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Chapter 9 Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review

Marla D. Schwarzfeld* Department of Biological Sciences, CW 405 Biological Sciences Centre, University of Alberta, Edmonton, AB, T6G 2E9 Email: [email protected]

Abstract. The parasitoid family Ichneumonidae is the largest family in the order Hymenoptera. This chapter provides a checklist of 1,160 ichneumonid species (299 genera) known from the Canadian Prairies Ecozone. The list is primarily drawn from literature records and also includes 35 newly recorded species from the ecozone. The number of species on the list is a vast underestimate of the number of ichneumonid species present, as many genera lack revisions and few biodiversity surveys have been conducted. Most species recorded from this ecozone are only known from the Nearctic region, but are not restricted to the Prairies Ecozone. Little is known about the ecology, habitat requirements, or host associations of most ichneumonid species, with 43% of the species on the checklist lacking any host records. Future research should include revisions of the many genera that have not been studied in the Nearctic region, as well as biodiversity surveys in prairie habitats, rearing of potential host species, and the creation of user-friendly identification resources.

Résumé. Les guêpes parasitoïdes de la famille des Ichneumonidae forment la plus grande famille de l’ordre des hyménoptères. Le présent chapitre dresse la liste des 1 160 espèces de cette famille, réparties en 299 genres, présentes dans l’écozone des prairies. Cette liste, établie principalement à partir de sources documentaires, fait état de 35 nouvelles mentions provenant de l’écozone. Elle sous-estime cependant très largement le nombre d’espèces d’ichneumonidés présentes puisque de nombreux genres n’ont toujours pas fait l’objet d’une révision et que les études sur la biodiversité restent peu nombreuses. La plupart des espèces recensées dans cette écozone ne sont connues que dans la région néarctique, mais ne sont pas limitées à l’écozone des prairies. On sait peu de choses sur l’écologie, les besoins en matière d’habitat ou les hôtes des ichneumonidés, et la liste n’a aucune information à fournir sur les hôtes de 43 % des espèces répertoriées. Les recherches futures devraient inclure des révisions des nombreux genres qui n’ont toujours pas été étudiés dans la région néarctique, l’étude de la biodiversité dans les habitats des prairies, l’élevage d’espèces hôtes possibles et la création de ressources documentaires d’identification facile à utiliser.

Introduction

Ichneumonidae is the largest family within the order Hymenoptera and may be among the largest insect families on earth (Gauld 1991). It consists of at least 24,268 described species (Yu et al. 2012) and has an estimated world total of over 100,000 species (Gauld 2002). The family is monophyletic and is one of two families (with Braconidae) that make up the superfamily Ichneumonoidea (Sharkey and Wahl 1992;

Schwarzfeld M. D. 2014. Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review. In Arthropods of Canadian Grasslands (Volume 4): Biodiversity and Systematics Part 2. Edited by D. J. Giberson and H. A. Cárcamo. Biological Survey of Canada. pp. 317-397. © 2014 Biological Survey of Canada. ISBN 978-0-9689321-7-9 doi: http://dx.doi.org/10.3752/9780968932179.ch9 Species checklist available at http://dx.doi.org/10.5886/5r772nsm *Current address: University of Northern British Columbia,3333 University Way, Prince George, BC, V2N 4Z9 318 M. D. Schwarzfeld

Belshaw et al. 1998). Ichneumonoidea has been hypothesized to be the sister group of Aculeata (Rasnitsyn 1988; Dowton and Austin 1994); more recent studies, however, have not supported this relationship. Instead, strong evidence suggests that the superfamily is sister to Proctotrupomorpha (Heraty et al. 2011; Sharkey et al. 2012), or to Aculeata + Proctotrupomorpha (Sharanowski et al. 2010). According to the most recent taxonomically comprehensive analysis, 38 subfamilies are currently recognized within Ichneumonidae (Quicke et al. 2009). Their boundaries and relationships are still highly unresolved, however, and several subfamilies are currently paraphyletic (Quicke et al. 2009). Parasitoids are insects that are free-living as adults, but whose larvae live and feed entirely on or in a single host arthropod, ultimately killing it (Godfray 1994). With the exception of a few species that are predators on spider eggs (Fitton et al. 1987) and a very few that may be partly herbivorous (Gauld 1983), all ichneumonids are parasitoids (Gauld and Bolton 1988). Within this basic lifestyle, however, there is a wide range of life histories. Most species parasitize Holometabola, in particular Lepidoptera and Symphyta, but a wide range of hosts is known, including several species that attack Araneae (Fitton et al. 1987; Gauld and Bolton 1988). Ichneumonidae species can be classified as either idiobionts (parasitoids that prevent their hosts from developing further) or koinobionts (parasitoids that allow the host to continue to develop) (Askew and Shaw 1986). Within each of these strategies, they can be further classified as ectoparasitoids or endoparasitoids. All host life stages (egg, larval, pupal, and rarely, adult) are prone to attack by different species of Ichneumonidae. Finally, ichneumonids can be primary parasitoids and either obligate or facultative hyperparasitoids (Gauld and Bolton 1988; Gauld 1991). Some ichneumonid species are highly host specific, whereas others are generalists. In general, koinobionts are thought more likely to be host specialists because of being highly physiologically intertwined with their hosts, whereas idiobionts may have a wider host range (Askew and Shaw 1986; Gauld 1988a), but are often specific in a particular niche (e.g., hosts in silk, or hosts in woody material) (Gauld 1988a; Godfray 1994).

Summary of Taxonomic and Biodiversity Studies in the Canadian Prairies Ecozone Despite the abundance, diversity, and ecological importance of Ichneumonidae, there is a dearth of ecological studies or biodiversity surveys on them in general, and a near absence of such studies in the Canadian Prairies. The information contained within this chapter is thus almost entirely drawn from taxonomic studies. In particular, a great many Nearctic groups were revised by H. K. Townes and C. E. Dasch, though many other workers made great contributions to the knowledge of Nearctic species. A preliminary list of the natural enemies of Canadian agricultural pests was drawn from the literature by Graham (1965), though no localities beyond province were provided. Bradley (1974, 1978) compiled lists of forest Lepidoptera parasitoids in three ichneumonid subfamilies (Ichneumoninae, Metopiinae, Pimplinae) from rearings and museum collections. Although these species were examined in the context of forest pest enemies, many records are from the Prairies Ecozone. Sharkey et al. (1987) studied the parasitoids associated with sunflower in Alberta and Saskatchewan. Finnamore (1994) conducted a survey of the Hymenoptera of Wagner Natural Area, a spring fen within the Aspen Parkland Ecoregion, which included an estimated 665 species of Ichneumonidae. Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 319

Extensive collecting by E.H. Strickland led to the publication of his checklist of Alberta Ichneumonoidea (1946, 1952), with many records from the Prairies Ecozone. In Manitoba, many ichneumonid species (as well as other taxa such as Carabidae; see Chapter 1, this volume) are known from the collecting efforts of N. Criddle and associates around Aweme. In this chapter, I provide a checklist of all Ichneumonidae species recorded from the Canadian Prairies Ecozone, with localities and a summary of host records. This list consists primarily of published records, with the addition of 35 new records for the region. I also summarize the biogeographical and ecological patterns of these species in the Canadian Prairies and discuss the gaps in our current knowledge.

Approach Region The Canadian Prairies Ecozone extends across the southern portions of Alberta, Saskatchewan, and Manitoba. It includes seven ecoregions, most of which are grassland dominated (Shorthouse 2010). Greater amounts of tree cover are found in the Aspen Parkland Ecoregion, which is a transitional zone between prairie grasslands and boreal forest; in the Cypress Upland Ecoregion; and in the Southwest Manitoba Uplands Ecoregion (Shorthouse 2010). This chapter lists all ichneumonid species that have been recorded from the Prairies Ecozone. Since habitat associations were not given for the vast majority of records, locality was used as the criterion for whether a species should be included in the checklist. When in doubt, I erred on the side of including species. For example, many records are from Waterton Lakes National Park, Alberta, with no further data. Since the park includes the ecologically important Fescue Grassland Ecoregion (Shorthouse 2010), I included these records, although they are equally likely to have come from non-prairie habitats at higher elevations. I also used a relaxed northern border; since the Aspen Parkland Ecoregion has been highly altered by agriculture, and this agricultural land extends beyond what would have historically been parkland habitat and into the boreal, I used the current agricultural limit as an approximate northern boundary. This limit is arbitrary since few species appear to be restricted to the Prairies Ecozone; most species found in the Aspen Parkland Ecoregion have ranges extending into the Boreal Plains Ecozone. I did not include records from the Prairie Provinces (Alberta, Saskatchewan, and Manitoba) that had no locality data beyond province; many of these are undoubtedly from the Prairies Ecozone as well. The Ichneumonidae subfamilies are summarized in Table 1, and a checklist of the Ichneumonidae of the Prairies Ecozone is provided in Table 2.

Literature Review This checklist consists almost entirely of previously published species records. To obtain the list, I extracted all published records of Ichneumonidae from Alberta, Saskatchewan, and Manitoba from the Taxapad database of Ichneumonoidea (Yu et al. 2012). Each reference was examined for locality data, and the list was pruned to include only those species that were recorded from the Prairies Ecozone. In some cases, records were excluded where subsequent revisions rendered them doubtful. For example, many species recorded from the Prairies Ecozone of Alberta by Strickland (1946, 1952) belong to genera that have since been revised by researchers who had access to his collections. I therefore excluded records that are, according to the more recent revisions, well outside 320 M. D. Schwarzfeld

the range of the species. In addition, to avoid duplicating records where a subsequent revision almost certainly referred to the same specimen(s) as an earlier revision or description, I included only the most recent revision in Table 2. For non-revisionary work (e.g., Strickland 1946, 1952; Bradley 1974, 1978), all records were included, though this may have resulted in multiple records of the same specimens. I excluded any records that were not identified to the species level. Finally, the checklist contains 113 species that were not part of the Taxapad-generated list (Yu et al. 2012). Seventy-eight of these records are from the literature and 35 are newly recorded from the Prairies Ecozone in this publication. With the exception of newly published records from my own work, I did not examine specimens to confirm any of the species identities included herein, as the scope of such a project would be monumental.

Unpublished Records Unpublished records were extracted from the database of the Wallis-Roughley Museum of Entomology at the University of Manitoba, Winnipeg, Manitoba. These consist of 149 new locality records and include 25 species that had not been previously recorded from the Prairies Ecozone. An additional 103 new locality records, including 10 new species records, are included from a survey of Pimplinae, Poemeniinae, and Rhyssinae in the Aspen Parkland Ecoregion of Alberta (MDS, unpublished). All previously unpublished localities and species records are in bold in Table 2.

Host Records A list of all recorded hosts for the species included in the checklist was extracted from the Taxapad Ichneumonoidea database (Yu et al. 2012), and the number of host species recorded for each ichenumonid species is included in Table 2. This includes all records from throughout the range of the parasitoid, rather than being limited to the Canadian Prairies. The host families and guilds parasitized by each species were summarized by J. Dombroskie (Cornell University Insect Collection). I did not distinguish between primary and secondary hosts; for most hyperparasitoids, records of both are included.

Diversity and Distribution Patterns Species and localities from 124 publications are recorded in the checklist (Table 2). Thirty- one of these publications are of an ecological or experimental nature; the remaining 93 references consist of revisions, checklists, and compilations of data. Species from 24 subfamilies of Ichneumonidae are recorded from the Canadian Prairies Ecozone (Table 1). The subfamily Adelognathinae (Adelognathus) has been collected in the Aspen Parkland Ecoregion of Alberta (Finnamore 1994; MDS, unpublished) and in Spruce Woods Provincial Park, Manitoba (B. Sharanowski, pers. comm.); however, no specimens have been identified to species. One species from the Palearctic subfamily Collyrinae (Collyria coxator (Villers)) was introduced to Swift Current, Saskatchewan, in the 1930s and to Lethbridge, Alberta, in 1960 to control the wheat stem sawfly (Cephus cinctus Norton) (Smith 1931, 1961). However, there is no evidence that the species became established at either location (Smith 1961; Shanower and Hoelmer 2004; Wahl et al. 2007; Cárcamo and Beres 2013). Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 321

Table 1. Summary of Ichneumonidae subfamilies recorded from the Canadian Prairies Ecozone. Abbreviations: K = Koinobiont; I = Idiobiont; En = Endoparasitoid; Ec = Ectoparasitoid; H = Hyperparasitoid.

Subfamily Biology1 Hosts1 Nearctic Identification Resources Genera Species Acaenitinae K, En Concealed Coleoptera Townes and Townes 1960 3 5 Anomaloninae K, En Lepidoptera, Dasch 1984 10 64 Coleoptera Banchinae K, En Lepidoptera Townes and Townes 1978 (Banchini 12 136 and Atrophini); Dasch 1988 (Glyptini) Campopleginae K, En Lepidoptera (most); Genera: Townes 1970b; many species 22 142 also Symphyta, described by Viereck (1925–1926a,b); Coleoptera, revisions include Walley 1947; Raphidioptera Sanborne 1984, 1986a,b,c; Wahl 1987 Cremastinae K, En Concealed Dasch 1979 6 36 Lepidoptera (most); also Coleoptera Cryptinae I, Ec Holometabola (most); Genera: Townes 1970a; Species: 58 143 (most) also Araneae egg sacs Townes and Townes 1962 (Cryptini); Townes and Gupta 1962 (Hemigasterini); Townes 1983 (some Phygadeuontini); Luhman 1990 (Endasys) Ctenopelmatinae K, En Symphyta (most); Genera: Townes 1970b; several genera 31 78 also Lepidoptera revised by Barron (1981, 1986, 1990, 1992, 1994, 1997) and Leblanc (1989, 1999) Cylloceriinae K, En Tipulidae Dasch 1992 2 3 Diacritinae Unknown Unknown Dasch 1992 1 1 Diplazontinae K, En Syrphidae Dasch 1964 11 45

Eucerotinae H2 Ichneumonoidea Barron 1976 1 6 Ichneumoninae3 K/I, En Lepidoptera Heinrich 1960a,b, 1961a,b, 1962a,b,c 46 154 Lycorininae K4, En?/ Small leaf-rolling Gauld and Wahl 2002 1 1 Ec4 Lepidoptera Mesochorinae K, H Ichneumonoidea Dasch 1971 4 49 (most); also Tachinidae Metopiinae K, En Lepidoptera Townes and Townes 1959 10 52 Ophioninae K, En Lepidoptera (most); Nearctic genera: Gauld and Wahl 3 5 also Scarabaeidae 2002; Enicospilus (part): Gauld 1988b Orthocentrinae K, En Mycetophilidae, Dasch 1992 15 34 Sciaridae Orthopelmatinae ?, En Cynipidae in galls Barron 1977 1 3 on Rubus, Rosa Pimplinae K/I, Ec/ Holometabola (most); Townes and Townes 1960 29 75 En also Araneae 322 M. D. Schwarzfeld

Subfamily Biology1 Hosts1 Nearctic Identification Resources Genera Species Poemeniinae I?, Ec Wood-boring Townes and Townes 1960 4 7 Coleoptera and Hymenoptera Rhyssinae I, Ec Wood-boring Townes and Townes 1960 3 6 Symphyta and possibly Coleoptera Tersilochinae3 K, En Coleoptera (most); Genera: Townes 1971; Nearctic 3 6 also Symphyta species: Horstmann 2010, 2012 Tryphoninae K, Ec Symphyta, Genera: Townes 1969; Nearctic 21 104 Lepidoptera species: Mason 1955, 1956, 1959; Townes et al. 1992; Townes and Townes 1949a,b; Loan 1981 Xoridinae I, Ec Wood-boring Townes and Townes 1960 2 5 Coleoptera (most) and possibly Symphyta

1 Wahl (1993), except where noted. 2 A summary of the unusual life history of Eucerotinae can be found in Barron (1976). 3 Several additional species listed from the Prairie Provinces with no localities. 4 Shaw (2004); early instars are possibly endoparasitic, but are probably concealed ectoparasitoids within the host’s anus.

A total of 1,160 species in 299 genera are included in the checklist of Ichneumonidae of the Prairies Ecozone (Table 2). This is a significant underestimate of the true number of species present since many genera have not been revised and a great many species await description. For example, the genus Ophion (Ophioninae) consists of 11 described Nearctic species, of which two have been recorded in the literature from Alberta, Saskatchewan, and Manitoba. However, an ongoing taxonomic study of Ophion, using morphological and molecular methods, has found a minimum of 21 species in the Prairies Ecozone of Alberta alone (MDS, unpublished). A similar pattern no doubt exists for many genera. Some examples of species-rich genera in desperate need of revision include Campoplex, Diadegma, and Hyposoter (Campopleginae); Gelis and Phygadeuon (Cryptinae); Alexeter and Mesoleius (Ctenopelmatinae); Orthocentrus (Orthocentrinae); Netelia (Tryphoninae); and many others. For genera such as these, this list is clearly inadequate, though it does provide a minimum starting point. The subfamily Ichneumoninae is among the most diverse ichneumonid subfamilies, with only Cryptinae having a greater number of described species; however, the species list provided here is preliminary. Ichneumoninae species were revised by Heinrich (1960a, 1960b, 1961a, 1961b, 1962a, 1962b, 1962c), but he focused on species from northeastern North America and in most cases provided only the province of collection rather than specific localities. I have thus included Strickland’s (1946, 1952) species records of Ichneumoninae, but since these were recorded before the publication of Heinrich’s synopses, the identifications are tentative. In addition to the lack of revisions for many taxa, there is a lack of biodiversity studies for the region. For example, Pimplinae, Poemeniinae, and Rhyssinae are among the best known and most easily identified ichneumonid subfamilies; however, a recent survey of Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 323

these three subfamilies in Alberta found 10 species that had not previously been recorded from the Prairies Ecozone. Nine of these species are new Alberta records and eight are newly recorded from the Prairie Provinces. All of these records are from the Aspen Parkland Ecoregion; a similar pattern would no doubt be seen if a survey were conducted in the prairie grasslands, and this would be greatly magnified for those subfamilies that are less well-known. This study emphasizes the importance of conducting biodiversity surveys and of publishing species records, as many commonly collected and easily identified species may be missing from the published record. Museum collections are another important yet underutilized source of biodiversity information. Twenty-five species are newly recorded from the Prairies Ecozone from the Wallis-Roughley Museum of Entomology, the only museum in the Prairies region to have databased all of its Ichneumonidae. Databasing specimens and creating publicly available databases of museum specimens is thus a high priority to increase the usability of museum data.

Biogeography Very few ichneumonid species are restricted to the Canadian Prairies Ecozone. Ninety- seven percent of the species listed in Table 2 (1,124 species) have also been recorded from outside the Prairies Ecozone, all except six having records from outside the three Prairie Provinces (Alberta, Saskatchewan, and Manitoba). Of the remaining 36 species, most (31 species) are known only from the type locality. Within the ecozone, however, little is known about the distribution of most species. Forty-four percent of the species in the checklist (512 species) have been recorded from only a single locality within the Prairies Ecozone (though, as discussed earlier, most are also found outside of this region). Only 6% of the species have been recorded from 10 or more localities within the region. Most species included in the checklist (75%, 873 species) have a Nearctic distribution, whereas 36 species are restricted to the Nearctic and Neotropical regions. A total of 182 species are Holarctic, with the remainder being distributed across multiple terrestrial ecozones. In several cases, Holarctic species are separated into Nearctic and Palearctic subspecies; few molecular studies have been done to test their conspecificity.

Habitat and Host Associations While in most cases definitive habitat associations are not given, few species of Ichneumonidae appear to be strongly associated with grassland habitats. Where habitat is indicated, the associations are often with trees or shrubs. Those species with notes that specifically indicate that they have been collected from grasslands or other prairie habitats (e.g., sand dunes, agricultural fields) are marked with an asterisk in Table 2. This does not imply, however, that other species lack this association, as in most cases no habitat association is recorded. The basic biology of most Ichneumonidae is known at the subfamily or genus level (e.g., Wahl 1993, Table 1). For many species, however, reliable rearing records are lacking. Of the species recorded from this region, 43% (493 species) have no host records, while many of the remaining species have been recorded from only one or two host species. Only 33% of the species in Table 2 have been recorded from three or more host species. While this could be an indication of host specificity, in many cases, it indicates a lack of rearing records, as most rearing studies have been conducted on a limited number of pest species (Askew and Shaw 1986). One or two host records provide useful insight into the biology 324 M. D. Schwarzfeld

of the species, but are clearly inadequate to understand the true host range. In addition, literature records are often unrepresentative or incorrect (Askew and Shaw 1986; Mills 1992). For example, the subfamily Diplazontinae is believed to be exclusively associated with Syphidae (Diptera) (Fitton and Rotheray 1982; Klopfstein et al. 2011); however, of the 24 diplazontine species included in the checklist that have host records, 17 have been recorded from other families and 16 from non-Diptera. Of the 667 species with host records, the vast majority (493 species, 74%) have been reared from Lepidoptera, particularly from Tortricidae (189 species), followed by Noctuidae (152) and Geometridae (133). Twenty-eight percent (186 species) are recorded from Hymenoptera, with 77% of these from sawflies (Symphyta). A total of 145 species have been recorded from more than one arthropod order, although, as in the case of Diplazontinae, some of these may be in error.

Future Research This chapter provides a preliminary checklist of Ichneumonidae of the Canadian Prairies Ecozone. While a useful starting point, the principal message is that Ichneumonidae are incredibly diverse in this region and that we know remarkably little about them. Among the most pressing research needs is an increase in taxonomic studies, particularly for the many genera that have never been properly revised. In addition, most existing revisions were conducted before the advent of molecular techniques; even among those taxa that are relatively well-known, there is great scope for discovery. In order to understand the roles that ichneumonids play in prairie ecosystems, there is a great need for both biodiversity surveys and ecological studies (including rearing of hosts). These would allow documentation of ichneumonid diversity, as well as habitat and host associations, which would in turn provide insight into host–parasitoid dynamics, biological control of agricultural pests, ichneumonid habitat requirements, impacts of ecological disturbance, and much more. Finally, a high priority should be placed on creating user-friendly keys and other identification resources, such as online species pages. These would serve to facilitate and encourage studies into the ecology and biodiversity of this important group of parasitoids.

Acknowledgements Thank you to Gary Anweiler and Felix Sperling for running Malaise traps in Edmonton and at Pigeon Lake, and to the many members of the Alberta Lepidopterists’ Guild who supplied me with Ophion. A huge thank you to Jason Dombroskie (Cornell University Insect Collection) for summarizing the host families and guilds. I am grateful to Andy Bennett (Canadian National Collection) for assistance with Pimplinae identifications and Barb Sharanowski (University of Manitoba) for access to the Wallis-Roughley Museum database. Thank you to Felix Sperling, Barb Sharanowski, and Andy Bennett for very helpful comments on earlier drafts of this manuscript. Finally, I would like to thank Dicky Yu and co-authors for their work in compiling the immensely valuable Taxapad Ichneumonoidea database, which formed the backbone of this manuscript. This work was supported by the Sustainable Forest Management Network, ACA Grants in Biodiversity (supported by the Alberta Conservation Association), Natural Sciences and Engineering Research Council (NSERC), and Alberta Ingenuity student scholarships to the author and an NSERC Discovery grant to Felix Sperling. Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 325 Borers Borers Leafrollers Host Family (No. spp.) Guild/Notes Noctuidae (10) Lepidoptera: Geometridae, Lycaenidae, Mostly Geometridae None None Coleoptera: Cerambycidae (1) None Coleoptera: Cerambycidae (2) Lepidoptera: Drepanidae, Erebidae, Geometridae, Hesperiidae, Noctuidae, Notodontidae, Nymphalidae, (14) Tortricidae Sphingidae, Lepidoptera: Drepanidae, Erebidae, Lasiocampidae, Notodontidae (13) Mostly setose None None (1) Tortricidae Lepidoptera: Hymenoptera: Cynipidae; Lepidoptera: Coleophoridae, Erebidae, Gelechiidae, Geometridae, Micropterigidae, Yponomeutidae, Notodontidae, Tortricidae, (34) Ypsolophidae (3) Lepidoptera: Lycaenidae

m , MB: South m b were estimated from mapped localities; localities in boldface type are previously unpublished locality records. Species type are previously unpublished locality in boldface localities from mapped localities; were estimated m Localities AB: Bilby; SK: Fort Pitt NA AB: Wagner MB: Treesbank NA; MB: Teulon AB: Wagner MB: Teulon AB: Southwest Wabamun, Taber, Orion, Jumping Pound Cr, AB: Edmonton, Elkwater P, Kenosee, Lloydmister, SK: Estevan, Great Deer, Waterton; NA, Wagner Binscarth, Glenboro, Keld, Aweme, Roche Percée, Saskatoon; MB: Shilo,Transcona Mt FR, 30 mi N Roblin, 5 SW Turtle Edmonton, Enilda, Seba Lakes NP, Waterton AB: Big Chief Hwy in Beach; MB: Concord, 5 mi N Minnedosa NA AB: Wagner Waterton AB: Mountain View, Shilo MB: Ninette, 5 mi SW Wabamun; AB: Edmonton, Elkwater L, Wagner AB: Elkwater L, Fort MacLeod, P; SK: Birch Cr; Waterton AB: Lundbreck, Red Rock Canyon in MB: Aweme 4,N 4,H,O 1,2,N 1,N 3,N 2,N 1,2,N 4,N 2,4,N 4,N 2,N,T 1,N http://dx.doi.org/10.5886/5r772nsm 4,N Townes, 1960 Townes, (Provancher, 1886) (Provancher, Cushman, 1920 (Cresson, 1869) (Ashmead, 1902) Schmid, 1970 (Say, 1829) (Say, a SUBFAMILY Species ACAENITINAE 1920 maurus Rohwer, Arotes Agrypon alpinum (Davis, 1898) Agrypon clandestinum (Gravenhorst, 1829) Coleocentrus manni formosum (Cresson, 1868) Spilopteron vicinum Spilopteron Agrypon anale Dasch, 1984 Agrypon annulare Agrypon carinifer Agrypon chlamidatum Agrypon cushmani Dasch, 1984 Spilopteron franclemonti Spilopteron ANOMALONINAE Agrypon alaskense al. et (2012) and are Yu defined followingfrom host records are and abbreviations) by superscript (indicated distributions the checklist. Species in bold are newly recorded from habitats. prairie other or dunes, sand fields, agricultural grasslands, from recorded specifically been have asterisk an with marked those and publication this in Ecozone Prairies the Species checklist available at Table 2. Checklist Table of Ichneumonidae recorded from the Canadian Prairies Ecozone. Locality references are identified bywith marked Localities checklist. following the listed superscript numbers following each species and are 326 M. D. Schwarzfeld Leafminers Leafrollers Cutworms Host Family (No. spp.) Guild/Notes (7) Tortricidae Lepidoptera: Geometridae, Noctuidae, Lepidoptera: Drepanidae, Geometridae (2) Loopers Lepidoptera: Crambidae, Notodontidae (2) Lepidoptera: Geometridae (1) (7) Tortricidae Lepidoptera: (1) Tenthredinidae Hymenoptera: Lepidoptera: Crambidae, Elachistidae, Erebidae, Geometridae, Nolidae, Nymphalidae, Pterophoridae, (21) Tortricidae Mostly leafrollers Lepidoptera: Notodontidae, Sphingidae (4) Most spp large None Lepidoptera: Erebidae, Geometridae, Micropterigidae, (20) Yponomeutidae Oecophoridae, Tortricidae, Mostly leafrollers None Lepidoptera: Erebidae, Geometridae, Noctuidae (5) Mostly loopers None Lepidoptera: Geometridae (1) Lepidoptera: Geometridae, Noctuidae (13) Mostly loopers Lepidoptera: Geometridae, Noctuidae (4) Mostly loopers Lepidoptera: Geometridae (4) None None Lepidoptera: Geometridae (1) Lepidoptera: Noctuidae (1) Lepidoptera: Erebidae, Noctuidae (2) Setose Lepidoptera: Erebidae (1) None Lepidoptera: Erebidae, Noctuidae (3) Lepidoptera: Erebidae, Noctuidae (4) Mostly cutworms Lepidoptera: Erebidae (5) Setose None Lepidoptera: Erebidae (1) None None Lepidoptera: Noctuidae (3) None None None None None None None None m , S of Lethbridge m , nr Waterton m b Localities NA AB: Wagner Shilo AB: Edmonton; MB: 5 mi SW AB: Wabamun Shilo NA; SK: Scout L; MB: 5 mi SW Wagner AB: Drumheller, Mt Turtle AB: Pincher; MB: Red View, AB: Derwent, Edmonton, Elk Point, Gull L, Mountain SK: Kenosee, Rockglen, Waterton; Wainwright, NA, Wagner Deer, Mt; Wood Bunch, Willow Saskatchewan Landing, Saskatoon, Scout L, Bog at 5 Tamarack FR, Woods Horton, Sandridge, Spruce Aweme, MB: Winnipeg Shilo, mi SW AB: nr Edmonton NA; SK: Snowden Wagner Edmonton, AB: Calgary, Kenosee, AB: Edgerton, Irvine, Lethbridge, Morrin; SK: Elbow, Douglas, Hartney Aweme, Mossbank, Saskatoon, Uren; MB: SK: Elbow Wabamun; AB: Edmonton, Jumping Pound Cr, AB: Brant, Brooks, Bow Island, Clymont, Edmonton, Hays, Irvine, SK: Dana, Dunblane, Wainwright; Lethbridge, Medicine Hat, Scandia, Albert, Redberry, Parkbeg, Prince Moose Jaw, Great Deer, Elbow, Saskatoon, Saskatchewan Landing, Strongfield, Swift Current; MB: Bald Hills at 13 mi N Glenboro Aweme, Treesbank Woods, Albert; MB: Spruce SK: Prince Shilo Bog at 5 mi SW Tamarack MB: AB: Edmonton; MB: Ninette 3 mi Aweme, Lakes NP; SK: Glaslyn, Pierceland; MB: Waterton AB: Treesbank SW Shilo, Spruce Woods, MB: Birds Hill, Pine Ridge AB: Edmonton MB: Winnipeg AB: MB: Aweme Waterton; MB: Ninette Victor; SK: St. SK: Tunstall Whitemouth Edmonton, Mayerthorpe; SK: Pelly; MB: AB: Drumheller, Winnipeg Arcola; MB: AB: Calgary; SK: AB: Waterton MB: Millwood Valley; Elkwater L; SK; Fox Elkwater, AB: Calgary, Great Sand Hills, Lancer, AB: Calahoo, Grassy L; SK: Earl Grey, Otterburne Aweme, Rosthern, Saskatoon; MB: AB: Jumping Pound Cr SK: Saskatoon Ninette Aweme, AB: Edmonton, Lethbridge; SK: Kenosee, Oxbow; MB: SK: Willows AB: Edmonton Lethbridge, Morrin, Edmonton, Grimshaw, Drumheller, AB: Czar, Albert Great Sand Hills, Prince SK: Elbow, Waterton; Onefour, AB: Edmonton, Ponoka SK: Saskatoon NA Wagner AB: Bilby, AB: Norton AB: Lethbridge AB: Clover Bar; SK: Saskatoon SK: Pike L Edmonton, Magrath AB: Calgary, 4,N 3,5,N 2,4,N,T 4,H,O 2,4,5,N 4,N 4,N 4,6,N 4,N 2,4,N 4,N 4,N 4,N 4,N 4,N 4,N 2,4,N 4,N 4,N 4,N 4,N 4,N 4,N 4,N 4,N 4,N 4,N 4,N 2,4,N 4,N 4,N 2,N,T 4,N 4,N 4,N 4,N 4,N 4,N 4,N 4,N Hopper, 1981 Hopper, Dasch, 1984 (Cresson, 1865) Dasch, 1984 Hopper, 1981 Hopper, (Dalla Torre, 1901) Torre, (Dalla Dasch, 1984 Hopper, 1981 Hopper, Dasch, 1984 (Norton, 1863) Hopper, 1981 Hopper, Hopper, 1981 Hopper, Dasch, 1984 (Davis, 1898) Dasch, 1984 (Cresson, 1872) Dasch, 1984 (Davis, 1898) (Provancher, 1886) (Provancher, Dasch, 1984 a Agrypon drepanae Dasch, 1984 Agrypon drepanae Agrypon metallicum Agrypon schizurae Dasch, 1984 SUBFAMILY Species Aphanistes heinrichi Agrypon extensor Dasch, 1984 Agrypon hyperetis Agrypon melleum Agrypon prismaticum (Norton, 1863) Agrypon provancheri Agrypon validum 1849) Agrypon varitarsum (Wesmael, Anomalon reticulatum Aphanistes coxatus Aphanistes decurvihastatus Aphanistes dreisbachi 1981 Hopper, Aphanistes enargiae Aphanistes hopperi Aphanistes manitobae 1981 Aphanistes masoni Hopper, Aphanistes walleyi Barylypa clavata Barylypa elongata Barylypa imitata Barylypa incompleta Barylypa sulcata ambiguus (Norton, 1863) *Erigorgus aquilonius Dasch, 1984 Erigorgus attenuatus Dasch, 1984 Erigorgus bilineatus Erigorgus Dasch, 1984 brooksi Erigorgus Dasch, 1984 compressus Erigorgus curtus (Norton, 1863) Erigorgus ferrugineus (Norton, 1863) Erigorgus niveus Dasch, 1984 Erigorgus nubilipennis Erigorgus planus Dasch, 1984 Erigorgus pumilus Dasch, 1984 Erigorgus (Davis, 1898) rotundus Erigorgus sinuosus Dasch, 1984 Erigorgus spinosus Dasch, 1984 Erigorgus Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 327 Leafminers Leafrollers Cutworms Host Family (No. spp.) Guild/Notes (7) Tortricidae Lepidoptera: Geometridae, Noctuidae, Lepidoptera: Drepanidae, Geometridae (2) Loopers Lepidoptera: Crambidae, Notodontidae (2) Lepidoptera: Geometridae (1) (7) Tortricidae Lepidoptera: (1) Tenthredinidae Hymenoptera: Lepidoptera: Crambidae, Elachistidae, Erebidae, Geometridae, Nolidae, Nymphalidae, Pterophoridae, (21) Tortricidae Mostly leafrollers Lepidoptera: Notodontidae, Sphingidae (4) Most spp large None Lepidoptera: Erebidae, Geometridae, Micropterigidae, (20) Yponomeutidae Oecophoridae, Tortricidae, Mostly leafrollers None Lepidoptera: Erebidae, Geometridae, Noctuidae (5) Mostly loopers None Lepidoptera: Geometridae (1) Lepidoptera: Geometridae, Noctuidae (13) Mostly loopers Lepidoptera: Geometridae, Noctuidae (4) Mostly loopers Lepidoptera: Geometridae (4) None None Lepidoptera: Geometridae (1) Lepidoptera: Noctuidae (1) Lepidoptera: Erebidae, Noctuidae (2) Setose Lepidoptera: Erebidae (1) None Lepidoptera: Erebidae, Noctuidae (3) Lepidoptera: Erebidae, Noctuidae (4) Mostly cutworms Lepidoptera: Erebidae (5) Setose None Lepidoptera: Erebidae (1) None None Lepidoptera: Noctuidae (3) None None None None None None None None m , S of Lethbridge m , nr Waterton m b Localities NA AB: Wagner Shilo AB: Edmonton; MB: 5 mi SW AB: Wabamun Shilo NA; SK: Scout L; MB: 5 mi SW Wagner AB: Drumheller, Mt Turtle AB: Pincher; MB: Red View, AB: Derwent, Edmonton, Elk Point, Gull L, Mountain SK: Kenosee, Rockglen, Waterton; Wainwright, NA, Wagner Deer, Mt; Wood Bunch, Willow Saskatchewan Landing, Saskatoon, Scout L, Bog at 5 Tamarack FR, Woods Horton, Sandridge, Spruce Aweme, MB: Winnipeg Shilo, mi SW AB: nr Edmonton NA; SK: Snowden Wagner Edmonton, AB: Calgary, Kenosee, AB: Edgerton, Irvine, Lethbridge, Morrin; SK: Elbow, Douglas, Hartney Aweme, Mossbank, Saskatoon, Uren; MB: SK: Elbow Wabamun; AB: Edmonton, Jumping Pound Cr, AB: Brant, Brooks, Bow Island, Clymont, Edmonton, Hays, Irvine, SK: Dana, Dunblane, Wainwright; Lethbridge, Medicine Hat, Scandia, Albert, Redberry, Parkbeg, Prince Moose Jaw, Great Deer, Elbow, Saskatoon, Saskatchewan Landing, Strongfield, Swift Current; MB: Bald Hills at 13 mi N Glenboro Aweme, Treesbank Woods, Albert; MB: Spruce SK: Prince Shilo Bog at 5 mi SW Tamarack MB: AB: Edmonton; MB: Ninette 3 mi Aweme, Lakes NP; SK: Glaslyn, Pierceland; MB: Waterton AB: Treesbank SW Shilo, Spruce Woods, MB: Birds Hill, Pine Ridge AB: Edmonton MB: Winnipeg AB: MB: Aweme Waterton; MB: Ninette Victor; SK: St. SK: Tunstall Whitemouth Edmonton, Mayerthorpe; SK: Pelly; MB: AB: Drumheller, Winnipeg Arcola; MB: AB: Calgary; SK: AB: Waterton MB: Millwood Valley; Elkwater L; SK; Fox Elkwater, AB: Calgary, Great Sand Hills, Lancer, AB: Calahoo, Grassy L; SK: Earl Grey, Otterburne Aweme, Rosthern, Saskatoon; MB: AB: Jumping Pound Cr SK: Saskatoon Ninette Aweme, AB: Edmonton, Lethbridge; SK: Kenosee, Oxbow; MB: SK: Willows AB: Edmonton Lethbridge, Morrin, Edmonton, Grimshaw, Drumheller, AB: Czar, Albert Great Sand Hills, Prince SK: Elbow, Waterton; Onefour, AB: Edmonton, Ponoka SK: Saskatoon NA Wagner AB: Bilby, AB: Norton AB: Lethbridge AB: Clover Bar; SK: Saskatoon SK: Pike L Edmonton, Magrath AB: Calgary, 4,N 3,5,N 2,4,N,T 4,H,O 2,4,5,N 4,N 4,N 4,6,N 4,N 2,4,N 4,N 4,N 4,N 4,N 4,N 4,N 2,4,N 4,N 4,N 4,N 4,N 4,N 4,N 4,N 4,N 4,N 4,N 4,N 2,4,N 4,N 4,N 2,N,T 4,N 4,N 4,N 4,N 4,N 4,N 4,N 4,N Hopper, 1981 Hopper, Dasch, 1984 (Cresson, 1865) Dasch, 1984 Hopper, 1981 Hopper, (Dalla Torre, 1901) Torre, (Dalla Dasch, 1984 Hopper, 1981 Hopper, Dasch, 1984 (Norton, 1863) Hopper, 1981 Hopper, Hopper, 1981 Hopper, Dasch, 1984 (Davis, 1898) Dasch, 1984 (Cresson, 1872) Dasch, 1984 (Davis, 1898) (Provancher, 1886) (Provancher, Dasch, 1984 a Agrypon drepanae Dasch, 1984 Agrypon drepanae Agrypon metallicum Agrypon schizurae Dasch, 1984 SUBFAMILY Species Aphanistes heinrichi Agrypon extensor Dasch, 1984 Agrypon hyperetis Agrypon melleum Agrypon prismaticum (Norton, 1863) Agrypon provancheri Agrypon validum 1849) Agrypon varitarsum (Wesmael, Anomalon reticulatum Aphanistes coxatus Aphanistes decurvihastatus Aphanistes dreisbachi 1981 Hopper, Aphanistes enargiae Aphanistes hopperi Aphanistes manitobae 1981 Aphanistes masoni Hopper, Aphanistes walleyi Barylypa clavata Barylypa elongata Barylypa imitata Barylypa incompleta Barylypa sulcata ambiguus (Norton, 1863) *Erigorgus aquilonius Dasch, 1984 Erigorgus attenuatus Dasch, 1984 Erigorgus bilineatus Erigorgus Dasch, 1984 brooksi Erigorgus Dasch, 1984 compressus Erigorgus curtus (Norton, 1863) Erigorgus ferrugineus (Norton, 1863) Erigorgus niveus Dasch, 1984 Erigorgus nubilipennis Erigorgus planus Dasch, 1984 Erigorgus pumilus Dasch, 1984 Erigorgus (Davis, 1898) rotundus Erigorgus sinuosus Dasch, 1984 Erigorgus spinosus Dasch, 1984 Erigorgus 328 M. D. Schwarzfeld Leafrollers Cutworms Cutworms Concealed feeders Lepidoptera: Erebidae, Geometridae, Lasiocampidae, Noctuidae, Notodontidae, Sphingidae (33) Host Family (No. spp.) Guild/Notes Lepidoptera: Erebidae, Noctuidae (10) Setose Hymenoptera: Pamphiliidae; Lepidoptera: Noctuidae (2) Lepidoptera: Elachistidae, Erebidae, Geometridae, Lasiocampidae, Noctuidae, Nolidae, Notodontidae, (26) Tortricidae Pyralidae, Saturniidae, Mostly leafrollers Lepidoptera: Tortricidae (1) Tortricidae Lepidoptera: None Lepidoptera: Noctuidae (3) (18) Tortricidae Lepidoptera: Geometridae, Pyralidae, Mostly leafrollers Lepidoptera: Geometridae (1) Lepidoptera: Saturniidae, Sphingidae (6) most spp social Large, Lepidoptera: Galacticidae, Gelechiidae, Olethreutidae, (7) Tortricidae Pyralidae, (1) Tortricidae Lepidoptera: Lepidoptera: Erebidae, Geometridae, Noctuidae, Notodontidae (16) None Lepidoptera: Erebidae, Geometridae, Noctuidae, Notodontidae, Nymphalidae, Sphingidae (15) Mostly setose Lepidoptera: Erebidae, Geometridae, Noctuidae (10) Mostly setose (6) Tortricidae Lepidoptera: Pyralidae, Concealed feeders Hymenoptera: Chrysididae, Diapriidae, Lepidoptera: Erebidae, Vespidae; Ichneumonidae, Noctuidae (10) Lepidoptera: Noctuidae (2) None Lepidoptera: Noctuidae (1) Cutworms None None Lepidoptera: Noctuidae (3) None None None Hymenoptera: Cynipidae; Lepidoptera: Choreutidae, Crambidae, Elachistidae, Erebidae, Galacticidae, Nolidae, Gelechiidae, Geometridae, Lycaenidae, (40) Tortricidae Psychidae, Pyralidae, Sesiidae,

, Brandon, Carberry, Horton, International Peace Horton, International Peace Birtle , Brandon, Carberry, 1 and 5 mi SW Shilo, Teulon, Winnipeg Teulon, Shilo, 1 and 5 mi SW

b Wainwright; SK: Attons L, Battle River, Bronson PF, Caron, Caron, Bronson PF, Attons L, Battle River, SK: Wainwright;

Cypress Hills PP, Prelate, Prince Albert, Radisson, Rutland; MB: Prelate, Prince Cypress Hills PP, Winnipeg Teulon, Ninette, Victor St. SK; Oxbow, Whyte, Sifton Ft. Clearwater, Aweme, MB: Altona , Localities Waterton; Tofield, L, Red Deer, Consort, Edmonton, George AB: Bilby, Regina, Saskatoon; MB: Birds Hill, at Cut Knife, Elbow, Attons L SK: Winnipeg Teulon, Ninette, Husavik, Killarney, Carberry, AB: Calgary Derwent, Edmonton, Lethbridge; SK: MacDowell, AB: Calgary, Winnipeg Ninette, 10 mi S Aweme, Saskatoon, Swift Current; MB: AB: 18 mi E Lacombe, Red Deer AB: Rimbey Thornhill MB: Valley, Pembina NA AB: Wagner Blackfoot Hills, Chedderville, Clymont, Aspen Beach, Bilby, AB: Edgerton, Edmonton, Parkland, Rockyford, Consort, Drumheller, Stavely, AB: Medicine Hat; Midcentral Canada (AB/SK) in sunflower fields Irvine, Lethbridge, Onefour, Aden, Edmonton, Elk Island P, AB: Treesbank Stockton, W 2 mi Aweme, Scandia; SK: Elbow; MB: Aspen Beach, Clymont, Cypress Hills, Edmonton, Elkwater, AB: NA; Wagner L, Golden Spike, Manyberries, 2 mi E Red Deer, George Mt; Wood Saltcoats, Saskatoon, Victor, St. Attons L, Neilburg, SK: AB: Cypress Hills, Green Court, Lethbridge, Red Deer; SK: Shilo MB: 5 mi SW Teulon Fort Garry, Aweme, MB: Wabamun; AB: Edmonton, Gull L, Riverton , St. Claude Aweme, NA; MB: Wagner AB: SK: Saskatoon, south in sunflower fields Waterton; AB: Bilby, AB: Edmonton Twin Butte; Tofield, Waterton, AB: Edmonton, Manyberries, Lethbridge, AB: Beauvallon, Blackfoot Hills, Elk Point, Hardisty, SK: Harris, Lebret, Rutland, Saskatoon; Wainwright; Medicine Hat, AB: Cochrane, Edmonton, Lethbridge, Medicine Hat; SK: Saskatoon MB: Aweme AB: Lethbridge SK: Saskatoon AB: Edmonton, Magrath; SK: Saskatoon Asquith, Clarkboro, AB: Edmonton, Lethbridge, Medicine Hat; SK: Saskatoon, White Fox AB: Medicine Hat NA Wagner AB: Edmonton, Lethbridge, Medicine Hat, AB: Lethbridge; SK: Saskatoon MB: Aweme, MB: Aweme, Mt FR, Millwood, 5 mi N Minnedosa, Ninette, Turtle Gardens at 3 mi S Shilo, sunflower fields Cut Knife, Donavon, Dunblane, Dundurn, Elbow, Ft a la Corne, Great Ft a la Corne, Great Cut Knife, Donavon, Dunblane, Dundurn, Elbow, Pike L, Great Sand Hills, Indian Head, Keppel, Lloydminster, Deer, White Fox; MB: Onah, Saskatoon, Snowden, Sutherland, Redberry, Midcentral Canada (AB/SK) in Winnipeg; Stockton, W 2 mi 4,8,N 4,N 4,5,7,8,H 5,11,N 4,N 4,N,T 4,N 5,9,12,N 5,11,H 5,11,13,N 2,5,11,N 2,N 11,N 2,4,N,T 4,N

4,N 4,N 11,N 5,11,N 2,4,N,T 11,N 2,4,9,10,N,T 11,N 11,14,N 4,8,N 11,N 4,N (Cresson, 1879) (Rohwer, 1915) (Rohwer, Cresson, 1868 Townes, 1978 Townes, (Cresson, 1870) (Holmgren, 1857) (Thunberg, 1824) (Thunberg, (Norton, 1863) (Cushman, 1922) (Provancher, 1874) (Provancher, Walley, 1931 Walley, Townes, 1978 Townes, Townes, 1978 Townes, Cresson, 1870 a 2,4,5,9,H,O Erigorgus stenotus Dasch, 1984 Erigorgus utahensis Dasch, 1984 Erigorgus Therion magnum Dasch, 1984 Apophua simplicipes canadensis *Arenetra SUBFAMILY Species Therion fuscipenne Erigorgus xanthopsis (Ashmead, 1890) Erigorgus aclerivorus Habronyx amoenus Dasch, 1984 Habronyx magniceps (Cresson, 1872) Habronyx Ophionellus foutsi Therion californicum (Linnaeus, Therion circumflexum 1758) Therion morio (Fabricius, 1781) 1917 Therion sassacus Viereck, maceratum (Cresson, 1879) *Trichomma BANCHINAE Alloplasta piceator Alloplasta superba criddlei Arenetra fumipennis *Arenetra hirsutula Arenetra nigrita Arenetra rufipes Cresson, 1870 *Arenetra 1978 Banchus apenes Townes, *Banchus canadensis Cresson, 1868 *Banchus ciliatus *Parania geniculata *Parania pulchra Dasch, 1984 Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 329 Leafrollers Cutworms Cutworms Concealed feeders Lepidoptera: Erebidae, Geometridae, Lasiocampidae, Noctuidae, Notodontidae, Sphingidae (33) Host Family (No. spp.) Guild/Notes Lepidoptera: Erebidae, Noctuidae (10) Setose Hymenoptera: Pamphiliidae; Lepidoptera: Noctuidae (2) Lepidoptera: Elachistidae, Erebidae, Geometridae, Lasiocampidae, Noctuidae, Nolidae, Notodontidae, (26) Tortricidae Pyralidae, Saturniidae, Mostly leafrollers Lepidoptera: Tortricidae (1) Tortricidae Lepidoptera: None Lepidoptera: Noctuidae (3) (18) Tortricidae Lepidoptera: Geometridae, Pyralidae, Mostly leafrollers Lepidoptera: Geometridae (1) Lepidoptera: Saturniidae, Sphingidae (6) most spp social Large, Lepidoptera: Galacticidae, Gelechiidae, Olethreutidae, (7) Tortricidae Pyralidae, (1) Tortricidae Lepidoptera: Lepidoptera: Erebidae, Geometridae, Noctuidae, Notodontidae (16) None Lepidoptera: Erebidae, Geometridae, Noctuidae, Notodontidae, Nymphalidae, Sphingidae (15) Mostly setose Lepidoptera: Erebidae, Geometridae, Noctuidae (10) Mostly setose (6) Tortricidae Lepidoptera: Pyralidae, Concealed feeders Hymenoptera: Chrysididae, Diapriidae, Lepidoptera: Erebidae, Vespidae; Ichneumonidae, Noctuidae (10) Lepidoptera: Noctuidae (2) None Lepidoptera: Noctuidae (1) Cutworms None None Lepidoptera: Noctuidae (3) None None None Hymenoptera: Cynipidae; Lepidoptera: Choreutidae, Crambidae, Elachistidae, Erebidae, Galacticidae, Nolidae, Gelechiidae, Geometridae, Lycaenidae, (40) Tortricidae Psychidae, Pyralidae, Sesiidae,

, Brandon, Carberry, Horton, International Peace Horton, International Peace Birtle , Brandon, Carberry, 1 and 5 mi SW Shilo, Teulon, Winnipeg Teulon, Shilo, 1 and 5 mi SW b Wainwright; SK: Attons L, Battle River, Bronson PF, Caron, Caron, Bronson PF, Attons L, Battle River, SK: Wainwright;

Cypress Hills PP, Prelate, Prince Albert, Radisson, Rutland; MB: Prelate, Prince Cypress Hills PP, Winnipeg Teulon, Ninette, Victor St. SK; Oxbow, Whyte, Sifton Ft. Clearwater, Aweme, MB: Altona , Localities Waterton; Tofield, L, Red Deer, Consort, Edmonton, George AB: Bilby, Regina, Saskatoon; MB: Birds Hill, at Cut Knife, Elbow, Attons L SK: Winnipeg Teulon, Ninette, Husavik, Killarney, Carberry, AB: Calgary Derwent, Edmonton, Lethbridge; SK: MacDowell, AB: Calgary, Winnipeg Ninette, 10 mi S Aweme, Saskatoon, Swift Current; MB: AB: 18 mi E Lacombe, Red Deer AB: Rimbey Thornhill MB: Valley, Pembina NA AB: Wagner Blackfoot Hills, Chedderville, Clymont, Aspen Beach, Bilby, AB: Edgerton, Edmonton, Parkland, Rockyford, Consort, Drumheller, Stavely, AB: Medicine Hat; Midcentral Canada (AB/SK) in sunflower fields Irvine, Lethbridge, Onefour, Aden, Edmonton, Elk Island P, AB: Treesbank Stockton, W 2 mi Aweme, Scandia; SK: Elbow; MB: Aspen Beach, Clymont, Cypress Hills, Edmonton, Elkwater, AB: NA; Wagner L, Golden Spike, Manyberries, 2 mi E Red Deer, George Mt; Wood Saltcoats, Saskatoon, Victor, St. Attons L, Neilburg, SK: AB: Cypress Hills, Green Court, Lethbridge, Red Deer; SK: Shilo MB: 5 mi SW Teulon Fort Garry, Aweme, MB: Wabamun; AB: Edmonton, Gull L, Riverton , St. Claude Aweme, NA; MB: Wagner AB: SK: Saskatoon, south in sunflower fields Waterton; AB: Bilby, AB: Edmonton Twin Butte; Tofield, Waterton, AB: Edmonton, Manyberries, Lethbridge, AB: Beauvallon, Blackfoot Hills, Elk Point, Hardisty, SK: Harris, Lebret, Rutland, Saskatoon; Wainwright; Medicine Hat, AB: Cochrane, Edmonton, Lethbridge, Medicine Hat; SK: Saskatoon MB: Aweme AB: Lethbridge SK: Saskatoon AB: Edmonton, Magrath; SK: Saskatoon Asquith, Clarkboro, AB: Edmonton, Lethbridge, Medicine Hat; SK: Saskatoon, White Fox AB: Medicine Hat NA Wagner AB: Edmonton, Lethbridge, Medicine Hat, AB: Lethbridge; SK: Saskatoon MB: Aweme, MB: Aweme, Mt FR, Millwood, 5 mi N Minnedosa, Ninette, Turtle Gardens at 3 mi S Shilo, sunflower fields Cut Knife, Donavon, Dunblane, Dundurn, Elbow, Ft a la Corne, Great Ft a la Corne, Great Cut Knife, Donavon, Dunblane, Dundurn, Elbow, Pike L, Great Sand Hills, Indian Head, Keppel, Lloydminster, Deer, White Fox; MB: Onah, Saskatoon, Snowden, Sutherland, Redberry, Midcentral Canada (AB/SK) in Winnipeg; Stockton, W 2 mi 4,8,N 4,N 4,5,7,8,H 5,11,N 4,N 4,N,T 4,N 5,9,12,N 5,11,H 5,11,13,N 2,5,11,N 2,N 11,N 2,4,N,T 4,N

4,N 4,N 11,N 5,11,N 2,4,N,T 11,N 2,4,9,10,N,T 11,N 11,14,N 4,8,N 11,N 4,N (Cresson, 1879) (Rohwer, 1915) (Rohwer, Cresson, 1868 Townes, 1978 Townes, (Cresson, 1870) (Holmgren, 1857) (Thunberg, 1824) (Thunberg, (Norton, 1863) (Cushman, 1922) (Provancher, 1874) (Provancher, Walley, 1931 Walley, Townes, 1978 Townes, Townes, 1978 Townes, Cresson, 1870 a 2,4,5,9,H,O Erigorgus stenotus Dasch, 1984 Erigorgus utahensis Dasch, 1984 Erigorgus Therion magnum Dasch, 1984 Apophua simplicipes canadensis *Arenetra SUBFAMILY Species Therion fuscipenne Erigorgus xanthopsis (Ashmead, 1890) Erigorgus aclerivorus Habronyx amoenus Dasch, 1984 Habronyx magniceps (Cresson, 1872) Habronyx Ophionellus foutsi Therion californicum (Linnaeus, Therion circumflexum 1758) Therion morio (Fabricius, 1781) 1917 Therion sassacus Viereck, maceratum (Cresson, 1879) *Trichomma BANCHINAE Alloplasta piceator Alloplasta superba criddlei Arenetra fumipennis *Arenetra hirsutula Arenetra nigrita Arenetra rufipes Cresson, 1870 *Arenetra 1978 Banchus apenes Townes, *Banchus canadensis Cresson, 1868 *Banchus ciliatus *Parania geniculata *Parania pulchra Dasch, 1984 330 M. D. Schwarzfeld Leafrollers Leafrollers Leafrollers Leafrollers Cutworms Cutworms Lepidoptera: Noctuidae (1) Host Family (No. spp.) Guild/Notes Lepidoptera: Noctuidae (5) Lepidoptera: Erebidae, Noctuidae (3) None None None Lepidoptera: Geometridae (1) None None None Lepidoptera: Noctuidae (1) None None None Hymenoptera: Cimbicidae; Lepidoptera: Erebidae, Noctuidae, Sphingidae (10) None Lepidoptera: Crambidae, Erebidae, Geometridae, Pieridae, Noctuidae, Saturniidae (14) Lepidoptera: Noctuidae (3) None None None Lepidoptera: Noctuidae (1) None (1) Tortricidae Lepidoptera: Tortricidae Hymenoptera: Diprionidae; Lepidoptera: (2) None (5) Tortricidae Lepidoptera: None (1) Tortricidae Lepidoptera: (3) Tortricidae Lepidoptera: Gelechiidae, Stem gall makers (2) Tortricidae Lepidoptera: Lepidoptera: Erebidae, Geometridae, Lasiocampidae, (11) Tortricidae Noctuidae, None None None None

