THAI FOR. BULL. (BOT.) 37: 64–106. 2009.

Towards a stable nomenclature for Thai ferns

STUART LINDSAY1, DAVID J. MIDDLETON1, THAWEESAKDI BOONKERD2 & SOMRAN SUDDEE3

ABSTRACT. There have been many changes to family and genus delimitation in Thai ferns since the publication of volume 3 (Pteridophytes) of the Flora of Thailand. These changes are discussed and current names for all taxa, including those recently described or recorded for Thailand, are presented. A new combination is made in Hymenasplenium.

KEY WORDS: Ferns, Flora of Thailand, nomenclature, Thailand.

INTRODUCTION

There are approximately 670 taxa of ferns in Thailand comprising about 5–7% of the total vascular flora (Middleton, 2003). The ferns were published in volume 3 of the Flora of Thailand (Tagawa & Iwatsuki, 1979, 1985, 1988, 1989) and 596 taxa were recognised as occurring in Thailand (excluding the lycophytes: Lycopodiaceae, Selaginellaceae and Isoetaceae). A number of papers have since been published in which new taxa have been described or new records have been added to the pteridophyte diversity of Thailand (Mitsuta, 1985; Iwatsuki et al., 1998; Parris, 1998a; Hovenkamp et al., 1998; Nooteboom, 1998; Boonkerd & Nooteboom, 2001; Boonkerd & Pollawatn 2000, 2002a, 2002b, 2006; Lindsay & Middleton, 2004, 2009c; Lindsay et al., 2004, 2008; Suksathan, 2004; Boonkerd et al., 2004; Boonkerd, 2006). Of particular significance amongst these are Boonkerd & Pollawatn (2000), which is a list of all the pteridophytes in Thailand (including an additional 27 taxa) with distribution maps and many photographs, and Boonkerd et al. (2004), in which many more species were added. A total of 71 fern taxa have been newly recorded or newly described for Thailand since the publication of the Flora of Thailand accounts. This rate of addition of new taxa and new records to a recently completed Flora account reflects the fact that pteridophytes had previously often been neglected on traditional collecting expeditions (and were therefore not available to Tagawa & Iwatsuki) and is testament to the relatively poor state of our taxonomic knowledge of them compared to many angiosperm plant groups. The taxonomy of ferns in Thailand has also been subject to considerable flux due to differences of opinion over the delimitation of families, genera and species. One need only glance at the lists of synonymy to see that very many species have had homotypic combinations made in many different genera (e.g. several species of Selliguea also have combinations in Polypodium, Phymatopsis, Pleopeltis, Phymatodes, Crypsinus

______1 Royal Botanic Garden Edinburgh, 20A Inverleith Row, Edinburgh, EH3 5LR, Scotland, UK. 2 Department of Botany, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. 3 The Forest Herbarium, Department of National Parks, Wildlife and Plant Conservation, Chatuchak, Bangkok 10900, Thailand. Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 65 and Phymatopteris - see Lindsay & Middleton, 2009a). This indecision about the best placement of species in genera is also manifest at the family level where the limits of several families have changed considerably over time and between contemporary Flora projects, leading, in some cases, to the abandonment of families altogether in some parts of Flora Malesiana (e.g. Kramer, 1971; Holttum, 1978, 1991) in favour of “groups” of genera. In recent years phylogenetic techniques, particularly using molecular data, have been used to attempt to solve the many problems in fern familial and generic delimitation (see Smith et al., 2006, 2008 and references cited therein). This has led to a greater consensus on the delimitation of families and genera. We are eventually beginning to see the possibility of stability in the names applied to the ferns of Thailand.

THE TABLES

In this paper we apply the family and genus concepts summarised by Smith et al. (2006, 2008) to the ferns of Thailand and present the data in two tables. For Davalliaceae we follow the more recent generic concepts of Kato & Tsutsumi (2008). In Table 1 the families and genera recognised by Smith et al. (2006, 2008) and Kato & Tsutsumi (2008) are compared to the families and genera recognised in the Flora of Thailand accounts. In Table 2 all fern taxa now known to occur in Thailand are listed using the up-to-date family and genus concepts from Smith et al. (2006, 2008) and Kato & Tsutsumi (2008), with occasional modifications from other recent works (e.g. Parris, 2007). These are compared to the names used for the same taxon from the Flora of Thailand and from Boonkerd & Pollawatn (2000). For species delimitation we follow Tagawa & Iwatsuki (1979, 1985, 1988, 1989), unless other pieces of taxonomic work have superseded these publications (e.g. Hovenkamp et al., 1998), or we have come to our own differing opinion in specific groups. In some cases the authorship of names is not the same as has been previously published. We have checked the problematic ones and only the correct authorships are used without highlighting the discrepancies. Likewise the spelling of specific epithets not in conformity with the International Code of Botanical Nomenclature (McNeill et al., 2006) has been corrected in all columns without comment. We include taxa which have been published as occurring in Thailand. We also include a small number of taxa which we provisionally record for Thailand from our own observations but which still await formal publication. We acknowledge that this list might be incomplete judging by the rate at which taxa have been newly recorded from Thailand in the last 20 years. However, this list has been compiled to facilitate the development of an on-line multi-access key to the ferns of Thailand (Lindsay & Middleton, 2009b) and new taxa can easily be added to this. The “nine new records” in Mitsuta (1985) were not included in the Flora of Thailand. Some of them are, however, attributable to species already known from Thailand. We have included a genuine additional record (Monachosorum henryi Christ) and as yet unconfirmed new records Thelypteris( laxa (Franch. & Sav.) Ching and Diplazium doerderleinii (Luerss.) Makino) in Table 2. The new record Diplazium okinawaense Tagawa has been synonymised to Diplazium virescens Kunze following Shieh et al. (1994). We have redetermined the other material cited in Mitsuta (1985) to taxa already reported in the Flora of Thailand, thus: Colysis flexiloba (Christ) Ching var. undulato- repanda (C.Chr.) Ching to Leptochilus ellipticus (Thunb.) Noot.; Crypsinus trisectus 66 THAI FOREST BULLETIN (BOTANY) 37

(Baker) Tagawa to Selliguea hirsuta (Baker) S.Linds.; Microlepia trichoclada Ching to Microlepia herbacea Ching & C.Chr. ex Tardieu & C.Chr.; Thelypteris angulariloba Ching to Thelypteris hirsutipes (C.B.Clarke) Ching; and Thelypteris esquirolii (Christ) Ching to Cyclosorus falcilobus (Hook.) Panigrahi.

Major changes in family delimitations Table 1 summarises the changes in family delimitation between the Flora of Thailand accounts and those currently recognised and lists the genera in each of the families. These changes include losing some familiar families altogether, such as Parkeriaceae (now in Pteridaceae) and Athyriaceae (now mostly in Woodsiaceae), and changing the delimitation of others quite radically, such as Polypodiaceae (which now includes Grammitidaceae) and Dryopteridaceae (which has lost Tectaria and its relatives and gained genera from Lomariopsidaceae, Davalliaceae and Athyriaceae). Pteridaceae has also undergone major changes and is now much larger with five subfamilies which do not correspond well to former families; e.g. Pteridaceae subfamily Pteridoideae includes only Pteris from the previously delimited Pteridaceae, with all of the other genera previously having been placed in Parkeriaceae.

Major changes in generic delimitation Table 2 lists all the currently used names for Thai ferns and compares them to Pteridophytes in Thailand (Boonkerd & Pollawatn, 2000) and Flora of Thailand (Tagawa & Iwatsuki, 1979, 1985, 1988, 1989). A total of 191 taxa have a name which is now different to the name used in the Flora of Thailand accounts, most of these as a result of changes in generic delimitation. Changes in generic delimitation are particularly extensive in Davalliaceae, Hymenophyllaceae, Polypodiaceae and Thelypteridaceae. Well known genera such as Grammitis, Polypodium, Trichomanes and Vittaria are no longer recognised in Thailand. Table 2 also provides references to the literature on new taxa, new records and in cases where the taxon epithet is different to that used in the Flora of Thailand account (usually due to synonymy). If there is no reference given then the change is made here. Black dots are used to indicate a taxon which is under a different name to that used in the original Flora of Thailand account; circles are used to indicate a taxon which is new to Thailand, either as a newly described taxon or a new record.

ACKNOWLEDGEMENTS

We thank The Leverhulme Trust for financial support for this work; the Curators and Collection Managers of herbaria that have loaned material or welcomed us on visits (notably A, BK, BKF, BM, C, CMU, E, K, SING); the library staff of the Royal Botanic Garden Edinburgh for their help in locating or obtaining protologues and other references; and Christopher Fraser-Jenkins, John McNeill, Barbara Parris, Alan Smith and John Thompson for advice and information. Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 67

), ), ) THYRIACEAE ), A ), RYOPTERIDACEAE , D Leucostegia ( Nephrolepis ( HEIROPLEURIACEAE ) ), C Bolbitis, Elaphoglossum, Lomagramma, Lomariopsis) Acrophorus, Arachniodes, Ctenitis, Acrophorus, ( ( ( AVALLIACEAE Dipteris Blechnum, Brainea, Woodwardia ( Stenochlaena ( ), D L EA ND RA C EAE ( ) ) ), O AE DTI A C EAE ARATTIACEAE SPL E NI A C EAE ICKSONI A C EAE AVALLIACEAE E NNST IPTERIDACEAE RYOPTERIDACEAE L E ICH NI A C EAE YM E NOPHYLL A C EAE L E CHN A C EAE YATHEACEAE OM AR IOPSID A C EAE QUISETACEAE INDS AEA C EAE OM AR IOPSID A C EAE TERIDACEAE CHIZAEACEAE Cheiropleuria Hypodematium Cyclopeltis Corresponding family (or families) in Flora of Thailand families) in Flora of family (or Corresponding & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa A B P D C D D D ( D Cyrtomium, Didymochlaena, Dryopteris, Polystichum ( L Teratophyllum E G H L L ( S M Wibelia Sticherus Vandenboschia , Gleichenia, Ptisana Diplopterygium , Davallia, Davallodes, Humata, , Callistopteris, Cephalomanes, Crepidomanes, Cephalomanes, Callistopteris, , Hymenasplenium , Didymochlaena, Dryopteris, Elaphoglossum, Genera currently recognized in Thailand. Those in Thailand. in recognized Genera currently Thailand in Flora of not recognized bold (26) were Asplenium, Blechnum, Brainea, Stenochlaena, Woodwardia Cibotium Cyathea Araiostegiella Dennstaedtia, Histiopteris, Hypolepis, Microlepia, Pteridium Dipteris Cheiropleuria, Arachniodes, Bolbitis, Ctenitis, Cyrtomium, Acrophorus, Peranema Hypodematium, Leucostegia, Lomagramma, Polystichum, Teratophyllum Equisetum Dicranopteris, Abrodictyum Didymoglossum, Hymenophyllum, Lindsaea, Sphenomeris, Tapeinidium Cyclopeltis, Lomariopsis, Nephrolepis Lygodium Angiopteris, Christensenia, AE DTI A C EAE ARATTIACEAE SPL E NI A C EAE AVALLIACEAE E NNST IPTERIDACEAE RYOPTERIDACEAE L E ICH NI A C EAE YM E NOPHYLL A C EAE L E CHN A C EAE IBOTIACEAE YATHEACEAE QUISETACEAE INDS AEA C EAE OM AR IOPSID A C EAE YGODIACEAE Smith et al. (2006, 2008) family A B C C D D D D E G H L L L M that within Pteridaceae the subfamily name Vittarioideae has priority over Adiantoideae (cf. Smith et al., 2006, 2008) and the subfamilial name Cryptogrammoideae, Cryptogrammoideae, name subfamilial the and 2008) 2006, al., et Smith (cf. Adiantoideae over priority & 1. Iwatsuki, Comparison 1979, of 1985, family 1988, delimitation and 1989). has composition Note between Smith et al. (2006, 2008) Thailand and (Tagawa Table Flora of Vittarioideae name subfamily the Pteridaceae within that which did not exist for Smith et al. (2006, 2008), has now been published (Lindsay & Middleton, 2009a). 68 THAI FOREST BULLETIN (BOTANY) 37

) ) ) ),

Azolla ( Arthropteris RA MMITID A C EAE ( ), G ) ARKERIACEAE ) ITTARIACEAE ZOLLACEAE ), P ), V ), Pteris ), A ), ( Heterogonium, Pleocnemia, Heterogonium, L EA ND RA C EAE ( Aglaomorpha, Arthromeris, Arthromeris, Aglaomorpha, ), O Onychium, Pityrogramma, Syngramma, Onychium, Pityrogramma, Adiantum ( Gymnogrammitis Salvinia ( ( ( Acrostichum ( ( ) TERIDACEAE ), P ARSILEACEAE L EA ND RA C EAE PHIOGLOSSACEAE SM U ND A C EAE RYOPTERIDACEAE THYRIACEAE AVALLIACEAE HELYPTERIDACEAE LAGIOGYRIACEAE OLYPODIACEAE SILOTACEAE ARKERIACEAE ARKERIACEAE TERIDACEAE ARKERIACEAE ARKERIACEAE A C EAE A LVINI CHIZAEACEAE Acrosorus, Calymmodon, Prosaptia, Scleroglossum Calymmodon, Prosaptia, Acrosorus, Ceratopteris Vittaria Vaginularia, Antrophyum, Corresponding family (or families) in Flora of Thailand families) in Flora of family (or Corresponding & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa M Absent O O O P P Belvisia, Christiopteris, Colysis, Drynaria, Lemmaphyllum, Lepisorus, Leptochilus, Loxogramme, Microsorum, Selliguea Platycerium, Pyrrosia, Neocheiropteris, D ( P P P P ( P Taenitis P ( S S D Pteridrys, Tectaria T A ,

Thelypteris Goniophlebium iphopterella Scleroglossum, Scleroglossum, Vaginularia , X , , Platycerium, , , Lemmaphyllum, Drynaria, Chrysogrammitis,

Haplopteris

Tomophyllum , Radiogrammitis Lecanopteris Oreogrammitis Pellaea , eris, Themelium

Genera currently recognized in Thailand. Those in bold Thailand. in recognized Genera currently Thailand in Flora of not recognized (26) were Marsilea Matonia Oleandra Botrychium, Helminthostachys, Ophioglossum Osmunda Plagiogyria Belvisia, Arthromeris, Aglaomorpha, Acrosorus, Calymmodon, Christiopteris, Ctenopterella, Dasygrammitis, Psilotum Cheilanthes, Doryopteris, Hemionitis, Notholaena, Parahemionitis Coniogramme Ceratopteris Acrostichum, Pteris, Syngramma, Taenitis Onychium, Pityrogramma, Antrophyum, Adiantum, Azolla, Salvinia Schizaea Pleocnemia, Pteridrys, Tectaria Heterogonium, Arthropteris, Pseudophegopteris, Macrothelypteris, Cyclosorus, Athyrium, Cornopteris, Deparia, Diplazium Gymnogrammitis, Lepisorus, Leptochilus, Loxogramme, Microsorum, Neocheiropt Pyrrosia, Prosaptia, Selliguea, subfam. Cheilanthoideae Cryptogrammoideae subfam. subfam. Parkerioideae subfam. Pteridoideae Vittarioideae subfam. A C EAE TONI OODSIACEAE ARSILEACEAE A L EA ND RA C EAE PHIOGLOSSACEAE SM U ND A C EAE ECTARIACEAE HELYPTERIDACEAE LAGIOGYRIACEAE OLYPODIACEAE SILOTACEAE TERIDACEAE TERIDACEAE TERIDACEAE TERIDACEAE TERIDACEAE A C EAE A LVINI CHIZAEACEAE Smith et al. (2006, 2008) family M M O O O P P P P P P P P S S T T W Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 69

1 37 Fraser- 11 . Basionym: Nooteboom Hovenkamp Boonkerd & 6 30 17 Adiantum philippense Adiantum

comb. nov R ödl-Linder (1990);

