioI. Mertens, 1967b. Die heJrpetol,o~l'sclle des Natur-Museums und FOJrsclhUIlgs-1nst1ltut{~S Sen(~ke:nberg 1m Frankfurt am einem Verseichnis ihrer Senck. 1969c. neue aIS'Welrlde:r -~;Ch.lldlcrote aus BioI. 50: 132. AIlllptHtJlen und J.'''''IVU!!~U We:st-Palosltans. 197:1-96. L. Mertens, R. and L. Milller. 1940. Die Arrlphtbtl;:n und ni;;IJUIIi;;U A.4U"V!J'«". Abh. Senck. GeselL 451:1-56. Wermuth. 1955. Die rezenten Schildkroten, Krokodile und Brilckenechsen. Abt. 1874. Bernhard tncmychld turtles: evidence from skull morptlol()~V Pal nAvy'«", P. 1987. The ph~rlo)tenletlc rel,itlonsi1nps of soft-shelled turtles Bull. Mus. Natur. Hist. 1989. The skeletal InOlrpl1()lo~:y {,l·\fnt{..rhr~n turtle, Adocus, and the relat! 1987. The chelonians the Laetolil Beds. Vh.(\r~'np site of Laetoli, northern Tanzania. of the Lake Mikan. J. C. 1820. Delectus Florae Faunae Brasiliensis. Antoinii Strauss, Miller, J. F. 1779. leones animalium et nhl,nh:.rn,m of natural wherein are deJllOf:atf:d animals and many curious Milstead. W. W. 1967. Fossil box turtles from central North ,,,u.v~,.vu. and box turtles of eastern Mexico. Milstead, W. W. 1969. Studies on the evolution of box turtles BulL Florida State Mus. BioI. Sci. 1-113. and D. W. Tinkle. 1967. prr'/Hl_"1'lP of western Mexico, with comments on the groups in the genus. Minton, S. A., Jr. 1966. A contribution to the h"'Y-nPlro!r,rru of West Pakistan. Bull. Amero' Mus. Natur. Hist. 134:27-184. Rhodin, F. Medem, P. Soini, M. S. HOO~lmoed, Distribution of the South American chelid turtle Pln"Un,nIH' Mittermeier, R. A.and R. Wilson. 1974. ArrlaZ(mUm pelOlUe(luSlld turtle. 1945. localities of American turtles. 1976. Handbuch Palae()herpe:tologJ,e. Part 7. Testudines. Fischer ZOloge:oglrapl11slcl1e Notizen fiber LJU'J.-L'''","'UH aus den Jahren 1886-1888. L>Ui§'''-''vU und Darda." Mitt. Ver. Steiermark, Graz Moll, 1986. The status, and level of eXI)lolltatlon of the freshwater turtle }prmnlpr11V"" mawei in Belize, Central America. BioL Conserv. O. 1980. Natural nn.nu••r baska Mlllla,rsum J. Sci. V"""" ....F·V...,. Moll, £~V~'AU~<. Annual meeilings 11. tortolse:s: In Vlrtffl""~'<' 1870. tuberosa, Art von Schildkroten aus uu.uma-Y'Ui<1Uta. Wiss. Wien 1990. Eine estudllneS, Pleurodira: Chlelldlae) of Vol. The China. Amer. Mus. hel:petollDglcal fauna of the Lobatski-Linokana area. 1932. Testudo verreauxii in variation. Afr.1. Sci. 29:466-472. Price, Turtles of the World. T. F. H. Pritchard,·P. 1979. H. Publ., 1980. 238:1-4. Pritchard, 1983. 108-111 L 1984. Further thc,ujt;hts on "Lonesome uaJapajt;os 39:20-23. Pritchard, P. 1986. A remferpiretatlon Testudo J?1J?jant~ul S(~hwell:1'!!:1'er. 1912. J. HeJrpetol. Pritchard, 1987. zuliae. Cat. Amer. r>"'U"'''_ 404:1-2. f\.U.Ul Pritchard, 1990. Several New Tortoise Tortoises and Turtles Newsletter and 1991. turtle from China. P. Trebbau. 1984. The turtles of Venezuela. Stud. ~lItUIIIIU. 2:1-403. J. R. and P. Gaimard. Autor du conformement aux instructions de s. exc. M. Ie Vicomte du tloucJuuze, uepat'tment de la Marine, k.J""'_""U~'" les la PhJrSl<::len.ne, .....v''''.u,!''..'''