SK: Dafoe, Dodsland, Radisson; MB: Aweme, Teulon Aweme, SK: Dafoe, Dodsland, Radisson; MB:

b Localities Chin, Clyde, Cypress Hills, Calgary, Bilby, AB: Belly River, White Fox AB: Edmonton; SK: AB: St. Paul; NA Wagner AB: Edmonton, Chin, St. Paul, Elkwater L, Wallwort AB: Edmonton; SK: MB: Aweme AB: Edmonton, Jumping Pound Cr Red Jumping Pound Cr, Cochrane, Edmonton, Elkwater, AB: Bilby, Albert; MB: Birch SK: Prince Waterton; NA, Wagner Butte, Twin Deer, Mt PP Turtle Shilo, W 5 mi River, Aweme Edmonton; SK: Dubuc; MB: AB: Calgary, Elkwater L, Red Deer; MB: AB: Cypress Hills, Edmonton, Elkwater, Teulon Rockglen SK: Kennedy, AB: Calgary; SK: Dafoe, Herschel, Melfort, Roche Percee, Saskatoon, Tyner AB: Medicine Hat; SK: Dundurn of Swift Current) SK: Great Sand Hills (W Cooking L, Edmonton, Lethbridge; SK: Earl Grey, Calgary, AB: Bilby, Marie; Val Pickthall, Regina, Rockglen, Swift Current, Unity, MB: Aweme AB: Calgary; SK: Saskatoon Waterton; Wabamun, AB: Edmonton, St. Paul, Aweme AB: Lethbridge; MB: White Fox SK: Gull L, Waterton; AB: Delia, AB: Medicine Hat, Scandia; SK: Bestville (nr Hazlet), Regina, Marie Val Secretan, Saskatchewan Landing, Uren, AB: Lethbridge MB: Aweme SK: Tilley; Chin, Edmonton, Lethbridge, Medicine Hat, AB: Calgary, Aweme Saskatoon, Swift Current; MB: Great Deer, AB: Cypress Hills, Elkwater P Woods Albert; MB: Spruce SK: Prince AB: Lethbridge Albert; MB: Ft. Garry Prince SK: Lloydminster, MB: Winnipeg SK: Saskatoon; MB: 2 mi N of Forrest Stn MB: Ninette MB: Ochre River Cypress NP, L Waterton Caroline, Chief Mt Hwy in AB: Bearberry, 5 mi N Star, Red Deer, Vermilion, 1 mi S Elkwater, Hills, Elkwater P, at gate; SK: Duck L, Frenchman Butte, Glaslyn, Lakes NP Waterton Aweme Saskatoon, Sheho; MB: Grassy L, Grenfell, Hillesden, Neilburg, Butte AB: Twin AB: Edmonton; SK: Snowden; MB: Ninette AB: Edmonton Butte; SK: Twin Cypress Hills, Stettler, AB: 30 mi NE Brooks, Calgary, MB: Carberry, Welby; Estevan, Kincaid, Minton, Rockglen, Saskatoon, Shilo 5 mi SW Edmonton, Elkwater, Elkwater L, Fawcett, Lethbridge, Medicine Hat, Elkwater L, Fawcett, Lethbridge, Medicine Hat, Edmonton, Elkwater, Lakes Waterton Waterton, Tabor, Morrin, Scandia, Sedgewick, St. Paul, Langham, Roche Percee, Earl Grey, Aylsham, SK: Winterburn; NP, Aweme, Arrow River, MB: Young; White Fox, Saskatoon, Stalwart, Birtle, Bowsman, Brandon, Woody, Benito, Beulah, Bield, Big Baldur, Gilbert Plains, Glenlea, Dauphin, Durban, Franklin, Grandview, Lenswood, Minitonas, Isle , Hamiota, Horton, Kenville, Laurier, Grosse Minnedosa, Neepawa, 20 mi N Roblin, Riverton, Russell, Thunder Hill, Ukraina, Amelie, Swan River, Silverton Stn, St. Whitewater Virden, 11,N 11,N 5,11,N 5,11,H,T,O 5,11,N 11,N

5,11,N 11,H,O 5,11,N 11,N 11,N 5,11,N 11,H 11,N,T 11,N,T 2,11,N

11,N 12,N 11,N 11,N 11,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N (Gravenhorst, 12,N 12,N 12,N Cushman, 1937 Cresson, 1865 (Walsh, 1873) (Walsh, (Thomson, 1877) (Walley, 1931) (Walley, Walsh, 1873 Walsh, Cushman, 1937 Provancher, 1874 Provancher, Cresson, 1865 (Fabricius, 1781) Dasch, 1988 Provancher, 1874 Provancher, Schmiedeknecht, Dasch, 1988 Dasch, 1988 Townes, 1978 Townes, Cresson, 1870 Gravenhorst, 1829 Dasch, 1988 Cushman, 1937 Cushman, 1933 Dasch, 1988 Dasch, 1988 Dasch, 1988 Dasch, 1988 Dasch, 1988 a 2,5,11,H 5,11,H Townes, 1978 Townes, Banchus nigroflavus *Banchus flavescens Cresson, 1868 Glypta albifaciens Cresson, 1870 Glypta borealis Glypta caulicola Glypta conflictanae SUBFAMILY Species *Exetastes illinoiensis Banchus hastator (Fabricius, 1793) 1874 *Banchus inermis Provancher, Banchus pallescens Banchus superbus Cresson, 1865 Cryptopimpla genalis Cryptopimpla quadrilineata 1829) Exetastes abdominalis *Exetastes albitarsis *Exetastes angustoralis Cushman, 1937 Exetastes brevicornis Exetastes caliginosus *Exetastes flavus Exetastes fornicator *Exetastes illusor *Exetastes obscurus Cresson, 1865 Exetastes pectinatus *Exetastes rempeli Exetastes scutellaris Exetastes suaveolens *Exetastes syriacus 1910 Glypta alberta Glypta albilineata Glypta albonotata Glypta canadensis Glypta cesta Glypta confusa Glypta convexa Glypta crassa Dasch, 1988 Glypta curta 5,9,11,15,16,17,18,19,N Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 331 Leafrollers Leafrollers Leafrollers Leafrollers Cutworms Cutworms Lepidoptera: Noctuidae (1) Host Family (No. spp.) Guild/Notes Lepidoptera: Noctuidae (5) Lepidoptera: Erebidae, Noctuidae (3) None None None Lepidoptera: Geometridae (1) None None None Lepidoptera: Noctuidae (1) None None None Hymenoptera: Cimbicidae; Lepidoptera: Erebidae, Noctuidae, Sphingidae (10) None Lepidoptera: Crambidae, Erebidae, Geometridae, Pieridae, Noctuidae, Saturniidae (14) Lepidoptera: Noctuidae (3) None None None Lepidoptera: Noctuidae (1) None (1) Tortricidae Lepidoptera: Tortricidae Hymenoptera: Diprionidae; Lepidoptera: (2) None (5) Tortricidae Lepidoptera: None (1) Tortricidae Lepidoptera: (3) Tortricidae Lepidoptera: Gelechiidae, Stem gall makers (2) Tortricidae Lepidoptera: Lepidoptera: Erebidae, Geometridae, Lasiocampidae, (11) Tortricidae Noctuidae, None None None None

SK: Dafoe, Dodsland, Radisson; MB: Aweme, Teulon Aweme, SK: Dafoe, Dodsland, Radisson; MB: b Localities Chin, Clyde, Cypress Hills, Calgary, Bilby, AB: Belly River, White Fox AB: Edmonton; SK: AB: St. Paul; NA Wagner AB: Edmonton, Chin, St. Paul, Elkwater L, Wallwort AB: Edmonton; SK: MB: Aweme AB: Edmonton, Jumping Pound Cr Red Jumping Pound Cr, Cochrane, Edmonton, Elkwater, AB: Bilby, Albert; MB: Birch SK: Prince Waterton; NA, Wagner Butte, Twin Deer, Mt PP Turtle Shilo, W 5 mi River, Aweme Edmonton; SK: Dubuc; MB: AB: Calgary, Elkwater L, Red Deer; MB: AB: Cypress Hills, Edmonton, Elkwater, Teulon Rockglen SK: Kennedy, AB: Calgary; SK: Dafoe, Herschel, Melfort, Roche Percee, Saskatoon, Tyner AB: Medicine Hat; SK: Dundurn of Swift Current) SK: Great Sand Hills (W Cooking L, Edmonton, Lethbridge; SK: Earl Grey, Calgary, AB: Bilby, Marie; Val Pickthall, Regina, Rockglen, Swift Current, Unity, MB: Aweme AB: Calgary; SK: Saskatoon Waterton; Wabamun, AB: Edmonton, St. Paul, Aweme AB: Lethbridge; MB: White Fox SK: Gull L, Waterton; AB: Delia, AB: Medicine Hat, Scandia; SK: Bestville (nr Hazlet), Regina, Marie Val Secretan, Saskatchewan Landing, Uren, AB: Lethbridge MB: Aweme SK: Tilley; Chin, Edmonton, Lethbridge, Medicine Hat, AB: Calgary, Aweme Saskatoon, Swift Current; MB: Great Deer, AB: Cypress Hills, Elkwater P Woods Albert; MB: Spruce SK: Prince AB: Lethbridge Albert; MB: Ft. Garry Prince SK: Lloydminster, MB: Winnipeg SK: Saskatoon; MB: 2 mi N of Forrest Stn MB: Ninette MB: Ochre River Cypress NP, L Waterton Caroline, Chief Mt Hwy in AB: Bearberry, 5 mi N Star, Red Deer, Vermilion, 1 mi S Elkwater, Hills, Elkwater P, at gate; SK: Duck L, Frenchman Butte, Glaslyn, Lakes NP Waterton Aweme Saskatoon, Sheho; MB: Grassy L, Grenfell, Hillesden, Neilburg, Butte AB: Twin AB: Edmonton; SK: Snowden; MB: Ninette AB: Edmonton Butte; SK: Twin Cypress Hills, Stettler, AB: 30 mi NE Brooks, Calgary, MB: Carberry, Welby; Estevan, Kincaid, Minton, Rockglen, Saskatoon, Shilo 5 mi SW Edmonton, Elkwater, Elkwater L, Fawcett, Lethbridge, Medicine Hat, Elkwater L, Fawcett, Lethbridge, Medicine Hat, Edmonton, Elkwater, Lakes Waterton Waterton, Tabor, Morrin, Scandia, Sedgewick, St. Paul, Langham, Roche Percee, Earl Grey, Aylsham, SK: Winterburn; NP, Aweme, Arrow River, MB: Young; White Fox, Saskatoon, Stalwart, Birtle, Bowsman, Brandon, Woody, Benito, Beulah, Bield, Big Baldur, Gilbert Plains, Glenlea, Dauphin, Durban, Franklin, Grandview, Lenswood, Minitonas, Isle , Hamiota, Horton, Kenville, Laurier, Grosse Minnedosa, Neepawa, 20 mi N Roblin, Riverton, Russell, Thunder Hill, Ukraina, Amelie, Swan River, Silverton Stn, St. Whitewater Virden, 11,N 11,N 5,11,N 5,11,H,T,O 5,11,N 11,N

5,11,N 11,H,O 5,11,N 11,N 11,N 5,11,N 11,H 11,N,T 11,N,T 2,11,N

11,N 12,N 11,N 11,N 11,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N (Gravenhorst, 12,N 12,N 12,N Cushman, 1937 Cresson, 1865 (Walsh, 1873) (Walsh, (Thomson, 1877) (Walley, 1931) (Walley, Walsh, 1873 Walsh, Cushman, 1937 Provancher, 1874 Provancher, Cresson, 1865 (Fabricius, 1781) Dasch, 1988 Provancher, 1874 Provancher, Schmiedeknecht, Dasch, 1988 Dasch, 1988 Townes, 1978 Townes, Cresson, 1870 Gravenhorst, 1829 Dasch, 1988 Cushman, 1937 Cushman, 1933 Dasch, 1988 Dasch, 1988 Dasch, 1988 Dasch, 1988 Dasch, 1988 a 2,5,11,H 5,11,H Townes, 1978 Townes, Banchus nigroflavus *Banchus flavescens Cresson, 1868 Glypta albifaciens Cresson, 1870 Glypta borealis Glypta caulicola Glypta conflictanae SUBFAMILY Species *Exetastes illinoiensis Banchus hastator (Fabricius, 1793) 1874 *Banchus inermis Provancher, Banchus pallescens Banchus superbus Cresson, 1865 Cryptopimpla genalis Cryptopimpla quadrilineata 1829) Exetastes abdominalis *Exetastes albitarsis *Exetastes angustoralis Cushman, 1937 Exetastes brevicornis Exetastes caliginosus *Exetastes flavus Exetastes fornicator *Exetastes illusor *Exetastes obscurus Cresson, 1865 Exetastes pectinatus *Exetastes rempeli Exetastes scutellaris Exetastes suaveolens *Exetastes syriacus 1910 Glypta alberta Glypta albilineata Glypta albonotata Glypta canadensis Glypta cesta Glypta confusa Glypta convexa Glypta crassa Dasch, 1988 Glypta curta 5,9,11,15,16,17,18,19,N 332 M. D. Schwarzfeld Mostly Concealed feeders Leafrollers Leafrollers Concealed feeders Leafrollers Concealed feeders Leafrollers Leafrollers Leafrollers Concealed feeders Concealed feeders Climbing cutworm Host Family (No. spp.) Guild/Notes None None None None (1) Tortricidae Lepidoptera: None Lepidoptera: Geometridae, Lasiocampidae, (28) Tortricidae Nymphalidae, Saturniidae, leafrollers (1) Tortricidae Lepidoptera: (14) Tortricidae Lepidoptera: Erebidae, Pyralidae, Mostly concealed conifer feeders None None None Lepidoptera: Noctuidae (1) None None None (1) Tortricidae Lepidoptera: None None None (3) Tortricidae Hymenoptera: Cephidae; Lepidoptera: Concealed feeders None None (1) Tortricidae Lepidoptera: (1) Tortricidae Lepidoptera: None (2) Tortricidae Lepidoptera: Lepidoptera: Crambidae, Gelechiidae, Noctuidae, (27) Tortricidae Mostly leafrollers (12) Tortricidae Lepidoptera: Momphidae, Concealed feeders Coleoptera: Cerambycidae (1) None (1) Tortricidae Lepidoptera: None (1) Tortricidae Lepidoptera: (5) Tortricidae Lepidoptera: (8) Tortricidae Lepidoptera: Gelechiidae, Leafrollers (3) Tortricidae Lepidoptera: (4) Tortricidae Lepidoptera: ,

b SW Shilo SW Localities Elkwater AB: Calgary, Waterton Butte, Twin View, Mountain Elkwater, AB: Cowley, L AB: George SK: Scott AB: Edmonton W Fawcett, Jumping Pound Cr at 20 mi AB: Edmonton, Elkwater P, SK: Wabamun; St. Paul, Sylvan L, Lethbridge, Red Deer, Calgary, Aweme, Saskatoon; MB: Lake Max, Lanigan, Lloydminster, Great Deer, Shilo, Gimli, Horton, Ninette, Pilot Mound, 30 mi N Roblin, 5 SW Virden Teulon, Swan River, Spruce Woods, Transcona SK: Kenosee; MB: Glenlea AB: Cypress Hills, Elkwater; SK: Glaslyn; MB: Beausejour, FR Spruce Woods AB: Clymont, Edmonton MB: 30 mi N Roblin Mt SK: Wood Elkwater L AB: Edmonton, Elkwater, SK: Tompkins MB: Carberry White Fox; MB: SK: Wainwright; Edmonton, Stettler, AB: Calmar, Dauphin, 2 mi E Douglas Stn, 9 N Forrest Gladstone, Ninette, 5 Shilo, Souris mi SW L; SK: Kenosee P AB: Edmonton, George Stockton W MB: 2 mi MB: Greenridge SK: Assiniboia AB: Edmonton, Elkwater L AB: Mountain View MB: Transcona Minton, Estevan, Kandahar, SK: Bienfait, Briercrest, Drinkwater, Altona Bunch; MB: Willow Torquay, Outram, Rockglen, L; SK: Kenosee AB: George Mt FR, Turtle in Elkwater L; MB: Max L AB: Edmonton, Elkwater, Transcona MB: Transcona Rainier, Lethbridge, Medicine Hat, Onefour, AB: Edmonton, Elkwater, Valley, Rose Lloydminster, Kisbey, Butte; SK: Dafoe, Elbow, Twin Snowden; MB: Boissevain, Horton, 5 mi N Minnedosa, Ninette, Spruce FR, Transcona Woods Elkwater Pk, Lethbridge, AB: "Blackfoot Coulee", Calgary, Kelso, SK: Kandahar, Wainwright; Vermilion, Manyberries, Scandia, Lumsden, Minton, Regina, Roche Percee, Rockglen, Saskatchewan Bunch; MB: Horton, Willow Marie, Val Landing, Saskatoon, Snowden, FR Spruce Woods Hays, Medicine Drumheller, AB: Lethbridge, Blackfoot Hills, Calgary, Lisieux, Regina, Gascoigne, Kennedy, Hat, Scandia; SK: Elbow, 5 mi Aweme, Bunch; MB: Willow Saskatoon, Victor, Rockglen, St. SK: Albert Prince SK: Saskatoon AB:Edmonton; SK: Lloydminster AB: Edmonton SK: Saskatoon Albert AB: Edmonton; SK: Prince Wasagaming MB: Camper, Bunch; MB: Willow Albert, SK: Prince Wainwright; AB: Edmonton, Dauphin Orion AB: Edmonton, Elkwater P, 5,12,H,T 9,12,20,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 5,12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 5,12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N Dasch, 1988 Cresson, 1870 Dasch, 1988 Cresson, 1870 (Viereck, 1912) (Viereck, Dasch, 1988 Ashmead, 1902 Dasch, 1988 Cresson, 1870 Dasch, 1988 Dasch, 1988 Dasch, 1988 Dasch, 1988 Dasch, 1988 Provancher, 1890 Provancher, (Provancher, 1883) (Provancher, Dasch, 1988 Cresson, 1870 Dasch, 1988 Cushman, 1919 Cresson, 1865 Dasch, 1988 Dalla Torre, 1901 Torre, Dalla Dasch, 1988 Dasch, 1988 Dasch, 1988 Dasch, 1988 Dasch, 1988 Dasch, 1988 a Glypta davisii Glypta elevata Glypta fumiferanae Glypta manitobae Glypta pulchripes Dasch, 1988 Glypta robusta Glypta rufiscutellaris Dasch, 1988 Glypta saperdae Glypta solida Glypta tortricis Glypta tricincta Glypta varipes SUBFAMILY Species Glypta occidentalis Glypta diminuta Glypta divisa Glypta dorsiatomanae Dasch, 1988 Glypta erratica Glypta flavomaculata Glypta fumosa Dasch, 1988 Glypta fuscata Glypta inusitata Glypta kukakensis Glypta linearis Glypta macilenta Glypta macra Cresson, 1870 Glypta maculata Glypta martini Glypta montana Dasch, 1988 Glypta mutica Glypta parallela Dasch, 1988 Glypta prognatha Glypta rubripes Cresson, 1870 Glypta rufofasciata Glypta rufula Glypta saskatchewan Glypta severa Glypta synnomae Dasch, 1988 Glypta truncata Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 333 Mostly Concealed feeders Leafrollers Leafrollers Concealed feeders Leafrollers Concealed feeders Leafrollers Leafrollers Leafrollers Concealed feeders Concealed feeders Climbing cutworm Host Family (No. spp.) Guild/Notes None None None None (1) Tortricidae Lepidoptera: None Lepidoptera: Geometridae, Lasiocampidae, (28) Tortricidae Nymphalidae, Saturniidae, leafrollers (1) Tortricidae Lepidoptera: (14) Tortricidae Lepidoptera: Erebidae, Pyralidae, Mostly concealed conifer feeders None None None Lepidoptera: Noctuidae (1) None None None (1) Tortricidae Lepidoptera: None None None (3) Tortricidae Hymenoptera: Cephidae; Lepidoptera: Concealed feeders None None (1) Tortricidae Lepidoptera: (1) Tortricidae Lepidoptera: None (2) Tortricidae Lepidoptera: Lepidoptera: Crambidae, Gelechiidae, Noctuidae, (27) Tortricidae Mostly leafrollers (12) Tortricidae Lepidoptera: Momphidae, Concealed feeders Coleoptera: Cerambycidae (1) None (1) Tortricidae Lepidoptera: None (1) Tortricidae Lepidoptera: (5) Tortricidae Lepidoptera: (8) Tortricidae Lepidoptera: Gelechiidae, Leafrollers (3) Tortricidae Lepidoptera: (4) Tortricidae Lepidoptera: , b SW Shilo SW Localities Elkwater AB: Calgary, Waterton Butte, Twin View, Mountain Elkwater, AB: Cowley, L AB: George SK: Scott AB: Edmonton W Fawcett, Jumping Pound Cr at 20 mi AB: Edmonton, Elkwater P, SK: Wabamun; St. Paul, Sylvan L, Lethbridge, Red Deer, Calgary, Aweme, Saskatoon; MB: Lake Max, Lanigan, Lloydminster, Great Deer, Shilo, Gimli, Horton, Ninette, Pilot Mound, 30 mi N Roblin, 5 SW Virden Teulon, Swan River, Spruce Woods, Transcona SK: Kenosee; MB: Glenlea AB: Cypress Hills, Elkwater; SK: Glaslyn; MB: Beausejour, FR Spruce Woods AB: Clymont, Edmonton MB: 30 mi N Roblin Mt SK: Wood Elkwater L AB: Edmonton, Elkwater, SK: Tompkins MB: Carberry White Fox; MB: SK: Wainwright; Edmonton, Stettler, AB: Calmar, Dauphin, 2 mi E Douglas Stn, 9 N Forrest Gladstone, Ninette, 5 Shilo, Souris mi SW L; SK: Kenosee P AB: Edmonton, George Stockton W MB: 2 mi MB: Greenridge SK: Assiniboia AB: Edmonton, Elkwater L AB: Mountain View MB: Transcona Minton, Estevan, Kandahar, SK: Bienfait, Briercrest, Drinkwater, Altona Bunch; MB: Willow Torquay, Outram, Rockglen, L; SK: Kenosee AB: George Mt FR, Turtle in Elkwater L; MB: Max L AB: Edmonton, Elkwater, Transcona MB: Transcona Rainier, Lethbridge, Medicine Hat, Onefour, AB: Edmonton, Elkwater, Valley, Rose Lloydminster, Kisbey, Butte; SK: Dafoe, Elbow, Twin Snowden; MB: Boissevain, Horton, 5 mi N Minnedosa, Ninette, Spruce FR, Transcona Woods Elkwater Pk, Lethbridge, AB: "Blackfoot Coulee", Calgary, Kelso, SK: Kandahar, Wainwright; Vermilion, Manyberries, Scandia, Lumsden, Minton, Regina, Roche Percee, Rockglen, Saskatchewan Bunch; MB: Horton, Willow Marie, Val Landing, Saskatoon, Snowden, FR Spruce Woods Hays, Medicine Drumheller, AB: Lethbridge, Blackfoot Hills, Calgary, Lisieux, Regina, Gascoigne, Kennedy, Hat, Scandia; SK: Elbow, 5 mi Aweme, Bunch; MB: Willow Saskatoon, Victor, Rockglen, St. SK: Albert Prince SK: Saskatoon AB:Edmonton; SK: Lloydminster AB: Edmonton SK: Saskatoon Albert AB: Edmonton; SK: Prince Wasagaming MB: Camper, Bunch; MB: Willow Albert, SK: Prince Wainwright; AB: Edmonton, Dauphin Orion AB: Edmonton, Elkwater P, 5,12,H,T 9,12,20,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 5,12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 5,12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N 12,N Dasch, 1988 Cresson, 1870 Dasch, 1988 Cresson, 1870 (Viereck, 1912) (Viereck, Dasch, 1988 Ashmead, 1902 Dasch, 1988 Cresson, 1870 Dasch, 1988 Dasch, 1988 Dasch, 1988 Dasch, 1988 Dasch, 1988 Provancher, 1890 Provancher, (Provancher, 1883) (Provancher, Dasch, 1988 Cresson, 1870 Dasch, 1988 Cushman, 1919 Cresson, 1865 Dasch, 1988 Dalla Torre, 1901 Torre, Dalla Dasch, 1988 Dasch, 1988 Dasch, 1988 Dasch, 1988 Dasch, 1988 Dasch, 1988 a Glypta davisii Glypta elevata Glypta fumiferanae Glypta manitobae Glypta pulchripes Dasch, 1988 Glypta robusta Glypta rufiscutellaris Dasch, 1988 Glypta saperdae Glypta solida Glypta tortricis Glypta tricincta Glypta varipes SUBFAMILY Species Glypta occidentalis Glypta diminuta Glypta divisa Glypta dorsiatomanae Dasch, 1988 Glypta erratica Glypta flavomaculata Glypta fumosa Dasch, 1988 Glypta fuscata Glypta inusitata Glypta kukakensis Glypta linearis Glypta macilenta Glypta macra Cresson, 1870 Glypta maculata Glypta martini Glypta montana Dasch, 1988 Glypta mutica Glypta parallela Dasch, 1988 Glypta prognatha Glypta rubripes Cresson, 1870 Glypta rufofasciata Glypta rufula Glypta saskatchewan Glypta severa Glypta synnomae Dasch, 1988 Glypta truncata 334 M. D. Schwarzfeld Stem borer Concealed feeders Host Family (No. spp.) Guild/Notes Argyresthiidae, Lepidoptera: Blastobasidae, Coleophoridae, Gelechiidae, Geometridae, Pyralidae, (28) Tortricidae Sesiidae, Mostly concealed feeders Lepidoptera: Crambidae, Gelechiidae, Tortricidae (14) Tortricidae Lepidoptera: Crambidae, Gelechiidae, Concealed feeders Argyresthiidae Lepidoptera: (1) Tineidae, Lepidoptera: Elachistidae, Gelechiidae, Sesiidae (6) Tortricidae, Mostly concealed feeders None Lepidoptera: Noctuidae (4) Stem borers (2) Tortricidae Lepidoptera: Stem borers Lepidoptera: Geometridae, Noctuidae, Notodontidae, (8) Ypsolophidae Sesiidae, None (5) Tortricidae Lepidoptera: Tineidae, Pyralidae, Concealed feeders Lepidoptera: Tenthredinidae; Hymenoptera: Crambidae, Notodontidae, Psychidae, Sesiidae, (11) Tortricidae Mostly concealed feeders None None Lepidoptera: Batrachedridae, Blastobasidae, Erebidae, Gelechiidae, Pyralidae (5) Mostly concealed feeders None None None Lepidoptera: Tenthredinidae; Hymenoptera: Cynipidae, Erebidae, Gelechiidae, Geometridae, Sesiidae, (21) Tortricidae Mostly concealed feeders None None Lepidoptera: Gelechiidae (2) None None None

Winnipeg Rosa b Localities MB: Aweme AB: southern AB White Fox; MB: Carberry SK: NA; SK: Kenosee, Snowden; MB: Wagner AB: Jumping Pound Cr, Mt, Turtle Boissevain, Aweme, SK: Indian Head NA; SK: Glaslyn, Pike L; MB: Wagner AB: Edmonton, Magrath, Altona , Aweme, AB: Edmonton NP, Waterton Rd in Cameron L Aspen Beach, Brooks, Calgary, AB: Edmonton, Elkwater L, Halcourt, Cypress Hills, Drumheller, Castor, Steveville, Lethbridge, Scandia, Stettler, Jumping Pound Cr, High River, Lakes NP; SK: Waterton NA, Wagner Vermilion, , Tofield Taber, Qu'Appelle, Kenosee, Adams, Canora, Dana, Meota, Moosomin, F. White Tunstall, Regina, Saskatoon, Strongfield, Swift Current, Redberry, Dauphin, Pilot Mound, Aweme, Mt; MB: Wood Bunch, Willow Fox, Pipestone, Swan River Edmonton, Drumheller, Aden, Blackfoot Hills, Calgary, AB: Lethbridge, Manyberries, Red Deer, Elkwater L, Jumping Pound Cr, "Blackfoot Coulee"; Wainwright, NA, Wagner Scandia, Stettler, AB: Jumping Pound Cr Lethbridge, Edmonton, Jumping Pound Cr, AB: Blackfoot Hills, Castor, at Attons L SK: Wainwright; NA, Wagner , Tofield Stettler, Red Deer, Cut Knife, Canora, Estevan, Fort Qu'Apelle, Good Spirit L, Kenosee Rutland, Albert, Redberry, Lebret, Lisieux, Moosomin, Prince P, Mt; MB: Brandon, Dauphin, 9 mi N Wood White Fox, Saskatoon, in Forrest, Gladstone, Horton, Husavick, 5 mi N Minnedosa, Max L Shilo, Mt FR, Ninette, Oak L, Pilot Mound, 3 mi S and 5 SW Turtle Winnipeg Whitewater, Whitewater L, 4 mi N Virden, Mt, Turtle nr Cut Knife, Elbow, Atton L AB: Edmonton, Jumping Pound Cr; SK: Ninette, 5 mi Aweme, Scout L; MB: Victor, Kenosee, Rockglen, St. Shilo SW NA AB: Wagner NA; SK: Moosomin; Wagner Lethbridge, AB: Jumping Pound Cr, MB: Brandon AB: Edmonton Shilo NA; MB: 5 mi SW Wagner AB: NA; SK: Canora, Wagner Manyberries, AB: Jumping Pound Cr, Aweme, White Fox; MB: Saskatoon, Good Spirit L, Great Deer, Brandon, Dauphin, Pilot Mound, Red Deer 6 mi NW Birch River, Winnipeg Stockton, W 3 mi S Shilo, 2 River, AB: Edmonton; MB: Ninette NA; SK: Canora Wagner AB: AB: Onefour; SK: Great Sand Hills Nisbet PF; MB: Bald Head Hills AB: Jumping Pound Cr; SK: Elbow, NA AB: Wagner NA; SK: Regina; Wagner Vermilion, AB: Edmonton, Oyen, Mt Turtle SK: Dubuc; MB: Waterton; NA, Wagner AB: SK: Bagley, Birch Hills, Dafoe, Dubuc, Earl Grey, Saskatoon, Birch Hills, Dafoe, Dubuc, Earl Grey, SK: Bagley, Melita, Aweme, Mt; MB: Wood Wiseton, White Fox, Marie, Val Winnipeg 5 mi N Minnedosa, Roblin, at 30 mi N Glenboro, Ninette Mt FR Turtle in MB: Max L 2,9,11,N 21,N 2,11,N 11,H 11,N 2,5,9,11,H 2,5,11,N 2,N 2,N 2,11,N 11,N 2,9,11,14,N 11,N 2,11,N 11,N 11,N 2,11,N 11,N 11,N 11,H 12,N 2,N (Walley, 1961) (Walley, Townes, 1978 Townes, Townes, 1978 Townes, Townes, 1978 Townes, (Gravenhorst, (Cresson, 1870) Townes, 1978 Townes, Cresson, 1870 (Cresson, 1870) Thomson, 1877 a 2,5,9,11,H 2,5,11,N Glypta vulgaris Lissonota accusator (Fabricius, 1793) 1978 Lissonota amphithyris Townes, SUBFAMILY Species *Lissonota coracina (Gmelin, 1790) Helotorus argyresthiae (Ashmead, 1896) Lissonota acrobasidis *Lissonota brunnea Cresson, 1868 Lissonota cephalotes *Lissonota clypeator 1820) *Lissonota coloradensis (Cresson, 1870) 1920 Lissonota conocola Rohwer, 1978 Lissonota cruralis Townes, 1978 Lissonota curticauda Townes, Lissonota dakrumae (Ashmead, 1896) 1978 Lissonota davisi Townes, Lissonota exigua (Cresson, 1870) Lissonota exilis Lissonota folii Lissonota fulvicornis Lissonota infulata 1942) Lissonota jacobi (Walley, 1978 Lissonota jaei Townes, Lissonota laevigata Lissonota leucoscelis Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 335 Stem borer Concealed feeders Host Family (No. spp.) Guild/Notes Argyresthiidae, Lepidoptera: Blastobasidae, Coleophoridae, Gelechiidae, Geometridae, Pyralidae, (28) Tortricidae Sesiidae, Mostly concealed feeders Lepidoptera: Crambidae, Gelechiidae, Tortricidae (14) Tortricidae Lepidoptera: Crambidae, Gelechiidae, Concealed feeders Argyresthiidae Lepidoptera: (1) Tineidae, Lepidoptera: Elachistidae, Gelechiidae, Sesiidae (6) Tortricidae, Mostly concealed feeders None Lepidoptera: Noctuidae (4) Stem borers (2) Tortricidae Lepidoptera: Stem borers Lepidoptera: Geometridae, Noctuidae, Notodontidae, (8) Ypsolophidae Sesiidae, None (5) Tortricidae Lepidoptera: Tineidae, Pyralidae, Concealed feeders Lepidoptera: Tenthredinidae; Hymenoptera: Crambidae, Notodontidae, Psychidae, Sesiidae, (11) Tortricidae Mostly concealed feeders None None Lepidoptera: Batrachedridae, Blastobasidae, Erebidae, Gelechiidae, Pyralidae (5) Mostly concealed feeders None None None Lepidoptera: Tenthredinidae; Hymenoptera: Cynipidae, Erebidae, Gelechiidae, Geometridae, Sesiidae, (21) Tortricidae Mostly concealed feeders None None Lepidoptera: Gelechiidae (2) None None None

Winnipeg Rosa b Localities MB: Aweme AB: southern AB White Fox; MB: Carberry SK: NA; SK: Kenosee, Snowden; MB: Wagner AB: Jumping Pound Cr, Mt, Turtle Boissevain, Aweme, SK: Indian Head NA; SK: Glaslyn, Pike L; MB: Wagner AB: Edmonton, Magrath, Altona , Aweme, AB: Edmonton NP, Waterton Rd in Cameron L Aspen Beach, Brooks, Calgary, AB: Edmonton, Elkwater L, Halcourt, Cypress Hills, Drumheller, Castor, Steveville, Lethbridge, Scandia, Stettler, Jumping Pound Cr, High River, Lakes NP; SK: Waterton NA, Wagner Vermilion, , Tofield Taber, Qu'Appelle, Kenosee, Adams, Canora, Dana, Meota, Moosomin, F. White Tunstall, Regina, Saskatoon, Strongfield, Swift Current, Redberry, Dauphin, Pilot Mound, Aweme, Mt; MB: Wood Bunch, Willow Fox, Pipestone, Swan River Edmonton, Drumheller, Aden, Blackfoot Hills, Calgary, AB: Lethbridge, Manyberries, Red Deer, Elkwater L, Jumping Pound Cr, "Blackfoot Coulee"; Wainwright, NA, Wagner Scandia, Stettler, AB: Jumping Pound Cr Lethbridge, Edmonton, Jumping Pound Cr, AB: Blackfoot Hills, Castor, at Attons L SK: Wainwright; NA, Wagner , Tofield Stettler, Red Deer, Cut Knife, Canora, Estevan, Fort Qu'Apelle, Good Spirit L, Kenosee Rutland, Albert, Redberry, Lebret, Lisieux, Moosomin, Prince P, Mt; MB: Brandon, Dauphin, 9 mi N Wood White Fox, Saskatoon, in Forrest, Gladstone, Horton, Husavick, 5 mi N Minnedosa, Max L Shilo, Mt FR, Ninette, Oak L, Pilot Mound, 3 mi S and 5 SW Turtle Winnipeg Whitewater, Whitewater L, 4 mi N Virden, Mt, Turtle nr Cut Knife, Elbow, Atton L AB: Edmonton, Jumping Pound Cr; SK: Ninette, 5 mi Aweme, Scout L; MB: Victor, Kenosee, Rockglen, St. Shilo SW NA AB: Wagner NA; SK: Moosomin; Wagner Lethbridge, AB: Jumping Pound Cr, MB: Brandon AB: Edmonton Shilo NA; MB: 5 mi SW Wagner AB: NA; SK: Canora, Wagner Manyberries, AB: Jumping Pound Cr, Aweme, White Fox; MB: Saskatoon, Good Spirit L, Great Deer, Brandon, Dauphin, Pilot Mound, Red Deer 6 mi NW Birch River, Winnipeg Stockton, W 3 mi S Shilo, 2 River, AB: Edmonton; MB: Ninette NA; SK: Canora Wagner AB: AB: Onefour; SK: Great Sand Hills Nisbet PF; MB: Bald Head Hills AB: Jumping Pound Cr; SK: Elbow, NA AB: Wagner NA; SK: Regina; Wagner Vermilion, AB: Edmonton, Oyen, Mt Turtle SK: Dubuc; MB: Waterton; NA, Wagner AB: SK: Bagley, Birch Hills, Dafoe, Dubuc, Earl Grey, Saskatoon, Birch Hills, Dafoe, Dubuc, Earl Grey, SK: Bagley, Melita, Aweme, Mt; MB: Wood Wiseton, White Fox, Marie, Val Winnipeg 5 mi N Minnedosa, Roblin, at 30 mi N Glenboro, Ninette Mt FR Turtle in MB: Max L 2,9,11,N 21,N 2,11,N 11,H 11,N 2,5,9,11,H 2,5,11,N 2,N 2,N 2,11,N 11,N 2,9,11,14,N 11,N 2,11,N 11,N 11,N 2,11,N 11,N 11,N 11,H 12,N 2,N (Walley, 1961) (Walley, Townes, 1978 Townes, Townes, 1978 Townes, Townes, 1978 Townes, (Gravenhorst, (Cresson, 1870) Townes, 1978 Townes, Cresson, 1870 (Cresson, 1870) Thomson, 1877 a 2,5,9,11,H 2,5,11,N Glypta vulgaris Lissonota accusator (Fabricius, 1793) 1978 Lissonota amphithyris Townes, SUBFAMILY Species *Lissonota coracina (Gmelin, 1790) Helotorus argyresthiae (Ashmead, 1896) Lissonota acrobasidis *Lissonota brunnea Cresson, 1868 Lissonota cephalotes *Lissonota clypeator 1820) *Lissonota coloradensis (Cresson, 1870) 1920 Lissonota conocola Rohwer, 1978 Lissonota cruralis Townes, 1978 Lissonota curticauda Townes, Lissonota dakrumae (Ashmead, 1896) 1978 Lissonota davisi Townes, Lissonota exigua (Cresson, 1870) Lissonota exilis Lissonota folii Lissonota fulvicornis Lissonota infulata 1942) Lissonota jacobi (Walley, 1978 Lissonota jaei Townes, Lissonota laevigata Lissonota leucoscelis 336 M. D. Schwarzfeld Mostly concealed feeders Concealed feeders Concealed feeders Concealed feeders Concealed feeders Concealed feeders Cutworms Cutworms Concealed conifer feeders Concealed feeders Host Family (No. spp.) Guild/Notes Lepidoptera: Gelechiidae (1) (6) Tortricidae Lepidoptera: None (1) Tortricidae Lepidoptera: None Lepidoptera: Sesiidae (2) Borers None Lepidoptera: Coleophoridae, Gelechiidae, Pyralidae, (7) Tortricidae Concealed feeders None None (1) Tineidae Lepidoptera: None Lepidoptera: Pyralidae (1) Lepidoptera: Crambidae (1) Lepidoptera: Gelechiidae, Sesiidae (2) Concealed feeders Lepidoptera: Gelechiidae (1) None None Lepidoptera: Noctuidae (1) None None None (7) Tortricidae Lepidoptera: Gelechiidae, Pyralidae, Concealed feeders Lepidoptera: Notodontidae, Pyralidae (5) Leafrollers Lepidoptera: Pyralidae (1) None None Coleoptera: Curculionidae (6) Vespidae; Coleoptera: Curculionidae; Hymenoptera: (6) Tischeriidae Lepidoptera: None None Lepidoptera: Tenthredinidae; Hymenoptera: Crambidae, Noctuidae, Nymphalidae, Plutellidae, (20) Tortricidae Lepidoptera: Noctuidae (1) None Lepidoptera: Noctuidae (1) (4) Tortricidae Lepidoptera: Crambidae, Noctuidae, None None Lepidoptera: Crambidae, Gelechiidae, Noctuidae, Pieridae, Sphingidae (25) None None

b MB; Ninette S wan River Localities Shilo MB: 5 mi SW NA Wagner AB: Edmonton, Jumping Pound Cr, NA Wagner AB: Jumping Pound Cr, SK: Saskatoon, Sutherland Marie Val AB: Elkwater L; SK: Carberry Aweme, AB: Elkwater P; SK: Pike L, Saskatoon; MB: Wagner NA Tofield, Edmonton, AB: Calgary, Winnipeg Shilo, W White Fox; MB: 5 mi NA; SK: Canora, Wagner AB: NA AB: Wagner NA; SK: Saskatoon Wagner AB: NA AB: Wagner MB: Transcona Bunch; MB: International Willow Aspen Beach, Edmonton; SK: AB: Shilo, Mt FR, Pilot Mound, 5 mi SW Turtle Peace Gardens in Wasagaming Waterton Red Deer, AB: Edmonton, Elkwater P, SK: Antelope NA AB: Wagner White Fox AB: Jumping Pound Cr; SK: NA Wagner AB: Elkwater, SK: Cypress Hills AB: Edmonton, Jumping Pound Cr Albert; Prince NA; SK: Elbow, Wagner AB: Jumping Pound Cr, Ninette Aweme, Albert; MB: SK: Dundurn, Prince Saskatoon; MB: 2 mi N Forrest, Avonlea, AB: Edmonton; SK: Stockton W MB: 2 mi Lethbridge AB: Calgary, Aweme SK: Rockglen; MB: AB: Bilby Orion, Sterling, Cranford AB: Manyberries, Milk River, AB: Lethbridge AB: Glenwood Whitewater AB: Elk Island; SK: Saskatoon; MB: AB: Edmonton; SK: Saskatoon SK: Wainwright; Edmonton, Lethbridge, AB: Calahoo, Calgary, Rosthern, Saskatoon AB: Edmonton AB: Lethbridge SK: Saskatoon AB: Edmonton; SK: south AB: Waterton Edmonton AB: Lethbridge, Red Deer, SK: Saskatoon; MB: Carberry Teulon AB: Lethbridge; MB: 5,H 11,N 2,N 14,29,N 22,N 5,22,N 2,11,N,T 11,N 22,N 5,N,T,O 2,11,N 2,11,N 11,N 2,11,N 11,N 5,N 22,N 2,N 2,11,N 11,N 11,N 2,11,N 22,N 28,N 2,11,N 11,N 2,11,N 2,N 11,N 11,N

11,N 11,N,T 2,11,N 11,N 11,N 25,N Aubert, 1977 (Thomson, 1887) (Weed, 1888) (Weed, (Viereck, (Viereck, (Viereck, 1925) (Viereck, Townes, 1978 Townes, (Viereck, 1925) (Viereck, (Cresson, 1870) Townes, 1978 Townes, Cushman, 1926 (Gravenhorst, 1829) (Ashmead, 1890) (Viereck, 1925) (Viereck, (Viereck, 1925) (Viereck, 1875) (Provancher, (Cresson, 1870) Walley, 1934 Walley, (Viereck, 1925) bryanti (Viereck, a 5,22,N,T 5,6,22,27,N Townes, 1978 Lissonota lirata Townes, 1978 Townes, Lissonota nigromacra *Lissonota scutellaris (Cresson, 1870) Syzeuctus elegans SUBFAMILY Species 2,5,11,N Meniscomorpha mirabilis (Cresson, 1870) 23,24,H,O 26,N (Provancher, 1873) Lissonota nigricornis (Provancher, Lissonota occidentalis 1978 Townes, Lissonota orophila Lissonota parva (Cresson, 1870) Lissonota pinguicula Lissonota punctata (Cresson, 1870) Lissonota punctiventrator 1978 Lissonota rasilis Townes, Lissonota reniculellae *Lissonota rubrica (Cresson, 1870) Lissonota sexcincta 1978 Lissonota stenostoma Townes, 1978 Lissonota subcalva Townes, *Lissonota tegularis (Cresson, 1870) 1978 Lissonota tetrazona Townes, 1978 Lissonota uncata Townes, 1978 Lissonota vidua Townes, 1978 Lissonota xanthophrys Townes, Sphelodon phoxopteridis Syzeuctus epischniae Syzeuctus eximius CAMPOPLEGINAE Bathyplectes *Bathyplectes curculionis Bathyplectes exiguus 1969 Callidora surata Tigner, 1879) Campoctonus carinatus (Provancher, (Cresson, Campoletis argentifrons 1864) *Campoletis atkinsoni 1925) Campoletis atypica 1903) Campoletis australis (Viereck, Campoletis clavata 1882) Campoletis distincta (Provancher, Campoletis diversa Campoletis flavicincta (Ashmead, 1890) Campoletis imperfecta Campoletis intermedia Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 337 Mostly concealed feeders Concealed feeders Concealed feeders Concealed feeders Concealed feeders Concealed feeders Cutworms Cutworms Concealed conifer feeders Concealed feeders Host Family (No. spp.) Guild/Notes Lepidoptera: Gelechiidae (1) (6) Tortricidae Lepidoptera: None (1) Tortricidae Lepidoptera: None Lepidoptera: Sesiidae (2) Borers None Lepidoptera: Coleophoridae, Gelechiidae, Pyralidae, (7) Tortricidae Concealed feeders None None (1) Tineidae Lepidoptera: None Lepidoptera: Pyralidae (1) Lepidoptera: Crambidae (1) Lepidoptera: Gelechiidae, Sesiidae (2) Concealed feeders Lepidoptera: Gelechiidae (1) None None Lepidoptera: Noctuidae (1) None None None (7) Tortricidae Lepidoptera: Gelechiidae, Pyralidae, Concealed feeders Lepidoptera: Notodontidae, Pyralidae (5) Leafrollers Lepidoptera: Pyralidae (1) None None Coleoptera: Curculionidae (6) Vespidae; Coleoptera: Curculionidae; Hymenoptera: (6) Tischeriidae Lepidoptera: None None Lepidoptera: Tenthredinidae; Hymenoptera: Crambidae, Noctuidae, Nymphalidae, Plutellidae, (20) Tortricidae Lepidoptera: Noctuidae (1) None Lepidoptera: Noctuidae (1) (4) Tortricidae Lepidoptera: Crambidae, Noctuidae, None None Lepidoptera: Crambidae, Gelechiidae, Noctuidae, Pieridae, Sphingidae (25) None None

b MB; Ninette S wan River Localities Shilo MB: 5 mi SW NA Wagner AB: Edmonton, Jumping Pound Cr, NA Wagner AB: Jumping Pound Cr, SK: Saskatoon, Sutherland Marie Val AB: Elkwater L; SK: Carberry Aweme, AB: Elkwater P; SK: Pike L, Saskatoon; MB: Wagner NA Tofield, Edmonton, AB: Calgary, Winnipeg Shilo, W White Fox; MB: 5 mi NA; SK: Canora, Wagner AB: NA AB: Wagner NA; SK: Saskatoon Wagner AB: NA AB: Wagner MB: Transcona Bunch; MB: International Willow Aspen Beach, Edmonton; SK: AB: Shilo, Mt FR, Pilot Mound, 5 mi SW Turtle Peace Gardens in Wasagaming Waterton Red Deer, AB: Edmonton, Elkwater P, SK: Antelope NA AB: Wagner White Fox AB: Jumping Pound Cr; SK: NA Wagner AB: Elkwater, SK: Cypress Hills AB: Edmonton, Jumping Pound Cr Albert; Prince NA; SK: Elbow, Wagner AB: Jumping Pound Cr, Ninette Aweme, Albert; MB: SK: Dundurn, Prince Saskatoon; MB: 2 mi N Forrest, Avonlea, AB: Edmonton; SK: Stockton W MB: 2 mi Lethbridge AB: Calgary, Aweme SK: Rockglen; MB: AB: Bilby Orion, Sterling, Cranford AB: Manyberries, Milk River, AB: Lethbridge AB: Glenwood Whitewater AB: Elk Island; SK: Saskatoon; MB: AB: Edmonton; SK: Saskatoon SK: Wainwright; Edmonton, Lethbridge, AB: Calahoo, Calgary, Rosthern, Saskatoon AB: Edmonton AB: Lethbridge SK: Saskatoon AB: Edmonton; SK: south AB: Waterton Edmonton AB: Lethbridge, Red Deer, SK: Saskatoon; MB: Carberry Teulon AB: Lethbridge; MB: 5,H 11,N 2,N 14,29,N 22,N 5,22,N 2,11,N,T 11,N 22,N 5,N,T,O 2,11,N 2,11,N 11,N 2,11,N 11,N 5,N 22,N 2,N 2,11,N 11,N 11,N 2,11,N 22,N 28,N 2,11,N 11,N 2,11,N 2,N 11,N 11,N

11,N 11,N,T 2,11,N 11,N 11,N 25,N Aubert, 1977 (Thomson, 1887) (Weed, 1888) (Weed, (Viereck, (Viereck, (Viereck, 1925) (Viereck, Townes, 1978 Townes, (Viereck, 1925) (Viereck, (Cresson, 1870) Townes, 1978 Townes, Cushman, 1926 (Gravenhorst, 1829) (Ashmead, 1890) (Viereck, 1925) (Viereck, (Viereck, 1925) (Viereck, 1875) (Provancher, (Cresson, 1870) Walley, 1934 Walley, (Viereck, 1925) bryanti (Viereck, a 5,22,N,T 5,6,22,27,N Townes, 1978 Lissonota lirata Townes, 1978 Townes, Lissonota nigromacra *Lissonota scutellaris (Cresson, 1870) Syzeuctus elegans SUBFAMILY Species 2,5,11,N Meniscomorpha mirabilis (Cresson, 1870) 23,24,H,O 26,N (Provancher, 1873) Lissonota nigricornis (Provancher, Lissonota occidentalis 1978 Townes, Lissonota orophila Lissonota parva (Cresson, 1870) Lissonota pinguicula Lissonota punctata (Cresson, 1870) Lissonota punctiventrator 1978 Lissonota rasilis Townes, Lissonota reniculellae *Lissonota rubrica (Cresson, 1870) Lissonota sexcincta 1978 Lissonota stenostoma Townes, 1978 Lissonota subcalva Townes, *Lissonota tegularis (Cresson, 1870) 1978 Lissonota tetrazona Townes, 1978 Lissonota uncata Townes, 1978 Lissonota vidua Townes, 1978 Lissonota xanthophrys Townes, Sphelodon phoxopteridis Syzeuctus epischniae Syzeuctus eximius CAMPOPLEGINAE Bathyplectes *Bathyplectes curculionis Bathyplectes exiguus 1969 Callidora surata Tigner, 1879) Campoctonus carinatus (Provancher, (Cresson, Campoletis argentifrons 1864) *Campoletis atkinsoni 1925) Campoletis atypica 1903) Campoletis australis (Viereck, Campoletis clavata 1882) Campoletis distincta (Provancher, Campoletis diversa Campoletis flavicincta (Ashmead, 1890) Campoletis imperfecta Campoletis intermedia 338 M. D. Schwarzfeld Mostly setose Loopers Loopers Loopers Loopers Concealed feeders Concealed feeders Concealed feeders Concealed feeders Concealed feeders Cutworms Concealed feeders Concealed feeders Lepidoptera: Erebidae, Geometridae, Lasiocampidae, Notodontidae, Nymphalidae (13) Host Family (No. spp.) Guild/Notes Lepidoptera: Crambidae, Gelechiidae, Plutellidae (5) Concealed feeders Lepidoptera: Choreutidae (1) None None None None None None Diptera: Syrphidae; Lepidoptera: Noctuidae (13) None (5) Tortricidae Lepidoptera: Crambidae, Pyralidae, Concealed feeders None Lepidoptera: Gelechiidae, Pyralidae (11) Lepidoptera: Noctuidae (1) None Lepidoptera: Noctuidae, Plutellidae, Pyralidae (4) (4) Tortricidae Lepidoptera: None None None (1) Tortricidae Lepidoptera: (4) Tortricidae Lepidoptera: Gelechiidae, Concealed feeders (3) Tortricidae Lepidoptera: None Lepidoptera: Geometridae (5) Lepidoptera: Geometridae, Lasiocampidae, Erebidae Mostly loopers (11) Hymenoptera: Diprionidae; Lepidoptera: Erebidae, Noctuidae (5) Mostly setose Lepidoptera: Erebidae, Geometridae, Nymphalidae (4) Diptera: Chloropidae; Lepidoptera: Geometridae (2) Lepidoptera: Geometridae (4) None None None Lepidoptera: Crambidae, Gelechiidae (2) Concealed feeders None Lepidoptera: Plutellidae (1) Lepidoptera: Pyralidae (2) Lepidoptera: Gelechiidae, Oecophoridae (3) Concealed feeders None None Lepidoptera: Geometridae (1) Lepidoptera: Noctuidae (1) Lepidoptera: Erebidae (1) Setose Lepidoptera: Geometridae (2) Lepidoptera: Geometridae (1) , LaSalle Minnedosa Winnipeg ; SK: Saskatoon; MB: Aweme, Glenlea Aweme, ; SK: Saskatoon; MB: Wabamun b AB: Wagner NA; MB: Aweme, Winnipeg Aweme, NA; MB: AB: Wagner Localities Lakes AB: Waterton SK: Saskatoon SK: Saskatoon SK: Saskatoon AB: Bilby Vermilion AB: Cypress Hills, Lethbridge, Medicine Hat, SK: Saskatoon MB: Aweme SK: Indian Head SK: Saskatoon Vermilion; AB: Edmonton, AB: Cowley SK: Kinistino AB: Brooks, Waterton AB: Cochrane AB: Lethbridge, Edmonton SK: Indian Head AB: Edmonton AB: Edmonton AB: Lethbridge AB: Lethbridge AB: Cypress Hills, Edmonton; SK: Saskatoon AB: Edmonton NA AB: Wagner NA AB: Wagner NA; SK: Saskatoon Wagner AB: Blackfoot Hills, Cochrane, Swift Current) NA; SK: Great Sand Hills (W Wagner AB: SK: Regina, Uren AB: Cypress Hills Ushta Siding SK: Nipawin, Prairie River, AB: Edmonton; SK: Elbow MB: Carberry Orion; MB: Carberry Lethbridge, Norton, Onefour, AB: Elkwater, AB: Lethbridge SK: Saskatoon Saskatoon SK: Elbow, Waterton; Onefour, AB: Elkwater, MB: Watson; Melfort, Saskatoon, Aylsham, AB: nr Lethbridge; SK: Franklin , Glenlea AB: Edmonton; midcentral Canada (AB/SK) in sunflower fields Bunch SK: Willow AB: Cochrane NA AB: Wagner NA Wagner AB: Bilby, AB: Cochrane NA Wagner AB: Edmonton, SK: Earl Grey AB: Edmonton,

