Records or the Western Australian Musellm Supplement No. 61: 77-154 (2000).

Flora and vegetation of the southern Carnarvon Basin, Western Australia

G.J. Keighery, N. Gibson, M.N. Lyons and Allan H. Burbidge Department of Conservation and Land Management, PO Box 51, Wanneroo, Western Australia 6065, Australia

Abstract This paper reports the first detailed study of the vascular flora of 2 the southern Carnarvon Basin, an area of c. 75000 km • A total flora of 2133 taxa of vascular plants was listed for the area. There are eight major conservation reserves which have 1559 taxa present in them. Most of the 574 unreserved taxa are wetland taxa, taxa of tropical affinities or those only present on the Acacia shrublands of the central basin. Vegetation patterning at a regional scale showed the major floristic boundary in the south west of the study area, which in turn reflected the major climatic gradients of the area. The other major influence on vegetation patterning was soil type.

INTRODUCTION temperate - arid change-over zone (Gibson, Despite Shark Bay being the site of very early Burbidge, Keighery and Lyons, 2000). visitation and study by several European expeditions (Beard, 1990; Keighery, 1990; George, 1999) the area was until recently still poorly known METHODS botanically. Beard (1975, 1976a) prepared structural vegetation maps for the whole area at a 1: 1 000000 Study area scale and Payne et al. (1987) have undertaken land The study area covered by the flora survey system maps (rangeland mapping) for the whole extended from 23°30'S to 28°00'S and from Dirk area at a 1: 250000 scale. More localised areas have Hartog Island and Shark Bay in the west to 115°30'E been mapped and published for the Kalbarri-Shark (Figure 1). The southwest of the study area Bay area by Beard (1976b, 1976c) at a scale of encompasses the change over from the South West 1: 250000. Botanical Province to the Eremaean Province and Limited floristic surveys have been undertaken in hence is an area of major vegetation change. Almost the north of the study area around Lake MacLeod the entire study area falls within the Carnarvon and by Tyler (1988), Rocky Pool near Carnarvon by Irwin botanical districts of Beard (1990). These Kenneally (1978) and Cape Range by Keighery and districts largely correspond to two IBRA regions ­ Gibson (1993). Other localised studies have Geraldton Sandplain and Carnarvon (Thackway involved the Shark Bay World Heritage Area by and Cresswell, 1995). Trudgen and Keighery (1995), Bernier and Dorre Islands by Royce (1962), Dirk Hartog Island by Field sampling and analysis Burbidge and George (1978), Kennedy Range by Sixty-three primary sites were sampled across the Newbey and Newbey (1992) and Toolonga Nature study area, stratified to cover the geographic range Reserve by Burbidge et al. (1980). of the major surface types of the Basin. Sites were The present work is part of a multi-disciplinary selected such that three to five different surface­ study to sample the biodiversity of a 75000 km2 types could be sampled in close proximity to region of Western Australia (the southern facilitate fauna sampling (Figure 1). Sites 30 m x 30 Carnarvon Basin) and identify biophysical factors m were established at each site and all vascular related to the observed patterns of species plants occurring in the sites were recorded. Sites distribution (Burbidge et al., 2000). were visited twice, in August 1994 and August The aim of this paper is to describe the flora and 1995. Some additional recordings were made when vegetation of the major landforms sampled as the sites were visited at other times in conjunction biodiversity sites and to discuss the composition, with fauna sampling. Sites were classified biogeographic patterns and conservation status of according to similarities in species composition the vascular plants of the study area. Related papers (presence/ absence data) using the Czekanowski in this volume deal with the wetIand flora (Gibson, coefficient and 'unweighted pair-group mean Keighery and Lyons, 2000) and the flora of the average' fusion method (UPGMA, Sneath and 78 G.]. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge

0 1130 /' I.' 14 1150 ' j'J ~~'~k )j t)( ~._-- ~I' )/ 't:--< ..~ ('- 24<>- Lake (\ '-~ f6 CU A3 Mac~ 5 J

5 4 11r: 4 3 1 j.M&5 ~2 I I BO MR la Kennedy Range National Park

Bemier Island Camarvon Nature Reserve

25 0

Dorre Island Nature Reserve Shark Bay World Heritage Area Pimbee

5 4 3 2 1a"1 WO

25 50

Kilometres

Existing reserves

Timber Reserve

CALM Leasehold

la Study sites

World Heritage Area Boundlll)'

1140

Figure 1 Location of 63 primary sampling sites and major conservation reserves in the study area, Flora and vegetation 79

