The Vertebrate Fauna of the New Mexico Permian Alfred S
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Distributions of Extinction Times from Fossil Ages and Tree Topologies: the Example of Some Mid-Permian Synapsid Extinctions Gilles Didier, Michel Laurin
Distributions of extinction times from fossil ages and tree topologies: the example of some mid-Permian synapsid extinctions Gilles Didier, Michel Laurin To cite this version: Gilles Didier, Michel Laurin. Distributions of extinction times from fossil ages and tree topologies: the example of some mid-Permian synapsid extinctions. 2021. hal-03258099v2 HAL Id: hal-03258099 https://hal.archives-ouvertes.fr/hal-03258099v2 Preprint submitted on 20 Sep 2021 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. Distributions of extinction times from fossil ages and tree topologies: the example of some mid-Permian synapsid extinctions Gilles Didier1 and Michel Laurin2 1 IMAG, Univ Montpellier, CNRS, Montpellier, France 2 CR2P (\Centre de Pal´eontologie { Paris"; UMR 7207), CNRS/MNHN/SU, Mus´eumNational d'Histoire Naturelle, Paris, France September 16, 2021 Abstract Given a phylogenetic tree that includes only extinct, or a mix of extinct and extant taxa, where at least some fossil data are available, we present a method to compute the distribution of the extinction time of a given set of taxa under the Fossilized-Birth-Death model. Our approach differs from the previous ones in that it takes into account (i) the possibility that the taxa or the clade considered may diversify before going extinct and (ii) the whole phylogenetic tree to estimate extinction times, whilst previous methods do not consider the diversification process and deal with each branch independently. -
Amphibians 1) Transition to Land A) Life on Terrestrial Earth Is a Major
Amphibians 1) Transition to land a) Life on terrestrial earth is a major theme for all non-fish vertebrates also known as Tetrapoda b) Of Tetrapoda there are two major groups Amphibians and amniotes c) The movement form water to land is one of the most complicated and dramatic events of the evolution of animals i) Land is physically hazardous for an animal that evolved in water, is made mostly of water, and all cellular activities occur in water. ii) Plants, snails, and many arthropods made the transition before vertebrates, which provided a plentiful food source. iii) With the transition to land, vertebrates had to adapt every organ system. d) Oxygen on land i) Atmospheric air contains 20 times more oxygen than water and diffuses more rapidly through air than water. ii) By the Devonian period (400+ million years ago) fish had diversified greatly. Some of these adaptations became useful for a terrestrial life (1) Fish had evolved an air sack within their body called a swim bladder. This would allow a space for gas exchange between an organism and air (a) These early fishes were most likely freshwater. Freshwater systems are more likely to evaporate or deoxygenate compares to salt water habitats. So having a vascularized swim bladder or lung would be beneficial. (b) To this day scientist still debate heavily on whether the swim bladder evolved for buoyancy control or lung first. (2) Fish also had evolved external nares for chemoreception. In a terrestrial environment these nares can be used to draw in air to the swim bladder/lung (3) Both of these structures show great examples of evolution utilizing existing structures to turn into something new and more adapted e) However both of the characteristics are shared among fishes and tetrapods, the big shift came in the bone structure of the limbs. -
Reptile Family Tree