. M.L o osservazioni de la Nature ou Tableau de rUnivers et des n1.pr"~Q soft shell turtles of North J:A Atlantic J. Friend 349 Ramo, C. en el Rau, R. flnllalJen uber die ge()mietnsclle LandschildkrOte, Rau, flU11:1UJeU uber ge()mletrisclle Lam:1schll Raw, L. R. G. 1978. tenrap'lUs, genus Pelusios, of Natal A. MltllenJ1leler and Cat. 1983. lK:mosterm.dale) ttlroll£hout its range. Rhodin, G. J. 1989. Ph'ylogel:letlc rl;~latlon:shll'S of the side-necked turtle Chelidae. Abstract. First World "'-'VjUl',!~:>:> of lieJrpetol()gy Rhodin, J., Uislribputl(,n of the turtles I>//""n,,,"'., raarlOtala Rhodin, A. J. of the South rU'i1vAAvU.U in lierpetol()gy A. G. J., R. and !. Buskirk. 1988. 439: 1-2. Mluermelcr, and Silva. taxonlomic status of Phl"un,(}n~ a from soulthe~astern BraziL Rhodin, A. G. J., R. Mittermeier, and C. H. Ernst. 1990. Cat. 481: Mrttermeler, and J. R. McMorris. and Pantanal JPrmn.rhp'IV.'i: coriacea. J. Richard, Richard, Richard, Lista slstenl1atlca y distribucion las Nal:ur!lchl;>-n J:;;mHlellung der Am,phll:>ten. Nova enlallLen von Sacalia bealei 1988. Die Chinesische Vlt~rallgencluldlla()te. Sacalia 1986. de Geochelone costarricensis continente americano. BioI. 1. HOlnb~lV Natur. 1901. a new land-tortoise from Ualapag()s Islands. Wirbelthiere zu ~cllunertler. Frankfurt Sachsse, W. 1975. and J. E. Lovich. 1988. irantf'~nnIS barbouri. Schmidt, kinosternid turtle Colombia. list of Historia Testudinum iconibus Illustrata. Palmii, ErJlanga{~I:::;: HTI,an",,,,n I 1801: 1955. The diamondback tefJrapms ~1Ylt'llat';te"rtys fo..,rnn'''' of perun:ml;ar Florida. Soc. 68: 157-164. UeoglraphlC variation in the chicken turtle JO"'"r,l1o/'i1" reticularia Latreille. F. 1956b. The rel,lhonstups and nomenclature of the soft-shelled turtles of southeastern United States. Mus. 26:1-21. Schwartz, F. and W. Henderson. 1985. Guide to fiCllJUlIC" of Indies, Exclusive of Milwaukee 1975. A checklist of Indian am,phibi,ms Prodomus M()nclgr;aptllae Cheloniorum. Arch. Naturwiss. Math. 1:271-368, ~ejelu~er. L M. 1945. Variation in Pacific mud 1978. Kinosternon flavescei'l!s. 216:1-4. from the and 1984. Revision of the West Indian TnJ!ch{~m11d PUJ'srr:aCl. Annual M{~etllng Amer. Soc. Univ. Uklahoma, cornp the trajche:m,rd turtles (Tc;~stl.lldiries: Em:vdidae cornmlrisons and ph'vlogelletJlc n;~latlon:shll)S in 1981. Stt?rnotllreYj'4S odoratus. 1979. The land tortoise of Aldabra, Geochelone QH!amea. Shaw, G. 1794. ~~'~~~,." of New Holland. Vol. I. Davis, London. Shaw, Vol. III. Part I. T. Da'vid!;on_ 14 vols. from 1800 to 1826, each Shea, G. M. 1987. Comment on the prc~po:sed sUJ'Pr,eSSlon nOlUelrlcl:atu:re hn..""•...,.",,,,,,· of three works Wells and C. Ross R. M. 1976. The natural Baur. in Alabama. BulL Florida State Mus. BioI. Sci. Shine, HerPetologlcal l;ontereOlce. and the 1984 Annual General and 28 to 2 Ch,ph,rnlc1::le von Australien: SchiIdkroten des ostlichen Hinterindien. Sitz. Akad. Wiss. Wien Math.-natur. Kl. Siebenrock, F. 1903b. zwei seltene und eine neue Schildkrote des Berliner Museums. Sitz. Acad. Wiss, Wien Math.-natur. Kl. Siebenrock, F. 1906a. von Ostafrika and Maida~~asJl\:ar, in Ost-Afrika in den Jahren 1903-1905 mit Mitteln der tlermlmn Wissenschaftliche Arbeiten. Siebenrock, 1906b. Zur Kenntnis der Schildkrotenfauna Insel Zool. Anz. 30:578-586. Siebenrock, 1906c. Eine neue Cinosternum-Art aus Florida. Zool. Anz. 30:727-728. Die Schildkrotenfamilie Cinosternidae. Sitz. Akad. Wiss. Wien Math.