Discovering Paleontology: Dinosaurs, Ice Age Mammals, Fossils & More Hints for Teachers Upper Elementary
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And the Origin and Evolution of the Ankylosaur Pelvis
Pelvis of Gargoyleosaurus (Dinosauria: Ankylosauria) and the Origin and Evolution of the Ankylosaur Pelvis Kenneth Carpenter1,2*, Tony DiCroce3, Billy Kinneer3, Robert Simon4 1 Prehistoric Museum, Utah State University – Eastern, Price, Utah, United States of America, 2 Geology Section, University of Colorado Museum, Boulder, Colorado, United States of America, 3 Denver Museum of Nature and Science, Denver, Colorado, United States of America, 4 Dinosaur Safaris Inc., Ashland, Virginia, United States of America Abstract Discovery of a pelvis attributed to the Late Jurassic armor-plated dinosaur Gargoyleosaurus sheds new light on the origin of the peculiar non-vertical, broad, flaring pelvis of ankylosaurs. It further substantiates separation of the two ankylosaurs from the Morrison Formation of the western United States, Gargoyleosaurus and Mymoorapelta. Although horizontally oriented and lacking the medial curve of the preacetabular process seen in Mymoorapelta, the new ilium shows little of the lateral flaring seen in the pelvis of Cretaceous ankylosaurs. Comparison with the basal thyreophoran Scelidosaurus demonstrates that the ilium in ankylosaurs did not develop entirely by lateral rotation as is commonly believed. Rather, the preacetabular process rotated medially and ventrally and the postacetabular process rotated in opposition, i.e., lateral and ventrally. Thus, the dorsal surfaces of the preacetabular and postacetabular processes are not homologous. In contrast, a series of juvenile Stegosaurus ilia show that the postacetabular process rotated dorsally ontogenetically. Thus, the pelvis of the two major types of Thyreophora most likely developed independently. Examination of other ornithischians show that a non-vertical ilium had developed independently in several different lineages, including ceratopsids, pachycephalosaurs, and iguanodonts. -
A Phylogenetic Analysis of the Basal Ornithischia (Reptilia, Dinosauria)
A PHYLOGENETIC ANALYSIS OF THE BASAL ORNITHISCHIA (REPTILIA, DINOSAURIA) Marc Richard Spencer A Thesis Submitted to the Graduate College of Bowling Green State University in partial fulfillment of the requirements of the degree of MASTER OF SCIENCE December 2007 Committee: Margaret M. Yacobucci, Advisor Don C. Steinker Daniel M. Pavuk © 2007 Marc Richard Spencer All Rights Reserved iii ABSTRACT Margaret M. Yacobucci, Advisor The placement of Lesothosaurus diagnosticus and the Heterodontosauridae within the Ornithischia has been problematic. Historically, Lesothosaurus has been regarded as a basal ornithischian dinosaur, the sister taxon to the Genasauria. Recent phylogenetic analyses, however, have placed Lesothosaurus as a more derived ornithischian within the Genasauria. The Fabrosauridae, of which Lesothosaurus was considered a member, has never been phylogenetically corroborated and has been considered a paraphyletic assemblage. Prior to recent phylogenetic analyses, the problematic Heterodontosauridae was placed within the Ornithopoda as the sister taxon to the Euornithopoda. The heterodontosaurids have also been considered as the basal member of the Cerapoda (Ornithopoda + Marginocephalia), the sister taxon to the Marginocephalia, and as the sister taxon to the Genasauria. To reevaluate the placement of these taxa, along with other basal ornithischians and more derived subclades, a phylogenetic analysis of 19 taxonomic units, including two outgroup taxa, was performed. Analysis of 97 characters and their associated character states culled, modified, and/or rescored from published literature based on published descriptions, produced four most parsimonious trees. Consistency and retention indices were calculated and a bootstrap analysis was performed to determine the relative support for the resultant phylogeny. The Ornithischia was recovered with Pisanosaurus as its basalmost member. -
A Revised Taxonomy of the Iguanodont Dinosaur Genera and Species