Soka!, 1973), while species were classified using the 5.0%), Papilionaceae (105, 4.9'?{J), Mimosaceae (95, 'two step' method of Austin and Belbin (1982) and 4.5%), Proteaceae (93, 4.4%), Goodeniaceae (71, the UPGMA fusion method. Only taxa identified to 3.3%), Cyperaceae (61, 2.9%), Orchidaceae (57, species level were included in the analysis (>95'J;, 2.7%), Amaranthaceae (50, 2.3°/.,), Myoporaceae (50, records). Annuals and geophytes were included in 2.3%) and Malvaceae (48, 2.3%). Together these 13 the analysis because all sites but one (NE2) were families comprise 51.7% of the total flora. considered to have had good rains prior to at least Of the remaining 109 families, only 12 contained one of the sampling periods. at least 20 species: Euphorbiaceae (36 taxa), A bulked soil sample (consisting of 24 regularly Brassicaceae (32), Solanaceae (32), Anthericaceae spaced subsamples across the site) of the top 10 cm (28), Zygophyllaceae (26), Apiaceae (22), was collected from each site, and soil mechanical Epacridaceae (22), Sterculiaceae (22), Boraginaceae analysis and macro nutrient analyses were (21), Chloanthaceae (21), Haemodoraceae (20) and undertaken on these samples [see McArthur (1991) Portulacaceae (20). These 12 families combined for details]. Eleven climatic parameters for each site contributed 14.2% of the total flora. were estimated from the climate modelling Several significant collections were made during program ANUCLIM (McMahon et al., 1995). Statist­ the survey, including the re-collection of Rumex ical relationships between site groups were tested crystallinus (the first record since 1877). Three using Kruskal - Wallis non-parametric analysis of apparently new taxa were first collected during the variance and Mann Whitney U-tests (Siege!, 1956). survey: Paracaleana lyonsii ms (Orchidaceae), In addition to the 63 primary sites, flora lists were Calandrinia sp. Coolcalalaya (GK and NG 698) available from 75 extra sites in the southern part of (Portulacaceae) and Eremophila glabra subsp. the study area (Gibson, Burbidge, Keighery and Zuytdorp (GK and NG 518) (Myoporaceae). Lyons, 2000) and from the wetland survey (Gibson, The broad composition of the flora reflects the Keighery and Lyons, 2000). Additional sampling major climatic influences affecting the study area. efforts in six mainland reserves were also There is a southern coastal winter rainfall zone and undertaken to compile comprehensive species lists an inland and northern arid zone. The whole study for the eight major reserves (Figure 1) in the study area is still dominated by winter rainfall (Wyrwoll, area. These lists also include previously published Courtney and Sandercock, 2000). Immediately north surveys and collections held in the Western of the study area, summer rainfall begins to Australian Herbarium (PERTH). The total known dominate. The majority of the study area is arid and vascular flora of the study area (Appendix 1) was forms part of the Eremaean Botanical Province compiled from the current and previous surveys, (Beard, 1975, 1976a). Thi5 pattern is also clearly seen and from collections held in PERTH. in the results for the vegetation analysis (see below). Nomenclature generally follows Green (1985) and This strong desert influence is clearly demon­ current usage at PERTH. Over 2200 voucher strated in the composition of the flora, with arid specimens have been lodged in PERTH. zone taxa dominating in six of the speciose families (Amaranthaceae, Asteraceae, Goodeniaceae, Malvaceae, Myoporaceae and Poaceae) and in RESULTS nearly all the smaller families (Euphorbiaceae, The results are given in two sections. The first Brassicaceae, Solanaceae, Zygophyllaceae, deals with the overall vascular flora recorded for Apiaceae, Sterculiaceae, Boraginaceae, the study area, its composition and conservation. It Chloanthaceae, and Portulacaceae). These desert provides a floristic context for the interpretation of influences are from predominantly temperate or the second part, which deals with quantitative winter rainfall semi-arid or arid areas, rather than a patterns in the floristic composition of the tropical desert flora. This can be illustrated using biodiversity sites and how this patterning relates to the Asteraceae, the largest family recorded from the the biophysical attributes of the study area. study area. The family has 492 native taxa present in Western Flora Australia, of which 181 were recorded from the study area. Species richness in the family is greatest Composition in the Avon-Wheatbelt and Coolgardie IBRA regions A total of 2133 taxa of vascular plants from 122 (areas of Mediterranean and arid-Mediterranean families were recorded from the study area climate) of Western Australia (Figure 2). The family, (Appendix 1). These comprised 9 ferns or fern allies, though spread widely through the arid zone 2 gymnosperms and 2122 flowering plants (of (providing the spectacular wildflower displays of the which 88 were naturalised aliens). Acacia shrublands) and tropics, is species rich in The largest families were the Myrtaceae (231 taxa, temperate Western Australia. 10.8% of the flora), Asteraceae (181 taxa, 8.5%), The Asteraceae of the study area are a reflection Poaceae (168 taxa, 7.9'Yo), Chenopodiaceae (106, of the above pattern, being comprised mainly of 80 G.]. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge

ericoid Myrtaceae, especially members of the genera Scholtzia, Baeckea and Pileanthus. There is also a considerable number of eucalypt and orchid taxa, especially on heavy soils in the Kalbarri area. The Haemodoraceae, Epacridaceae (Keighery, 1996) and the genus Acacia (Hopper and Maslin, 1978) are also species diverse on the Kalbarri sandplains.

Weeds In total, 88 species of naturalised vascular plant species (weeds) were recorded for the area. These weeds are nearly all annuals from the families Poaceae (41 taxa), Asteraceae (19), Brassicaceae (12) and Caryophyllaceae (10). This is a low number compared to other areas of Western Australia (Keighery, 1995), where some 1032 taxa are recorded as naturalised (ca. 9% of the total flora). But it is comparable to Cape Range, where 30 weeds were recorded by Keighery and Gibson (1993) and the Kimberley where 184 taxa have been recorded (Hussey et al., 1997). The largely intact nature of the native vegetation of the study area lowers the number of weeds. It is likely that further collecting around the townsites of Kalbarri and Carnarvon will increase the number of naturalised taxa, but it is considered that the major environmental weeds have been Figure 2 Species richness patterns of the native documented. Asteraceae of Western Australia. Certain weeds have already had a severe impact on parts of the study area. Foremost are riverine sites, especially along the Gascoyne, Wooramel annuals (52 taxa) that are widespread in the winter and Irwin Rivers where the fringing understorey rainfall areas of the semi-arid and arid zone. There vegetation is often dominated by Buffel Grass are also 20 taxa of the temperate zone that end (Cenchrus ciliaris), Mosman River Grass (Cenchrus their ranges in the Basin (not necessarily in the setigerus) and Castor Oil Plants (Ricinus communis). study area, e.g. Brachycome iberidifolia, Numerous herbaceous weeds such as Cape Weed Chthonocephalus pseudevax and Cratystylis (Arctotheca calendula) dominate the Murchison subspinescens) and 13 tropical taxa that are at their River foreshore in Kalbarri National Park, southern limits (e.g. Blumea tenella, Launaea probably due to past grazing. The other major sarmentosa, Pluchea rubellifiora and Streptoglossa areas of weed invasion are under the Acacia bubakii). There is also a component of 12 taxa that shrublands of Zuytdorp and Cooloomia National are either endemics or species centred on the Basin Park where Brassica tournefortii has replaced (e.g. Chthonocephalus oldfieldianus, Haptotrichion Parietaria debilis, after sheep, rabbit and now goat I colwillii, Rhodanthe cremea and Rhodanthe grazing pressure. There are similar, but more oppositifolia subsp. ornata). Many of these are localised problems in the Shark Bay World recently described or still unnamed. Heritage Area. The pattern shown in the Asteraceae is repeated Several other weeds are potentially serious. in the Poaceae (Keighery, 1984), Chenopodiaceae Patterson's Curse (Echium plantagineum) is (Wilson, 1984), Amaranthaceae and Goodeniaceae spreading along road verges through the southern (Carolin et al., 1992). The desert elements of the flora arid zone, including North West Coastal Highway, are largely of southern affinities rather than arid and has the potential to replace the mass displays tropical, as would be found further north. of everlastings in these areas. Alford et al. (2000) The other major influence is that the study area have reported Boxthorn (Lycium ferocissimum) on contains a portion of the northern heathlands of the islands in the Shark Bay World Heritage area. This South-West Botanical Province, a large section of species is already a major pest on other offshore desert to the north and east, and the interzone islands. Athel Pine (Tamarix aphylla), a serious weed between them (Beard, 1990). The northern heath­ of the Finke River system in the Northern Territory, lands are diverse in members of the Proteaceae, is spreading along the Gascoyne River at especially Grevillea (Gibson et al. 1997), and the Camarvon. Flora and vegetation 81

