GEOL 104 Dinosaurs: a Natural History
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The Stratigraphic Importance of the Brontothere (Cf. Diplacodon Elatus) in the Brennan Basin Member of the Duchesne River Formation of Utah
Foss. Rec., 17, 69–74, 2014 www.foss-rec.net/17/69/2014/ doi:10.5194/fr-17-69-2014 © Author(s) 2014. CC Attribution 3.0 License. The stratigraphic importance of the brontothere (cf. Diplacodon elatus) in the Brennan Basin Member of the Duchesne River Formation of Utah B. J. Burger and L. Tackett II Department of Geology, Utah State University, Uintah Basin Regional Campus 320 North Aggie Blvd. Vernal, UT 84078, USA Correspondence to: B. J. Burger ([email protected]) Received: 10 June 2014 – Revised: 6 August 2014 – Accepted: 12 August 2014 – Published: 27 August 2014 Abstract. We report on the first occurrence of an early southern Mississippia (Gazin and Sullivan, 1942), and the horned brontothere in the Brennan Basin Member of the Pacific Northwest (Mihlbacher, 2007). But nowhere is the Duchesne River Formation in northeastern Utah. This is the record of brontotheres as diverse as the fossil record ob- first record of a brontothere from the Brennan Basin Member. tained from the middle Eocene depositional basins located in Previously, brontotheres have been reported from the higher northeastern Utah, southwestern Wyoming and northwestern stratigraphic La Point Member (Duchesneodus uintensis) Colorado (Lull, 1905; Cook, 1926; Douglass, 1909; Gregory, and the lower stratigraphic Uinta Formation (Sphenocoelus 1912; Gunnell and Yarborough, 2000; Hatcher, 1895; Lucas uintensis, Fossendorhinus diploconus, Metarhinus fluviatilis, et al., 2004; Lucas and Holbrook, 2004; Lucas and Schoch, Metarhinus abbotti, Sthenodectes incisivum, Metatelmath- 1982; Mader, 2000, 2009a, b, Mihlbacher, 2008, 2011; Os- erium ultimum, Protitanotherium emarginatum, Pollyosbor- born, 1889, 1895, 1908, 1913, 1929; Peterson, 1914a, b, nia altidens, Diplacodon elatus). -
The Evolution of Micro-Cursoriality in Mammals
© 2014. Published by The Company of Biologists Ltd | The Journal of Experimental Biology (2014) 217, 1316-1325 doi:10.1242/jeb.095737 RESEARCH ARTICLE The evolution of micro-cursoriality in mammals Barry G. Lovegrove* and Metobor O. Mowoe* ABSTRACT Perissodactyla) in response to the emergence of open landscapes and In this study we report on the evolution of micro-cursoriality, a unique grasslands following the Eocene Thermal Maximum (Janis, 1993; case of cursoriality in mammals smaller than 1 kg. We obtained new Janis and Wilhelm, 1993; Yuanqing et al., 2007; Jardine et al., 2012; running speed and limb morphology data for two species of elephant- Lovegrove, 2012b; Lovegrove and Mowoe, 2013). shrews (Elephantulus spp., Macroscelidae) from Namaqualand, Loosely defined, cursorial mammals are those that run fast. South Africa, which we compared with published data for other However, more explicit definitions of cursoriality remain obscure mammals. Elephantulus maximum running speeds were higher than because locomotor performance is influenced by multiple variables, those of most mammals smaller than 1 kg. Elephantulus also including behaviour, biomechanics, physiology and morphology possess exceptionally high metatarsal:femur ratios (1.07) that are (Taylor et al., 1970; Garland, 1983a; Garland, 1983b; Garland and typically associated with fast unguligrade cursors. Cursoriality evolved Janis, 1993; Stein and Casinos, 1997; Carrano, 1999). In an in the Artiodactyla, Perissodactyla and Carnivora coincident with evaluation of these definition problems, Carrano (Carrano, 1999) global cooling and the replacement of forests with open landscapes argued that ‘…morphology should remain the fundamental basis for in the Oligocene and Miocene. The majority of mammal species, making distinctions between locomotor performance…’. -
Rethinking the Evolution of the Human Foot: Insights from Experimental Research Nicholas B