Reptile Family Tree - Peters 2015 Distribution of Scales, Scutes, Hair and Feathers Fish scales 100 Ichthyostega Eldeceeon 1990.7.1 Pederpes 91 Eldeceeon holotype Gephyrostegus watsoni Eryops 67 Solenodonsaurus 87 Proterogyrinus 85 100 Chroniosaurus Eoherpeton 94 72 Chroniosaurus PIN3585/124 98 Seymouria Chroniosuchus Kotlassia 58 94 Westlothiana Casineria Utegenia 84 Brouffia 95 78 Amphibamus 71 93 77 Coelostegus Cacops Paleothyris Adelospondylus 91 78 82 99 Hylonomus 100 Brachydectes Protorothyris MCZ1532 Eocaecilia 95 91 Protorothyris CM 8617 77 95 Doleserpeton 98 Gerobatrachus Protorothyris MCZ 2149 Rana 86 52 Microbrachis 92 Elliotsmithia Pantylus 93 Apsisaurus 83 92 Anthracodromeus 84 85 Aerosaurus 95 85 Utaherpeton 82 Varanodon 95 Tuditanus 91 98 61 90 Eoserpeton Varanops Diplocaulus Varanosaurus FMNH PR 1760 88 100 Sauropleura Varanosaurus BSPHM 1901 XV20 78 Ptyonius 98 89 Archaeothyris Scincosaurus 77 84 Ophiacodon 95 Micraroter 79 98 Batropetes Rhynchonkos Cutleria 59 Nikkasaurus 95 54 Biarmosuchus Silvanerpeton 72 Titanophoneus Gephyrostegeus bohemicus 96 Procynosuchus 68 100 Megazostrodon Mammal 88 Homo sapiens 100 66 Stenocybus hair 91 94 IVPP V18117 69 Galechirus 69 97 62 Suminia Niaftasuchus 65 Microurania 98 Urumqia 91 Bruktererpeton 65 IVPP V 18120 85 Venjukovia 98 100 Thuringothyris MNG 7729 Thuringothyris MNG 10183 100 Eodicynodon Dicynodon 91 Cephalerpeton 54 Reiszorhinus Haptodus 62 Concordia KUVP 8702a 95 59 Ianthasaurus 87 87 Concordia KUVP 96/95 85 Edaphosaurus Romeria primus 87 Glaucosaurus Romeria texana Secodontosaurus -
Reptiles A. Cladistics 1. Many Groups of Organisms
Reptiles A. Cladistics 1. Many groups of organisms are “polyphyletic” a. This means that the group combines 2 or more lineages - example=fish 2. Cladistics follows only pure lineages going back in time - example Osteichthys B. Reptile Classifiecation - looks like a polyphyletic group 1. Dry skin - no loss of water through skin like amphibians 2. Aminotic egg - an egg that can survive on dry land - in contrast with the amphibian egg C. Mammals and Birds are derived from different lineages of reptiles (We will see below) D. Stem Reptiles 1. Different lineages based on the temporal region of their skulls - number of holes (or bars) a. These holes are necessary to accommodate large jaw muscles b. Anapsid Skull - no holes in temporal - jaws can move fast, but with little force 1. Muscles that move the jaw are small 2. There is no good paleotological evidence for the transition between amphibians and reptiles - no fossil intermediates a. Fossil amphibians have lots of dermal bones in skull b. Amphibians have no temporal openings in skull 1. (Aside) both fossil amphibians and primitive reptiles have a parietal “eye” that senses light and dark (“third” eye in middle of head) c. Reptile skull is higher than amphibian to accomodate larger jaw muscles d. Of the modern reptiles only turtles are anapsids 2. Diapsid Skull - has holes in the temporal region a. Diapsid reptiles gave rise to lizards and snakes - they have a diapsid skull 1. Also Tuatara, crocodiles, dinosaurs and pterydactyls Reptiles b. One group of diapsids also had a pre-orbital hole in the skull in front of eye - this hole is still preserved in the birds - this anatomy suggests strongly that the birds are derived from the diapsid reptiles 3. -
Morphology, Phylogeny, and Evolution of Diadectidae (Cotylosauria: Diadectomorpha)
Morphology, Phylogeny, and Evolution of Diadectidae (Cotylosauria: Diadectomorpha) by Richard Kissel A thesis submitted in conformity with the requirements for the degree of doctor of philosophy Graduate Department of Ecology & Evolutionary Biology University of Toronto © Copyright by Richard Kissel 2010 Morphology, Phylogeny, and Evolution of Diadectidae (Cotylosauria: Diadectomorpha) Richard Kissel Doctor of Philosophy Graduate Department of Ecology & Evolutionary Biology University of Toronto 2010 Abstract Based on dental, cranial, and postcranial anatomy, members of the Permo-Carboniferous clade Diadectidae are generally regarded as the earliest tetrapods capable of processing high-fiber plant material; presented here is a review of diadectid morphology, phylogeny, taxonomy, and paleozoogeography. Phylogenetic analyses support the monophyly of Diadectidae within Diadectomorpha, the sister-group to Amniota, with Limnoscelis as the sister-taxon to Tseajaia + Diadectidae. Analysis of diadectid interrelationships of all known taxa for which adequate specimens and information are known—the first of its kind conducted—positions Ambedus pusillus as the sister-taxon to all other forms, with Diadectes sanmiguelensis, Orobates pabsti, Desmatodon hesperis, Diadectes absitus, and (Diadectes sideropelicus + Diadectes tenuitectes + Diasparactus zenos) representing progressively more derived taxa in a series of nested clades. In light of these results, it is recommended herein that the species Diadectes sanmiguelensis be referred to the new genus -