43 36 Christ Lindsay & Middleton (2009c); Middleton & Lindsay Hance 22 Tardieu & Ching Tardieu Wall. ex Hook. & Grev. Wall. Alderw. (Franch.) Copel. Wall. ex Hook. & Grev. & Hook. ex Wall. Nooteboom (1997); Nooteboom Boonkerd et al. (2004); Bedd. 29 10 Sw. (C.Presl) S.Linds., Hovenkamp et al. (1998); 16 Ebihara et al. (2006). Boonkerd & Pollawatn (2002a); 5 47 has been used in recent literature we can find no J. Thompson, pers. comm.; pers. Thompson, J. Nakaike (1992); Nakaike NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Asplenium affine Asplenium antrophyoides Asplenium batuense Asplenium confusum Asplenium crinicaule Absent Asplenium delavayi Asplenium ensiforme Asplenium exiguum Absent Asplenium grevillei 42 28 Lindsay & Middleton (2004); Middleton & Lindsay Petchsri, Boonkerd & Baum (2009); 21 Hovenkamp (1986); 35 Hymenasplenium excisum 15 Boonkerd & Suksathan (2009); 45 9 Murdock (2008); Murdock Hymenasplenium excisum 27 B. Parris, pers. comm.; pers. Parris, B. 41 Boonkerd & Pollawatn (2000); Khullar (1994); Khullar 4 20 Christ taxonomic change by current authors; 46 Hance Wall. ex Hook. & Grev. Wall. Tardieu & Ching Tardieu Alderw. (Franch.) Copel. Parris & Latiff (1997); Bedd. Wall. ex Hook. & Grev. Wall. Mitsuta (1985); Mitsuta 34 Sw. 26 Zhang (1999b); Zhang Holttum & Grimes (1980); 40 14 Boonkerd & Pollawatn (2006); 8 Lu (1999); Lu 25 Parris (2007); Kato & Tsutsumi (2008); Tsutsumi & Kato NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Asplenium affine Asplenium antrophyoides Asplenium batuense Asplenium confusum Asplenium crinicaule Absent Asplenium delavayi Asplenium ensiforme Asplenium exiguum Absent Asplenium grevillei 33 19 Zhang (1999a); Zhang Boonkerd & Nooteboom (2001); 3 39 Holttum (1991); 10 13 Provisional new records, currently under investigation; 44 Parris (1998b); 32 Lindsay et al. (2008); al. et Lindsay 44 24 Wall. ex Hook. Wall. Iwatsuki et al. (1998); al. et Iwatsuki Christ 18 Boonkerd & Pollawatn (2004); Thompson (2008); Thompson 7 Boonkerd (in press); Kunze Hance C.Presl, Epimel. Bot. 74 (1851 [‘1849’]). Although the name Holttum (1982); 38 2 Wall. ex Hook. & Grev. Wall. Tardieu & Ching Tardieu is reported for Thailand by Boonkerd & Pollawatn (2006) but the combination has not been validly published and the status of the varieties of this of varieties the of status the and published validly been not has combination the but (2006) Pollawatn & Boonkerd by Thailand for reported is

(Franch.) Copel. Alderw. Bedd. Wall. ex Hook. & Grev. Wall. 12 Sw. Parris (1998a); 31 subjunonicum Lindsay et al. (2004); al. et Lindsay Flora of Thailand (Tagawa & Iwatsuki, 1979, 1985, 1988, 1989). Two names for a & single Iwatsuki, entry 1979, indicate in that 1985, the a 1988, Two third change 1989). column Flora Thailand was linked of (Tagawa made by a ‘/’ to the name in the “ A dditions and Corrections” section in part 4 of volume 3 of the Flora of Thailand account. or Two more a indicate that ‘+’ names these for taxa are a now single treated as entry synonyms. Black linked dots by indicate species whose names have changed since the publication of the Flora of Thailand or were changed in the “ A dditions and Corrections” section. Circles indicate species that have been discovered or described since the publication of the Flora of The reference Thailand. beside a name refers to the place(s) of publication of the new taxon or record, or the source of information about the change of a name: Boonkerd (2006); Pollawatn (2002b); Jenkins (1997); (2005); Miyamoto & 23 Asplenium excisum evidence that the combination has previously been validly published; (1998); (2004); Suksathan var. species need further research; CURRENT NAME CURRENT ASPLENIACEAE Asplenium affine Asplenium antrophyoides Asplenium batuense Asplenium confusum Asplenium crinicaule Asplenium decrescens Asplenium delavayi Asplenium ensiforme Asplenium exiguum Asplenium finlaysonianum Asplenium grevillei ○ ○ Table 2. Table Comparison of the current names for Thai fern species with the names used for the same taxa in Pteridophytes in Thailand (Boonkerd & Pollawatn, 2000) and

70 THAI FOREST BULLETIN (BOTANY) 37 (Mett.) C.Chr. phyllitidis Sw. Hook. nidus musifolium Blume Bedd. Blume Christ Lam. D.Don. subsp. D.Don. subsp. C.G.Matthew & Christ & C.G.Matthew Tardieu Blume Tagawa & K.Iwats. Tagawa G.Forst. Lam. D.Don. subsp. D.Don Sw. Wall. ex Hook. & Grev. Wall. L. var. L. var. C.Chr. L. var. L. var. Holttum Asplenium phyllitidis NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Absent Absent Asplenium humbertii Absent Asplenium interjectum Absent Asplenium varians Asplenium longissimum Asplenium macrophyllum Asplenium nidus Asplenium nidus Asplenium nitidum Asplenium normale Asplenium paradoxum Asplenium pellucidum Asplenium perakense Asplenium phyllitidis malesicum Asplenium falcatum Asplenium rockii Asplenium salignum Asplenium scortechinii Asplenium siamense Asplenium simonsianum Asplenium tenerum phyllitidis (Mett.) C.Chr. Sw. Hook. nidus musifolium Blume Bedd. Blume Christ Lam. C.G.Matthew & Christ D.Don. subsp. D.Don. subsp. Blume Tardieu Lam. Tagawa & K.Iwats. Tagawa D.Don G.Forst. Sw. Wall. ex Hook. & Grev. Wall. C.Chr. L. var. L. var. L. var. Holttum NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Absent Absent Asplenium humbertii Absent Asplenium interjectum Absent Asplenium varians Asplenium longissimum Asplenium macrophyllum Asplenium nidus Asplenium nidus Asplenium nitidum Asplenium normale Asplenium paradoxum Asplenium pellucidum Asplenium perakense Asplenium phyllitidis Asplenium phyllitidis malesicum Asplenium falcatum Asplenium rockii Asplenium salignum Asplenium scortechinii Asplenium siamense Asplenium simonsianum Asplenium tenerum 9 phyllitidis (Mett.) C.Chr. 2 44 44 20 34 Willd. Sw. Hook. nidus musifolium Hook. Mett. ex Kuhn. Blume Bedd. Blume Christ Lam. Sledge. D.Don. D.Don. subsp. C.G.Matthew & Christ D.Don. subsp. Blume Tardieu

G.Forst. Tagawa & K.Iwats. Tagawa D.Don Sw. Sw. G.Forst. L. var. L. var. C.Chr. C.Chr. L. var. L. var. Holttum CURRENT NAME CURRENT Asplenium griffithianum Asplenium gueinzianum Asplenium humbertii Asplenium inaequilaterale Asplenium interjectum Asplenium khasianum Asplenium laciniatum Asplenium longissimum Asplenium macrophyllum Asplenium nidus Asplenium nidus Asplenium nitidum Asplenium normale Asplenium paradoxum Asplenium pellucidum Asplenium perakense Asplenium phyllitidis Asplenium phyllitidis malesicum Asplenium polyodon Asplenium rockii Asplenium salignum Asplenium scortechinii Asplenium siamense Asplenium simonsianum Asplenium tenerum ○ ○ ○ ○ ● ● Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 71 Ching Wall. ex Hook. & Grev. Wall. Blume Kunze ex Mett. (Burm.f.) Bedd. Makino D.Don N.Murak. & Hatan. auct. non Lam. / Kunze Blume auct. non G.Forst. (L.) J.Sm. L. C.Presl Burm.f. (Hook.) J.Sm. NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Asplenium tenuifolium Asplenium thunbergii Asplenium caudatum Asplenium squamulatum Asplenium yoshinagae Asplenium unilaterale Asplenium apogamum Asplenium cheilosorum Asplenium excisum Absent Asplenium obscurum Blechnum finlaysonianum Blechnum indicum Blechnum orientale Brainea insignis Stenochlaena palustris Absent cochin-chinensis Woodwardia Cibotium barometz N.Murak. & J.Yokoy. Wall. ex Hook. & Grev. Wall. Blume Kunze ex Mett. (Burm.f.) Bedd. Makino Hook. N.Murak. & Hatan. D.Don Kunze (L.f.) Sm. Blume auct. non G.Forst. C.Presl L. (L.) J.Sm. Burm.f. (Hook.) J.Sm. NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Asplenium tenuifolium Asplenium thunbergii Asplenium caudatum Asplenium squamulatum Asplenium yoshinagae Asplenium apogamum Asplenium cheilosorum Asplenium excisum Hymenasplenium inthanonense Asplenium obscurum Blechnum finlaysonianum Blechnum indicum Blechnum orientale Brainea insignis Stenochlaena palustris harlandii Woodwardia japonica Woodwardia Cibotium barometz 18 45 N.Murak. & J.Yokoy. (Kunze ex Mett.) Tagawa (Kunze ex Mett.) 46 4,7 (N.Murak. & Hatan.) (Blume) Tagawa (Blume) 34 (C.Presl) S.Linds. Wall. ex Hook. & Grev. ex Hook. & Grev. Wall. Makino Hook. (Burm.f.) Bedd. D.Don Cav. (L.f.) Sm. Kunze Blume.

L. (L.) J.Sm. Burm.f. (Hook.) J.Sm. CURRENT NAME CURRENT Asplenium tenuifolium Asplenium thunbergii Asplenium truncatum Asplenium vittaeforme Asplenium yoshinagae Hymenasplenium apogamum Nakaike Hymenasplenium cheilosorum Hymenasplenium excisum Hymenasplenium inthanonense Hymenasplenium obscurum BLECHNACEAE Blechnum finlaysonianum Blechnum indicum Blechnum orientale Brainea insignis Stenochlaena palustris harlandii Woodwardia japonica Woodwardia CIBOTIACEAE Cibotium barometz ● ● ● ● ○ ● ○ ● 72 THAI FOREST BULLETIN (BOTANY) 37

lorrainii lorrainii (Hook.) Copel. (Hook.) Blume var. Blume var. Blume var. (C.Chr.) Ching (C.Chr.) Ching (Wall. ex Hook.) Copel. (Wall. (Hook.) Copel. Copel. Wall. ex Hook. Wall. (Wall. ex Hook.) Copel. (Wall. Copel. Tardieu & C.Chr. Tardieu (Wall. ex Hook.) Holttum Hook.) ex (Wall. (G.Forst.) Sw. (Hance) Holttum Absent Absent Araiostegia imbricata Davallodes membranulosum NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Cyathea borneensis Cyathea chinensis Cyathea contaminans Cyathea gigantea Absent Absent Cyathea latebrosa Absent Cyathea podophylla Cyathea spinulosa Araiostegia faberiana Davallia petelotii Davallia solida Davallia trichomanoides trichomanoides Davallia trichomanoides

lorrainii (Hook.) Copel. Blume var. Blume var. Blume var. Ching (C.Chr.) Ching (C.Chr.) (Wall. ex Hook.) Copel. (Wall. Copel. (Hook.) Copel. Wall. ex Hook. Wall. (Wall. ex Hook.) Copel. (Wall. Copel. (Wall. ex Hook.) Holttum (Wall. Tardieu & C.Chr. Tardieu (G.Forst.) Sw. NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Cyathea borneensis Cyathea chinensis Cyathea contaminans Cyathea gigantea Absent Absent Cyathea latebrosa Absent Cyathea podophylla Cyathea spinulosa Araiostegia faberiana Davallia petelotii Davallia solida Davallia trichomanoides trichomanoides Davallia trichomanoides (Hance) Holttum Absent Absent Araiostegia imbricata Davallodes membranulosum 30 lorrainii 44 (Blume) M.Kato & (Hook.) Copel. 44 10 Blume var. Blume var. (C.Chr.) M.Kato & Tsutsumi Tsutsumi M.Kato & (C.Chr.) Blume var. Blume var. (Ching) M.Kato & Tsutsumi (Ching) M.Kato & (Hook.) Copel. (Wall. ex Hook.) Copel. (Wall.

(Hook.) Copel. Mett. R.Br. Copel. Copel. Wall. ex Hook. Wall. (Wall. ex Hook.) Copel. (Wall. Tardieu & C.Chr. & C.Chr. Tardieu (Wall. ex Hook.) Holttum (Wall. (Blume) Copel. (G.Forst.) Sw. (G.Forst.) Sw. 30,19 Davallodes borneense Davallodes hymenophylloides Tsutsumi Davallodes imbricatum Davallodes membranulosum CURRENT NAME CURRENT CYATHEACEAE Cyathea borneensis Cyathea chinensis Cyathea contaminans Cyathea gigantea Cyathea glabra Cyathea hymenodes Cyathea latebrosa Cyathea moluccana Cyathea podophylla Cyathea spinulosa DAVALLIACEAE Araiostegiella faberiana Davallia petelotii Davallia solida Davallia trichomanoides trichomanoides Davallia trichomanoides (Hance) Holttum ○ ● ○ ○ ○ ● ○ Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 73 (Kunze) Copel. (Wall. ex Hook. & (Wall. (Sm.) C.Presl (Wall. ex Hook.) C.Presl (Wall. (Kuhn) Alderw. (Kuhn) Sm. (Wall. ex Hook.) T.Moore ex Hook.) (Wall. (L.f.) M.Kato (Sm.) Desv. / (Sm.) Desv. Ching & C.Chr. ex Ching & C.Chr. T.Moore (D.Don) Copel. (Burm.f.) Mett. ex Kuhn (Blume) C.Presl Blume (Thunb.) Mett. ex Kuhn (Wall. ex Hook. & Grev.) J.Sm. ex Hook. & Grev.) (Wall. (Thunb.) J.Sm. (Sm.) Desv. / (Sm.) Desv. Mett. ex Kuhn (L.f.) J.Sm ex Diels / (Blume) T.Moore / T.Moore (Blume) Pachypleuria angustata NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Araiostegia pseudocystopteris Araiostegia pulchra Davallodes viscidulum Humata angustata / C.Presl Grev.) Davallia corniculata Humata heterophylla Davallia heterophylla Humata pectinata Pachypleuria pectinata Humata repens Pachypleuria repens Humata vestita Pachypleuria vestita Davallia denticulata Davallia divaricata Absent Dennstaedtia scabra Histiopteris incisa Hypolepis punctata calvescens Microlepia firma Microlepia herbacea Microlepia & C.Chr. Tardieu (Kunze) Copel. (Kuhn) Alderw. (Kuhn) (Wall. ex Hook.) C.Presl (Wall. (Wall. ex Hook.) T.Moore ex Hook.) (Wall. (Sm.) Desv. Ching & C.Chr. ex Ching & C.Chr. T.Moore (D.Don) Copel. (Burm.f.) Mett. ex Kuhn Blume (Thunb.) Mett. ex Kuhn (Wall. ex Hook. & Grev.) J.Sm. ex Hook. & Grev.) (Wall. (Thunb.) J.Sm. (Sm.) Desv. Mett. ex Kuhn (L.f.) J.Sm ex Diels (Blume) T.Moore (Blume) NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Araiostegia pseudocystopteris Araiostegia pulchra Davallodes viscidulum Humata angustata Davallia corniculata Humata heterophylla Humata pectinata Humata repens Humata vestita Davallia denticulata Davallia divaricata Absent Dennstaedtia scabra Histiopteris incisa Hypolepis punctata calvescens Microlepia firma Microlepia herbacea Microlepia & C.Chr. Tardieu 10,19 (Kunze) M.Kato & (Wall. ex Hook.) C.Presl (Wall. (Kuhn) Alderw. Alderw. (Kuhn) (Copel.) M.Kato & Tsutsumi (Copel.) M.Kato & Ching & C.Chr. ex Ching & C.Chr. (Wall. ex Hook.) T.Moore T.Moore ex Hook.) (Wall. (Sm.) Desv. (Sm.) Desv.

(D.Don) M.Kato & Tsutsumi (D.Don) M.Kato & (T.Moore) M.Kato & Tsutsumi M.Kato & (T.Moore) ( Burm.f.) M.Kato & Tsutsumi ( Burm.f.) M.Kato & (Thunb.) Mett. ex Kuhn (Blume) M.Kato & Tsutsumi (Blume) M.Kato & (Thunb.) J.Sm. (Sm.) Desv. (Sm.) Desv. (Wall. ex Hook. & Grev.) J.Sm. ex Hook. & Grev.) (Wall. Mett. ex Kuhn (Blume) T.Moore (Blume) (L.f.) Diels CURRENT NAME CURRENT Davallodes pseudocystopteris Tsutsumi Davallodes pulchra Davallodes viscidulum Humata angustata Humata corniculata Humata heterophylla Humata pectinata Humata repens Humata vestita denticulata Wibelia divaricata Wibelia embolostegia Wibelia DENNSTAEDTIACEAE Dennstaedtia scabra Histiopteris incisa Hypolepis punctata calvescens Microlepia firma Microlepia herbacea Microlepia & C.Chr. Tardieu ● ● ● ● ● ○ 74 THAI FOREST BULLETIN (BOTANY) 37 (J. A gardh) Domin (Roxb.) Kuhn Tagawa T.Moore, nom. nud. T.Moore, (Kuhn) Ohwi (Wall. ex Hook.) (Wall. (Blume) C.Presl (D.Don) J.Sm. (L.) T.Moore (L.) (L.) Kuhn subsp. (L.) Kuhn subsp. Alderw. (Thunb.) C.Presl Reinw. Copel. (C.B.Clarke) Bedd. wightianum yarrabense var. var. var. var. NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa hookeriana Microlepia C.Presl kurzii Microlepia platyphylla Microlepia puberula Microlepia ridleyi Microlepia speluncae Microlepia strigosa Microlepia taiwaniana Microlepia trapeziformis Microlepia Absent Absent Pteridium aquilinum aquilinum R .M.Tryon Pteridium aquilinum caudatum bicuspis Cheiropleuria Dipteris conjugata stipellatus Acrophorus Arachniodes assamica

latiusculum wightianum yarrabense (Roxb.) Kuhn T.Moore, nom. nud. T.Moore, (Wall. ex Hook.) (Wall. (Blume) C.Presl Tagawa (D.Don) J.Sm. (Kuhn) Ohwi (L.) T.Moore (L.) (L.) Kuhn var. (L.) Kuhn var. (L.) Kuhn var. (L.) Kuhn var. Alderw. (Thunb.) C.Presl Reinw. Copel. (C.B.Clarke) Bedd. NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) hookeriana Microlepia C.Presl kurzii Microlepia platyphylla Microlepia puberula Microlepia ridleyi Microlepia speluncae Microlepia strigosa Microlepia taiwaniana Microlepia trapeziformis Microlepia Absent Pteridium aquilinum A.Heller ex Underw. (Desv.) Pteridium aquilinum (J. A gardh) R .M.Tryon Pteridium aquilinum Domin bicuspis Cheiropleuria Dipteris conjugata stipellatus Acrophorus Arachniodes assamica

japonicum wightianum

26 4,6,38,42 (Wall. ex J. A gardh) (Wall. (Roxb.) Kuhn Christ (Wall. ex Hook.) (Wall. (Blume) C.Presl Tagawa Tagawa (D.Don) J.Sm. (Kuhn) Ohwi 38,42 (L.) T.Moore T.Moore (L.) (L.) Kuhn subsp. (L.) Kuhn subsp. Alderw. Alderw.