Site Groups bbl 002 1-- nal I 1 bb3 1- cul I pel I I I bb2 I bb4 1- I I IIld3 I-- I ho4 I I 2.. hoS I 1- I 1Ild1 1 I 1Ild2 I 1- 1 001 I I 003 1- I I wo3 II I 2b woS 1- I I I wo4 1--1--1 I 1Ild4 I I IIldS I-- I 1 wo1 I 1 I 2c wo2 I 1- I I na2 II 1 pt!4 - 1-1--1- I I I I bbS I cu6 I 1 3.. cu2 I I cu3 1- II cu4 I 3b cuS J-I I I ,,11 -- I e12 _.1 J J I 3c pe2 I I I I peS -I-- III 1 3d pe3 --I 1--1-1--1-- I I gj1 I ke3 I 1 1<,,4 1--1- II gj2 I I 1 gj3 I-- I 1 1 4.. gj4 I I I I gjS 1--1 I I mr1 LI 1 I keS I II mr2 I I II mrS --1- I I I I 4b mr4 I-I I 1-1- I I ne1 I ne3 -- I I 5 ne4 I I I neS I-I I 1-

kel mr3 I 6 ke2 I I I ne2 1- 7 I I n..3 I na4 1- I 8 naS I I-- I I zu1 -- I zu2 -I I 9.. I zu4 I I zuS I I I 9b zu3 I 1--1 II I II I 0.30 0.48 0.67 0.86 1.04 1. 23

Dis·similarity

Figure 3 Dendrogram of the nine major site groups defined by the classification of the vascular flora. 82 G.]. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge

Table 1 Number of taxa recorded for the eight major regarding the property (Anonymous, 1997) will reserves of the study area. see nearly all of these placed in an expanded Zuytdorp Nature Reserve. Reserve Number of taxa

Kalbarri National Park 1071 Rare and Priority Flora: Toolonga Nature Reserve 200 There are 11 species of declared rare flora and 144 Zuytdorp National Park 314 Cooloomia Nature Reserve 226 taxa of priority flora recorded in the study area Francois Peran National Park 279 (Table 2; CALM, 1998). Of these priority taxa 33 are Bemier Island Nature Reserve 59 PI, 62 are P2, 39 are P3 and 10 are P4. (See Dorre Island Nature Reserve 143 Appendix 1 for definitions of conservation codes). Kennedy Range National Park 314 Of the declared rare species, eight are confined to riverine cliffs or heavy soils in the Kalbarri area (Beyeria lepidopetala, Caladenia bryceana subsp. cracens, Caladenia wanosa, Drakaea concolor, Reservation status of species Eremophila microtheca, Hypocalymma longifolium, There are eight major conservation reserves in this Lechenaultia chlorantha and Leucopogon marginatus). area (Figure 1). A major aspect of the present study was to attempt to produce an indication of the New records and range ends reservation status of individual plant taxa in the Numerous new records and range extensions study area. Species lists for all eight major reserves were noted during the survey. One notable example were compiled. The flora of the numerous small was the recording of Eremophila occidens ms at island nature reserves in the Shark Bay World Nanga, this species having been previously Heritage Area have been compiled separately regarded as endemic to Cape Range. Taxa endemic (Alford et al., 2000; A.A. Burbidge pers. comm.). to all or part of the study area and taxa at their Total numbers of taxa recorded for the eight major northern or southern limits in the study area are reserves are shown in Table 1 with full lists in annotated in Appendix 1. Appendix 1. Most notable is the listing of 223 taxa at their These parks and reserves contained a total of 1559 northern limit in Kalbarri National Park, being over taxa leaving 574 taxa (37% of the total) that were 20% of the Park's known flora. This is a comparable not recorded for any conservation area (Appendix figure to another conservation area with numerous 1). These unreserved taxa were composed of six range ends, the Stirling Range. Another 229 taxa distinctive groups. They were: end their range within the World Heritage area to • Marine angiosperms (Cymodocea, Halophila and the north of Kalbarri (Trudgen and Keighery, 1995). Posidonia). These taxa are largely covered by This is further evidence that the area is a major marine reserves proposed for the W orId transition zone for the vascular flora of Western Heritage Area. Australia. • Wetland taxa of tropical or arid affinities (e.g. Peplidium species, Potamogeton species, Vegetation patterning Schoenoplectus species, Cyperus difformis). These A total of 626 taxa were recorded from the 63 species occur in the northern sector of the study sites (over 25% of the known flora of the study area, around Lake MacLeod, major rivers and area). Of these 626 taxa, 218 (35%) were recorded in in springs near the Kennedy Range. only one site and were excluded because • Terrestrial taxa of tropical affinities, especially preliminary analysis of the total dataset indicated those occurring on clay or alluvial soils that that these 'singletons' added little information. The penetrate the study area in the north, usually final dataset included 408 taxa from 63 sites. Species along the major rivers or drainage lines (e.g. richness varied from nine to 66 taxa per site (with Crinum flaccidum) or on coastal limestones singletons excluded), with individual taxa occurring (Grevillea variifolia). in between two and 40 sites. • Red sand taxa confined to the Acacia shrublands In this analysis site groups are partitioned at the of the central region of the study area. nine group level (Figure 3) and species assemblages • Species confined to freshwater claypans (e.g. at the 20 group level (Table 3). Some site groups Lythrum sp. Towrana (Cranfield 2183) and showed distinct pattern below this level and these Rumex crystallinus) were designated as subgroups. In all, 15 sites • Endemics of the World Heritage Area. There are groups and subgroups were defined. 45 endemics of the Shark Bay World Heritage The major floristic boundary in the southwest of Area that are largely found on Tamala and the study area is clearly evident in the analysis with Nanga Stations. Although these are not yet in a site groups 1 to 6 representing typical Carnarvon conservation reserve, current proposals vegetation types and site groups 8 and 9 l-. _ -- -~-~~~------

Flora and vegetation 83

J 13"

24°

Bernier I t

25°

DOrrell

27'

'< F: Cl

0 o 115 o

Figure 4 Distribution of the nine major site groups across the study area~ 84 G.]. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge

Table 2 Rare and Priority Flora recorded for the study area. [ms indicates a manuscript name currently used by Western Australian Herbarium (PERTH); see Appendix 1 for definitions of conservation codes].