Distributions of Extinction Times from Fossil Ages and Tree Topologies: the Example of Some Mid-Permian Synapsid Extinctions
bioRxiv preprint doi: https://doi.org/10.1101/2021.06.11.448028; this version posted June 11, 2021. The copyright holder for this preprint (which was not certified by peer review) is the author/funder. All rights reserved. No reuse allowed without permission. Distributions of extinction times from fossil ages and tree topologies: the example of some mid-Permian synapsid extinctions Gilles Didier1 and Michel Laurin2 1IMAG, Univ Montpellier, CNRS, Montpellier, France 2CR2P (“Centre de Recherches sur la Paléobiodiversité et les Paléoenvironnements”; UMR 7207), CNRS/MNHN/UPMC, Sorbonne Université, Muséum National d’Histoire Naturelle, Paris, France June 11, 2021 Abstract Given a phylogenetic tree of extinct and extant taxa with fossils where the only temporal infor- mation stands in the fossil ages, we devise a method to compute the distribution of the extinction time of a given set of taxa under the Fossilized-Birth-Death model. Our approach differs from the previous ones in that it takes into account the possibility that the taxa or the clade considered may diversify before going extinct, whilst previous methods just rely on the fossil recovery rate to estimate confidence intervals. We assess and compare our new approach with a standard previous one using simulated data. Results show that our method provides more accurate confidence intervals. This new approach is applied to the study of the extinction time of three Permo-Carboniferous synapsid taxa (Ophiacodontidae, Edaphosauridae, and Sphenacodontidae) that are thought to have disappeared toward the end of the Cisuralian, or possibly shortly thereafter. The timing of extinctions of these three taxa and of their component lineages supports the idea that a biological crisis occurred in the late Kungurian/early Roadian. -
The Global Distribution of Tetrapods Reveals a Need for Targeted Reptile
1 The global distribution of tetrapods reveals a need for targeted reptile 2 conservation 3 4 Uri Roll#1,2, Anat Feldman#3, Maria Novosolov#3, Allen Allison4, Aaron M. Bauer5, Rodolphe 5 Bernard6, Monika Böhm7, Fernando Castro-Herrera8, Laurent Chirio9, Ben Collen10, Guarino R. 6 Colli11, Lital Dabool12 Indraneil Das13, Tiffany M. Doan14, Lee L. Grismer15, Marinus 7 Hoogmoed16, Yuval Itescu3, Fred Kraus17, Matthew LeBreton18, Amir Lewin3, Marcio Martins19, 8 Erez Maza3, Danny Meirte20, Zoltán T. Nagy21, Cristiano de C. Nogueira19, Olivier S.G. 9 Pauwels22, Daniel Pincheira-Donoso23, Gary Powney24, Roberto Sindaco25, Oliver Tallowin3, 10 Omar Torres-Carvajal26, Jean-François Trape27, Enav Vidan3, Peter Uetz28, Philipp Wagner5,29, 11 Yuezhao Wang30, C David L Orme6, Richard Grenyer✝1 and Shai Meiri✝*3 12 13 # Contributed equally to the paper 14 ✝ Contributed equally to the paper 15 * Corresponding author 16 17 Affiliations: 18 1 School of Geography and the Environment, University of Oxford, Oxford, OX13QY, UK. 19 2 Mitrani Department of Desert Ecology, The Jacob Blaustein Institutes for Desert Research, 20 Ben-Gurion University, Midreshet Ben-Gurion 8499000, Israel. (Current address) 21 3 Department of Zoology, Tel-Aviv University, Tel-Aviv 6997801, Israel. 22 4 Hawaii Biological Survey, 4 Bishop Museum, Honolulu, HI 96817, USA. 23 5 Department of Biology, Villanova University, Villanova, PA 19085, USA. 24 6 Department of Life Sciences, Imperial College London, Silwood Park Campus Silwood Park, 25 Ascot, Berkshire, SL5 7PY, UK 26 7 Institute of Zoology, Zoological Society of London, London NW1 4RY, UK. 27 8 School of Basic Sciences, Physiology Sciences Department, Universidad del Valle, Colombia. -
Early Tetrapod Relationships Revisited