35,N 22,N 32,H 5,8,N 35,N 14,N 14,N 35,N 2,42,N 40,N 30,N 22,N 5,22,N 5,N 14,N 22,N 22,N 14,N 2,32,N 7,N 34,N 34,N,T 22,N 5,N 2,N 22,N

33,N 5,N 22,N 2,N 14,N 2,41,N 42,N 22,N 31,N 34,N 2,32,N 2,14,32,N 22,N 22,N 42,N 9,42,N 2,H (Viereck, 1925) (Viereck, (Viereck, (Viereck, Sanborne, 1986 Viereck, 1912 Viereck, Cushman, 1915 (Viereck, 1925) (Viereck, (Howard, 1889) (Cresson, 1864) Walley, 1970 Walley, (Gravenhorst, 1829) (Cresson, 1864) Walley, 1947 Walley, (Viereck, 1925) (Viereck, (Cresson, 1864) (Viereck, 1925) (Viereck, (Cresson, 1864) (Provancher, 1875) (Provancher, (Viereck, 1925) (Viereck, (Viereck, 1925) (Viereck, a 5,22,N Casinaria petiolaris Campoletis julia Campoletis kingi 1925 Viereck, Campoplex erythromerus 1883) Campoplex mellipes (Provancher, 1947 Casinaria canadensis Walley, 1917) *Diadegma openangorum (Viereck, SUBFAMILY Species 22,N Casinaria forcipata 5,9,36,37,38,39,H,T,O,Oc Campoletis linearis 1925) Campoletis lipomerus (Viereck, Campoletis nigricoxa Campoletis nigripes 1925) Campoletis nigriscaposa (Viereck, *Campoletis oxylus 1927) Campoletis parasignata (Walley, Campoletis patsuiketorum Campoletis plena Campoletis pyralidis Campoletis septentrionalis Campoletis tibialis Campoletis tibiator 1925) Campoplex crassatus (Viereck, 1874) Campoplex fusiformis (Provancher, 1925) Campoplex hexagonalis (Viereck, 1874) Campoplex infumatus (Provancher, Campoplex tortricidis Casinaria eupitheciae Casinaria genuina (Norton, 1863) Casinaria limenitidis 1912 Casinaria scabriformis Viereck, 1941 Casinaria semiothisae Walley, 1925 Cymodusa melanocera Viereck, 1925) *Cymodusa partis (Viereck, 1912 *Cymodusopsis aristoteliae Viereck, Cymodusopsis latifacies Cymodusopsis variabilis Sanborne, 1986 1925) Diadegma stenosomum (Viereck, 1925) Diadegma trachas (Viereck, 1868) Dusona annexa (Förster, 1901) Torre, Dusona bellula (Dalla 1886) Dusona crassicornis (Provancher, 1940) Dusona deceptor (Walley, *Cymodusa distincta (Cresson, 1865) *Diadegma insulare 1940) Dusona canadensis (Walley, Dusona diversa (Norton, 1863) 1917) Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 339 Mostly setose Loopers Loopers Loopers Loopers Concealed feeders Concealed feeders Concealed feeders Concealed feeders Concealed feeders Cutworms Concealed feeders Concealed feeders Lepidoptera: Erebidae, Geometridae, Lasiocampidae, Notodontidae, Nymphalidae (13) Host Family (No. spp.) Guild/Notes Lepidoptera: Crambidae, Gelechiidae, Plutellidae (5) Concealed feeders Lepidoptera: Choreutidae (1) None None None None None None Diptera: Syrphidae; Lepidoptera: Noctuidae (13) None (5) Tortricidae Lepidoptera: Crambidae, Pyralidae, Concealed feeders None Lepidoptera: Gelechiidae, Pyralidae (11) Lepidoptera: Noctuidae (1) None Lepidoptera: Noctuidae, Plutellidae, Pyralidae (4) (4) Tortricidae Lepidoptera: None None None (1) Tortricidae Lepidoptera: (4) Tortricidae Lepidoptera: Gelechiidae, Concealed feeders (3) Tortricidae Lepidoptera: None Lepidoptera: Geometridae (5) Lepidoptera: Geometridae, Lasiocampidae, Erebidae Mostly loopers (11) Hymenoptera: Diprionidae; Lepidoptera: Erebidae, Noctuidae (5) Mostly setose Lepidoptera: Erebidae, Geometridae, Nymphalidae (4) Diptera: Chloropidae; Lepidoptera: Geometridae (2) Lepidoptera: Geometridae (4) None None None Lepidoptera: Crambidae, Gelechiidae (2) Concealed feeders None Lepidoptera: Plutellidae (1) Lepidoptera: Pyralidae (2) Lepidoptera: Gelechiidae, Oecophoridae (3) Concealed feeders None None Lepidoptera: Geometridae (1) Lepidoptera: Noctuidae (1) Lepidoptera: Erebidae (1) Setose Lepidoptera: Geometridae (2) Lepidoptera: Geometridae (1) , LaSalle Minnedosa Winnipeg ; SK: Saskatoon; MB: Aweme, Glenlea Aweme, ; SK: Saskatoon; MB: Wabamun b AB: Wagner NA; MB: Aweme, Winnipeg Aweme, NA; MB: AB: Wagner Localities Lakes AB: Waterton SK: Saskatoon SK: Saskatoon SK: Saskatoon AB: Bilby Vermilion AB: Cypress Hills, Lethbridge, Medicine Hat, SK: Saskatoon MB: Aweme SK: Indian Head SK: Saskatoon Vermilion; AB: Edmonton, AB: Cowley SK: Kinistino AB: Brooks, Waterton AB: Cochrane AB: Lethbridge, Edmonton SK: Indian Head AB: Edmonton AB: Edmonton AB: Lethbridge AB: Lethbridge AB: Cypress Hills, Edmonton; SK: Saskatoon AB: Edmonton NA AB: Wagner NA AB: Wagner NA; SK: Saskatoon Wagner AB: Blackfoot Hills, Cochrane, Swift Current) NA; SK: Great Sand Hills (W Wagner AB: SK: Regina, Uren AB: Cypress Hills Ushta Siding SK: Nipawin, Prairie River, AB: Edmonton; SK: Elbow MB: Carberry Orion; MB: Carberry Lethbridge, Norton, Onefour, AB: Elkwater, AB: Lethbridge SK: Saskatoon Saskatoon SK: Elbow, Waterton; Onefour, AB: Elkwater, MB: Watson; Melfort, Saskatoon, Aylsham, AB: nr Lethbridge; SK: Franklin , Glenlea AB: Edmonton; midcentral Canada (AB/SK) in sunflower fields Bunch SK: Willow AB: Cochrane NA AB: Wagner NA Wagner AB: Bilby, AB: Cochrane NA Wagner AB: Edmonton, SK: Earl Grey AB: Edmonton,

35,N 22,N 32,H 5,8,N 35,N 14,N 14,N 35,N 2,42,N 40,N 30,N 22,N 5,22,N 5,N 14,N 22,N 22,N 14,N 2,32,N 7,N 34,N 34,N,T 22,N 5,N 2,N 22,N

33,N 5,N 22,N 2,N 14,N 2,41,N 42,N 22,N 31,N 34,N 2,32,N 2,14,32,N 22,N 22,N 42,N 9,42,N 2,H (Viereck, 1925) (Viereck, (Viereck, (Viereck, Sanborne, 1986 Viereck, 1912 Viereck, Cushman, 1915 (Viereck, 1925) (Viereck, (Howard, 1889) (Cresson, 1864) Walley, 1970 Walley, (Gravenhorst, 1829) (Cresson, 1864) Walley, 1947 Walley, (Viereck, 1925) (Viereck, (Cresson, 1864) (Viereck, 1925) (Viereck, (Cresson, 1864) (Provancher, 1875) (Provancher, (Viereck, 1925) (Viereck, (Viereck, 1925) (Viereck, a 5,22,N Casinaria petiolaris Campoletis julia Campoletis kingi 1925 Viereck, Campoplex erythromerus 1883) Campoplex mellipes (Provancher, 1947 Casinaria canadensis Walley, 1917) *Diadegma openangorum (Viereck, SUBFAMILY Species 22,N Casinaria forcipata 5,9,36,37,38,39,H,T,O,Oc Campoletis linearis 1925) Campoletis lipomerus (Viereck, Campoletis nigricoxa Campoletis nigripes 1925) Campoletis nigriscaposa (Viereck, *Campoletis oxylus 1927) Campoletis parasignata (Walley, Campoletis patsuiketorum Campoletis plena Campoletis pyralidis Campoletis septentrionalis Campoletis tibialis Campoletis tibiator 1925) Campoplex crassatus (Viereck, 1874) Campoplex fusiformis (Provancher, 1925) Campoplex hexagonalis (Viereck, 1874) Campoplex infumatus (Provancher, Campoplex tortricidis Casinaria eupitheciae Casinaria genuina (Norton, 1863) Casinaria limenitidis 1912 Casinaria scabriformis Viereck, 1941 Casinaria semiothisae Walley, 1925 Cymodusa melanocera Viereck, 1925) *Cymodusa partis (Viereck, 1912 *Cymodusopsis aristoteliae Viereck, Cymodusopsis latifacies Cymodusopsis variabilis Sanborne, 1986 1925) Diadegma stenosomum (Viereck, 1925) Diadegma trachas (Viereck, 1868) Dusona annexa (Förster, 1901) Torre, Dusona bellula (Dalla 1886) Dusona crassicornis (Provancher, 1940) Dusona deceptor (Walley, *Cymodusa distincta (Cresson, 1865) *Diadegma insulare 1940) Dusona canadensis (Walley, Dusona diversa (Norton, 1863) 1917) 340 M. D. Schwarzfeld Concealed feeders Loopers Loopers Loopers Loopers Loopers Loopers Loopers Loopers Leafrollers Cutworms Host Family (No. spp.) Guild/Notes Lepidoptera: Noctuidae (1) None Lepidoptera: Geometridae (2) None None Lepidoptera: Noctuidae (1) Lepidoptera: Geometridae (2) Lepidoptera: Geometridae, Notodontidae (2) Hymenoptera: Diprionidae Lepidoptera: Geometridae (3) Conifer feeders None Lepidoptera: Geometridae (1) Lepidoptera: Geometridae (8) None (1) Tortricidae Lepidoptera: Lepidoptera: Notodontidae (1) None None None Lepidoptera: Geometridae (1) Lepidoptera: Geometridae (2) Lepidoptera: Geometridae (1) Lepidoptera: Geometridae, Lasiocampidae (4) Mostly loopers Lepidoptera: Noctuidae (1) Lepidoptera: Erebidae (2) Bristly lichen feeders Lepidoptera: Coleophoridae, Gelechiidae, Pyralidae, (13) Tortricidae Concealed feeders (12) Tortricidae Lepidoptera: Pyralidae, Concealed feeders Lepidoptera: Geometridae (1) Lepidoptera: Crambidae, Coleophoridae, Erebidae, Gelechiidae, Lasiocampidae, Noctuidae, Nymphalidae, (34) Tortricidae Pieridae, Pyralidae, None None None Lepidoptera: Erebidae, Geometridae, Noctuidae, Nymphalidae, Pieridae, Sphingidae (24) Lepidoptera: Erebidae, Geometridae, Hesperiidae, Lasiocampidae, Limacodidae, Lycaenidae, Megalopygidae, Noctuidae, Notodontidae, Nymphalidae, Pyralidae, Saturniidae (31) Lepidoptera: Pterophoridae (1) None Lepidoptera: Erebidae (8) Setose None None None None None Lepidoptera: Geometridae, Noctuidae (9) Lepidoptera: Erebidae, Geometridae, Lasiocampidae, Notodontidae, Saturniidae, Sphingidae (16) , Oak Bluff, St. Claude Gladstone , Oak Bluff, , Roblin Swan River b Localities NA, Waterton AB: Wagner Aweme AB: Bilby; MB: NA Wagner AB: Gull L, Red Deer, Aweme AB: Edmonton; MB: MB: Teulon SK: Radison, Waterton; Edmonton, Lethbridge, Calgary, AB: Bilby, Saskatoon; Transcona MB: Aweme, MB: Aweme AB: Waterton NA Wagner AB: Cypress Hills, Edmonton, Lethbridge, NA AB: Wagner NA Wagner AB: Edmonton, NA Wagner AB: Edmonton, NA Wagner AB: Edmonton, Aweme MB: Waterton; NA, Wagner AB: Edmonton, MB: Transcona Transcona Aweme, SK: Earl Grey; MB: AB: Waterton NA Wagner AB: Edmonton, NA Wagner Edmonton, AB: Calgary, NA AB: Wagner AB: Calgary NA Wagner AB: Edmonton, SK: Saskatoon Waterton; AB: Edmonton, SK: St. Louis, Cut Knife AB: Edmonton MB: Kenville AB: nr Calgary Waterton Lethbridge, AB: Calgary, Lakes AB: Waterton SK: Indian Head SK: St. Louis SK: Kinistino MB: Gladstone , Piney Lakes AB: Waterton AB: Edmonton MB: Carman, AB: Edmonton MB: Transcona MB: Winnipeg MB: Winnipeg AB: Edmonton AB, Southern SK Southern AB: sites in parkland and Peace River area 44,N 46,N 5,N 5,N 22,N 5,41,43,N 22,N 22,N 42,N 5,43,N 2,42,N 9,H 2,5,H,O 22,N 42,N 22,N 22,N,T,Oc 2,N 5,N 2,N 2,5,42,H 9,10,N 2,42,N 2,5,42,N 2,5,42,N 2,5,N ) 5,42,N 29,H 2,42,N 22,N 42,N 22,N 9,N,T 14,N 42,N 42,N 14,42,N 5,N 2,5,42,N 5,N 45,N 42,N 2,5,N (Weed, 1903) (Weed, (Ashmead, 1902) (Viereck, 1925) (Viereck, (Viereck, 1925) (Viereck, (Davis, 1898) (Say, 1835) (Say, (Viereck, 1925) (Viereck, (Viereck, 1912) (Viereck, (Viereck, 1925) (Viereck, (Cresson, 1865 (Norton, 1863) (Viereck, 1925) (Viereck, (Viereck, 1925) (Viereck, (Gravenhorst, 1829) a (Viereck, 1925) Dusona downesi (Viereck, 1882) Dusona semirufa (Provancher, SUBFAMILY Species (Ashmead, 1890) Enytus eureka (Viereck, 1926) Dusona fossata (Viereck, Dusona glauca (Norton, 1863) 1940) Dusona laminata (Walley, Dusona laticincta 1940) Dusona lobata (Walley, 1879) Dusona minor (Provancher, Dusona occidentalis 1940) Dusona pectoralis (Walley, 1925) Dusona seamansi (Viereck, 1925) Dusona signata (Viereck, 1925) Dusona stricklandi (Viereck, 1940) Dusona vara (Walley, 1926) Dusona varicoxa (Viereck, 1906) Dusona wyomingensis (Viereck, Cushman, 1924 ochreofrons Echthronomas Enytus montanus (Ashmead, 1890) Enytus oculus 1925) Hyposoter asper (Viereck, 1925) Hyposoter degrysei (Viereck, 1925) Hyposoter erythrinus (Viereck, Hyposoter exiguae Hyposoter fugitivus Hyposoter occidentali Hyposoter popofensis (Ashmead, 1902) Hyposoter rivalis (Cresson, 1872) Olesicampe alaskensis Olesicampe banffensis Olesicampe lata Olesicampe patula Olesicampe typica Phobocampe bicingulata Phobocampe clisiocampae (Walley, 1940) Dusona insolita (Walley, Dusona major (Cresson, 1879) Dusona petiolator (Fabricius, 1804) 1886) Dusona scalaria (Provancher, 1874) Dusona vicina (Provancher, Dusona villosa (Norton, 1863) Dusona vitticollis Hyposoter annulipes (Cresson, 1864) Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 341 Concealed feeders Loopers Loopers Loopers Loopers Loopers Loopers Loopers Loopers Leafrollers Cutworms Host Family (No. spp.) Guild/Notes Lepidoptera: Noctuidae (1) None Lepidoptera: Geometridae (2) None None Lepidoptera: Noctuidae (1) Lepidoptera: Geometridae (2) Lepidoptera: Geometridae, Notodontidae (2) Hymenoptera: Diprionidae Lepidoptera: Geometridae (3) Conifer feeders None Lepidoptera: Geometridae (1) Lepidoptera: Geometridae (8) None (1) Tortricidae Lepidoptera: Lepidoptera: Notodontidae (1) None None None Lepidoptera: Geometridae (1) Lepidoptera: Geometridae (2) Lepidoptera: Geometridae (1) Lepidoptera: Geometridae, Lasiocampidae (4) Mostly loopers Lepidoptera: Noctuidae (1) Lepidoptera: Erebidae (2) Bristly lichen feeders Lepidoptera: Coleophoridae, Gelechiidae, Pyralidae, (13) Tortricidae Concealed feeders (12) Tortricidae Lepidoptera: Pyralidae, Concealed feeders Lepidoptera: Geometridae (1) Lepidoptera: Crambidae, Coleophoridae, Erebidae, Gelechiidae, Lasiocampidae, Noctuidae, Nymphalidae, (34) Tortricidae Pieridae, Pyralidae, None None None Lepidoptera: Erebidae, Geometridae, Noctuidae, Nymphalidae, Pieridae, Sphingidae (24) Lepidoptera: Erebidae, Geometridae, Hesperiidae, Lasiocampidae, Limacodidae, Lycaenidae, Megalopygidae, Noctuidae, Notodontidae, Nymphalidae, Pyralidae, Saturniidae (31) Lepidoptera: Pterophoridae (1) None Lepidoptera: Erebidae (8) Setose None None None None None Lepidoptera: Geometridae, Noctuidae (9) Lepidoptera: Erebidae, Geometridae, Lasiocampidae, Notodontidae, Saturniidae, Sphingidae (16) , Oak Bluff, St. Claude Gladstone , Oak Bluff, , Roblin Swan River b Localities NA, Waterton AB: Wagner Aweme AB: Bilby; MB: NA Wagner AB: Gull L, Red Deer, Aweme AB: Edmonton; MB: MB: Teulon SK: Radison, Waterton; Edmonton, Lethbridge, Calgary, AB: Bilby, Saskatoon; Transcona MB: Aweme, MB: Aweme AB: Waterton NA Wagner AB: Cypress Hills, Edmonton, Lethbridge, NA AB: Wagner NA Wagner AB: Edmonton, NA Wagner AB: Edmonton, NA Wagner AB: Edmonton, Aweme MB: Waterton; NA, Wagner AB: Edmonton, MB: Transcona Transcona Aweme, SK: Earl Grey; MB: AB: Waterton NA Wagner AB: Edmonton, NA Wagner Edmonton, AB: Calgary, NA AB: Wagner AB: Calgary NA Wagner AB: Edmonton, SK: Saskatoon Waterton; AB: Edmonton, SK: St. Louis, Cut Knife AB: Edmonton MB: Kenville AB: nr Calgary Waterton Lethbridge, AB: Calgary, Lakes AB: Waterton SK: Indian Head SK: St. Louis SK: Kinistino MB: Gladstone , Piney Lakes AB: Waterton AB: Edmonton MB: Carman, AB: Edmonton MB: Transcona MB: Winnipeg MB: Winnipeg AB: Edmonton AB, Southern SK Southern AB: sites in parkland and Peace River area 44,N 46,N 5,N 5,N 22,N 5,41,43,N 22,N 22,N 42,N 5,43,N 2,42,N 9,H 2,5,H,O 22,N 42,N 22,N 22,N,T,Oc 2,N 5,N 2,N 2,5,42,H 9,10,N 2,42,N 2,5,42,N 2,5,42,N 2,5,N ) 5,42,N 29,H 2,42,N 22,N 42,N 22,N 9,N,T 14,N 42,N 42,N 14,42,N 5,N 2,5,42,N 5,N 45,N 42,N 2,5,N (Weed, 1903) (Weed, (Ashmead, 1902) (Viereck, 1925) (Viereck, (Viereck, 1925) (Viereck, (Davis, 1898) (Say, 1835) (Say, (Viereck, 1925) (Viereck, (Viereck, 1912) (Viereck, (Viereck, 1925) (Viereck, (Cresson, 1865 (Norton, 1863) (Viereck, 1925) (Viereck, (Viereck, 1925) (Viereck, (Gravenhorst, 1829) a (Viereck, 1925) Dusona downesi (Viereck, 1882) Dusona semirufa (Provancher, SUBFAMILY Species (Ashmead, 1890) Enytus eureka (Viereck, 1926) Dusona fossata (Viereck, Dusona glauca (Norton, 1863) 1940) Dusona laminata (Walley, Dusona laticincta 1940) Dusona lobata (Walley, 1879) Dusona minor (Provancher, Dusona occidentalis 1940) Dusona pectoralis (Walley, 1925) Dusona seamansi (Viereck, 1925) Dusona signata (Viereck, 1925) Dusona stricklandi (Viereck, 1940) Dusona vara (Walley, 1926) Dusona varicoxa (Viereck, 1906) Dusona wyomingensis (Viereck, Cushman, 1924 ochreofrons Echthronomas Enytus montanus (Ashmead, 1890) Enytus oculus 1925) Hyposoter asper (Viereck, 1925) Hyposoter degrysei (Viereck, 1925) Hyposoter erythrinus (Viereck, Hyposoter exiguae Hyposoter fugitivus Hyposoter occidentali Hyposoter popofensis (Ashmead, 1902) Hyposoter rivalis (Cresson, 1872) Olesicampe alaskensis Olesicampe banffensis Olesicampe lata Olesicampe patula Olesicampe typica Phobocampe bicingulata Phobocampe clisiocampae (Walley, 1940) Dusona insolita (Walley, Dusona major (Cresson, 1879) Dusona petiolator (Fabricius, 1804) 1886) Dusona scalaria (Provancher, 1874) Dusona vicina (Provancher, Dusona villosa (Norton, 1863) Dusona vitticollis Hyposoter annulipes (Cresson, 1864) 342 M. D. Schwarzfeld Web-spinning sawflies Web-spinning Loopers Concealed feeders Conifer shoot borers Conifer shoot borers Loopers Host Family (No. spp.) Guild/Notes None Lepidoptera: Erebidae, Geometridae, Noctuidae (10) Coleoptera: Elateridae (1) (3) Tortricidae Lepidoptera: Artematopodidae Coleoptera: (1) Coleoptera: Cerambycidae, Curculionidae (2) Borers None None None Coleoptera: Cerambycidae (1) None Lepidoptera: Noctuidae (2) (3) Tortricidae Lepidoptera: Noctuidae, None Lepidoptera: Elachistidae, Noctuidae (2) None Lepidoptera: Noctuidae (3) Lepidoptera: Noctuidae (1) None None Hymenoptera: Pamphiliidae (2) (2) Tortricidae Lepidoptera: Lepidoptera: Geometridae (1) None (1) Tortricidae Lepidoptera: None Coleoptera: Cerambycidae; Lepidoptera: Crambidae (2) Lepidoptera: Crambidae, Erebidae, Geometridae, (9) Tortricidae Pyralidae, Mostly concealed feeders Lepidoptera: Notodontidae (2) Lepidoptera: Pyralidae (1) Hymenoptera: Ichneumonidae, Coleophoridae; Lepidoptera: Crambidae, Erebidae, Geometridae, Lasiocampidae, Noctuidae, Notodontidae, Pyralidae, (32) Tortricidae Lepidoptera: Noctuidae (1) Argidae Hymenoptera: (1) Lepidoptera: Geometridae, Noctuidae, Nolidae, (5) Tortricidae Lepidoptera: Erebidae, Crambidae, Gelechiidae, Tortricidae, Noctuidae, Tineidae, Pyralidae, (22) Yponomeutidae Mostly concealed feeders Lepidoptera: Pyralidae (1) None None b Localities AB, Southern SK Southern MB: Winnipeg SK: Asquith AB: Edmonton NA AB: Wagner AB: Edmonton AB: Bilby Villeneuve Edmonton, AB: Calgary, AB: Edmonton AB: Edmonton AB: Drumheller; SK: Elbow MB: Winnipeg AB: Lethbridge, Medicine Hat; SK: Saskatoon, Swift Current AB: Waterton AB: Waterton Edmonton, Butte AB: Twin Bunch Willow Stettler; SK: Lisieux, Regina, Rockglen, AB: Drumheller, SK: Tyner AB: Oyen AB: Waterton AB: Delburne Great Sand Hills, Swift Edmonton, Gleichen; SK: Elbow, AB: Calgary, Current AB: Three Creeks SK: Elbow MB: Shilo SK: Tunstall Glenwood, Edmonton, Elkwater, Coaldale, Drumheller, AB: Calgary, Orion, Manyberries, Medicine Hat, Milk River, Lethbridge, Lost River, SK: Winnifred; Waterton, Three Hills, Red Deer, Oyen, Pearce, Rainier, Regina, Rosthern, Rutland, Saskatoon, Buchanan, Estevan, Great Deer, White Fox; Tunstall, Torquay, Stoughton, Strongfield, Swift Current, MB: Brandon, Minnedosa Kinsella, Stettler, Calgary, AB: Edmonton, Jumping Pond Cr, Waterton Wainwright, Lethbridge AB: Oldman River, Aweme MB: Waterton; Lethbridge, AB: Edmonton, Jumping Pound Cr, Tangleflags Calgary); SK: W AB: Jumping Pound Cr (20 mi AB: Edmonton AB: Edmonton Throughout Nearctic, indoors, attacking caterpillars on AB: Lethbridge; stored products MB: Whitemouth MB: Morris, Treesbank NA Wagner AB: Edmonton, Midcentral Canada (AB/SK) in sunflower fields 2,5,N 50,H 49,N 9,22,N 49,N 49,N 49,N 5,49,N 5,51,W 29,H 49,N 49,N 14,N 49,N 5,H 2,H 49,H 49,N 48,N 49,N,T 49,H,O 49,N 49,N 5,49,N 49,N 49,N 49,N 9,N 49,H 51,N

51,N Horstmann, (Sanborne, (Viereck, 1925) (Viereck, (Davis, 1898) Sanborne, 1984 (Thomson, 1887) (Brischke, 1880) (Provancher, 1874) (Provancher, Barron & Walley, 1983 Walley, Barron & Wahl, 1987 Wahl, (Viereck, 1905) (Viereck, (Holmgren, 1860) 14,N 48,N (Provancher, 1874) (Provancher, (Gravenhorst, 1829) 48,N Wahl, 1987 Wahl, a 48,N 48,N Phobocampe confusa Pyracmon sepiellus Sinophorus orientis Sanborne, 1984 Sinophorus sabulus Sanborne, 1984 SUBFAMILY Species 47,N Sinophorus nitidus (Brischke, 1880) Phobocampe flavipes Pyracmon ctenicerae Pyracmon hyalinus Rhimphoctona aldrichi Rhimphoctona brevicauda 1986) Rhimphoctona longicauda 1980 Rhimphoctona macdunnoughi 1925) (Viereck, Rhimphoctona macrocephala Rhimphoctona vancouverensis (Harrington, 1894) 1944) Scirtetes canadensis (Wally, 1912) Sinophorus caradrinae (Viereck, Sinophorus constrictus Sanborne, 1984 Sinophorus costalis (Thomson, 1887) Sanborne, 1984 Sinophorus erectus 1958) Sinophorus eruficinctus (Walkley, Sinophorus heliothidis Sinophorus infimus Sanborne, 1984 Sinophorus masoni Sanborne, 1984 Sinophorus megalodontis Sanborne, 1984 Sinophorus relativus *Sinophorus sulcatellus 1887) *Sinophorus teratis (Weed, Sinophorus townesorum Sanborne, 1984 *Sinophorus tumidus Sanborne, 1984 8,N Sinophorus validus (Cresson, 1864) Sinophorus vericulus Sanborne, 1984 carbonellum (Thomson, 1887) Tranosema coxalis Tranosemella canescens Venturia finlaysonae Venturia scitula Venturia 1944) Xanthocampoplex orbitalis (Walley, (Provancher, 1874) (Provancher, Sinophorus platycephalus Sanborne, 1984 Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 343 Web-spinning sawflies Web-spinning Loopers Concealed feeders Conifer shoot borers Conifer shoot borers Loopers Host Family (No. spp.) Guild/Notes None Lepidoptera: Erebidae, Geometridae, Noctuidae (10) Coleoptera: Elateridae (1) (3) Tortricidae Lepidoptera: Artematopodidae Coleoptera: (1) Coleoptera: Cerambycidae, Curculionidae (2) Borers None None None Coleoptera: Cerambycidae (1) None Lepidoptera: Noctuidae (2) (3) Tortricidae Lepidoptera: Noctuidae, None Lepidoptera: Elachistidae, Noctuidae (2) None Lepidoptera: Noctuidae (3) Lepidoptera: Noctuidae (1) None None Hymenoptera: Pamphiliidae (2) (2) Tortricidae Lepidoptera: Lepidoptera: Geometridae (1) None (1) Tortricidae Lepidoptera: None Coleoptera: Cerambycidae; Lepidoptera: Crambidae (2) Lepidoptera: Crambidae, Erebidae, Geometridae, (9) Tortricidae Pyralidae, Mostly concealed feeders Lepidoptera: Notodontidae (2) Lepidoptera: Pyralidae (1) Hymenoptera: Ichneumonidae, Coleophoridae; Lepidoptera: Crambidae, Erebidae, Geometridae, Lasiocampidae, Noctuidae, Notodontidae, Pyralidae, (32) Tortricidae Lepidoptera: Noctuidae (1) Argidae Hymenoptera: (1) Lepidoptera: Geometridae, Noctuidae, Nolidae, (5) Tortricidae Lepidoptera: Erebidae, Crambidae, Gelechiidae, Tortricidae, Noctuidae, Tineidae, Pyralidae, (22) Yponomeutidae Mostly concealed feeders Lepidoptera: Pyralidae (1) None None b Localities AB, Southern SK Southern MB: Winnipeg SK: Asquith AB: Edmonton NA AB: Wagner AB: Edmonton AB: Bilby Villeneuve Edmonton, AB: Calgary, AB: Edmonton AB: Edmonton AB: Drumheller; SK: Elbow MB: Winnipeg AB: Lethbridge, Medicine Hat; SK: Saskatoon, Swift Current AB: Waterton AB: Waterton Edmonton, Butte AB: Twin Bunch Willow Stettler; SK: Lisieux, Regina, Rockglen, AB: Drumheller, SK: Tyner AB: Oyen AB: Waterton AB: Delburne Great Sand Hills, Swift Edmonton, Gleichen; SK: Elbow, AB: Calgary, Current AB: Three Creeks SK: Elbow MB: Shilo SK: Tunstall Glenwood, Edmonton, Elkwater, Coaldale, Drumheller, AB: Calgary, Orion, Manyberries, Medicine Hat, Milk River, Lethbridge, Lost River, SK: Winnifred; Waterton, Three Hills, Red Deer, Oyen, Pearce, Rainier, Regina, Rosthern, Rutland, Saskatoon, Buchanan, Estevan, Great Deer, White Fox; Tunstall, Torquay, Stoughton, Strongfield, Swift Current, MB: Brandon, Minnedosa Kinsella, Stettler, Calgary, AB: Edmonton, Jumping Pond Cr, Waterton Wainwright, Lethbridge AB: Oldman River, Aweme MB: Waterton; Lethbridge, AB: Edmonton, Jumping Pound Cr, Tangleflags Calgary); SK: W AB: Jumping Pound Cr (20 mi AB: Edmonton AB: Edmonton Throughout Nearctic, indoors, attacking caterpillars on AB: Lethbridge; stored products MB: Whitemouth MB: Morris, Treesbank NA Wagner AB: Edmonton, Midcentral Canada (AB/SK) in sunflower fields 2,5,N 50,H 49,N 9,22,N 49,N 49,N 49,N 5,49,N 5,51,W 29,H 49,N 49,N 14,N 49,N 5,H 2,H 49,H 49,N 48,N 49,N,T 49,H,O 49,N 49,N 5,49,N 49,N 49,N 49,N 9,N 49,H 51,N

51,N Horstmann, (Sanborne, (Viereck, 1925) (Viereck, (Davis, 1898) Sanborne, 1984 (Thomson, 1887) (Brischke, 1880) (Provancher, 1874) (Provancher, Barron & Walley, 1983 Walley, Barron & Wahl, 1987 Wahl, (Viereck, 1905) (Viereck, (Holmgren, 1860) 14,N 48,N (Provancher, 1874) (Provancher, (Gravenhorst, 1829) 48,N Wahl, 1987 Wahl, a 48,N 48,N Phobocampe confusa Pyracmon sepiellus Sinophorus orientis Sanborne, 1984 Sinophorus sabulus Sanborne, 1984 SUBFAMILY Species 47,N Sinophorus nitidus (Brischke, 1880) Phobocampe flavipes Pyracmon ctenicerae Pyracmon hyalinus Rhimphoctona aldrichi Rhimphoctona brevicauda 1986) Rhimphoctona longicauda 1980 Rhimphoctona macdunnoughi 1925) (Viereck, Rhimphoctona macrocephala Rhimphoctona vancouverensis (Harrington, 1894) 1944) Scirtetes canadensis (Wally, 1912) Sinophorus caradrinae (Viereck, Sinophorus constrictus Sanborne, 1984 Sinophorus costalis (Thomson, 1887) Sanborne, 1984 Sinophorus erectus 1958) Sinophorus eruficinctus (Walkley, Sinophorus heliothidis Sinophorus infimus Sanborne, 1984 Sinophorus masoni Sanborne, 1984 Sinophorus megalodontis Sanborne, 1984 Sinophorus relativus *Sinophorus sulcatellus 1887) *Sinophorus teratis (Weed, Sinophorus townesorum Sanborne, 1984 *Sinophorus tumidus Sanborne, 1984 8,N Sinophorus validus (Cresson, 1864) Sinophorus vericulus Sanborne, 1984 carbonellum (Thomson, 1887) Tranosema coxalis Tranosemella canescens Venturia finlaysonae Venturia scitula Venturia 1944) Xanthocampoplex orbitalis (Walley, (Provancher, 1874) (Provancher, Sinophorus platycephalus Sanborne, 1984 344 M. D. Schwarzfeld Mostly Borers Borers Concealed feeders Concealed feeders Concealed feeders Concealed feeders Concealed feeders Host Family (No. spp.) Guild/Notes None None None Coleoptera: Cerambycidae (1) None Lepidoptera: Gelechiidae (1) Stem gall makers Lepidoptera: Noctuidae (1) (1) Tephritidae Diptera: Coleoptera: Mordellidae (3) Coleoptera: Cerambycidae (1) None Lepidoptera: Coleophoridae (1) None None Lepidoptera: Coleophoridae, Cosmopterigidae, (6) Tortricidae Gelechiidae, Concealed feeders None Coleoptera: Curculionidae; Lepidoptera: Cosmopterigidae, Elachistidae, Gelechiidae, (22) Tortricidae Nymphalidae, Pyralidae, Mostly stem borers None Lepidoptera: Crambidae, Erebidae, Gelechiidae, (28) Tortricidae Noctuidae, Pieridae, Pyralidae, Lepidoptera: Crambidae (2) (2) Tortricidae Lepidoptera: None None None Coleoptera: Cerambycidae; Lepidoptera: (18) Tortricidae Coleophoridae, Gelechiidae, concealed feeders None None Lepidoptera: Choreutidae, Gelechiidae, Momphidae, (21) Tortricidae Pyralidae, (1) Tortricidae Lepidoptera: (8) Tortricidae Lepidoptera: Coleophoridae, Concealed feeders Lepidoptera: Gelechiidae (1) None b Localities SK: Saskatoon MB: Brandon AB: Calgary MB: Brandon SK: Saskatoon; MB: Ninette AB: Medicine Hat SK: Elbow Edmonton, Elkwater, Aden, Coaldale, Cypress Hills, Drumheller, AB: Lethbridge, Magrath, Manyberries, Medicine Irvine, Jumping Pound Cr, Attons L, Duck SK: Wayne; Taber, Morrin, Steveville, Hat, Milk River, Albert, Redberry, Parkbeg, Prince Grenfell, Lloydminster, L, Elbow, Carberry, Aweme, MB: Tompkins; Roadene, Saskatoon, Swift Current, Vita, Shilo, Bog at 5 mi SW Tamarack Ninette, Pierson, 3 mi S Shilo, Winnipeg Wawanesa, Aweme, AB: Edmonton, Lethbridge, Medicine Hat; SK: Saskatoon; MB: Brandon SK: Indian Head SK: Elbow AB: Onefour; SK: Swift Current AB: Edmonton, Ponoka SK: White Fox SK: Great Sand Hills, Saskatchewan Landing AB: Clymont SK: Indian Head, Saskatoon MB: Ninette "Cameron Ranch", Cassils, Chin, AB: Brooks, Burdett, Calgary, Glenwood, Clarinda, Cochrane, Cypress Hills, Edmonton, Elkwater P, Lethbridge Manyberries, Medicine Hat, Gull L, Jumping Pound Cr, Rosebud, Orion, Oyen, Picture Butte, Red Deer, Morrin, Onefour, Wild Horse, Wardlow, Vermilion, Tofield, Taber, Scandia, Stavely, Antelope L, Consul, Dollard, Duck Dunblane, Earl SK: Winnifred; Great Sand Hills, Indian Estevan, Garrick, Great Deer, Elbow, Grey, Moosomin, Nipawin, Parkbeg, Regina, Moose Jaw, Head, Kenosee P, Roadene, Roche Percee, Rosetown, Saskatoon, Strongfield, Swift Treesbank Current; MB: Aweme, AB: Wabamun Edmonton, AB: Rosemary AB: Calgary; SK: Saskatoon AB: Edmonton MB: 9 mi N Forrest Stn Ninette Aweme, SK: Saskatoon; MB: SK: Birch Hills Uren, Saskatoon SK: Redberry, AB: Lethbridge, Onefour; SK: Estevan, Great Sand Hills, Kenosee, Treesbank Aweme, Swift Current, Uren; MB: MB: Dauphin, Morris Moosomin, Pike L, Rosetown, AB: Coaldale, Onefour; SK: Elbow, Shilo Saskatoon; MB: Ninette, 5 mi SW Elbow SK: Bounty, SK: Great Sand Hills 52,N 52,N 52,N 52,N 52,N 52,N 52,N 52,N 14,52,N 52,N 52,N 52,N 52,N 52,N 52,N,T 52,N 52,N 52,N 52,N 52,N 52,N 52,N 52,N 52,N 52,N 52,N 52,N 52,N Ashmead, (Cushman, 1930) (Cushman, 1917) Cushman, 1917 Dasch, 1979 Cushman, 1920 Dasch, 1979 Dasch, 1979 (Cushman, 1935) Dasch, 1979 Dasch, 1979 (Weed, 1887) (Weed, (Weed, 1888) (Weed, Dasch, 1979 (Provancher, 1874) (Provancher, a 5,52,N 52,N 5,43,52,53,N,T,Oc 7,52,N,T CREMASTINAE albertensis Dasch, 1979 Cremastus cookii Temelucha Dasch, 1979 crepera Temelucha rhyacioniae Temelucha SUBFAMILY Species Pristomerus euryptychiae 1896 Cremastus canadensis Dasch, 1979 Cremastus Kerrich, 1959 cressoni Cremastus Cushman, 1920 dorcaschemae Cremastus gallaecola Cremastus hyalinipennis (Cresson, 1872) Cremastus incompletus (Provancher, Cremastus 1875) Cushman, 1917 mordellistenae Cremastus 1965 Townes, nordi Cremastus Cushman, 1935 protractus Cremastus Neleothymus pusillus mellipes (Provancher, Nothocremastus 1875) Pristomerus baumhoferi Cushman, 1930 Pristomerus coracinus Dasch, 1979 Pristomerus laticeps *Pristomerus spinator (Fabricius, 1804) chilonis Temelucha cymosa Temelucha dakotae Temelucha forbesi Temelucha fuscalata Temelucha inflata Temelucha platynotae Temelucha undulata Temelucha angulosa Dasch, 1979 Trathala Cremastus flavopictus Dasch, 1979 Cremastus politus Dasch, 1979 Cremastus recta Temelucha Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 345 Mostly Borers Borers Concealed feeders Concealed feeders Concealed feeders Concealed feeders Concealed feeders Host Family (No. spp.) Guild/Notes None None None Coleoptera: Cerambycidae (1) None Lepidoptera: Gelechiidae (1) Stem gall makers Lepidoptera: Noctuidae (1) (1) Tephritidae Diptera: Coleoptera: Mordellidae (3) Coleoptera: Cerambycidae (1) None Lepidoptera: Coleophoridae (1) None None Lepidoptera: Coleophoridae, Cosmopterigidae, (6) Tortricidae Gelechiidae, Concealed feeders None Coleoptera: Curculionidae; Lepidoptera: Cosmopterigidae, Elachistidae, Gelechiidae, (22) Tortricidae Nymphalidae, Pyralidae, Mostly stem borers None Lepidoptera: Crambidae, Erebidae, Gelechiidae, (28) Tortricidae Noctuidae, Pieridae, Pyralidae, Lepidoptera: Crambidae (2) (2) Tortricidae Lepidoptera: None None None Coleoptera: Cerambycidae; Lepidoptera: (18) Tortricidae Coleophoridae, Gelechiidae, concealed feeders None None Lepidoptera: Choreutidae, Gelechiidae, Momphidae, (21) Tortricidae Pyralidae, (1) Tortricidae Lepidoptera: (8) Tortricidae Lepidoptera: Coleophoridae, Concealed feeders Lepidoptera: Gelechiidae (1) None b Localities SK: Saskatoon MB: Brandon AB: Calgary MB: Brandon SK: Saskatoon; MB: Ninette AB: Medicine Hat SK: Elbow Edmonton, Elkwater, Aden, Coaldale, Cypress Hills, Drumheller, AB: Lethbridge, Magrath, Manyberries, Medicine Irvine, Jumping Pound Cr, Attons L, Duck SK: Wayne; Taber, Morrin, Steveville, Hat, Milk River, Albert, Redberry, Parkbeg, Prince Grenfell, Lloydminster, L, Elbow, Carberry, Aweme, MB: Tompkins; Roadene, Saskatoon, Swift Current, Vita, Shilo, Bog at 5 mi SW Tamarack Ninette, Pierson, 3 mi S Shilo, Winnipeg Wawanesa, Aweme, AB: Edmonton, Lethbridge, Medicine Hat; SK: Saskatoon; MB: Brandon SK: Indian Head SK: Elbow AB: Onefour; SK: Swift Current AB: Edmonton, Ponoka SK: White Fox SK: Great Sand Hills, Saskatchewan Landing AB: Clymont SK: Indian Head, Saskatoon MB: Ninette "Cameron Ranch", Cassils, Chin, AB: Brooks, Burdett, Calgary, Glenwood, Clarinda, Cochrane, Cypress Hills, Edmonton, Elkwater P, Lethbridge Manyberries, Medicine Hat, Gull L, Jumping Pound Cr, Rosebud, Orion, Oyen, Picture Butte, Red Deer, Morrin, Onefour, Wild Horse, Wardlow, Vermilion, Tofield, Taber, Scandia, Stavely, Antelope L, Consul, Dollard, Duck Dunblane, Earl SK: Winnifred; Great Sand Hills, Indian Estevan, Garrick, Great Deer, Elbow, Grey, Moosomin, Nipawin, Parkbeg, Regina, Moose Jaw, Head, Kenosee P, Roadene, Roche Percee, Rosetown, Saskatoon, Strongfield, Swift Treesbank Current; MB: Aweme, AB: Wabamun Edmonton, AB: Rosemary AB: Calgary; SK: Saskatoon AB: Edmonton MB: 9 mi N Forrest Stn Ninette Aweme, SK: Saskatoon; MB: SK: Birch Hills Uren, Saskatoon SK: Redberry, AB: Lethbridge, Onefour; SK: Estevan, Great Sand Hills, Kenosee, Treesbank Aweme, Swift Current, Uren; MB: MB: Dauphin, Morris Moosomin, Pike L, Rosetown, AB: Coaldale, Onefour; SK: Elbow, Shilo Saskatoon; MB: Ninette, 5 mi SW Elbow SK: Bounty, SK: Great Sand Hills 52,N 52,N 52,N 52,N 52,N 52,N 52,N 52,N 14,52,N 52,N 52,N 52,N 52,N 52,N 52,N,T 52,N 52,N 52,N 52,N 52,N 52,N 52,N 52,N 52,N 52,N 52,N 52,N 52,N Ashmead, (Cushman, 1930) (Cushman, 1917) Cushman, 1917 Dasch, 1979 Cushman, 1920 Dasch, 1979 Dasch, 1979 (Cushman, 1935) Dasch, 1979 Dasch, 1979 (Weed, 1887) (Weed, (Weed, 1888) (Weed, Dasch, 1979 (Provancher, 1874) (Provancher, a 5,52,N 52,N 5,43,52,53,N,T,Oc 7,52,N,T CREMASTINAE albertensis Dasch, 1979 Cremastus cookii Temelucha Dasch, 1979 crepera Temelucha rhyacioniae Temelucha SUBFAMILY Species Pristomerus euryptychiae 1896 Cremastus canadensis Dasch, 1979 Cremastus Kerrich, 1959 cressoni Cremastus Cushman, 1920 dorcaschemae Cremastus gallaecola Cremastus hyalinipennis (Cresson, 1872) Cremastus incompletus (Provancher, Cremastus 1875) Cushman, 1917 mordellistenae Cremastus 1965 Townes, nordi Cremastus Cushman, 1935 protractus Cremastus Neleothymus pusillus mellipes (Provancher, Nothocremastus 1875) Pristomerus baumhoferi Cushman, 1930 Pristomerus coracinus Dasch, 1979 Pristomerus laticeps *Pristomerus spinator (Fabricius, 1804) chilonis Temelucha cymosa Temelucha dakotae Temelucha forbesi Temelucha fuscalata Temelucha inflata Temelucha platynotae Temelucha undulata Temelucha angulosa Dasch, 1979 Trathala Cremastus flavopictus Dasch, 1979 Cremastus politus Dasch, 1979 Cremastus recta Temelucha 346 M. D. Schwarzfeld Leafminers Conifer sawflies Loopers Aquatic stem borer Host Family (No. spp.) Guild/Notes (12) Vespidae Hymenoptera: Megachilidae, Sphecidae, Lepidoptera: Elachistidae, Gelechiidae, Pyralidae, (14) Tortricidae Concealed feeders Tortricidae Lepidoptera: Gelechiidae, Gracillariidae, (7) Concealed feeders Coleoptera: Cerambycidae, Cleridae, Curculionidae, Hymenoptera: Cynipidae (7) Trogossitidae; Mostly wood borers None None Amaurobiidae, Linyphiidae; Coleoptera: Araneae: Anthomyiidae; Lepidoptera: Curculionidae; Diptera: (7) Tortricidae Hymenoptera: Braconidae; Lepidoptera: Limacodidae, Noctuidae (5) Hymenoptera: Cimbicidae, Diprionidae, Lepidoptera: Coleophoridae, Tenthredinidae; Incurvariidae, Lasiocampidae, Notodontidae, Zygaenidae (40) Tortricidae, Psychidae, Saturniidae, Hymenoptera: Cimbicidae (1) Hymenoptera: Diprionidae (8) Hymenoptera: Cimbicidae, Diprionidae; Lepidoptera: (14) Tortricidae Erebidae, Noctuidae, Lepidoptera: Crambidae (1) Lepidoptera: Crambidae (2) Grass stem borers None None (1) Tenthredinidae Hymenoptera: None None None None None Hymenoptera: Braconidae, Diprionidae, Ichneumonidae (31) Lepidoptera: Geometridae (2) Lepidoptera: Geometridae, Lasiocampidae, Noctuidae (6) None None None None None None None Neuroptera: Hemerobiidae (1) None Franklin m b Localities White Fox; MB: Indian Head, Snowden, SK: Clemenceau, Elbow, Shilo Ninette, 5 mi SW SK: Indian Head SK: Saskatoon NA; MB: Ninette Wagner AB: AB: Medicine Hat, Edmonton AB: Edmonton AB: Edmonton AB: Waterton Edmonton; SK: Nipawin; MB: Drumheller, AB: Bilby, AB: Edmonton, Drumheller Hamiota MB: Aweme, AB: Edmonton Aweme SK: Pike L, Regina; MB: Wabamun; AB: AB: Wabamun Edmonton, AB: Waterton Edmonton, AB: Wabamun AB: Edmonton; SK: Regina AB: Lethbridge AB: Edmonton AB: Edmonton NA AB: Wagner AB: Edmonton NA AB: Wagner AB: Cochrane Cypress Hills, Drumheller, Chin, Cochrane, Cowley, Calgary, AB: Bilby, St. Paul, Lethbridge, Onefour, Edmonton, Glenwood, High River, Regina, Moose Jaw, W SK: 10 mi Welling; Waterton, Wabamun, Teulon Bunch; MB: Bird's Hill, Willow Saskatoon, AB: Lethbridge Rush L NA; SK: Redberry, Wagner AB: NA AB: Wagner NA AB: Wagner NA AB: Wagner NA; SK: Regina Wagner AB: NA Wagner AB: Wabamun, SK: Lloydminster NP) AB: Cypress Hills, Red Rock Canyon (Waterton 55,H 57,N 2,57,N 2,N 14,H 57,H 14,H 54,55,H 55,N 55,N

5,55,H 55,N 2,N 55,N 5,55,N 55,N 14,56,N 7,52,N 5,N 52,N 52,N 56,N (Walkley, (Walkley, 56,N 2,52,N Townes, 1983 Townes, (Cushman, (Fabricius, Forster, 1876 Forster, (Cresson, 1868) (Ashmead, 1890) (Cook & Davis, (Norton, 1869) Townes, 1983 Townes, Dasch, 1979 (Davis, 1897) Townes, 1962 Townes, (Provancher, 1875) (Provancher, (Cresson, 1872) (Gravenhorst, 1807) Townes, 1962 Townes, (Cushman, 1927) a 5,N 9,55,H 5,55,N 14,N 2,57,N 2,N 2,57,N 5,55,N Trathala delicata Trathala abbreviatus Agrothereutes 1794) 1925) Apsilops sericatus (Viereck, SUBFAMILY Species *Aptesis gracilis 2,N 2,N 1902) Trathala enigmatica Trathala granulata (Davis, 1898) Trathala Dasch, 1979 oregona Trathala CRYPTINAE Aclastus flagellatus Aclastus micator nigricapitata Acrolyta 1891) 1824) stylator (Thunberg, Acroricnus cimbicivorus Agrothereutes lophyri Agrothereutes mandator (Linnaeus, 1758) Agrothereutes Apsilops bicolor *Aptesis fastigata 1877) Aptesis segnis (Provancher, 1962) (Townes, Aritranis freemani Atractodes angustipennis Torre, (Dalla Atractodes brevissimus Bathythrix eurypyga Bathythrix sericea Bathythrix triangularis Buathra dorsicarinata (Pratt, 1945) 1962) Caenocryptus erasus (Townes, (Townes, Ceratophygadeuon brevacus 1944) 1983 Ceratophygadeuon crassidens Townes, Ceratophygadeuon limatulus (Provancher, Ceratophygadeuon perditus 1886) Ceratophygadeuon provancheri 1983 Ceratophygadeuon rugifer Townes, Charitopes mellicornis (Cresson, *Compsocryptus resolutus 1879) 1924) 1958) (Thunberg, 1824) Buathra laborator (Thunberg, Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 347 Leafminers Conifer sawflies Loopers Aquatic stem borer Host Family (No. spp.) Guild/Notes (12) Vespidae Hymenoptera: Megachilidae, Sphecidae, Lepidoptera: Elachistidae, Gelechiidae, Pyralidae, (14) Tortricidae Concealed feeders Tortricidae Lepidoptera: Gelechiidae, Gracillariidae, (7) Concealed feeders Coleoptera: Cerambycidae, Cleridae, Curculionidae, Hymenoptera: Cynipidae (7) Trogossitidae; Mostly wood borers None None Amaurobiidae, Linyphiidae; Coleoptera: Araneae: Anthomyiidae; Lepidoptera: Curculionidae; Diptera: (7) Tortricidae Hymenoptera: Braconidae; Lepidoptera: Limacodidae, Noctuidae (5) Hymenoptera: Cimbicidae, Diprionidae, Lepidoptera: Coleophoridae, Tenthredinidae; Incurvariidae, Lasiocampidae, Notodontidae, Zygaenidae (40) Tortricidae, Psychidae, Saturniidae, Hymenoptera: Cimbicidae (1) Hymenoptera: Diprionidae (8) Hymenoptera: Cimbicidae, Diprionidae; Lepidoptera: (14) Tortricidae Erebidae, Noctuidae, Lepidoptera: Crambidae (1) Lepidoptera: Crambidae (2) Grass stem borers None None (1) Tenthredinidae Hymenoptera: None None None None None Hymenoptera: Braconidae, Diprionidae, Ichneumonidae (31) Lepidoptera: Geometridae (2) Lepidoptera: Geometridae, Lasiocampidae, Noctuidae (6) None None None None None None None Neuroptera: Hemerobiidae (1) None Franklin m b Localities White Fox; MB: Indian Head, Snowden, SK: Clemenceau, Elbow, Shilo Ninette, 5 mi SW SK: Indian Head SK: Saskatoon NA; MB: Ninette Wagner AB: AB: Medicine Hat, Edmonton AB: Edmonton AB: Edmonton AB: Waterton Edmonton; SK: Nipawin; MB: Drumheller, AB: Bilby, AB: Edmonton, Drumheller Hamiota MB: Aweme, AB: Edmonton Aweme SK: Pike L, Regina; MB: Wabamun; AB: AB: Wabamun Edmonton, AB: Waterton Edmonton, AB: Wabamun AB: Edmonton; SK: Regina AB: Lethbridge AB: Edmonton AB: Edmonton NA AB: Wagner AB: Edmonton NA AB: Wagner AB: Cochrane Cypress Hills, Drumheller, Chin, Cochrane, Cowley, Calgary, AB: Bilby, St. Paul, Lethbridge, Onefour, Edmonton, Glenwood, High River, Regina, Moose Jaw, W SK: 10 mi Welling; Waterton, Wabamun, Teulon Bunch; MB: Bird's Hill, Willow Saskatoon, AB: Lethbridge Rush L NA; SK: Redberry, Wagner AB: NA AB: Wagner NA AB: Wagner NA AB: Wagner NA; SK: Regina Wagner AB: NA Wagner AB: Wabamun, SK: Lloydminster NP) AB: Cypress Hills, Red Rock Canyon (Waterton 55,H 57,N 2,57,N 2,N 14,H 57,H 14,H 54,55,H 55,N 55,N