ARTICLE IN PRESS + MODEL Cretaceous Research xx (2007) 1e25 www.elsevier.com/locate/CretRes A revised taxonomy of the iguanodont dinosaur genera and species Gregory S. Paul 3109 North Calvert Station, Side Apartment, Baltimore, MD 21218-3807, USA Received 20 April 2006; accepted in revised form 27 April 2007 Abstract Criteria for designating dinosaur genera are inconsistent; some very similar species are highly split at the generic level, other anatomically disparate species are united at the same rank. Since the mid-1800s the classic genus Iguanodon has become a taxonomic grab-bag containing species spanning most of the Early Cretaceous of the northern hemisphere. Recently the genus was radically redesignated when the type was shifted from nondiagnostic English Valanginian teeth to a complete skull and skeleton of the heavily built, semi-quadrupedal I. bernissartensis from much younger Belgian sediments, even though the latter is very different in form from the gracile skeletal remains described by Mantell. Currently, iguanodont remains from Europe are usually assigned to either robust I. bernissartensis or gracile I. atherfieldensis, regardless of lo- cation or stage. A stratigraphic analysis is combined with a character census that shows the European iguanodonts are markedly more morpho- logically divergent than other dinosaur genera, and some appear phylogenetically more derived than others. Two new genera and a new species have been or are named for the gracile iguanodonts of the Wealden Supergroup; strongly bipedal Mantellisaurus atherfieldensis Paul (2006. Turning the old into the new: a separate genus for the gracile iguanodont from the Wealden of England. In: Carpenter, K. (Ed.), Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs. -
Phylogeny and Biogeography of Iguanodontian Dinosaurs, with Implications from Ontogeny and an Examination of the Function of the Fused Carpal-Digit I Complex
Phylogeny and Biogeography of Iguanodontian Dinosaurs, with Implications from Ontogeny and an Examination of the Function of the Fused Carpal-Digit I Complex By Karen E. Poole B.A. in Geology, May 2004, University of Pennsylvania M.A. in Earth and Planetary Sciences, August 2008, Washington University in St. Louis A Dissertation submitted to The Faculty of The Columbian College of Arts and Sciences of The George Washington University in partial fulfillment of the requirements for the degree of Doctor of Philosophy August 31, 2015 Dissertation Directed by Catherine Forster Professor of Biology The Columbian College of Arts and Sciences of The George Washington University certifies that Karen Poole has passed the Final Examination for the degree of Doctor of Philosophy as of August 10th, 2015. This is the final and approved form of the dissertation. Phylogeny and Biogeography of Iguanodontian Dinosaurs, with Implications from Ontogeny and an Examination of the Function of the Fused Carpal-Digit I Complex Karen E. Poole Dissertation Research Committee: Catherine A. Forster, Professor of Biology, Dissertation Director James M. Clark, Ronald Weintraub Professor of Biology, Committee Member R. Alexander Pyron, Robert F. Griggs Assistant Professor of Biology, Committee Member ii © Copyright 2015 by Karen Poole All rights reserved iii Dedication To Joseph Theis, for his unending support, and for always reminding me what matters most in life. To my parents, who have always encouraged me to pursue my dreams, even those they didn’t understand. iv Acknowledgements First, a heartfelt thank you is due to my advisor, Cathy Forster, for giving me free reign in this dissertation, but always providing valuable commentary on any piece of writing I sent her, no matter how messy. -
Postcranial Anatomy of Tanius Sinensis Wiman, 1929 (Dinosauria; Hadrosauroidea) Postkraniala Anatomin Hos Tanius Sinensis Wiman, 1929 (Dinosauria; Hadrosauroidea)
Examensarbete vid Institutionen för geovetenskaper Degree Project at the Department of Earth Sciences ISSN 1650-6553 Nr 320 Postcranial Anatomy of Tanius Sinensis Wiman, 1929 (Dinosauria; Hadrosauroidea) Postkraniala anatomin hos Tanius sinensis Wiman, 1929 (Dinosauria; Hadrosauroidea) Niclas H. Borinder INSTITUTIONEN FÖR GEOVETENSKAPER DEPARTMENT OF EARTH SCIENCES Examensarbete vid Institutionen för geovetenskaper Degree Project at the Department of Earth Sciences ISSN 1650-6553 Nr 320 Postcranial Anatomy of Tanius Sinensis Wiman, 1929 (Dinosauria; Hadrosauroidea) Postkraniala anatomin hos Tanius sinensis Wiman, 1929 (Dinosauria; Hadrosauroidea) Niclas H. Borinder ISSN 1650-6553 Copyright © Niclas H. Borinder and the Department of Earth Sciences, Uppsala University Published at Department of Earth Sciences, Uppsala University (www.geo.uu.se), Uppsala, 2015 Abstract Postcranial Anatomy of Tanius Sinensis Wiman, 1929 (Dinosauria; Hadrosauroidea) Niclas H. Borinder Tanius sinensis Wiman, 