Taxon Conservation Code Taxon Conservation Code

Beyeria lepidopetala R Calytrix paucicostata 2 Caladenia bryceana subsp. cracens ms R Calytrix purpllrea 2 Caladenia hoffmanii ms R Chamelallcium marchantii 2 Caladenia wanosa R Chthonocephalus muellerianlls 2 Drakaea concolor rns R Chthonocephalus tomentellus 2 Drakonorchis barbarella ms R Cryptandra glabriflora 2 Eremophila microtheca R Cryptandra scoparia var. microcephala 2 Eucalyptus beardiana R Dampiera krauseana 2 Hypocalymma longifolium R Eremaea acutifolia 2 Lechenaultia chlorantha R Eremophila glabra subsp. psammophora ms 2 Leucopogon marginatus R Eremophila occidens ms 2 Eucalyptus diminuta ms 2 Beyeria gardneri 1 Frankenia confusa 2 Chamelaucium oenanthum rns 1 Grevillea annulifera 2 Chthonocephalus oldfteldianus 1 Grevillea costata 2 Chthonocephalus spathulatus 1 Grevillea rogersoniana 2 Eremophila cuneata rns 1 Grevillea stenomera 2 Eremophila lanata rns 1 Grevillea triloba 2 Eremophila physocalyx ms 1 Hemiandra sp. Kalbarri (D.Bellairs 1505) 2 Labichea eremaea 1 Malleostemon sp. Cooloomia (S.D.Hopper 1353) 2 Lepidium scandens 1 Malleostemon sp. Kalbarri (L.ACraven 7083) 2 Levenhookia octomaculata 1 Malleostemon sp. Moonyoonooka (RJ.Cranfield Macarthuria georgeana 1 2947) 2 Malleostemon sp. Hardabutt Rapids (D.Bellairs Melaleuca huegelii subsp. pristicensis 2 1654A) 1 Microcorys tenuifolia 2 Millotia depauperata 1 Millotia jacksonii 2 Persoonia brachystylis 1 Murchisonia fragrans 2 Persoonia papillosa 1 Olearia occidentissima 2 Pityrodia axillaris 1 Pityrodia glutinosa 2 Ptilotus beardii 1 Platysace sp. Kalbarri (D.& B.Bellairs 1383) 2 Ptilotus stipitatlls 1 Platysace sp. Kennedy (P.G.Wilson 8450) 2 Ptilotus stirlingii var. pumilus 1 Ptilotus alexandri 2 Rhodanthe sp. Overlander (P.5.5hort 2096) 1 Rhodanthe oppositifolia subsp. ornata 2 Schoenia ftlifolia subsp. arenicola 1 Rumex crystallinus 2 Scholtzia cordata ms 1 Scaevola chrysopogon 2 Scholtzia sp. Eurardy O.5.Beard 6886) 1 Schoenus sp. Kalbarri (K.RNewbey 9352) 2 Scholtzia sp. Folly Hill (M.E.Trudgen 12097) 1 Scholtzia sp. East Yuna (AC.Burns 6) 2 Scholtzia sp. Murchison River (AS.George 7098) 1 Scholtzia sp. Galena (W.E.Blacka1l4728) 2 Sclerolaena stylosa 1 Scholtzia sp. Ross Graham Lookout (S.Maley 6) 2 Tetragonia coronata 1 Scholtzia sp. Z-Bend (Bellairs-Kalflora 912a) 2 Thryptomene ninghanensis rns 1 Sondottia glabrata 2 Thryptomene sp. Carrarang (M.E.Trudgen 7420) 1 Thryptomene sp. Eagle Gorge (AG.Gunness 2360) 2 Thryptomene sp. Steep Point (M.E.Trudgen 7421) 1 Thryptomene sp. Eurardy (D.& B.Bellairs) 2 Thryptomene sp. Tamala (M.E.Trudgen 7384) 1 Thryptomene sp. Junga Dam (AG.Gunness 2383A) 2 Verticordia lepidophylla var. quantula 1 Thryptomene stenophylla 2 Vittadinia cervicularis var. oldfteldii 1 Thysanotus kalbarriensis ms 2 Verticordia dasystylis subsp. kalbarriensis 2 Abutilon sp. Hamelin (AM.Ashby 2196) 2 Verticordia galeata 2 Abutilon sp. Quobba (H.Demarz 3858) 2 Acacia gelasina 2 Acacia didyma 3 Acacia leptospermoides subsp. obovata 2 Acacia drepanophylla 3 Acacia stereophylla var. cylindrata 2 Acacia isoneura subsp. nimia ms 3 Acacia subrigida 2 Acacia plautella rns 3 Acanthocarplls parviflorus 2 Acacia sclerosperma subsp. glaucescens 3 Angianthus microcephalus 2 Anthocercis intricata 3 Anthotroche myoporoides 2 Anthotroche walcottii 3 Baeckea subcuneata 2 Arnocrinum dnlmmondii 3 Bergia auriculata 2 Calocephalus aervoides 3 Beyeria cygnorum 2 Calycopeplus marginatus 3 Calytrix drummondii 2 Centrolepis cephaloformis subsp. murrayi 3 Calytrix formosa 2 Chamelaucium conostigmum ms 3 Calytrix harvestiana 2 Comesperma acerosum 3 Flora and vegetation 85

Table 2 (cont.)