5,55,H 55,N 2,N 55,N 5,55,N 55,N 14,56,N 7,52,N 5,N 52,N 52,N 56,N (Walkley, (Walkley, 56,N 2,52,N Townes, 1983 Townes, (Cushman, (Fabricius, Forster, 1876 Forster, (Cresson, 1868) (Ashmead, 1890) (Cook & Davis, (Norton, 1869) Townes, 1983 Townes, Dasch, 1979 (Davis, 1897) Townes, 1962 Townes, (Provancher, 1875) (Provancher, (Cresson, 1872) (Gravenhorst, 1807) Townes, 1962 Townes, (Cushman, 1927) a 5,N 9,55,H 5,55,N 14,N 2,57,N 2,N 2,57,N 5,55,N Trathala delicata Trathala abbreviatus Agrothereutes 1794) 1925) Apsilops sericatus (Viereck, SUBFAMILY Species *Aptesis gracilis 2,N 2,N 1902) Trathala enigmatica Trathala granulata (Davis, 1898) Trathala Dasch, 1979 oregona Trathala CRYPTINAE Aclastus flagellatus Aclastus micator nigricapitata Acrolyta 1891) 1824) stylator (Thunberg, Acroricnus cimbicivorus Agrothereutes lophyri Agrothereutes mandator (Linnaeus, 1758) Agrothereutes Apsilops bicolor *Aptesis fastigata 1877) Aptesis segnis (Provancher, 1962) (Townes, Aritranis freemani Atractodes angustipennis Torre, (Dalla Atractodes brevissimus Bathythrix eurypyga Bathythrix sericea Bathythrix triangularis Buathra dorsicarinata (Pratt, 1945) 1962) Caenocryptus erasus (Townes, (Townes, Ceratophygadeuon brevacus 1944) 1983 Ceratophygadeuon crassidens Townes, Ceratophygadeuon limatulus (Provancher, Ceratophygadeuon perditus 1886) Ceratophygadeuon provancheri 1983 Ceratophygadeuon rugifer Townes, Charitopes mellicornis (Cresson, *Compsocryptus resolutus 1879) 1924) 1958) (Thunberg, 1824) Buathra laborator (Thunberg, 348 M. D. Schwarzfeld Borers Stem borer Host Family (No. spp.) Guild/Notes None None None None Lepidoptera: Crambidae, Erebidae, Noctuidae, (12) Tortricidae Nymphalidae, Pyralidae, Sesiidae, Hymenoptera: Cimbicidae (2) None None None None Tortricidae Lepidoptera: Tenthredinidae; Hymenoptera: (2) None Coleoptera: Buprestidae (1) None None None None None None None Coleoptera: Curculionidae (1) None Coleoptera: Cerambycidae, Curculionidae Hymenoptera: Crabronidae (4) None Neuroptera: Chrysopidae, Hemerobiidae Lepidoptera: (9) Tortricidae Geometridae, None (1) Tachinidae Diptera: Lepidoptera: Sesiidae (1) Borers Coleoptera: Cerambycidae (2) None None None None Argidae Hymenoptera: (2) (1) Tenthredinidae Hymenoptera: (5) Tenthredinidae Hymenoptera: Diprionidae, Mostly conifer sawflies None None None

Sanford , Winnipeg Swan River Franklin m m m m b Localities Duchess, Edmonton, Cassils, Chin, Drumheller, AB: Brooks, Calgary, Elkwater L, Lethbridge, Manyberries, Medicine Hat, Elkwater, Lethbridge; SK: Love High River, AB: Calgary, SK: Big Waterton; Taber, AB: Edmonton, Gleichen, Lethbridge, Saskatoon Muddy, AB: Waterton AB: Cypress Hills, Edmonton, Irvine, Medicine Hat; SK: Estevan, Transcona Aweme, MB: Weyburn; Regina, Secretan, Uren, Aweme MB: Weyburn; AB: Lethbridge; SK: Minton, AB: Edmonton, Red Deer; MB: AB: Bilby AB: Edmonton, Elkwater L; MB: Shilo W MB: 5 mi Bunch SK: Willow AB: Edmonton AB: Bilby SK: Sutherland AB: Pincher AB: Edmonton; SK: Saskatoon Edmonton AB: Wabamun, MB: McCreary AB: Edmonton AB: Vermilion AB: Lethbridge; SK: Moosomin; MB: NA AB: Wagner NA; SK: Lloydminster Wagner AB: Lakes NP; Waterton AB: Calgary, Aweme Lakes NP; MB: Waterton AB: Edmonton, AB: Elkwater at Cut Knife Atton's L SK: AB: Waterton AB: nr Edmonton AB: Pincher AB: approx Cypress Hills SK: nr Regina AB: nr Edmonton AB: Elkwater P nr Edmonton, Elkwater; SK: Cypress Hills AB: Drumheller, AB: nr Edmonton NA AB: Wagner NA AB: Wagner Scandia, Waterton, Welling; SK: Dodsland, Maple Cr, Qu'Appelle, SK: Dodsland, Maple Cr, Welling; Waterton, Scandia, Bunch, Rosthern, Willow White Fox, Regina, Rockglen, Saskatoon, Franklin , Souris Spruce Woods, Aweme, Swift Current; MB: Teulon Transcona, AB: Edmonton 14,N 56,N 14,58,N 5,9,57,N 5,58,N 58,N 5,9,56,N 58,N 56,N 58,N 58,N 2,N 56,N 58,N 56,N 58,N 2,N 5,9,55,N,T 14,55,N 58,N 2,N 58,N 55,H 58,N 14,55,N 5,55,N 55,N 55,N 14,55,N (Fahringer, (Fahringer, (Provancher, (Provancher, Townes & Gupta, Townes Townes, 1962 Townes, (Cresson, Townes & Gupta, Townes (Dalla Torre, 1902) Torre, (Dalla (Provancher, (Provancher, Townes, 1944 Townes, Townes, 1962 Townes, (Cresson, 1864) Townes, 1962 Townes, (Provancher, 1882) (Provancher, Luhman, 1990 (Dalla Torre, 1902) Torre, (Dalla Davis, 1895 a 9,14,56,N 56,N 5,56,N 2,57,H 56,N 56,N 56,N Cryptus albitarsis (Cresson, 1864) Endasys hexamerus Luhman, 1990 Endasys monticola 1879) (Provancher, Endasys mucronatus 1983 dorsatus Townes, Ethelurgus SUBFAMILY Species 56,N Cubocephalus occidentalis 1875) 126,H 1877) Cryptus luctuosus Cresson, 1864 *Cryptus minimus Pratt, 1945 Cryptus mutatus Pratt, 1945 *Cryptus persimilis Cresson, 1864 Cryptus ruralis Pratt, 1945 Cubocephalus alacris Cubocephalus annectus Cubocephalus annulatus 1864) Cubocephalus atriclunis 1962 Cubocephalus baldaufii Cubocephalus contractus Cubocephalus dreisbachi Cubocephalus hebes Cubocephalus hirtipes Cubocephalus inhabilis 1865) Demopheles corruptor (Taschenberg, Diaglyptidea lavoiei 1882) crassa (Provancher, Dichrogaster 1983 nigriceps Townes, Dichrogaster schimitscheki Dichrogaster 1935) Echthrus adillae Echthrus niger Cresson, 1868 Endasys aurigena Luhman, 1990 Luhman, 1990 Endasys bicolorescens Endasys chrysoleptus Luhman, 1990 Endasys michiganensis Luhman, 1990 Endasys nemati Luhman, 1990 1874) Endasys pubescens (Provancher, Endasys xanthostomus Luhman, 1990 1983 57,N Townes, fuscidens Ethelurgus 1983 opacus Townes, Ethelurgus 1962 Townes & Gupta, Townes Cubocephalus prolixus 1962 Endasys aurantifex Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 349 Borers Stem borer Host Family (No. spp.) Guild/Notes None None None None Lepidoptera: Crambidae, Erebidae, Noctuidae, (12) Tortricidae Nymphalidae, Pyralidae, Sesiidae, Hymenoptera: Cimbicidae (2) None None None None Tortricidae Lepidoptera: Tenthredinidae; Hymenoptera: (2) None Coleoptera: Buprestidae (1) None None None None None None None Coleoptera: Curculionidae (1) None Coleoptera: Cerambycidae, Curculionidae Hymenoptera: Crabronidae (4) None Neuroptera: Chrysopidae, Hemerobiidae Lepidoptera: (9) Tortricidae Geometridae, None (1) Tachinidae Diptera: Lepidoptera: Sesiidae (1) Borers Coleoptera: Cerambycidae (2) None None None None Argidae Hymenoptera: (2) (1) Tenthredinidae Hymenoptera: (5) Tenthredinidae Hymenoptera: Diprionidae, Mostly conifer sawflies None None None

Sanford , Winnipeg Swan River Franklin m m m m b Localities Duchess, Edmonton, Cassils, Chin, Drumheller, AB: Brooks, Calgary, Elkwater L, Lethbridge, Manyberries, Medicine Hat, Elkwater, Lethbridge; SK: Love High River, AB: Calgary, SK: Big Waterton; Taber, AB: Edmonton, Gleichen, Lethbridge, Saskatoon Muddy, AB: Waterton AB: Cypress Hills, Edmonton, Irvine, Medicine Hat; SK: Estevan, Transcona Aweme, MB: Weyburn; Regina, Secretan, Uren, Aweme MB: Weyburn; AB: Lethbridge; SK: Minton, AB: Edmonton, Red Deer; MB: AB: Bilby AB: Edmonton, Elkwater L; MB: Shilo W MB: 5 mi Bunch SK: Willow AB: Edmonton AB: Bilby SK: Sutherland AB: Pincher AB: Edmonton; SK: Saskatoon Edmonton AB: Wabamun, MB: McCreary AB: Edmonton AB: Vermilion AB: Lethbridge; SK: Moosomin; MB: NA AB: Wagner NA; SK: Lloydminster Wagner AB: Lakes NP; Waterton AB: Calgary, Aweme Lakes NP; MB: Waterton AB: Edmonton, AB: Elkwater at Cut Knife Atton's L SK: AB: Waterton AB: nr Edmonton AB: Pincher AB: approx Cypress Hills SK: nr Regina AB: nr Edmonton AB: Elkwater P nr Edmonton, Elkwater; SK: Cypress Hills AB: Drumheller, AB: nr Edmonton NA AB: Wagner NA AB: Wagner Scandia, Waterton, Welling; SK: Dodsland, Maple Cr, Qu'Appelle, SK: Dodsland, Maple Cr, Welling; Waterton, Scandia, Bunch, Rosthern, Willow White Fox, Regina, Rockglen, Saskatoon, Franklin , Souris Spruce Woods, Aweme, Swift Current; MB: Teulon Transcona, AB: Edmonton 14,N 56,N 14,58,N 5,9,57,N 5,58,N 58,N 5,9,56,N 58,N 56,N 58,N 58,N 2,N 56,N 58,N 56,N 58,N 2,N 5,9,55,N,T 14,55,N 58,N 2,N 58,N 55,H 58,N 14,55,N 5,55,N 55,N 55,N 14,55,N (Fahringer, (Fahringer, (Provancher, (Provancher, Townes & Gupta, Townes Townes, 1962 Townes, (Cresson, Townes & Gupta, Townes (Dalla Torre, 1902) Torre, (Dalla (Provancher, (Provancher, Townes, 1944 Townes, Townes, 1962 Townes, (Cresson, 1864) Townes, 1962 Townes, (Provancher, 1882) (Provancher, Luhman, 1990 (Dalla Torre, 1902) Torre, (Dalla Davis, 1895 a 9,14,56,N 56,N 5,56,N 2,57,H 56,N 56,N 56,N Cryptus albitarsis (Cresson, 1864) Endasys hexamerus Luhman, 1990 Endasys monticola 1879) (Provancher, Endasys mucronatus 1983 dorsatus Townes, Ethelurgus SUBFAMILY Species 56,N Cubocephalus occidentalis 1875) 126,H 1877) Cryptus luctuosus Cresson, 1864 *Cryptus minimus Pratt, 1945 Cryptus mutatus Pratt, 1945 *Cryptus persimilis Cresson, 1864 Cryptus ruralis Pratt, 1945 Cubocephalus alacris Cubocephalus annectus Cubocephalus annulatus 1864) Cubocephalus atriclunis 1962 Cubocephalus baldaufii Cubocephalus contractus Cubocephalus dreisbachi Cubocephalus hebes Cubocephalus hirtipes Cubocephalus inhabilis 1865) Demopheles corruptor (Taschenberg, Diaglyptidea lavoiei 1882) crassa (Provancher, Dichrogaster 1983 nigriceps Townes, Dichrogaster schimitscheki Dichrogaster 1935) Echthrus adillae Echthrus niger Cresson, 1868 Endasys aurigena Luhman, 1990 Luhman, 1990 Endasys bicolorescens Endasys chrysoleptus Luhman, 1990 Endasys michiganensis Luhman, 1990 Endasys nemati Luhman, 1990 1874) Endasys pubescens (Provancher, Endasys xanthostomus Luhman, 1990 1983 57,N Townes, fuscidens Ethelurgus 1983 opacus Townes, Ethelurgus 1962 Townes & Gupta, Townes Cubocephalus prolixus 1962 Endasys aurantifex 350 M. D. Schwarzfeld Borers Host Family (No. spp.) Guild/Notes Araneae: Gnaphosidae (1) None Coleoptera: Curculionidae; Lepidoptera: Erebidae, Hesperiidae, Lasiocampidae, Noctuidae, Pieridae, (13) Tortricidae Saturniidae, Coleoptera: Curculionidae; Hymenoptera: Lepidoptera: Erebidae, Tenthredinidae; Ichneumonidae, Gelechiidae, Hesperiidae, Lasiocampidae, Noctuidae, (32) Tortricidae Pieridae, Psychididae, Pyralidae, Hymenoptera: Braconidae, Diprionidae, Ichneumonidae, Tenthredinidae; Pteromalidae, Lepidoptera: Bucculatricidae, Coleophoridae, Cosmopterigidae, Erebidae, Lasiocampidae, Notodontidae, Nymphalidae, Papilionidae, Pyralidae, Neuroptera: Yponomeutidae; Saturniidae, Tortricidae, Chrysopidae (64) None None Coleoptera: Cerambycidae (1) None Lepidoptera: Crambidae (1) Stem borer Hymenoptera: Megachilidae (2) None Vespidae Hymenoptera: Tenthredinidae, Megachilidae, (6) Coleoptera: Cerambycidae; Araneae: Lycosidae; Hymenoptera: Diprionidae, Lepidoptera: Erebidae (7) Hymenoptera: Ichneumonidae; Lepidoptera: Elachistidae, Erebidae, Geometridae, Lasiocampidae, (21) Tortricidae Mostly concealed feeders None (1) Tortricidae Lepidoptera: Crambidae, Concealed feeders None Lepidoptera: Notodontidae (1) Hymenoptera: Braconidae Lepidoptera: Tortricidae Cosmopterigidae, Erebidae, Gracillaridae, (11) Hymenoptera: Braconidae (2) Hymenoptera: Braconidae, Diprionidae, Ichneumonidae; Lepidoptera: Gelechiidae, (19) Tortricidae Geometridae, Notodontidae, Hymenoptera: Braconidae, Diprionidae, Tortricidae Lepidoptera: Geometridae, Tenthredinidae; (15) None Agromyzidae, Muscidae; Hymenoptera: Diptera: (9) Tineidae Cynipidae; Lepidoptera: Plutellidae, None Anthomyiidae, Diptera: Hymenoptera: Tachinidae; Ichneumonidae; Lepidoptera: Noctuidae, Nymphalidae, (7) Tortricidae Lepidoptera: Notodontidae, Pieridae (2) None Hymenoptera: Sphecidae; Lepidoptera: Saturniidae (3)

, Gnadenthal Glenlea b Localities SK: Dundurn, Redberry (near Cut Attons L parkland region; SK: Waterton, AB: Edmonton, Swan River , Teulon Aweme, Knife), Pike L, 5 mi E Swift Current; MB: MB: Franklin AB: Lethbridge; MB: MB: Swan River , Winnipeg SK: Grenfell AB: Edmonton AB: Edmonton NA AB: Wagner MB: Minnedosa AB: Clymont, Lethbridge White Fox AB: Edmonton; SK: AB: Edmonton, Lethbridge Aweme at Cut Knife, Pickthall; MB: Attons L AB: Cypress Hills; SK: Saint Paul, Consort, Edmonton, Lacombe, Red Deer, AB: Calgary, Lakes NP; SK: Glaslyn, Kenosee L, Waterton Sylvan L, Saskatoon; MB: Aweme AB: Edmonton Waterton AB: Edmonton, Medicine Hat, AB: Waterton MB: Aweme NA Wagner AB: Edmonton, NA AB: Wagner AB: Cochrane, Lethbridge, Edmonton FR Woods MB: Spruce Regina AB: Cooking L; SK: Redberry, NA AB: Wagner NA AB: Wagner NA AB: Wagner Douglas Aweme, AB: Edmonton; MB: Vermilion, Consort, Cypress Hills, AB: Blackfoot Hills, Calgary, Saskatchewan Landing, Saskatoon SK: Great Deer, Waterton; Lakes Waterton Waterton, Cypress Hills, AB: Calgary, 2,H,O 55,N 2,5,N 59,N 55,N 5,55,N 5,57,N 55,N 14,55,N 5,N 2,H 55,N 55,N 5,55,H,O 5,55,H 2,N 9,N,T,Oc 2,N 14,55,N 57,N 60,N 2,H (Cresson, 9,N,T,Oc 5,9,N (Gravenhorst, (Provancher, (Provancher, Townes, 1983 Townes, (Townes, 1962) (Townes, Mason, 1968 (Gravenhorst, 1807) (Say, 1835) (Say, Townes, 1983 Townes, a 5,9,46,55,N 5,N 5,55,N 56,H 5,55,H,O (Provancher, 1874) Gambrus apicatus (Provancher, Gelis tenellus SUBFAMILY Species 1776) Ischnus inquisitorius (Müller, 5,55,N (Provancher, Gambrus canadensis (Provancher, 1875) Gambrus ultimus (Cresson, 1864) 1892) Gelis drassi (Riley, Glyphicnemis californica (Cresson, 1879) Glyphicnemis mandibularis Helcostizus yukonensis (Ashmead, 1890) Hoplocryptus albicollaris (Cresson, 1872) Hoplocryptus imitator 1877) Hoplocryptus linnae 1874) Hoplocryptus notatus (Provancher, Idiolispa analis 1874) Ischnus latus (Provancher, albomaculatus (Cresson, 1864) Listrognathus 1962 Townes, nigrescens Listrognathus paludatus (Cresson, 1872) Listrognathus 1875) mandibularis (Provancher, Lysibia tenax Lysibia 1886) Mastrus aciculatus (Provancher, Mastrus laplantei 1983 latus Townes, Medophron Megacara hortulana (Gravenhorst, 1829) 1983 Townes, Megacara impressa Megacara vagans (Gravenhorst, 1829) Megaplectes monticola 1829) Meringopus asymmetricus (Pratt, 1945) Meringopus calescens (Gravenhorst, 1829) 1864) Hemiteles amboniger Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 351 Borers Host Family (No. spp.) Guild/Notes Araneae: Gnaphosidae (1) None Coleoptera: Curculionidae; Lepidoptera: Erebidae, Hesperiidae, Lasiocampidae, Noctuidae, Pieridae, (13) Tortricidae Saturniidae, Coleoptera: Curculionidae; Hymenoptera: Lepidoptera: Erebidae, Tenthredinidae; Ichneumonidae, Gelechiidae, Hesperiidae, Lasiocampidae, Noctuidae, (32) Tortricidae Pieridae, Psychididae, Pyralidae, Hymenoptera: Braconidae, Diprionidae, Ichneumonidae, Tenthredinidae; Pteromalidae, Lepidoptera: Bucculatricidae, Coleophoridae, Cosmopterigidae, Erebidae, Lasiocampidae, Notodontidae, Nymphalidae, Papilionidae, Pyralidae, Neuroptera: Yponomeutidae; Saturniidae, Tortricidae, Chrysopidae (64) None None Coleoptera: Cerambycidae (1) None Lepidoptera: Crambidae (1) Stem borer Hymenoptera: Megachilidae (2) None Vespidae Hymenoptera: Tenthredinidae, Megachilidae, (6) Coleoptera: Cerambycidae; Araneae: Lycosidae; Hymenoptera: Diprionidae, Lepidoptera: Erebidae (7) Hymenoptera: Ichneumonidae; Lepidoptera: Elachistidae, Erebidae, Geometridae, Lasiocampidae, (21) Tortricidae Mostly concealed feeders None (1) Tortricidae Lepidoptera: Crambidae, Concealed feeders None Lepidoptera: Notodontidae (1) Hymenoptera: Braconidae Lepidoptera: Tortricidae Cosmopterigidae, Erebidae, Gracillaridae, (11) Hymenoptera: Braconidae (2) Hymenoptera: Braconidae, Diprionidae, Ichneumonidae; Lepidoptera: Gelechiidae, (19) Tortricidae Geometridae, Notodontidae, Hymenoptera: Braconidae, Diprionidae, Tortricidae Lepidoptera: Geometridae, Tenthredinidae; (15) None Agromyzidae, Muscidae; Hymenoptera: Diptera: (9) Tineidae Cynipidae; Lepidoptera: Plutellidae, None Anthomyiidae, Diptera: Hymenoptera: Tachinidae; Ichneumonidae; Lepidoptera: Noctuidae, Nymphalidae, (7) Tortricidae Lepidoptera: Notodontidae, Pieridae (2) None Hymenoptera: Sphecidae; Lepidoptera: Saturniidae (3)

, Gnadenthal Glenlea b Localities SK: Dundurn, Redberry (near Cut Attons L parkland region; SK: Waterton, AB: Edmonton, Swan River , Teulon Aweme, Knife), Pike L, 5 mi E Swift Current; MB: MB: Franklin AB: Lethbridge; MB: MB: Swan River , Winnipeg SK: Grenfell AB: Edmonton AB: Edmonton NA AB: Wagner MB: Minnedosa AB: Clymont, Lethbridge White Fox AB: Edmonton; SK: AB: Edmonton, Lethbridge Aweme at Cut Knife, Pickthall; MB: Attons L AB: Cypress Hills; SK: Saint Paul, Consort, Edmonton, Lacombe, Red Deer, AB: Calgary, Lakes NP; SK: Glaslyn, Kenosee L, Waterton Sylvan L, Saskatoon; MB: Aweme AB: Edmonton Waterton AB: Edmonton, Medicine Hat, AB: Waterton MB: Aweme NA Wagner AB: Edmonton, NA AB: Wagner AB: Cochrane, Lethbridge, Edmonton FR Woods MB: Spruce Regina AB: Cooking L; SK: Redberry, NA AB: Wagner NA AB: Wagner NA AB: Wagner Douglas Aweme, AB: Edmonton; MB: Vermilion, Consort, Cypress Hills, AB: Blackfoot Hills, Calgary, Saskatchewan Landing, Saskatoon SK: Great Deer, Waterton; Lakes Waterton Waterton, Cypress Hills, AB: Calgary, 2,H,O 55,N 2,5,N 59,N 55,N 5,55,N 5,57,N 55,N 14,55,N 5,N 2,H 55,N 55,N 5,55,H,O 5,55,H 2,N 9,N,T,Oc 2,N 14,55,N 57,N 60,N 2,H (Cresson, 9,N,T,Oc 5,9,N (Gravenhorst, (Provancher, (Provancher, Townes, 1983 Townes, (Townes, 1962) (Townes, Mason, 1968 (Gravenhorst, 1807) (Say, 1835) (Say, Townes, 1983 Townes, a 5,9,46,55,N 5,N 5,55,N 56,H 5,55,H,O (Provancher, 1874) Gambrus apicatus (Provancher, Gelis tenellus SUBFAMILY Species 1776) Ischnus inquisitorius (Müller, 5,55,N (Provancher, Gambrus canadensis (Provancher, 1875) Gambrus ultimus (Cresson, 1864) 1892) Gelis drassi (Riley, Glyphicnemis californica (Cresson, 1879) Glyphicnemis mandibularis Helcostizus yukonensis (Ashmead, 1890) Hoplocryptus albicollaris (Cresson, 1872) Hoplocryptus imitator 1877) Hoplocryptus linnae 1874) Hoplocryptus notatus (Provancher, Idiolispa analis 1874) Ischnus latus (Provancher, albomaculatus (Cresson, 1864) Listrognathus 1962 Townes, nigrescens Listrognathus paludatus (Cresson, 1872) Listrognathus 1875) mandibularis (Provancher, Lysibia tenax Lysibia 1886) Mastrus aciculatus (Provancher, Mastrus laplantei 1983 latus Townes, Medophron Megacara hortulana (Gravenhorst, 1829) 1983 Townes, Megacara impressa Megacara vagans (Gravenhorst, 1829) Megaplectes monticola 1829) Meringopus asymmetricus (Pratt, 1945) Meringopus calescens (Gravenhorst, 1829) 1864) Hemiteles amboniger 352 M. D. Schwarzfeld Setose Concealed feeders Host Family (No. spp.) Guild/Notes Lepidoptera: Lasiocampidae (2) Lepidoptera: Sesiidae (1) Borers Lepidoptera: Noctuidae, Sesiidae (2) Diptera: Sciomyzidae (3) Aquatic snail parasitoids Lepidoptera: Pyralidae (1) (13) Tortricidae Lepidoptera: Pyralidae, Concealed feeders None None None Lepidoptera: Crambidae, Pyralidae (12) Concealed feeders Hymenoptera: Pompilidae (1) (1) Tenthredinidae Hymenoptera: (1) Tenthredinidae Hymenoptera: Hymenoptera: Diprionidae (1) (7) Vespidae Hymenoptera: (1) Tenthredinidae Hymenoptera: (4) Tenthredinidae Hymenoptera: Cynipidae, (1) Tortricidae Lepidoptera: Anthomyiidae, Coleoptera: Cerambycidae; Diptera: Muscidae; Lepidoptera: Erebidae, Noctuidae (8) None (4) Tachinidae Diptera: None (15) Tenthredinidae Hymenoptera: Diprionidae, None None None None Lepidoptera: Erebidae (1) Diptera: Syrphidae (2) Hymenoptera: Cynipidae; Lepidoptera: Zygaenidae (3) None Lepidoptera: Pyralidae (14) Vespidae; Hymenoptera: Mostly Vespidae Anthomyiidae, Fanniidae (3) Diptera: Anthomyiidae Diptera: (1) Thomisidae (1) Araneae: None None None Lepidoptera: Gelechiidae, Geometridae, Tineidae (5) Tenthredinidae; Hymenoptera: Braconidae, Cynipidae, Tineidae, Lepidoptera: Elachistidae Psychidae, Neuroptera: Chrysopidae (10) Tortricidae; None Coleoptera: Buprestidae, Cerambycidae (3) Coleoptera: Buprestidae, Cerambycidae (5)

, Swan River Swan River , Seven , Narcisse La Broquerie Birtle Swan River ; midcentral Canada (AB/SK) in b Localities Lakes: Waterton Waterton, AB: Cochrane, Foremost, Lethbridge, Waterton Foremost, AB: Calgary, AB: Lethbridge AB: Edmonton AB: Edmonton AB: Lethbridge; MB: sunflower fields SK: Pickthall AB: Calgary AB: Lethbridge Aweme AB: Edmonton; MB: AB: Medicine Hat Tofield AB: Edmonton, MB: Aweme AB: Edmonton AB: Medicine Hat AB: Edmonton AB: Lethbridge; SK: Cut Knife; MB: AB: Edmonton Glenlea, Grunthal, MB: Beausejour, MB: Glenlea AB: Lethbridge, Edmonton; MB: AB: Edmonton AB: Albert Prince Waterton AB: Wabamun, Edmonton, NA AB: Wagner SK: Antelope L AB: Edmonton, Gull L NA AB: Wagner NA; SK: Regina Wagner Wabamun, Edmonton, Saint Paul, AB: Bilby, AB: Edmonton AB: Wabamun NA AB: Wagner AB: Lethbridge NA AB: Wagner AB: Cypress Hills, Edmonton, Oldman River nr Lethbridge AB: Onefour Aweme MB: Waldheim; SK: Wabamun; AB: Edmonton, Lethbridge, AB: Edmonton AB: Edmonton SK: Lloydminster AB: Duchess AB: Calgary SK: Saskatoon SK: Katepwa L nr Indian Head SK: Katepwa L Kleefeld Winnipeg, Sisters , Steinbach Whitemouth, 2,H 5,N 57,H 2,N 5,55,N 5,55,N 55,N 14,N 5,9,56,H 14,N 2,H 55,N 14,56,N 55,N 2,H,Oc 5,55,N 5,55,N 5,9,N 9,N ) 57,N 5,56,N 5,55,N 56,N 56,N 5,8,9,N,T 55,N,T 56,N ) 5,H,T 55,N,T 57,N 5,W 55,N 56,N 14,55,N 55,N 55,N (Brischke, 1881) Gravenhorst, (Provancher, (Provancher, (Provancher, (Provancher, (Ashmead, 1890) (Provancher, (Provancher, Provancher, 1882 Provancher, (Cresson, 1879) (Cresson, 1864) (Gravenhorst, Townes, 1962 Townes, (Cresson, 1879) Gravenhorst, Townes, 1983 Townes, (Cresson, 1879) (Provancher, 1886) (Provancher, (Cresson, 1864) (Gmelin, 1970) (Provancher, 1875) (Provancher, Townes, 1962 Townes, Townes, 1962 Townes, (Gravenhorst, 1807 a 14,56,N 56,N 5,56,N 2,5,56,N,Oc 14,56,H 55,H 9,61,62,H Meringopus genatus (Pratt, 1945) Meringopus relativus SUBFAMILY Species Phygadeuon subfuscus Cresson, 1864 Meringopus tejonensis Mesoleptus declivus *Mesostenus albinotatus 1829 *Mesostenus gracilis Cresson, 1864 1962 Mesostenus liogaster Townes, Mesostenus melanurus Cushman, 1929 1962 *Mesostenus sicarius Townes, Mesostenus thoracicus Cresson, 1864 Messatoporus rufiventris Cushman, 1929 albicoxus Oresbius 1962) parallelus (Townes, Oresbius tsugae (Cushman, 1939) Oresbius Pachysomoides fulvus Parmortha circumcincta 1879) Parmortha parvula (Gravenhorst, 1829) Phygadeuon aciculatus Phygadeuon fumator 1829 1875 Phygadeuon ovalis Provancher, Pleolophus coriaceus Pleolophus indistinctus 1886) Pleolophus rubrocinctus 1874) Polyaulon bimaculatus (Ashmead, 1890 Polyaulon erythropa *Polytribax crotchii Pygocryptus brevicornis Rhembobius abdominalis (Provancher, 1874) Schenkia graminicola 1829) Schenkia iridescens Sphecophaga vesparum (Curtis, 1828) Stilpnus gagates Sulcarius nigricornis (Thomson, 1884) cyperia Trychosis latidens Trychosis 1877) montivaga (Provancher, Trychosis 1944 semirubra Townes, Trychosis Xenolytus bitinctus Xiphulcus floricolator (Gravenhorst, 1807) Xiphulcus parallelus Xylophrurus bicolor Cushman, 1919 Xylophrurus nubilipennis (Cresson, 1864) Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 353 Setose Concealed feeders Host Family (No. spp.) Guild/Notes Lepidoptera: Lasiocampidae (2) Lepidoptera: Sesiidae (1) Borers Lepidoptera: Noctuidae, Sesiidae (2) Diptera: Sciomyzidae (3) Aquatic snail parasitoids Lepidoptera: Pyralidae (1) (13) Tortricidae Lepidoptera: Pyralidae, Concealed feeders None None None Lepidoptera: Crambidae, Pyralidae (12) Concealed feeders Hymenoptera: Pompilidae (1) (1) Tenthredinidae Hymenoptera: (1) Tenthredinidae Hymenoptera: Hymenoptera: Diprionidae (1) (7) Vespidae Hymenoptera: (1) Tenthredinidae Hymenoptera: (4) Tenthredinidae Hymenoptera: Cynipidae, (1) Tortricidae Lepidoptera: Anthomyiidae, Coleoptera: Cerambycidae; Diptera: Muscidae; Lepidoptera: Erebidae, Noctuidae (8) None (4) Tachinidae Diptera: None (15) Tenthredinidae Hymenoptera: Diprionidae, None None None None Lepidoptera: Erebidae (1) Diptera: Syrphidae (2) Hymenoptera: Cynipidae; Lepidoptera: Zygaenidae (3) None Lepidoptera: Pyralidae (14) Vespidae; Hymenoptera: Mostly Vespidae Anthomyiidae, Fanniidae (3) Diptera: Anthomyiidae Diptera: (1) Thomisidae (1) Araneae: None None None Lepidoptera: Gelechiidae, Geometridae, Tineidae (5) Tenthredinidae; Hymenoptera: Braconidae, Cynipidae, Tineidae, Lepidoptera: Elachistidae Psychidae, Neuroptera: Chrysopidae (10) Tortricidae; None Coleoptera: Buprestidae, Cerambycidae (3) Coleoptera: Buprestidae, Cerambycidae (5)

, Swan River Swan River , Seven , Narcisse La Broquerie Birtle Swan River ; midcentral Canada (AB/SK) in b Localities Lakes: Waterton Waterton, AB: Cochrane, Foremost, Lethbridge, Waterton Foremost, AB: Calgary, AB: Lethbridge AB: Edmonton AB: Edmonton AB: Lethbridge; MB: sunflower fields SK: Pickthall AB: Calgary AB: Lethbridge Aweme AB: Edmonton; MB: AB: Medicine Hat Tofield AB: Edmonton, MB: Aweme AB: Edmonton AB: Medicine Hat AB: Edmonton AB: Lethbridge; SK: Cut Knife; MB: AB: Edmonton Glenlea, Grunthal, MB: Beausejour, MB: Glenlea AB: Lethbridge, Edmonton; MB: AB: Edmonton AB: Albert Prince Waterton AB: Wabamun, Edmonton, NA AB: Wagner SK: Antelope L AB: Edmonton, Gull L NA AB: Wagner NA; SK: Regina Wagner Wabamun, Edmonton, Saint Paul, AB: Bilby, AB: Edmonton AB: Wabamun NA AB: Wagner AB: Lethbridge NA AB: Wagner AB: Cypress Hills, Edmonton, Oldman River nr Lethbridge AB: Onefour Aweme MB: Waldheim; SK: Wabamun; AB: Edmonton, Lethbridge, AB: Edmonton AB: Edmonton SK: Lloydminster AB: Duchess AB: Calgary SK: Saskatoon SK: Katepwa L nr Indian Head SK: Katepwa L Kleefeld Winnipeg, Sisters , Steinbach Whitemouth, 2,H 5,N 57,H 2,N 5,55,N 5,55,N 55,N 14,N 5,9,56,H 14,N 2,H 55,N 14,56,N 55,N 2,H,Oc 5,55,N 5,55,N 5,9,N 9,N ) 57,N 5,56,N 5,55,N 56,N 56,N 5,8,9,N,T 55,N,T 56,N ) 5,H,T 55,N,T 57,N 5,W 55,N 56,N 14,55,N 55,N 55,N (Brischke, 1881) Gravenhorst, (Provancher, (Provancher, (Provancher, (Provancher, (Ashmead, 1890) (Provancher, (Provancher, Provancher, 1882 Provancher, (Cresson, 1879) (Cresson, 1864) (Gravenhorst, Townes, 1962 Townes, (Cresson, 1879) Gravenhorst, Townes, 1983 Townes, (Cresson, 1879) (Provancher, 1886) (Provancher, (Cresson, 1864) (Gmelin, 1970) (Provancher, 1875) (Provancher, Townes, 1962 Townes, Townes, 1962 Townes, (Gravenhorst, 1807 a 14,56,N 56,N 5,56,N 2,5,56,N,Oc 14,56,H 55,H 9,61,62,H Meringopus genatus (Pratt, 1945) Meringopus relativus SUBFAMILY Species Phygadeuon subfuscus Cresson, 1864 Meringopus tejonensis Mesoleptus declivus *Mesostenus albinotatus 1829 *Mesostenus gracilis Cresson, 1864 1962 Mesostenus liogaster Townes, Mesostenus melanurus Cushman, 1929 1962 *Mesostenus sicarius Townes, Mesostenus thoracicus Cresson, 1864 Messatoporus rufiventris Cushman, 1929 albicoxus Oresbius 1962) parallelus (Townes, Oresbius tsugae (Cushman, 1939) Oresbius Pachysomoides fulvus Parmortha circumcincta 1879) Parmortha parvula (Gravenhorst, 1829) Phygadeuon aciculatus Phygadeuon fumator 1829 1875 Phygadeuon ovalis Provancher, Pleolophus coriaceus Pleolophus indistinctus 1886) Pleolophus rubrocinctus 1874) Polyaulon bimaculatus (Ashmead, 1890 Polyaulon erythropa *Polytribax crotchii Pygocryptus brevicornis Rhembobius abdominalis (Provancher, 1874) Schenkia graminicola 1829) Schenkia iridescens Sphecophaga vesparum (Curtis, 1828) Stilpnus gagates Sulcarius nigricornis (Thomson, 1884) cyperia Trychosis latidens Trychosis 1877) montivaga (Provancher, Trychosis 1944 semirubra Townes, Trychosis Xenolytus bitinctus Xiphulcus floricolator (Gravenhorst, 1807) Xiphulcus parallelus Xylophrurus bicolor Cushman, 1919 Xylophrurus nubilipennis (Cresson, 1864) 354 M. D. Schwarzfeld Hymenoptera: Diprionidae (1) None None None None Host Family (No. spp.) Guild/Notes Hymenoptera: Cimbicidae (1) None None None None None None None None Hymenoptera: Pamphiliidae (1) None None None None None None None None None None None Hymenoptera: Pamphiliidae (1) Hymenoptera: Pamphiliidae (2) (16) Tenthredinidae Hymenoptera: Diprionidae, None Hymenoptera: Diprionidae (4) None None None None (10) Tenthredinidae Argidae, Hymenoptera: (2) Tenthredinidae Hymenoptera: None None (1) Tenthredinidae Hymenoptera: (1) Tenthredinidae Hymenoptera: None None Hymenoptera: Tenthredinidae (1) Tenthredinidae Hymenoptera: None m , Winnipeg Fawcett , St. Paul Strathclair b AB: Drumheller, George L, Gull L, Jumping Pound Cr (20 mi W W L, Gull Jumping Pound Cr (20 mi George AB: Drumheller, Calgary), Red Deer; SK: Great Sand Hills, Saskatoon AB: Calgary, Edmonton AB: Calgary, AB: Bilby, Calgary, Carstairs, Demmitt, Drumheller, Edmonton, Carstairs, Demmitt, Drumheller, Calgary, AB: Bilby, Waterton Calgary), W L, Jumping Pound Cr (20mi George Elkwater P, Victor, Albert, Rockglen, St. Prince Elbow, NP; SK: Broadview, Saskatoon; MB: Southwest AB: Edmonton; MB: Ninette AB: Red Deer Localities AB: Edmonton AB: Edmonton AB: Edmonton , Red Deer AB: Southwest AB: Edmonton AB: Waterton AB: Edmonton AB: Red Deer AB: Edmonton Mt Turtle AB: Edmonton, Elkwater L; MB: AB: Edmonton AB: Edmonton; SK: Pike L AB: Cypress Hills, Edmonton AB: Edmonton AB: Edmonton, Lethbridge AB: Edmonton, Red Deer AB: Edmonton, Lethbridge AB: Edmonton, AB: Edmonton Red Deer; SK: L, Jumping Pound Cr, Edmonton, George AB: Calgary, Rockglen, Brooks SK: Redberry Wabamun; L, Edmonton, George AB: Bilby, White SK: 23 km E Waterton; AB: MB: Winnipeg SK: Resource, Wadena; AB: Winnipeg AB: Edmonton MB: Riverton MB: Aweme, Waterton at View Calgary), Mountain W AB: Jumping Pound Cr (20 mi Waterton Lakes NP, AB: Edmonton Calgary) W AB: Jumping Pound Cr (20 mi Wabamun Calgary), W AB: Edmonton, Jumping Pound Cr (20 mi Calgary) W L, Jumping Pound Cr (20 mi AB: Edmonton, George AB: Edmonton AB: Edmonton Rockglen SK: Elbow, AB: Edmonton W Jumping Pound Cr (32 km Edmonton, Elkwater, Cowley, AB: Bilby, Aweme MB: Victor; NP; SK: St. L Waterton Waterton, Calgary), AB: Cypress Hills, Edmonton, Red Deer SK: Willow Bunch SK: Willow AB: Edmonton, Fawcett, Waterton AB: Edmonton, Fawcett, 5,N 5,N 5,9,N 71,N 70,H 5,N 5,68,N 5,N 9,67,N 5,N

63,N 5,N 5,N 5,N

65,N 5,N 68,N 68,N 5,N

68,N 9,N 5,N 5,63,N 63,N 5,N 64,N 64,N 68,N

68,N 14,N 64,N 5,64,N 29,N 5,N 5,64,N 64,N 64,N 65,N 64,N 5,N (Davis, 1897) (Provancher, (Provancher, (Provancher, (Provancher, Walley, 1941 Walley, (Thomson, 1883) Davis, 1897 (Gravenhorst, (Gravenhorst, Barron, 1994 (Provancher, (Ashmead, 1902) (Thomson, 1883) Cushman, 1922 Barron, 1981 Barron, 1994 Cushman, 1933 Leblanc, 1989 Leblanc, 1989 (Davis, 1897) Davis, 1897 Leblanc, 1989 Leblanc, 1989 a 5,63,N 5,9,N 66,H 5,68,N 68,69,H,Oc Lathrolestes aquilus Lathrolestes Homaspis albipes Davis, 1897 Himerta pumila Leblanc, 1989 Himerta lineola (Provancher, 1886) Zoophthorus gigas (Provancher, Himerta foxleei SUBFAMILY Species Himerta carinata Leblanc, 1989 CTENOPELMATINAE Cushman, 1924 Absyrtus arealis 1874) Alexeter canaliculatus (Provancher, Alexeter difficilis (Cresson, 1868) Alexeter luteifrons Alexeter notatus Anisotacrus popofensis (Ashmead, 1902) Campodorus picens (Davis, 1897) Ctenopelma crassicorne 1941 Walley, Ctenopelma croceum 1886) Ctenopelma nigricorne (Provancher, Ctenopelma ruficeps Ctenopelma sanguineum 1875) bituminosus Davis, 1897 Euryproctus ramis Davis, 1897 Euryproctus coxatus Hadrodactylus femoratus Hadrodactylus seminiger Hadrodactylus Himerta annulata (Davis, 1895) Himerta atra (Cushman, 1924) Himerta bicolorata Homaspis slossonae Hypamblys albopictus 1829) 1933) Hypamblys conformis (Walley, Lamachus lophyri Ashmead, 1898 Lamachus ruficoxalis (Cushman, 1919) bulbus Barron, 1994 Lathrolestes carinatus Lathrolestes caudatus Lathrolestes luteolator Lathrolestes 1829) nigricollis Lathrolestes pictus Lathrolestes Barron, 1994 protrusus Lathrolestes 2010 thomsoni Reshchikov, Lathrolestes truncatus Lathrolestes 1888) Mesoleius insidiosus (Cresson, 1864) 1874) Himerta rubiginosa (Cresson, 1879) Himerta epicnemia Hadrodactylus tibialis Hadrodactylus Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 355 Hymenoptera: Diprionidae (1) None None None None Host Family (No. spp.) Guild/Notes Hymenoptera: Cimbicidae (1) None None None None None None None None Hymenoptera: Pamphiliidae (1) None None None None None None None None None None None Hymenoptera: Pamphiliidae (1) Hymenoptera: Pamphiliidae (2) (16) Tenthredinidae Hymenoptera: Diprionidae, None Hymenoptera: Diprionidae (4) None None None None (10) Tenthredinidae Argidae, Hymenoptera: (2) Tenthredinidae Hymenoptera: None None (1) Tenthredinidae Hymenoptera: (1) Tenthredinidae Hymenoptera: None None Hymenoptera: Tenthredinidae (1) Tenthredinidae Hymenoptera: None m , Winnipeg Fawcett , St. Paul Strathclair b AB: Drumheller, George L, Gull L, Jumping Pound Cr (20 mi W W L, Gull Jumping Pound Cr (20 mi George AB: Drumheller, Calgary), Red Deer; SK: Great Sand Hills, Saskatoon AB: Calgary, Edmonton AB: Calgary, AB: Bilby, Calgary, Carstairs, Demmitt, Drumheller, Edmonton, Carstairs, Demmitt, Drumheller, Calgary, AB: Bilby, Waterton Calgary), W L, Jumping Pound Cr (20mi George Elkwater P, Victor, Albert, Rockglen, St. Prince Elbow, NP; SK: Broadview, Saskatoon; MB: Southwest AB: Edmonton; MB: Ninette AB: Red Deer Localities AB: Edmonton AB: Edmonton AB: Edmonton , Red Deer AB: Southwest AB: Edmonton AB: Waterton AB: Edmonton AB: Red Deer AB: Edmonton Mt Turtle AB: Edmonton, Elkwater L; MB: AB: Edmonton AB: Edmonton; SK: Pike L AB: Cypress Hills, Edmonton AB: Edmonton AB: Edmonton, Lethbridge AB: Edmonton, Red Deer AB: Edmonton, Lethbridge AB: Edmonton, AB: Edmonton Red Deer; SK: L, Jumping Pound Cr, Edmonton, George AB: Calgary, Rockglen, Brooks SK: Redberry Wabamun; L, Edmonton, George AB: Bilby, White SK: 23 km E Waterton; AB: MB: Winnipeg SK: Resource, Wadena; AB: Winnipeg AB: Edmonton MB: Riverton MB: Aweme, Waterton at View Calgary), Mountain W AB: Jumping Pound Cr (20 mi Waterton Lakes NP, AB: Edmonton Calgary) W AB: Jumping Pound Cr (20 mi Wabamun Calgary), W AB: Edmonton, Jumping Pound Cr (20 mi Calgary) W L, Jumping Pound Cr (20 mi AB: Edmonton, George AB: Edmonton AB: Edmonton Rockglen SK: Elbow, AB: Edmonton W Jumping Pound Cr (32 km Edmonton, Elkwater, Cowley, AB: Bilby, Aweme MB: Victor; NP; SK: St. L Waterton Waterton, Calgary), AB: Cypress Hills, Edmonton, Red Deer SK: Willow Bunch SK: Willow AB: Edmonton, Fawcett, Waterton AB: Edmonton, Fawcett, 5,N 5,N 5,9,N 71,N 70,H 5,N 5,68,N 5,N 9,67,N 5,N

63,N 5,N 5,N 5,N

65,N 5,N 68,N 68,N 5,N

68,N 9,N 5,N 5,63,N 63,N 5,N 64,N 64,N 68,N

68,N 14,N 64,N 5,64,N 29,N 5,N 5,64,N 64,N 64,N 65,N 64,N 5,N (Davis, 1897) (Provancher, (Provancher, (Provancher, (Provancher, Walley, 1941 Walley, (Thomson, 1883) Davis, 1897 (Gravenhorst, (Gravenhorst, Barron, 1994 (Provancher, (Ashmead, 1902) (Thomson, 1883) Cushman, 1922 Barron, 1981 Barron, 1994 Cushman, 1933 Leblanc, 1989 Leblanc, 1989 (Davis, 1897) Davis, 1897 Leblanc, 1989 Leblanc, 1989 a 5,63,N 5,9,N 66,H 5,68,N 68,69,H,Oc Lathrolestes aquilus Lathrolestes Homaspis albipes Davis, 1897 Himerta pumila Leblanc, 1989 Himerta lineola (Provancher, 1886) Zoophthorus gigas (Provancher, Himerta foxleei SUBFAMILY Species Himerta carinata Leblanc, 1989 CTENOPELMATINAE Cushman, 1924 Absyrtus arealis 1874) Alexeter canaliculatus (Provancher, Alexeter difficilis (Cresson, 1868) Alexeter luteifrons Alexeter notatus Anisotacrus popofensis (Ashmead, 1902) Campodorus picens (Davis, 1897) Ctenopelma crassicorne 1941 Walley, Ctenopelma croceum 1886) Ctenopelma nigricorne (Provancher, Ctenopelma ruficeps Ctenopelma sanguineum 1875) bituminosus Davis, 1897 Euryproctus ramis Davis, 1897 Euryproctus coxatus Hadrodactylus femoratus Hadrodactylus seminiger Hadrodactylus Himerta annulata (Davis, 1895) Himerta atra (Cushman, 1924) Himerta bicolorata Homaspis slossonae Hypamblys albopictus 1829) 1933) Hypamblys conformis (Walley, Lamachus lophyri Ashmead, 1898 Lamachus ruficoxalis (Cushman, 1919) bulbus Barron, 1994 Lathrolestes carinatus Lathrolestes caudatus Lathrolestes luteolator Lathrolestes 1829) nigricollis Lathrolestes pictus Lathrolestes Barron, 1994 protrusus Lathrolestes 2010 thomsoni Reshchikov, Lathrolestes truncatus Lathrolestes 1888) Mesoleius insidiosus (Cresson, 1864) 1874) Himerta rubiginosa (Cresson, 1879) Himerta epicnemia Hadrodactylus tibialis Hadrodactylus 356 M. D. Schwarzfeld Host Family (No. spp.) Guild/Notes Hymenoptera: Cimbicidae (2) None (11) Tenthredinidae Hymenoptera: Diprionidae, None None None (2) Tenthredinidae Hymenoptera: Diprionidae, None (1) Tenthredinidae Hymenoptera: Hymenoptera: Cimbicidae; Lepidoptera: Erebidae, Lasiocampidae, Sphingidae (13) (1) Tetragnathidae Araneae: (2) Tenthredinidae Hymenoptera: Lepidoptera: Erebidae (1) None (1) Tenthredinidae Hymenoptera: None (1) Tenthredinidae Hymenoptera: (1) Tenthredinidae Hymenoptera: Hymenoptera: Cimbicidae (1) Hymenoptera: Cimbicidae (1) Hymenoptera: Cimbicidae (5) Hymenoptera: Cimbicidae (7) (1) Tenthridinidae Hymenoptera: Argidae, Hymenoptera: (4) Tenthredinidae Argidae, Hymenoptera: Lepidoptera: Tenthredinidae; Sphingidae (11) None None (2) Tenthredinidae Hymenoptera: (2) Tenthredinidae Hymenoptera: None None None None None None Hymenoptera: Tenthredinidae; Pamphiliidae, Lepidoptera: Crambidae, Pyralidae, (5) Tortricidae None None ; m

m

, 3 central localities m m Wabamun , S of Saskatoon m Fawcett m b , Glenboro, Hodgson, Horton, Millwood, 5 mi N Franklin , Glenboro, Hodgson, Horton, Millwood, 5 mi N Localities AB: Edmonton AB: Red Deer MB: Treesbank AB: Edmonton Attons L, Cut Knife, Bapaume, Great Edmonton; SK: Wabamun, AB: Swift Current), Elbow; MB: Ninette, Duck Mt FR Sand Hills (W Franklin Aweme, Edmonton; MB: AB: Calgary, AB: Edmonton AB: Edmonton, Lethbridge Calgary); MB: Bald Head Hills (21 W AB: Jumping Pound Cr (32 km km N Glenboro), Shilo (tamarack bog), Swift Current) (Cut Knife), Great Sand Hills (W Attons L SK: AB: Drumheller; SK: Kenosee; MB: Ninette W L, Jumping Pound Cr (32 km AB: Bilby , Edmonton, George Cut Knife, Prince Calgary), Lundbreck, Red Deer; SK: Big River, Albert; MB: Ninette, Morton Calgary); SK: Great Sand Hills; MB: W AB: Jumping Pound Cr (32km Shilo (tamarack bog) AB: Edmonton AB: Edmonton AB: Edmonton AB: Edmonton AB: Cameron L NA AB: Wagner nr Regina, Fort Qu'Appelle; MB: SK: nr Lloydminster, Riding Mt P SK: nr N Battleford; MB: Boissevain, Portage la Prairie SK: Duck Mt Waterton nr AB: Cochrane, Edmonton, Red Deer, NA Wagner AB: Edmonton, AB: Edmonton, Red Deer AB: Cypress Hills, Edmonton, at Cut Knife Attons L SK: AB: Bilby SK: Sutherland AB: Edmonton, AB: Edmonton NA Wagner AB: Edmonton, AB: Edmonton AB: Edmonton NP Waterton Rd in AB: Cameron L Albert AB: Gull L; SK: Prince Edmonton, Clymont, Drumheller, Aspen Beach, Bilby, AB: Gull L, Jumping Pound Cr, Egremont, Elkwater L, P, Wainwright, Wabamun, Lethbridge, Manyberries, Stettler, SK: Birch Hills, Canora, Dubuc, Estevan, Gull L, Holbein, Waterton; Regina, Albert, Redberry, Kenosee, Lebret, Prince Kandahar, Boissevain, Carberry, Aweme, Saskatoon, 5 mi E Swift Current; MB: Dauphin, Shilo, Minnedosa, Ninette, Oak L, Pilot Mound, Roblin, 5 mi SW Stockton, Swan River W Souris, 3 mi AB: Edmonton SK: nr Saskatoon 5,N 5,N 75,N 2,5,N 2,5,H 76,H 5,9,43,H 78,N 76,N 78,H 5,H 5,N 5,9,75,N 79,H 73,N 14,N 72,H 78,H 14,N 75,N 5,9,N 76,H 75,N 79,H 5,N 77,N 5,9,N 5,77,N 5,N 9,N 2,N 5,N,T 14,N 75,N 77,N (Ashmead, 1896) (Gravenhorst, (Fabricius, 1804) (Linnaeus, 1758) (Cresson, 1864) (Thunberg, 1824) (Thunberg, (Cresson, 1868) (Holmgren, 1880) (Provancher, 1886) (Provancher, (Cresson, 1864) 1874) (Provancher, (Gravenhorst, 1829) (Cresson, 1864) Walley, 1935 Walley, (Davis, 1897) (Cresson, 1868) (Davis, 1897) (Davis, 1897) (Provancher, 1876) (Provancher, a 5,N 5,9,79,H (Cresson, 1868) Mesoleius submarginatus 1886 Rhorus tristis Provancher, Scolobates auriculatus Syndipnus conformis (Holmgren, 1857) SUBFAMILY Species 74,H Rhorus pilosus (Davis, 1897) Mesoleius tarsalis (Cresson, 1868) 1912 Morley, Mesoleius tenthredinis Mesoleptidea decens Oetophorus pleuralis (Cresson, 1864) Opheltes glaucopterus Otlophorus carbonarius (Gravenhorst, 1829) xanthostomus (Gravenhorst, Pantorhaestes 1829) Perilissus araius Burks, 1952 Perilissus coloradensis (Ashmead, 1896) Perilissus concolor Perilissus decoloratus Perilissus discolor (Cresson, 1864) Phobetes striatus Phobetes uniformis mellipes Protarchus 1844) sorbi (Ratzeburg, Protarchus 1822) testatorius (Thunberg, Protarchus Rhinotorus ovalis Rhorus clapini (Cresson, 1868) Rhorus varifrons Scopesis gesticulator Sympherta fucata Syndipnus lateralis 1940 Syndipnus rubiginosus Walley, Synoecetes festivus semirufus (Cresson, 1864) Trematopygus Xenoschesis cinctiventris Xenoschesis nitida CYLLOCERIINAE Allomacrus arcticus (Roman, 1925) Cylloceria borealis Cylloceria melancholica 1820) DIACRITINAE Diacritus muliebris (Cresson, 1868) Synomelix obesa Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 357 Host Family (No. spp.) Guild/Notes Hymenoptera: Cimbicidae (2) None (11) Tenthredinidae Hymenoptera: Diprionidae, None None None (2) Tenthredinidae Hymenoptera: Diprionidae, None (1) Tenthredinidae Hymenoptera: Hymenoptera: Cimbicidae; Lepidoptera: Erebidae, Lasiocampidae, Sphingidae (13) (1) Tetragnathidae Araneae: (2) Tenthredinidae Hymenoptera: Lepidoptera: Erebidae (1) None (1) Tenthredinidae Hymenoptera: None (1) Tenthredinidae Hymenoptera: (1) Tenthredinidae Hymenoptera: Hymenoptera: Cimbicidae (1) Hymenoptera: Cimbicidae (1) Hymenoptera: Cimbicidae (5) Hymenoptera: Cimbicidae (7) (1) Tenthridinidae Hymenoptera: Argidae, Hymenoptera: (4) Tenthredinidae Argidae, Hymenoptera: Lepidoptera: Tenthredinidae; Sphingidae (11) None None (2) Tenthredinidae Hymenoptera: (2) Tenthredinidae Hymenoptera: None None None None None None Hymenoptera: Tenthredinidae; Pamphiliidae, Lepidoptera: Crambidae, Pyralidae, (5) Tortricidae None None ; m