C.Presl (Thunb.) C.Presl Reinw. Reinw. Copel. (C.B.Clarke) Bedd. 42 CURRENT NAME CURRENT hookeriana Microlepia C.Presl kurzii Microlepia platyphylla Microlepia puberula Microlepia ridleyi Microlepia speluncae Microlepia strigosa Microlepia taiwaniana Microlepia trapeziformis Microlepia Monachosorum henryi Pteridium aquilinum (Nakai) Á.Löve & D.Löve Pteridium aquilinum W.C.Shieh (J. A gardh) Pteridium semihastatum S.B.Andrews DIPTERIDACEAE bicuspis Cheiropleuria Dipteris conjugata DRYOPTERIDACEAE nodosus Acrophorus Arachniodes assamica ○ ○ ● ● ● Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 75

sinensis costulata (Ching) Ching (Willd.) K.Iwats. subsp. (Willd.) (Rosenst.) Ching (D.Don) Ching (Christ) Ohwi (Hayata) H.Ito (Fée) Ching (C.Chr. ex Ching) K.Iwats (C.Chr. K.Iwats. (Christ) Ching (C.Presl) Ching ex C.Chr. ex Ching (C.Presl) Ching ex C.Chr. & Tardieu & Ching ex C.Chr. (Bedd.) C.Chr. (C.Presl) Hennipman (Baker) K.Iwats. var. (Baker) K.Iwats. var. (Baker) K.Iwats. var. (C.Presl) Ching ex C.Chr. (Wall. ex Hook. & Grev.) Schott ex Hook. & Grev.) (Wall. Hennipman (Wall. ex Hook. & Grev.) Schott ex Hook. & Grev.) (Wall. virens compacta NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Arachniodes cavalerii Arachniodes chinensis Absent Arachniodes henryi Arachniodes speciosa Arachniodes spectabilis Absent Bolbitis angustipinna Bolbitis appendiculata appendiculata Bolbitis copelandii Bolbitis costata Bolbitis deltigera Bolbitis heteroclita Bolbitis hookeriana Bolbitis scalpturata Bolbitis sinensis Bolbitis sinensis & K.Iwats Tagawa (Hook.) Bolbitis sinuata Bolbitis tonkinensis Bolbitis virens var. Bolbitis virens var. sinensis costulata (Buch.-Ham. ex Hook.) Bolbitis appendiculata (Ogata) Ching vivipara (Ching) Ching (Willd.) K.Iwats. subsp. (Willd.) (Rosenst.) Ching (Christ) Ohwi (D.Don) Ching (Hayata) H.Ito (Fée) Ching (C.Chr. ex Ching) K.Iwats. (C.Chr. K.Iwats. + (C.Presl) Ching ex C.Chr. (Christ) Ching Ching ex C.Chr. & Tardieu & Ching ex C.Chr. (Bedd.) C.Chr. (Baker) K.Iwats. var. (Baker) K.Iwats. var. (Baker) K.Iwats. var. (Baker) K.Iwats. var. (C.Presl) Hennipman (C.Presl) Ching ex C.Chr. (Wall. ex Hook. & Grev.) Schott ex Hook. & Grev.) (Wall. (Wall. ex Hook. & Grev.) Schott ex Hook. & Grev.) (Wall. Hennipman virens compacta NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Arachniodes cavalerii Arachniodes chinensis Absent Arachniodes henryi Arachniodes speciosa Arachniodes spectabilis Arachniodes subreflexipinna Bolbitis angustipinna Bolbitis appendiculata appendiculata Bolbitis copelandii Bolbitis costata Bolbitis deltigera Bolbitis heteroclita Bolbitis hookeriana K.Iwats. subsp. (Willd.) Hennipman Bolbitis scalpturata Bolbitis sinensis Bolbitis sinensis & K.Iwats. Tagawa (Hook.) Bolbitis sinuata Bolbitis tonkinensis Bolbitis virens var. Bolbitis virens var.

4 8,10 sinensis costulata (Ogata) Ching (Ching) Ching (T.Moore) Ching (T.Moore) (Willd.) K.Iwats. subsp. (Willd.) (Rosenst.) Ching (Christ) Ohwi (D.Don) Ching (Hayata) H.Ito

(Christ) Ching (Fée) Ching K.Iwats. (C.Chr. ex Ching) K.Iwats. (C.Chr. (C.Presl) Ching ex C.Chr. (C.Presl) Ching ex C.Chr. Ching ex C.Chr. & Tardieu Tardieu & Ching ex C.Chr. (Bedd.) C.Chr. (Bedd.) C.Chr. (Baker) K.Iwats. var. (Baker) K.Iwats. var. (Baker) K.Iwats. var. (Baker) K.Iwats. var. (C.Presl) Ching ex C.Chr. (C.Presl) Ching ex C.Chr. (C.Presl) Hennipman Hennipman (Wall. ex Hook. & Grev.) Schott ex Hook. & Grev.) (Wall. Schott ex Hook. & Grev.) (Wall. virens compacta CURRENT NAME CURRENT Arachniodes cavalerii Arachniodes chinensis Arachniodes coniifolia Arachniodes henryi Arachniodes speciosa Arachniodes spectabilis Arachniodes subreflexipinna Bolbitis angustipinna Bolbitis appendiculata appendiculata Bolbitis copelandii Bolbitis costata Bolbitis deltigera Bolbitis heteroclita Bolbitis hookeriana Bolbitis scalpturata Bolbitis sinensis Bolbitis sinensis & K.Iwats. Tagawa (Hook.) Bolbitis sinuata Bolbitis tonkinensis Bolbitis virens var. Bolbitis virens var. ○ ○ 76 THAI FOREST BULLETIN (BOTANY) 37 (Sw.) J.Sm. (Sw.) Ching (C.Hope) C.Chr. (Blume) T.Moore (Blume) Holttum Tagawa & K.Iwats. Tagawa Ching Ching (Baker) C.Chr. (Bedd.) C.Chr. C.Chr. (Blume) Ching (D.Don) C.Chr. Mitsuta ex Darnaedi, (Christ) Holttum J.Sm. (Blume) Kuntze Ching Tagawa & K.Iwats. Tagawa (Bedd.) Ching (D.Don) Kuntze Rosenst. auct. non (C.Hope) Ching / (Kunze) Ching NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Ctenitis dumrongii Ctenitis mannii Ctenitis subobscura Ctenitis vilis Cyrtomium fortunei Didymochlaena truncatula Dryopteris cochleata Absent Dryopteris gymnophylla Arachniodes hasseltii Dryopteris hendersonii Dryopteris hirtipes Dryopteris integriloba Dryopteris neoassamensis Dryopteris neochrysocoma Dryopteris polita Dryopteris porosa Dryopteris pseudosparsa Dryopteris rheophila M.Kato & K.Iwats. Dryopteris scottii Dryopteris sparsa Dryopteris subtriangularis Elaphoglossum angulatum Elaphoglossum dumrongii Elaphoglossum malayense ex Darnaedi,

Ching (C.Hope) C.Chr. Holttum (Sw.) J.Sm. (Sw.) (Blume) T.Moore (Blume) Tagawa & K.Iwats. Tagawa Ching Ching (Baker) C.Chr. (Bedd.) C.Chr. C.Chr. (D.Don) C.Chr. Mitsuta (Blume) Ching (Christ) Holttum J.Sm. (Blume) Kuntze Tagawa & K.Iwats. Tagawa Ching (D.Don) Kuntze (Bedd.) Ching Rosenst. (Kunze) Ching NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Ctenitis dumrongii Ctenitis subobscura Ctenitis vilis Cyrtomium fortunei Didymochlaena truncatula Dryopteris cochleata Absent Dryopteris gymnophylla Arachniodes hasseltii Dryopteris hendersonii Dryopteris hirtipes Dryopteris integriloba Dryopteris neoassamensis Dryopteris neochrysocoma Dryopteris polita Dryopteris porosa Dryopteris pseudosparsa Dryopteris rheophila M.Kato & K.Iwats. Dryopteris scottii Dryopteris sparsa Dryopteris subtriangularis Elaphoglossum angulatum Elaphoglossum dumrongii Elaphoglossum malayense 10 (Blume) T.Moore T.Moore (Blume) Ching (C.Hope) C.Chr. (C.Hope) C.Chr. (Sw.) J.Sm. (Sw.) Holttum Tagawa & K.Iwats. Tagawa Ching Ching (Baker) C.Chr. (Baker) C.Chr. (Bedd.) C.Chr. (Bedd.) C.Chr. C.Chr. C.Chr. (D.Don) C.Chr. (D.Don) C.Chr. Mitsuta ex Darnaedi, (Baker) C.Chr.

(Christ) Holttum J.Sm. (Blume) C.Chr. (Blume) Kuntze Tagawa & K.Iwats. Tagawa (D.Don) Kuntze Ching Rosenst. (Bedd.) Ching Kunze) Ching CURRENT NAME CURRENT Ctenitis dumrongii Ctenitis subobscura Ctenitis vilis ( Cyrtomium fortunei Didymochlaena truncatula Dryopteris cochleata Dryopteris diffracta Dryopteris gymnophylla Dryopteris hasseltii Dryopteris hendersonii Dryopteris hirtipes Dryopteris integriloba Dryopteris neoassamensis Dryopteris neochrysocoma Dryopteris polita Dryopteris porosa Dryopteris pseudosparsa Dryopteris rheophila M.Kato & K.Iwats. Dryopteris scottii Dryopteris sparsa Dryopteris subtriangularis Elaphoglossum angulatum Elaphoglossum dumrongii Elaphoglossum malayense ● ○ ● Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 77 (Tagawa) (Tagawa) (Blume) T.Moore (Blume) Rosenst. Holttum (Fée) T.Moore (Fée) (Wall. ex Baker) (Wall. (Yatabe) Makino (Yatabe) Scort. ex Ridl. (Blume) Mett. ex Kuhn (Forssk.) Kuhn (Blume) T.Moore (Blume) Rosenst. Tagawa & K.Iwats. Tagawa Alderw. (C.B.Clarke) Ching Blume (Fée) Holttum (Kuhn) C.Chr. (Wall. ex Hook.) C.Presl (Wall. A lston & Bonner NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Elaphoglossum marginatum Elaphoglossum melanostictum Elaphoglossum stelligerum ex T.Moore Elaphoglossum subellipticum Elaphoglossum yoshinagae Hypodematium crenatum Hypodematium glanduloso-pilosum Ohwi Leucostegia immersa Lomagramma grossoserrata Diacalpe aspidioides Polystichum attenuatum Polystichum biaristatum Polystichum eximium Polystichum lindsaeifolium Polystichum prolificans Absent Absent Polystichum semifertile Polystichum tenggerense aculeatum Teratophyllum ludens Teratophyllum Absent (Tagawa) (Tagawa) (Blume) T.Moore (Blume) Rosenst. Holttum (Fée) T.Moore (Fée) (Yatabe) Makino (Yatabe) (Wall. ex Baker) (Wall. Scort. ex. Ridl. (Blume) Mett. ex Kuhn (Forssk.) Kuhn (Blume) T.Moore (Blume) Rosenst. Tagawa & K.Iwats. Tagawa Alderw. (C.B.Clarke) Ching (Fée) Holttum (Kuhn) C.Chr. (Wall. ex Hook.) C.Presl (Wall. Blume D.Don A lston & Bonner NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Elaphoglossum marginatum Elaphoglossum melanostictum Elaphoglossum stelligerum ex T.Moore Elaphoglossum subellipticum Elaphoglossum yoshinagae Hypodematium crenatum Hypodematium glanduloso-pilosum Ohwi Leucostegia immersa Lomagramma grossoserrata Diacalpe aspidioides Polystichum attenuatum Polystichum biaristatum Polystichum eximium Polystichum lindsaeifolium Polystichum prolificans Absent Absent Polystichum semifertile Polystichum tenggerense aculeatum Teratophyllum ludens Teratophyllum Equisetum diffusum 8,10 8 (Tagawa) (Tagawa) Ching (Blume) T.Moore T.Moore (Blume) Rosenst. Holttum (Fée) T.Moore T.Moore (Fée) Scort. ex Ridl. (Yatabe) Makino (Yatabe) (Wall. ex Baker) (Wall. 4 Rosenst. (Blume) Mett. ex Kuhn (Forssk.) Kuhn Tagawa & K.Iwats. Tagawa (Blume) T.Moore T.Moore (Blume) Alderw. Alderw. (C.B.Clarke) Ching ( R oxb.) C.V.Morton (Blume) Mett. (Wall. ex Hook.) C.Presl (Wall. (Fée) Holttum (Kuhn) C.Chr. (Kuhn) C.Chr.

D.Don A lston & Bonner CURRENT NAME CURRENT Elaphoglossum marginatum Elaphoglossum melanostictum Elaphoglossum stelligerum ex T.Moore Elaphoglossum subellipticum Elaphoglossum yoshinagae Hypodematium crenatum Hypodematium glanduloso-pilosum Ohwi Leucostegia immersa Lomagramma grossoserrata Peranema aspidioides Polystichum attenuatum Polystichum biaristatum Polystichum eximium Polystichum lindsaeifolium Polystichum prolificans Polystichum pseudotsus-simense Polystichum scariosum Polystichum semifertile Polystichum tenggerense aculeatum Teratophyllum ludens Teratophyllum EQUISETACEAE Equisetum diffusum ● ○ ○ ○ 78 THAI FOREST BULLETIN (BOTANY) 37 (J.Sm. ex Baker) (Blume) Copel. (Hand.-Mazz.) Tagawa Tagawa (Hand.-Mazz.) (C.Presl) Holttum R.Br. (Burm.f.) Underw. (Burm.f.) Underw. Copel. (J.Sm. ex Baker) Copel. Blume (Burm.f.) Underw. (Burm.f.) Underw. (Burm.f.) Underw. (Willd.) Spreng. (Willd.) C.Chr. Mett. Roxb. ex Vaucher Roxb. ex (Christ) Holttum ( R osenst.) Nakai linearis subpectinata tetraphylla Cephalomanes gemmatum NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Equisetum debile Dicranopteris curranii Dicranopteris linearis var. Absent Dicranopteris linearis var. Dicranopteris linearis var. Dicranopteris speciosa Dicranopteris splendida Gleichenia blotiana Gleichenia longissima Gleichenia norrisii Gleichenia microphylla Absent Gleichenia truncata gemmata Macroglena / K.Iwats. Selenodesmium obscurum (J.Sm. ex Baker) (Blume) K.Iwats. (Hand.-Mazz.) Tagawa Tagawa (Hand.-Mazz.) R.Br. (C.Presl) Holttum (Burm.f.) Underw. (Burm.f.) Underw. (Burm.f.) Underw. Copel. Blume (Burm.f.) Underw. (Burm.f.) Underw. (Burm.f.) Underw. (Willd.) Spreng. (Willd.) C.Chr. Mett. Roxb. ex Vaucher Roxb. ex (Christ) Holttum ( R osenst.) Nakai Holttum montana subpectinata tetraphylla obscurum linearis NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Equisetum debile Dicranopteris curranii Dicranopteris linearis var. Dicranopteris linearis var. Dicranopteris linearis var. Dicranopteris linearis var. Dicranopteris speciosa Dicranopteris splendida Gleichenia blotiana Gleichenia longissima Gleichenia norrisii Gleichenia microphylla Absent Gleichenia truncata Cephalomanes gemmatum K.Iwats. Cephalomanes obscurum var. (Roxb. ex debile 10 (Blume) Nakai (C.Chr.) Nakai (C.Chr.) Desf. subsp. (Hand.-Mazz.) Tagawa Tagawa (Hand.-Mazz.) (Mett.) Nakai (C.Presl) Holttum R.Br. R.Br. (Blume) Ebihara & K.Iwats. (Burm.f.) Underw. (Burm.f.) Underw. (Burm.f.) Underw. Copel. (C.V.Morton) (C.V.Morton) (Burm.f) Underw. (Burm.f) Underw. (Burm.f.) Underw.