1929 was one of the first hadrosauroid or “duck-billed” taxa erected from China, indeed one of the very first non-avian dinosaur taxa to be erected based on material from the country. Since the original description by Wiman in 1929, the anatomy of T. sinensis has received relatively little attention in the literature since then. This is unfortunate given the importance of T. sinensis as a possible non-hadrosaurid hadrosauroid i.e. a member of Hadrosauroidea outside the family of Hadrosauridae, living in the Late Cretaceous, at a time when most non-hadrosaurid hadro- sauroids had become replaced by the members of Hadrosauridae. To gain a better understanding of the anatomy of T. sinensis and its phylogenetic relationships, the postcranial anatomy of it is redescribed. T. sinensis is found to have a mosaic of basal traits like strongly opisthocoelous cervical vertebrae, the proximal end of scapula being dorsoventrally wider than the distal end, the ratio between the proximodistal length of the metatarsal III and the mediolateral width of this element being greater than 4.5. -
Dinosaur (DK Eyewitness Books)
Eyewitness DINOSAUR www.ketabha.org Eyewitness DINOSAUR www.ketabha.org Magnolia flower Armored Polacanthus skin Rock fragment with iridium deposit Corythosaurus Tyrannosaurus coprolite (fossil dropping) Megalosaurus jaw www.ketabha.org Eyewitness Troodon embryo DINOSAUR Megalosaurus tooth Written by DAVID LAMBERT Kentrosaurus www.ketabha.org LONDON, NEW YORK, Ammonite mold MELBOURNE, MUNICH, AND DELHI Ammonite cast Consultant Dr. David Norman Senior editor Rob Houston Editorial assistant Jessamy Wood Managing editors Julie Ferris, Jane Yorke Managing art editor Owen Peyton Jones Art director Martin Wilson Gila monster Associate publisher Andrew Macintyre Picture researcher Louise Thomas Production editor Melissa Latorre Production controller Charlotte Oliver Jacket designers Martin Wilson, Johanna Woolhead Jacket editor Adam Powley DK DELHI Editor Kingshuk Ghoshal Designer Govind Mittal DTP designers Dheeraj Arora, Preetam Singh Project editor Suchismita Banerjee Design manager Romi Chakraborty Troodon Iguanodon hand Production manager Pankaj Sharma Head of publishing Aparna Sharma First published in the United States in 2010 by DK Publishing 375 Hudson Street, New York, New York 10014 Copyright © 2010 Dorling Kindersley Limited, London 10 11 12 13 14 10 9 8 7 6 5 4 3 2 1 175403—12/09 All rights reserved under International and Pan-American Copyright Conventions. No part of this publication may be reproduced, stored in a retrieval system, or transmitted in any form or by any means, electronic, mechanical, photocopying, recording, or otherwise, without the prior written permission of the copyright owner. Published in Great Britain by Dorling Kindersley Limited. A catalog record for this book is available from the Library of Congress. ISBN 978-0-7566-5810-6 (Hardcover) ISBN 978-0-7566-5811-3 (Library Binding) Color reproduction by MDP, UK, and Colourscan, Singapore Printed and bound by Toppan Printing Co. -
The PT Interview
Microraptors © Gregory S. Paul The Prehistoric Times Interview: Gregory S. Paul ByVince 1.J. Curley Vince Curley: Your Book, Predatory Dinosaurs of the World is one of the greatest dinosaur hooks ever written. I've been told it's no longer in print, which I feel is a great injustice. Is this true, and if so, is there any possibility of another edition coming out again? Gregory S. Paul: Just one of the greatest dinosaur books ever written? But seriously, the book is too out of date to be in print. And, someday .... V.C.: The Scientific pieces at the Lambeosaurus © Gregory S. Paul American book of Dinosaurs Dinosaur National is a huge undertaking, with Monument. His excellent artwork and infor- snorkeling mative contributions from so Diplodocus ·pair is many good scientists. How the most evocative did this project come about; expression of the was it your brainchild or view of sauropods something Scientific as semi-aquatic American asked you to ever achieved. accomplish? Meanwhile the Allosaurus chas- GSP: The publisher of the ing Camptosaurus book was kind enough to ask would not look out me to be the editor. That is of place in a cur- not really a huge undertak- rent book on ing, since as editor I let oth- dinosaurs. It is a ers do most of the work. shame they are filed away rather V.C.: Whom would you list than be displayed as being your biggest influ- on tbe side some- ences in your dinosaur art, in -- ..:- __ ..::.c_.-" where. I was delighted to get inditermsvidualofs?past and present '"======================================~r them published in Camptosaurus & Dryosaurus © Gregory S. -