m

, 3 central localities m m Wabamun , S of Saskatoon m Fawcett m b , Glenboro, Hodgson, Horton, Millwood, 5 mi N Franklin , Glenboro, Hodgson, Horton, Millwood, 5 mi N Localities AB: Edmonton AB: Red Deer MB: Treesbank AB: Edmonton Attons L, Cut Knife, Bapaume, Great Edmonton; SK: Wabamun, AB: Swift Current), Elbow; MB: Ninette, Duck Mt FR Sand Hills (W Franklin Aweme, Edmonton; MB: AB: Calgary, AB: Edmonton AB: Edmonton, Lethbridge Calgary); MB: Bald Head Hills (21 W AB: Jumping Pound Cr (32 km km N Glenboro), Shilo (tamarack bog), Swift Current) (Cut Knife), Great Sand Hills (W Attons L SK: AB: Drumheller; SK: Kenosee; MB: Ninette W L, Jumping Pound Cr (32 km AB: Bilby , Edmonton, George Cut Knife, Prince Calgary), Lundbreck, Red Deer; SK: Big River, Albert; MB: Ninette, Morton Calgary); SK: Great Sand Hills; MB: W AB: Jumping Pound Cr (32km Shilo (tamarack bog) AB: Edmonton AB: Edmonton AB: Edmonton AB: Edmonton AB: Cameron L NA AB: Wagner nr Regina, Fort Qu'Appelle; MB: SK: nr Lloydminster, Riding Mt P SK: nr N Battleford; MB: Boissevain, Portage la Prairie SK: Duck Mt Waterton nr AB: Cochrane, Edmonton, Red Deer, NA Wagner AB: Edmonton, AB: Edmonton, Red Deer AB: Cypress Hills, Edmonton, at Cut Knife Attons L SK: AB: Bilby SK: Sutherland AB: Edmonton, AB: Edmonton NA Wagner AB: Edmonton, AB: Edmonton AB: Edmonton NP Waterton Rd in AB: Cameron L Albert AB: Gull L; SK: Prince Edmonton, Clymont, Drumheller, Aspen Beach, Bilby, AB: Gull L, Jumping Pound Cr, Egremont, Elkwater L, P, Wainwright, Wabamun, Lethbridge, Manyberries, Stettler, SK: Birch Hills, Canora, Dubuc, Estevan, Gull L, Holbein, Waterton; Regina, Albert, Redberry, Kenosee, Lebret, Prince Kandahar, Boissevain, Carberry, Aweme, Saskatoon, 5 mi E Swift Current; MB: Dauphin, Shilo, Minnedosa, Ninette, Oak L, Pilot Mound, Roblin, 5 mi SW Stockton, Swan River W Souris, 3 mi AB: Edmonton SK: nr Saskatoon 5,N 5,N 75,N 2,5,N 2,5,H 76,H 5,9,43,H 78,N 76,N 78,H 5,H 5,N 5,9,75,N 79,H 73,N 14,N 72,H 78,H 14,N 75,N 5,9,N 76,H 75,N 79,H 5,N 77,N 5,9,N 5,77,N 5,N 9,N 2,N 5,N,T 14,N 75,N 77,N (Ashmead, 1896) (Gravenhorst, (Fabricius, 1804) (Linnaeus, 1758) (Cresson, 1864) (Thunberg, 1824) (Thunberg, (Cresson, 1868) (Holmgren, 1880) (Provancher, 1886) (Provancher, (Cresson, 1864) 1874) (Provancher, (Gravenhorst, 1829) (Cresson, 1864) Walley, 1935 Walley, (Davis, 1897) (Cresson, 1868) (Davis, 1897) (Davis, 1897) (Provancher, 1876) (Provancher, a 5,N 5,9,79,H (Cresson, 1868) Mesoleius submarginatus 1886 Rhorus tristis Provancher, Scolobates auriculatus Syndipnus conformis (Holmgren, 1857) SUBFAMILY Species 74,H Rhorus pilosus (Davis, 1897) Mesoleius tarsalis (Cresson, 1868) 1912 Morley, Mesoleius tenthredinis Mesoleptidea decens Oetophorus pleuralis (Cresson, 1864) Opheltes glaucopterus Otlophorus carbonarius (Gravenhorst, 1829) xanthostomus (Gravenhorst, Pantorhaestes 1829) Perilissus araius Burks, 1952 Perilissus coloradensis (Ashmead, 1896) Perilissus concolor Perilissus decoloratus Perilissus discolor (Cresson, 1864) Phobetes striatus Phobetes uniformis mellipes Protarchus 1844) sorbi (Ratzeburg, Protarchus 1822) testatorius (Thunberg, Protarchus Rhinotorus ovalis Rhorus clapini (Cresson, 1868) Rhorus varifrons Scopesis gesticulator Sympherta fucata Syndipnus lateralis 1940 Syndipnus rubiginosus Walley, Synoecetes festivus semirufus (Cresson, 1864) Trematopygus Xenoschesis cinctiventris Xenoschesis nitida CYLLOCERIINAE Allomacrus arcticus (Roman, 1925) Cylloceria borealis Cylloceria melancholica 1820) DIACRITINAE Diacritus muliebris (Cresson, 1868) Synomelix obesa 358 M. D. Schwarzfeld Mostly Host Family (No. spp.) Guild/Notes None Coleoptera: Chrysomelidae, Curculionidae; Diptera: Anthomyiidae, Muscidae, Syrphidae; Hymenoptera: Diprionidae, Pamphiliidae; Lepidoptera: Crambidae, Elachistidae, Erebidae, Gelechiidae, Noctuidae, Plutellidae (74) None None Diptera: Syrphidae (3) Coleoptera: Curculionidae; Diptera: Syrphidae, Tenthredinidae; Hymenoptera: Tachinidae; (17) Tortricidae Lepidoptera: Noctuidae, Mostly Syrphidae Diptera: Syrphidae; Lepidoptera: Noctuidae (5) Syrphidae Coleoptera: Curculionidae Diptera: Syrphidae (16) Mostly Syrphidae Coleoptera: Curculionidae; Diptera: Syrphidae (9) Mostly Syrphidae Diptera: Syrphidae; Lepidoptera: Noctuidae (13) Mostly Syrphidae Diptera: Syrphidae None Diptera: Syrphidae; Lepidoptera: Lasiocampidae (4) Mostly Syrphidae (5) Tachinidae Diptera: Syrphidae, Mostly Syrphidae Diptera: Cecidomyiidae, Syrphidae; Lepidoptera: (5) Tortricidae Mostly concealed feeders Anthomyiidae, Syrphidae; Lepidoptera: Diptera: (14) Tortricidae Mostly Syrphidae None Diptera: Syrphidae (2) None Anthomyiidae, Syrphidae; Lepidoptera: Diptera: (11) Tortricidae Pieridae, Plutellidae, None Anthomyiidae, Syrphidae; Lepidoptera: Diptera: Choreutidae (4) Anthomyiidae, Coleoptera: Bruchidae; Diptera: Chloropidae, Syrphidae; Lepidoptera: Crambidae, Noctuidae (12) None Diptera: Syrphidae; Lepidoptera: Oecophoridae (2) Anthomyiidae, Syrphidae (2) Diptera: None None None Diptera: Syrphidae (6) None None , Ninette, , Morris Glenlea Winnipeg b AB: Wagner NA; MB: 5 mi SW Shilo NA; MB: 5 mi SW Wagner AB: Wawanesa, Mt Russell, Sanford , Shilo, Swan River Turtle FR, Virden, Whitewater L, Localities NA AB: Wagner NA AB: Wagner Cypress Hills, Edmonton, Elkwater L, Aspen Beach, Bilby, AB: Rockyford, Scandia, Lethbridge, Onefour, Medicine Hat, Milk River, Assiniboia, Belle Plaine, Chaplin, Elbow, SK: Wabamun; Standard, Minton, Moosomin, Pike L, Redberry, Maple Cr, Estevan, Kandahar, Marie, Val Regina, Rockglen, Rutland, Saskatoon, Swift Current, south; MB: Boissevain, Brandon, Douglas, NA; Wagner Wabamun, Edmonton, High River, AB: Bilby, SK: Swift Current Waterton Wabamun, Sylvan L, Edmonton, Gull L, Red Deer, AB: Bilby, AB: Medicine Hat NA; SK: Wagner Wabamun, Consort, Edmonton, Sylvan L, AB: Bilby, Shilo 5 mi SW Aweme, Bunch; MB: Willow Antelope L, Snowden, Clymont, Edmonton, Lethbridge Calgary, AB: Bilby, Edmonton; SK: Cookson, Indian Head Calgary, AB: Bilby, Vermilion AB: Consort, Edmonton, White Fox NA; SK: Saskatoon, Wagner AB: Edmonton, Duchess, Edmonton, Lethbridge, Medicine Hat, Calgary, AB: Bilby, Aweme SK: Dunblane, Moosomin, Saskatoon; MB: Waterton; Onefour, NA AB: Wagner NP Waterton Rd in Cameron L NA Wagner AB: Edmonton, NA; Wagner AB: Brooks, Cooking L, Duchess, Edmonton, Lethbridge, Ninette, Starbuck Treesbank, SK: Moosomin, Regina, Saskatoon; MB: AB: Blackfoot Hills, Duchess, Edmonton, Lethbridge Aspen Beach, Pincher AB: AB: Edmonton Treesbank NA; SK: Saskatoon; MB: Ninette, Wagner AB: Edmonton, NA AB: Wagner NA; SK: Regina; MB: Wagner AB: Bilby , Edmonton Elkwater, Treesbank Assiniboine River nr Edmonton, Gull L, Lethbridge, Medicine Hat, Onefour, AB: Calgary, SK: Waterton; NA, Wagner Tilley, Shaughnessy, Patricia, Pincher Cr, Cypress Hills, Great Sand Saskatoon, Swift Current NA AB: Wagner NA AB: Wagner AB: Edmonton AB: Bilby AB: Iron Springs, Lethbridge; SK: Moosomin NA Wagner AB: Edmonton, AB: Wabamun Lethbridge, AB: Edmonton, Consort Lethbridge; SK: Regina, Edmonton, High River, AB: Calgary, Bunch Saskatoon, Willow 80,H 80,H 2,H 2,H 80,H,O 2,N 2,N,T 2,5,80,H,O 5,80,H,T 5,80,N,T 2,5,80,H,O 2,9,80,H 80,H 2,80,H,T 2,H 80,H 5,80,N 2,H (Dasch, 1964) (Dasch, 1964) (Holmgren, (Davis, 1895) (Thomson, 1890) (Fabricius, (Say, 1835) (Say, (Cresson, 1868) 1824) (Thunberg, Brues, 1908 80,N Dasch, 1964 2,80,N (Ruthe, 1859) (Holmgren, 1858) (Thomson, 1890) a 5,9,80,81,W 5,80,H,T,O 2,80,H 5,80,H 2,80,H,T 2,5,80,H,O DIPLAZONTINAE Diplazon bradleyi Diplazon deletus Diplazon orbitalis (Cresson, 1865) SUBFAMILY Species pacificus (Cresson, 1879) Homotropus 2,5,9,80,H 1858) Diplazon hyperboreus (Marshall, 1877) Diplazon hyperboreus *Diplazon laetatorius 1781) Diplazon pectoratorius (Thunberg, 1824) Diplazon scutellaris 1824) Diplazon tetragonus (Thunberg, Diplazon tibiatorius Enizemum ornatum (Gravenhorst, 1829) Enizemum petiolatum Fossatyloides gracilentus nigritarsus (Gravenhorst, Homotropus 1829) pallipes (Gravenhorst, Homotropus 1829) signatus (Gravenhorst, 1829) Homotropus (Dasch, 1964) vitreus Homotropus (Desvignes, 1856) Phthorima compressa melanaspis (Thomson, 1890) Promethes sulcator (Gravenhorst, Promethes 1829) Sussaba aciculata Sussaba dorsalis (Holmgren, 1858) Sussaba pulchella Sussaba punctiventris (Brues, 1908) 80,N,T Sussaba rohweri Syrphoctonus albopictus 2,80,N Syrphoctonus columbiensis 80,N Syrphoctonus fraudulentus 80,N Syrphoctonus laevis (Cresson, 1865) Syrphoctonus maculifrons 5,80,N,T,Oc Syrphoctonus minimus (Cresson, 1864) Syrphoctonus nudus (Dasch, 1964) Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 359 Mostly Host Family (No. spp.) Guild/Notes None Coleoptera: Chrysomelidae, Curculionidae; Diptera: Anthomyiidae, Muscidae, Syrphidae; Hymenoptera: Diprionidae, Pamphiliidae; Lepidoptera: Crambidae, Elachistidae, Erebidae, Gelechiidae, Noctuidae, Plutellidae (74) None None Diptera: Syrphidae (3) Coleoptera: Curculionidae; Diptera: Syrphidae, Tenthredinidae; Hymenoptera: Tachinidae; (17) Tortricidae Lepidoptera: Noctuidae, Mostly Syrphidae Diptera: Syrphidae; Lepidoptera: Noctuidae (5) Syrphidae Coleoptera: Curculionidae Diptera: Syrphidae (16) Mostly Syrphidae Coleoptera: Curculionidae; Diptera: Syrphidae (9) Mostly Syrphidae Diptera: Syrphidae; Lepidoptera: Noctuidae (13) Mostly Syrphidae Diptera: Syrphidae None Diptera: Syrphidae; Lepidoptera: Lasiocampidae (4) Mostly Syrphidae (5) Tachinidae Diptera: Syrphidae, Mostly Syrphidae Diptera: Cecidomyiidae, Syrphidae; Lepidoptera: (5) Tortricidae Mostly concealed feeders Anthomyiidae, Syrphidae; Lepidoptera: Diptera: (14) Tortricidae Mostly Syrphidae None Diptera: Syrphidae (2) None Anthomyiidae, Syrphidae; Lepidoptera: Diptera: (11) Tortricidae Pieridae, Plutellidae, None Anthomyiidae, Syrphidae; Lepidoptera: Diptera: Choreutidae (4) Anthomyiidae, Coleoptera: Bruchidae; Diptera: Chloropidae, Syrphidae; Lepidoptera: Crambidae, Noctuidae (12) None Diptera: Syrphidae; Lepidoptera: Oecophoridae (2) Anthomyiidae, Syrphidae (2) Diptera: None None None Diptera: Syrphidae (6) None None , Ninette, , Morris Glenlea Winnipeg b AB: Wagner NA; MB: 5 mi SW Shilo NA; MB: 5 mi SW Wagner AB: Wawanesa, Mt Russell, Sanford , Shilo, Swan River Turtle FR, Virden, Whitewater L, Localities NA AB: Wagner NA AB: Wagner Cypress Hills, Edmonton, Elkwater L, Aspen Beach, Bilby, AB: Rockyford, Scandia, Lethbridge, Onefour, Medicine Hat, Milk River, Assiniboia, Belle Plaine, Chaplin, Elbow, SK: Wabamun; Standard, Minton, Moosomin, Pike L, Redberry, Maple Cr, Estevan, Kandahar, Marie, Val Regina, Rockglen, Rutland, Saskatoon, Swift Current, south; MB: Boissevain, Brandon, Douglas, NA; Wagner Wabamun, Edmonton, High River, AB: Bilby, SK: Swift Current Waterton Wabamun, Sylvan L, Edmonton, Gull L, Red Deer, AB: Bilby, AB: Medicine Hat NA; SK: Wagner Wabamun, Consort, Edmonton, Sylvan L, AB: Bilby, Shilo 5 mi SW Aweme, Bunch; MB: Willow Antelope L, Snowden, Clymont, Edmonton, Lethbridge Calgary, AB: Bilby, Edmonton; SK: Cookson, Indian Head Calgary, AB: Bilby, Vermilion AB: Consort, Edmonton, White Fox NA; SK: Saskatoon, Wagner AB: Edmonton, Duchess, Edmonton, Lethbridge, Medicine Hat, Calgary, AB: Bilby, Aweme SK: Dunblane, Moosomin, Saskatoon; MB: Waterton; Onefour, NA AB: Wagner NP Waterton Rd in Cameron L NA Wagner AB: Edmonton, NA; Wagner AB: Brooks, Cooking L, Duchess, Edmonton, Lethbridge, Ninette, Starbuck Treesbank, SK: Moosomin, Regina, Saskatoon; MB: AB: Blackfoot Hills, Duchess, Edmonton, Lethbridge Aspen Beach, Pincher AB: AB: Edmonton Treesbank NA; SK: Saskatoon; MB: Ninette, Wagner AB: Edmonton, NA AB: Wagner NA; SK: Regina; MB: Wagner AB: Bilby , Edmonton Elkwater, Treesbank Assiniboine River nr Edmonton, Gull L, Lethbridge, Medicine Hat, Onefour, AB: Calgary, SK: Waterton; NA, Wagner Tilley, Shaughnessy, Patricia, Pincher Cr, Cypress Hills, Great Sand Saskatoon, Swift Current NA AB: Wagner NA AB: Wagner AB: Edmonton AB: Bilby AB: Iron Springs, Lethbridge; SK: Moosomin NA Wagner AB: Edmonton, AB: Wabamun Lethbridge, AB: Edmonton, Consort Lethbridge; SK: Regina, Edmonton, High River, AB: Calgary, Bunch Saskatoon, Willow 80,H 80,H 2,H 2,H 80,H,O 2,N 2,N,T 2,5,80,H,O 5,80,H,T 5,80,N,T 2,5,80,H,O 2,9,80,H 80,H 2,80,H,T 2,H 80,H 5,80,N 2,H (Dasch, 1964) (Dasch, 1964) (Holmgren, (Davis, 1895) (Thomson, 1890) (Fabricius, (Say, 1835) (Say, (Cresson, 1868) 1824) (Thunberg, Brues, 1908 80,N Dasch, 1964 2,80,N (Ruthe, 1859) (Holmgren, 1858) (Thomson, 1890) a 5,9,80,81,W 5,80,H,T,O 2,80,H 5,80,H 2,80,H,T 2,5,80,H,O DIPLAZONTINAE Diplazon bradleyi Diplazon deletus Diplazon orbitalis (Cresson, 1865) SUBFAMILY Species pacificus (Cresson, 1879) Homotropus 2,5,9,80,H 1858) Diplazon hyperboreus (Marshall, 1877) Diplazon hyperboreus *Diplazon laetatorius 1781) Diplazon pectoratorius (Thunberg, 1824) Diplazon scutellaris 1824) Diplazon tetragonus (Thunberg, Diplazon tibiatorius Enizemum ornatum (Gravenhorst, 1829) Enizemum petiolatum Fossatyloides gracilentus nigritarsus (Gravenhorst, Homotropus 1829) pallipes (Gravenhorst, Homotropus 1829) signatus (Gravenhorst, 1829) Homotropus (Dasch, 1964) vitreus Homotropus (Desvignes, 1856) Phthorima compressa melanaspis (Thomson, 1890) Promethes sulcator (Gravenhorst, Promethes 1829) Sussaba aciculata Sussaba dorsalis (Holmgren, 1858) Sussaba pulchella Sussaba punctiventris (Brues, 1908) 80,N,T Sussaba rohweri Syrphoctonus albopictus 2,80,N Syrphoctonus columbiensis 80,N Syrphoctonus fraudulentus 80,N Syrphoctonus laevis (Cresson, 1865) Syrphoctonus maculifrons 5,80,N,T,Oc Syrphoctonus minimus (Cresson, 1864) Syrphoctonus nudus (Dasch, 1964) 360 M. D. Schwarzfeld Mostly Loopers Cutworms Host Family (No. spp.) Guild/Notes Lepidoptera: Erebidae, Geometridae, Noctuidae, Notodontidae (8) None Diptera: Syrphidae (3) None None Diptera: Syrphidae (16) Anthomyiidae, Coleoptera: Curculionidae; Diptera: Syrphidae (9) Mostly Syrphidae None (4) Tortricidae Diptera: Syrphidae; Lepidoptera: Syrphidae None Diptera: Syrphidae (2) None Coleoptera: Curculionidae; Diptera: Syrphidae (5) Mostly Syrphidae Diptera: Syrphidae; Lepidoptera: Coleophoridae, (28) Tortricidae Crambidae, Erebidae, Mostly Syrphidae None Hymenoptera: Diprionidae, Ichneumonidae, (3) Tenthredinidae Lepidoptera: Erebidae (1) Hymenoptera: Diprionidae, Ichneumonidae; Lepidoptera: Geometridae (15) Mostly conifer feeders None Hymenoptera: Ichneumonidae; Lepidoptera: Erebidae, Geometridae, Noctuidae (20) Mostly loopers on conifers (2) Lepidoptera: Lycaenidae Lepidoptera: Notodontidae (2) Lepidoptera: Pyralidae (1) Lepidoptera: Geometridae, Noctuidae (5) Mostly loopers Lepidoptera: Geometridae (1) Lepidoptera: Geometridae, Notodontidae (4) Mostly loopers None None None None Hymenoptera, Lepidoptera: Diprionidae, Erebidae, Geometridae, Noctuidae, Notodontidae, Nymphalidae, (14) Tortricidae Sphingidae, None Lepidoptera: Noctuidae (1) None m , Edmonton, nr Lethbridge m b Localities NA Wagner AB: Edmonton, Edmonton AB: Bilby, NA Wagner AB: Edmonton, Edmonton AB: Bilby, AB: Bilby NA; SK: Cypress Hills Wagner AB: Cooking L , Edmonton, NA, Vermilion AB: Edmonton , Wagner SK: Saskatoon Waterton; AB: NA, Waterton AB: Wagner NA; MB: Shilo Wagner AB: Cooking L, Edmonton, NA AB: Wagner NA AB: Wagner AB: Edmonton, Lethbridge Aweme MB: Wabamun; Calgary), W AB: Jumping Pound Cr (20 mi Treesbank FR, Woods Spruce Aweme, NA; MB: Wagner AB: AB: Edmonton, Lundbreck; SK: Cut Knife MB: Aweme Attons L, Cut Knife SK: Waterton; AB: Edmonton, Treesbank Alberta; MB: AB: Glendon, Lunbreck; SK: Prince AB: Edmonton SK: Cypress Hills AB: Waterton AB: Edmonton, Red Deer AB: Edmonton, Gull L Edmonton AB: Drumheller, NA AB: Wagner AB: Edmonton AB: Edmonton AB: Edmonton Aweme SK: Great Deer; MB: AB: nr Calgary AB: Cypress Hills SK: Indian Head Waterton; AB: Edmonton, SK: Pike L; MB: Swan River 5,N 5,83,H 5,N 2,80,H 5,80,N 5,83,N 80,H,O 83,N 2,N

5,N 5,14,83,H 2,N 5,N 14,N 5,80,N 82,N 82,N 83,84,H 82,N 82,N 82,N 2,82,N (Provancher, (Provancher, 14,N (Linnaeus, (Thunberg, (Thunberg, (Gravenhorst, (Cresson, 1877) 2,H (Provancher, (Provancher, (Provancher, (Provancher, (Cresson, 1868) (Cresson, 1864) (Cresson, 1877) (Ashmead, 1890) Walley, 1932 Walley, (Cresson, 1864) a 2,5,H 2,80,N 2,9,14,80,H,O 2,5,9,N 80,H,O 2,5,80,H 5,80,H,T,O,Oc 5,N 85,H Syrphoctonus pectoralis Coelichneumon histricus (Cresson, 1867) SUBFAMILY Species 1874) frigidus Cresson, 1869 Euceros 1824) 1829) 1758) Syrphoctonus pleuralis Syrphoctonus quadrangularis (Dasch, 1964) Davis, 1895 Syrphoctonus robustus 1809) Syrphoctonus tarsatorius (Panzer, Syrphophilus bizonarius (Gravenhorst, 1829) Syrphophilus ichneumonoides 1874) Syrphophilus tricinctorius Syrphophilus tricinctus (Ashmead, 1902) (Forster, erythrozonus Tymmophorus 1850) fasciventris Dasch, 1964 Tymmophorus citropectoralis Woldstedtius (Schmiedeknecht, 1926) flavolineatus Woldstedtius EUCEROTINAE Barron, 1976 congregatus Euceros 1932 decorus Walley, Euceros digitalis Euceros sanguineus Davis, 1897 Euceros semiothisae Barron, 1976 Euceros ICHNEUMONINAE Anisobas texensis Anisopygus americanus Heinrich, 1961 Aoplus confirmatus (Cresson, 1877) Aoplus ruficeps (Gravenhorst, 1829) 1857) Aoplus torpidus (Wesmael, Asthenolabus agilis Asthenolabus canadensis Asthenolabus scutellatus 1875) Coelichneumon ater Coelichneumon deliratorius Coelichneumon eximius (Stephens, 1835) Coelichneumon maurus (Cresson, 1864) Heinrich, Coelichneumon neocretatus 1960 9,86,H Aoplus velox Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 361 Mostly Loopers Cutworms Host Family (No. spp.) Guild/Notes Lepidoptera: Erebidae, Geometridae, Noctuidae, Notodontidae (8) None Diptera: Syrphidae (3) None None Diptera: Syrphidae (16) Anthomyiidae, Coleoptera: Curculionidae; Diptera: Syrphidae (9) Mostly Syrphidae None (4) Tortricidae Diptera: Syrphidae; Lepidoptera: Syrphidae None Diptera: Syrphidae (2) None Coleoptera: Curculionidae; Diptera: Syrphidae (5) Mostly Syrphidae Diptera: Syrphidae; Lepidoptera: Coleophoridae, (28) Tortricidae Crambidae, Erebidae, Mostly Syrphidae None Hymenoptera: Diprionidae, Ichneumonidae, (3) Tenthredinidae Lepidoptera: Erebidae (1) Hymenoptera: Diprionidae, Ichneumonidae; Lepidoptera: Geometridae (15) Mostly conifer feeders None Hymenoptera: Ichneumonidae; Lepidoptera: Erebidae, Geometridae, Noctuidae (20) Mostly loopers on conifers (2) Lepidoptera: Lycaenidae Lepidoptera: Notodontidae (2) Lepidoptera: Pyralidae (1) Lepidoptera: Geometridae, Noctuidae (5) Mostly loopers Lepidoptera: Geometridae (1) Lepidoptera: Geometridae, Notodontidae (4) Mostly loopers None None None None Hymenoptera, Lepidoptera: Diprionidae, Erebidae, Geometridae, Noctuidae, Notodontidae, Nymphalidae, (14) Tortricidae Sphingidae, None Lepidoptera: Noctuidae (1) None m , Edmonton, nr Lethbridge m b Localities NA Wagner AB: Edmonton, Edmonton AB: Bilby, NA Wagner AB: Edmonton, Edmonton AB: Bilby, AB: Bilby NA; SK: Cypress Hills Wagner AB: Cooking L , Edmonton, NA, Vermilion AB: Edmonton , Wagner SK: Saskatoon Waterton; AB: NA, Waterton AB: Wagner NA; MB: Shilo Wagner AB: Cooking L, Edmonton, NA AB: Wagner NA AB: Wagner AB: Edmonton, Lethbridge Aweme MB: Wabamun; Calgary), W AB: Jumping Pound Cr (20 mi Treesbank FR, Woods Spruce Aweme, NA; MB: Wagner AB: AB: Edmonton, Lundbreck; SK: Cut Knife MB: Aweme Attons L, Cut Knife SK: Waterton; AB: Edmonton, Treesbank Alberta; MB: AB: Glendon, Lunbreck; SK: Prince AB: Edmonton SK: Cypress Hills AB: Waterton AB: Edmonton, Red Deer AB: Edmonton, Gull L Edmonton AB: Drumheller, NA AB: Wagner AB: Edmonton AB: Edmonton AB: Edmonton Aweme SK: Great Deer; MB: AB: nr Calgary AB: Cypress Hills SK: Indian Head Waterton; AB: Edmonton, SK: Pike L; MB: Swan River 5,N 5,83,H 5,N 2,80,H 5,80,N 5,83,N 80,H,O 83,N 2,N

5,N 5,14,83,H 2,N 5,N 14,N 5,80,N 82,N 82,N 83,84,H 82,N 82,N 82,N 2,82,N (Provancher, (Provancher, 14,N (Linnaeus, (Thunberg, (Thunberg, (Gravenhorst, (Cresson, 1877) 2,H (Provancher, (Provancher, (Provancher, (Provancher, (Cresson, 1868) (Cresson, 1864) (Cresson, 1877) (Ashmead, 1890) Walley, 1932 Walley, (Cresson, 1864) a 2,5,H 2,80,N 2,9,14,80,H,O 2,5,9,N 80,H,O 2,5,80,H 5,80,H,T,O,Oc 5,N 85,H Syrphoctonus pectoralis Coelichneumon histricus (Cresson, 1867) SUBFAMILY Species 1874) frigidus Cresson, 1869 Euceros 1824) 1829) 1758) Syrphoctonus pleuralis Syrphoctonus quadrangularis (Dasch, 1964) Davis, 1895 Syrphoctonus robustus 1809) Syrphoctonus tarsatorius (Panzer, Syrphophilus bizonarius (Gravenhorst, 1829) Syrphophilus ichneumonoides 1874) Syrphophilus tricinctorius Syrphophilus tricinctus (Ashmead, 1902) (Forster, erythrozonus Tymmophorus 1850) fasciventris Dasch, 1964 Tymmophorus citropectoralis Woldstedtius (Schmiedeknecht, 1926) flavolineatus Woldstedtius EUCEROTINAE Barron, 1976 congregatus Euceros 1932 decorus Walley, Euceros digitalis Euceros sanguineus Davis, 1897 Euceros semiothisae Barron, 1976 Euceros ICHNEUMONINAE Anisobas texensis Anisopygus americanus Heinrich, 1961 Aoplus confirmatus (Cresson, 1877) Aoplus ruficeps (Gravenhorst, 1829) 1857) Aoplus torpidus (Wesmael, Asthenolabus agilis Asthenolabus canadensis Asthenolabus scutellatus 1875) Coelichneumon ater Coelichneumon deliratorius Coelichneumon eximius (Stephens, 1835) Coelichneumon maurus (Cresson, 1864) Heinrich, Coelichneumon neocretatus 1960 9,86,H Aoplus velox 362 M. D. Schwarzfeld Loopers Loopers Loopers Concealed feeders Cutworms Cutworms Cutworms Cutworms Concealed feeders Host Family (No. spp.) Guild/Notes Lepidoptera: Geometridae, Notodontidae (4) Mostly loopers Lepidoptera: Erebidae, Noctuidae, Saturniidae, Sesiidae (11) Mostly setose Lepidoptera: Geometridae, Noctuidae (2) Lepidoptera: Noctuidae (3) Lepidoptera: Erebidae, Noctuidae (4) Mostly setose None Lepidoptera: Sphingidae (1) Lepidoptera: Saturniidae (1) None None Lepidoptera: Geometridae (1) None Lepidoptera: Pyralidae (1) Hymenoptera: Diprionidae, Ichneumonidae;Lepidoptera: Erebidae, Geometridae, Lasiocampidae, Noctuidae, Pieridae (28) None None None None Lepidoptera: Geometridae (1) None Lepidoptera: Geometridae (3) (2) Tortricidae Lepidoptera: None Acrolepiidae, Coleoptera: Cerambycidae; Lepidoptera: Noctuidae, Pyralidae (4) None None Lepidoptera: Noctuidae (1) Lepidoptera: Coleophoridae, Noctuidae (3) None None None Lepidoptera: Noctuidae (2) None Lepidoptera: Noctuidae (1) None None None Lepidoptera: Noctuidae (3) (2) Tortricidae Lepidoptera: Geometridae, Conifer feeders Tortricidae Hymenoptera: Diprionidae; Lepidoptera: (10) Mostly Tortricidae None Lepidoptera: Noctuidae (1) None Lepidoptera: Noctuidae (5) MB:

; Wabamun Wabamun m , Red Deer, Vermilion, Vermilion, Gull L , Red Deer, m m b Southwest Localities Aweme AB: Edmonton; MB: MB: Aweme Aweme MB: Waterton; AB: Calgary, SK: Albert; Prince MB: Aweme AB: Edmonton AB: Edmonton Treesbank AB: Edmonton; MB: corner MB: 2 localities in SW MB: Glenlea AB: Edmonton AB: nr Calgary AB: Edmonton, Several localities in southwest SK and southeast MB SK: Saskatoon AB: Calgary MB: Aweme AB: Drumheller AB: Edmonton AB: Cochrane, Edmonton AB: Fawcett NA Wagner AB: Edmonton, AB: Lethbridge MB: Swan River AB: Edmonton Edmonton, Lethbridge AB: Bilby , Calgary, AB: Edmonton, Lethbridge, Red Deer AB: Calgary; MB: Husavick Edmonton, Lethbridge, AB: Calgary, AB: Cochrane, Edmonton AB: Edmonton AB: Calgary St. Paul AB: Edmonton, Red Deer, AB: Calgary; SK: Roche Percee AB: Edmonton AB: Edmonton Edmonton, Lethbridge AB: Calgary, Edmonton , Lethbridge AB: Calgary, AB: Edmonton MB: Stead SK: Weyburn MB: Portage MB: Dauphin zone" AB, SK: "Transition AB: Edmonton; AB: Edmonton 14,N 9,H 5,N 9,N 89,N 14,N 88,N 89,N 14,N 5,83,N 5,N 9,N 5,N 14,N 89,N 14,N 5,89,N 87,N

9,N 83,N 83,H 83,N 83,89,N

5,N 14,N 5,N

2,14,N 5,9,92,N

9,14,N 90,N

5,N 5,N 5,N 5,N 5,N 5,9,13,N 5,91,N Cushman, 1931 (Ashmead, 1890) Heinrich, 1961 (Cresson, 1864) (Cresson, 1867) (Cushman, 1925) Heinrich, 1962 (Cresson, 1865) (Walkley, 1958) (Walkley, a 83,N 5,83,N 9,N 83,N 5,9,83,N Diphyus hudsonicus (Cresson, 1877) 1877) Coelichneumon orpheus (Cresson, 1864) Cratichneumon rubricus (Provancher, Cratichneumon vescus (Provancher, Cratichneumon viator (Scopoli, 1763) 1886) (Provancher, marginatus Diadromus SUBFAMILY Species 5,N 1882) 1874) Cyclolabus signatus (Provancher, 1874) 1877) 1825) Conocalama manitobae Heinrich, 1962 Conocalama occidentalis (Cresson, 1868) Conocalama quebecensis (Provancher, 1875) Cratichneumon pilosulus (Provancher, Cratichneumon popofensis Ashmead, 1902 (Cresson, Cratichneumon promptus Cratichneumon scitulus (Cresson, 1864) Cratichneumon unifasciatorius (Say, Ctenichneumon caeruleops Heinrich, 1961 Ctenichneumon excultus (Cresson, 1867) Ctenichneumon punctiscuta Heinrich, 1961 Ctenichneumon syphax (Cresson, 1864) Cyclolabus lobatus 1875) Dentilabus rufipes (Provancher, Diadromus subtilicornis Diphyus allapsus (Cresson, 1865) Diphyus animosus (Cresson, 1864) Diphyus apiculatus Diphyus comes (Cresson, 1864) 9,N Diphyus discus (Cresson, 1864) *Diphyus euxoae Heinrich, 1969 Diphyus macilentus Diphyus nuncius (Cresson, 1877) Diphyus ormenus (Cresson, 1864) Diphyus populorum (Heinrich, 1961) Diphyus provancheri (Cresson, 1867) Diphyus robustus Diphyus rubellus (Cresson, 1865) Diphyus subfuscus (Cresson, 1864) 1882 gaspesianus Provancher, Dirophanes Dirophanes hariolus Ectopimorpha hiulcops Ectopimorpha luperinae Ectopimorpha wilsoni (Cresson, 1867) Eutanyacra improvisa Townes, 1944 Cratichneumon pteridis Townes, helvolus (Cresson, 1867) Diadromus Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 363 Loopers Loopers Loopers Concealed feeders Cutworms Cutworms Cutworms Cutworms Concealed feeders Host Family (No. spp.) Guild/Notes Lepidoptera: Geometridae, Notodontidae (4) Mostly loopers Lepidoptera: Erebidae, Noctuidae, Saturniidae, Sesiidae (11) Mostly setose Lepidoptera: Geometridae, Noctuidae (2) Lepidoptera: Noctuidae (3) Lepidoptera: Erebidae, Noctuidae (4) Mostly setose None Lepidoptera: Sphingidae (1) Lepidoptera: Saturniidae (1) None None Lepidoptera: Geometridae (1) None Lepidoptera: Pyralidae (1) Hymenoptera: Diprionidae, Ichneumonidae;Lepidoptera: Erebidae, Geometridae, Lasiocampidae, Noctuidae, Pieridae (28) None None None None Lepidoptera: Geometridae (1) None Lepidoptera: Geometridae (3) (2) Tortricidae Lepidoptera: None Acrolepiidae, Coleoptera: Cerambycidae; Lepidoptera: Noctuidae, Pyralidae (4) None None Lepidoptera: Noctuidae (1) Lepidoptera: Coleophoridae, Noctuidae (3) None None None Lepidoptera: Noctuidae (2) None Lepidoptera: Noctuidae (1) None None None Lepidoptera: Noctuidae (3) (2) Tortricidae Lepidoptera: Geometridae, Conifer feeders Tortricidae Hymenoptera: Diprionidae; Lepidoptera: (10) Mostly Tortricidae None Lepidoptera: Noctuidae (1) None Lepidoptera: Noctuidae (5) MB:

; Wabamun Wabamun m , Red Deer, Vermilion, Vermilion, Gull L , Red Deer, m m b Southwest Localities Aweme AB: Edmonton; MB: MB: Aweme Aweme MB: Waterton; AB: Calgary, SK: Albert; Prince MB: Aweme AB: Edmonton AB: Edmonton Treesbank AB: Edmonton; MB: corner MB: 2 localities in SW MB: Glenlea AB: Edmonton AB: nr Calgary AB: Edmonton, Several localities in southwest SK and southeast MB SK: Saskatoon AB: Calgary MB: Aweme AB: Drumheller AB: Edmonton AB: Cochrane, Edmonton AB: Fawcett NA Wagner AB: Edmonton, AB: Lethbridge MB: Swan River AB: Edmonton Edmonton, Lethbridge AB: Bilby , Calgary, AB: Edmonton, Lethbridge, Red Deer AB: Calgary; MB: Husavick Edmonton, Lethbridge, AB: Calgary, AB: Cochrane, Edmonton AB: Edmonton AB: Calgary St. Paul AB: Edmonton, Red Deer, AB: Calgary; SK: Roche Percee AB: Edmonton AB: Edmonton Edmonton, Lethbridge AB: Calgary, Edmonton , Lethbridge AB: Calgary, AB: Edmonton MB: Stead SK: Weyburn MB: Portage MB: Dauphin zone" AB, SK: "Transition AB: Edmonton; AB: Edmonton 14,N 9,H 5,N 9,N 89,N 14,N 88,N 89,N 14,N 5,83,N 5,N 9,N 5,N 14,N 89,N 14,N 5,89,N 87,N

9,N 83,N 83,H 83,N 83,89,N

5,N 14,N 5,N

2,14,N 5,9,92,N

9,14,N 90,N

5,N 5,N 5,N 5,N 5,N 5,9,13,N 5,91,N Cushman, 1931 (Ashmead, 1890) Heinrich, 1961 (Cresson, 1864) (Cresson, 1867) (Cushman, 1925) Heinrich, 1962 (Cresson, 1865) (Walkley, 1958) (Walkley, a 83,N 5,83,N 9,N 83,N 5,9,83,N Diphyus hudsonicus (Cresson, 1877) 1877) Coelichneumon orpheus (Cresson, 1864) Cratichneumon rubricus (Provancher, Cratichneumon vescus (Provancher, Cratichneumon viator (Scopoli, 1763) 1886) (Provancher, marginatus Diadromus SUBFAMILY Species 5,N 1882) 1874) Cyclolabus signatus (Provancher, 1874) 1877) 1825) Conocalama manitobae Heinrich, 1962 Conocalama occidentalis (Cresson, 1868) Conocalama quebecensis (Provancher, 1875) Cratichneumon pilosulus (Provancher, Cratichneumon popofensis Ashmead, 1902 (Cresson, Cratichneumon promptus Cratichneumon scitulus (Cresson, 1864) Cratichneumon unifasciatorius (Say, Ctenichneumon caeruleops Heinrich, 1961 Ctenichneumon excultus (Cresson, 1867) Ctenichneumon punctiscuta Heinrich, 1961 Ctenichneumon syphax (Cresson, 1864) Cyclolabus lobatus 1875) Dentilabus rufipes (Provancher, Diadromus subtilicornis Diphyus allapsus (Cresson, 1865) Diphyus animosus (Cresson, 1864) Diphyus apiculatus Diphyus comes (Cresson, 1864) 9,N Diphyus discus (Cresson, 1864) *Diphyus euxoae Heinrich, 1969 Diphyus macilentus Diphyus nuncius (Cresson, 1877) Diphyus ormenus (Cresson, 1864) Diphyus populorum (Heinrich, 1961) Diphyus provancheri (Cresson, 1867) Diphyus robustus Diphyus rubellus (Cresson, 1865) Diphyus subfuscus (Cresson, 1864) 1882 gaspesianus Provancher, Dirophanes Dirophanes hariolus Ectopimorpha hiulcops Ectopimorpha luperinae Ectopimorpha wilsoni (Cresson, 1867) Eutanyacra improvisa Townes, 1944 Cratichneumon pteridis Townes, helvolus (Cresson, 1867) Diadromus 364 M. D. Schwarzfeld Spiny Loopers Loopers Concealed feeders Cutworms Cutworms Cutworms Cutworms Concealed feeders Lepidoptera: Noctuidae (1) Host Family (No. spp.) Guild/Notes Lepidoptera: Geometridae (4) Lepidoptera: Erebidae (1) Setose Lepidoptera: Cossidae, Hepialidae, Sesiidae (5) Borers Lepidoptera: Pyralidae (3) Lepidoptera: Crambidae, Erebidae, Noctuidae (8) Nymphalidae, Sesiidae (4) Lepidoptera: Lycaenidae, Lepidoptera: Noctuidae (8) None None Lepidoptera: Pterophoridae (1) None None Lepidoptera: Geometridae, Noctuidae (7) Mostly loopers Lepidoptera: Noctuidae (9) Lepidoptera: Coleophoridae, Noctuidae (2) None None None Lepidoptera: Erebidae, Noctuidae (3) None None Lepidoptera: Sesiidae (1) Borers None None Lepidoptera: Geometridae, Hepialidae, Noctuidae (4) None (1) Tortricidae Lepidoptera: None None None Lepidoptera: Erebidae, Geometridae, Nymphalidae (3) Lepidoptera: Noctuidae (1) Lepidoptera: Erebidae (2) Setose lichen feeders None None None None None None None Lepidoptera: Geometridae (1) Lepidoptera: Noctuidae (1) None None Lepidoptera: Erebidae (1) Lepidoptera: Noctuidae (1) Lepidoptera: Nymphalidae (6) m ; m

m Altona , m Winnipeg ; MB: Franklin ; MB: 1 locality m m Grosse Isle Grosse , Edmonton, Red Deer; SK: nr Prince Albert , Edmonton, Red Deer; SK: nr Prince m Altona , Glenlea, m m b in southwest Localities Lethbridge, Edmonton; MB: Drumheller, AB: Calgary, Lenswood , Swan River MB: Glenlea AB: Edmonton AB: Edmonton Sprucewoods, five localities in south MB Treesbank, MB: nr Calgary AB: Bilby, SK: Saskatoon; MB: Beulah SK: Secretan, Swift Current AB: Edmonton AB: Medicine Hat; MB: Birtle AB: Cochrane, Edmonton L AB: Waterton MB: Aweme Aweme AB: Cochrane; MB: AB: Edmonton; MB: SK: nr Lloydminster; MB: 2 localities in southwest AB: Edmonton AB: Edmonton Dauphin L Aweme, MB: AB: Edmonton; SK: 1 locality in southeast AB: Edmonton; SK: 1 locality in southeast Albert, 1 locality in southwest AB: 1 locality in southeast; SK: nr Prince MB: Swan River AB: Edmonton AB: Bilby , Edmonton Aweme AB: Midnapore; MB: AB: Edmonton AB: Edmonton MB: Dauphin L Edmonton AB: Calgary, AB: Cypress Hills, Edmonton Edmonton, Red Deer; MB: AB: Bilby , Calgary, Edmonton, Lethbridge AB: Bilby , Calgary, AB: Edmonton Aweme MB: Wabamun; AB: Edmonton, AB: Edmonton AB: Edmonton Edmonton AB: Calgary, AB: Edmonton AB: Edmonton, Lethbridge MB: Dauphin L AB: Edmonton AB: Calgary AB: Calgary; MB: Ninette AB: Edmonton MB: Swan River AB: Edmonton SK: nr Regina MB: Swan River , Winnipeg 85,N 85,N 5,N 5,N 14,H 5,N 5,83,N 83,94,N 85,N 85,N 5,93,N 85,N 85,N 85,N 14,N 84,N 83,N 5,9,17,N 5,N 5,N 9,N 84,N 5,N 9,N 5,N 5,N 5,N 5,N 85,N 5,9,83,N 9,N 5,9,N 85,N 5,9,N 5,9,83,95,N 5,N 5,N 5,9,N,T 5,N 5,N 85,N 5,9,N 5,N 83,N 5,83,N Heinrich, 1961 Heinrich, 1961 Heinrich, 1961 Heinrich, 1961 Heinrich, 1956 Heinrich, 1961 (Provancher, 1874) (Provancher, Heinrich, 1961 Cresson, 1877 Cresson, 1877 83, N (Say, 1835) (Say, Heinrich, 1961 Provancher, 1875 Provancher, r Say, 1825 r Say, Cresson, 1864 Davis, 1898 a 14,H,O 14,N Ichneumon devincto Eutanyacra suturalis 1829) Hoplismenus morulus (Say, 1875 Ichneumon cervulus Provancher, Ichneumon dioryctriae Ichneumon gracilicornops Ichneumon longulus Cresson, 1864 belangeri (Cresson, 1877) Limerodops SUBFAMILY Species Ichneumon eurypus Eutanyacra trivittata Exephanes terminalis 1875) (Provancher, Heterischnus huardi Homotherus porcelariae townesi Hybophorellus Hypomecus quadriannulatus (Gravenhorst, 1829) Ichneumon ambulatorius Fabricius, 1775 Ichneumon annulatorius Fabricius, 1775 1902) Ichneumon artemis (Viereck, Cresson, 1877 Ichneumon atrox 1877 Ichneumon bimembris Provancher, *Ichneumon canadensis Cresson, 1867 1825 Ichneumon centrator Say, Heinrich, 1956 Ichneumon cessatorops Ichneumon chasmodops Heinrich, 1961 Cresson, 1867 Ichneumon creperus Ichneumon faciens Ichneumon feralis Cresson, 1867 Ichneumon feriens Heinrich, 1961 Ichneumon gestuosus Cresson, 1877 Ichneumon hippisleyae Ichneumon inurbanus Cresson, 1867 1875 Ichneumon lachrymans Provancher, Ichneumon laetus Brulle, 1846 Ichneumon maius Cresson, 1867 Heinrich, 1961 Ichneumon microferiens (Provancher, Ichneumon nigrovariegatus 1874) Ichneumon parvus Cresson, 1864 Ichneumon pedalis Cresson, 1864 Ichneumon pictifrons Ichneumon placidus Ichneumon pseudowinkleyi Ichneumon putus Cresson, 1877 Ichneumon suburbanus Heinrich, 1961 Ichneumon uncinatus Ichneumon ultimus Cresson, 1877 Ichneumon versabilis Ichneumon subdolus Cresson, 1867 Hoplismenus rutilus (Cresson, 1864) Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 365 Spiny Loopers Loopers Concealed feeders Cutworms Cutworms Cutworms Cutworms Concealed feeders Lepidoptera: Noctuidae (1) Host Family (No. spp.) Guild/Notes Lepidoptera: Geometridae (4) Lepidoptera: Erebidae (1) Setose Lepidoptera: Cossidae, Hepialidae, Sesiidae (5) Borers Lepidoptera: Pyralidae (3) Lepidoptera: Crambidae, Erebidae, Noctuidae (8) Nymphalidae, Sesiidae (4) Lepidoptera: Lycaenidae, Lepidoptera: Noctuidae (8) None None Lepidoptera: Pterophoridae (1) None None Lepidoptera: Geometridae, Noctuidae (7) Mostly loopers Lepidoptera: Noctuidae (9) Lepidoptera: Coleophoridae, Noctuidae (2) None None None Lepidoptera: Erebidae, Noctuidae (3) None None Lepidoptera: Sesiidae (1) Borers None None Lepidoptera: Geometridae, Hepialidae, Noctuidae (4) None (1) Tortricidae Lepidoptera: None None None Lepidoptera: Erebidae, Geometridae, Nymphalidae (3) Lepidoptera: Noctuidae (1) Lepidoptera: Erebidae (2) Setose lichen feeders None None None None None None None Lepidoptera: Geometridae (1) Lepidoptera: Noctuidae (1) None None Lepidoptera: Erebidae (1) Lepidoptera: Noctuidae (1) Lepidoptera: Nymphalidae (6) m ; m m Altona , m Winnipeg ; MB: Franklin ; MB: 1 locality m m Grosse Isle Grosse , Edmonton, Red Deer; SK: nr Prince Albert , Edmonton, Red Deer; SK: nr Prince m Altona , Glenlea, m m b in southwest Localities Lethbridge, Edmonton; MB: Drumheller, AB: Calgary, Lenswood , Swan River MB: Glenlea AB: Edmonton AB: Edmonton Sprucewoods, five localities in south MB Treesbank, MB: nr Calgary AB: Bilby, SK: Saskatoon; MB: Beulah SK: Secretan, Swift Current AB: Edmonton AB: Medicine Hat; MB: Birtle AB: Cochrane, Edmonton L AB: Waterton MB: Aweme Aweme AB: Cochrane; MB: AB: Edmonton; MB: SK: nr Lloydminster; MB: 2 localities in southwest AB: Edmonton AB: Edmonton Dauphin L Aweme, MB: AB: Edmonton; SK: 1 locality in southeast AB: Edmonton; SK: 1 locality in southeast Albert, 1 locality in southwest AB: 1 locality in southeast; SK: nr Prince MB: Swan River AB: Edmonton AB: Bilby , Edmonton Aweme AB: Midnapore; MB: AB: Edmonton AB: Edmonton MB: Dauphin L Edmonton AB: Calgary, AB: Cypress Hills, Edmonton Edmonton, Red Deer; MB: AB: Bilby , Calgary, Edmonton, Lethbridge AB: Bilby , Calgary, AB: Edmonton Aweme MB: Wabamun; AB: Edmonton, AB: Edmonton AB: Edmonton Edmonton AB: Calgary, AB: Edmonton AB: Edmonton, Lethbridge MB: Dauphin L AB: Edmonton AB: Calgary AB: Calgary; MB: Ninette AB: Edmonton MB: Swan River AB: Edmonton SK: nr Regina MB: Swan River , Winnipeg 85,N 85,N 5,N 5,N 14,H 5,N 5,83,N 83,94,N 85,N 85,N 5,93,N 85,N 85,N 85,N 14,N 84,N 83,N 5,9,17,N 5,N 5,N 9,N 84,N 5,N 9,N 5,N 5,N 5,N 5,N 85,N 5,9,83,N 9,N 5,9,N 85,N 5,9,N 5,9,83,95,N 5,N 5,N 5,9,N,T 5,N 5,N 85,N 5,9,N 5,N 83,N 5,83,N Heinrich, 1961 Heinrich, 1961 Heinrich, 1961 Heinrich, 1961 Heinrich, 1956 Heinrich, 1961 (Provancher, 1874) (Provancher, Heinrich, 1961 Cresson, 1877 Cresson, 1877 83, N (Say, 1835) (Say, Heinrich, 1961 Provancher, 1875 Provancher, r Say, 1825 r Say, Cresson, 1864 Davis, 1898 a 14,H,O 14,N Ichneumon devincto Eutanyacra suturalis 1829) Hoplismenus morulus (Say, 1875 Ichneumon cervulus Provancher, Ichneumon dioryctriae Ichneumon gracilicornops Ichneumon longulus Cresson, 1864 belangeri (Cresson, 1877) Limerodops SUBFAMILY Species Ichneumon eurypus Eutanyacra trivittata Exephanes terminalis 1875) (Provancher, Heterischnus huardi Homotherus porcelariae townesi Hybophorellus Hypomecus quadriannulatus (Gravenhorst, 1829) Ichneumon ambulatorius Fabricius, 1775 Ichneumon annulatorius Fabricius, 1775 1902) Ichneumon artemis (Viereck, Cresson, 1877 Ichneumon atrox 1877 Ichneumon bimembris Provancher, *Ichneumon canadensis Cresson, 1867 1825 Ichneumon centrator Say, Heinrich, 1956 Ichneumon cessatorops Ichneumon chasmodops Heinrich, 1961 Cresson, 1867 Ichneumon creperus Ichneumon faciens Ichneumon feralis Cresson, 1867 Ichneumon feriens Heinrich, 1961 Ichneumon gestuosus Cresson, 1877 Ichneumon hippisleyae Ichneumon inurbanus Cresson, 1867 1875 Ichneumon lachrymans Provancher, Ichneumon laetus Brulle, 1846 Ichneumon maius Cresson, 1867 Heinrich, 1961 Ichneumon microferiens (Provancher, Ichneumon nigrovariegatus 1874) Ichneumon parvus Cresson, 1864 Ichneumon pedalis Cresson, 1864 Ichneumon pictifrons Ichneumon placidus Ichneumon pseudowinkleyi Ichneumon putus Cresson, 1877 Ichneumon suburbanus Heinrich, 1961 Ichneumon uncinatus Ichneumon ultimus Cresson, 1877 Ichneumon versabilis Ichneumon subdolus Cresson, 1867 Hoplismenus rutilus (Cresson, 1864) 366 M. D. Schwarzfeld Loopers Loopers Loopers Loopers Loopers Loopers Loopers Concealed feeders Cutworms Host Family (No. spp.) Guild/Notes (6) Tortricidae Lepidoptera: Erebidae, Mostly setose None Lepidoptera: Coleophoridae, Noctuidae (3) Lepidoptera: Noctuidae (8) (3) Tortricidae Lepidoptera: Erebidae, Geometridae, Lepidoptera: Noctuidae, Nymphalidae (5) Mostly spiny Lepidoptera: Erebidae, Nymphalidae (3) Setose or spiny None Lepidoptera: Geometridae (1) (2) Lepidoptera: Lycaenidae Hymenoptera: Ichneumonidae; Lepidoptera: Sphingidae (10) Lycaenidae, None None None Lepidoptera: Plutellidae, Pterophoridae (2) Concealed feeders (9) Tortricidae Lepidoptera: Geometridae, Noctuidae, Mostly loopers (7) Tortricidae Lepidoptera: Noctuidae, Conifer feeders Tortricidae Hymenoptera: Diprionidae; Lepidoptera: (3) Conifer feeders (2) Tortricidae Lepidoptera: None None Lepidoptera: Geometridae (3) Lepidoptera: Geometridae (1) Lepidoptera: Geometridae (4) Lepidoptera: Geometridae (1) None Lepidoptera: Geometridae (2) Lepidoptera: Geometridae (1) None Lepidoptera: Saturniidae (1) None Hymenoptera: Diprionidae; Lepidoptera: Crambidae, Erebidae, Noctuidae (6) Lepidoptera: Noctuidae, Sesiidae (3) Lepidoptera: Noctuidae (1) None Lepidoptera: Erebidae, Lasiocampidae, Noctuidae, Nymphalidae, Pieridae (9) None None Lepidoptera: Nymphalidae, Pieridae (5) Lepidoptera: Notodontidae, Nymphalidae (3) Setose or spiny SK, MB: across Winnipeg

m , Swan River ; m Birtle Domain , Minnedosa m Glenlea several south and central localities;