4

47 (Willd.) Nakai (Willd.) (Christ) Holttum (Blume) Ching ( R osenst.) Nakai Holttum linearis montana subpectinata tetraphylla obscurum CURRENT NAME CURRENT Equisetum ramosissimum Hauke Vaucher) GLEICHENIACEAE Dicranopteris curranii Dicranopteris linearis var. Dicranopteris linearis var. Dicranopteris linearis var. Dicranopteris linearis var. Dicranopteris speciosa Dicranopteris splendida Diplopterygium blotianum Diplopterygium longissimum Diplopterygium norrisii Gleichenia microphylla Sticherus hirtus Sticherus truncatus HYMENOPHYLLACEAE idoneum Abrodictyum Ebihara & K.Iwats. obscurum Abrodictyum var. ● ○ ● ● ● ○ ● ● ● Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 79 (Baker) (Blume) Trichomanes Trichomanes (Copel.) Copel. (D.C.Eaton) (C.Presl) Bosch + Bosch / (Poir.) Copel. + (Poir.) (Bory ex Willd.) Willd.) (Bory ex (Bosch) Copel. & Tagawa (Baker) (Blume) Copel., (Blume) C.Presl (C.Presl) K.Iwats. (Nees & Blume) Copel. Blume) & (Nees (Blume) K.Iwats. Tagawa & K.Iwats. / Tagawa (C.Presl) Copel. (C.Presl) Copel. / (Copel.) Copel. (Christ) K.Iwats. (G.Forst.) Bosch (Bory ex Willd.) Copel. Willd.) (Bory ex (Blume) Prantl + (Bedd.) Tagawa & K.Iwats. Tagawa (Bedd.) (G.Forst.) Pic.Serm. / (Bosch) Bosch siamense Cephalomanes obscurum Crepidomanes parvifolium Crepidomanes Cephalomanes meifolium NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Selenodesmium obscurum pro parte / K.Iwats. var. meifolia Macroglena / K.Iwats. Callistopteris apiifolia Cephalomanes apiifolium Cephalomanes javanicum bipunctatum Crepidomanes bilabiatum Crepidomanes brevipes Crepidomanes christii Crepidomanes Reediella humilis humile Crepidomanes kurzii Crepidomanes latealatum Crepidomanes latemarginale Crepidomanes Copel. megistostomum Crepidomanes Gonocormus prolifer Gonocormus saxifragoides Gonocormus siamensis minutum Crepidomanes parvifolium Microgonium K.Iwats. / K.Iwats. bimarginatum Microgonium bimarginatum (Copel.) Copel. (D.C.Eaton) (Bosch) Bosch (Poir.) Copel. (Poir.) (Baker) K.Iwats. (Blume) C.Presl (Bosch) Copel. (C.Presl) K.Iwats. (Bory ex Willd.) K.Iwats. Willd.) (Bory ex (Blume) K.Iwats. (Blume) K.Iwats. (C.Presl) Copel. (G.Forst.) Bosch (Copel.) Copel. (Bedd.) Tagawa & K.Iwats. Tagawa (Bedd.) (Christ) K.Iwats. siamense NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Cephalomanes obscurum var. Cephalomanes meifolium Cephalomanes apiifolium Cephalomanes javanicum bipunctatum Crepidomanes Crepidomanes brevipes Crepidomanes christii Crepidomanes humile Crepidomanes kurzii Crepidomanes latealatum Crepidomanes latemarginale Crepidomanes Copel. megistostomum Crepidomanes minutum Crepidomanes parvifolium Crepidomanes bimarginatum Trichomanes

47 (Copel.) Copel. (Bosch) Ebihara & (D.C.Eaton) (Poir.) Copel. (Poir.) (Baker) K.Iwats. (Blume) C.Presl (Bosch) Copel. (Blume) K.Iwats. (C.Presl) Copel. (C.Presl) Copel. (Blume) Ebihara & K.Iwats. (Copel.) Copel. (G.Forst.) Bosch (Bedd.) Tagawa & K.Iwats. Tagawa (Bedd.)

(Hook.) Ebihara & K.Iwats. (Christ) K.Iwats. siamense CURRENT NAME CURRENT obscurum Abrodictyum var. pluma Abrodictyum Callistopteris apiifolia Cephalomanes javanicum bipunctatum Crepidomanes Crepidomanes brevipes Crepidomanes christii Crepidomanes humile Crepidomanes kurzii Crepidomanes latealatum Crepidomanes latemarginale Crepidomanes Copel. megistostomum Crepidomanes minutum Crepidomanes parvifolium Crepidomanes Didymoglossum bimarginatum K.Iwats. ● ● ● ● ● ● ● 80 THAI FOREST BULLETIN (BOTANY) 37 Trichomanes Trichomanes Sw. Sw. (Bosch) Rosenst. Copel. (Sw.) Copel. / (Sw.) (Bosch) C.Chr. (Bosch) C.Chr. (Sw.) Sw. Sw. (Sw.) (Müll.Berol.) Bosch / Spreng. (Bosch) Baker Müll.Berol. Wall. ex Hook. Wall. Racib. (Bedd.) Copel. / Hook. & Grev. (Sw.) Copel. / (Sw.) (Sw.) K.Iwats. (Sw.) (Bosch) Copel. / (Blume) K.Iwats. (Bosch) Copel. / (Blume) C.Presl. / (Copel.) Copel. / (Bedd.) Baker (Sw.) Copel. / (Sw.) Bosch / (Spreng.) Copel. / (Racib.) Copel. / (Wall. ex Hook.) Copel. / (Wall. (Hook. & Grev.) Copel. / (Hook. & Grev.) (Bosch) Bosch NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Didymoglossum exiguum exiguum Trichomanes motleyi Microgonium motleyi sublimbatum Microgonium sublimbatum Trichomanes Meringium acanthoides Hymenophyllum acanthoides Mecodium badium Hymenophyllum badium Hymenophyllum barbatum Meringium blandum Hymenophyllum blandum Meringium bontocense Hymenophyllum bontocense Meringium denticulatum Hymenophyllum denticulatum digitatum Microtrichomanes digitatum Crepidomanes Mecodium exsertum Hymenophyllum exsertum Meringium holochilum Hymenophyllum holochilum Mecodium javanicum Hymenophyllum javanicum pallidum Pleuromanes pallidum Crepidomanes Mecodium polyanthos Hymenophyllum polyanthos Sw. (Bosch) Rosenst. (Sw.) Sw. (Sw.) (Bosch) C.Chr. Copel. Spreng. (Bosch) Baker Müll.Berol. Wall. ex Hook. Wall. Racib. (Sw.) K.Iwats. (Sw.) Hook. & Grev. (Blume) K.Iwats. (Bedd.) Baker (Bosch) Bosch NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) exiguum Trichomanes motleyi Trichomanes sublimbatum Trichomanes Hymenophyllum acanthoides Hymenophyllum badium Hymenophyllum barbatum Hymenophyllum blandum Hymenophyllum bontocense Hymenophyllum denticulatum digitatum Crepidomanes Hymenophyllum exsertum Hymenophyllum holochilum Hymenophyllum javanicum pallidum Crepidomanes Hymenophyllum polyanthos Sw. Sw. (Bosch) Rosenst. (Müll.Berol.) E bihara & (Bosch) C.Chr. (Bosch) C.Chr. Copel. (Sw.) Sw. Sw. (Sw.) Spreng. (Bosch) Baker (Sw.) Fosberg Fosberg (Sw.) Wall. ex Hook. Wall. (Blume) Ebihara & K.Iwats. Racib. (Bedd.) Copel. Hook. & Grev. Hook. & Grev. (Bosch) Ebihara & K.Iwats.

CURRENT NAME CURRENT Didymoglossum exiguum Didymoglossum motleyi Didymoglossum sublimbatum K.Iwats. Hymenophyllum acanthoides Hymenophyllum badium Hymenophyllum barbatum Hymenophyllum blandum Hymenophyllum bontocense Hymenophyllum denticulatum Hymenophyllum digitatum Hymenophyllum exsertum Hymenophyllum holochilum Hymenophyllum javanicum Hymenophyllum pallidum Hymenophyllum polyanthos ● ● ● ● ● ● ● ● ● ● ● ● ● Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 81 Crepidomanes Crepidomanes Crepidomanes Crepidomanes Crepidomanes Crepidomanes Christ (C.Presl) C.Chr. Kunze Blume / Bedd. / C.Presl / Blume / Dryand. (Kunze) Copel. / (Christ) Copel. / K.U.Kramer Holttum Blume Christ Hook. & Grev. Sw. (Willd.) Sw. (Willd.) Ching Alderw. Holttum Blume (Blume) K.Iwats. (Bedd.) K.Iwats. (Blume) K.Iwats. NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Mecodium productum Hymenophyllum productum Mecodium riukiuense Hymenophyllum riukiuense Meringium meyenianum Hymenophyllum serrulatum auriculatum Trichomanes auriculatum birmanicum Trichomanes birmanicum maximum Trichomanes maximum Lindsaea bouillodii Lindsaea chienii Lindsaea cultrata Absent Lindsaea divergens Lindsaea doryphora Lindsaea ensifolia Lindsaea heterophylla Lindsaea integra Lindsaea javanensis Lindsaea lucida Lindsaea malayensis Lindsaea napaea (C.Presl) C.Chr. Kunze Christ (Blume) K.Iwats. (Bedd.) K.Iwats. (Blume) K.Iwats. Dryand. K.U.Kramer Holttum Blume Christ Hook. & Grev. Sw. (Willd.) Sw. (Willd.) Holttum Alderw. Blume Ching NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Hymenophyllum productum Hymenophyllum riukiuense Hymenophyllum serrulatum auriculatum Crepidomanes birmanicum Crepidomanes maximum Crepidomanes Lindsaea bouillodii Lindsaea chienii Lindsaea cultrata Absent Lindsaea divergens Lindsaea doryphora Lindsaea ensifolia Lindsaea heterophylla Lindsaea integra Lindsaea javanensis Lindsaea lucida Lindsaea malayensis Lindsaea napaea 10 Christ (C.Presl) C.Chr. (C.Presl) C.Chr. Kunze (Bedd.) Ching (Blume) Copel. Ching (Blume) Copel. Dryand. Holttum

K.U.Kramer Blume Christ Hook. & Grev. Sw. Sw. (Willd.) Sw. Sw. (Willd.) Holttum Alderw. Alderw. Ching Blume CURRENT NAME CURRENT Hymenophyllum productum Hymenophyllum riukiuense Hymenophyllum serrulatum auriculata Vandenboschia birmanica Vandenboschia maxima Vandenboschia LINDSAEACEAE Lindsaea bouillodii Lindsaea chienii Lindsaea cultrata Lindsaea dissectiformis Lindsaea divergens Lindsaea doryphora Lindsaea ensifolia Lindsaea heterophylla Lindsaea integra Lindsaea javanensis Lindsaea lucida Lindsaea malayensis Lindsaea napaea ● ● ● ● ● ● ○ 82 THAI FOREST BULLETIN (BOTANY) 37 pectinata Alderw. (Sw.) Kunze (Sw.) (Hook.) K.U.Kramer (L.) Maxon (L.) Maxon Maxon (L.) Alderw. (L.) C.Presl (Cav.) C.Chr. (Cav.) (Desv.) Christ (Desv.) (Sw.) Schott (Sw.) (G.Forst.) C.Presl (Burm.f.) Kuhn (Lam.) Mett. ex Kuhn (C.Presl) Holttum (Lam.) Mett. ex Kuhn (Fée) C.Chr. (Roxb. ex Griff.) Hieron. (Roxb. ex Griff.) auct. non (Cav.) C.Chr. auct. non (Cav.) Roxb. (Bory) Thwaites var. Thwaites var. (Bory) (Christ) K.U.Kramer (Tagawa) K.U.Kramer (Tagawa) commixta chinensis divaricata divaricata orbiculata NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Lindsaea oblanceolata Lindsaea odorata Lindsaea orbiculata var. Lindsaea orbiculata var. Lindsaea parallelogramma Lindsaea parasitica Lindsaea repens (Blume) Mett. ex Kuhn Absent Sphenomeris chinensis var. Sphenomeris chinensis var. luzonicum Tapeinidium pinnatum Tapeinidium Cyclopeltis crenata Lomariopsis lineata acutifolia Nephrolepis biserrata Nephrolepis cordifolia Nephrolepis davallioides Nephrolepis falcata Nephrolepis hirsutula Nephrolepis radicans Nephrolepis pectinata Alderw. (Sw.) Kunze (Sw.) (Hook.) K.U.Kramer Alderw. (L.) Maxon (L.) Maxon (L.) C.Presl (Cav.) C.Chr. (Cav.) (Desv.) Christ (Desv.) (Sw.) Schott (Sw.) (G.Forst.) C.Presl (Burm.f.) Kuhn (Lam.) Mett. ex Kuhn (Roxb. ex Griff.) Hieron. (Roxb. ex Griff.) (Lam.) Mett. ex Kuhn (C.Presl) Holttum auct. non (Cav.) C.Chr. auct. non (Cav.) (Fée) C.Chr. Roxb. (Bory) Thwaites var. Thwaites var. (Bory) Dryand. (Christ) K.U.Kramer (Tagawa) K.U.Kramer (Tagawa) orbiculata commixta chinensis divaricata Tapeinidium luzonicum Tapeinidium pinnatum Tapeinidium Cyclopeltis crenata Lomariopsis lineata acutifolia Nephrolepis biserrata Nephrolepis cordifolia Nephrolepis davallioides Nephrolepis falcata Nephrolepis hirsutula Nephrolepis radicans Nephrolepis NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Lindsaea oblanceolata Lindsaea odorata Lindsaea orbiculata var. Lindsaea orbiculata var. Lindsaea parallelogramma Lindsaea parasitica Lindsaea repens (Blume) Mett. ex Kuhn Lindsaea tenera Sphenomeris chinensis var. Sphenomeris chinensis var. pectinata 16 Alderw. Alderw. (Sw.) Kunze (Sw.) 4 J.Sm. (Hook.) K.U.Kramer Alderw. Alderw. (L.) C.Presl (L.) Maxon (L.) Maxon (Cav.) C.Chr. C.Chr. (Cav.) (Desv.) Christ (Desv.) (Sw.) Schott (Sw.) (G.Forst.) C.Presl (Burm.f.) Kuhn

(Lam.) Mett. ex Kuhn (Lam.) Mett. ex Kuhn (C.Presl) Holttum (Fée) C.Chr. (Fée) C.Chr. (Roxb. ex Griff.) Hieron. (Roxb. ex Griff.) Roxb. (Bory) Thwaites var. Thwaites var. (Bory) Dryand. (Christ) K.U.Kramer (Tagawa) K.U.Kramer (Tagawa) ommixta orbiculata chinensis divaricata CURRENT NAME CURRENT Lindsaea oblanceolata Lindsaea odorata Lindsaea orbiculata var. Lindsaea orbiculata c var. Lindsaea parallelogramma Lindsaea parasitica Lindsaea repens (Blume) Mett. ex Kuhn Lindsaea tenera Sphenomeris chinensis var. Sphenomeris chinensis var. luzonicum Tapeinidium pinnatum Tapeinidium LOMARIOPSIDACEAE Cyclopeltis crenata Lomariopsis lineata acutifolia Nephrolepis biserrata Nephrolepis cordifolia Nephrolepis davallioides Nephrolepis falciformis Nephrolepis hirsutula Nephrolepis radicans Nephrolepis ● ○ Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 83 (Blume) Maxon Wall. ex T.Moore ex Wall. (Cav.) R.Br. (Cav.) (Decne.) Pic.Serm. C.Presl (Burm.f.) Sw. Tagawa & K.Iwats. Tagawa (Thunb.) Sw. (L.) Sw. Blume (Blume) C.Presl (G.Forst.) Hoffm. C.Presl musifolia

NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa delicatula Nephrolepis circinatum Lygodium flexuosum Lygodium giganteum Lygodium japonicum Lygodium microphyllum Lygodium polystachyum Lygodium salicifolium Lygodium Absent Angiopteris evecta Christensenia aesculifolia Marattia sambucina Marsilea crenata Absent Absent Oleandra (Blume) Maxon (Cav.) R.Br. (Cav.) Wall. ex T.Moore ex Wall. (Decne.) Pic.Serm. C.Presl (Burm.f.) Sw. Tagawa & K.Iwats. Tagawa (Thunb.) Sw. (L.) Sw. Blume (Blume) C.Presl (G.Forst.) Hoffm. C.Presl NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) delicatula Nephrolepis circinatum Lygodium flexuosum Lygodium giganteum Lygodium japonicum Lygodium microphyllum Lygodium polystachyum Lygodium salicifolium Lygodium Absent Angiopteris evecta Christensenia aesculifolia Marattia sambucina Marsilea crenata Absent Absent Oleandra musifolia 27 16 44 10 (Blume) Maxon (Cav.) R.Br. R.Br. (Cav.) 23 Wall. ex T.Moore T.Moore ex Wall. C.Presl C.Presl ( A fzel.) J.Sm. (Thunb.) Sw. (Thunb.) Sw. (Burm.f.) Sw. (Burm.f.) Sw. Tagawa & K.Iwats. Tagawa