© 2018. Published by The Company of Biologists Ltd | Journal of Experimental Biology (2018) 221, jeb174425. doi:10.1242/jeb.174425 REVIEW Rethinking the evolution of the human foot: insights from experimental research Nicholas B. Holowka* and Daniel E. Lieberman* ABSTRACT presumably owing to their lack of arches and mobile midfoot joints Adaptive explanations for modern human foot anatomy have long for enhanced prehensility in arboreal locomotion (see Glossary; fascinated evolutionary biologists because of the dramatic differences Fig. 1B) (DeSilva, 2010; Elftman and Manter, 1935a). Other studies between our feet and those of our closest living relatives, the great have documented how great apes use their long toes, opposable apes. Morphological features, including hallucal opposability, toe halluces and mobile ankles for grasping arboreal supports (DeSilva, length and the longitudinal arch, have traditionally been used to 2009; Holowka et al., 2017a; Morton, 1924). These observations dichotomize human and great ape feet as being adapted for bipedal underlie what has become a consensus model of human foot walking and arboreal locomotion, respectively. However, recent evolution: that selection for bipedal walking came at the expense of biomechanical models of human foot function and experimental arboreal locomotor capabilities, resulting in a dichotomy between investigations of great ape locomotion have undermined this simple human and great ape foot anatomy and function. According to this dichotomy. Here, we review this research, focusing on the way of thinking, anatomical features of the foot characteristic of biomechanics of foot strike, push-off and elastic energy storage in great apes are assumed to represent adaptations for arboreal the foot, and show that humans and great apes share some behavior, and those unique to humans are assumed to be related underappreciated, surprising similarities in foot function, such as to bipedal walking. -
Morphology, Phylogeny, and Evolution of Diadectidae (Cotylosauria: Diadectomorpha)
Morphology, Phylogeny, and Evolution of Diadectidae (Cotylosauria: Diadectomorpha) by Richard Kissel A thesis submitted in conformity with the requirements for the degree of doctor of philosophy Graduate Department of Ecology & Evolutionary Biology University of Toronto © Copyright by Richard Kissel 2010 Morphology, Phylogeny, and Evolution of Diadectidae (Cotylosauria: Diadectomorpha) Richard Kissel Doctor of Philosophy Graduate Department of Ecology & Evolutionary Biology University of Toronto 2010 Abstract Based on dental, cranial, and postcranial anatomy, members of the Permo-Carboniferous clade Diadectidae are generally regarded as the earliest tetrapods capable of processing high-fiber plant material; presented here is a review of diadectid morphology, phylogeny, taxonomy, and paleozoogeography. Phylogenetic analyses support the monophyly of Diadectidae within Diadectomorpha, the sister-group to Amniota, with Limnoscelis as the sister-taxon to Tseajaia + Diadectidae. Analysis of diadectid interrelationships of all known taxa for which adequate specimens and information are known—the first of its kind conducted—positions Ambedus pusillus as the sister-taxon to all other forms, with Diadectes sanmiguelensis, Orobates pabsti, Desmatodon hesperis, Diadectes absitus, and (Diadectes sideropelicus + Diadectes tenuitectes + Diasparactus zenos) representing progressively more derived taxa in a series of nested clades. In light of these results, it is recommended herein that the species Diadectes sanmiguelensis be referred to the new genus -
Morphology and Evolutionary Significance of the Atlas−Axis Complex in Varanopid Synapsids
Morphology and evolutionary significance of the atlas−axis complex in varanopid synapsids NICOLÁS E. CAMPIONE and ROBERT R. REISZ Campione, N.E. and Reisz, R.R. 2011. Morphology and evolutionary significance of the atlas−axis complex in varanopid synapsids. Acta Palaeontologica Polonica 56 (4): 739–748. The atlas−axis complex has been described in few Palaeozoic taxa, with little effort being placed on examining variation of this structure within a small clade. Most varanopids, members of a clade of gracile synapsid predators, have well pre− served atlas−axes permitting detailed descriptions and examination of morphological variation. This study indicates that the size of the transverse processes on the axis and the shape of the axial neural spine vary among members of this clade. In particular, the small mycterosaurine varanopids possess small transverse processes that point posteroventrally, and the axial spine is dorsoventrally short, with a flattened dorsal margin in lateral view. The larger varanodontine varanopids have large transverse processes with a broad base, and a much taller axial spine with a rounded dorsal margin in lateral view. Based on outgroup comparisons, the morphology exhibited by the transverse processes is interpreted as derived in varanodontines, whereas the morphology of the axial spine is derived in mycterosaurines. The axial spine anatomy of Middle Permian South African varanopids is reviewed and our interpretation is consistent with the hypothesis that at least two varanopid taxa are present in South Africa, a region overwhelmingly dominated by therapsid synapsids and parareptiles. Key words: Synapsida, Varanopidae, Mycterosaurinae, Varanodontinae, atlas−axis complex, axial skeleton, Middle Permian, South Africa. -