Winnipeg m , Vermilion; MB: , Vermilion; Wabamun m m b Localities AB: Vermilion MB: Treesbank AB: Edmonton AB: Edmonton AB: Lethbridge Winnipeg AB: Calgary; MB: SK: White Fox Albert SK: nr Prince SK: Indian Head Waterton AB: Edmonton, Red Deer, AB: Cypress Hills NA Wagner AB: Vermilion, Edmonton, AB: Edmonton; SK: Glaslyn AB: Edmonton SK: White Fox AB: Edmonton Lakes NP Waterton AB: AB: 1 locality in southwest AB: Edmonton AB: Edmonton AB: Breton, Vermilion SK: Fort-à-la-Corne MB: Russell Edmonton; MB: AB: Calgary, NA AB: Wagner MB: Altona SK: Tyner AB: Cypress Hills, Medicine Hat; MB: Cypress Hills, Edmonton, Lethbridge; AB: Calgary, Cochrane, Cypress Hills AB: Calgary, AB: Calahoo, Edmonton, Lethbridge; MB: MB: Altona , Roland Rosa AB: Bilby , Edmonton, AB: Cypress Hills, Edmonton, Medicine Hat AB: Edmonton Albert; MB: 2 localities nr Dauphin SK: nr Prince AB: Edmonton, Lethbridge Edmonton, Lethbridge; MB: AB: Calgary, AB: Cypress Hills, Edmonton, Red Deer SK: nr Regina; MB: several localities in south MB: 1 locality in southwest AB, SK, MB Across southern ; Winnipeg Across southern AB, SK, MB Across southern southern part 2,N 5,N 5,9,H 9,14,N 87,N 5,83,N 84,N

86,N 5,9,N 5,83,N

5,20,N 9,N,T 2,5,N 5,N 83,N 96,N 97,N 83,N 83,N 83,N 14,N 14,N 83,N 14,83,N 84,N 5,H 83,H (Cresson, (Provancher, (Provancher, (Viereck, (Viereck, Heinrich, Heinrich, (Cresson, (Thunberg, (Thunberg, (Provancher, (Provancher, Provancher, Provancher, (Schrank, 1802) Heinrich, 1961 Viereck, 1924 Viereck, Cushman, 1920 (Gravenhorst, (Cresson, 1867) Heinrich, 1968 Heinrich, 1962 Heinrich, 1962 (Cresson, 1867) Heinrich, 1961 Heinrich, 1962 (Cresson, 1867) (Cresson, 1867) a 14,N 14,H,O 94,N 83,H 5,N 5,83,N 5,6,9,13,83,N,Oc 5,N 5,N 5,9,83,N,Oc 83,84,N 14,N 9,N,Oc 1961 Spilichneumon taos (Cresson, 1877) Melanichneumon absconditus vincibilis Oronotus Phaeogenes hebe Spilichneumon inconstans 1864) Spilichneumon nubivagus (Cresson, 1867) Thyrateles instabilis Thyrateles mormonus (Cresson, 1877) SUBFAMILY Species 1886) Platylabus rubricapensis 1882 1902) Melanichneumon flavicarina Neotypus nobilitator 1807) Netanyacra dacotae Obtusodonta manitobae Heinrich, 1962 Obtusodonta obscuricolor (Heinrich, 1961) calcatorius Orgichneumon montanus (Cresson, 1864) Patrocloides perluctuosus Patrocloides Phaeogenes arcticus Phaeogenes cacoeciae Platylabops pecki Platylabus clarus bakeri (Davis, 1898) Pristicerops grandis (Brulle, 1846) Protichneumon Heinrich, 1962 Rubicundiella deuteromelas Rubicundiella perturbatrix 1962 Rubicundiella simplicior Heinrich, 1961 (Cresson, 1864) Spilichneumon bronteus *Spilichneumon superbus (Provancher, 1886) Stenichneumon culpator Stenichneumon pallidipennis Stenobarichneumon saundersii 1877) Syspasis tauma (Heinrich, 1951) Thyrateles lugubrator (Gravenhorst, 1807) (Cresson, 1877) Thyrateles procax 1824) 1877) Uchida, 1930 Neotypus coreensis Platylabus foxleei Platylabus imitans Platylabus luteatae lascivus (Cresson, 1867) Pristiceros effigies Protichneumon Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 367 Loopers Loopers Loopers Loopers Loopers Loopers Loopers Concealed feeders Cutworms Host Family (No. spp.) Guild/Notes (6) Tortricidae Lepidoptera: Erebidae, Mostly setose None Lepidoptera: Coleophoridae, Noctuidae (3) Lepidoptera: Noctuidae (8) (3) Tortricidae Lepidoptera: Erebidae, Geometridae, Lepidoptera: Noctuidae, Nymphalidae (5) Mostly spiny Lepidoptera: Erebidae, Nymphalidae (3) Setose or spiny None Lepidoptera: Geometridae (1) (2) Lepidoptera: Lycaenidae Hymenoptera: Ichneumonidae; Lepidoptera: Sphingidae (10) Lycaenidae, None None None Lepidoptera: Plutellidae, Pterophoridae (2) Concealed feeders (9) Tortricidae Lepidoptera: Geometridae, Noctuidae, Mostly loopers (7) Tortricidae Lepidoptera: Noctuidae, Conifer feeders Tortricidae Hymenoptera: Diprionidae; Lepidoptera: (3) Conifer feeders (2) Tortricidae Lepidoptera: None None Lepidoptera: Geometridae (3) Lepidoptera: Geometridae (1) Lepidoptera: Geometridae (4) Lepidoptera: Geometridae (1) None Lepidoptera: Geometridae (2) Lepidoptera: Geometridae (1) None Lepidoptera: Saturniidae (1) None Hymenoptera: Diprionidae; Lepidoptera: Crambidae, Erebidae, Noctuidae (6) Lepidoptera: Noctuidae, Sesiidae (3) Lepidoptera: Noctuidae (1) None Lepidoptera: Erebidae, Lasiocampidae, Noctuidae, Nymphalidae, Pieridae (9) None None Lepidoptera: Nymphalidae, Pieridae (5) Lepidoptera: Notodontidae, Nymphalidae (3) Setose or spiny SK, MB: across Winnipeg

m , Swan River ; m Birtle Domain , Minnedosa m Glenlea several south and central localities;

Winnipeg m , Vermilion; MB: , Vermilion; Wabamun m m b Localities AB: Vermilion MB: Treesbank AB: Edmonton AB: Edmonton AB: Lethbridge Winnipeg AB: Calgary; MB: SK: White Fox Albert SK: nr Prince SK: Indian Head Waterton AB: Edmonton, Red Deer, AB: Cypress Hills NA Wagner AB: Vermilion, Edmonton, AB: Edmonton; SK: Glaslyn AB: Edmonton SK: White Fox AB: Edmonton Lakes NP Waterton AB: AB: 1 locality in southwest AB: Edmonton AB: Edmonton AB: Breton, Vermilion SK: Fort-à-la-Corne MB: Russell Edmonton; MB: AB: Calgary, NA AB: Wagner MB: Altona SK: Tyner AB: Cypress Hills, Medicine Hat; MB: Cypress Hills, Edmonton, Lethbridge; AB: Calgary, Cochrane, Cypress Hills AB: Calgary, AB: Calahoo, Edmonton, Lethbridge; MB: MB: Altona , Roland Rosa AB: Bilby , Edmonton, AB: Cypress Hills, Edmonton, Medicine Hat AB: Edmonton Albert; MB: 2 localities nr Dauphin SK: nr Prince AB: Edmonton, Lethbridge Edmonton, Lethbridge; MB: AB: Calgary, AB: Cypress Hills, Edmonton, Red Deer SK: nr Regina; MB: several localities in south MB: 1 locality in southwest AB, SK, MB Across southern ; Winnipeg Across southern AB, SK, MB Across southern southern part 2,N 5,N 5,9,H 9,14,N 87,N 5,83,N 84,N

86,N 5,9,N 5,83,N

5,20,N 9,N,T 2,5,N 5,N 83,N 96,N 97,N 83,N 83,N 83,N 14,N 14,N 83,N 14,83,N 84,N 5,H 83,H (Cresson, (Provancher, (Provancher, (Viereck, (Viereck, Heinrich, Heinrich, (Cresson, (Thunberg, (Thunberg, (Provancher, (Provancher, Provancher, Provancher, (Schrank, 1802) Heinrich, 1961 Viereck, 1924 Viereck, Cushman, 1920 (Gravenhorst, (Cresson, 1867) Heinrich, 1968 Heinrich, 1962 Heinrich, 1962 (Cresson, 1867) Heinrich, 1961 Heinrich, 1962 (Cresson, 1867) (Cresson, 1867) a 14,N 14,H,O 94,N 83,H 5,N 5,83,N 5,6,9,13,83,N,Oc 5,N 5,N 5,9,83,N,Oc 83,84,N 14,N 9,N,Oc 1961 Spilichneumon taos (Cresson, 1877) Melanichneumon absconditus vincibilis Oronotus Phaeogenes hebe Spilichneumon inconstans 1864) Spilichneumon nubivagus (Cresson, 1867) Thyrateles instabilis Thyrateles mormonus (Cresson, 1877) SUBFAMILY Species 1886) Platylabus rubricapensis 1882 1902) Melanichneumon flavicarina Neotypus nobilitator 1807) Netanyacra dacotae Obtusodonta manitobae Heinrich, 1962 Obtusodonta obscuricolor (Heinrich, 1961) calcatorius Orgichneumon montanus (Cresson, 1864) Patrocloides perluctuosus Patrocloides Phaeogenes arcticus Phaeogenes cacoeciae Platylabops pecki Platylabus clarus bakeri (Davis, 1898) Pristicerops grandis (Brulle, 1846) Protichneumon Heinrich, 1962 Rubicundiella deuteromelas Rubicundiella perturbatrix 1962 Rubicundiella simplicior Heinrich, 1961 (Cresson, 1864) Spilichneumon bronteus *Spilichneumon superbus (Provancher, 1886) Stenichneumon culpator Stenichneumon pallidipennis Stenobarichneumon saundersii 1877) Syspasis tauma (Heinrich, 1951) Thyrateles lugubrator (Gravenhorst, 1807) (Cresson, 1877) Thyrateles procax 1824) 1877) Uchida, 1930 Neotypus coreensis Platylabus foxleei Platylabus imitans Platylabus luteatae lascivus (Cresson, 1867) Pristiceros effigies Protichneumon 368 M. D. Schwarzfeld

Host Family (No. spp.) Guild/Notes Lepidoptera: Erebidae (1) Lepidoptera: Papilionidae (6) None Lepidoptera: Nymphalidae, Papilionidae (8) Lepidoptera: Tenthredinidae; Hymenoptera: Erebidae (5) Mostly setose None Lepidoptera: Crambidae, Erebidae, Geometridae, Noctuidae, Notodontidae (16) (7) Tortricidae Lepidoptera: Gelechiidae, Pyralidae, Concealed feeders None None None Hymenoptera: Braconidae, Ichneumonidae, Lepidoptera: Geometridae (7) Tenthredinidae; None Hymenoptera: Braconidae, Diprionidae, Ichneumonidae; Lepidoptera: Erebidae, Geometridae, Lasiocampidae, Noctuidae (33) None Coleoptera: Curculionidae; Hymenoptera: Braconidae, Ichneumonidae; Lepidoptera: Erebidae (5) Lepidoptera: Tenthredinidae; Hymenoptera: Geometridae (4) None None None None None Coleoptera: Chrysomelidae; Hymenoptera: Braconidae; Lepidoptera: Crambidae, Geometridae, Noctuidae, Pieridae (11) None Argidae Hymenoptera: (1) Hymenoptera: Diprionidae, Ichneumonidae; Lepidoptera: Notodontidae (10) Tenthredinidae; Hymenoptera: Braconidae; Lepidoptera: Bucculatricidae, Erebidae, Geometridae (82) Mostly Braconidae None Lepidoptera: Geometridae (1) Hymenoptera: Cimbicidae, Ichneumonidae (2) None Coleoptera: Chrysomelidae (1) Hymenoptera: Braconidae; Lepidoptera: Crambidae, Gelechiidae, Geometridae, Lasiocampidae, Lycaenidae, (21) Tortricidae Hymenoptera: Braconidae; Lepidoptera: Bucculatricidae, Crambidae, Erebidae, Geometridae, Noctuidae, Pieridae, Sphingidae (27)

; m Franklin , Swan River m Swan River m b Localities AB: Edmonton AB: Edmonton; MB: 2 southern localities AB, SK, MB: across Across southern Aweme; MB: Waterton; AB: southern Canada NP Waterton Nevis, AB: Buck Mt, Drumheller, MB: nr Brandon, west of Brandon AB: Red Deer AB: Edmonton; MB: Glenlea MB: Aweme L AB: George L AB: George NA AB: Wagner MB: Gilbert Plains, Minitonas MB: Ninette NA; Wagner Wabamun, L, George AB: Cooking L, Edmonton, Elkwater, SK: Calder; MB: Ninette, NA; SK: Broadview, Wagner Edmonton, Elkwater P, AB: Bilby, White Fox; MB: Albert, Prince Esterhazy, L, Lethbridge, Onefour, Edmonton, Elkwater L, George AB: Bilby, Lakes NP; SK: Great Sand Hills, Swift Current, Saskatoon, Waterton Shilo White Fox; MB: 5 mi SW near Swift Current Yorkton, NA; SK: Wagner AB: Edmonton, AB: Edmonton White Fox NA; SK: Wagner AB: MB: Aweme Attons L, Swift Current NA; SK: Wagner AB: Edmonton, Aweme L; MB: AB: George Shilo NA; SK: Saskatoon; MB: 5 mi SW Wagner AB: NA; MB: Brandon, Glenlea Wagner AB: Elkwater L, Lethbridge, NA Wagner AB: Wabamun, AB: Edmonton AB: Bilby AB: Gleichen, Moon L; MB: Gull L Calgary), W L, Jumping Pond Cr (20 mi AB: Edmonton, George AB: Edmonton SK: Saskatoon Wayne; NA, Wagner AB: Edmonton, Elkwater L, Teulon AB: Bilby; SK: Estevan; MB: AB: Onefour; SK: Saskatoon AB: Cochrane Wagner NA Wagner , 5 mi SW Shilo MB: Dauphin, Lenswood , 5 mi SW 5,17,N 91,N 100,H 2,100,N 5,N 2,9,100,H 5,N 100,H 2,5,9,100,H 5,83,N 100,N 100,N 100,N 100,N 2,100,N 100,N 100,N 100,H,T,A,O 2,100,N 99,H,O 2,N 100,N 100,N 2,100,N 100,N 2,100,N

100,H,T 2,100,N 100,N 5,97,98,N (Say, 1825) (Say, Dasch, 1971 (Cresson, 1877) Dasch, 1971 Cushman, Dasch, 1971 (Cresson, 1867) (Provancher, 1877) (Provancher, Cresson, 1868 a 83,N 100,H 2,100,102,H,T 2,5,9,100,101,N Tricholabus citatus Tricholabus fulvipes Trogus lapidator (Fabricius, 1787) Trogus SUBFAMILY Species Mesochorus americanus Cresson, 1872 Tricholabus nortonii Tricholabus subcyaneus (Cresson, Virgichneumon 1864) texanus Virgichneumon brevicinctor *Vulgichneumon LYCORININAE (Cresson, 1870) albomarginata Lycorina MESOCHORINAE alsium Dasch, 1971 Astiphromma Dasch, 1971 cordatum Astiphromma exitiale Astiphromma leucogrammum (Holmgren, Astiphromma 1860) Dasch, 1971 perditum Astiphromma splenium (Curtis, 1833) Astiphromma Cidaphus occidentalis 1924 Mesochorus agilis Cresson, 1865 1883 Provancher, Mesochorus areolatus Mesochorus bellus Dasch, 1971 1917 Mesochorus calais Viereck, Dasch, 1971 Mesochorus coronatus Dasch, 1971 Mesochorus cupreatus *Mesochorus curvulus Thomson, Mesochorus deletus Dasch, 1971 Mesochorus dentatus Dasch, 1971 Mesochorus dimidiatus Holmgren, 1860 1835) (Say, Mesochorus discitergus Mesochorus dreisbachi Mesochorus erythraeus Dasch, 1971 Mesochorus exsertus Dasch, 1971 Mesochorus foersteri Dasch, 1971 Mesochorus fuliginatus Mesochorus gemellus Holmgren, 1860 1886 Mesochorus applanatus Dasch, 1971 Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 369

Host Family (No. spp.) Guild/Notes Lepidoptera: Erebidae (1) Lepidoptera: Papilionidae (6) None Lepidoptera: Nymphalidae, Papilionidae (8) Lepidoptera: Tenthredinidae; Hymenoptera: Erebidae (5) Mostly setose None Lepidoptera: Crambidae, Erebidae, Geometridae, Noctuidae, Notodontidae (16) (7) Tortricidae Lepidoptera: Gelechiidae, Pyralidae, Concealed feeders None None None Hymenoptera: Braconidae, Ichneumonidae, Lepidoptera: Geometridae (7) Tenthredinidae; None Hymenoptera: Braconidae, Diprionidae, Ichneumonidae; Lepidoptera: Erebidae, Geometridae, Lasiocampidae, Noctuidae (33) None Coleoptera: Curculionidae; Hymenoptera: Braconidae, Ichneumonidae; Lepidoptera: Erebidae (5) Lepidoptera: Tenthredinidae; Hymenoptera: Geometridae (4) None None None None None Coleoptera: Chrysomelidae; Hymenoptera: Braconidae; Lepidoptera: Crambidae, Geometridae, Noctuidae, Pieridae (11) None Argidae Hymenoptera: (1) Hymenoptera: Diprionidae, Ichneumonidae; Lepidoptera: Notodontidae (10) Tenthredinidae; Hymenoptera: Braconidae; Lepidoptera: Bucculatricidae, Erebidae, Geometridae (82) Mostly Braconidae None Lepidoptera: Geometridae (1) Hymenoptera: Cimbicidae, Ichneumonidae (2) None Coleoptera: Chrysomelidae (1) Hymenoptera: Braconidae; Lepidoptera: Crambidae, Gelechiidae, Geometridae, Lasiocampidae, Lycaenidae, (21) Tortricidae Hymenoptera: Braconidae; Lepidoptera: Bucculatricidae, Crambidae, Erebidae, Geometridae, Noctuidae, Pieridae, Sphingidae (27)

; m Franklin , Swan River m Swan River m b Localities AB: Edmonton AB: Edmonton; MB: 2 southern localities AB, SK, MB: across Across southern Aweme; MB: Waterton; AB: southern Canada NP Waterton Nevis, AB: Buck Mt, Drumheller, MB: nr Brandon, west of Brandon AB: Red Deer AB: Edmonton; MB: Glenlea MB: Aweme L AB: George L AB: George NA AB: Wagner MB: Gilbert Plains, Minitonas MB: Ninette NA; Wagner Wabamun, L, George AB: Cooking L, Edmonton, Elkwater, SK: Calder; MB: Ninette, NA; SK: Broadview, Wagner Edmonton, Elkwater P, AB: Bilby, White Fox; MB: Albert, Prince Esterhazy, L, Lethbridge, Onefour, Edmonton, Elkwater L, George AB: Bilby, Lakes NP; SK: Great Sand Hills, Swift Current, Saskatoon, Waterton Shilo White Fox; MB: 5 mi SW near Swift Current Yorkton, NA; SK: Wagner AB: Edmonton, AB: Edmonton White Fox NA; SK: Wagner AB: MB: Aweme Attons L, Swift Current NA; SK: Wagner AB: Edmonton, Aweme L; MB: AB: George Shilo NA; SK: Saskatoon; MB: 5 mi SW Wagner AB: NA; MB: Brandon, Glenlea Wagner AB: Elkwater L, Lethbridge, NA Wagner AB: Wabamun, AB: Edmonton AB: Bilby AB: Gleichen, Moon L; MB: Gull L Calgary), W L, Jumping Pond Cr (20 mi AB: Edmonton, George AB: Edmonton SK: Saskatoon Wayne; NA, Wagner AB: Edmonton, Elkwater L, Teulon AB: Bilby; SK: Estevan; MB: AB: Onefour; SK: Saskatoon AB: Cochrane Wagner NA Wagner , 5 mi SW Shilo MB: Dauphin, Lenswood , 5 mi SW 5,17,N 91,N 100,H 2,100,N 5,N 2,9,100,H 5,N 100,H 2,5,9,100,H 5,83,N 100,N 100,N 100,N 100,N 2,100,N 100,N 100,N 100,H,T,A,O 2,100,N 99,H,O 2,N 100,N 100,N 2,100,N 100,N 2,100,N

100,H,T 2,100,N 100,N 5,97,98,N (Say, 1825) (Say, Dasch, 1971 (Cresson, 1877) Dasch, 1971 Cushman, Dasch, 1971 (Cresson, 1867) (Provancher, 1877) (Provancher, Cresson, 1868 a 83,N 100,H 2,100,102,H,T 2,5,9,100,101,N Tricholabus citatus Tricholabus fulvipes Trogus lapidator (Fabricius, 1787) Trogus SUBFAMILY Species Mesochorus americanus Cresson, 1872 Tricholabus nortonii Tricholabus subcyaneus (Cresson, Virgichneumon 1864) texanus Virgichneumon brevicinctor *Vulgichneumon LYCORININAE (Cresson, 1870) albomarginata Lycorina MESOCHORINAE alsium Dasch, 1971 Astiphromma Dasch, 1971 cordatum Astiphromma exitiale Astiphromma leucogrammum (Holmgren, Astiphromma 1860) Dasch, 1971 perditum Astiphromma splenium (Curtis, 1833) Astiphromma Cidaphus occidentalis 1924 Mesochorus agilis Cresson, 1865 1883 Provancher, Mesochorus areolatus Mesochorus bellus Dasch, 1971 1917 Mesochorus calais Viereck, Dasch, 1971 Mesochorus coronatus Dasch, 1971 Mesochorus cupreatus *Mesochorus curvulus Thomson, Mesochorus deletus Dasch, 1971 Mesochorus dentatus Dasch, 1971 Mesochorus dimidiatus Holmgren, 1860 1835) (Say, Mesochorus discitergus Mesochorus dreisbachi Mesochorus erythraeus Dasch, 1971 Mesochorus exsertus Dasch, 1971 Mesochorus foersteri Dasch, 1971 Mesochorus fuliginatus Mesochorus gemellus Holmgren, 1860 1886 Mesochorus applanatus Dasch, 1971 370 M. D. Schwarzfeld

Concealed feeders Concealed feeders Loopers Loopers Concealed feeders Mostly conifer loopers Concealed feeders Host Family (No. spp.) Guild/Notes Diprionidae, Elachistidae, Erebidae, Geometridae, (21) Mostly Tortricidae Noctuidae, Sphingidae, conifer feeders Hymenoptera: Braconidae, Ichneumonidae; Lepidoptera: Bucculatricidae, Choreutidae, Erebidae, Gelechiidae, Geometridae, Noctuidae, Nolidae, Pieridae, Plutellidae (78) (3) Tortricidae Lepidoptera: (8) Tortricidae Lepidoptera: Geometridae, Mostly concealed feeders Coleoptera: Chrysomelidae; Diptera: Tachinidae; Tachinidae; Coleoptera: Chrysomelidae; Diptera: Hymenoptera: Braconidae, Ichneumonidae, Lepidoptera: Gelechiidae, Noctuidae, Tenthredinidae; (43) Tortricidae Notodontidae, Psychidae, Saturniidae, None None Lepidoptera: Erebidae, Geometridae, Noctuidae (12) Mostly conifer loopers (1) Ypsolophidae Lepidoptera: None Hymenoptera: Diprionidae, Ichneumonidae, (7) Tenthredinidae Mostly conifer sawflies None None Hymenoptera, Lepidoptera: Braconidae, Crambidae, (5) Yponomeutidae Geometridae, Lepidoptera: Geometridae (1) None Erebidae, Tachinidae, Diptera, Lepidoptera: Geometridae (15) Mostly conifer loopers and Tachinidae Geometridae (2) Tachinidae, Diptera, Lepidoptera: Hymenoptera, Lepidoptera: Braconidae, None Hymenoptera: Ichneumonidae (1) Yponomeutidae Lepidoptera: Crambidae, Tortricidae, (4) Concealed feeders Diptera: Muscidae; Hymenoptera: Braconidae; Lepidoptera: Crambidae, Gracillariidae, Nolidae, Plutellidae (9) Lepidoptera: Gracillariidae (1) Lepidoptera: Tortricidae, Gracillariidae, (5) Yponomeutidae Hymenoptera: Braconidae; Lepidoptera: Gelechiidae, (4) Tortricidae Mostly concealed feeders None Lepidoptera: Geometridae (1) (3) Tortricidae Lepidoptera: Elachistidae, Concealed feeders

m m Swan River b Localities Wagner NA, Duffield, Edmonton, Nampa, Ponoka, Cr, AB: Buffalo FR, Woods Shilo, Spruce Lakes NP; MB: Ninette, 5 mi SW Waterton Mt Turtle AB: Elkwater L AB: Edmonton SK: Kindersley NA Wagner Cochrane, Edmonton, Elkwater, AB: Bilby, AB: Edmonton AB: Beaver Mines, Ferry Saskatchewan S of Stony Plain, Hayter, Provost, Red Deer Ranger Stn, Rivercours; SK: Canora, Cloan, Humboldt, Imperial; MB: Crandall, Hamiota, Mulvihill, Shoal L Transcona NA; MB: Wagner L, Lethbridge, George AB: Bilby, AB: Edmonton, Elkwater P Edmonton, Coaldale, Cypress Hills, Drumheller, AB: Calgary, Newell, Lethbridge, Manyberries, L Glenwood, Jumping Pound Cr, Oyen, Picture Butte, Rosebud, Scandia, Medicine Hat, Milk River, Horse; SK: Wild Lakes NP, Waterton NA, Wagner Vauxhall, Tilley, Cadillac, Consul, at Cut Knife, Battle River, Attons L Antelope L, Great Sand Hills, Indian Head, Minton, Dunblane, Estevan, Great Deer, Aweme, MB: Tompkins; Parkbeg, Regina, Saskatoon, Swift Current, Brandon, Morris AB: Edmonton AB: Edmonton AB: Edmonton, Elk Island P Mt FR Turtle in MB: Max L Winfield; AB: Belly River Ranger Stn, Edmonton, Lethbridge, Indian Head, Saskatoon SK: Barrier River, AB: Orion L, Lethbridge AB: George Edmonton, NP, L Waterton at Cameron L Burdett, Calgary, AB: Bilby, Rosemary, Onefour, L, Lethbridge, Lost River, George Elkwater, Moose SK: Dunblane, Great Deer, Winfield; Walsh, 21 mi S Stavely, Aweme, MB: Watrous; Albert, Saskatoon, Swift Current, Prince Mt P, Shilo, Starbuck , Brandon, 5 mi SW SK: Saskatoon Saskatoon AB: Lethbridge, Manyberries, Onefour; SK: Great Deer, AB: Edmonton, Elkwater L AB: Edmonton AB: Edmonton, Rockyford; SK: Cypress Hills Calgary W AB: Jumping Pd Cr 20 mi View) Ho (Mountain Westward AB: Gull L, nr Brooks, SK: Saskatoon; MB: 1 locality in southwest AB: Edmonton MB: 2 localities (southwest and southeast) 100,N

9,100,H,T 2,100,N 100,N

100,N 100,N

100,N 100,N,T,O 100,N 105,N 100,N 100,N 100,N 100,N 100,N

100,N 100,N,T 100,N 100,N 100,N 103,N Dasch, 1971 (Thomson, (Provancher, 1879) (Provancher, Ashmead, 1898 (Zetterstedt, 1838) Townes & Townes, & Townes, Townes a 2,100,N 100,N,T,A 103,104,105,N 104,105,N 104,N 2,14,53,100,N (Thunberg, 1822) Mesochorus giberius (Thunberg, Mesochorus validus Dasch, 1971 Dasch, 1971 Stictopisthus argaleus & Townes, Chorinaeus aequalis Townes 1959 SUBFAMILY Species 2,14,100,H,T,O Mesochorus uniformis Cresson, 1872 Mesochorus hesperus Dasch, 1971 Mesochorus inaequidens Dasch, 1971 Mesochorus incultus Dasch, 1971 Mesochorus longiscutatus Dasch, 1971 Mesochorus masoni Dasch, 1971 Mesochorus ottawaensis (Harrington, 1892) Mesochorus peltatus Dasch, 1971 *Mesochorus perniciosus Viereck, 1911 Mesochorus personatus Dasch, 1971 Mesochorus tachinae Mesochorus unicarinatus Dasch, 1971 Mesochorus varianus Dasch, 1971 Mesochorus vittator Stictopisthus bilineatus Dasch, 1971 Stictopisthus crenatus Stictopisthus flaviceps Stictopisthus lanceolatus METOPIINAE 1966 bicolor Walley, Bothromus & Townes, gibbus Townes Bothromus 1959 Chorinaeus excessorius Davis, 1897 1886) Mesochorus maleficus Dasch, 1971 Dasch, 1971 Mesochorus prolatus Carria dreisbachi 1959 Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 371

Concealed feeders Concealed feeders Loopers Loopers Concealed feeders Mostly conifer loopers Concealed feeders Host Family (No. spp.) Guild/Notes Diprionidae, Elachistidae, Erebidae, Geometridae, (21) Mostly Tortricidae Noctuidae, Sphingidae, conifer feeders Hymenoptera: Braconidae, Ichneumonidae; Lepidoptera: Bucculatricidae, Choreutidae, Erebidae, Gelechiidae, Geometridae, Noctuidae, Nolidae, Pieridae, Plutellidae (78) (3) Tortricidae Lepidoptera: (8) Tortricidae Lepidoptera: Geometridae, Mostly concealed feeders Coleoptera: Chrysomelidae; Diptera: Tachinidae; Tachinidae; Coleoptera: Chrysomelidae; Diptera: Hymenoptera: Braconidae, Ichneumonidae, Lepidoptera: Gelechiidae, Noctuidae, Tenthredinidae; (43) Tortricidae Notodontidae, Psychidae, Saturniidae, None None Lepidoptera: Erebidae, Geometridae, Noctuidae (12) Mostly conifer loopers (1) Ypsolophidae Lepidoptera: None Hymenoptera: Diprionidae, Ichneumonidae, (7) Tenthredinidae Mostly conifer sawflies None None Hymenoptera, Lepidoptera: Braconidae, Crambidae, (5) Yponomeutidae Geometridae, Lepidoptera: Geometridae (1) None Erebidae, Tachinidae, Diptera, Lepidoptera: Geometridae (15) Mostly conifer loopers and Tachinidae Geometridae (2) Tachinidae, Diptera, Lepidoptera: Hymenoptera, Lepidoptera: Braconidae, None Hymenoptera: Ichneumonidae (1) Yponomeutidae Lepidoptera: Crambidae, Tortricidae, (4) Concealed feeders Diptera: Muscidae; Hymenoptera: Braconidae; Lepidoptera: Crambidae, Gracillariidae, Nolidae, Plutellidae (9) Lepidoptera: Gracillariidae (1) Lepidoptera: Tortricidae, Gracillariidae, (5) Yponomeutidae Hymenoptera: Braconidae; Lepidoptera: Gelechiidae, (4) Tortricidae Mostly concealed feeders None Lepidoptera: Geometridae (1) (3) Tortricidae Lepidoptera: Elachistidae, Concealed feeders

m m Swan River b Localities Wagner NA, Duffield, Edmonton, Nampa, Ponoka, Cr, AB: Buffalo FR, Woods Shilo, Spruce Lakes NP; MB: Ninette, 5 mi SW Waterton Mt Turtle AB: Elkwater L AB: Edmonton SK: Kindersley NA Wagner Cochrane, Edmonton, Elkwater, AB: Bilby, AB: Edmonton AB: Beaver Mines, Ferry Saskatchewan S of Stony Plain, Hayter, Provost, Red Deer Ranger Stn, Rivercours; SK: Canora, Cloan, Humboldt, Imperial; MB: Crandall, Hamiota, Mulvihill, Shoal L Transcona NA; MB: Wagner L, Lethbridge, George AB: Bilby, AB: Edmonton, Elkwater P Edmonton, Coaldale, Cypress Hills, Drumheller, AB: Calgary, Newell, Lethbridge, Manyberries, L Glenwood, Jumping Pound Cr, Oyen, Picture Butte, Rosebud, Scandia, Medicine Hat, Milk River, Horse; SK: Wild Lakes NP, Waterton NA, Wagner Vauxhall, Tilley, Cadillac, Consul, at Cut Knife, Battle River, Attons L Antelope L, Great Sand Hills, Indian Head, Minton, Dunblane, Estevan, Great Deer, Aweme, MB: Tompkins; Parkbeg, Regina, Saskatoon, Swift Current, Brandon, Morris AB: Edmonton AB: Edmonton AB: Edmonton, Elk Island P Mt FR Turtle in MB: Max L Winfield; AB: Belly River Ranger Stn, Edmonton, Lethbridge, Indian Head, Saskatoon SK: Barrier River, AB: Orion L, Lethbridge AB: George Edmonton, NP, L Waterton at Cameron L Burdett, Calgary, AB: Bilby, Rosemary, Onefour, L, Lethbridge, Lost River, George Elkwater, Moose SK: Dunblane, Great Deer, Winfield; Walsh, 21 mi S Stavely, Aweme, MB: Watrous; Albert, Saskatoon, Swift Current, Prince Mt P, Shilo, Starbuck , Brandon, 5 mi SW SK: Saskatoon Saskatoon AB: Lethbridge, Manyberries, Onefour; SK: Great Deer, AB: Edmonton, Elkwater L AB: Edmonton AB: Edmonton, Rockyford; SK: Cypress Hills Calgary W AB: Jumping Pd Cr 20 mi View) Ho (Mountain Westward AB: Gull L, nr Brooks, SK: Saskatoon; MB: 1 locality in southwest AB: Edmonton MB: 2 localities (southwest and southeast) 100,N

9,100,H,T 2,100,N 100,N

100,N 100,N

100,N 100,N,T,O 100,N 105,N 100,N 100,N 100,N 100,N 100,N

100,N 100,N,T 100,N 100,N 100,N 103,N Dasch, 1971 (Thomson, (Provancher, 1879) (Provancher, Ashmead, 1898 (Zetterstedt, 1838) Townes & Townes, & Townes, Townes a 2,100,N 100,N,T,A 103,104,105,N 104,105,N 104,N 2,14,53,100,N (Thunberg, 1822) Mesochorus giberius (Thunberg, Mesochorus validus Dasch, 1971 Dasch, 1971 Stictopisthus argaleus & Townes, Chorinaeus aequalis Townes 1959 SUBFAMILY Species 2,14,100,H,T,O Mesochorus uniformis Cresson, 1872 Mesochorus hesperus Dasch, 1971 Mesochorus inaequidens Dasch, 1971 Mesochorus incultus Dasch, 1971 Mesochorus longiscutatus Dasch, 1971 Mesochorus masoni Dasch, 1971 Mesochorus ottawaensis (Harrington, 1892) Mesochorus peltatus Dasch, 1971 *Mesochorus perniciosus Viereck, 1911 Mesochorus personatus Dasch, 1971 Mesochorus tachinae Mesochorus unicarinatus Dasch, 1971 Mesochorus varianus Dasch, 1971 Mesochorus vittator Stictopisthus bilineatus Dasch, 1971 Stictopisthus crenatus Stictopisthus flaviceps Stictopisthus lanceolatus METOPIINAE 1966 bicolor Walley, Bothromus & Townes, gibbus Townes Bothromus 1959 Chorinaeus excessorius Davis, 1897 1886) Mesochorus maleficus Dasch, 1971 Dasch, 1971 Mesochorus prolatus Carria dreisbachi 1959 372 M. D. Schwarzfeld

Loopers Loopers Concealed feeders Concealed feeders Concealed feeders Concealed feeders Leafrollers Leafrollers Leafrollers Concealed feeders Concealed feeders Concealed feeders Concealed feeders Lepidoptera: Pyralidae, Tortricidae (10) Tortricidae Lepidoptera: Pyralidae, Concealed conifer feeders Lepidoptera: Elachistidae, Geometridae, Tortricidae, Tortricidae, Lepidoptera: Elachistidae, Geometridae, (14) Yponomeutidae Mostly conifer loopers Host Family (No. spp.) Guild/Notes (15) Tortricidae Lepidoptera: Gelechiidae, Erebidae, Mostly concealed feeders (2) Tortricidae Lepidoptera: (1) Tortricidae Lepidoptera: (11) Saturniidae Notodontidae, Noctuidae, Lepidoptera: Lepidoptera: Noctuidae (2) Lepidoptera: Geometridae (2) Lepidoptera: Pyralidae (2) (2) Tortricidae Lepidoptera: None None None (1) Tortricidae Lepidoptera: (1) Tortricidae Lepidoptera: None Hymenoptera: Cynipidae; Lepidoptera: Noctuidae, (6) Yponomeutidae Pyralidae, Tortricidae, Mostly concealed feeders None (6) Tortricidae Lepidoptera: Lepidoptera: Crambidae Tenthredinidae; Hymenoptera: (2) Lepidoptera: Gelichiidae (2) Lepidoptera: Pyralidae (2) ( Lepidoptera: Gelechiidae (1) (1) Tortricidae Lepidoptera: None (2) Tortricidae Anobiidae; Lepidoptera: Coleoptera: None Lepidoptera: Geometridae (1) None Hymenoptera: Ichneumonidae; Lepidoptera: Noctuidae (3) None Lepidoptera: Elachistidae, Gelechiidae, Geometridae, Heterobathmiidae, Tortricidae, Pyralidae, (25) Ypsolophidae Yponomeutidae, Concealed feeders (17) Tortricidae Lepidoptera: Hesperiidae, Concealed feeders None Lepidoptera: Bucculatricidae, Gelechiidae, Tortricidae Tortricidae Lepidoptera: Bucculatricidae, Gelechiidae, (9) Concealed feeders , 2

m

, Waterton; , Waterton; ; MB: Aweme, ; MB: Aweme,

m m m m ; MB: Agassiz, m ; SK: nr Prince Albert ; SK: nr Prince m , southeast m Spruce Woods FR Woods , Spruce m , Wagner NA , Wagner m m m , Sutherland m m ; MB: Deepdale , Lethbridge NA , Wagner m m m m b m across southern MB locations in south southwest Localities AB: Edmonton Albert AB: 1 locality in southeast; SK: nr Prince Wainwright NA, nr Wagner AB: Blackfoot Hills, AB: Edmonton AB: Edmonton MB: Aweme AB: Chin; SK: Mossbank NA AB: Wagner NA Wagner AB: Edmonton, NA, Waterton AB: Wagner NA AB: Wagner AB: Edmonton NA AB: Wagner NA AB: Wagner Aweme AB: Lethbridge; MB: AB: Edmonton NA Wagner AB: Edmonton, NA AB: Wagner AB: Edmonton AB: Edmonton; MB: nr Brandon NA AB: Wagner NA AB: Wagner SK: Saskatoon SK: nr Saskatoon NA Wagner AB: Edmonton, NA AB: Wagner AB: Edmonton AB: Edmonton MB: Pine Ridge SK: Southeast Wainwright; AB: Lloydminster, Tofield AB: southwest Wainwright, AB: Fawcett, Waterton AB: Cypress Hills, Cochrane, Lethbridge AB: nr Calgary, SK: Roche Percee; MB: Transcona SK: Roche Percee; MB: SK: Kinistino, several locations in parkland 2,N 2,N 2,N 2,N 2,N 104,N

2,104,N

14,W

5,104,H

5,104,H 2,104,105,H 2,104,105,N 5,104,105,N 104,N 104,105,H 104,N 2,5,104,105,N 9,14,105,N 104,105,N 5,H 5,104,105,N 2,104,N 105,N 104,N (Davis, 1897) (Say, 1835) (Say, Townes & Townes, & Townes, Townes Thomson, 1887 Davis, 1897 Walsh, 1873 Walsh, Townes & Townes, & Townes, Townes Townes & Townes, & Townes, Townes Davis, 1897 Cresson, 1864 Townes & Townes, & Townes, Townes Townes & Townes, 1959 Townes, & Townes Townes & Townes, 1959 Townes, & Townes Walsh, 1873 Walsh, (Cresson, 1868) Townes & Townes, 1959 Townes, & Townes Holmgren, 1858 a 104,105,H 105,H 104,105,N 2,104,N 2,104,105,N 2,N 2,N 104,105,N 2,N 45,104,105,H Exochus washingtonensis Cresson, 1868 Exochus albifrons Chorinaeus funebris (Gravenhorst, 1829) Exochus annulicrus & Townes, Exochus armillosus Townes 1959 1959 & Townes, Exochus brutus Townes Exochus denotatus Davis, 1897 *Exochus flavifrontalis 1959 Townes, & Exochus mesodon Townes Exochus pictus Hypsicera cuneata Periope aethiops Spudaeus scaber (Gravenhorst, 1829) SUBFAMILY Species Exochus nigripalpis 1959 Exochus atriceps Exochus bryanti Exochus canidens Exochus cnemidotus Exochus mitratus Gravenhorst, 1829 1959 Townes, & Townes Exochus peroniae Exochus pleuralis Exochus spilotus & Townes, Exochus transversus Townes 1959 Holmgren, 1858 Exochus turgidus 1785) Hypsicera femoralis (Geoffroy, Cresson, 1879 Metopius edwardsii Metopius errantia Spudaeus indigus (Davis, 1897) brunnipes (Cresson, 1879) 104,N Triclistus & Townes, crassus Townes Triclistus 1959 1959 Metopius pollinctorius Chorinaeus subcarinatus Holmgren, 1858 Exochus ostentatus Townes & Townes, & Townes, Exochus megadon Townes 1959 Exochus signifer 1959 1959 Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 373

Loopers Loopers Concealed feeders Concealed feeders Concealed feeders Concealed feeders Leafrollers Leafrollers Leafrollers Concealed feeders Concealed feeders Concealed feeders Concealed feeders Lepidoptera: Pyralidae, Tortricidae (10) Tortricidae Lepidoptera: Pyralidae, Concealed conifer feeders Lepidoptera: Elachistidae, Geometridae, Tortricidae, Tortricidae, Lepidoptera: Elachistidae, Geometridae, (14) Yponomeutidae Mostly conifer loopers Host Family (No. spp.) Guild/Notes (15) Tortricidae Lepidoptera: Gelechiidae, Erebidae, Mostly concealed feeders (2) Tortricidae Lepidoptera: (1) Tortricidae Lepidoptera: (11) Saturniidae Notodontidae, Noctuidae, Lepidoptera: Lepidoptera: Noctuidae (2) Lepidoptera: Geometridae (2) Lepidoptera: Pyralidae (2) (2) Tortricidae Lepidoptera: None None None (1) Tortricidae Lepidoptera: (1) Tortricidae Lepidoptera: None Hymenoptera: Cynipidae; Lepidoptera: Noctuidae, (6) Yponomeutidae Pyralidae, Tortricidae, Mostly concealed feeders None (6) Tortricidae Lepidoptera: Lepidoptera: Crambidae Tenthredinidae; Hymenoptera: (2) Lepidoptera: Gelichiidae (2) Lepidoptera: Pyralidae (2) ( Lepidoptera: Gelechiidae (1) (1) Tortricidae Lepidoptera: None (2) Tortricidae Anobiidae; Lepidoptera: Coleoptera: None Lepidoptera: Geometridae (1) None Hymenoptera: Ichneumonidae; Lepidoptera: Noctuidae (3) None Lepidoptera: Elachistidae, Gelechiidae, Geometridae, Heterobathmiidae, Tortricidae, Pyralidae, (25) Ypsolophidae Yponomeutidae, Concealed feeders (17) Tortricidae Lepidoptera: Hesperiidae, Concealed feeders None Lepidoptera: Bucculatricidae, Gelechiidae, Tortricidae Tortricidae Lepidoptera: Bucculatricidae, Gelechiidae, (9) Concealed feeders , 2

m

, Waterton; , Waterton; ; MB: Aweme, ; MB: Aweme, m m m m ; MB: Agassiz, m ; SK: nr Prince Albert ; SK: nr Prince m , southeast m Spruce Woods FR Woods , Spruce m , Wagner NA , Wagner m m m , Sutherland m m ; MB: Deepdale , Lethbridge NA , Wagner m m m m b m across southern MB locations in south southwest Localities AB: Edmonton Albert AB: 1 locality in southeast; SK: nr Prince Wainwright NA, nr Wagner AB: Blackfoot Hills, AB: Edmonton AB: Edmonton MB: Aweme AB: Chin; SK: Mossbank NA AB: Wagner NA Wagner AB: Edmonton, NA, Waterton AB: Wagner NA AB: Wagner AB: Edmonton NA AB: Wagner NA AB: Wagner Aweme AB: Lethbridge; MB: AB: Edmonton NA Wagner AB: Edmonton, NA AB: Wagner AB: Edmonton AB: Edmonton; MB: nr Brandon NA AB: Wagner NA AB: Wagner SK: Saskatoon SK: nr Saskatoon NA Wagner AB: Edmonton, NA AB: Wagner AB: Edmonton AB: Edmonton MB: Pine Ridge SK: Southeast Wainwright; AB: Lloydminster, Tofield AB: southwest Wainwright, AB: Fawcett, Waterton AB: Cypress Hills, Cochrane, Lethbridge AB: nr Calgary, SK: Roche Percee; MB: Transcona SK: Roche Percee; MB: SK: Kinistino, several locations in parkland 2,N 2,N 2,N 2,N 2,N 104,N

2,104,N

14,W

5,104,H

5,104,H 2,104,105,H 2,104,105,N 5,104,105,N 104,N 104,105,H 104,N 2,5,104,105,N 9,14,105,N 104,105,N 5,H 5,104,105,N 2,104,N 105,N 104,N (Davis, 1897) (Say, 1835) (Say, Townes & Townes, & Townes, Townes Thomson, 1887 Davis, 1897 Walsh, 1873 Walsh, Townes & Townes, & Townes, Townes Townes & Townes, & Townes, Townes Davis, 1897 Cresson, 1864 Townes & Townes, & Townes, Townes Townes & Townes, 1959 Townes, & Townes Townes & Townes, 1959 Townes, & Townes Walsh, 1873 Walsh, (Cresson, 1868) Townes & Townes, 1959 Townes, & Townes Holmgren, 1858 a 104,105,H 105,H 104,105,N 2,104,N 2,104,105,N 2,N 2,N 104,105,N 2,N 45,104,105,H Exochus washingtonensis Cresson, 1868 Exochus albifrons Chorinaeus funebris (Gravenhorst, 1829) Exochus annulicrus & Townes, Exochus armillosus Townes 1959 1959 & Townes, Exochus brutus Townes Exochus denotatus Davis, 1897 *Exochus flavifrontalis 1959 Townes, & Exochus mesodon Townes Exochus pictus Hypsicera cuneata Periope aethiops Spudaeus scaber (Gravenhorst, 1829) SUBFAMILY Species Exochus nigripalpis 1959 Exochus atriceps Exochus bryanti Exochus canidens Exochus cnemidotus Exochus mitratus Gravenhorst, 1829 1959 Townes, & Townes Exochus peroniae Exochus pleuralis Exochus spilotus & Townes, Exochus transversus Townes 1959 Holmgren, 1858 Exochus turgidus 1785) Hypsicera femoralis (Geoffroy, Cresson, 1879 Metopius edwardsii Metopius errantia Spudaeus indigus (Davis, 1897) brunnipes (Cresson, 1879) 104,N Triclistus & Townes, crassus Townes Triclistus 1959 1959 Metopius pollinctorius Chorinaeus subcarinatus Holmgren, 1858 Exochus ostentatus Townes & Townes, & Townes, Exochus megadon Townes 1959 Exochus signifer 1959 1959 374 M. D. Schwarzfeld

Concealed feeders Concealed feeder Concealed feeders Fungus gnats Fungus gnats Fungus gnats Concealed feeders Loopers Bioluminescent fungus gnat Mostly concealed feeders Host Family (No. spp.) Guild/Notes Lepidoptera: Bucculatricidae, Choreutidae, Crambidae, Geometridae, Gracillariidae, Nolidae, Pyralidae, (27) Tortricidae Tortricidae, Lepidoptera: Crambidae, Geometridae, (16) Yponomeutidae Concealed feeders Lepidoptera: Choreutidae, Elachistidae, Gelechiidae, (22) Tortricidae Gracillariidae, Pyralidae, Concealed feeders Lepidoptera: Elachistidae (1) (3) Tortricidae Lepidoptera: Lepidoptera: Pyralidae, Tortricidae (2) Tortricidae Lepidoptera: Pyralidae, Concealed feeders Yponomeutidae Lepidoptera: Crambidae, Tortricidae, (7) Concealed feeders Lepidoptera: Erebidae (1) None (7) Tortricidae Lepidoptera: Elachistidae, Pyralidae, Concealed feeders (1) Yponomeutidae Lepidoptera: None Lepidoptera: Crambidae, Tephritidae; Diptera: Drepanidae, Erebidae, Noctuidae, Notodontidae, Saturniidae (19) Apatelodidae, Saturniidae (12) Lepidoptera: Setose or spiny Lepidoptera: Erebidae, Noctuidae, Notodontidae, Saturniidae (14) Mostly cutworms Lepidoptera: Noctuidae (1) Lepidoptera: Sphingidae (2) None Diptera: Mycetophilidae (3) Diptera: Mycetophilidae (5) None None Diptera: Mycetophilidae (3) Diptera: Keroplatidae (1) None None None None None None None None (1) Yponomeutidae Lepidoptera: Diptera: Keroplatidae (1) Fungus gnats Diptera: Keroplatidae; Hymenoptera: (3) Tenthredinidae None m m m m m , southwest m , Wagner NA; MB: south of Winnipeg NA; MB: south of Wagner , m b Localities AB: Cochrane, Edmonton AB: Manyberries, Brooks; SK: Indian Head AB: Edmonton; SK: nr Lloydminster NA; SK: Saskatoon Wagner AB: Edmonton, NA AB: Wagner MB: Pine Ridge Shilo, southwest MB: Ninette, Pierson, 5 mi SW southwest Treesbank, Assiniboine River at MB: SK: Saskatoon southwest MB: Baldur, NA Wagner Aspen Beach, AB: MB: Aweme Winnipeg Lethbridge; MB: AB: Calgary, MB: Aweme MB: Winnipeg L, Lacombe, Pigeon Edmonton, Erskine, George AB: Calgary, Wyndham- Winfield, Sherwood Park, Spruce Grove, Rochon Sands PP, south Winnipeg, Carseland PP; SK: Saskatoon; MB: Brandon, MB: South AB: Edmonton AB: Edmonton, Jumping Pound Cr Elbow, Milk River; SK: Bredenbury, AB: Edmonton, Jumping Pound Cr, Snowden, White Fox at 1,300m Lakes NP Waterton AB: NP Waterton Rd in AB: Edmonton, Cameron L AB: Aspen Beach AB: Edmonton, Jumping Pound Cr SK: Rockglen, Scout L White Fox Alton L, Saskatoon, Aspen Beach, Battle River; SK: AB: AB: Elkwater L L George AB: Elkwater P, AB: Jumping Pound Cr AB: Edmonton AB: Jumping Pound Cr (15 mi E Calgary); MB: Ninette 1300m at NP L Waterton Waterton, Pk, Jumping Pound Cr, AB: Elkwater AB: Jumping Pound Cr AB: Edmonton, Jumping Pound Cr SK: Tofield; Lundbreck, Onefour, AB: Edmonton, Jumping Pound Cr, Saskatoon Elbow, Bredenbury, MB: Brandon 104,N 79,H 104,N 91,N,T 79,H 79,H 79,H 2,H,O

79,H 79,H 29,N 79,H 104,105,H,O 2,5,104,105,N 79,H 79,H 79,H 5,107,N,T,Oc 79,H 79,H 79,H 79,H 105,106,N 29,109,N 105,106,N 79,N 108,N,T 105,106,N 79,H (Gravenhorst, (Forster, 1871) (Forster, Schiødte, 1838 Rossem, 1981 (Ashmead, 1899) Forster, 1871 Forster, (Ashmead, 1890) (Strobl, 1901) Forster, 1871 Forster, Say, 1829 Say, Dasch, 1992 Holmgren, 1873 Forster, 1871 Forster, Walley, 1969 Walley, Townes & Townes, & Townes, Townes Dasch, 1992 Townes & Townes, 1959 & Townes, Townes (Davis, 1897) a 105,106,N 2,104,105,H 79,H 79,H 79,H,O 104,105,N 105,106,N Triclistus evexus Triclistus 1959 (Say, 1829) (Say, emarginalus Triclistus podagricus (Gravenhorst, Triclistus 1829) pygmaeus (Cresson, 1864) Triclistus 1969 confusus Walley, Trieces epinotiae Trieces integer Trieces Aperileptus albipalpus 1871 Catastenus femoralis Forster, Eusterinx bispinosa SUBFAMILY Species Ophion bilineatus 1829) 1959 Triclistus pallipes Triclistus & Townes, Townes arcuatus Trieces onitis Trieces & Townes, sapineus Townes Trieces 1959 2, 1959 & Townes, Townes teres Trieces OPHIONINAE 1835) (Say, Enicospilus purgatus Enicospilus texanus 1905 Ophion idoneus Viereck, 1811) atricolor (Olivier, Thyreodon ORTHOCENTRINAE latus Dasch, 1992 Aniseres pallipes Aniseres Dialipsis dissimilis Dasch, 1992 1871 Forster, Entypoma robustum Eusterinx oligomera Eusterinx subdola Eusterinx tenuicincta Eusterinx trifasciata Gnathochorisis crassulus (Thomson, 1888) (Holmgren, 1857) Helictes borealis Helictes paucus Hyperacmus crassicornis (Gravenhorst, 1829) Schiødte, 1838 Megastylus impressor Megastylus orbitator Megastylus similis Eusterinx inaequalis Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 375