Taxon Conservation Code Taxon Conservation Code

Cryptandra nudiflora 3 Scaevola oldfieldli 3 Dicrastylis linearifolia 3 ScllOltzia sp. Ajana (T.A.Halliday 137) 3 Eucalyptus foecunda subsp. Coolimba Stenantfzemum divaricatum 3 (M.I.H.Brooker 9556) 3 Verticordia cooloomia 3 Geleznowia verrucosa 3 Verticordia densiflora var. roseostella 3 Goodenia sericostacfzya 3 Verticordia dicfzroma var. dicfzroma 3 Grevillea candical1s 3 Verticordia dicfzroma var. syntoma 3 Grevillea leptopoda 3 Villarsia congestiflora 3 Grevillea stel10stacfzya 3 Gymnantfzera cllnningfzamii 3 Calotfzamnus pacfzystacfzyus 4 Hemigel1ia pimelifolia 3 Diuris recurva 4 Lasiopetalum oldfieldii subsp. oldfieldii 3 Goodenia neogoodenia 4 Lasiopetalum oppositifolium 3 JackBonia velutina 4 Lepidobolus densus ms 3 Lepidium puberulllm 4 Macartfzuria intricata 3 Lepidosperma rupestre 4 Mirbelia sp. Kalbarri (MoO.Crisp 6261) 3 Plectracfzne bromoides 4 Petropfzile biternata 3 Verticordia capillaris 4 Pfzysopsis cfzrysopfzylla 3 Verticordia polytricfza 4 Pfzlegmatospermum drllmmondii 3 Wllrmbea murcfzisoniana 4

representing typical Irwin (Geraldton Sandplains) (MD3) had an overstorey of Eucalyptus victrix while vegetation types (Figures 3 and 4). Site group 7 is the understorey was typical of the Acacia represented by only a single site and its affinities shrublands. Another site (B01) was dominated by are not clear. Site groups 1 to 4 were the most Atriplex, it differed from sites in site group 1 in widespread across the study area. being rich in species belonging to assemblage K Site group 1 comprised the sites with a major (especially in terms of herbs), an assemblage largely saline influence. Species assemblage 0 is highly lacking in the site group 1. This site was classified faithful (i.e. largely restricted) to this site group most similar to B03 (a typical Acacia shrubland). with low to moderate levels of constancy (referring Species assemblage A was moderately faithful to to the degree of occurrence of a species assemblage both site group 2 and site group 4, while species in the site group). Species in this assemblage assemblages 0, E, and J differentiated site group 2 include Halosarcia indica and H. halocnemoides from other site groups. Species assemblage G was (samphire species), Frankenia aff. pauciflora (a largely confined to site groups 1, 2a and 2b, species species generally restricted to saline calcareous assemblage I was largely restricted to site group 2a substrates) and a series of annual taxa which are and site group 6 and species assemblage L was most restricted to this site group. This group faithful to site group 2c and site group 5. encompasses a range of saline plots ranging from The different subgroups within site group 2 Halosarcia and Sclerostegia shrublands to an Atriplex reflected differences in Acacia species found on shrubland upslope from a saline flat. Sites occurred these sites. Acacia tetragonophylla was the most in coastal areas from Nanga to Lake MacLeod widespread Acacia occurring in all three subgroups, (Figure 4) and species richness averaged 23.7 taxa/ A. linophylla was largely restricted to subgroups 2a plot. and 2b as was A. victoriae while A. grasbyi was Acacia shrublands are one of the most widespread restricted to site group 2b and A. coolgardiensis and typical communities of the Carnarvon subsp. effusa was largely restricted to subgroup 2c Bioregion. Site groups 2 and 4 represent the Acacia and site group 5 (Table 3). Weedy annuals such as shrublands on red sands. No single Acacia species *Hypochaeris glabra, *Bromus arenarius, *Cenchrus dominated sites in site group 2 but Acacia ciliata, *Sonchus oleraceus and *Asphodelus fistulosa sclerosperma and A. linophylla were the most are more common in subgroups 2a and 2b common; sites in site group 4 were generally compared to 2e. dominated by Acacia aneura (Mulga). Site group 2 The coastal Acacia and Melaleuca shrublands of was a very widespread group in the centre of the site group 3 are largely defined by species study area, stretching from Nanga to Boolathana assemblages Q and R which are highly faithful to and east to Meedo (Figure 4). It had the highest site group 3 but with generally only low to average species richness of 46.6 taxa / plot. moderate consistency. Site group 3 as a whole has a While most sites in site group 2 were dominated coastal distribution extending from Edel Land to by Acacia spp., one site along the Wooramel River Cuvier; average species richness is high at 45.6 taxa Table 3 Sorted two way table showing the major site groups and species assemblages. Sites appear as columns, species as rows. 0'QC Site Groups 1 2a 2b 2c 3a 3b 3c 3d 4a 4b 5 6 7 8 9a 9b