-natur. KL Siebenrock, F. 1907b. zum Teil seltene SchildkrOten aus SUdchina. Sitz. Acad. Wiss. Wien. Math.- natur. KL 1 Siebenrock, 1909a. der rezenten Schildkroten. mit BerucKsIChl:lglmg der in hi<:tnri<:t'h"r Zeit au~;ge:stOl,belt1en Arten. ZooL Jahrb., 10:427-618. Sieben.rock, F. 1909b. die HeI'ect:ltlglung ~el])stamllg1celt von Sternothems .,"~ .. ,~~"'~ seV'Ch,eUi:mSIS Siebenrock. Zoo!' Anz. Siebenrock, F. 1914. Eine neue Chelodina Art aus Anz. Akad. Wiss. Wien Math.- natur. KL Sites, J. W., 1. Bickham, reconstruction of turtle Ph'Vlo,geltues: H,elslhonslillps Smith, H. name of the snapper turtle. Helrpetololl?,lCa 11: 16. Smith, 1947. musk from southeastern United States. J. 353 Smith, and Brandon. 1968. Data nova heI-peltol()gu:::a Mexicana. Trans. Acad. 71:49-61. and Grant 1958. NoteVllorlthv hel:-Ptl.les from Jalisco, ..A ....h.VV. Smith, 1952. new turtle from Texas. Wasmann J. AUmOrSh]lp of scientific name the leatherback sea turtle. J. Smith, B. Smith. 1973. Mexico. Volume II. literature exclusive of the Mexican Helnmngton, Vermont Smith. 1979. of the herpetlJtalllna of Mexico. VoL to Mt:~XH;an turtles. Hell1mngt.on, Vermont Smith, M., 1980. Sternolhernus 1830, DrolDos~~d Bull. 1950a. localities of Mexican and amphIbIans. Univ. Kansas. Sci. 1950b. annotated checklist and to the of Mexico px(~ln,,:jvp of Mus. 199:1-253. Smith, 1916. list of the crocodiles, tortoises, turtles lizards at present known to inhabit Siam. J. Natur. Hist Soc. Siam 2:48-57. Smith, M. 1931. The fauna of British and fiUlPllilUlii. Vol. 1. Loricata and Testudines. Smith, 1951. new new turtle from western Illinois sand Bull...... UJ,'-'(ll';V Sci. 9:189-199. Zootaxonomica J. B. von. 1824. Animalla nova, novae Testudinum et Ranarum quas in itinere Hr:3lsl1llam anms 1817-1820 HUbsc:hrrlanll1, Munchen. conservation pf()gr,amme for the Aldabra tortoise Geochelone InternatL 28 :66-69. Stebbins, R. 1985. field to western and amphlltnatns. Second edition. tlCtugntoln Mifflin, Boston. H. 1902a. Proc. BioI. Soc. aShlugt L. H. 1902b. Some turtles. Proc. BioI. Soc. L. 1907. Hplrrv>·!O!',\O'V of Bull. U. NatL L. H. 1918. DescflPtion of a new smlppmg turtle and a new lizard from Florida. BioI. Soc. 31:89-92. L. 1925. and sullspecl,es of American turtles. J. W BtSnllngmn Sci. 1~(:'W):4bL·4bj. 1J~H 1917. list of North American amphlol,ms and ...... i). Harvard Univ. IN: Walbamn, J. J. Friedrich Walbaum, J. J. 1785. Be:schreilburlg der lpe:nglenlscllen Schildkrote. Schrift. BerL Gesellsch...... "..,rl.<- 13 Waller, T. Distribuc1on, habitat donosobarrosi, Notas sobre la distribuci6n de Ac,anllwlche'lys en la Arj2:entma Wallin, 1977. The Linnean tvpe-speClml~n of Zoon Carella caretta of the Pubt Waterhouse, Pmc. Zoot Soc. London Weaver, :sy~;ternatlcs, fossil evelfuttOn of Stud. Zoot softsheU turtle from the soulthe:astem United PubL Webb, of the Gulf softshell Webb, R. G. .u""" .",un Recent soft-shelled turtles Univ. 137:1. Webb, 1973b. Cat. ArneI'. tllllUIJIlU. Webb, G. 1973c. muticus. Cat. ArneI'. tlU:UJlJIHJ. 1973d. Webb, £1 ..",u,,,,,n« of Indian softshell turtles Bull. Mus. Natl. Webb, 1981. narrow-head J:)Vl1k:)1I~;1l turtle, Chilra indica Zoot Surv. India 79:203-204. Webb. R. G. 1982. Taxonomic COfiCel:llUl); the tnC)llVChld. turtle LJ£.'