Center of Pressure Trajectory During Gait: a Comparison of Four Foot Positions
Gait & Posture 40 (2014) 719–722 Contents lists available at ScienceDirect Gait & Posture journal homepage: www.elsevier.com/locate/gaitpost Short Communication Center of pressure trajectory during gait: A comparison of four foot positions Vipul Lugade, Kenton Kaufman * Motion Analysis Laboratory, Division of Orthopedic Research, Mayo Clinic, Rochester, MN 55905, USA ARTICLE INFO ABSTRACT Article history: Knowledge of the center of pressure (COP) trajectory during stance can elucidate possible foot pathology, Received 16 May 2013 provide comparative effectiveness of foot orthotics, and allow for appropriate calculation of balance Received in revised form 20 May 2014 control and joint kinetics during gait. Therefore, the goal of this study was to investigate the COP Accepted 1 July 2014 movement when walking at self-selected speeds with plantigrade, equinus, inverted, and everted foot positions. A total of 13 healthy subjects were asked to walk barefoot across an 8-m walkway with Keywords: embedded force plates. The COP was computed for each stance limb using the ground reaction forces and Center of pressure moments collected from three force plates. Results demonstrated that the COP excursion was 83% of the Gait analysis foot length and 27% of the foot width in the anterior–posterior and medial lateral directions for Foot pathology Regression plantigrade walking, respectively. Regression equations explained 94% and 44% of the anterior–posterior and medial–lateral COP variability during plantigrade walking, respectively. While the range of motion and COP velocity were similar for inverted and everted walking, the COP remained on the lateral and medial aspects of the foot for these two walking conditions, respectively. -
Normal and Abnormal Gaits in Dogs
Pagina 1 di 12 Normal And Abnormal Gait Chapter 91 David M. Nunamaker, Peter D. Blauner z Methods of Gait Analysis z The Normal Gaits of the Dog z Effects of Conformation on Locomotion z Clinical Examination of the Locomotor System z Neurologic Conditions Associated With Abnormal Gait z Gait Abnormalities Associated With Joint Problems z References Methods of Gait Analysis Normal locomotion of the dog involves proper functioning of every organ system in the body, up to 99% of the skeletal muscles, and most of the bony structures.(1-75) Coordination of these functioning parts represents the poorly understood phenomenon referred to as gait. The veterinary literature is interspersed with only a few reports addressing primarily this system. Although gait relates closely to orthopaedics, it is often not included in orthopaedic training programs or orthopaedic textbooks. The current problem of gait analysis in humans and dogs is the inability of the study of gait to relate significantly to clinical situations. Hundreds of papers are included in the literature describing gait in humans, but up to this point there has been little success in organizing the reams of data into a useful diagnostic or therapeutic regime. Studies on human and animal locomotion commonly involve the measurement and analysis of the following: Temporal characteristics Electromyographic signals Kinematics of limb segments Kinetics of the foot-floor and joint resultants The analyses of the latter two types of measurements require the collection and reduction of voluminous amounts of data, but the lack of a rapid method of processing this data in real time has precluded the use of gait analysis as a routine clinical tool, particularly in animals. -
SUPPLEMENTARY INFORMATION: Tables, Figures and References