(L.) Sw. (L.) Sw. D.M.Johnson (Blume) C.Presl (Blume) Murdock (G.Forst.) Hoffm. (G.Forst.) Hoffm. R.Br. C.Presl CURRENT NAME CURRENT undulata Nephrolepis LYGODIACEAE circinatum Lygodium flexuosum Lygodium giganteum Lygodium japonicum Lygodium microphyllum Lygodium polystachyum Lygodium salicifolium Lygodium MARATTIACEAE Angiopteris angustifolia Angiopteris evecta Christensenia aesculifolia Ptisana sambucina MARSILEACEAE Marsilea crenata Marsilea scalaripes MATONIACEAE Matonia pectinata OLEANDRACEAE Oleandra musifolia ● ○ ● ○ ○ 84 THAI FOREST BULLETIN (BOTANY) 37

(L.) Hook. Willd. Hook. Wall. ex Hook. & Grev. ex Hook. & Wall. L. R.Br. L. (Scort. ex Bedd.) Copel. Ching Ching lanuginosum

(Sw.) C.Chr. (Sw.) (Blume) Bedd. (Willd.) Ching (Willd.) (Hook.) C.Presl Hook. zeylanica

costatum gramineum pendulum petiolatum

adnata communis lanuginosum

triangularis angustifolia cinnamomea vachellii pistillaris undulata wallichii

Japanobotrychium NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Oleandra Oleandra Oleandra Botrychium / Tagawa ex Nishida Grev.) & Hook. ex (Wall. Helminthostachys Ophioglossum Ophioglossum Ophioglossum Ophioglossum Osmunda Osmunda Absent Osmunda Plagiogyria Plagiogyria Acrosorus (L.) Hook. Willd. Hook. Wall. ex Hook. & Grev. Wall. L. R.Br. L. (Scort. ex Bedd.) Copel. Ching Ching (Sw.) C.Chr. (Sw.) (Blume) Bedd. (Willd.) Ching (Willd.) (Hook.) C.Presl (C.Presl) Blume Hook. zeylanica

costatum gramineum pendulum petiolatum

adnata communis lanuginosum

triangularis angustifolia cinnamomea javanica vachellii pistillaris undulata wallichii

NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Oleandra Oleandra Oleandra Botrychium Helminthostachys Ophioglossum Ophioglossum Ophioglossum Ophioglossum Osmunda Osmunda Osmunda Osmunda Plagiogyria Plagiogyria Acrosorus 31 4,10 (L.) Hook. (Christ) Copel. Willd. Willd. Hook. Wall. ex Hook. & Grev. ex Hook. & Grev. Wall. L. R.Br. R.Br. L. Ching Ching

(Sw.) C.Chr. C.Chr. (Sw.) (Hook.) C.Presl (Blume) Bedd. (Willd.) Ching (Willd.) (C.Presl) Blume Hook. adnata communis

friderici-et-pauli cinnamomea javanica vachellii

CURRENT NAME CURRENT Oleandra pistillaris Oleandra undulata Oleandra wallichii OPHIOGLOSSACEAE Botrychium lanuginosum Helminthostachys zeylanica Ophioglossum costatum Ophioglossum gramineum Ophioglossum pendulum Ophioglossum petiolatum OSMUNDACEAE Osmunda angustifolia Osmunda Osmunda Osmunda PLAGIOGYRIACEAE Plagiogyria Plagiogyria POLYPODIACEAE Acrosorus ○ ● Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 85 Tagawa & K.Iwats. Tagawa Tagawa & K.Iwats. Tagawa (Christ) Ching (Willd.) C.V.Morton (Willd.) (Franch. & Bureau auct. non (Hook.) Christ (Wall. ex Mett.) Copel. (Wall. Copel. (Nees. & Blume) Kunze (Mett.) Ching (Copel.) C.Chr. & Tardieu & (Copel.) C.Chr. (C.Chr.) Tagawa (C.Chr.) (Fée) Copel. (Blume) Copel. (Hieron. ex C.Chr.) Tagawa (Hieron. ex C.Chr.) Christ coronans acuminata

amplexifolia lehmanni phuluangensis

NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Absent Photinopteris Aglaomorpha Absent Absent Arthromeris Arthromeris Arthromeris Absent tatsienensis Arthromeris ex Christ) Ching Belvisia annamensis Belvisia henryi Belvisia mucronata Belvisia revoluta Calymmodon asiaticus Calymmodon cucullatus (Nees & Blume) C.Presl Christiopteris tricuspis Absent Ctenopteris moultonii Xiphopteris khaoluangensis Absent Ctenopteris mollicoma Drynaria bonii Tagawa & K.Iwats. Tagawa Tagawa & K.Iwats. Tagawa (Christ) Ching (Franch. & Bureau auct. non (Wall. ex Mett.) Copel. (Wall. (Hook.) Christ (Kunze) Copel. Copel. (Blume) C.Presl (Nees. & Blume) Kunze (Mett.) Ching (Copel.) C.Chr. & Tardieu & (Copel.) C.Chr. (C.Chr.) Tagawa (C.Chr.) (Fée) Copel. (Blume) Copel. Christ (Hieron. ex C.Chr.) Tagawa (Hieron. ex C.Chr.) coronans heraclea speciosa

amplexifolia lehmanni phuluangensis

NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Absent Photinopteris Aglaomorpha Absent Aglaomorpha Arthromeris Arthromeris Arthromeris Absent tatsienensis Arthromeris ex Christ) Ching Belvisia annamensis Belvisia henryi Belvisia mucronata Belvisia revoluta Calymmodon asiaticus Calymmodon cucullatus (Nees & Blume) C.Presl Christiopteris tricuspis Absent Ctenopteris moultonii Xiphopteris khaoluangensis Absent Ctenopteris mollicoma Drynaria bonii 33 32 41,44 10 33 31 16 4 16 (Baker) Parris (Tagawa & K.Iwats.) (Tagawa (Alderw.) Parris (Alderw.) Tagawa & K.Iwats. Tagawa (Hook.) Roos (Nees & Blume) Parris 41 (Grev.) Parris (Grev.) (Christ) Ching (Willd.) Hovenkamp (Willd.) (Baker) Copel. (Franch. & Bureau (Wall. ex Mett.) Copel. (Wall. (Hook.) Christ (Kunze) Copel. Copel. (Mett.) Ching (C.Chr.) Tagawa Tagawa (C.Chr.) Parris

(Copel.) Tagawa (Copel.) (Fée) Copel. (L.f.) Mirbel ex Copel. Christ (Hieron. ex C.Chr.) Tagawa Tagawa (Hieron. ex C.Chr.) acuminata coronans drynarioides heraclea

amplexifolia lehmanni phuluangensis proteus

streptophyllus

33 CURRENT NAME CURRENT Acrosorus Aglaomorpha Aglaomorpha Aglaomorpha Aglaomorpha Arthromeris Arthromeris Arthromeris Arthromeris tatsienensis Arthromeris ex Christ) Ching Belvisia annamensis Belvisia henryi Belvisia mucronata Belvisia spicata Calymmodon asiaticus Calymmodon curtus Christiopteris tricuspis Chrysogrammitis musgraviana blechnoides Ctenopterella khaoluangensis Ctenopterella Parris Dasygrammitis brevivenosa Dasygrammitis mollicoma Drynaria bonii ○ ● ○ ○ ○ ● ● ○ ● ● ○ ● 86 THAI FOREST BULLETIN (BOTANY) 37 (Hook.) Blume + C.B.Clarke ex Baker Desv. (Copel.) Tagawa (Copel.) (Blume) Donk Christ (Hook.) Mett. (Hook.) Copel. / (Wall. ex Hook.) Nakai (Wall. Wall. ex Mett. Wall. Baker (J.Sm. ex Hook. & Grev.) (J.Sm. ex Hook. & Grev.) C.H.Wright (L.) J.Sm. (Desv.) T.Moore (Desv.) (Wall. ex Mett.) J.Sm. (Wall. (Sw.) Bedd. (Sw.) (Kunze ex Mett.) J.Sm. (Bedd.) Bedd. NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Drynaria fortunei Drynaria parishii Drynaria propinqua ex Bedd. Drynaria quercifolia Drynaria rigidula Drynaria sparsisora Polypodium amoenum Polypodium garrettii Polypodium manmeiense Polypodium argutum Hook. Polypodium microrhizoma Polypodium verrucosum Polypodium persicifolium Polypodium subauriculatum Polypodium beddomei Araiostegia dareiformis Gymnogrammitis dareiformis & C.Chr. Tardieu Ching ex Myrmecophila crustacea Absent Myrmecophila sinuosa ex H.Ito Lemmaphyllum accedens (Blume) C.Presl + (C.B.Clarke ex Baker) (Hook.) Ching ex Tardieu Tardieu (Hook.) Ching ex (Desv.) Bedd. (Desv.) (Hook.) J.Sm. (Wall. ex Mett.) Bedd. (Wall. (Blume) Donk Christ J.Sm. ex Hook. & Grev. Copel. Bedd. (Wall. ex Hook.) Copel. (Wall. C.H.Wright (L.) J.Sm. (Desv.) T.Moore (Desv.) (Wall. ex Mett.) J.Sm. (Wall. (Kunze ex Mett.) J.Sm. (Bedd.) Bedd. (Sw.) Bedd. (Sw.) NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Drynaria fortunei Drynaria parishii Drynaria propinqua ex Bedd. Drynaria quercifolia Drynaria rigidula Drynaria sparsisora Goniophlebium amoenum Polypodium garrettii Polypodium manmeiense Goniophlebium argutum Goniophlebium microrhizoma Bedd. Goniophlebium verrucosum Goniophlebium persicifolium Goniophlebium subauriculatum Goniophlebium molle Gymnogrammitis dareiformis & C.Chr. Lecanopteris crustacea Absent Lecanopteris sinuosa Lemmaphyllum accedens 16 7 36 (Blume) C.Presl (C.B.Clarke ex Baker) (Hook.) Ching ex Tardieu Tardieu (Hook.) Ching ex (Desv.) Bedd. (Desv.) (Christ) Rödl-Linder (Christ) Rödl-Linder 22 J.Sm. (Cav.) Wagner & Grether Wagner (Cav.) (Wall. ex Mett.) Bedd. (Wall. (Blume) Donk 28 Copel. (L.) J.Sm. (Wall. ex Hook.) Copel. (Wall. (Desv.) T.Moore T.Moore (Desv.) Blume

(Wall. ex Mett.) J.Sm. (Wall. (Bedd.) Bedd. (Sw.) Bedd. (Sw.) Nakaike CURRENT NAME CURRENT Drynaria roosii Drynaria parishii Drynaria propinqua ex Bedd. Drynaria quercifolia Drynaria rigidula Drynaria sparsisora Goniophlebium amoenum Goniophlebium lachnopus Goniophlebium manmeiense Goniophlebium mengtzeense Goniophlebium microrhizoma Bedd. Goniophlebium percussum Goniophlebium persicifolium Goniophlebium subauriculatum Gymnogrammitis dareiformis & C.Chr. Lecanopteris crustacea Lecanopteris pumila Lecanopteris sinuosa Lemmaphyllum accedens ● ● ● ● ● ● ● ● ● ● ● ○ ● Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 87 Ching (Ching) (Hook.) C.Presl Ching Blume (C.Chr.) Ching (C.Chr.) (Baker ex Takeda) Ching Takeda) (Baker ex (Rosenst.) Ching (Blume) Holttum C.Presl (Hook. & Grev.) Ching + (Hook. & Grev.) (Cav.) Kaulf. (Cav.) (Christ) Ching (Blume) C.Presl (Baker) Ching + (Christ) Ching Ching (D.Don) C.Presl (Hook.) Ching (C.Chr.) Ching (C.Chr.) NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Lemmaphyllum carnosum Lepisorus bicolor Lepisorus contortus Lepisorus heterolepis Lepisorus longifolius Lepisorus nudus Lepisorus oosphaerus Lepisorus scolopendrium Mehra & Bir Lepisorus sinensis Lepisorus subconfluens Lepisorus sublinearis Lepisorus suboligolepidus Leptochilus axillaris Leptochilus decurrens Colysis pentaphylla Colysis pothifolia Colysis hemionitidea Colysis macrophylla Absent Colysis pedunculata Colysis wui Absent Absent Colysis pothifolia Ching (Ching) (Hook.) C.Presl (Christ) C.Chr. Ching Ching Blume (C.Chr.) Ching (C.Chr.) (Baker ex Takeda) Ching Takeda) (Baker ex (Rosenst.) Ching (Cav.) Kaulf. (Cav.) (Blume) Holttum C.Presl (Blume) C.Presl (Hook. & Grev.) Ching + (Hook. & Grev.) (Christ) Ching (Baker) Ching + (Christ) Ching Ching (Hook.) Ching (C.Chr.) Ching (C.Chr.) NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Lemmaphyllum carnosum Lepisorus bicolor Lepisorus contortus Lepisorus heterolepis Lepisorus longifolius Lepisorus nudus Lepisorus oosphaerus Lepisorus scolopendrium Mehra & Bir Lepisorus sinensis Lepisorus subconfluens Lepisorus sublinearis Lepisorus suboligolepidus Leptochilus axillaris Leptochilus decurrens Colysis pentaphylla (D.Don) C.Presl Colysis hemionitidea Colysis macrophylla Absent Colysis pedunculata Colysis wui Loxogramme acroscopa Loxogramme assimilis

4 29 4 Ching (C.Presl) Noot. (Ching) (Hook.) C.Presl (Blume) Noot. (Hook. & Grev.) Fraser-Jenk. Fraser-Jenk. (Hook. & Grev.) (Christ) C.Chr. Ching Ching 5 Blume (C.Chr.) Ching (C.Chr.) (Blume) Holttum (Baker ex Takeda) Ching Takeda) (Baker ex (Rosenst.) Ching (Thunb.) Noot. (Cav.) Kaulf. (Cav.)

(Christ) Ching (Christ) Ching Fée Ching (Hook.) Ching pedunculatus

CURRENT NAME CURRENT Lemmaphyllum carnosum Lepisorus bicolor Lepisorus contortus Lepisorus heterolepis Lepisorus longifolius Lepisorus nudus Lepisorus oosphaerus Lepisorus scolopendrium Mehra & Bir Lepisorus sinensis Lepisorus subconfluens Lepisorus sublinearis Lepisorus suboligolepidus Leptochilus axillaris Leptochilus decurrens Leptochilus ellipticus Leptochilus hemionitideus Leptochilus macrophyllus Leptochilus minor Leptochilus Loxogramme acroscopa Loxogramme assimilis ● ● ● ○ ● ○ ○ 88 THAI FOREST BULLETIN (BOTANY) 37 (D.Don) (D.Don) (Bory) (Blume) Ching (Rosenst.) (Hook.) C.Chr. (D.Don) Tagawa (D.Don) (Blume) Ching Christ (L.) Copel. Ching (D.Don) C.Presl (Bedd.) Sledge (Blume) Copel. (Blume) C.Presl Blume (Blume) Ching auct. non Blume NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Loxogramme avenia Absent Loxogramme chinensis Absent Loxogramme duclouxii Loxogramme involuta Loxogramme lankokiensis C.Chr. Absent Loxogramme scolopendrina C.Presl Loxogramme subecostata heterocarpum Microsorum dilatatum Microsorum membranaceum Microsorum Ching Absent pteropus Microsorum punctatum Microsorum Absent superficiale Microsorum Absent zippelii Microsorum normalis Neocheiropteris Grammitis adspersa Grammitis setosa (D.Don) (Bory) (Blume) Ching (Hook.) C.Chr. (Rosenst.) (D.Don) Tagawa (D.Don) (Blume) Ching (Zenker) M.G.Price (L.) Copel. Ching Christ M.G.Price (Bedd.) Sledge (D.Don) C.Presl (Blume) Copel. M.G.Price (Blume) C.Presl Blume (Blume) Ching auct. non Blume sp. NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Loxogramme avenia Loxogramme centicola Loxogramme chinensis Loxogramme cuspidata Loxogramme duclouxii Loxogramme involuta Loxogramme lankokiensis C.Chr. Loxogramme porcata Loxogramme scolopendrina C.Presl Loxogramme subecostata heterocarpum Microsorum dilatatum Microsorum membranaceum Microsorum Ching Absent pteropus Microsorum punctatum Microsorum Absent superficiale Microsorum Microsorum zippelii Microsorum normalis Neocheiropteris Grammitis adspersa Grammitis setosa 34 3 4 41,33 33 29 4,8 4 1 (Gaudich.) C.V.Morton 35 (D.Don) (Blume) Ching (Rosenst.) Boonkerd & Noot. (Hook.) C.Chr. (Hook.) C.Chr. (D.Don) Tagawa Tagawa (D.Don) (Blume) Ching (Blume) Parris (Blume) Parris Copel. Christ (Zenker) M.G.Price (L.) Copel. Ching M.G.Price M.G.Price Boonkerd (D.Don) C.Presl (Blume) Copel. (Blume) C.Presl