Dinosaurs Found in Utah Record Chart
UTAH’S DINOSAUR RECORD GEOLOGIC UTAH’S MESOZOIC TIME SCALE ROCK RECORD Southern Tyrannosaurus Torosaurus & Alamosaurus fauna Tyrannosaurus Alamosaurus ankylosaurid teeth long-horned 65 NORTH HORN FORMATION centrosaurines Maastrichtian Ornithomimus new chasmosaurines Hagryphus DINOSAUR Highest diversity North America ceratopsid & KAIPAROWITS FORMATION hadrosaurid fauna Gryposaurus Edmontonia EXTINCTION 75 Campanian new tyrannosaurids Parasaurolophus new tyrannosaurid First centrosaurines Diabloceratops WAHWEAP FORMATION - MASUK FORMATION & hadrosaurids new hadrosaurine pachycephalosaurs nodosaurids Santonian Upper STRAIGHT CLIFFS FORMATION Diverse tooth fauna Blue Gate Member MANCOS SHALE First ceratopsomorphs Coniacian Zuniceratops advanced nodosaurid National Monument Ferron Sandstone - Smoky Hollow Member Nothronychus hadrosauromorphs teeth Turonian Local extinction Pachycephalosaurid TROPIC SHALE. - Tununk Member MANCOS SHALE of sauropods Grand Sraircase - Escalante teeth ornithomimid DAKOTA FORMATION Asian immigration tiny Cenomanian ornithopod Eolambia Mussentuchit Member induced extinction tyrannosaurid and origin of North teeth tiny pleurocoelid 100 troodontid teeth American Late neoceratopsian teeth Animantarx brachiosaurid teeth Cretaceous dinosaur Albian fauna Ruby Ranch Cedarpelta Member Isolation of North Tennontosaurus nodosaurids America resulting Abydosaurus in local extinctions pleurocoelid Deinonychus and increased Acrocanthosaurus brachiosaurids Aptian FORMATION endemism Nedcolbertia new camarasaurid Poison Strip Sandstone -
Osteology of a Near-Complete Skeleton of Tenontosaurus Tilletti (Dinosauria: Ornithopoda) from the Cloverly Formation, Montana, USA
Osteology of a Near-Complete Skeleton of Tenontosaurus tilletti (Dinosauria: Ornithopoda) from the Cloverly Formation, Montana, USA A report submitted in partial fulfilment of the requirements for the degree of MEarthSci Geology (Hons.), University of Manchester Jonathan Tennant 70406912 MEarthSci Geology (Hons.) School of Earth, Atmospheric and Environmental Sciences University of Manchester Williamson Building Oxford Road Manchester M13 9PL Osteology of Tenontosaurus tilletti Contents Figures and Tables..............................................................................................................................................3 1.0 Abstract .............................................................................................................................................................6 2.0 Introduction ......................................................................................................................................................7 2.1 Aims and Objectives....................................................................................................................................7 2.2 Institutional Abbreviations ........................................................................................................................9 3.0 Previous Work ............................................................................................................................................... 11 3.1 The Cloverly Formation .......................................................................................................................... -
Partial Skull and Endocranial Cast of the Ankylosaurian Dinosaur Hungarosaurus from the Late Cretaceous of Hungary: Implications for Locomotion
Palaeontologia Electronica palaeo-electronica.org Partial skull and endocranial cast of the ankylosaurian dinosaur Hungarosaurus from the Late Cretaceous of Hungary: implications for locomotion Attila Ősi, Xabier Pereda Suberbiola, and Tamás Földes ABSTRACT A partial skull of ankylosaur from the Upper Cretaceous (Santonian) Csehbánya Formation in Iharkút and the endocranial cast taken from it are described. The mor- phology of the exoccipital, the elongated ‘neck’ region of the basioccipital, the shape of the occipital condyle, and the different flexure of the medulla relative to the forebrain unambiguously differentiate this specimen from the basicranium of Struthiosaurus, so it is assigned to Hungarosaurus sp. Whereas the endocranial cast reflects a brain gener- ally similar to those of other ankylosaurs, the dorsally hypertrophied