Samuels et al. Evolution of the patellar sesamoid bone in mammals SUPPLEMENTARY INFORMATION: Tables, Figures and References Supplementary Table S1: Mammals$ Higher taxa Genus sp. Estimated. age of Patellar Comments# (partial) specimen, location state 0/1/2 (absent/ ‘patelloid’/ present) Sinoconodonta Sinoconodon Jurassic 0 Patellar groove absent, suggests no rigneyi (Kielan- patella Jaworowska, Cifelli & Luo, Sinoconodon is included on our 2004) phylogeny within tritylodontids. Morganucodonta Megazostrodon Late Triassic, southern 0 rudnerae (Jenkins Africa & Parrington, 1976) Morganucodonta Eozostrodon sp. Late Triassic, Wales 0 Asymmetric patellar groove, (Jenkins et al., specimens disarticulated so it is hard 1976) to assess the patella but appears absent Docodonta Castorocauda 164 Mya, mid-Jurassic, 0 Semi-aquatic adaptations lutrasimilis (Ji, China Luo, Yuan et al., 2006) Docodonta Agilodocodon 164 Mya, mid-Jurassic, 0 scansorius China (Meng, Ji, Zhang et al., 2015) Docodonta Docofossor 160 Mya 0 brachydactylus (Luo, Meng, Ji et al., 2015) Docodonta Haldanodon 150-155 Mya, Late 0 Shallow patellar groove exspectatus Jurassic, Portugal (Martin, 2005b) Australosphenida Asfaltomylos Mid-Jurassic, South ? Postcranial material absent patagonicus America (Martin, 2005a) Australosphenida Ornithorhynchus Extant 2 Platypus, genome sequenced Monotremata anatinus (Warren, Hillier, Marshall Graves et (Herzmark, 1938; al., 2008) Rowe, 1988) Samuels et al. Australosphenida Tachyglossus + Extant 2 Echidnas Monotremata Zaglossus spp. (Herzmark, 1938; Rowe, 1988) Mammaliaformes Fruitafossor 150 Mya, Late Jurassic, 0 Phylogenetic status uncertain indet. windscheffeli (Luo Colorado & Wible, 2005) Mammaliaformes Volaticotherium Late Jurassic/Early ? Hindlimb material incomplete indet. antiquus (Meng, Cretaceous Hu, Wang et al., 2006) Eutriconodonta Jeholodens 120-125 Mya, Early 0 Poorly developed patellar groove jenkinsi (Ji, Luo Cretaceous, China & Ji, 1999) Eutriconodonta Gobiconodon spp. -
The Influence of Foot Posture on the Cost of Transport in Humans
790 The Journal of Experimental Biology 213, 790-797 © 2010. Published by The Company of Biologists Ltd doi:10.1242/jeb.038984 The influence of foot posture on the cost of transport in humans C. B. Cunningham1, N. Schilling2, C. Anders3 and D. R. Carrier1,* 1Department of Biology, University of Utah, 257S 1400E, Salt Lake City, UT, 84112, USA, 2Friedrich-Schiller-Universität, Institut für Spezielle Zoologie und Evolutionsbiologie mit Phyletischem Museum, Erbertstrasse 1, 07743 Jena, Germany and 3Universitätsklinikum Jena, Klinik für Unfall-, Hand- und Wiederherstellungschirurgie, FB Motorik, Pathophysiologie und Biomechanik, Erfurter Straße 35, 07743 Jena, Germany *Author for correspondence ([email protected]) Accepted 29 November 2009 SUMMARY Although humans appear to be specialized for endurance running, the plantigrade posture of our feet, in which the heel contacts the substrate at the beginning of a step, seems incompatible with economical running. In this study, we tested the hypothesis that plantigrade foot posture reduces the energetic cost of transport (COT) during walking in humans. When human subjects walked with their heels slightly elevated in a ‘low-digitigrade’ posture, COT increased by 53% above that of normal plantigrade walking. By contrast, there was no difference in COT when subjects ran with digitigrade versus plantigrade foot posture. Stride frequency increased and stride length decreased when subjects switched to digitigrade walking; however, this change did not influence the COT. Additionally, we found that possible reductions in postural stability appear not to have caused the elevated cost of digitigrade walking. Digitigrade walking, however, did (1) increase the external mechanical work performed by the limbs; (2) reduce the pendular exchange of kinetic and potential energy of the center of mass; (3) increase the average ground reaction force moment at the ankle joint; and (4) increase the recruitment of major extensor muscles of the ankle, knee, hip and back. -
Spinal Motor Outputs During Step-To-Step Transitions of Diverse Human Gaits
ORIGINAL RESEARCH ARTICLE published: 15 May 2014 HUMAN NEUROSCIENCE doi: 10.3389/fnhum.2014.00305 Spinal motor outputs during step-to-step transitions of diverse human gaits Valentina La Scaleia 1,2,3, Yuri P.Ivanenko 3, Karl E. Zelik 3 and Francesco Lacquaniti 1,2,3* 1 Department of Systems Medicine, University of Rome Tor Vergata, Rome, Italy 2 Centre of Space Bio-medicine, University of Rome Tor Vergata, Rome, Italy 3 Laboratory of Neuromotor Physiology, Santa Lucia Foundation, Rome, Italy Edited by: Aspects of human motor control can be inferred from the coordination of muscles Leonardo Gizzi, Pain Clinic Center during movement. For instance, by combining multimuscle electromyographic (EMG) for Anesthesiology, Emergency and recordings with human