(Blume) Ching (Blume) Copel. CURRENT NAME CURRENT Loxogramme avenia Loxogramme centicola Loxogramme chinensis Loxogramme cuspidata Loxogramme duclouxii Loxogramme involuta Loxogramme lankokiensis C.Chr. Loxogramme porcata Loxogramme scolopendrioides Loxogramme subecostata heterocarpum Microsorum insigne Microsorum membranaceum Microsorum Ching musifolium Microsorum pteropus Microsorum punctatum Microsorum siamense Microsorum superficiale Microsorum thailandicum Microsorum zippelii Microsorum normalis Neocheiropteris adspersa Oreogrammitis congener Oreogrammitis ○ ○ ○ ● ● ○ ○ ○ ● ● Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 89 (Burm.f.) Copel. (J.Koenig) Desv. (D.Don) Tagawa (D.Don) (Blume) Copel. (Copel.) Copel. (Giesenh. ex Diels) Hook. de Jonch. & Hennipman & Jonch. de (Kunze) Copel. (Blume) Mett. auct. non (Hook.) C.Chr. & auct. non (Hook.) C.Chr. (Christ) C.Chr. & (Christ) C.Chr. (Blume) Ching (G.Forst.) C.Presl (D.Don) Ching (Sw.) Ching (Sw.) (Blume) Tagawa & Tagawa (Blume) (C.Presl ex Bedd.) (Blume) Christ (Baker) Ching NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Grammitis dorsipila Tardieu Grammitis bongoensis cuspidatum Microsorum rubidum Microsorum nigrescens Microsorum scolopendria Microsorum Platycerium coronarium Platycerium holttumii Absent Platycerium wallichii alata Prosaptia khasyana Prosaptia Tardieu celebica Prosaptia K.Iwats. contigua Prosaptia obliquata Prosaptia leysii Prosaptia floccigera Pyrrosia angustata Pyrrosia angustissima Pyrrosia & K.Iwats. Tagawa costata Pyrrosia & K.Iwats. Tagawa flocculosa Pyrrosia (Burm.f.) Pic.Serm. (Blume) Pic.Serm. (D.Don) Pic.Serm. (Blume) Pic.Serm. (J.Koenig) Desv. (Copel.) Copel. de Jonch. & Hennipman (Giesenh. ex Diels) Hook. (Christ) C.Chr. & (Christ) C.Chr. Christ (Blume) Mett. auct. non (Hook.) C.Chr. & auct. non (Hook.) C.Chr. (D.Don) Ching (Blume) Tagawa & Tagawa (Blume) (Blume) Ching (Sw.) Ching (Sw.) (G.Forst.) C.Presl (C.Presl ex Bedd.) (Blume) Christ (Baker) Ching NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Grammitis dorsipila Tardieu Grammitis bongoensis Phymatosorus cuspidatus Phymatosorus longissimus Phymatosorus nigrescens Phymatosorus scolopendria Platycerium coronarium Platycerium holttumii Platycerium ridleyi Platycerium wallichii alata Prosaptia khasyana Prosaptia Tardieu celebica Prosaptia K.Iwats. contigua Prosaptia obliquata Prosaptia leysii Prosaptia floccigera Pyrrosia angustata Pyrrosia angustissima Pyrrosia & K.Iwats. Tagawa costata Pyrrosia & K.Iwats. Tagawa flocculosa Pyrrosia

25 41 33 33 15 (R.Br.) S.G.Lu (R.Br.) 41 (Burm.f.) Pic.Serm. (Blume) Parris (Blume) Pic.Serm. 7 (D.Don) Pic.Serm. (Baker) M.G.Price (Christ) Parris (J.Koenig) Desv. (J.Koenig) Desv. (Giesenh. ex Diels) de Jonch. & Hennipman Hook.

T.Moore Christ (Blume) Mett. (D.Don) Ching (Sw.) Ching (Sw.) (G.Forst.) C.Presl (Blume) Tagawa & Tagawa (Blume) (Blume) Ching (C.Presl ex Bedd.) (Blume) Christ CURRENT NAME CURRENT dorsipila Oreogrammitis reinwardtii Oreogrammitis Phymatosorus cuspidatus Phymatosorus longissimus Phymatosorus membranifolius Phymatosorus scolopendria Platycerium coronarium Platycerium holttumii Platycerium ridleyi Platycerium wallichii alata Prosaptia barathrophylla Prosaptia celebica Prosaptia K.Iwats. contigua Prosaptia obliquata Prosaptia pectinata Prosaptia albicans Pyrrosia angustata Pyrrosia angustissima Pyrrosia & K.Iwats. Tagawa costata Pyrrosia & K.Iwats. Tagawa flocculosa Pyrrosia ● ● ● ● ● ● ○ ● ● ● 90 THAI FOREST BULLETIN (BOTANY) 37 (Mett. ex (L.) C.Presl (Sw.) Ching (Sw.) (C.Chr.) Ching (C.Chr.) auct. non (Blume) (Ching) Tagawa (Ching) (Mett.) Tagawa (Mett.) (Hook.) Holttum + auct. non (Fée) C.Chr. (Giesenh.) Ching (Christ) Ching + (Sw.) Ching + (Sw.) (L.) Farw. + (L.) Farw. (Burm.f.) C.V.Morton (Mett. ex Kuhn) Ching auct. non (Blume) (Sw.) Ching (Sw.) minor Pyrrosia heteractis Pyrrosia (Giesenh.) Ching (Kunze) Ching + (Kaulf.) Farw. (Kaulf.) Farw. (Giesenh.) Ching + heteractis NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa lanceolata Pyrrosia adnascens Pyrrosia nuda Pyrrosia varia Pyrrosia Absent eberhardtii Pyrrosia heteractis Pyrrosia var. Kuhn) Ching var. longifolia Pyrrosia mannii Pyrrosia nummularifolia Pyrrosia penangiana Pyrrosia mollis Pyrrosia Drymoglossum piloselloides Absent tonkinensis Pyrrosia Absent stigmosa Pyrrosia Grammitis jagoriana Grammitis hirtella Tuyama minus Scleroglossum pusillum Scleroglossum Alderw. Crypsinus cruciformis (Mett. ex lingua heteractis (Sw.) Ching (Sw.) auct. non (Blume) (Ching) Tagawa Tagawa (Ching) (L.) M.G.Price (Mett.) Tagawa (Mett.) (Giesenh.) Ching auct. non (Fée) C.Chr. (Hook.) Holttum + (Sw.) Ching + (Sw.) (L.) Farw. + (L.) Farw. (Burm.f.) C.V.Morton auct. non (Blume) (Sw.) Ching (Sw.) (Thunb.) Farw. var. var. (Thunb.) Farw. (Giesenh.) Ching (Thunb.) Farw. var. var. (Thunb.) Farw. (Kunze) Ching (Kaulf.) Farw. (Kaulf.) Farw. (Giesenh.) Ching + NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) lanceolata Pyrrosia adnascens Pyrrosia nuda Pyrrosia varia Pyrrosia lingua Pyrrosia lingua Pyrrosia Kuhn) Hovenkamp longifolia Pyrrosia mannii Pyrrosia nummularifolia Pyrrosia penangiana Pyrrosia mollis Pyrrosia piloselloides Pyrrosia Absent tonkinensis Pyrrosia Absent stigmosa Pyrrosia Grammitis jagoriana Grammitis hirtella Tuyama minus Scleroglossum pusillum Scleroglossum Alderw. Crypsinus cruciformis 23 33 (Mett. ex porosa tonkinensis 4 41,33 41 10,15 41 heteractis lingua 15 (Mett. ex Kuhn) Parris (Copel.) Parris (Sw.) Ching (Sw.) (Kuhn) Alderw. (Kuhn) (Blume) Alderw. (Blume) (L.) M.G.Price (Ching) Fraser-Jenk. (Hook.) Holttum

(Racib.) K.H. Shing (L.) Farw. (Burm.f.) C.V.Morton (Burm.f.) C.V.Morton (Sw.) Ching (Sw.) (C.Presl) Hovenkamp var. (C.Presl) Hovenkamp var. (Thunb.) Farw. var. (Thunb.) Farw. (Giesenh.) Ching (C.Presl) Hovenkamp var. (C.Presl) Hovenkamp var. (Thunb.) Farw. var. var. (Thunb.) Farw. CURRENT NAME CURRENT lanceolata Pyrrosia lingua Pyrrosia lingua Pyrrosia Kuhn) Hovenkamp longifolia Pyrrosia mannii Pyrrosia nummularifolia Pyrrosia penangiana Pyrrosia piloselloides Pyrrosia porosa Pyrrosia porosa Pyrrosia (Giesenh.) Ching rasamalae Pyrrosia stigmosa Pyrrosia Radiogrammitis jagoriana Radiogrammitis multifolia pusillum Scleroglossum sulcatum Scleroglossum Selliguea cruciformis ● ○ ● ● ● ○ ● ○ ● ● ● ● ● Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 91 (Hook.) Copel. (C.Chr.) Holttum (C.Chr.) (Hook.) Copel. (Blume) Holttum Tagawa & K.Iwats. Tagawa Sw. (Blume) Kunze (Blume) J.Sm. (Bory) C.Chr. Blume (C.Presl) Holttum (Wall. ex Kunze) (Wall. (Hook.) Copel. Tagawa & K.Iwats. Tagawa (Houtt.) Copel. (Cav.) Copel. (Cav.) (L.) Beauv. NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Crypsinus ebenipes Crypsinus enervis Crypsinus griffithianus Selliguea heterocarpa Crypsinus hirsutus Crypsinus laciniatus Absent Crypsinus oxylobus Sledge Crypsinus rhynchophyllus Crypsinus stenophyllus Crypsinus trilobus Ctenopteris tenuisecta Ctenopteris subfalcata Xiphopteris hieronymusii Psilotum complanatum Psilotum nudum Absent Cheilanthes belangeri Cheilanthes delicatula (Hook.) Copel. (C.Chr.) Holttum (C.Chr.) Sw. (Blume) Holttum Tagawa & K.Iwats. Tagawa (Hook.) Copel. (Blume) Kunze Blume (Blume) J.Sm. (Bory) C.Chr. (S.G.Gmel) Kunze (C.Presl) Holttum (Wall. ex Kunze) (Wall. (Hook.) Copel. Tagawa & K.Iwats. Tagawa (Houtt.) Copel. (Cav.) Copel. (Cav.) (L.) Beauv. NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Crypsinus ebenipes Crypsinus enervis Crypsinus griffithianus Selliguea heterocarpa Crypsinus hirsutus Crypsinus laciniatus Absent Crypsinus oxylobus Sledge Crypsinus rhynchophyllus Crypsinus stenophyllus Crypsinus trilobus Ctenopteris tenuisecta Ctenopteris subfalcata Xiphopteris hieronymusii Psilotum complanatum Psilotum nudum Cheilanthes argentea Cheilanthes belangeri Cheilanthes delicatula 33 33 4 16 (C.Chr.) Parris (C.Chr.) (Blume) Parris (Hook.) Fraser-Jenk. (Blume) Parris Sw. Sw. Tagawa & K.Iwats. Tagawa (Bory) C.Chr. (Bory) C.Chr. Blume (Blume) Parris (S.G.Gmel) Kunze (Hook.) Fraser-Jenk.

(C.Presl) Parris (Hook.) S.Linds. (Baker) Hovenkamp (Wall. ex Kunze) Fraser-Jenk. (Wall. (Cav.) Ching (Cav.) (Tagawa & K.Iwats.) S.Linds. (Tagawa (Houtt.) M.G.Price (L.) Beauv. (L.) Beauv. CURRENT NAME CURRENT Selliguea ebenipes Selliguea enervis Selliguea griffithiana Selliguea heterocarpa Selliguea hirsuta Selliguea laciniata Selliguea lateritia Selliguea oxyloba Selliguea rhynchophylla Selliguea stenophylla Selliguea triloba Themelium tenuisectum subfalcatum Tomophyllum hieronymusii Xiphopterella PSILOTACEAE Psilotum complanatum Psilotum nudum PTERIDACEAE subfam. Cheilanthoideae Cheilanthes argentea Cheilanthes belangeri Cheilanthes delicatula ● ● ● ● ● ○ ● ● ● ● ● ● ● ○ 92 THAI FOREST BULLETIN (BOTANY) 37

(Ching & Hayata (S.K.Wu) (S.K.Wu) Franch. & Sav. Cheilanthes subrufa (D.Don) Diels (Burm.f.) Sw. auct. non (Forssk.) Kaulf., auct. non (Forssk.) Kaulf., auct. non (Forssk.) auct. non (Forssk.) Kaulf., auct. non (Forssk.) Kaulf., (Wall. ex Hook.) Underw. (Wall. Hook. Tardieu & C.Chr. Tardieu (Wall. ex Hook.) J.Sm. (Wall. (Burm.f.) T.Moore (Burm.f.) Cheilanthes pseudoargentea D.Don + (Blume) Hieron. Cheilanthes formosana Cheilanthes krameri Cheilanthes pseudofarinosa Cheilanthes siamensis serrulata NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Cheilanthes farinosa pro parte / Cheilanthes fragilis Cheilanthes farinosa pro parte / Cheilanthes farinosa Kaulf., pro parte / K.Iwats. (S.K.Wu) Cheilanthes farinosa pro parte / K.Iwats. S.K.Wu) Cheilanthes rufa auct. non Baker Cheilanthes farinosa pro parte / K.Iwats. Cheilanthes tenuifolia Absent Doryopteris ludens Gymnopteris vestita Notholaena velutina Hemionitis arifolia Absent Coniogramme fraxinea var. (S.K.Wu) K.Iwats. (S.K.Wu) (Ching & S.K.Wu) K.Iwats. (Ching & S.K.Wu) (D.Don) Diels Hayata (S.K.Wu) K.Iwats. (S.K.Wu) (Burm.f.) Sw. Tardieu & C.Chr. Tardieu Franch. & Sav. (Wall. ex Hook.) Underw. (Wall. Hook. (Wall. ex Hook.) J.Sm. (Wall. (Burm.f.) T.Moore (Burm.f.) Alderw. D.Don (Blume) Hieron. serrulata NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Cheilanthes formosana Cheilanthes fragilis Cheilanthes krameri Cheilanthes pseudoargentea Cheilanthes pseudofarinosa Cheilanthes rufa Cheilanthes siamensis Cheilanthes tenuifolia Absent Doryopteris ludens Gymnopteris vestita Notholaena velutina Hemionitis arifolia Pellaea timorensis Coniogramme fraxinea var.