cerebellum (also present is Struthiosaurus transylvanicus) is quite unusual within the group suggesting a more sophisticated cerebral coordination of posture and movement, and perhaps a more cursorial locomotary habit than predicted for other ankylosaurs. Attila Ősi. Hungarian Academy of Sciences – Eötvös Loránd University, Lendület Dinosaur Research Group, Pázmány Péter sétány 1/c, Budapest, 1117, Hungary [email protected] Xabier Pereda Suberbiola. Universidad del País Vasco/Euskal Herriko Unibertsitatea, Facultad de Ciencia y Tecnología, Dpto. Estratigrafía y Paleontología, Apartado 644, 48080 Bilbao, Spain [email protected] Tamás Földes. Institute of Diagnostic Imaging and Radiation Oncology of the University of Kaposvár, Kaposvár, 7400 Guba S. 40, Hungary [email protected] Keywords: Santonian, Iharkút, Ankylosauria, Hungarosaurus, skull, endocranial cast INTRODUCTION gary, which includes disarticulated cranial bones (Ősi, 2005; Ősi and Makádi, 2009), mainly the type Cranial remains of ankylosaurs are among the material of Struthiosaurus, Struthiosaurus austria- rarest fossils from the Late Cretaceous of Europe. -
Matthew Carl Lamanna
Curriculum Vitae Matthew Carl Lamanna Assistant Curator Section of Vertebrate Paleontology Carnegie Museum of Natural History 4400 Forbes Avenue Pittsburgh, Pennsylvania 15213-4080 (412) 578-2696 (Office) (412) 622-8837 (Fax) Email: [email protected] Internet: http://www.carnegiemnh.org/vp/lamanna.html Education 2004 Ph.D., University of Pennsylvania, Department of Earth and Environmental Science. 1999 M.Sc., University of Pennsylvania, Department of Earth and Environmental Science. 1997 B.Sc., Hobart College, Departments of Geoscience and Biology, cum laude. Research Interests Mesozoic (principally Cretaceous) vertebrate faunas, paleoecology, and paleobiogeography; non-avian and avian dinosaur anatomy, systematics, and phylogeny. Academic and Professional Positions 2013–present Research Associate, Cleveland Museum of Natural History. 2012–present Principal Investigator and Project Director, Antarctic Peninsula Paleontology Project (AP3). 2005–present Adjunct Assistant Professor, Department of Geology and Planetary Science, University of Pittsburgh. 2004–present Assistant Curator, Section of Vertebrate Paleontology, Carnegie Museum of Natural History. 1999–present Paleontologist, Bahariya Dinosaur Project. 1997–present Research Associate, Academy of Natural Sciences of Drexel University (Philadelphia). 1997–1998 Exhibit Design Consultant, Dinosaur Hall, Academy of Natural Sciences (Philadelphia). 1995 Research Assistant, University of New Orleans Lance Dinosaur Project. Field Experience 2016 Unnamed formation, Robertson Island, -
An Inventory of Non-Avian Dinosaurs from National Park Service Areas
Lucas, S.G. and Sullivan, R.M., eds., 2018, Fossil Record 6. New Mexico Museum of Natural History and Science Bulletin 79. 703 AN INVENTORY OF NON-AVIAN DINOSAURS FROM NATIONAL PARK SERVICE AREAS JUSTIN S. TWEET1 and VINCENT L. SANTUCCI2 1National Park Service, 9149 79th Street S., Cottage Grove, MN 55016 -email: [email protected]; 2National Park Service, Geologic Resources Division, 1849 “C” Street, NW, Washington, D.C. 20240 -email: [email protected] Abstract—Dinosaurs have captured the interest and imagination of the general public, particularly children, around the world. Paleontological resource inventories within units of the National Park Service have revealed that body and trace fossils of non-avian dinosaurs have been documented in at least 21 National Park Service areas. In addition there are two historically associated occurrences, one equivocal occurrence, two NPS areas with dinosaur tracks in building stone, and one case where fossils have been found immediately outside of a monument’s boundaries. To date, body fossils of non- avian dinosaurs are documented at 14 NPS areas, may also be present at another, and are historically associated with two other parks. Dinosaur trace fossils have been documented at 17 NPS areas and are visible in building stone at two parks. Most records of NPS dinosaur fossils come from park units on the Colorado Plateau, where body fossils have been found in Upper Jurassic and Lower Cretaceous rocks at many locations, and trace fossils are widely distributed in Upper Triassic and Jurassic rocks. Two NPS units are particularly noted for their dinosaur fossils: Dinosaur National Monument (Upper Triassic through Lower Cretaceous) and Big Bend National Park (Upper Cretaceous).