neuroanatomy, it is possible to estimate alpha-motoneuron (MN) Intensive Care Medicine University Hospital Göttingen, Germany pool activations along the spinal cord. It has previously been shown that the spinal motor Reviewed by: output fluctuates with the body’s center-of-mass motion, with bursts of activity around Massimo Sartori, Medical University foot-strike and foot lift-off during walking. However, it is not known whether these MN Göttingen, Germany bursts are generalizable to other ambulation tasks, nor is it clear if the spatial locus of Anderson Souza Oliveira, Aalborg the activity (along the rostrocaudal axis of the spinal cord) is fixed or variable. Here we University, Denmark sought to address these questions by investigating the spatiotemporal characteristics of *Correspondence: Francesco Lacquaniti, Department the spinal motor output during various tasks: walking forward, backward, tiptoe and uphill. of Systems Medicine, University of We reconstructed spinal maps from 26 leg muscle EMGs, including some intrinsic foot Rome Tor Vergata, IRCCS Santa muscles. -
FNR-417-W Animal Diversity and Tracking
FNR-417-W UNIT 1 Animal Diversity and Tracking Animal tracks are useful to reveal the diversity of organisms within different environments. Overview ....................................................2 Teachers’ Notes ........................................ 4 Lesson 1: Animal Tracks .......................... 5 Lesson 2: Scent Stations ......................... 6 Activity: Scent Station Data Sheet ........8 Lesson 3: Indoor Track Casting ............. 9 Lesson 4: Outdoor Track Casting ........ 10 Lesson 5: Animal Tracking .....................11 Activity: Animal Tracking Data Sheet ................................................12 Activity: Habitat Sketch .........................13 AUTHORS Rod N. Williams, Jarred Brooke, Robert N. Chapman, and Rebecca Busse, Department of Forestry and Natural Resources, Purdue University, West Lafayette, Indiana Animal Diversity and Tracking OVERVIEW LESSON PLAN ESTIMATED TIME Lesson 3 Three 30–90 minute lessons Next Generation Science Standards K-2-ETS1-2 3-LS4-3 4-LS1-1 VOCABULARY English/Language Arts Unguligrade • Habitat • RI.K.1 RI.1.1 RI.2.2 RI.3.7 SL.4.2 • Generalist • Stride RI.K.2 RI.1.2 RI.2.4 SL.3.1 RI.5.1 • Specialist • Straddle RI.K.3 RI.1.3 RI.2.7 SL.3.2 RI.5.2 • Plantigrade • Gait RI.K.4 RI.1.4 SL.2.1 RI.4.1 RI.5.4 W.K.2 RI.1.7 SL.2.2 RI.4.2 SL.5.1 • Digitigrade • Track W.K.3 SL.1.1 RI.3.1 RI.4.3 SL.5.2 UNIT OBJECTIVES SL.K.1 SL.1.2 RI.3.2 RI.4.4 SL.K.2 RI.2.1 RI.3.4 SL.4.1 Students will be able to: • Identify wildlife species using tracks Math • Recognize that animal diversity can -
The Vertebrate Fauna of the New Mexico Permian Alfred S
New Mexico Geological Society Downloaded from: http://nmgs.nmt.edu/publications/guidebooks/11 The vertebrate fauna of the New Mexico Permian Alfred S. Romer, 1960, pp. 48-54 in: Rio Chama Country, Beaumont, E. C.; Read, C. B.; [eds.], New Mexico Geological Society 11th Annual Fall Field Conference Guidebook, 129 p. This is one of many related papers that were included in the 1960 NMGS Fall Field Conference Guidebook. Annual NMGS Fall Field Conference Guidebooks Every fall since 1950, the New Mexico Geological Society (NMGS) has held an annual Fall Field Conference that explores some region of New Mexico (or surrounding states). Always well attended, these conferences provide a guidebook to participants. Besides detailed road logs, the guidebooks contain many well written, edited, and peer-reviewed geoscience papers. These books have set the national standard for geologic guidebooks and are an essential geologic reference for anyone working in or around New Mexico. Free Downloads NMGS has decided to make peer-reviewed papers from our Fall Field Conference guidebooks available for free download. Non-members will have access to guidebook papers two years after publication. Members have access to all papers. This is in keeping with our mission of promoting interest, research, and cooperation regarding geology in New Mexico. However, guidebook sales represent a significant proportion of our operating budget. Therefore, only research papers are available for download. Road logs, mini-papers, maps, stratigraphic charts, and other selected content are available only in the printed guidebooks. Copyright Information Publications of the New Mexico Geological Society, printed and electronic, are protected by the copyright laws of the United States.