10 (S.K.Wu) K.Iwats. (S.K.Wu) (Ching & S.K.Wu) K.Iwats. (Ching & S.K.Wu) 4 (Roxb. ex Hook. & Grev.) (Roxb. ex Hook. & Grev.) Hayata (D.Don) Diels (S.K.Wu) K.Iwats. (S.K.Wu) (Burm.f.) Sw. (Burm.f.) Sw. R .M.Tryon

Franch. & Sav. Franch. & Sav. Hook. (Wall. ex Hook.) J.Sm. (Wall. Alderw. (Wall. ex Hook.) J.Sm. (Wall. D.Don (Blume) Hieron. 11 errulata CURRENT NAME CURRENT Cheilanthes formosana Cheilanthes fragilis Cheilanthes krameri Cheilanthes pseudoargentea Cheilanthes pseudofarinosa Cheilanthes rufa Cheilanthes siamensis Cheilanthes tenuifolia Doryopteris alleniae Doryopteris ludens Hemionitis vestita & C.Chr. Notholaena velutina Tardieu Parahemionitis cordata Fraser-Jenk. Pellaea timorensis PTERIDACEAE subfam. Cryptogrammoideae Coniogramme fraxinea s var. ● ● ● ● ● ● ○ ● ● ○ Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 93 (L.) Link (L.) Brongn. Tardieu Willd. Fée (Desv.) C.Chr. (Desv.) C.Hope L. Fée Wall. ex J. A gardh Wall. Wall. ex J. A gardh Wall. Hook. Christ Burm.f. J. A gardh J.Sm. ex Hieron. Poir. L. L. Tagawa NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Coniogramme petelotii Coniogramme procera aureum Acrostichum speciosum Acrostichum Ceratopteris thalictroides Onychium contiguum Onychium siliculosum calomelanos Pityrogramma Pteris aspericaulis Pteris asperula Pteris bella Pteris biaurita Pteris blumeana Pteris cretica Pteris dalhousiae Pteris decrescens Pteris ensiformis Pteris grevilleana Pteris heteromorpha Pteris linearis (L.) Link (L.) Brongn. Willd. Tardieu Fée (Desv.) C.Chr. (Desv.) C.Hope L. Fée Wall. ex J. A gardh Wall. Wall. ex J. A gardh Wall. Hook. Christ J. A gardh Burm.f. J.Sm. ex Hieron. Poir. L. L. Tagawa NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Coniogramme petelotii Coniogramme procera aureum Acrostichum speciosum Acrostichum Ceratopteris thalictroides Onychium contiguum Onychium siliculosum calomelanos Pityrogramma Pteris aspericaulis Pteris asperula Pteris bella Pteris biaurita Pteris blumeana Pteris cretica Pteris dalhousiae Pteris decrescens Pteris ensiformis Pteris grevilleana Pteris heteromorpha Pteris linearis (L.) Link (L.) Brongn. Tardieu Tardieu Willd. Willd. Fée (Desv.) C.Chr. C.Chr. (Desv.) C.Hope L. Fée

Wall. ex J. A gardh Wall. Wall. ex J. A gardh Wall. Christ Hook. Burm.f. J. A gardh J.Sm. ex Hieron. Poir. Poir. L. L. Tagawa Tagawa CURRENT NAME CURRENT Coniogramme petelotii Coniogramme procera PTERIDACEAE subfam. Parkerioideae aureum Acrostichum speciosum Acrostichum Ceratopteris thalictroides PTERIDACEAE subfam. Pteridoideae Onychium contiguum Onychium siliculosum calomelanos Pityrogramma Pteris aspericaulis Pteris asperula Pteris bella Pteris biaurita Pteris blumeana Pteris cretica Pteris dalhousiae Pteris decrescens Pteris ensiformis Pteris grevilleana Pteris heteromorpha Pteris linearis 94 THAI FOREST BULLETIN (BOTANY) 37 L. (C.Presl) J.Sm. Hook. Willd. Tagawa & K.Iwats. Tagawa L. (Willd.) Sw. (Willd.) Wall. ex J. A gardh Wall. L. J. A gardh Wall. ex J. A gardh Wall. Wall. ex Hook. & Grev. Wall. H.Ito Wall. ex J. A gardh Wall. Poir. Sw. Christ D.Don Kunze L. Masam. stenophylla

NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Pteris longipes Pteris longipinnula Pteris mertensioides Pteris multifida Pteris nepalensis Pteris phuluangensis Pteris plumbea Pteris scabripes Pteris semipinnata Pteris Pteris subquinata Pteris tokioi Pteris tripartita Pteris venusta Pteris vittata Pteris wallichiana Syngramma alismifolia blechnoides Taenitis Absent Adiantum capillus-veneris Adiantum caudatum Adiantum edgeworthii L. (C.Presl) J.Sm. Hook. Tagawa & K.Iwats. Tagawa L. Willd. (Willd.) Sw. (Willd.) Wall. ex J. A gardh Wall. L. J. A gardh Wall. ex Hook. & Grev. Wall. Wall. ex J. A gardh Wall. H.Ito Wall. ex J. A gardh Wall. Poir. Sw. Christ D.Don Kunze L. Masam. NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Pteris longipes Pteris longipinnula Pteris mertensioides Pteris multifida Pteris nepalensis Pteris phuluangensis Pteris plumbea Pteris scabripes Pteris semipinnata Pteris stenophylla Pteris subquinata Pteris tokioi Pteris tripartita Pteris venusta Pteris vittata Pteris wallichiana Syngramma alismifolia blechnoides Taenitis Absent Adiantum capillus-veneris Adiantum caudatum Adiantum edgeworthii 10 L. (C.Presl) J.Sm. Hook. Tagawa & K.Iwats. Tagawa

Willd. Willd. (Willd.) Sw. Sw. (Willd.) L. Wall. ex J. A gardh Wall. Hook. & Grev. L. J. A gardh Wall. ex Hook. & Grev. ex Hook. & Grev. Wall. Wall. ex J. A gardh Wall. H.Ito Wall. ex J. A gardh Wall. Sw. Sw. Poir. Poir. Christ D.Don Kunze L. Masam. CURRENT NAME CURRENT Pteris longipes Pteris longipinnula Pteris mertensioides Pteris multifida Pteris nepalensis Pteris phuluangensis Pteris plumbea Pteris scabripes Pteris semipinnata Pteris stenophylla Pteris subquinata Pteris tokioi Pteris tripartita Pteris venusta Pteris vittata Pteris wallichiana Syngramma alismifolia blechnoides Taenitis interrupta Taenitis Vittarioideae PTERIDACEAE subfam. Adiantum capillus-veneris Adiantum caudatum Adiantum edgeworthii ○ Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 95 Vittaria Vittaria Baker Baker, pro parte Baker, Blume L. Baker, pro parte Baker, Baker Wall. ex Hook. Wall. (Bory) Thwaites & (Bory) L. Blume Fée + Blume Copel. Kuhn Tagawa & K.Iwats. Tagawa Mett. ex Kuhn Sw. C.Chr. & Tardieu & C.Chr. Sw. Fée auct. non Ching NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Adiantum erylliae Adiantum flabellulatum Adiantum stenochlamys Absent Absent Adiantum philippense Adiantum siamense Adiantum soboliferum Adiantum stenochlamys Absent Adiantum zollingeri callifolium Antrophyum obovatum Antrophyum parvulum Antrophyum stenophyllum Antrophyum amboinensis Vittaria forrestiana angustifolia Vittaria elongata Vittaria ensiformis Vittaria flexuosa Vittaria scolopendrina Vittaria Hook. sikkimensis Vittaria taeniophylla Vittaria Vittaria Vittaria Baker Blume Baker, pro parte Baker, pro parte Baker, L. Baker (Bory) Thwaites & (Bory) Blume Wall. ex Hook. Wall. L. Fée + Copel. Mett. ex Kuhn Blume Tagawa & K.Iwats. Tagawa Kuhn Sw. C.Chr. & Tardieu & C.Chr. Sw. Fée auct. non Ching NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Adiantum erylliae Adiantum flabellulatum Adiantum stenochlamys Absent Absent Adiantum philippense Adiantum siamense Adiantum soboliferum Adiantum stenochlamys Absent Adiantum zollingeri callifolium Antrophyum obovatum Antrophyum parvulum Antrophyum stenophyllum Antrophyum amboinensis Vittaria forrestiana angustifolia Vittaria elongata Vittaria ensiformis Vittaria flexuosa Vittaria scolopendrina Vittaria Hook. sikkimensis Vittaria taeniophylla Vittaria 21 40 37 39 46 C.Presl 10 (Blume) E.H.Crane (Fée) X.C.Zhang (Copel.) E.H.Crane Baker 43 L. (Kuhn) E.H.Crane Baker Blume S.Linds. & D.J.Middleton Baker C.Chr. (Sw.) E.H.Crane (Sw.) Wall. ex Hook. Wall. Blume Suksathan L. (Sw.) E.H.Crane (Sw.) (Fée) E.H.Crane

Tagawa & K.Iwats. Tagawa Lam. Mett. ex Kuhn C.Chr. & Tardieu Tardieu & C.Chr. CURRENT NAME CURRENT Adiantum erylliae Adiantum flabellulatum Adiantum fragiliforme Adiantum latifolium Adiantum phanomensis Adiantum philippense Adiantum siamense Adiantum soboliferum Adiantum stenochlamys Adiantum thongthamii Adiantum zollingeri callifolium Antrophyum obovatum Antrophyum parvulum Antrophyum vittarioides Antrophyum Haplopteris amboinensis Haplopteris angustifolia Haplopteris elongata Haplopteris ensiformis Haplopteris flexuosa Haplopteris scolopendrina Haplopteris sikkimensis Haplopteris taeniophylla ● ○ ○ ○ ● ● ● ● ● ● ● ● ● 96 THAI FOREST BULLETIN (BOTANY) 37 (Hayata) Tagawa & Tagawa (Hayata) (C.Chr.) Holttum (C.Chr.) (Racib.) Holttum (Racib.) Holttum Tagawa & K.Iwats. Tagawa (Mett.) Holttum (Copel.) Holttum Fée (C.Presl) Holttum (Fée) Mett. (Desv.) Alston (Desv.) (L.) Sm. Tagawa & K.Iwats. Tagawa Roxb. ex Bory Willd. (L.) Sw. (L.) All. (L.) R.Br. NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa winitii Antrophyum paradoxa Vaginularia trichoidea Vaginularia Azolla caroliniana Azolla pinnata Salvinia cucullata Salvinia natans Schizaea dichotoma Schizaea digitata palisotii Arthropteris alderwereltii Heterogonium gurupahense Heterogonium hennipmanii Heterogonium pinnatum Heterogonium sagenioides Heterogonium Pleocnemia hemiteliiformis Pleocnemia irregularis Pleocnemia submembranacea K.Iwats. (Hayata) Tagawa & Tagawa (Hayata) (C.Chr.) Holttum (C.Chr.) Tagawa & K.Iwats. Tagawa Holttum (Racib.) Holttum (Mett.) Holttum (Copel.) Holttum (C.Presl) Holttum Fée (Fée) Mett. (Desv.) Alston (Desv.) (L.) Sm. Tagawa & K.Iwats. Tagawa Willd. Roxb. ex Bory (L.) Sw. (L.) All. (L.) R.Br. NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) winitii Antrophyum paradoxa Vaginularia trichoidea Vaginularia Azolla caroliniana Azolla pinnata Salvinia cucullata Salvinia natans Schizaea dichotoma Schizaea digitata palisotii Arthropteris alderwereltii Heterogonium gurupahense Heterogonium hennipmanii Heterogonium pinnatum Heterogonium sagenioides Heterogonium Pleocnemia hemiteliiformis Pleocnemia irregularis Pleocnemia submembranacea K.Iwats. Hayata) Tagawa & Tagawa Hayata) (C.Chr.) Holttum (C.Chr.) Holttum (Racib.) Holttum Tagawa & K.Iwats. Tagawa (Mett.) Holttum (Copel.) Holttum (C.Presl) Holttum Fée (Fée) Mett. (Desv.) Alston (Desv.) (L.) Sm.

(Tagawa & K.Iwats.) S.Linds. (Tagawa Roxb. ex Bory Willd. Willd. (L.) Sw. (L.) Sw. (L.) All. (L.) R.Br. R.Br. CURRENT NAME CURRENT Haplopteris winitii paradoxa Vaginularia trichoidea Vaginularia SALVINIACEAE Azolla caroliniana Azolla pinnata Salvinia cucullata Salvinia natans SCHIZAEACEAE Schizaea dichotoma Schizaea digitata TECTARIACEAE palisotii Arthropteris alderwereltii Heterogonium gurupahense Heterogonium hennipmanii Heterogonium pinnatum Heterogonium sagenioides Heterogonium Pleocnemia hemiteliiformis Pleocnemia irregularis Pleocnemia submembranacea ( K.Iwats. ● Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 97 Holttum (Christ) C.Chr. & (Christ) C.Chr. Holttum (Ridl.) C.Chr. (C.Presl) Holttum (Willd.) C.Chr. & Ching & C.Chr. (Willd.) (Alderw.) C.Chr. (Alderw.) (J.Sm.) C.Chr., pro parte pro C.Chr., (J.Sm.) (J.Sm.) C.Chr., pro parte pro C.Chr., (J.Sm.) (C.Presl) Copel. Ching (Wall. ex Hook.) C.Chr. (Wall. (Willd.) C.Chr. (Willd.) (Baker) C.Chr. Cav. (Hook.) Copel. (Wall. ex Bedd.) C.Chr. (Wall. Tardieu & C.Chr. Tardieu (Kunze) Copel. Tagawa NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Pteridrys australis Pteridrys cnemidaria Ching Pteridrys syrmatica amplifolia Tectaria angulata Tectaria barberi Tectaria Absent coadunata Tectaria coadunata Tectaria crenata Tectaria decurrens Tectaria devexa Tectaria fauriei Tectaria rumicifolia Tectaria fuscipes Tectaria griffithii Tectaria gymnosora Tectaria herpetocaulos Tectaria variolosa Tectaria Absent laotica Tectaria Ctenitis manilensis (C.Presl) Holttum Holttum (Christ) C.Chr. & (Christ) C.Chr. (Ridl.) C.Chr. Holttum (Willd.) C.Chr. & Ching C.Chr. (Willd.) (C.Presl) Holttum (Alderw.) C.Chr. (Alderw.) (C.Presl) Copel. Ching (Zoll. & Moritzi) C.V.Morton (Ridl.) Holttum (Willd.) C.Chr. (Willd.) (Fée) Holttum (Baker) C.Chr. Tectaria manilensis Tectaria Cav. (Hook.) Copel. (Wall. ex Bedd.) C.Chr. (Wall. Copel. Tardieu & C.Chr. Tardieu (Kunze) Copel. Tagawa (Luerss.) C.Chr. + manilensis chupengensis NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Pteridrys australis Pteridrys cnemidaria Ching Pteridrys syrmatica amplifolia Tectaria angulata Tectaria barberi Tectaria brachiata Tectaria christii Tectaria absent crenata Tectaria decurrens Tectaria devexa Tectaria fauriei Tectaria rumicifolia Tectaria fuscipes Tectaria griffithii Tectaria gymnosora Tectaria herpetocaulos Tectaria impressa Tectaria keckii Tectaria laotica Tectaria manilensis Tectaria var. var. 4 4 13 13 (Christ) C.Chr. & (Christ) C.Chr. os Holttum

Holttum (Willd.) C.Chr. & Ching C.Chr. (Willd.) (C.Presl) Holttum (J.Sm.) C.Chr. (Alderw.) C.Chr. C.Chr. (Alderw.) (C.Presl) Copel. Ching (Zoll. & Moritzi) C.V.Morton (Fée) Holttum (Willd.) C.Chr. C.Chr. (Willd.) (Baker) C.Chr. (Baker) C.Chr. (Wall. ex Bedd.) C.Chr. ex Bedd.) C.Chr. (Wall. Tardieu & C.Chr. & C.Chr. Tardieu Cav. Cav. (Hook.) Copel. Copel. (Kunze) Copel. Tagawa Tagawa (Luerss.) C.Chr. (Kunze) Holttum CURRENT NAME CURRENT Pteridrys australis Pteridrys cnemidaria Ching Pteridrys syrmatica amplifolia Tectaria angulata Tectaria barberi Tectaria brachiata Tectaria christii Tectaria coadunata Tectaria crenata Tectaria decurrens Tectaria devexa Tectaria fauriei Tectaria fissa Tectaria fuscipes Tectaria griffithii Tectaria gymnosora Tectaria herpetocaul Tectaria impressa Tectaria keckii Tectaria laotica Tectaria manilensis Tectaria ○ ● ● ● ○ ● 98 THAI FOREST BULLETIN (BOTANY) 37 (Wall. ex Hook. & (Wall. (Blume) Copel. (Houlston & T.Moore) T.Moore) (Houlston & (Rosenst.) C.Chr. (Wall. ex Hook.) (Wall. (Houtt.) Copel. (Hook.) Ching (C.Hope) Ching (C.Presl) K.Iwats. (Wall. ex Benth.) Ching (Wall. (Alderw.) C.Chr. (Alderw.) (Baker) C.Chr. (Baker) Ching (D.Don) C.V.Morton C.Chr. (Blume) Copel. NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa melanocaulis Tectaria phaeocaulis Tectaria Absent Absent polymorpha Tectaria Copel. rockii Tectaria maingayi Tectaria Absent Absent simonsii Tectaria singaporeana Tectaria Ching Grev.) tenerifrons Tectaria ternifolia Tectaria vasta Tectaria zeilanica Quercifilix Thelypteris arida Thelypteris articulata & K.Iwats. Tagawa Thelypteris aspera Thelypteris repens Thelypteris ciliata (Houlston & T.Moore) T.Moore) (Houlston & (Wall. ex Hook. & (Wall. (C.Presl) Holttum (Blume) Copel. (Wall. ex Benth.) Holttum (Wall. ( R oxb.) C.V.Morton (Wall. ex Hook.) (Wall. (Rosenst.) C.Chr. (Houtt.) Copel. (Hook.) Ching (C.Hope) Ching (Alderw.) C.Chr. (Alderw.) (Baker) Ching (D.Don) Holttum (Willd.) Copel. (Willd.) (Fée) R .C.Moran C.Chr. (Blume) Copel. NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) melanocaulis Tectaria phaeocaulis Tectaria Absent pilosa Tectaria polymorpha Tectaria Copel. rockii Tectaria semipinnata Tectaria Absent siifolia Tectaria simonsii Tectaria singaporeana Tectaria Ching Grev.) tenerifrons Tectaria ternifolia Tectaria vasta Tectaria zeilanica Quercifilix Christella arida articulatum Pronephrium Holttum asperum Pronephrium Thelypteris repens ciliata Trigonospora 14 4 4 24 44 Rusea (Houlston & T.Moore) T.Moore) (Houlston & (Wall. ex Hook. & (Wall. (Blume) Copel. S.Linds. ( R oxb.) C.V.Morton (Wall. ex Hook.) (Wall. (Rosenst.) C.Chr. (Rosenst.) C.Chr.