Concealed feeders Concealed feeder Concealed feeders Fungus gnats Fungus gnats Fungus gnats Concealed feeders Loopers Bioluminescent fungus gnat Mostly concealed feeders Host Family (No. spp.) Guild/Notes Lepidoptera: Bucculatricidae, Choreutidae, Crambidae, Geometridae, Gracillariidae, Nolidae, Pyralidae, (27) Tortricidae Tortricidae, Lepidoptera: Crambidae, Geometridae, (16) Yponomeutidae Concealed feeders Lepidoptera: Choreutidae, Elachistidae, Gelechiidae, (22) Tortricidae Gracillariidae, Pyralidae, Concealed feeders Lepidoptera: Elachistidae (1) (3) Tortricidae Lepidoptera: Lepidoptera: Pyralidae, Tortricidae (2) Tortricidae Lepidoptera: Pyralidae, Concealed feeders Yponomeutidae Lepidoptera: Crambidae, Tortricidae, (7) Concealed feeders Lepidoptera: Erebidae (1) None (7) Tortricidae Lepidoptera: Elachistidae, Pyralidae, Concealed feeders (1) Yponomeutidae Lepidoptera: None Lepidoptera: Crambidae, Tephritidae; Diptera: Drepanidae, Erebidae, Noctuidae, Notodontidae, Saturniidae (19) Apatelodidae, Saturniidae (12) Lepidoptera: Setose or spiny Lepidoptera: Erebidae, Noctuidae, Notodontidae, Saturniidae (14) Mostly cutworms Lepidoptera: Noctuidae (1) Lepidoptera: Sphingidae (2) None Diptera: Mycetophilidae (3) Diptera: Mycetophilidae (5) None None Diptera: Mycetophilidae (3) Diptera: Keroplatidae (1) None None None None None None None None (1) Yponomeutidae Lepidoptera: Diptera: Keroplatidae (1) Fungus gnats Diptera: Keroplatidae; Hymenoptera: (3) Tenthredinidae None m m m m m , southwest m , Wagner NA; MB: south of Winnipeg NA; MB: south of Wagner , m b Localities AB: Cochrane, Edmonton AB: Manyberries, Brooks; SK: Indian Head AB: Edmonton; SK: nr Lloydminster NA; SK: Saskatoon Wagner AB: Edmonton, NA AB: Wagner MB: Pine Ridge Shilo, southwest MB: Ninette, Pierson, 5 mi SW southwest Treesbank, Assiniboine River at MB: SK: Saskatoon southwest MB: Baldur, NA Wagner Aspen Beach, AB: MB: Aweme Winnipeg Lethbridge; MB: AB: Calgary, MB: Aweme MB: Winnipeg L, Lacombe, Pigeon Edmonton, Erskine, George AB: Calgary, Wyndham- Winfield, Sherwood Park, Spruce Grove, Rochon Sands PP, south Winnipeg, Carseland PP; SK: Saskatoon; MB: Brandon, MB: South AB: Edmonton AB: Edmonton, Jumping Pound Cr Elbow, Milk River; SK: Bredenbury, AB: Edmonton, Jumping Pound Cr, Snowden, White Fox at 1,300m Lakes NP Waterton AB: NP Waterton Rd in AB: Edmonton, Cameron L AB: Aspen Beach AB: Edmonton, Jumping Pound Cr SK: Rockglen, Scout L White Fox Alton L, Saskatoon, Aspen Beach, Battle River; SK: AB: AB: Elkwater L L George AB: Elkwater P, AB: Jumping Pound Cr AB: Edmonton AB: Jumping Pound Cr (15 mi E Calgary); MB: Ninette 1300m at NP L Waterton Waterton, Pk, Jumping Pound Cr, AB: Elkwater AB: Jumping Pound Cr AB: Edmonton, Jumping Pound Cr SK: Tofield; Lundbreck, Onefour, AB: Edmonton, Jumping Pound Cr, Saskatoon Elbow, Bredenbury, MB: Brandon 104,N 79,H 104,N 91,N,T 79,H 79,H 79,H 2,H,O

79,H 79,H 29,N 79,H 104,105,H,O 2,5,104,105,N 79,H 79,H 79,H 5,107,N,T,Oc 79,H 79,H 79,H 79,H 105,106,N 29,109,N 105,106,N 79,N 108,N,T 105,106,N 79,H (Gravenhorst, (Forster, 1871) (Forster, Schiødte, 1838 Rossem, 1981 (Ashmead, 1899) Forster, 1871 Forster, (Ashmead, 1890) (Strobl, 1901) Forster, 1871 Forster, Say, 1829 Say, Dasch, 1992 Holmgren, 1873 Forster, 1871 Forster, Walley, 1969 Walley, Townes & Townes, & Townes, Townes Dasch, 1992 Townes & Townes, 1959 & Townes, Townes (Davis, 1897) a 105,106,N 2,104,105,H 79,H 79,H 79,H,O 104,105,N 105,106,N Triclistus evexus Triclistus 1959 (Say, 1829) (Say, emarginalus Triclistus podagricus (Gravenhorst, Triclistus 1829) pygmaeus (Cresson, 1864) Triclistus 1969 confusus Walley, Trieces epinotiae Trieces integer Trieces Aperileptus albipalpus 1871 Catastenus femoralis Forster, Eusterinx bispinosa SUBFAMILY Species Ophion bilineatus 1829) 1959 Triclistus pallipes Triclistus & Townes, Townes arcuatus Trieces onitis Trieces & Townes, sapineus Townes Trieces 1959 2, 1959 & Townes, Townes teres Trieces OPHIONINAE 1835) (Say, Enicospilus purgatus Enicospilus texanus 1905 Ophion idoneus Viereck, 1811) atricolor (Olivier, Thyreodon ORTHOCENTRINAE latus Dasch, 1992 Aniseres pallipes Aniseres Dialipsis dissimilis Dasch, 1992 1871 Forster, Entypoma robustum Eusterinx oligomera Eusterinx subdola Eusterinx tenuicincta Eusterinx trifasciata Gnathochorisis crassulus (Thomson, 1888) (Holmgren, 1857) Helictes borealis Helictes paucus Hyperacmus crassicornis (Gravenhorst, 1829) Schiødte, 1838 Megastylus impressor Megastylus orbitator Megastylus similis Eusterinx inaequalis 376 M. D. Schwarzfeld

Setose Conifer sawflies Conifer sawflies Fungus gnats Fungus gnats Fungus gnats Fungus gnats Hymenoptera: Braconidae; Lepidoptera: Crambidae, Gracillariidae, Lasiocampidae, Noctuidae, Psychidae, (19) Tortricidae Pyralidae, Mostly concealed feeders Host Family (No. spp.) Guild/Notes Hymenoptera: Cynipidae (13) Gall makers Coleoptera: Chrysomelidae; Hymenoptera: Lepidoptera: Erebidae, Tenthredinidae; concealed feeders None Hymenoptera: Diprionidae (1) Diptera: Mycetophilidae (1) Diptera: Mycetophilidae (3) None None None None None None None Diptera: Bolitophilidae (1) Diptera: Mycetophilidae (5) Diptera: Diadocidiidae (1) Fungus gnats None None Hymenoptera: Cynipidae (3) Gall makers Diptera: Cecidomyiidae; Hymenoptera: Cynipidae, Lepidoptera: Noctuidae, Tenthredinidae; Saturniidae (24) Gall makers Araneae: Theridiidae Linyphiidae, Araneae, Araneidae, Lepidoptera: Tetragnathidae, Elachistidae, Gracillariidae (8) Araneidae Araneae: (2) Mostly Gracillariidae, Incurvariidae (11) Hymenoptera: Ichneumonidae; Lepidoptera: Erebidae, (21) Tortricidae Geometridae, Noctuidae, Pyralidae, Mostly conifer feeders Lepidoptera: Lasiocampidae (1) None Hymenoptera: Diprionidae (10) None Lepidoptera: Tenthredinidae; Hymenoptera: Lasiocampidae (2) Lepidoptera: Erebidae, Geometridae, Pyralidae, (9) Tortricidae Mostly concealed feeders Lepidoptera: Erebidae, Geometridae, Papilionidae, (8) Tortricidae Pieridae, , Pigeon , , Pigeon L Pembina PP

, Pembina PP ; MB: Aweme Pembina PP m , Lethbridge George L m , Pigeon L ; SK: Conquest , Lethbridge, , Pembina PP Pigeon L George L m , Wagner NA , Wagner ; MB: Glenlea b m , Wagner NA, Waterton; SK: nr Regina Waterton; NA, Wagner L , Localities AB: Red Deer Waterton NP, Waterton Rd in Aspen Beach, Edmonton, Cameron L AB: Edmonton, NP, Waterton Rd in Cameron L Aspen Beach, Bilby, AB: Butte, Twin View, Mountain L, Jumping Pound Cr, Elkwater L, George Attons L, Cypress Hills, Moosomin, Nipawin, Saskatoon; SK: Waterton; Stockton W Shilo, 2 mi MB: Horton, Ninette, Ninga, 5 mi SW AB: Cowley AB: Jumping Pound Cr AB: Jumping Pound Cr Edmonton NP, Waterton Rd in AB: Cameron L AB: Elkwater L Shilo AB: Edmonton; MB: 5 mi SW AB: Edmonton, Elkwater L, Jumping Pound Cr; SK: Elbow, White Fox; MB: Ninette Lakes NP Waterton AB: AB: Edmonton AB: Jumping Pound Cr MB: Ninette at 4,300' Lakes NP Waterton AB: NA AB: Wagner America" AB: Medicine Hat; "Throughout N. AB, MB, SK: Numerous locations across prairie and AB: Edmonton; parkland region NA Wagner AB: Edmonton, Lethbridge, AB: Lethbridge, Cooking L , Edmonton, AB: Calgary, MB: Aweme Winnipeg nr MB: Birds Hill PP, AB: Edmonton, AB: Edmonton , George L George L AB: Edmonton, Elkwater P, NA; SK: Cut Knife, Swift Current Wagner AB: Edmonton , AB: George L NA AB: Wagner Sand Hills Aweme, AB: Edmonton; MB: AB: George L AB: Edmonton , George L southeast 5,H 2,N 109,H 5,110,N 5,110,H 5,109,N 2,109,N 5,112,N 1,2,H 79,H 79,H 79,H 2,H 79,H 79,H 5,109,N 14,79,H 1,N 79,H 79,N 79,N 79,N 79,N 79,H 79,N (Gravenhorst, Ashmead, 1890 Ashmead, 1890 Kasparyan 1973 Cushman, (Cresson, 1870) Forster, 1871 Forster, Provancher, Provancher, Roman, 1923 (Cresson, (Cresson, (Haliday, 1838) (Haliday, (Gravenhorst, Forster, 1871 Forster, Rossem, 1981 Dasch, 1992 Dasch, 1992 Dasch, 1992 (Cresson, 1870) Dasch, 1992 (Cresson, 1870) (Haliday, 1838) (Haliday, Dasch, 1992 a 14,79,H 5,109,H,O,Au 1,5,9,105,109,N 1,2,5,105,109,N 109,112,H 105,111,N 1,2,109,H Acrotaphus wiltii Acrotaphus (Zetterstedt, 1838) Orthocentrus frontator Plectiscidea collaris 1829) Plectiscidea communis Symplecis breviuscula PIMPLINAE degener Acrodactyla Apechthis annulicornis Cushman 1921 Clistopyga maculifrons 1960 Walkley, Delomerista borealis SUBFAMILY Species Orthopelma occidentale 1933 Forster, 1871 Pantisarthrus lubricus Forster, Plectiscidea inflata Dasch, 1992 Plectiscidea prolata bicolor Proclitus extensor Proclitus foveatus Proclitus fulvicornis Proclitus 1838) paganus (Haliday, Proclitus praetor Proclitus Symplecis glabra Symplecis invisitata ORTHOPELMATINAE Orthopelma californicum 1824) Orthopelma mediator (Thunberg, quadrisculpta Acrodactyla alboricta Acropimpla 1870) Alophosternum foliicola Apechthis picticornis 1870) Delomerista diprionis Cushman, 1939 Delomerista longicauda Delomerista mandibularis (Gravenhorst, 1829) 1820) 1880 Clistopyga canadensis Apechthis ontario Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 377

Setose Conifer sawflies Conifer sawflies Fungus gnats Fungus gnats Fungus gnats Fungus gnats Hymenoptera: Braconidae; Lepidoptera: Crambidae, Gracillariidae, Lasiocampidae, Noctuidae, Psychidae, (19) Tortricidae Pyralidae, Mostly concealed feeders Host Family (No. spp.) Guild/Notes Hymenoptera: Cynipidae (13) Gall makers Coleoptera: Chrysomelidae; Hymenoptera: Lepidoptera: Erebidae, Tenthredinidae; concealed feeders None Hymenoptera: Diprionidae (1) Diptera: Mycetophilidae (1) Diptera: Mycetophilidae (3) None None None None None None None Diptera: Bolitophilidae (1) Diptera: Mycetophilidae (5) Diptera: Diadocidiidae (1) Fungus gnats None None Hymenoptera: Cynipidae (3) Gall makers Diptera: Cecidomyiidae; Hymenoptera: Cynipidae, Lepidoptera: Noctuidae, Tenthredinidae; Saturniidae (24) Gall makers Araneae: Theridiidae Linyphiidae, Araneae, Araneidae, Lepidoptera: Tetragnathidae, Elachistidae, Gracillariidae (8) Araneidae Araneae: (2) Mostly Gracillariidae, Incurvariidae (11) Hymenoptera: Ichneumonidae; Lepidoptera: Erebidae, (21) Tortricidae Geometridae, Noctuidae, Pyralidae, Mostly conifer feeders Lepidoptera: Lasiocampidae (1) None Hymenoptera: Diprionidae (10) None Lepidoptera: Tenthredinidae; Hymenoptera: Lasiocampidae (2) Lepidoptera: Erebidae, Geometridae, Pyralidae, (9) Tortricidae Mostly concealed feeders Lepidoptera: Erebidae, Geometridae, Papilionidae, (8) Tortricidae Pieridae, , Pigeon , , Pigeon L Pembina PP

, Pembina PP ; MB: Aweme Pembina PP m , Lethbridge George L m , Pigeon L ; SK: Conquest , Lethbridge, , Pembina PP Pigeon L George L m , Wagner NA , Wagner ; MB: Glenlea b m , Wagner NA, Waterton; SK: nr Regina Waterton; NA, Wagner L , Localities AB: Red Deer Waterton NP, Waterton Rd in Aspen Beach, Edmonton, Cameron L AB: Edmonton, NP, Waterton Rd in Cameron L Aspen Beach, Bilby, AB: Butte, Twin View, Mountain L, Jumping Pound Cr, Elkwater L, George Attons L, Cypress Hills, Moosomin, Nipawin, Saskatoon; SK: Waterton; Stockton W Shilo, 2 mi MB: Horton, Ninette, Ninga, 5 mi SW AB: Cowley AB: Jumping Pound Cr AB: Jumping Pound Cr Edmonton NP, Waterton Rd in AB: Cameron L AB: Elkwater L Shilo AB: Edmonton; MB: 5 mi SW AB: Edmonton, Elkwater L, Jumping Pound Cr; SK: Elbow, White Fox; MB: Ninette Lakes NP Waterton AB: AB: Edmonton AB: Jumping Pound Cr MB: Ninette at 4,300' Lakes NP Waterton AB: NA AB: Wagner America" AB: Medicine Hat; "Throughout N. AB, MB, SK: Numerous locations across prairie and AB: Edmonton; parkland region NA Wagner AB: Edmonton, Lethbridge, AB: Lethbridge, Cooking L , Edmonton, AB: Calgary, MB: Aweme Winnipeg nr MB: Birds Hill PP, AB: Edmonton, AB: Edmonton , George L George L AB: Edmonton, Elkwater P, NA; SK: Cut Knife, Swift Current Wagner AB: Edmonton , AB: George L NA AB: Wagner Sand Hills Aweme, AB: Edmonton; MB: AB: George L AB: Edmonton , George L southeast 5,H 2,N 109,H 5,110,N 5,110,H 5,109,N 2,109,N 5,112,N 1,2,H 79,H 79,H 79,H 2,H 79,H 79,H 5,109,N 14,79,H 1,N 79,H 79,N 79,N 79,N 79,N 79,H 79,N (Gravenhorst, Ashmead, 1890 Ashmead, 1890 Kasparyan 1973 Cushman, (Cresson, 1870) Forster, 1871 Forster, Provancher, Provancher, Roman, 1923 (Cresson, (Cresson, (Haliday, 1838) (Haliday, (Gravenhorst, Forster, 1871 Forster, Rossem, 1981 Dasch, 1992 Dasch, 1992 Dasch, 1992 (Cresson, 1870) Dasch, 1992 (Cresson, 1870) (Haliday, 1838) (Haliday, Dasch, 1992 a 14,79,H 5,109,H,O,Au 1,5,9,105,109,N 1,2,5,105,109,N 109,112,H 105,111,N 1,2,109,H Acrotaphus wiltii Acrotaphus (Zetterstedt, 1838) Orthocentrus frontator Plectiscidea collaris 1829) Plectiscidea communis Symplecis breviuscula PIMPLINAE degener Acrodactyla Apechthis annulicornis Cushman 1921 Clistopyga maculifrons 1960 Walkley, Delomerista borealis SUBFAMILY Species Orthopelma occidentale 1933 Forster, 1871 Pantisarthrus lubricus Forster, Plectiscidea inflata Dasch, 1992 Plectiscidea prolata bicolor Proclitus extensor Proclitus foveatus Proclitus fulvicornis Proclitus 1838) paganus (Haliday, Proclitus praetor Proclitus Symplecis glabra Symplecis invisitata ORTHOPELMATINAE Orthopelma californicum 1824) Orthopelma mediator (Thunberg, quadrisculpta Acrodactyla alboricta Acropimpla 1870) Alophosternum foliicola Apechthis picticornis 1870) Delomerista diprionis Cushman, 1939 Delomerista longicauda Delomerista mandibularis (Gravenhorst, 1829) 1820) 1880 Clistopyga canadensis Apechthis ontario 378 M. D. Schwarzfeld Leafcutter bees Wood borers Wood Host Family (No. spp.) Guild/Notes None Coleoptera: Curculionidae; Lepidoptera: (39) Tortricidae Gelechiidae, Pyralidae, Argyresthiidae, Conifer borers None Anthomyiidae, Coleoptera: Curculionidae; Diptera: Chamaemyiidae; Hymenoptera: Cephidae, Lepidoptera: Erebidae, Tenthredinidae; Cimbicidae, Gelechiidae, Geometridae, Lasiocampidae, Noctuidae, Tortricidae, Nolidae, Notodontidae, Pieridae, Pyralidae, (58) Yponomeutidae None Coleoptera: Cerambycidae; Hymenoptera: Xiphydriidae (4) Coleoptera: Buprestidae, Cerambycidae, Curculionidae; Tenthredinidae; Hymenoptera: Tortricidae Lepidoptera: Cossidae, Pyralidae, Sesiidae, (29) Mostly wood borers Coleoptera: Buprestidae, Cerambycidae, Curculionidae; Lepidoptera: Erebidae, Prodoxidae, Pyralidae, Sesiidae (18) Mostly wood borers Coleoptera: Cerambycidae; Lepidoptera: Gelechiidae, (9) Tortricidae Borers Coleoptera: Cerambycidae; Lepidoptera: Gelechiidae (2) Borers Coleoptera: Buprestidae, Cerambycidae (3) Borers Curculionidae, Cerambycidae, Buprestidae, Coleoptera: Monotomidae; Hymenoptera: Cynipidae; Lepidoptera: Erebidae, Pyralidae, Sesiidae (52) Argyresthiidae, Mostly wood borers Coleoptera: Buprestidae (1) borers Wood None Coleoptera: Curculionidae, Rhynchitidae; Diptera: Chamaemyiidae, Chloropidae; Hymenoptera: Tenthredinidae; Cephidae, Cynipidae, Eurytomidae, Argyresthiidae, Lepidoptera: Coleophoridae, Crambidae, Gelechiidae, Lasiocampidae, Noctuidae, Tortricidae, Pterophoridae, Pyralidae, Sesiidae, Uraniidae (43) Mostly concealed feeders None Hymenoptera: Megachilidae (2) Coleoptera: Buprestidae, Cerambycidae; Hymenoptera: (3) Vespidae Mostly wood borers Coleoptera: Buprestidae, Cerambycidae, Cleridae, Curculionidae, Melyridae; Hymenoptera: Cephidae, Crabronidae, Cynipidae, Megachilidae, Siricidae, Xiphydriidae; Lepidoptera: Vespidae, Tenthredinidae, Geometridae, Pyralidae, Sesiidae, Sphingidae, (46) Tortricidae Mostly concealed feeders Coleoptera: Curculionidae; Hymenoptera: Diprionidae; (6) Tortricidae Lepidoptera: Erebidae, Pyralidae, Mostly concealed feeders Coleoptera: Buprestidae, Cerambycidae, Curculionidae; Lepidoptera: Sesiidae (24) borers Wood m Pigeon L Fort Garry , Jenner Lethbridge, , Vermilion, Wabamun, Waterton Lakes NP; Waterton Wabamun, , Vermilion, , Waterton; MB: Aweme, Onah Aweme, MB: Waterton; NA , , Wagner m parkland (no localities), Wagner NA; SK: Cypress Wagner parkland (no localities),

Pigeon L George L , Wagner NA; SK: Indian Head; MB: across south Wagner , , m m , Pigeon L ; SK: Regina, Saskatoon Edmonton , George L b Localities AB: Edmonton AB: Edmonton, AB: George L SK: Vawn Wainwright; AB: Hardisty, AB: Edmonton, Aweme SK: Indian Head; MB: SK: Earl Grey AB: Edmonton, George L NA Wagner AB: George L, NA AB: Wagner George L AB: Calgary, MB: Aweme NA AB: Cooking L, George Wagner AB: Wagner, AB: George L, Pembina PP Aweme White Fox; MB: River, Torch NA; SK: Wagner AB: Aweme SK: Leslie; MB: MB: Aweme AB: Southwest AB: Edmonton , Medicine Hat; MB: AB: Edmonton Hills, nr Saskatoon MB: Onah 1,H 1,H 109,N 114,H,O 109,N 109,112,N 9,14,109,N 2,H 1,2,105,N,T 2,109,N 1,H,O 1,N 1,2,5,46,105,H 5,109,H 109,H 109,N 2,114,H (Walsh, 1873) (Walsh, (Holmgren, (Walsh, 1873) (Walsh, (Say, 1829) (Say, Townes, 1960 Townes, (Hartig, 1847) Heinrich, 1949 Gupta, 1982 (Cresson, 1880) (Cresson 1870) a 1,5,H 1,5,109,H 1,5,109,H 1,2,109,113,H,O Delomerista masoni Dolichomitus cephalotes 1860) Ephialtes duplicauda Flacopimpla nigriceps (Fabricius, 1793) stercorator Iseropus SUBFAMILY Species Dolichomitus vitticrus Delomerista novita Dolichomitus imperator (Kriechbaumer, 1854) 1848) Dolichomitus populneus (Ratzeburg, Dolichomitus pterelas Dolichomitus pygmaeus Dolichomitus sericeus (Ratzeburg, Dolichomitus terebrans 1844) slossonae (Davis, 1898) Dreisbachia detrita (Holmgren, Endromopoda 1860) 1960) Endromopoda producta (Walley, 1914 Morley, Ephialtes brevis Ephialtes manifestato r (Linnaeus, 1758) Exeristes comstockii Delomerista townesorum Gupta, 1982 Dolichomitus messor (Gravenhorst, 1829) Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 379 Leafcutter bees Wood borers Wood Host Family (No. spp.) Guild/Notes None Coleoptera: Curculionidae; Lepidoptera: (39) Tortricidae Gelechiidae, Pyralidae, Argyresthiidae, Conifer borers None Anthomyiidae, Coleoptera: Curculionidae; Diptera: Chamaemyiidae; Hymenoptera: Cephidae, Lepidoptera: Erebidae, Tenthredinidae; Cimbicidae, Gelechiidae, Geometridae, Lasiocampidae, Noctuidae, Tortricidae, Nolidae, Notodontidae, Pieridae, Pyralidae, (58) Yponomeutidae None Coleoptera: Cerambycidae; Hymenoptera: Xiphydriidae (4) Coleoptera: Buprestidae, Cerambycidae, Curculionidae; Tenthredinidae; Hymenoptera: Tortricidae Lepidoptera: Cossidae, Pyralidae, Sesiidae, (29) Mostly wood borers Coleoptera: Buprestidae, Cerambycidae, Curculionidae; Lepidoptera: Erebidae, Prodoxidae, Pyralidae, Sesiidae (18) Mostly wood borers Coleoptera: Cerambycidae; Lepidoptera: Gelechiidae, (9) Tortricidae Borers Coleoptera: Cerambycidae; Lepidoptera: Gelechiidae (2) Borers Coleoptera: Buprestidae, Cerambycidae (3) Borers Curculionidae, Cerambycidae, Buprestidae, Coleoptera: Monotomidae; Hymenoptera: Cynipidae; Lepidoptera: Erebidae, Pyralidae, Sesiidae (52) Argyresthiidae, Mostly wood borers Coleoptera: Buprestidae (1) borers Wood None Coleoptera: Curculionidae, Rhynchitidae; Diptera: Chamaemyiidae, Chloropidae; Hymenoptera: Tenthredinidae; Cephidae, Cynipidae, Eurytomidae, Argyresthiidae, Lepidoptera: Coleophoridae, Crambidae, Gelechiidae, Lasiocampidae, Noctuidae, Tortricidae, Pterophoridae, Pyralidae, Sesiidae, Uraniidae (43) Mostly concealed feeders None Hymenoptera: Megachilidae (2) Coleoptera: Buprestidae, Cerambycidae; Hymenoptera: (3) Vespidae Mostly wood borers Coleoptera: Buprestidae, Cerambycidae, Cleridae, Curculionidae, Melyridae; Hymenoptera: Cephidae, Crabronidae, Cynipidae, Megachilidae, Siricidae, Xiphydriidae; Lepidoptera: Vespidae, Tenthredinidae, Geometridae, Pyralidae, Sesiidae, Sphingidae, (46) Tortricidae Mostly concealed feeders Coleoptera: Curculionidae; Hymenoptera: Diprionidae; (6) Tortricidae Lepidoptera: Erebidae, Pyralidae, Mostly concealed feeders Coleoptera: Buprestidae, Cerambycidae, Curculionidae; Lepidoptera: Sesiidae (24) borers Wood m Pigeon L Fort Garry , Jenner Lethbridge, , Vermilion, Wabamun, Waterton Lakes NP; Waterton Wabamun, , Vermilion, , Waterton; MB: Aweme, Onah Aweme, MB: Waterton; NA , , Wagner m parkland (no localities), Wagner NA; SK: Cypress Wagner parkland (no localities),

Pigeon L George L , Wagner NA; SK: Indian Head; MB: across south Wagner , , m m , Pigeon L ; SK: Regina, Saskatoon Edmonton , George L b Localities AB: Edmonton AB: Edmonton, AB: George L SK: Vawn Wainwright; AB: Hardisty, AB: Edmonton, Aweme SK: Indian Head; MB: SK: Earl Grey AB: Edmonton, George L NA Wagner AB: George L, NA AB: Wagner George L AB: Calgary, MB: Aweme NA AB: Cooking L, George Wagner AB: Wagner, AB: George L, Pembina PP Aweme White Fox; MB: River, Torch NA; SK: Wagner AB: Aweme SK: Leslie; MB: MB: Aweme AB: Southwest AB: Edmonton , Medicine Hat; MB: AB: Edmonton Hills, nr Saskatoon MB: Onah 1,H 1,H 109,N 114,H,O 109,N 109,112,N 9,14,109,N 2,H 1,2,105,N,T 2,109,N 1,H,O 1,N 1,2,5,46,105,H 5,109,H 109,H 109,N 2,114,H (Walsh, 1873) (Walsh, (Holmgren, (Walsh, 1873) (Walsh, (Say, 1829) (Say, Townes, 1960 Townes, (Hartig, 1847) Heinrich, 1949 Gupta, 1982 (Cresson, 1880) (Cresson 1870) a 1,5,H 1,5,109,H 1,5,109,H 1,2,109,113,H,O Delomerista masoni Dolichomitus cephalotes 1860) Ephialtes duplicauda Flacopimpla nigriceps (Fabricius, 1793) stercorator Iseropus SUBFAMILY Species Dolichomitus vitticrus Delomerista novita Dolichomitus imperator (Kriechbaumer, 1854) 1848) Dolichomitus populneus (Ratzeburg, Dolichomitus pterelas Dolichomitus pygmaeus Dolichomitus sericeus (Ratzeburg, Dolichomitus terebrans 1844) slossonae (Davis, 1898) Dreisbachia detrita (Holmgren, Endromopoda 1860) 1960) Endromopoda producta (Walley, 1914 Morley, Ephialtes brevis Ephialtes manifestato r (Linnaeus, 1758) Exeristes comstockii Delomerista townesorum Gupta, 1982 Dolichomitus messor (Gravenhorst, 1829) 380 M. D. Schwarzfeld Mostly concealed Underbark Clerids Mostly spiders Hymenoptera: Braconidae, Diprionidae, Lepidoptera: Tenthredinidae; Ichneumonidae, Choreutidae, Coleophoridae, Erebidae, Gelechiidae, Geometridae, Gracillariidae, Lasiocampidae, Tortricidae, Oecophoridae, Psychidae, Pyralidae, Zygaenidae (63) Yponomeutidae, feeders Host Family (No. spp.) Guild/Notes Coleoptera: Cerambycidae, Chrysomelidae, Hymenoptera: Tachinidae; Curculionidae; Diptera: Braconidae, Diprionidae, Ichneumonidae; Lepidoptera: Choreutidae, Coleophoridae, Cosmopterigidae, Crambidae, Epermeniidae, Erebidae, Gelechiidae, Geometridae, Gracillariidae, Hesperiidae, Lasiocampidae, Noctuidae, Nolidae, Notodontidae, Nymphalidae, Papilionidae, Pieridae, Plutellidae, Psychidae, Pterophoridae, Pyralidae, Saturniidae, (155) Tortricidae Sesiidae, Mostly concealed feeders Araneidae, Araneae: Theridiidae; Coleoptera: Curculionidae; Lepidoptera: Coleophoridae, Elachistidae, Erebidae, Gelechiidae, Geometridae, (19) Tortricidae Lasiocampidae, Coleoptera: Cerambycidae; Hymenoptera: Diprionidae; Lepidoptera: Erebidae, Geometridae, Hesperiidae, Lasiocampidae, Noctuidae, Notodontidae, Pieridae, (42) Tortricidae None Curculionidae; Buprestidae, Cerambycidae, Coleoptera: Hymenoptera: Tephritidae; Diptera: Scathophagidae, Tenthredinidae; Braconidae, Cynipidae, Diprionidae, Lepidoptera: Choreutidae, Coleophoridae, Elachistidae, Erebidae, Gelechiidae, Geometridae, Gracillariidae, Lasiocampidae, Momphidae, Noctuidae, Pieridae, Tischeriidae, Plutellidae, Pterophoridae, Pyralidae, (127) Tortricidae Mostly concealed feeders Hymenoptera: Braconidae; Lepidoptera: Gelechiidae, Zygaenidae (5) None Tortricidae Lepidoptera: Coleophoridae, Gelechiidae, (10) Concealed feeders Coleoptera: Cleridae (2) None Lepidoptera: Erebidae, Sesiidae, Tachinidae; Diptera: (6) Tortricidae Mostly concealed feeders None Araneidae Araneae: (1) Coleoptera: Curculionidae; Hymenoptera: Cimbicidae; Lepidoptera: Crambidae, Erebidae, Gelechiidae, Lasiocampidae, Noctuidae, Nymphalidae, Pieridae, Plutellidae, Psychidae, Pyralidae, Saturniidae, Sesiidae, Zygaenidae (55) Yponomeutidae, Tortricidae, Coleoptera: Cerambycidae; Lepidoptera: Tortricidae (2) Tortricidae Lepidoptera: Cerambycidae; Coleoptera: Araneae: Araneidae, Araneae: Linyphiidae, Tetragnathidae, Thomisidae; Coleoptera: Cerambycidae, Theridiidae, Lepidoptera: Tenthredinidae; Hymenoptera: Geometridae (12) Apidae, Coleoptera: Cerambycidae; Hymenoptera: Cephidae, Chrysididae, Crabronidae, Cynipidae, Xiphydriidae; Lepidoptera: Lasiocampidae Vespidae, (27) Mostly Crabronidae ,

Canwood , Pembina PP , Pigeon L Red , Wagner , Pigeon L Wagner Jenner , Lethbridge, Wagner NA , Wagner Glenlea George L , Vermilion; SK: , Vermilion; ; MB: Aweme, Swan River, Swan River, Aweme, ; MB: ; MB: Winnipeg m m , Pembina PP ; SK: Pike L, Saskatoon m George L , Pembina PP m George L , Pakowki dunes NA; Wagner , Wabamun, Lethbridge, George L , Eagle Hill Cr, Regina, Val Val Regina, Hills , Eagle Hill Cr, Cypress m , Katepwa P, Saskatoon, Swift Current; MB: , Katepwa P, Hill , , Stead Spruce Woods Franklin , Red Deer River, , Sylvan L, Waterton , Sylvan L, George L George L George L m , Red Deer, Wagner NA, Waterton, Waterton Waterton NA, Waterton, Wagner Pigeon L , Red Deer, b m ; SK: Indian Head, southwest m Lakes; SK: Bodmin Russell, Kenville , Red Deer River, Aweme, White Fox; MB: Marie, across region, especially in parkland Widespread ; Winnipeg Swan River, Manitou, Regina at Kandahar, NA; SK: Dafoe, Quill L Localities AB: Edmonton, NA AB: Cooking L, Wagner AB: Edmonton , George L Lakes, several locations in prairie/ Waterton NA, Wagner Vermilion, AB: parkland NA AB: Edmonton, Wagner AB: Lethbridge George L AB: Bilby, Transcona AB: Blackfoot Hills, Drumheller; MB: AB: Edmonton, Waterton, AB: Edmonton, Red Deer, Cooking L , Edmonton, Calgary, AB: Bilby, Edmonton, George L , Calgary, Armena, Bilby, AB: Pembina PP , AB: Clymont, Edmonton, AB: Cooking L , Edmonton, NA; SK: Cypress Hills; MB: Deepdale, Keld, Wagner , Wabamun Deer, Treesbank Swan River , Teulon, MB: Waterton; NA, Wagner AB: Edmonton, AB: Pigeon L AB: Edmonton, Lakes NP Wateron AB: , Waterton Lakes NA , Waterton Pigeon L , Wagner southwest Teulon; Widespread across prairies Widespread Teulon; Cypress Hills P PF , Cypress Brandon, Aweme, MB: Aweme, Teulon MB: Aweme, 1,109,H 1,5,105,H 1,2,5,9,N 1,109,N 109,N 1,N 1,2,109,N

1,2,5,9,109,N 109,N 1,5,105,109,H 5,109,H 2,109,N 1,2,5,9,105,109,N (Provancher, (Provancher, (Say, 1835) (Say, (Gravenhorst, Howard, 1892 (Kriechbaumer, (Kriechbaumer, (Ashmead, 1902) (Townes, 1960) (Townes, Cresson, 1865 (Townes, 1960) (Townes, Cresson, 1870 (Townes 1960) (Townes (Gravenhorst, 1829) Cresson, 1865 (Townes, 1960) (Townes, a 2,109,H 1,2,5,9,105,H 1,2,5,109,H 1,105,H 1,5,9,105,109,N,T,Oc Itoplectis vesca *Itoplectis conquisitor Itoplectis curticauda 1887) 1960) Liotryphon masoni (Townes, *Pimpla pedalis Pimpla stricklandi Pimpla tenuicornis SUBFAMILY Species carbonator (Gravenhorst, Oxyrrhexis 1807) Itoplectis fustiger Itoplectis quadricingulata 1880) Itoplectis viduata Cresson, 1870 Polysphincta burgessii Polysphincta koebeli Scambus atrocoxalis Liotryphon dentatus Scambus brevicornis Scambus brevicornis 1829) (Panzer, 1804) Perithous scurra (Panzer, Pimpla aquilonia Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 381 Mostly concealed Underbark Clerids Mostly spiders Hymenoptera: Braconidae, Diprionidae, Lepidoptera: Tenthredinidae; Ichneumonidae, Choreutidae, Coleophoridae, Erebidae, Gelechiidae, Geometridae, Gracillariidae, Lasiocampidae, Tortricidae, Oecophoridae, Psychidae, Pyralidae, Zygaenidae (63) Yponomeutidae, feeders Host Family (No. spp.) Guild/Notes Coleoptera: Cerambycidae, Chrysomelidae, Hymenoptera: Tachinidae; Curculionidae; Diptera: Braconidae, Diprionidae, Ichneumonidae; Lepidoptera: Choreutidae, Coleophoridae, Cosmopterigidae, Crambidae, Epermeniidae, Erebidae, Gelechiidae, Geometridae, Gracillariidae, Hesperiidae, Lasiocampidae, Noctuidae, Nolidae, Notodontidae, Nymphalidae, Papilionidae, Pieridae, Plutellidae, Psychidae, Pterophoridae, Pyralidae, Saturniidae, (155) Tortricidae Sesiidae, Mostly concealed feeders Araneidae, Araneae: Theridiidae; Coleoptera: Curculionidae; Lepidoptera: Coleophoridae, Elachistidae, Erebidae, Gelechiidae, Geometridae, (19) Tortricidae Lasiocampidae, Coleoptera: Cerambycidae; Hymenoptera: Diprionidae; Lepidoptera: Erebidae, Geometridae, Hesperiidae, Lasiocampidae, Noctuidae, Notodontidae, Pieridae, (42) Tortricidae None Curculionidae; Buprestidae, Cerambycidae, Coleoptera: Hymenoptera: Tephritidae; Diptera: Scathophagidae, Tenthredinidae; Braconidae, Cynipidae, Diprionidae, Lepidoptera: Choreutidae, Coleophoridae, Elachistidae, Erebidae, Gelechiidae, Geometridae, Gracillariidae, Lasiocampidae, Momphidae, Noctuidae, Pieridae, Tischeriidae, Plutellidae, Pterophoridae, Pyralidae, (127) Tortricidae Mostly concealed feeders Hymenoptera: Braconidae; Lepidoptera: Gelechiidae, Zygaenidae (5) None Tortricidae Lepidoptera: Coleophoridae, Gelechiidae, (10) Concealed feeders Coleoptera: Cleridae (2) None Lepidoptera: Erebidae, Sesiidae, Tachinidae; Diptera: (6) Tortricidae Mostly concealed feeders None Araneidae Araneae: (1) Coleoptera: Curculionidae; Hymenoptera: Cimbicidae; Lepidoptera: Crambidae, Erebidae, Gelechiidae, Lasiocampidae, Noctuidae, Nymphalidae, Pieridae, Plutellidae, Psychidae, Pyralidae, Saturniidae, Sesiidae, Zygaenidae (55) Yponomeutidae, Tortricidae, Coleoptera: Cerambycidae; Lepidoptera: Tortricidae (2) Tortricidae Lepidoptera: Cerambycidae; Coleoptera: Araneae: Araneidae, Araneae: Linyphiidae, Tetragnathidae, Thomisidae; Coleoptera: Cerambycidae, Theridiidae, Lepidoptera: Tenthredinidae; Hymenoptera: Geometridae (12) Apidae, Coleoptera: Cerambycidae; Hymenoptera: Cephidae, Chrysididae, Crabronidae, Cynipidae, Xiphydriidae; Lepidoptera: Lasiocampidae Vespidae, (27) Mostly Crabronidae ,

Canwood , Pembina PP , Pigeon L Red , Wagner , Pigeon L Wagner Jenner , Lethbridge, Wagner NA , Wagner Glenlea George L , Vermilion; SK: , Vermilion; ; MB: Aweme, Swan River, Swan River, Aweme, ; MB: ; MB: Winnipeg m m , Pembina PP ; SK: Pike L, Saskatoon m George L , Pembina PP m George L , Pakowki dunes NA; Wagner , Wabamun, Lethbridge, George L , Eagle Hill Cr, Regina, Val Val Regina, Hills , Eagle Hill Cr, Cypress m , Katepwa P, Saskatoon, Swift Current; MB: , Katepwa P, Hill , , Stead Spruce Woods Franklin , Red Deer River, , Sylvan L, Waterton , Sylvan L, George L George L George L m , Red Deer, Wagner NA, Waterton, Waterton Waterton NA, Waterton, Wagner Pigeon L , Red Deer, b m ; SK: Indian Head, southwest m Lakes; SK: Bodmin Russell, Kenville , Red Deer River, Aweme, White Fox; MB: Marie, across region, especially in parkland Widespread ; Winnipeg Swan River, Manitou, Regina at Kandahar, NA; SK: Dafoe, Quill L Localities AB: Edmonton, NA AB: Cooking L, Wagner AB: Edmonton , George L Lakes, several locations in prairie/ Waterton NA, Wagner Vermilion, AB: parkland NA AB: Edmonton, Wagner AB: Lethbridge George L AB: Bilby, Transcona AB: Blackfoot Hills, Drumheller; MB: AB: Edmonton, Waterton, AB: Edmonton, Red Deer, Cooking L , Edmonton, Calgary, AB: Bilby, Edmonton, George L , Calgary, Armena, Bilby, AB: Pembina PP , AB: Clymont, Edmonton, AB: Cooking L , Edmonton, NA; SK: Cypress Hills; MB: Deepdale, Keld, Wagner , Wabamun Deer, Treesbank Swan River , Teulon, MB: Waterton; NA, Wagner AB: Edmonton, AB: Pigeon L AB: Edmonton, Lakes NP Wateron AB: , Waterton Lakes NA , Waterton Pigeon L , Wagner southwest Teulon; Widespread across prairies Widespread Teulon; Cypress Hills P PF , Cypress Brandon, Aweme, MB: Aweme, Teulon MB: Aweme, 1,109,H 1,5,105,H 1,2,5,9,N 1,109,N 109,N 1,N 1,2,109,N

1,2,5,9,109,N 109,N 1,5,105,109,H 5,109,H 2,109,N 1,2,5,9,105,109,N (Provancher, (Provancher, (Say, 1835) (Say, (Gravenhorst, Howard, 1892 (Kriechbaumer, (Kriechbaumer, (Ashmead, 1902) (Townes, 1960) (Townes, Cresson, 1865 (Townes, 1960) (Townes, Cresson, 1870 (Townes 1960) (Townes (Gravenhorst, 1829) Cresson, 1865 (Townes, 1960) (Townes, a 2,109,H 1,2,5,9,105,H 1,2,5,109,H 1,105,H 1,5,9,105,109,N,T,Oc Itoplectis vesca *Itoplectis conquisitor Itoplectis curticauda 1887) 1960) Liotryphon masoni (Townes, *Pimpla pedalis Pimpla stricklandi Pimpla tenuicornis SUBFAMILY Species carbonator (Gravenhorst, Oxyrrhexis 1807) Itoplectis fustiger Itoplectis quadricingulata 1880) Itoplectis viduata Cresson, 1870 Polysphincta burgessii Polysphincta koebeli Scambus atrocoxalis Liotryphon dentatus Scambus brevicornis Scambus brevicornis 1829) (Panzer, 1804) Perithous scurra (Panzer, Pimpla aquilonia 382 M. D. Schwarzfeld

Concealed feeders Mostly gall makers Mostly Mostly concealed feeders Lepidoptera: Pyralidae, Tortricidae (2) Tortricidae Lepidoptera: Pyralidae, Concealed conifer feeders Host Family (No. spp.) Guild/Notes Hymenoptera: Diprionidae; Lepidoptera: Coleophoridae, Gelechiidae, Gracillariidae, (18) Tortricidae Incurvariidae, concealed feeders Coleoptera: Chrysomelidae; Hymenoptera: Braconidae, Cynipidae, Diprionidae, Ichneumonidae, Lepidoptera: Choreutidae, Tenthredinidae; Coleophoridae, Cosmopterigidae, Crambidae, Gelechiidae, Geometridae, Gracillariidae, Erebidae, Nolidae, Psychidae, Lasiocampidae, Lyonetiidae, Tischeriidae, Pyralidae, Saturniidae, Sesiidae, (93) Tortricidae Anthomyiidae; Lepidoptera: Pyralidae, Diptera: (6) Tortricidae Concealed conifer feeders Hymenoptera: Braconidae, Tachinidae; Diptera: Ichneumonidae; Lepidoptera: Crambidae, Drepanidae, Erebidae, Geometridae, Hesperiidae, Lasiocampidae, Noctuidae, Notodontidae, Nymphalidae, Lycaenidae, Papilionidae, Pieridae, Psychidae, Saturniidae, (90) Yponomeutidae Tortricidae, Araneidae, Clubionidae, Miturgidae, Araneae: Acrolepiidae, Elachistidae, Salticidae; Lepidoptera: Lasiocampidae, Noctuidae (9) Hymenoptera: Diprionidae; Lepidoptera: (5) Tortricidae None Lepidoptera: Momphidae (1) Stem borers Coleoptera: Curculionidae; Hymenoptera: Lepidoptera: Cosmopterigidae, Tenthredinidae; Crambidae, Gelechiidae, Momphidae, Noctuidae, (23) Tortricidae Pterophoridae, Concealed feeders (3) Tortricidae Lepidoptera: Coleoptera: Curculionidae; Hymenoptera: Cynipidae, Lepidoptera: Tenthredinidae; Ichneumonidae, Tortricidae; Gracillariidae, Tischeriidae, Sesiidae, Raphidioptera: Raphidiidae (76) Araneae: Clubionidae (3) Araneae: Clubionidae (2) Araneidae, Araneae: Linyphiidae, Theridiidae, Curculionidae (9) Mostly spiders Araneidae, Linyphiidae, Philodromidae, Araneae: Hymenoptera: Chrysomelidae; Coleoptera: Theridiidae; Lepidoptera: Drepanidae, Tenthredinidae; Diprionidae, Erebidae, Gelechiidae, Geometridae, Noctuidae, (36) Tortricidae Notodontidae, Papilionidae, Mostly spiders Araneidae, Araneae: Lepidoptera: Tetragnathidae; (6) Tortricidae Elachistidae, Araneae: Dictynidae, Linyphiidae; Coleoptera: Scolytidae (5) Mostly spiders Theriidae Araneae: (1) Lepidoptera: Yponomeutidae (1) Yponomeutidae Lepidoptera: None m , Pigeon L , Bodmin Hills , Wabamun, , Pigeon L Wabamun, , Pigeon L ; SK: , Roblin ; midcentral , Lethbridge, ; MB: Aweme, nr Aweme, NA ; MB: , Wagner NA , Wagner , Red Deer, , Pigeon L Red Deer, George L ; SK: Indian Head, Sutherland , Pembina PP m m George L , Pembina PP , Pembina PP , Red Deer, Rockyford, Waterton Waterton Rockyford, , Pigeon L Red Deer, George L , southwest Edmonton , Pembina PP m , nr Lloydminster m George L George L George L Jenner m Pembina PP , southwest , Winnipeg b m Lethbridge, Kenville Aweme, White Fox; MB: Lakes; SK: Canada (AB, SK) in sunflower fields; several locations across prairies NA , Wagner Wabamun Localities AB: Elkwater P AB: Edmonton , Wagner AB: Edmonton, AB: nr Medicine Hat AB: Wagner Edmonton, George L Edmonton, Elkwater, AB: Cooking L , Cochrane,Cowley, AB: Edmonton, SK: Indian Head SK: Great Sand Hills Wagner; Vermilion, AB: Edmonton, Lethbridge, White Fox; of Swift Current), Rutland, Saskatoon, Current, (W Winnipeg Shoal L, Aweme, MB: NA AB: Wagner AB: Blackfoot Hills, MB: Stockton Edmonton, AB: Bilby, NA, Waterton Wagner AB: Edmonton, SK: Waterton; Edmonton, nr Cooking L, AB: Calgary, Cooking L , Edmonton, AB: Bilby, SK: Cypress Hills, Swift Waterton; NA, Wagner Wabamun, Sylvan L, Aweme White Fox; MB: Current, Atton's L, NA; SK: Wagner Wabamun, AB: Edmonton, Red Deer, Bateman, Saskatoon, Swift Current AB: Edmonton, AB: Edmonton, Lethbridge, NA Wagner AB: Edmonton, MB: Winnipeg Winnipeg Snowden; MB: Winnipeg, Birtle Winnipeg, Snowden; MB: 1,2,5,H 1,5,109,H 1,109,H 1,5,109,H 2,5,9,109,H 1,2,5,N 1,H 2,14,H,T 1,105,N 1,2,N 1,105,N 1,105,N 1,2,5,109,H 9,N 2,N 2,109,N 1,5,9,8,105,109,N 1,109,N 1,9,105,109,N (Holmgren, 1856) Walley, 1960 Walley, (Holmgren, 1860) (Boheman, 1821) (Poda, 1761) Townes, 1960 Townes, Cresson, 1870 a Walley, 1960 Scambus granulosus Walley, Walley, 1960 Scambus imparis Walley, Walley, 1960 Scambus canadensis Walley, Scambus dioryctriae (Ashmead, 1890) Scambus pterophori 1800) (Panzer, Schizopyga circulator ovivora Tromatobia Zatypota anomala (Holmgren, 1860) SUBFAMILY Species 1844) Scambus vesicarius (Ratzeburg, 5,9,46,105,109,115,H,O 1,2,5,109,H,T,Oc Walley, 1960 Scambus deceptor Walley, 1960 Scambus decorus Walley, *Scambus hispae (Harris, 1835) 1960 Walley Scambus tenebrosus Schizopyga frigida Sinarachna pallipes atalantae Theronia variabilis Tromatobia *Zabrachypus primus Cushman, 1920 Zatypota crassipes Walley, 1960 Scambus longicorpus Walley, Zaglyptus varipes (Gravenhorst, 1829) Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 383

Concealed feeders Mostly gall makers Mostly Mostly concealed feeders Lepidoptera: Pyralidae, Tortricidae (2) Tortricidae Lepidoptera: Pyralidae, Concealed conifer feeders Host Family (No. spp.) Guild/Notes Hymenoptera: Diprionidae; Lepidoptera: Coleophoridae, Gelechiidae, Gracillariidae, (18) Tortricidae Incurvariidae, concealed feeders Coleoptera: Chrysomelidae; Hymenoptera: Braconidae, Cynipidae, Diprionidae, Ichneumonidae, Lepidoptera: Choreutidae, Tenthredinidae; Coleophoridae, Cosmopterigidae, Crambidae, Gelechiidae, Geometridae, Gracillariidae, Erebidae, Nolidae, Psychidae, Lasiocampidae, Lyonetiidae, Tischeriidae, Pyralidae, Saturniidae, Sesiidae, (93) Tortricidae Anthomyiidae; Lepidoptera: Pyralidae, Diptera: (6) Tortricidae Concealed conifer feeders Hymenoptera: Braconidae, Tachinidae; Diptera: Ichneumonidae; Lepidoptera: Crambidae, Drepanidae, Erebidae, Geometridae, Hesperiidae, Lasiocampidae, Noctuidae, Notodontidae, Nymphalidae, Lycaenidae, Papilionidae, Pieridae, Psychidae, Saturniidae, (90) Yponomeutidae Tortricidae, Araneidae, Clubionidae, Miturgidae, Araneae: Acrolepiidae, Elachistidae, Salticidae; Lepidoptera: Lasiocampidae, Noctuidae (9) Hymenoptera: Diprionidae; Lepidoptera: (5) Tortricidae None Lepidoptera: Momphidae (1) Stem borers Coleoptera: Curculionidae; Hymenoptera: Lepidoptera: Cosmopterigidae, Tenthredinidae; Crambidae, Gelechiidae, Momphidae, Noctuidae, (23) Tortricidae Pterophoridae, Concealed feeders (3) Tortricidae Lepidoptera: Coleoptera: Curculionidae; Hymenoptera: Cynipidae, Lepidoptera: Tenthredinidae; Ichneumonidae, Tortricidae; Gracillariidae, Tischeriidae, Sesiidae, Raphidioptera: Raphidiidae (76) Araneae: Clubionidae (3) Araneae: Clubionidae (2) Araneidae, Araneae: Linyphiidae, Theridiidae, Curculionidae (9) Mostly spiders Araneidae, Linyphiidae, Philodromidae, Araneae: Hymenoptera: Chrysomelidae; Coleoptera: Theridiidae; Lepidoptera: Drepanidae, Tenthredinidae; Diprionidae, Erebidae, Gelechiidae, Geometridae, Noctuidae, (36) Tortricidae Notodontidae, Papilionidae, Mostly spiders Araneidae, Araneae: Lepidoptera: Tetragnathidae; (6) Tortricidae Elachistidae, Araneae: Dictynidae, Linyphiidae; Coleoptera: Scolytidae (5) Mostly spiders Theriidae Araneae: (1) Lepidoptera: Yponomeutidae (1) Yponomeutidae Lepidoptera: None m , Pigeon L , Bodmin Hills , Wabamun, , Pigeon L Wabamun, , Pigeon L ; SK: , Roblin ; midcentral , Lethbridge, ; MB: Aweme, nr Aweme, NA ; MB: , Wagner NA , Wagner , Red Deer, , Pigeon L Red Deer, George L ; SK: Indian Head, Sutherland , Pembina PP m m George L , Pembina PP , Pembina PP , Red Deer, Rockyford, Waterton Waterton Rockyford, , Pigeon L Red Deer, George L , southwest Edmonton , Pembina PP m , nr Lloydminster m George L George L George L Jenner m Pembina PP , southwest , Winnipeg b m Lethbridge, Kenville Aweme, White Fox; MB: Lakes; SK: Canada (AB, SK) in sunflower fields; several locations across prairies NA , Wagner Wabamun Localities AB: Elkwater P AB: Edmonton , Wagner AB: Edmonton, AB: nr Medicine Hat AB: Wagner Edmonton, George L Edmonton, Elkwater, AB: Cooking L , Cochrane,Cowley, AB: Edmonton, SK: Indian Head SK: Great Sand Hills Wagner; Vermilion, AB: Edmonton, Lethbridge, White Fox; of Swift Current), Rutland, Saskatoon, Current, (W Winnipeg Shoal L, Aweme, MB: NA AB: Wagner AB: Blackfoot Hills, MB: Stockton Edmonton, AB: Bilby, NA, Waterton Wagner AB: Edmonton, SK: Waterton; Edmonton, nr Cooking L, AB: Calgary, Cooking L , Edmonton, AB: Bilby, SK: Cypress Hills, Swift Waterton; NA, Wagner Wabamun, Sylvan L, Aweme White Fox; MB: Current, Atton's L, NA; SK: Wagner Wabamun, AB: Edmonton, Red Deer, Bateman, Saskatoon, Swift Current AB: Edmonton, AB: Edmonton, Lethbridge, NA Wagner AB: Edmonton, MB: Winnipeg Winnipeg Snowden; MB: Winnipeg, Birtle Winnipeg, Snowden; MB: 1,2,5,H 1,5,109,H 1,109,H 1,5,109,H 2,5,9,109,H 1,2,5,N 1,H 2,14,H,T 1,105,N 1,2,N 1,105,N 1,105,N 1,2,5,109,H 9,N 2,N 2,109,N 1,5,9,8,105,109,N 1,109,N 1,9,105,109,N (Holmgren, 1856) Walley, 1960 Walley, (Holmgren, 1860) (Boheman, 1821) (Poda, 1761) Townes, 1960 Townes, Cresson, 1870 a Walley, 1960 Scambus granulosus Walley, Walley, 1960 Scambus imparis Walley, Walley, 1960 Scambus canadensis Walley, Scambus dioryctriae (Ashmead, 1890) Scambus pterophori 1800) (Panzer, Schizopyga circulator ovivora Tromatobia Zatypota anomala (Holmgren, 1860) SUBFAMILY Species 1844) Scambus vesicarius (Ratzeburg, 5,9,46,105,109,115,H,O 1,2,5,109,H,T,Oc Walley, 1960 Scambus deceptor Walley, 1960 Scambus decorus Walley, *Scambus hispae (Harris, 1835) 1960 Walley Scambus tenebrosus Schizopyga frigida Sinarachna pallipes atalantae Theronia variabilis Tromatobia *Zabrachypus primus Cushman, 1920 Zatypota crassipes Walley, 1960 Scambus longicorpus Walley, Zaglyptus varipes (Gravenhorst, 1829) 384 M. D. Schwarzfeld Wood borers Wood Host Family (No. spp.) Guild/Notes None Coleoptera, Hymenoptera: Buprestidae, Cerambycidae, Siricidae (4) borers Wood Hymenoptera: Siricidae (3) borers Wood Coleoptera: Buprestidae, Cerambycidae; Lepidoptera: (6) Tortricidae Mostly wood borers None Coleoptea: Cerambycidae; Hymenoptera: Megachilidae (2) Hymenoptera: Ichneumonidae, Siricidae, Xiphydriidae (7) borers Wood Hymenoptera: Siricidae; Lepidoptera: Notodontidae (3) Hymenoptera: Siricidae (1) Hymenoptera: Ichneumonidae, Siricidae, Xiphydriidae (7) borers Wood Hymenoptera: Xiphydriidae (3) None None Hymenoptera: Cynipidae (1) Gall makers None None Attelabidae Coleoptera: (1) (3) Tenthredinidae Hymenoptera: Diprionidae, Argidae Hymenoptera: (2) None None (3) Tenthredinidae Hymenoptera: None (1) Tenthredinidae Hymenoptera: None None None None None None (4) Tenthredinidae Hymenoptera: Coleoptera: Cerambycidae, Melandryidae; Hymenoptera: Siricidae (10) borers Wood Araneae: Araneidae, Araneae: Dictynidae, Theridiidae, (11) Tortricidae Gracillariidae; Lepidoptera: Mostly spiders Crabronidae, Hymenoptera: Cerambycidae; Coleoptera: (6) Tortricidae Megachilidae; Lepidoptera: Mostly cone borers Winnipeg , Wagner NA , Wagner NA , Wagner , Lethbridge, Waterton; MB: Waterton; , Lethbridge, Franklin George L George L George L George L , Wagner NA , Pigeon L Wagner , Pembina PP b Localities AB: Edmonton, MB: Aweme AB: George L AB: George L NA; SK: Indian Head Wagner AB: Edmonton, Midnapore, AB: Red Deer AB: Edmonton, AB: Waterton AB: Winnipeg George L AB: Calgary, NA, Waterton AB: Wagner AB: Edmonton, AB: Cochrane, AB: George L Lethbridge, Onefour, Aden, Oldman River, AB: Gilchrist Ranch, Scandia; SK: Elbow; MB: Shilo AB: Elkwater AB: Edmonton; MB: Bunch Willow PP; SK: Writing-on-Stone AB: Cameron L, Edmonton, Irvine, Lethbridge, Scandia; SK: AB: Calgary, Saskatoon Lloydminster, Cypress Hills PP, SK: Saskatoon SK: MacDowall MB: 13 mi N Glenboro, Onah Mt; MB: Horton Wood Katepwa, SK: Bagley, Edmonton, Lethbridge Wainwright, AB: Blackfoot Hills nr AB: Edmonton; SK: Saskatoon NA AB: Wagner Edmonton, Morrin; SK: Earl Grey AB: Bilby, NA Wagner Wabamun, Edmonton, AB: Bilby, Edmonton, Gleichen; SK: Dafoe, Regina Calgary, AB: Bilby, NA AB: Wagner NA AB: Wagner NA AB: Wagner Waterton Edmonton, Orkney District, AB: Bilby, NA Wagner Edmonton, AB: Bilby, 117,N 1,5,H 2,74,H 109,N 116,N 2,N 14,117,N