Biol. Rev. (2003), 78, pp. 251–345. f Cambridge Philosophical Society 251 DOI: 10.1017/S1464793102006103 Printed in the United Kingdom Early tetrapod relationships revisited MARCELLO RUTA1*, MICHAEL I. COATES1 and DONALD L. J. QUICKE2 1 The Department of Organismal Biology and Anatomy, The University of Chicago, 1027 East 57th Street, Chicago, IL 60637-1508, USA ([email protected]; [email protected]) 2 Department of Biology, Imperial College at Silwood Park, Ascot, Berkshire SL57PY, UK and Department of Entomology, The Natural History Museum, Cromwell Road, London SW75BD, UK ([email protected]) (Received 29 November 2001; revised 28 August 2002; accepted 2 September 2002) ABSTRACT In an attempt to investigate differences between the most widely discussed hypotheses of early tetrapod relation- ships, we assembled a new data matrix including 90 taxa coded for 319 cranial and postcranial characters. We have incorporated, where possible, original observations of numerous taxa spread throughout the major tetrapod clades. A stem-based (total-group) definition of Tetrapoda is preferred over apomorphy- and node-based (crown-group) definitions. This definition is operational, since it is based on a formal character analysis. A PAUP* search using a recently implemented version of the parsimony ratchet method yields 64 shortest trees. Differ- ences between these trees concern: (1) the internal relationships of aı¨stopods, the three selected species of which form a trichotomy; (2) the internal relationships of embolomeres, with Archeria -
Morphology and Evolutionary Significance of the Atlas−Axis Complex in Varanopid Synapsids
Morphology and evolutionary significance of the atlas−axis complex in varanopid synapsids NICOLÁS E. CAMPIONE and ROBERT R. REISZ Campione, N.E. and Reisz, R.R. 2011. Morphology and evolutionary significance of the atlas−axis complex in varanopid synapsids. Acta Palaeontologica Polonica 56 (4): 739–748. The atlas−axis complex has been described in few Palaeozoic taxa, with little effort being placed on examining variation of this structure within a small clade. Most varanopids, members of a clade of gracile synapsid predators, have well pre− served atlas−axes permitting detailed descriptions and examination of morphological variation. This study indicates that the size of the transverse processes on the axis and the shape of the axial neural spine vary among members of this clade. In particular, the small mycterosaurine varanopids possess small transverse processes that point posteroventrally, and the axial spine is dorsoventrally short, with a flattened dorsal margin in lateral view. The larger varanodontine varanopids have large transverse processes with a broad base, and a much taller axial spine with a rounded dorsal margin in lateral view. Based on outgroup comparisons, the morphology exhibited by the transverse processes is interpreted as derived in varanodontines, whereas the morphology of the axial spine is derived in mycterosaurines. The axial spine anatomy of Middle Permian South African varanopids is reviewed and our interpretation is consistent with the hypothesis that at least two varanopid taxa are present in South Africa, a region overwhelmingly dominated by therapsid synapsids and parareptiles. Key words: Synapsida, Varanopidae, Mycterosaurinae, Varanodontinae, atlas−axis complex, axial skeleton, Middle Permian, South Africa. -
Marine Reptiles Arne R
Virginia Commonwealth University VCU Scholars Compass Study of Biological Complexity Publications Center for the Study of Biological Complexity 2011 Marine Reptiles Arne R. Rasmessen The Royal Danish Academy of Fine Arts John D. Murphy Field Museum of Natural History Medy Ompi Sam Ratulangi University J. Whitfield iG bbons University of Georgia Peter Uetz Virginia Commonwealth University, [email protected] Follow this and additional works at: http://scholarscompass.vcu.edu/csbc_pubs Part of the Life Sciences Commons Copyright: © 2011 Rasmussen et al. This is an open-access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. Downloaded from http://scholarscompass.vcu.edu/csbc_pubs/20 This Article is brought to you for free and open access by the Center for the Study of Biological Complexity at VCU Scholars Compass. It has been accepted for inclusion in Study of Biological Complexity Publications by an authorized administrator of VCU Scholars Compass. For more information, please contact [email protected]. Review Marine Reptiles Arne Redsted Rasmussen1, John C. Murphy2, Medy Ompi3, J. Whitfield Gibbons4, Peter Uetz5* 1 School of Conservation, The Royal Danish Academy of Fine Arts, Copenhagen, Denmark, 2 Division of Amphibians and Reptiles, Field Museum of Natural History, Chicago, Illinois, United States of America, 3 Marine Biology Laboratory, Faculty of Fisheries and Marine Sciences, Sam Ratulangi University, Manado, North Sulawesi, Indonesia, 4 Savannah River Ecology Lab, University of Georgia, Aiken, South Carolina, United States of America, 5 Center for the Study of Biological Complexity, Virginia Commonwealth University, Richmond, Virginia, United States of America Of the more than 12,000 species and subspecies of extant Caribbean, although some species occasionally travel as far north reptiles, about 100 have re-entered the ocean. -
C.-Shoulder Joint
59.14.98,1:81 Article X.-THE LOCOMOTOR APPARATUS OF CERTAIN PRIMITIVE AND MAMMAL-LIKE REPTILES' BY ALFRED S. ROMER PLATES XXVII TO XLVI TABLE OF CONTENTS PAGE INTRODIUCTION ......................................................... 519 FORE LIMB, MIUSCULATURE ............................................. 524 1.-Trapezius (and sterno-cleido-mastoideus) ........................ 524 2.-Axial muscles acting on the girdle ............................... 525 a.-Dorsal: levator scapul& (and atlanto-scapularis, atlanto- acromialis or omo-trachelius and rhomboideuA), serratus anterior .............................................. 525 b.-Ventral: sterno- (episterno) hyoideus and omo-hyoideus, sterno-costo-coracoidei (subclavius) ................... 526 3.-Latissimus dorsi and teres major ................................ 527 4.-Pectoralis ..................................................... 528 5.-Deltoid ....................................................... 529 6.-Subcoraco-scapularis and scap'ilo-humeralis posterior (subscapularis) 530 7.-Scapulo-humeralis anterior (teres minor), supracoracoideus (supra- and infraspinatus), and coraco-brachialis ..................... 532 8.-Biceps and brachialis .......................................... 536 9.-Triceps ............................... A 537 10.-Brachio- (humero-) radialis (supinator longus) and supinator ...... 538 11.-Forearm: extensors............................................. 539 12.-Forearm: flexors (includingpronators) ............................ 540 13.-Groupingof forelimb muscles -
Xenacanthus (Chondrichthyes: Xenacanthiformes) from North America
Acta Geologica Polonica, Vol. 49 (J 999), No.3, pp. 215-266 406 IU S UNES 0 I Dentitions of Late Palaeozoic Orthacanthus species and new species of ?Xenacanthus (Chondrichthyes: Xenacanthiformes) from North America GARY D. JOHNSON Department of Earth Sciences and Physics, University of South Dakota; 414 East Clark Street, Vermillion, SD 57069-2390, USA. E-mail: [email protected] ABSTRACT: JOHNSON, G.D. 1999. Dentitions of Late Palaeozoic Orthacanthus species and new species of ?Xenacanthus (Chondrichthyes: Xenacanthiformes) from North America. Acta Geologica Polonica, 49 (3),215-266. Warszawa. Orthacanthus lateral teeth have paired, variably divergent, smooth, usually carinated labio-lingually compressed principal cusps separated by a central foramen; one or more intermediate cusps; and an api cal button on the base isolated from the cusps. Several thousand isolated teeth from Texas Artinskian bulk samples are used to define the heterodont dentitions of O. texensis and O. platypternus. The O. tex ensis tooth base has a labio-Iingual width greater than the anteromedial-posterolateral length, the basal tubercle is restricted to the thick labial margin, the principal cusps are serrated to varying degrees, and the posterior cusp is larger. The O. platypternus tooth base is longer than wide, its basal tubercle extends to the center, the labial margin is thin, serrations are absent on the principal cusps, the anterior cusp is larger, and a single intermediate cusp is present. More than two hundred isolated teeth from Nebraska (Gzhelian) and Pennsylvania (Asselian) provide a preliminary description of the heterodont dentition of O. compress us . The principal cusps are similar to O.