(C.Presl) B.K.Nayar & Kaur (Wall. ex Benth.) Panigrahi (Wall. (Hook.) Ching (D.Don) Ching (Houtt.) Sledge (Alderw.) C.Chr. C.Chr. (Alderw.) (Baker) S.Linds. (Baker) Ching (Willd.) Copel. (Willd.) (Fée) R .C.Moran C.Chr. C.Chr. (Blume) Copel. CURRENT NAME CURRENT melanocaulis Tectaria phaeocaulis Tectaria phanomensis Tectaria pilosa Tectaria polymorpha Tectaria Copel. rockii Tectaria semipinnata Tectaria shahidaniana Tectaria siifolia Tectaria simonsii Tectaria singaporeana Tectaria Ching Grev.) tenerifrons Tectaria ternifolia Tectaria vasta Tectaria zeilanica Tectaria THELYPTERIDACEAE Cyclosorus aridus Cyclosorus articulatus Panigrahi Cyclosorus asperus Cyclosorus canus Cyclosorus ciliatus ○ ○ ● ○ ○ ● ● ● ● ● ● Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 99 (Rosenst.) (Blume) (Rosenst.) (Blume) (Blume) (Mett.) K.Iwats. (Blume) Tagawa & Tagawa (Blume) (Willd.) K.Iwats. (Willd.) (Blume) Ching (Baker) K.Iwats. (Bedd.) Tagawa & Tagawa (Bedd.) (Hook.) Ching (Hook.) K.Iwats. (Blume) Ching (Forssk.) E .P.St.John (C.B.Clarke & Baker) (Baker) Panigrahi (Blume) Tagawa & Tagawa (Blume) NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Thelypteris crassifolia Thelypteris crinipes Thelypteris cylindrothrix K.Iwats. Thelypteris dentata Thelypteris evoluta & K.Iwats Tagawa Thelypteris exsculpta Thelypteris falciloba Thelypteris ferox K.Iwats. Thelypteris glandulosa K.Iwats., nom. illeg. Thelypteris heterocarpa C.V.Morton Thelypteris immersa Thelypteris interrupta Thelypteris lakhimpurensis K.Iwats. Thelypteris larutensis K.Iwats. Thelypteris lebeufii Thelypteris megaphylla Thelypteris menisciicarpa K.Iwats. (Blume) (Blume) (Rosenst.) Holttum (Bedd.) C.Chr. Holttum, (Blume) Holttum (Blume) Holttum (Rosenst.) Ching (Baker) Holttum (Blume) Holttum (Willd.) H.Ito (Willd.) (Hook.) Ching (Forssk.) Brownsey & Jermy (Hook.) Holttum (C.B.Clarke & Baker) (Baker) Holttum (Blume) Holttum NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Mesophlebion crassifolium Christella crinipes Cyclosorus cylindrothrix Christella dentata Christella evoluta Holttum exsculptum Pronephrium Thelypteris falciloba Chingia ferox glandulosum Pronephrium heterocarpus Sphaerostephanos Holttum immersum Amphineuron Cyclosorus interruptus lakhimpurense Pronephrium larutensis Sphaerostephanos Christella lebeufii penniger Sphaerostephanos nom. nud. menisciicarpon Pronephrium Holttum 11 ( R osenst.) B.K.Nayar & Kaur (Blume) (Blume) (Blume) Ching (Mett.) Ching (Willd.) H.Ito (Willd.) (Blume) S.Linds. (Baker) S.Linds. (Bedd.) Ching (Hook.) Panigrahi (Blume) S.Linds.

(Forssk.) Ching (C.B.Clarke & Baker) (Hook.) Ching (Bedd.) Ching (Baker) W.M.Chu W.M.Chu (Baker) (Blume) Ching clarkei

CURRENT NAME CURRENT Cyclosorus crassifolius Cyclosorus crinipes Cyclosorus Cyclosorus dentatus Cyclosorus evolutus Ching Cyclosorus exsculptus Cyclosorus falcilobus Cyclosorus ferox Cyclosorus glandulosus Cyclosorus heterocarpus Ching Cyclosorus immersus Cyclosorus interruptus Cyclosorus lakhimpurense Cyclosorus larutensis Cyclosorus lebeufii Cyclosorus megaphyllus Cyclosorus menisciicarpus Holttum ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 100 THAI FOREST BULLETIN (BOTANY) 37 (Blume) K.Iwats. + (C.Chr.) Ching (C.Chr.) (Hook.) C.F.Reed (Hook.) C.F.Reed (Kuhn) K.Iwats. (Retz.) Sw. (Retz.) Sw. (Alderw.) Tagawa & Tagawa (Alderw.) (J.Sm. ex Hook.) Tagawa & K.Iwats. Tagawa (L.) Tardieu (L.) (Blume) K.Iwats. (Sw.) K.Iwats. (Sw.) (Baker) K.Iwats. (Hook.) K.Iwats. + (Kaulf.) Fosberg (Sw.) K.Iwats. (Sw.) (Poir.) K.Iwats. (Poir.) (Fée) C.V.Morton ( R oxb.) C.V.Morton (C.Hope) K.Iwats. (Bedd.) K.Iwats. triphylla parishii NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Thelypteris molliuscula Thelypteris nudata Thelypteris opulenta Thelypteris papilio Thelypteris parasitica Thelypteris penangiana Thelypteris polycarpa Meniscium proliferum Thelypteris repanda Absent Absent Thelypteris siamensis Thelypteris subelata Thelypteris subpubescens Thelypteris latipinna Thelypteris sumatrana K.Iwats. Thelypteris terminans & K.Iwats. Tagawa Absent Thelypteris triphylla var. Thelypteris triphylla var. Thelypteris truncata Thelypteris tuberculifera (Blume) Copel. (Hook.) Holttum (Alderw.) Holttum (Alderw.) (Wall. ex Hook.) Holttum (Wall. (Blume) Holttum + (C.Presl) Holttum (Poir.) Holttum (Poir.) (C.Chr.) Ching (C.Chr.) (J.Sm. ex Hook.) (Kaulf.) Holttum (Fée) Holttum (Sw.) Holttum (Sw.) Holttum (Sw.) (Roxb.) Holttum (Retz.) Sw. (Retz.) Sw. (L.) H.Lév. (Tagawa & K.Iwats.) Holttum (Tagawa (Benth.) H.Lév. (Baker) Holttum (C.Hope) Holttum (Bedd.) Nakaike parishii . triphyllum NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Christella appendiculata nudatum Pronephrium opulentum Amphineuron Christella papilio Christella parasitica penangianum Pronephrium polycarpa Sphaerostephanos Meniscium proliferum repandum Pronephrium rubicundum Pronephrium salicifolium Pronephrium Christella siamensis Christella subelata Christella subpubescens Christella latipinna terminans Amphineuron Holttum Absent triphyllum Pronephrium var triphyllum Pronephrium var. Pneumatopteris truncata Thelypteris tuberculifera 4 4 12 24 (Blume) Ching (Hook.) Copel. (Kuhn) Ching S.Linds. (C.Chr.) Panigrahi (C.Chr.) (Alderw.) S.Linds. (Alderw.) (L.) Farw. (L.) Farw. (Blume) Holttum (Wall. ex Hook.) Copel. (Wall. (J.Sm. ex Hook.) (Baker) Ching (Tagawa & K.Iwats.) Panigrahi (Tagawa (Kaulf.) Nakaike (Sw.) Tardieu ex Tardieu & C.Chr. & C.Chr. Tardieu ex Tardieu (Sw.) & C.Chr. Tardieu ex Tardieu (Sw.) (Poir.) Farw. Farw. (Poir.) (Fée) B.K.Nayar & Kaur

( R oxb.) B.K.Nayar & Kaur (Retz.) Tardieu ex Tardieu & C.Chr. & C.Chr. Tardieu ex Tardieu (Retz.) (C.Hope) Ching (Bedd.) S.Linds. parishii triphyllus CURRENT NAME CURRENT Cyclosorus molliusculus Cyclosorus nudatus Cyclosorus opulentus Cyclosorus papilio Cyclosorus parasiticus Cyclosorus penangianus Cyclosorus polycarpus Cyclosorus prolifer Cyclosorus repandus Cyclosorus rubicundus Cyclosorus salicifolius Cyclosorus siamensis Cyclosorus subelatus Cyclosorus subpubescens Cyclosorus terminans Panigrahi Cyclosorus thailandicus Cyclosorus triphyllus var. Cyclosorus triphyllus var. Cyclosorus truncatus Cyclosorus tuberculifer ● ● ● ● ● ● ● ● ● ○ ○ ● ● ● ● ○ ● ● ● ● Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 101 C.Presl (Kunze) M.Kato (Baker) Ching (Bedd.) Pic.Serm. (Gaudich.) Alston (Gaudich.) Christ (Baker) C.Chr. (C.Hope) C.Chr. (Thunb.) C.V.Morton (Clarke) Ching (C.Chr.) K.Iwats. + (C.Chr.) (Baker) Ching C.Chr. (Blume) Ching (Kunze) Ching (D.Don) Tagawa (D.Don) (Wall. ex Bedd.) Ching (Wall. (Christ) Tagawa & Tagawa (Christ) auct. non (Thunb.) (L.) C.V.Morton Deparia petersenii NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Thelypteris xylodes Thelypteris unita Thelypteris hirtisora Thelypteris valida K.Iwats. Thelypteris ornata Thelypteris torresiana Absent Thelypteris confluens Thelypteris flaccida Thelypteris hirsutipes Absent Thelypteris singalanensis Thelypteris viscosa Athyrium anisopterum Anisocampium cumingianum Kuniwatsukia cuspidata Athyrium dissitifolium Athyrium mackinnonii Athyrium setiferum Cornopteris opaca Deparia japonica M.Kato / Holttum (Baker) Ching C.Presl (Gaudich.) Ching (L.) Holttum (C.B.Clarke) Holttum (Wall. ex Bedd.) Ching (Wall. (Blume) Ching (Bedd.) Pichi-Serm. (Baker) Holttum Christ (Baker) C.Chr. (C.Hope) C.Chr. (C.Chr.) Ching (C.Chr.) (Thunb.) C.V.Morton auct. non (Thunb.) Copel. (Kunze) Ching C.Chr. (D.Don) Tagawa (D.Don) NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Thelypteris xylodes unitus Sphaerostephanos Cyclosorus hirtisorus ornata Macrothelypteris torresiana Macrothelypteris Pseudophegopteris sumatrana Thelypteris confluens Metathelypteris flaccida Coryphopteris hirsutipes Absent Metathelypteris singalanensis Coryphopteris viscosa Athyrium anisopterum Anisocampium cumingianum Kuniwatsukia cuspidata Athyrium dissitifolium Athyrium mackinnonii Athyrium setiferum Cornopteris opaca Athyrium japonicum 26 14 Holttum (Gaudich.) Ching (Baker) Ching (Wall. ex Bedd.) Ching (Wall. (C.Presl) Milde (C.Hope) C.Chr. Christ (Baker) C.Chr. (Baker) C.Chr. (Clarke) Ching (Bedd.) M.Kato (Thunb.) C.V.Morton (Thunb.) C.V.Morton

(Blume) Ching (Christ) Ching (Kunze) M.Kato (Baker) Ching C.Chr. (Kunze) Panigrahi (D.Don) Tagawa Tagawa (D.Don) (L.) Ching (Franch. & Sav.) Ching (Franch. & Sav.) CURRENT NAME CURRENT Cyclosorus tylodes Cyclosorus unitus Cyclosorus validus ornata Macrothelypteris torresiana Macrothelypteris Pseudophegopteris sumatrana Thelypteris confluens Thelypteris flaccida Thelypteris hirsutipes Thelypteris laxa Thelypteris singalanensis Thelypteris viscosa WOODSIACEAE Athyrium anisopterum Athyrium cumingianum Athyrium cuspidatum Athyrium dissitifolium Athyrium mackinnonii Athyrium setiferum Cornopteris opaca Deparia petersenii ● ● ● ● ● ● ● ○ ● ● ● ● 102 THAI FOREST BULLETIN (BOTANY) 37

A lderw. (Blume) (Mett. ex Baker) (Wall. ex Hook.) (Wall. Blume (Baker) Christ Christ Christ (Miq.) C.Chr. Blume (Retz.) Sw. C.Presl Blume (Mett.) (Mett.) Tardieu (Mett.) Blume (Hayata) M.Kato Blume Tardieu auct. non (Willd.) M.Kato / auct. non (Willd.) Tardieu NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Deparia boryana Deparia subfluvialis Diplazium accedens Diplazium bantamense Diplazium conterminum Diplazium cordifolium Diplazium crenato-serratum T.Moore Diplazium dilatatum Absent Diplazium donianum Diplazium esculentum Diplazium heterophlebium Diels Diplazium leptophyllum Diplazium malaccense Diplazium megaphyllum Diplazium mettenianum Diplazium muricatum Diplazium petelotii Diplazium petri Diplazium polypodioides Diplazium prescottianum T.Moore Absent A lderw. (Blume) (Mett. ex Baker) (Wall. ex Hook.) (Wall. Blume Christ (Baker) Christ Christ (Miq.) C.Chr. Blume C.Presl Blume (Retz.) Sw. (Mett.) (Mett.) Tardieu (Mett.) Blume Blume auct. non (Willd.) Tagawa auct. non (Willd.) Tardieu Tardieu NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Athyrium boryanum Diplazium accedens Diplazium bantamense Diplazium conterminum Diplazium cordifolium Diplazium crenato-serratum T.Moore Diplazium dilatatum Absent Diplazium donianum Diplazium esculentum Diplazium heterophlebium Diels Diplazium leptophyllum Diplazium malaccense Diplazium megaphyllum Diplazium mettenianum Diplazium muricatum Diplazium petelotii Diplazium petri Diplazium polypodioides Diplazium prescottianum T.Moore Absent 26 10 lderw. A lderw. (Blume) (Mett. ex Baker) (Wall. ex Hook.) (Wall. (Baker) Christ Blume (Miq.) C.Chr. (Miq.) C.Chr. Christ Christ Holttum (Luerss.) Makino Blume C.Presl Blume (Retz.) Sw. (Retz.) Sw. (Mett.) (Mett.) Tardieu Tardieu (Mett.) Blume Blume

(Hayata) M.Kato Tardieu Tardieu Tardieu Tardieu CURRENT NAME CURRENT Deparia subfluvialis Diplazium accedens Diplazium bantamense Diplazium conterminum Diplazium cordifolium Diplazium crenato-serratum T.Moore Diplazium dilatatum Diplazium doederleinii Diplazium donianum Diplazium esculentum Diplazium heterophlebium Diels Diplazium leptophyllum Diplazium malaccense Diplazium megaphyllum Diplazium mettenianum Diplazium muricatum Diplazium petelotii Diplazium petri Diplazium polypodioides Diplazium prescottianum T.Moore Diplazium procumbens ● ○ ○ Towards a stable nomenclature for Thai ferns (S. LINDSAY, D.J. MIDDLETON, T. BOONKERD & S. SUDDEE) 103 Holttum (C.Presl) C.Presl (Baker) C.Chr. (Wall. ex Hook. & Grev.) ex Hook. & Grev.) (Wall. Blume Holttum Blume Tagawa (Bory) Sw. C.Chr. Holttum NAME IN FLORA OF THAILAND OF NAME IN FLORA & Iwatsuki, 1979, 1985, 1988, 1989) (Tagawa Diplazium riparium Diplazium siamense Diplazium silvaticum Diplazium simplicivenium Diplazium sorzogonense Diplazium subintegrum Diplazium subserratum Diplazium subsinuatum Tagawa Diplazium taiwanense Diplazium tomentosum Absent Diplazium xiphophyllum Holttum (C.Presl) C.Presl (Baker) C.Chr. (Wall. ex Hook. & Grev.) ex Hook. & Grev.) (Wall. Blume Holttum Blume Tagawa (Bory) Sw. C.Chr. Holttum NAME IN PTERIDOPHYTES IN THAILAND NAME IN PTERIDOPHYTES (Boonkerd & Pollawatn, 2000) Diplazium riparium Diplazium siamense Diplazium silvaticum Diplazium simplicivenium Diplazium sorzogonense Diplazium subintegrum Diplazium subserratum Diplazium subsinuatum Tagawa Diplazium taiwanense Diplazium tomentosum Absent Diplazium xiphophyllum

26 Holttum (C.Presl) C.Presl (Baker) C.Chr. (Baker) C.Chr. (Wall. ex Hook. & Grev.) ex Hook. & Grev.) (Wall. Blume Holttum Blume Tagawa Tagawa (Bory) Sw. (Bory) Sw.

Kunze C.Chr. C.Chr. Holttum CURRENT NAME CURRENT Diplazium riparium Diplazium siamense Diplazium silvaticum Diplazium simplicivenium Diplazium sorzogonense Diplazium subintegrum Diplazium subserratum Diplazium subsinuatum Tagawa Diplazium taiwanense Diplazium tomentosum Diplazium virescens Diplazium xiphophyllum ○ 104 THAI FOREST BULLETIN (BOTANY) 37

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