bbnbcplbbmbbmmlbbwwwlmmwwnplbclccccleelppplgkkggggmlkmmmlnnnnlkmklnlnnnlzzlzzz boabuelbbdooddlooooolddooaelbuluuuul11leeeljeejjjjrlerrrleeeelerelelaaaluuluuu 121311124345121133541451224156123451121253113423451152541134511321213451121453 1 1 1 11111 1111111 ------+------+-----+------+--+----+--+---+------+----+----+---+-+---+--+--- Species Assemblage A 1 Abutilon otocarpum **** ** ** ** * 1 Phyllanthus maderaspatensis ***** *** *** 1 Bulbostylis barbata *** * * **** 1 Calandrinia sp. (GJK/NG 1495) *** ** ** * ** *** 1 Cassia helmsii ** * * ****** ** * 1 Eremophila subfloccosa *** ** * *** 1 Maireana planifolia * *** *** 1 Asphodelus fistulosus * ***** ** ** 1 Chenopodium cristatum ** **** **** 1 Chthonocephalus spathulatus *** ** ** * * ** 1 Rhodanthe charsleyae * ***** ***** *** 1 Stenopetalum pedicellare * 1* * * ** * * ** ** 1 Acacia linophylla ** * ** ** 1 Spartothamnella teucriiflora * ** ** * 1 Myriocephalus guerinae * 1 *** * * 1 Acacia victoriae *** ** * ** ** 1 Tragus australianus ** ** ** ***** * 1 III C'l Sclerolaena densiflora ** I ** *** ** 1 1 1 1 ..;-. Boerhavia gardneri 1 1 1 1 ~ * ** ** ** III Hakea preissii ** * ** 1 1 1 1 ~. Corchorus walcottii **** ::r ** **** 1 1 1 1 III Eragrostis dielsii I *** *** ***** * ** *** ** ** * 1 1 1 1 ~ Ptilotus villosiflorus ** ** *** * **** * 1 1 1 1 Triraphis mollis * 1 ** ** ** * * *** 1 1 1 1 1 :z Eriachne aristidea ** **** ** 1 1 1 1 ;:C'l Acacia aneura 1 1 1 **** *** * 1 1 1 1 Cl> Eriachne 0 pulchella 1 * * *** ** 1 1 1 1 .? Ptilotus helipteroides 1 ****** * ** 1 1 1 1 Tribulus astrocarpus 1 1 ** **** ** 1 1 1 1 :: Tripogon loliiformis 1 1 * **** *** 1 1 1 1 Z r-' Paspalidium basicladum 1 * *1 1 * ** * *1* * 1 1 1 1 '< 0 Goodenia havilandii 1 1 1 * ***** 1 * 1 1 1 1 ::l Enneapogon caerulescens * 1 1 1 *** *1* 1 1 1 1 !:!' Haloragis trigonocarpa 1 * 1 1 1 ** *1* 1 1 1 1 > ptilotus aervoides 1* * 1 1 * 1 ** *1* 1 1 1 1 ~ Stenopetalum sphaerocarpum 1* ~ * 1 * 1 *1 1 *** 1 1 1 1 l: Cleome viscosa ... 1 1 1 1 1 * *1 1 1 1 1 0- Lepidium oxytrichum 1 1 1 1 1 * *1 1 1 1 1 c: Eriachne helmsii (JQ 1 1 1 1 1 * *1 *1 1 1 1 III , . Indigofera monophylla 1 1 1 I11II***I 'TI ,.,0- Eriachne dominii 1 1* I * I 1 I 1 I **1 * l'ol l'ol Sida kingii 1 I* I 1 1 1 1 1 * * I* ::l ---- -+------+--- -+-- ---+--+----+--+---+------+----+----+---+-+- -+--+--- Q.. 0( Species Assemblage B ~ * ** *** ()q Eriachne avenacea ~ Waitzia acuminata ** ;, Menkea villosula *** o Thyridolepis mitchelliana * * ** ::l Eremophila setacea ms * ** Plectrachne rigidissima sida rohlenae * Hibiscus coatesii * *** - ----+------+-- -+------+--+----+--+---+------+----+----+---+-+---+--+- - Species Assemblage C Acacia xiphophylla * * * * ptilotus exaltatus * ** Cassia sturtii ** ** ptilotus polakii ** Streptoglossa liatroides *** Panicum effusum ** Plectrachne schinzii * * Sauropus crassifolius ** Sida corrugata * * Atriplex lindleyi subsp. inflata * Marsdenia viridiflora ** Sida aff. intricata (GJK/NG 1134) ** Yakirra australiensis ** ------+------+-----+------+--+- -+--+---+------+----+----+---+-+---+--+-- Species Assemblage D Acacia grasbyi *** Sarcostemma viminale ** Cassia sp. ** * Zygophyllum ovatum * * *** Atriplex semilunaris * ** Sisymbrium iri *** Thysanotus speckii * ** * ---+------+-----+------+ -+-- +--+- -+------+ -+- -+---+-+---+--+-- Species Assemblage E Brachyscome ciliaris * * *** Podolepis capillaris * * ** *** * Tetragonia cristata *** Enteropogon acicularis * ** * ** Eremophila latrobei ** * Maireana carnosa ** * Eremophila clarkei ** Hibiscus burtonii * * Eremophila leucophylla ** Ptilotus grandiflorus *** 00 Gyrostemon ramulosus * * '-J - ----+- ----+-- --+- ---+ -+ ---+--+---+------+-- -+----+---+-+---+ -+--- Table 3 (cont.)