.u:;,nv.' punctata of the Sierra Madre UCCldlenltal, UA"''''AVV. L. and N. L. LUS,chlll); Univ. Kansas Mus. Natur. R. G. 1985. Taxonomic status rostrata A n1"unAOA,;;, R. G. 1990a. sinensis Wle);lmann, Vrr.nn"",ri conservation of the names. Zool. Nomencl R. G. 1990b. Cat. Amer. FUIIU"'AU. 487:1-7. 1960. A new i>VA',",U\.,U turtle from Coahuila, Mexico. M. 1989. A chelonian The Indo-Chinese box turtle in carlhv'ltv Plastron Wf.~lssmg;er. H. 1987. Testudo graeca anamurensis ssp. nov. aus KIl~maSl'en. Wien Wells. R. W. and C. R. WeUm);ton. 1983. A svnonS1S of the in Au:,tralua. Australian J. tielrpetol. 1985. A classification of the Anlpl'UOlia and '''''''If/UA,''' of Australia. Australian J. Wermuth, H. 1952. Testudo hermanni robertmertensi n. und vorkommen in VlJ"UU"-". 157-164. H. 1958. Status und Nomenklatur der Maurischen LandschildkrOte, Testudo graeca, in SW-Asien und Afrika. Senck. BioI. n. ssp. D. Wermuth, H. and R. Mertens. 1961. Schildkroten, Krokodile, HI1uck:enecllsen. G. Fischer, Jena. H. and R. Mertens. 1977. Liste der rezenten Alflph:uncm und L\.I;IJUIJl\;;U. Testudines, <;rc)codvlla Das Tierreich, Berlin 100:1-174. 1901a. 1.''''I"nuA''''' des Kom~:sbl~r);ler Zl[)olog:lsclhen Museums. Anz. 24:297-301. Werner, F. 1901b. Uber A''''lfJUAA",n und Batrachier aus Ecuador und Neu-Guinea. Verh. Bot. Ges. Wien 51 :593- 603. from the M. 1839. Jahren 1832 bis 1834. Hoaelscher, Coblenz. im nordlichen America beobachtet wurden. Isis von Oken 21:364",383. L.."V'JAV,F,A"-. ges:ammelt auf einer .LJ\A,'IJ'..HU.-..... 1950a. Variation and sellectlon of cervical central articulations of turtles. BulL Amer. Hist. PT.JV.J-JVk. Williams, E. CU/7e11!SIS and' the evolution of Natur. 1952. West Indies, and a tentative amllnJ.t.ement torl'Ols~es of the 99:541~560. Williams, E. E. 1954b. of Pelusios and the evolution of that genus. survey of Williams, E. 1958. Australian chelid P Williams, E. 1960. Two of tortoises in northern South America. Breviora 120:1-13. G. Grandison, and 1967. Chelonia l1el'Jressa Garman re~·mvestJtiZalted. Breviora 271:1-15. V. and G. 1991. 509:1-8. 1979. The Turtles Thailand. Siamfarm ZooL blCllolUC:lll data on the hawksbill turtle, 'retmoclu~lvs irnbricata \LJlHUctt;:lJt::>, Wood, C. 1977. Evolution of the errtvdme turtles J. Herpet()l. Wood, C. 1984. side-necked turtle Pelusios adansonii in Lake Abstract. Annual MeettrliZ lcntn\'ol. nerp(~!ojl., Univ. Norman. some turtle blood albumins with Hendrickson. 1988. recles(;nJ't!cm of the Australian flatback sea turtle, BulL Zoo!. 1: 1-69. atr,acFlem!vs) dahli, from Colol1t1bla. ppj'n/",tlp/'>l<' from . Zhao Herpetol(JiZV ofChina. Soc. Stud. nUJIUllll.J. Oxford, and Chinese Soc. Stud. G.R. 1971. relicularia. 107:1-3. carinatum, KIJrlmltelrnCln carinatus, Pelusios carolina, 8 Plrrnnp'1P 104 189 250 4 Chelinae 4 Chelodina 10 Chelonia 83 Cheloniidae 80 Chelonoidis 247, Chelus 20 .TiP-J.ljnTfl 92 92 25 243 chichiriviche, lrlacltenlVS 208 chilensis, (jf~oche,tone bealei, bechuanicus, Pelusios becki, (je,OC!:letcme 17 86 360 crassicollis, 166 creaseri, Kinosternon 221 cruentatum, J{uwsifenwn ('rfl>,nt,(}t/f,O<' 1 Rhinoclemmvs l1u,U:WrlaruJ 162 131 157 246 168 53 304 208 262 hoodensis, Geochelone 289 161 146 casl'anOlaes 61 186 189 271 Nilssonia 312 6 13 276 231 oblron1W. Chelodina 14 arachnoides o 17 1- ISBN 0-9617431-0-5