1,2,109,H 14,74,H 120,N 9,H,O

117,N 5,N 1,N ) 2,5,109,N 5,9,H 9,N 2,N 14,120,N 2,14,74,N 1,2,109,N 118,N 5,74,N 120,N 109,N 2,109,N 1,109,N,Oc Mason, 1955) Horstmann, 2010 (Müller, 1776) (Müller, Townes & Townes, & Townes, Townes (Davis, 1897) Horstmann, 2012 (Cresson, 1870 (Cresson, 1879) Cresson, 1868 (Davis, 1897) Townes & Townes, & Townes, Townes Townes 1960 Townes (Riley & Howard, (Mason, 1955) (Cresson, 1864) (Cresson, 1870) (Kirby, 1837) (Kirby, (Cresson, 1864) a 5,109,H,O,Oc,Au 119,N 1,5,9,109,N,Au,Oc 119,N 2,N 2,N Zatypota percontatoria Podoschistus vittifrons (Cresson, 1868) Poemenia albipes Rhyssa alaskensis Ashmead, 1902 SUBFAMILY Species Rhyssella nitida 1949 Neoxorides pilulus Poemenia thoracica Poemenia americana RHYSSINAE Megarhyssa atrata Fabricius, 1781 nortoni (Cresson, Megarhyssa 1864) Rhyssa lineolata TERSILOCHINAE Aneuclis interstitialis Barycnemis brevicauda Barycnemis harpura Schrank, 1802) Barycnemis rufipes Horstmann, 2010 1879) Barycnemis rugosa (Provancher, 1956) Diaparsis baldufi (Walkley, TRYPHONINAE 1879) Aderaeon bedardi (Provancher, Boethus schizoceri & Gupta, Townes Cosmoconus posticatus 1992 Cteniscus leptonyx (Mason, 1955) Cteniscus promediatus Cteniscus scaphuloides ( Cteniscus vitticollis Ctenochira analis (Holmgren, 1857) Ctenochira arcuata Ctenochira debilis Ctenochira deplanata 1949 Ctenochira extricata Ctenochira gillettei 1838) Ctenochira haemosterna (Haliday, 1888) Rhyssa persuasoria (Linnaeus, 1758) POEMENIINAE (Cresson, 1870) Neoxorides borealis alpestris (Holmgren, 1860) Pseudorhyssa Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 385 Wood borers Wood Host Family (No. spp.) Guild/Notes None Coleoptera, Hymenoptera: Buprestidae, Cerambycidae, Siricidae (4) borers Wood Hymenoptera: Siricidae (3) borers Wood Coleoptera: Buprestidae, Cerambycidae; Lepidoptera: (6) Tortricidae Mostly wood borers None Coleoptea: Cerambycidae; Hymenoptera: Megachilidae (2) Hymenoptera: Ichneumonidae, Siricidae, Xiphydriidae (7) borers Wood Hymenoptera: Siricidae; Lepidoptera: Notodontidae (3) Hymenoptera: Siricidae (1) Hymenoptera: Ichneumonidae, Siricidae, Xiphydriidae (7) borers Wood Hymenoptera: Xiphydriidae (3) None None Hymenoptera: Cynipidae (1) Gall makers None None Attelabidae Coleoptera: (1) (3) Tenthredinidae Hymenoptera: Diprionidae, Argidae Hymenoptera: (2) None None (3) Tenthredinidae Hymenoptera: None (1) Tenthredinidae Hymenoptera: None None None None None None (4) Tenthredinidae Hymenoptera: Coleoptera: Cerambycidae, Melandryidae; Hymenoptera: Siricidae (10) borers Wood Araneae: Araneidae, Araneae: Dictynidae, Theridiidae, (11) Tortricidae Gracillariidae; Lepidoptera: Mostly spiders Crabronidae, Hymenoptera: Cerambycidae; Coleoptera: (6) Tortricidae Megachilidae; Lepidoptera: Mostly cone borers Winnipeg , Wagner NA , Wagner NA , Wagner , Lethbridge, Waterton; MB: Waterton; , Lethbridge, Franklin George L George L George L George L , Wagner NA , Pigeon L Wagner , Pembina PP b Localities AB: Edmonton, MB: Aweme AB: George L AB: George L NA; SK: Indian Head Wagner AB: Edmonton, Midnapore, AB: Red Deer AB: Edmonton, AB: Waterton AB: Winnipeg George L AB: Calgary, NA, Waterton AB: Wagner AB: Edmonton, AB: Cochrane, AB: George L Lethbridge, Onefour, Aden, Oldman River, AB: Gilchrist Ranch, Scandia; SK: Elbow; MB: Shilo AB: Elkwater AB: Edmonton; MB: Bunch Willow PP; SK: Writing-on-Stone AB: Cameron L, Edmonton, Irvine, Lethbridge, Scandia; SK: AB: Calgary, Saskatoon Lloydminster, Cypress Hills PP, SK: Saskatoon SK: MacDowall MB: 13 mi N Glenboro, Onah Mt; MB: Horton Wood Katepwa, SK: Bagley, Edmonton, Lethbridge Wainwright, AB: Blackfoot Hills nr AB: Edmonton; SK: Saskatoon NA AB: Wagner Edmonton, Morrin; SK: Earl Grey AB: Bilby, NA Wagner Wabamun, Edmonton, AB: Bilby, Edmonton, Gleichen; SK: Dafoe, Regina Calgary, AB: Bilby, NA AB: Wagner NA AB: Wagner NA AB: Wagner Waterton Edmonton, Orkney District, AB: Bilby, NA Wagner Edmonton, AB: Bilby, 117,N 1,5,H 2,74,H 109,N 116,N 2,N 14,117,N

1,2,109,H 14,74,H 120,N 9,H,O

117,N 5,N 1,N ) 2,5,109,N 5,9,H 9,N 2,N 14,120,N 2,14,74,N 1,2,109,N 118,N 5,74,N 120,N 109,N 2,109,N 1,109,N,Oc Mason, 1955) Horstmann, 2010 (Müller, 1776) (Müller, Townes & Townes, & Townes, Townes (Davis, 1897) Horstmann, 2012 (Cresson, 1870 (Cresson, 1879) Cresson, 1868 (Davis, 1897) Townes & Townes, & Townes, Townes Townes 1960 Townes (Riley & Howard, (Mason, 1955) (Cresson, 1864) (Cresson, 1870) (Kirby, 1837) (Kirby, (Cresson, 1864) a 5,109,H,O,Oc,Au 119,N 1,5,9,109,N,Au,Oc 119,N 2,N 2,N Zatypota percontatoria Podoschistus vittifrons (Cresson, 1868) Poemenia albipes Rhyssa alaskensis Ashmead, 1902 SUBFAMILY Species Rhyssella nitida 1949 Neoxorides pilulus Poemenia thoracica Poemenia americana RHYSSINAE Megarhyssa atrata Fabricius, 1781 nortoni (Cresson, Megarhyssa 1864) Rhyssa lineolata TERSILOCHINAE Aneuclis interstitialis Barycnemis brevicauda Barycnemis harpura Schrank, 1802) Barycnemis rufipes Horstmann, 2010 1879) Barycnemis rugosa (Provancher, 1956) Diaparsis baldufi (Walkley, TRYPHONINAE 1879) Aderaeon bedardi (Provancher, Boethus schizoceri & Gupta, Townes Cosmoconus posticatus 1992 Cteniscus leptonyx (Mason, 1955) Cteniscus promediatus Cteniscus scaphuloides ( Cteniscus vitticollis Ctenochira analis (Holmgren, 1857) Ctenochira arcuata Ctenochira debilis Ctenochira deplanata 1949 Ctenochira extricata Ctenochira gillettei 1838) Ctenochira haemosterna (Haliday, 1888) Rhyssa persuasoria (Linnaeus, 1758) POEMENIINAE (Cresson, 1870) Neoxorides borealis alpestris (Holmgren, 1860) Pseudorhyssa 386 M. D. Schwarzfeld Concealed feeders Leafmining sawflies Needle miners Host Family (No. spp.) Guild/Notes None None (2) Tenthredinidae Hymenoptera: None None None None (5) Tenthredinidae Hymenoptera: None None (3) Tenthredinidae Hymenoptera: None (4) Tenthredinidae Hymenoptera: (6) Tenthredinidae Hymenoptera: (5) Tenthredinidae Hymenoptera: None None (3) Tenthredinidae Hymenoptera: None None None None None None (3) Tenthredinidae Hymenoptera: None None None None (3) Tenthredinidae Hymenoptera: None (18) Tortricidae Lepidoptera: Gracillariidae, Pyralidae, Concealed feeders None None None Hymenoptera, Tenthredinidae, Lepidoptera: Blastobasidae, Coleophoridae, Elachistidae, Gelechiidae, Pyralidae (10) Mostly concealed feeders None None Lepidoptera: Gelechiidae (3) None None b Localities NA AB: Wagner AB: Edmonton NA AB: Wagner NA Wagner AB: Edmonton, AB: Bilby MB: Beausejour AB: Edmonton, Red Deer NA Wagner Jumping Pound Cr, Aspen Beach, Elkwater P, AB: NA Wagner AB: Jumping Pound Cr, SK: Waterton; Medicine Hat, Onefour, AB: Blairmore, Elkwater P, Marie; MB:Carberry Val Cypress Hills, Saskatoon, Scout L, Jumping Pound Cr, Edmonton, Elkwater P, Aspen Beach, Bilby, AB: Albert White Fox, Prince Sundre; SK: Kenosee, NA AB: Wagner NA; SK: Elbow Wagner AB: Edmonton, NA Wagner AB: Red Deer, Milk Frank, Jumping Pound Cr, Edmonton, Elkwater P, AB: Drumheller, at Allen L SK: Waterton; NA, Wagner Red Deer, View, Mountain River, Albert Cut Knife, Prince Jumping Pound Cr, Edmonton, Elkwater L, P, AB: Bilby, 9 mi N Forrest, Aweme, SK: Saskatoon; MB: Waterton; Wabamun, Ninette Gull L AB: Bilby, AB: Edmonton, Lacombe, Red Deer; SK: Dahlton, Sutherland Edmonton; SK: Regina AB: Bilby, AB: Cowley Drumheller; SK: Saskatoon, Great Sand Hills, Roche Percee AB: Bilby, MB: Winnipeg Assiniboia, Cut Knife, Regina, Qu'Appelle SK: Swift Current, Bunch; SK: Roche Percee Willow Waterton, AB: Edmonton, L; SK: Snowden Waterton NA, Wagner AB: Edmonton, Gull L, AB: Gull L; SK: Saskatoon nr Cut Knife, Indian Head Attons L NA; SK: Wagner AB: SK: Wabamun; Edmonton, Wainwright, Blackfoot Coulee nr AB: Bilby, Harlan, White Fox SK: Roche Percee AB: Edmonton Jumping Pound Cr AB: Calgary, Calgary); SK: Canora, W AB: Edmonton, Jumping Pound Cr (20 mi Ninette Saskatoon; MB: E. Selkirk, Fort Garry, Kenosee, Lloydminster, Aweme MB: Victor; Frank, Jumping Pound Cr; SK: St. AB: Elkwater P, Acme, Drumheller; MB: Dunrae AB: Cochrane, Cypress H, Edmonton, Elkwater P, Calgary, AB: Bilby, Aweme SK: Snowden; MB: Waterton; Jumping Pound Cr, Bunch; MB: Shell River Willow AB: Edmonton; SK: Elbow, AB: Edmonton Butte AB: Twin Calgary) W AB: Jumping Pound Cr (20 mi NA Wagner AB: Edmonton, AB: Edmonton 2,N 122,N 5,119,H 5,119,N 2,14,H

74,N

74,N

14,119,N 122,N 119,N 121,N 122,N

2,5,N 119,N 5,N 5,119,H 14,119,N 119,N 122,N 122,N 119,N 14,70,119,H 122,N 122,N 119,N 2,14,N 122,N 119,N 2,119,H 5,N 5,121,N 122,N 5,121,N 122,N (Hall, 1919) Townes & Townes, & Townes, Townes Townes, 1992 Townes, (Ashmead, 1902) (Ruthe, 1855) Mason, 1959 Mason, 1959 Cushman, 1935 (Cresson, 1868) Cushman, 1922 (Ashmead, 1902) (Cresson, 1864) (Holmgren, 1857) Mason, 1959 (Cresson, 1865) (Walley, 1937) (Walley, Provancher, 1877 5, Provancher, (Ashmead, 1902) (Cushman, 1924) a 14,119,N,T 2,N 1,119,N 2,N 2,14,119,N 119,N Townes & Townes, 1949 Townes, & Ctenochira infans Townes SUBFAMILY Species 2,14,119,H Davis, 1897 *Exyston boreotis Ctenochira niveicola Ctenochira pikonematis 1949 Ctenochira rufa Ctenochira subcrassa (Cresson, 1868) 1874) Dyspetes rufus (Provancher, Eridolius clypeatus analis Brischke, 1871 Erromenus & Townes, caelator Townes Erromenus 1949 Davis, 1897 glabrosus Erromenus 1883 Provancher, marginatus Erromenus & Townes, nasalis Townes Erromenus & Townes, planus Townes Erromenus 1949 1878) punctatus (Woldstedt, Erromenus punctulatus Holmgren, 1857 Erromenus zonarius (Gravenhorst, 1820) Erromenus (Mason, 1956) Excavarus etrocaulus Excavarus velox Exyston californicus Exyston chamaeleon *Exyston clavatus Mason, 1959 Exyston hadros Exyston lophotos 1875) 2, Exyston maculosus (Provancher, 1886 Provancher, Exyston marginatus Exyston spinulosus Mason, 1959 2, Exyston variatus Exyston venustus Grypocentrus albipes Ruthe, 1855 & Townes, Grypocentrus barbatus Townes 1951 fontinalis Hercus Monoblastus ferius (Davis, 1898) Monoblastus kaniacensis Monoblastus montezuma (Cameron, 1886) Neliopisthus elegans Neliopisthus fulvicoxis Neliopisthus niger Neliopisthus piceae Netelia alaskensis Netelia barberi 1949 Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 387 Concealed feeders Leafmining sawflies Needle miners Host Family (No. spp.) Guild/Notes None None (2) Tenthredinidae Hymenoptera: None None None None (5) Tenthredinidae Hymenoptera: None None (3) Tenthredinidae Hymenoptera: None (4) Tenthredinidae Hymenoptera: (6) Tenthredinidae Hymenoptera: (5) Tenthredinidae Hymenoptera: None None (3) Tenthredinidae Hymenoptera: None None None None None None (3) Tenthredinidae Hymenoptera: None None None None (3) Tenthredinidae Hymenoptera: None (18) Tortricidae Lepidoptera: Gracillariidae, Pyralidae, Concealed feeders None None None Hymenoptera, Tenthredinidae, Lepidoptera: Blastobasidae, Coleophoridae, Elachistidae, Gelechiidae, Pyralidae (10) Mostly concealed feeders None None Lepidoptera: Gelechiidae (3) None None b Localities NA AB: Wagner AB: Edmonton NA AB: Wagner NA Wagner AB: Edmonton, AB: Bilby MB: Beausejour AB: Edmonton, Red Deer NA Wagner Jumping Pound Cr, Aspen Beach, Elkwater P, AB: NA Wagner AB: Jumping Pound Cr, SK: Waterton; Medicine Hat, Onefour, AB: Blairmore, Elkwater P, Marie; MB:Carberry Val Cypress Hills, Saskatoon, Scout L, Jumping Pound Cr, Edmonton, Elkwater P, Aspen Beach, Bilby, AB: Albert White Fox, Prince Sundre; SK: Kenosee, NA AB: Wagner NA; SK: Elbow Wagner AB: Edmonton, NA Wagner AB: Red Deer, Milk Frank, Jumping Pound Cr, Edmonton, Elkwater P, AB: Drumheller, at Allen L SK: Waterton; NA, Wagner Red Deer, View, Mountain River, Albert Cut Knife, Prince Jumping Pound Cr, Edmonton, Elkwater L, P, AB: Bilby, 9 mi N Forrest, Aweme, SK: Saskatoon; MB: Waterton; Wabamun, Ninette Gull L AB: Bilby, AB: Edmonton, Lacombe, Red Deer; SK: Dahlton, Sutherland Edmonton; SK: Regina AB: Bilby, AB: Cowley Drumheller; SK: Saskatoon, Great Sand Hills, Roche Percee AB: Bilby, MB: Winnipeg Assiniboia, Cut Knife, Regina, Qu'Appelle SK: Swift Current, Bunch; SK: Roche Percee Willow Waterton, AB: Edmonton, L; SK: Snowden Waterton NA, Wagner AB: Edmonton, Gull L, AB: Gull L; SK: Saskatoon nr Cut Knife, Indian Head Attons L NA; SK: Wagner AB: SK: Wabamun; Edmonton, Wainwright, Blackfoot Coulee nr AB: Bilby, Harlan, White Fox SK: Roche Percee AB: Edmonton Jumping Pound Cr AB: Calgary, Calgary); SK: Canora, W AB: Edmonton, Jumping Pound Cr (20 mi Ninette Saskatoon; MB: E. Selkirk, Fort Garry, Kenosee, Lloydminster, Aweme MB: Victor; Frank, Jumping Pound Cr; SK: St. AB: Elkwater P, Acme, Drumheller; MB: Dunrae AB: Cochrane, Cypress H, Edmonton, Elkwater P, Calgary, AB: Bilby, Aweme SK: Snowden; MB: Waterton; Jumping Pound Cr, Bunch; MB: Shell River Willow AB: Edmonton; SK: Elbow, AB: Edmonton Butte AB: Twin Calgary) W AB: Jumping Pound Cr (20 mi NA Wagner AB: Edmonton, AB: Edmonton 2,N 122,N 5,119,H 5,119,N 2,14,H

74,N

74,N

14,119,N 122,N 119,N 121,N 122,N

2,5,N 119,N 5,N 5,119,H 14,119,N 119,N 122,N 122,N 119,N 14,70,119,H 122,N 122,N 119,N 2,14,N 122,N 119,N 2,119,H 5,N 5,121,N 122,N 5,121,N 122,N (Hall, 1919) Townes & Townes, & Townes, Townes Townes, 1992 Townes, (Ashmead, 1902) (Ruthe, 1855) Mason, 1959 Mason, 1959 Cushman, 1935 (Cresson, 1868) Cushman, 1922 (Ashmead, 1902) (Cresson, 1864) (Holmgren, 1857) Mason, 1959 (Cresson, 1865) (Walley, 1937) (Walley, Provancher, 1877 5, Provancher, (Ashmead, 1902) (Cushman, 1924) a 14,119,N,T 2,N 1,119,N 2,N 2,14,119,N 119,N Townes & Townes, 1949 Townes, & Ctenochira infans Townes SUBFAMILY Species 2,14,119,H Davis, 1897 *Exyston boreotis Ctenochira niveicola Ctenochira pikonematis 1949 Ctenochira rufa Ctenochira subcrassa (Cresson, 1868) 1874) Dyspetes rufus (Provancher, Eridolius clypeatus analis Brischke, 1871 Erromenus & Townes, caelator Townes Erromenus 1949 Davis, 1897 glabrosus Erromenus 1883 Provancher, marginatus Erromenus & Townes, nasalis Townes Erromenus & Townes, planus Townes Erromenus 1949 1878) punctatus (Woldstedt, Erromenus punctulatus Holmgren, 1857 Erromenus zonarius (Gravenhorst, 1820) Erromenus (Mason, 1956) Excavarus etrocaulus Excavarus velox Exyston californicus Exyston chamaeleon *Exyston clavatus Mason, 1959 Exyston hadros Exyston lophotos 1875) 2, Exyston maculosus (Provancher, 1886 Provancher, Exyston marginatus Exyston spinulosus Mason, 1959 2, Exyston variatus Exyston venustus Grypocentrus albipes Ruthe, 1855 & Townes, Grypocentrus barbatus Townes 1951 fontinalis Hercus Monoblastus ferius (Davis, 1898) Monoblastus kaniacensis Monoblastus montezuma (Cameron, 1886) Neliopisthus elegans Neliopisthus fulvicoxis Neliopisthus niger Neliopisthus piceae Netelia alaskensis Netelia barberi 1949 388 M. D. Schwarzfeld Concealed feeders Concealed feeders Concealed feeders Concealed feeders Concealed feeders Concealed feeders Concealed feeders Mostly cutworms Cutworms Concealed feeders Concealed feeders Loopers Host Family (No. spp.) Guild/Notes (9) Tenthredinidae Hymenoptera: Lepidoptera: Noctuidae (2) Lepidoptera: Noctuidae (6) Lepidoptera: Noctuidae, Notodontidae (2) None Lepidoptera: Erebidae, Geometridae, Notodontidae (8) Lepidoptera: Erebidae (1) Lepidoptera: Noctuidae (1) Lepidoptera: Erebidae Lepidoptera: Noctuidae (8) Lepidoptera: Drepanidae, Geometridae (16) Loopers (23) Tortricidae Lepidoptera: Elachistidae, Pyralidae, Concealed feeders (2) Tortricidae Lepidoptera: (1) Tortricidae Lepidoptera: (3) Tortricidae Lepidoptera: None (6) Tortricidae Lepidoptera: Gelechiidae, Concealed feeders Diptera, Lepidoptera: Cecidomyiidae, Crambidae, (10) Tortricidae Noctuidae, Lepidoptera: Pyralidae (3) (10) Tortricidae Lepidoptera: Gelechiidae, Concealed feeders None (1) Tortricidae Lepidoptera: (3) Tortricidae Lepidoptera: (1) Tortricidae Lepidoptera: (27) Tortricidae Lepidoptera: Pyralidae, Concealed feeders Hymenoptera: Ichneumonidae (1) None None (7) Tenthredinidae Hymenoptera: None None (1) Tenthredinidae Hymenoptera: (1) Tenthredinidae Hymenoptera: (3) Tenthredinidae Hymenoptera: (4) Tenthredinidae Hymenoptera: None None , 5 mi W W , 5 mi

Franklin b Shilo, Souris, Treesbank Localities AB: Edmonton, Lethbridge AB: Treesbank MB: Franklin AB: Edmonton AB: Edmonton MB: Swan River AB: Lethbridge; MB: MB: Treesbank MB: Franklin AB: Edmonton NA; SK: Glaslyn, Rockglen; Wagner Chin, Lethbridge, AB: Calgary, MB: Deepdale AB: Hespero, Cypress Hills; SK: Mossbank AB: Elkwater Prairie 4200' Waterton Butte, Twin AB: Edmonton, Elkwater P, AB: Mountain View AB: Elkwater Edmonton, Drumheller, AB: Blackfoot Coulee, Brooks, Calgary, Calgary), W Jumping Pound Cr (20 mi Lethbridge, Lloydminster, Assiniboia, Battle River, SK: Wainwright; Vermilion, Morrin, Indian Head, Kenosee, Mossbank, Great Deer, Elbow, Earl Grey, Regina, Saskatoon, Swift Current; Albert, Redberry, Nipawin, Prince MB: 5 mi N Minnedosa Transcona SK: Fort a la Corne; MB: Waterton; Sylvan L, Red Deer, AB: Edmonton, Onefour, Mt FR, Turtle MB: Waldron; SK: Balgonie, Cut Knife, Hubbard, Shilo, Stonewall 5 mi SW AB: Orion; MB: Glenboro Ninette MB: Aweme, MB: Ninette AB: Chedderville NA; SK: Nipawin, Pike L, Poplar Cr; Wagner Edmonton, AB: Calgary, Shilo MB: Dauphin, Durban, 5 mi SW Brandon AB: Edmonton; MB: 6 mi NW NA; SK: Wagner Wabamun, Edmonton, Elkwater P, AB: Bilby, Shilo, Souris Lloydminster; MB: 5 mi SW Shilo 5 mi SW Aweme, AB: Lundbreck; MB: SK: Canora; MB: Minnedosa AB: Edmonton Vermilion Jumping Pound Cr, AB: Elkwater L, P, NA Wagner AB: Longview, Gull L, Elkwater P, Cypress Hills, Edmonton, Elkwater, AB: Castor, SK: Waterton; NA, Wagner Wabamun, Red Deer, Jumping Pound Cr, Rockglen, Saskatoon, Mossbank, Redberry, Earl Grey, Canora, Elbow, Dawson Rd, Gilbert Plains Carberry, Aweme, Scout L; MB: Oldman Jumping Pound Cr, AB: Edmonton, Elkwater L, P, SK: Saskatoon, Snowden, Wainwright; NA, Wagner River nr Lethbridge, Stockton W Shilo, 2 mi Aweme, White Fox; MB: Grassy L; SK: Katepwa; Jumping Pound Cr, AB: Edmonton, Elkwater P, Stockton W MB: 2 mi AB: Edmonton AB: Elkwater P Calgary); W AB: Edmonton, Jumping Pound Cr (20 mi Mt FR Turtle MB: 119,H 5,124,N 119,N

2,5,20,124,N 14,119,H 119,H 5,123,N,T 124,N 124,N 124,N 5,124,N 2,5,9,119,H 14,119,N 5,119,N 2,5,124,N 14,H 124,N 9,N 124,N 14,N 124,N 124,N 119,H 124,N 9,N 43,N 5,H 5,9,N,T 124,N 124,N 97,N,T ) 9,N,T Holmgren, Loan, 1981 Kasparyan, Loan, 1981 Loan, 1981 Loan, 1981 Loan, 1981 Schiødte, 1838 Townes & Townes, & Townes, Townes (Ratzeburg, 1852) (Ratzeburg, Cresson, 1865 Loan, 1981 Cresson, 1870 (Cresson, 1864) Townes, 1939 Townes, Loan, 1981 Rohwer, 1920 Rohwer, (Cresson, 1864) s Holmgren, 1857 (Hartig, 1837) Loan, 1981 Townes & Townes, & Townes, Townes Townes & Townes, & Townes, Townes (Thomson, 1888) (Morley, 1913) (Morley, (Ashmead, 1900) (Viereck, 1909) (Viereck, (Cushman, 1924) (Brischke, 1880) (Olivier, 1811) 1811) (Olivier, (Cushman, 1924 a 2,14,119,N 119,N 119,N 2,119,H 2,5,119,N (Cushman, 1924) Netelia brevicornis Netelia ocellata Phytodietus coxanotus Phytodietus criddleanae Phytodietus fuscus Phytodietus plesius Phytodietus solicitanae Phytodietus varianae Phytodietus vulgaris Polyblastus provancheri 1970 SUBFAMILY Species Phytodietus rufosus Netelia chloris Netelia deceptor Netelia johnsoni Netelia latungula Netelia longipalpus Netelia pallens Netelia sayi Netelia tarsata (Cresson, 1868) Phytodietus burgessi Phytodietus conflictanae Loan, 1981 Phytodietus improbanae Phytodietus obscurus Phytodietus pleuralis Phytodietus semirufus Polyblastus alternans Polyblastus bimacula 1949 Polyblastus botrys Polyblastus cancer Polyblastus flexus Polyblastus gloriosus (Davis, 1897) Polyblastus melanostigmus 1857 Polyblastus pedalis Polyblastus pumilu Polyblastus stenocentrus Holmgren, 1857 1992 Polyblastus subterminus Townes, Polyblastus tibialis 1949 1949 Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 389 Concealed feeders Concealed feeders Concealed feeders Concealed feeders Concealed feeders Concealed feeders Concealed feeders Mostly cutworms Cutworms Concealed feeders Concealed feeders Loopers Host Family (No. spp.) Guild/Notes (9) Tenthredinidae Hymenoptera: Lepidoptera: Noctuidae (2) Lepidoptera: Noctuidae (6) Lepidoptera: Noctuidae, Notodontidae (2) None Lepidoptera: Erebidae, Geometridae, Notodontidae (8) Lepidoptera: Erebidae (1) Lepidoptera: Noctuidae (1) Lepidoptera: Erebidae Lepidoptera: Noctuidae (8) Lepidoptera: Drepanidae, Geometridae (16) Loopers (23) Tortricidae Lepidoptera: Elachistidae, Pyralidae, Concealed feeders (2) Tortricidae Lepidoptera: (1) Tortricidae Lepidoptera: (3) Tortricidae Lepidoptera: None (6) Tortricidae Lepidoptera: Gelechiidae, Concealed feeders Diptera, Lepidoptera: Cecidomyiidae, Crambidae, (10) Tortricidae Noctuidae, Lepidoptera: Pyralidae (3) (10) Tortricidae Lepidoptera: Gelechiidae, Concealed feeders None (1) Tortricidae Lepidoptera: (3) Tortricidae Lepidoptera: (1) Tortricidae Lepidoptera: (27) Tortricidae Lepidoptera: Pyralidae, Concealed feeders Hymenoptera: Ichneumonidae (1) None None (7) Tenthredinidae Hymenoptera: None None (1) Tenthredinidae Hymenoptera: (1) Tenthredinidae Hymenoptera: (3) Tenthredinidae Hymenoptera: (4) Tenthredinidae Hymenoptera: None None , 5 mi W W , 5 mi

Franklin b Shilo, Souris, Treesbank Localities AB: Edmonton, Lethbridge AB: Treesbank MB: Franklin AB: Edmonton AB: Edmonton MB: Swan River AB: Lethbridge; MB: MB: Treesbank MB: Franklin AB: Edmonton NA; SK: Glaslyn, Rockglen; Wagner Chin, Lethbridge, AB: Calgary, MB: Deepdale AB: Hespero, Cypress Hills; SK: Mossbank AB: Elkwater Prairie 4200' Waterton Butte, Twin AB: Edmonton, Elkwater P, AB: Mountain View AB: Elkwater Edmonton, Drumheller, AB: Blackfoot Coulee, Brooks, Calgary, Calgary), W Jumping Pound Cr (20 mi Lethbridge, Lloydminster, Assiniboia, Battle River, SK: Wainwright; Vermilion, Morrin, Indian Head, Kenosee, Mossbank, Great Deer, Elbow, Earl Grey, Regina, Saskatoon, Swift Current; Albert, Redberry, Nipawin, Prince MB: 5 mi N Minnedosa Transcona SK: Fort a la Corne; MB: Waterton; Sylvan L, Red Deer, AB: Edmonton, Onefour, Mt FR, Turtle MB: Waldron; SK: Balgonie, Cut Knife, Hubbard, Shilo, Stonewall 5 mi SW AB: Orion; MB: Glenboro Ninette MB: Aweme, MB: Ninette AB: Chedderville NA; SK: Nipawin, Pike L, Poplar Cr; Wagner Edmonton, AB: Calgary, Shilo MB: Dauphin, Durban, 5 mi SW Brandon AB: Edmonton; MB: 6 mi NW NA; SK: Wagner Wabamun, Edmonton, Elkwater P, AB: Bilby, Shilo, Souris Lloydminster; MB: 5 mi SW Shilo 5 mi SW Aweme, AB: Lundbreck; MB: SK: Canora; MB: Minnedosa AB: Edmonton Vermilion Jumping Pound Cr, AB: Elkwater L, P, NA Wagner AB: Longview, Gull L, Elkwater P, Cypress Hills, Edmonton, Elkwater, AB: Castor, SK: Waterton; NA, Wagner Wabamun, Red Deer, Jumping Pound Cr, Rockglen, Saskatoon, Mossbank, Redberry, Earl Grey, Canora, Elbow, Dawson Rd, Gilbert Plains Carberry, Aweme, Scout L; MB: Oldman Jumping Pound Cr, AB: Edmonton, Elkwater L, P, SK: Saskatoon, Snowden, Wainwright; NA, Wagner River nr Lethbridge, Stockton W Shilo, 2 mi Aweme, White Fox; MB: Grassy L; SK: Katepwa; Jumping Pound Cr, AB: Edmonton, Elkwater P, Stockton W MB: 2 mi AB: Edmonton AB: Elkwater P Calgary); W AB: Edmonton, Jumping Pound Cr (20 mi Mt FR Turtle MB: 119,H 5,124,N 119,N

2,5,20,124,N 14,119,H 119,H 5,123,N,T 124,N 124,N 124,N 5,124,N 2,5,9,119,H 14,119,N 5,119,N 2,5,124,N 14,H 124,N 9,N 124,N 14,N 124,N 124,N 119,H 124,N 9,N 43,N 5,H 5,9,N,T 124,N 124,N 97,N,T ) 9,N,T Holmgren, Loan, 1981 Kasparyan, Loan, 1981 Loan, 1981 Loan, 1981 Loan, 1981 Schiødte, 1838 Townes & Townes, & Townes, Townes (Ratzeburg, 1852) (Ratzeburg, Cresson, 1865 Loan, 1981 Cresson, 1870 (Cresson, 1864) Townes, 1939 Townes, Loan, 1981 Rohwer, 1920 Rohwer, (Cresson, 1864) s Holmgren, 1857 (Hartig, 1837) Loan, 1981 Townes & Townes, & Townes, Townes Townes & Townes, & Townes, Townes (Thomson, 1888) (Morley, 1913) (Morley, (Ashmead, 1900) (Viereck, 1909) (Viereck, (Cushman, 1924) (Brischke, 1880) (Olivier, 1811) 1811) (Olivier, (Cushman, 1924 a 2,14,119,N 119,N 119,N 2,119,H 2,5,119,N (Cushman, 1924) Netelia brevicornis Netelia ocellata Phytodietus coxanotus Phytodietus criddleanae Phytodietus fuscus Phytodietus plesius Phytodietus solicitanae Phytodietus varianae Phytodietus vulgaris Polyblastus provancheri 1970 SUBFAMILY Species Phytodietus rufosus Netelia chloris Netelia deceptor Netelia johnsoni Netelia latungula Netelia longipalpus Netelia pallens Netelia sayi Netelia tarsata (Cresson, 1868) Phytodietus burgessi Phytodietus conflictanae Loan, 1981 Phytodietus improbanae Phytodietus obscurus Phytodietus pleuralis Phytodietus semirufus Polyblastus alternans Polyblastus bimacula 1949 Polyblastus botrys Polyblastus cancer Polyblastus flexus Polyblastus gloriosus (Davis, 1897) Polyblastus melanostigmus 1857 Polyblastus pedalis Polyblastus pumilu Polyblastus stenocentrus Holmgren, 1857 1992 Polyblastus subterminus Townes, Polyblastus tibialis 1949 1949 390 M. D. Schwarzfeld Gupta Walley Walley Graham 112 44 MacNay 72 24 Turnock and Turnock Mason 1956, Barron 1976, Criddle 1921, 16 82 Mitchell 1950, Mitchell 121 92 54 Wallis-Roughley Wallis-Roughley Sanborne 1986 a , 9 34 et al. 2010, et al. 1959, et Wylie and Loan 1984, and Loan Wylie McKay and Galloway McKay and Criddle 1925, Criddle 62 102 43 Wood borers Wood et al. 2002, Mason 1955, Turnock 1988, Turnock Hobbs et al. 1987, Still and Wong 1973, Wong and Still Viereck 1926, Viereck Swailes 1960, 15 23 120 91 111 53 Walley 1941, Walley Olfert Reschikov 33 81 71 Lee 1991, Lee 101 Sharkey 8 Walley 1940 a , Walley 42 et al. 1992, Dasch 1979, Barron 1977, Barron 52 et al. 2000, Dasch 1964, Walley 1947, Walley Strickland 1952, Strickland 110 80 32 Heinrich 1962 b , 14 Dasch 1971, 90 Townes Townes 100 119 Viereck 1925–1926 a , Viereck McKay and Galloway 1999, Galloway McKay and 22 61 Peterson 1958, Langor Host Family (No. spp.) Guild/Notes (9) Tenthredinidae Hymenoptera: Hymenoptera: Tenthredinidae (4) Tenthredinidae Hymenoptera: None (1) Tenthredinidae Hymenoptera: (1) Tenthredinidae Hymenoptera: None None None None None None None None Coleoptera: Buprestidae (2) borers Wood Coleoptera: Buprestidae, Cerambycidae (4) borers Wood Coleoptera: Buprestidae, Cerambycidae (8) borers Wood Coleoptera: Cerambycidae (2) 7 70 Wahl 1987, Wahl 51 et al. 1993, Dasch 1992, Walley 1970, Walley 79 Viereck 1925–1926 b , Viereck 31 41 Byers MDS, unpublished, Mitchell 1985, Mitchell Heinrich 1961 b , Walkley 1956, Walkley Mason 1968, 13 99 89 Schaaf 1972, 109 60 118 6 Digweed 1998, 69 Horstmann 1987, Eidt and Sippell 1961, and Sippell Eidt Walley 1927, Walley Walley 1940 b , Walley 50 21 30 78 Walley 1967, Walley 40 Dasch 1988, Norton 1863, Hopper 1939, 12 Luhman 1986, Luhman 88 108 59 Strickland 1946, Strickland Horstmann 2010, 5 Barron 1994, 68 117 Wahl and Sime 2006, and Sime Wahl 98 Barron 1986, 2010, al. et Carlson 1979, Prentice 1955, Prentice 77 Sanborne 1984, Australasian. 29 20 49 m Felt 1904, Felt Au Townes 1963 Townes 107 126 Sarfraz Dasch 1984, Luhman 1990, Luhman 4 Heinrich 1962 c , 39 58 87 Cushman 1919, Horstmann 2012, 67 Treherne 1916, Treherne Leblanc 1999, Leblanc 116 97 Afrotropical, 76 Townes and Townes 1978, Townes and Townes A Cushman 1930, 11 Walley 1969, Walley 28 Sanborne 1986 c , 106 48 Barron 1989, 3 Putnam 1982, Putnam Townes 1983, Townes b 38 57 Oceanic, Wylie and Bucher 1977, Bucher and Wylie Wong 1955, Wong Heinrich 1960 a , Oc 19 66 86 Barron 1992, Heinrich 1968, Heinrich 75 96 Townes and Townes 1949 b , Townes and Townes Localities Elkwater L, Cassils, Edmonton, Elkwater, Alder Flats, Bilby, AB: Lethbridge, Manyberries, Onefour, Gull L, Lancaster P, Elkwater P, Assiniboia, Canora, Antelope L, SK: Waterton; Orion, Red Deer, Albert, Redberry Grenfell, Kenosee, Mossbank, Pike L, Prince Elbow, Mt; MB: 6 Wood Marie, Val Saskatoon, Scout L, Victor, L, Rutland, St. Brandon mi NW AB: Edmonton, nr Medicine Hat AB: Edmonton AB: Edmonton, Lethbridge AB: Consort, Edmonton, Fawcet, Gull L, Red Deer; SK: Cut Knife, Battle River north of Cut Knife SK: Saskatoon Transcona NA; MB: Wagner AB: Edmonton, Lethbridge, Red Deer, AB: Edmonton NA Wagner Edmonton, AB: Bilby, AB: Fawcett; SK: Nipawin, Saskatoon NA AB: Wagner NA; SK: Dafoe Wagner AB: Cochrane, AB: Waterton AB: Edmonton, Red Deer SK: Indian Head AB: Calgary Edmonton AB: Bilby, Bradley 1974, Bradley 125 Oriental, 105 Stehr and Cook 1968, O 115 Putnam 1973, Putnam Finnamore 1994, 2 37 Barron 1990, 1,N 14,119,N 5,N 5,121,N 65 King and Atkinson 1928, King and Swift 1946,

Heinrich 1961 a ,

27 1,N Brown and Lindquist 1953, Barron and Walley 1983, Walley Barron and 95

85 2,5,125,N 10 47 125,N 5,N Loan 1981, 1,5,N 119,N,T 5,N 2,N 124 Wylie and Ayre 1979, Ayre and Wylie Neotropical, Townes and Gupta 1962, Gupta and Townes 18 T 56 Walley, Walley, 2010, al. et Horstmann 2008, Leblanc 1989, Leblanc 114 Townes and Townes 1949 a , Townes and Townes Parry 1995, (Cresson, 1870) Walley 1977, Walley (Cresson, 1865) 64 74 46 26 Batista Holarctic, 36 Heinrich 1960 b , Heinrich H Townes and Townes 1959, Townes and Townes Heinrich 1962 a , Townes 1938, Townes 94 Wylie 1977, Wylie Cr = Creek; FR = Forest Reserve; L = Lake; Mt = Mountain; NA = Natural Area; NP = National Park; P = Park; PP = Provincial Park; PF = Provincial Forest; Stn Forest; PF = Provincial Park; = Provincial PP = Park; P Park; = National NP Area; = Natural NA Mt = Mountain; = Lake; L Reserve; Forest FR = Cr = Creek; 84 (Provancher, 1886) (Provancher, 17 104 123 Townes and Townes 1960, Townes and Townes 1 Nearctic, Holmes 1953, a N Barron 1981, Tigner 1969, Tigner Barron 1997, Brown 1962, 119,H 63 25 73 113 45 14,121,N 14,125,N 2,14,125,N 2,14,125,N Walley 1966, Walley Mason 1959, Sanborne 1986 b , Sanborne 1962, Townes and Townes Bradley 1978, Cushman 1927, Abbreviations: Abbreviations: : Parentheses indicate the species was originally described in a different genus. Species authorities : Parentheses indicate the species was originally described in a different Polyblastus varitarsus (Gravenhorst, 1829) 1873) Scapnetes ornatus (Walsh, SUBFAMILY Species Xorides calidus 1937 1950 1950 Holmgren, 1857 Polyblastus wahlbergi incompletus Smicroplectrus 1937 Walley, robustus Smicroplectrus Thymaris kansensis (Brue, 1907) communis Cresson, 1868 Tryphon & Townes, mystax Townes Tryphon & Townes, Townes rempeli Tryphon seminiger Cresson, 1864 Tryphon 1958 townesi Walkley, Tryphon & Townes, viator Townes Tryphon 1950 XORIDINAE Odontocolon aethiops Odontocolon vicinum Xorides insularis (Cresson, 1879) 1824) Xorides stigmapterus (Say, 1931, = Station. 2000, 1982, 1954, . (2012) by J. Dombroskie (Cornell Museum of (Cornell al . (2012) by J. Dombroskie et Yu from were summarized host families The species. ichneumonid per of host species recorded Hosts: No. spp. = number families, listed of the certain or from larvae), setose or spiny having (e.g., or morphology leafrollers) (e.g., guild particular a of or mostly entirely hosts were the Where Entomology). these notes were also provided by J. Dombroskie. Museum of Entomology database, Museum of Entomology 35 55 83 93 103 R eferences: 122 a b 1944, Range: Bilodeau 1984, Bilodeau Ichneumonidae (Hymenoptera) of the Canadian Prairies Ecozone: A Review 391 Gupta Walley Walley Graham 112 44 MacNay 72 24 Turnock and Turnock Mason 1956, Barron 1976, Criddle 1921, 16 82 Mitchell 1950, Mitchell 121 92 54 Wallis-Roughley Wallis-Roughley Sanborne 1986 a , 9 34 et al. 2010, et al. 1959, et Wylie and Loan 1984, and Loan Wylie McKay and Galloway McKay and Criddle 1925, Criddle 62 102 43 Wood borers Wood et al. 2002, Mason 1955, Turnock 1988, Turnock Hobbs et al. 1987, Still and Wong 1973, Wong and Still Viereck 1926, Viereck Swailes 1960, 15 23 120 91 111 53 Walley 1941, Walley Olfert Reschikov 33 81 71 Lee 1991, Lee 101 Sharkey 8 Walley 1940 a , Walley 42 et al. 1992, Dasch 1979, Barron 1977, Barron 52 et al. 2000, Dasch 1964, Walley 1947, Walley Strickland 1952, Strickland 110 80 32 Heinrich 1962 b , 14 Dasch 1971, 90 Townes Townes 100 119 Viereck 1925–1926 a , Viereck McKay and Galloway 1999, Galloway McKay and 22 61 Peterson 1958, Langor Host Family (No. spp.) Guild/Notes (9) Tenthredinidae Hymenoptera: Hymenoptera: Tenthredinidae (4) Tenthredinidae Hymenoptera: None (1) Tenthredinidae Hymenoptera: (1) Tenthredinidae Hymenoptera: None None None None None None None None Coleoptera: Buprestidae (2) borers Wood Coleoptera: Buprestidae, Cerambycidae (4) borers Wood Coleoptera: Buprestidae, Cerambycidae (8) borers Wood Coleoptera: Cerambycidae (2) 7 70 Wahl 1987, Wahl 51 et al. 1993, Dasch 1992, Walley 1970, Walley 79 Viereck 1925–1926 b , Viereck 31 41 Byers MDS, unpublished, Mitchell 1985, Mitchell Heinrich 1961 b , Walkley 1956, Walkley Mason 1968, 13 99 89 Schaaf 1972, 109 60 118 6 Digweed 1998, 69 Horstmann 1987, Eidt and Sippell 1961, and Sippell Eidt Walley 1927, Walley Walley 1940 b , Walley 50 21 30 78 Walley 1967, Walley 40 Dasch 1988, Norton 1863, Hopper 1939, 12 Luhman 1986, Luhman 88 108 59 Strickland 1946, Strickland Horstmann 2010, 5 Barron 1994, 68 117 Wahl and Sime 2006, and Sime Wahl 98 Barron 1986, 2010, al. et Carlson 1979, Prentice 1955, Prentice 77 Sanborne 1984, Australasian. 29 20 49 m Felt 1904, Felt Au Townes 1963 Townes 107 126 Sarfraz Dasch 1984, Luhman 1990, Luhman 4 Heinrich 1962 c , 39 58 87 Cushman 1919, Horstmann 2012, 67 Treherne 1916, Treherne Leblanc 1999, Leblanc 116 97 Afrotropical, 76 Townes and Townes 1978, Townes and Townes A Cushman 1930, 11 Walley 1969, Walley 28 Sanborne 1986 c , 106 48 Barron 1989, 3 Putnam 1982, Putnam Townes 1983, Townes b 38 57 Oceanic, Wylie and Bucher 1977, Bucher and Wylie Wong 1955, Wong Heinrich 1960 a , Oc 19 66 86 Barron 1992, Heinrich 1968, Heinrich 75 96 Townes and Townes 1949 b , Townes and Townes Localities Elkwater L, Cassils, Edmonton, Elkwater, Alder Flats, Bilby, AB: Lethbridge, Manyberries, Onefour, Gull L, Lancaster P, Elkwater P, Assiniboia, Canora, Antelope L, SK: Waterton; Orion, Red Deer, Albert, Redberry Grenfell, Kenosee, Mossbank, Pike L, Prince Elbow, Mt; MB: 6 Wood Marie, Val Saskatoon, Scout L, Victor, L, Rutland, St. Brandon mi NW AB: Edmonton, nr Medicine Hat AB: Edmonton AB: Edmonton, Lethbridge AB: Consort, Edmonton, Fawcet, Gull L, Red Deer; SK: Cut Knife, Battle River north of Cut Knife SK: Saskatoon Transcona NA; MB: Wagner AB: Edmonton, Lethbridge, Red Deer, AB: Edmonton NA Wagner Edmonton, AB: Bilby, AB: Fawcett; SK: Nipawin, Saskatoon NA AB: Wagner NA; SK: Dafoe Wagner AB: Cochrane, AB: Waterton AB: Edmonton, Red Deer SK: Indian Head AB: Calgary Edmonton AB: Bilby, Bradley 1974, Bradley 125 Oriental, 105 Stehr and Cook 1968, O 115 Putnam 1973, Putnam Finnamore 1994, 2 37 Barron 1990, 1,N 14,119,N 5,N 5,121,N 65 King and Atkinson 1928, King and Swift 1946,

Heinrich 1961 a ,

27 1,N Brown and Lindquist 1953, Barron and Walley 1983, Walley Barron and 95

85 2,5,125,N 10 47 125,N 5,N Loan 1981, 1,5,N 119,N,T 5,N 2,N 124 Wylie and Ayre 1979, Ayre and Wylie Neotropical, Townes and Gupta 1962, Gupta and Townes 18 T 56 Walley, Walley, 2010, al. et Horstmann 2008, Leblanc 1989, Leblanc 114 Townes and Townes 1949 a , Townes and Townes Parry 1995, (Cresson, 1870) Walley 1977, Walley (Cresson, 1865) 64 74 46 26 Batista Holarctic, 36 Heinrich 1960 b , Heinrich H Townes and Townes 1959, Townes and Townes Heinrich 1962 a , Townes 1938, Townes 94 Wylie 1977, Wylie Cr = Creek; FR = Forest Reserve; L = Lake; Mt = Mountain; NA = Natural Area; NP = National Park; P = Park; PP = Provincial Park; PF = Provincial Forest; Stn Forest; PF = Provincial Park; = Provincial PP = Park; P Park; = National NP Area; = Natural NA Mt = Mountain; = Lake; L Reserve; Forest FR = Cr = Creek; 84 (Provancher, 1886) (Provancher, 17 104 123 Townes and Townes 1960, Townes and Townes 1 Nearctic, Holmes 1953, a N Barron 1981, Tigner 1969, Tigner Barron 1997, Brown 1962, 119,H 63 25 73 113 45 14,121,N 14,125,N 2,14,125,N 2,14,125,N Walley 1966, Walley Mason 1959, Sanborne 1986 b , Sanborne 1962, Townes and Townes Bradley 1978, Cushman 1927, Abbreviations: Abbreviations: : Parentheses indicate the species was originally described in a different genus. Species authorities : Parentheses indicate the species was originally described in a different Polyblastus varitarsus (Gravenhorst, 1829) 1873) Scapnetes ornatus (Walsh, SUBFAMILY Species Xorides calidus 1937 1950 1950 Holmgren, 1857 Polyblastus wahlbergi incompletus Smicroplectrus 1937 Walley, robustus Smicroplectrus Thymaris kansensis (Brue, 1907) communis Cresson, 1868 Tryphon & Townes, mystax Townes Tryphon & Townes, Townes rempeli Tryphon seminiger Cresson, 1864 Tryphon 1958 townesi Walkley, Tryphon & Townes, viator Townes Tryphon 1950 XORIDINAE Odontocolon aethiops Odontocolon vicinum Xorides insularis (Cresson, 1879) 1824) Xorides stigmapterus (Say, 1931, = Station. 2000, 1982, 1954, . (2012) by J. Dombroskie (Cornell Museum of (Cornell . (2012) by J. Dombroskie al et Yu from were summarized host families The species. ichneumonid per of host species recorded Hosts: No. spp. = number families, listed of the certain or from larvae), setose or spiny having (e.g., or morphology leafrollers) (e.g., guild particular a of or mostly entirely hosts were the Where Entomology). these notes were also provided by J. Dombroskie. Museum of Entomology database, Museum of Entomology 35 55 83 93 103 R eferences: 122 a b 1944, Range: Bilodeau 1984, Bilodeau 392 M. D. Schwarzfeld

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