Site Groups 1 2a 2b 2c 3& 3b 3c 3d 4a 4b 5 6 7 8 9a 9b

bbnbcplbbmbbmmlbbwwwlmmwwnplbclccccleelppplgkkggggmlkmmmlnnnnlkmkInlnnnlzzlzzz boabuelbbdooddlooooolddooaelbuluuuulllleeeljeejjjjrlerrrleeeelerelelaaaluuluuu 121311124345121133541451224156123451121253113423451152541134511321213451 12 1453 1 1 1 11111 1111111 ------+------+-----+------+--+----+--+---+------+----+----+---+-+---+--+--- Species Assemblage F 1 1 1 11111 1111111 Acanthocarpus verticillatus 1* 1*1 11111 1111111 Glycine canescens 1** 1*1 11 111 *1 1 11111 Gymnema granitica ms 1* 1*1 11111 1111111 Angianthus tomentosus **1 1**1 11 111 1 1 11I11 Lawrencia spicata ** 1 1*1 11111 111 I 1 1 1 Maireana georgei * 1 1*1 11111 1111111 Zygophyllum kochii *1 1*1 11111 1111111 Chorizema racemosum 1 1*1 11 111 1*1 1*111 I Eriachne benthamii 1 1*1 11111 1*111111 Calandrinia eremaea 1 1**1 11 111 1 1*11111 Calandrinia liniflora 1 1*1 11 11*1 1 1 11111* Lepidium linifolium 1 1*1 11 1*11 *1 1 11111 ------+------+-----+------+--+----+--+---+------+----+----+---+-+---+--+--- Species Assemblage G 1 1 1 11111 1111111 Amaranthus mitchellii 1*** **1* 1 11111 1111111 Commicarpus australis 1 ** **1* 1 *1 *1 1 1* 1 1 1 1 1 1 1 1 Portulaca oleracea 1**1* 1 11 111 *1 1 11111 Trianthema triquetra 1 * ** 1 1 11111*11 11111 Maireana tomentosa *1 *1* 1 11 11*1 1 1 11111 Atriplex vesicaria **** 1* 1**1 11 1*11 1 1 11111 Swainsona pterostylis ** 1* * 1** 1 1 1 1 *1 1 1 1 1 1 1 1 1 Calotis multicaulis *** 1 ** **1** **1 1 1 1 1 1 *1 1 1 1 1 1 1 Cephalipterum drummondii ** 1 1***** 1 11 111 *1 1 11111 Sclerolaena recurvicuspis * 1 **1* ** 1 1 1 1 1 1 1 1 I 1 1 1 1 Senecio glossanthus ** * 1 * * 1 * I 11111 11 11111 Maireana trichoptera ** 1 1*1 1111*1 11 11111 Pogonolepis muelleriana **1 *1 1 11 111* 1 1 11111 Convolvulus erubescens 1 *** 1** 1 11 111 *1 1 1111I Heliotropium undulatum 1* ** * 1** 1 11*111 I I 11111 Hypochaeris glabra 1**** 1**1 11 111 1 1*11111 Lotus cruentus ** 1** * 1* 1 1 *1 1 1 1 1 1 1 1 1 1 1 Rhodanthe psammophila * 1** * 1 1 1 1** 1 1 1 1 1 1 1 1 1 1 Rhodanthe chlorocephala 1*1* 1 11111 1111111 Tribulus forrestii * 1 * 1** 1* 11 111 1 1 11I11 Dactyloctenium radulans 1 1* 1 11111 *111I11I Eragrostis basedowii 1 1* 1 11111 *1111111 Tribulus terrestris 1 1* 1 11111* 1111111 Lawrencia densiflora 1 1* 1 11*111* 1 1 11111 Wahlenbergia tumidifructa 1* 1* 1 11 111* 1 1 11111 ------+------+-----+------+--+----+--+---+------+----+----+---+-+---+--+--- ..., 1 1 Species Assemblage H 1 1 1 1 1 1 ...0' Calandrinia ptychosperma 1 * 1 1 1 1 * 1 1 1 1 l'I Eucalyptus victrix 1 1 1 * 1 1 1 1 l'I 1 * 1 ::l Indigofera colutea 1 1 1 1 1 *1 1 1 1 Q. 1 1 <: Emex australis 1 * 1 1 *1 1 1 1 ~ ()Q Sisymbrium erysimoides 1* * 1 1 1 1 1 1 1 1 ~ Lotus australis * 1 * 1 1 1 1 1 1 1 1 £ ------+------+-----+------+--+----+--+---+------+----+----+---+-+---+--+--- o' Species Assemblage I 1 1 1 1 1 1 1 :l Acacia murrayana * 1 1 1 1 1 1* 1 Pityrodia paniculata * 1 1 1 1 1 1 * 1 Adriana tomentosa ** 1 1 1 1 1 1* 1 Trichodesma zeylanicum *** * 1 1 1 1 * 1 1* 1 Anthobolus foveolatus * 1 1 1 * 1 1 ! 1 Indigofera occidentalis ms * 1 1 1 1 1 1 * 1 Acacia ramulosa * 1 1 * 1 1 1 1 1 Alectryon oleifolius * 1 1 1 1 * 1 1 1 Corynotheca pungens ** 1 1 1 1 1 1 * 1 Psammagrostis wiseana * 1 1 1 1 1 1 * 1 Pityrodia loxocarpa ** 1 1 1 1 *1 1** 1 Tephrosia gardneri (GJK/NG 1074) 1 ** 1 1 1 1 1 1 * 1 Eremophila crenulata ms 1 **1 1 1 1 1 1 1 Tribulus cistoides 1 **1 1 1 1 1 1 1 Rhodanthe citrina 1 ***1 1 * 1 1 1 1 1 Goodenia tenuiloba 1 * 1 1 1 1 * 1 1 1 Rhagodia drummondii 1 * 1 1 1 1 *1 1 1 1 1 * ------+------+-----+------+--+----+--+---+------+----+----+---+-+---+--+--- Species Assemblage J 1 1 1 1 1 1 1 1 1 1 1 Calotis hispidula 1 ** 1 1 1 1 1 1 * *1 1 1 1 Solanum cleistogamum 1 * 1 1 1 1 *1 1 *1 1 1 1 Exocarpos sparteus 1 1 1 1 1 * 1 1 1 1 1 1 Lobelia winfridae 1 1 1 1 1 1 1 1 1 1 1 Leichardtia australis 1 ** 1 1 * 1 1 1 1 1 1 1 1 Rhagodia eremaea 1 ** 1 1 1 1 1 1 1* 1 1 1 Senecio gregorii 1 ** 1 1 1 1 1 1 1 1 1 1 Jasminum calcarium 1 * 1 1 1 1 1 1 1 1 1 1* 1 * ------+------+-----+------+--+----+--+---+------+----+----+---+-+---+--+--- Species Assemblage K 1 1 1 1 1 1 1 1 1 1 Abutilon oxycarpum 1 1** 1 1 * 1 1 ** 1 1 1 1 Euphorbia australis 1* * *1** * 1 1 ** 1 * 1 1 * 1 1 1 Heterodendrum oleifolium 1 * 1 ** 1 *1 * ** 1 1 *1 1 1 1 Cassia chatelainiana 1 ****1 *1 1 * ** 1 *1 ** 1 1 * 1** 1 Lobelia heterophylla 1 ** 1 1 1 *** 1 *1 1 1 * 1 1 Acacia sclerosperma 1 *** * 1*** *1 1* 1 1 ** **1 1 1 1 Cenchrus ciliaris 1**** * 1** * 1 1 * ** 1** 1* * ***1 * *1 1 1 Acacia tetragonophylla 1 ******1*****1 * ***1 **** 1***1 ****** 1 *1 1 1 Enchylaena tomentosa 1*******1*****1 ** 1 * ** 1***1 ** 1 1 1 1 Euphorbia boophthona * * 1*******1** * 1* 1* **** *1***1 **** 1* **1 1* 1 Paractaenum novae-hollandiae 1****** 1**** 1* ** 1** ** * *1* 1 * ** 1 *1 1 * 1 Chenopodium gaudichaudianum 1 * ****1*****1* * *1 * 1 1 ******1 1 1* 1 '" Table 3 (cont.)

Site Groups 1 2a 2b 2c 3a 3b 3c 3d ~a ~b 5 6 7 8 9a 9b

bbnbcplbbmbbmmlbbwwwlmmwwnplbclccccleelppplgkkggggmlkmmmlnnnnlkmklnlnnnlzzlzzz boabuelbbdooddlooooolddooaelbuluuuulllleeeljeejjjjrlerrrleeeelerelelaaaluuluuu 121311124345121133541451224156123451121253113423451152541134511321213451 12 145 3 1 1 1 11111 1111111 ------+------+-----+------+--+----+--+---+------+----+----+---+-+---+--+--- Tetragonia diptera ** 1** **1***** * *** * I ** * I 11***1 I Aristida holathera I **** I* *** ** ** ** I * ******** **** I * *** III I Ptilotus polystachyus I******* I *** ****** * I *** * **** I ** II*** II Euphorbia drummondii * I ** ** I * * ** ** 1** *** * ** ** * ** I ** IIII Solanum lasiophyllum * 1* *** I * ** * * ** *1 * *** ******** ****1 ***1 1* II Murchisonia volubilis 1** *** 1 * * * *1 * * * * 1 **1 1 *1 1 Eragrostis lanipes 1 * ** 1 * ** * 1 ****** **** 1 * 1 1 1 1 Brachyscome cheilocarpa *** 1* * ** 1 * * * * 1 ** 1 1 1 1 1 Triglochin calcitrapum *** * 1 *** 1* * ** * 1 1 1 1 1 1 * Brachyscome iberidifolia I ** I* * ** *** * 1** ** **** *** III I Calandrinia polyandra ** * 1** **** I***** ****** * ** * I** *** **** ** * II*** II Erodium cygnorum ** 1* *** * **** ***** **** I** *** ** **** IIII * Pimelea microcephala I **** ** ** * ** * ** *** ** ** * IIII * Crassula colorata *** ** * *** ** ** ** ** ** *** 1 1 1 1 * ptilotus gaudichaudii ** ***** * ** * ** ** * ****** *** ** 1 1 1 1 ptilotus obovatus * * * ***** ****** * *** *** ******** * *** 1 1 1 1 * Sida calyxhyrnenia ** * * * ** * **** * *** ** * * * 1 1* 1 1 Rhagodia latifolia *** * * * * * ** * * **** 1 1 * 1 1 * Stylobasium spathulatum ** *** ** * ** * * * 1 1*** 1 1 * Brachyscome latisquamea * ** * * ** *** 1 1 1 1 Sclerolaena diacantha ** * * ** ** ** 1 1 1 1 Brassica tournefortii * *** ** ** ** * *** 1 1** 1 1 Threlkeldia diffusa * ** * * ** ** ** *** 1 1 * 1 1 ptilotus divaricatus * **** ** *** * *** 1 1* 1 1 Rhagodia preissii * **** * ** * *** 1 1 1 1 * Nicotiana occidentalis ** **** ** ** * * *** ** * * * * 1 1 1 1 Rostraria pumila ***** * ** * * ** * ** *** ** *** * 1 1 1 1 Sonchus oleraceus * * ** * ** * * * * *** 1 1 1 1 Scaevola spinescens **** * * ** *** * 1 1 1 1 1 Scaevola tomentosa ***** * * **** * *** 1 ** 1 1 1 1 Zygophyllum fruticulosum * *** * * * * *** * 1 1 1 1 1 Cuscuta epithyrnum * ** * * * * 1 1 1 1 1 Lepidium rotundum ** * * ** ** * * * 1 1 1 1 1 Salsola kali *** ** *** ** ** * * ** 1 1 1 1 1 Dysphania rhadinostachya ** * * ** ** * *** 1 1 1 1 1 Goodenia berardiana * ** * *** * * ** * * * ** 1 1 1 1 1 Gnephosis arachnoidea * * * ** * * * * * 1 1 1 I I Parietaria debilis * ** * ** * * * 1 1 1 1 I ------+------+-----+------+--+----+--+---+------+----+----+---+-+---+--+--- Species Assemblage L 1 1 1 1 1 1 1 1 1 1 1 Acacia coolgardiensis subsp. effusa 1 1 * 1* *** 1 1 * 1 *** 1 1 1 1 1 Schoenia cassiniana * ** 1* ** I * Lawrencella davenportii ** * I * * * *** Uldinia ceratocarpa * 1 * wai tzia ni tida * * * **** * *1 * * **** Chthonocephalus tomentellus * ** **** 1 * * * Eremophila maitlandii * * *** * * I * Porana sericea * * *** I ** Rhyncharrhena linearis * ** *** I ** Podolepis canescens * * * I * Actinobole uliginosum * * I ** Solanum orbiculatum * * I *** *** * Chthonocephalus pseudevax I * Marsdenia australis * I * * * * ** Dianella revoluta * I **** * Stipa elegantissima * * ** I ** **** ------+------+-----+------+--+----+--+---+------+- --+- -+-- +-+- -+--+- - Species Assemblage M I III Acanthocarpus aff. robustus (Hopper 1367) I I 1* **I *** Thysanotus patersonii I I* 1* **I * Mirbelia ramulosa I * * * I * II Melaleuca sp. (Beard 6768) I * III * * Plectrachne bromoides I 1* II ** Acanthocarpus robustus I 1** II Loxocarya aspera ms I I * II * ------+------+-----+------+--+----+--+---+------+----+-- -+---+ +---+--+--- Species Assemblage N I I Acacia latipes I 1 * * Lechenaultia linarioides I I * * Acacia subrigida I I ** Lasiopetalum oppositifolium I I ** Melaleuca aff. quadrifidus I I Eremophila occidens ms I I Wurmbea cernua I I Pityrodia verbascina I I * Acacia longispinea I I * Tricoryne aff. corynothecoides (GJK/NG 1274) I I ** Eucalyptus eudesmioides I I * Lamarchea hakeifolia var. brevifolia I I Stenanthemum complicatum 1 I * * Baeckea sp. (A.S.George 11346) 1 * I Brachychiton gregorii I 1 Grevillea stenobotrya I I * Verticordia forrestii I I Allocasuarina acutivalvis I I ** * Grevillea eriostachya I I * ** * Acacia rostellifera I * **I* * Acacia spathulifolia I * * I *** Calothamnus borealis I * * I * Halgania cyanea I * I Alyxia buxifolia I I ------+------+-- -+--- --+--+--- +--+---+------+----+----+---+-+---+ -+--- Table 3 (cant.) Site Groups 1 2a 2b 2c 3a 3b 3c 3d ~a ~b 5 6 7 8 9a 9b

bbnbcplbbmbbmmlbbwwwlmmwwnplbclccccleelppplgkkggggmlkrnmrnlnnnnlkmklnlnnnlzzlzzz boabuelbbdooddlooooolddooaelbuluuuulllleeeljeejjjjrlerrrleeeelerelelaaaluuluuu 12131112434512113354145122415612345112125311342345115254113451 13212134 5112 1453 1 1 1 11111 1111111 ------+------+-----+------+--+----+--+---+------+----+----+---+-+---+--+--- Species Assemblage 0 1 1 1 11111 1111111 Carpobrotus sp. (GJK/NG 1707) **1 1 1 11111 1111111 Rhodanthe stricta **1 1 1 11111 1111111 Frankenia aff. pauciflora **1 1 1 11111 I I I1111 Triglochin centrocarpum **1 1 1 11111 111I111 Eragrostis pergracilis * ** 1 1 1 11111 1111111 Gunniopsis septifraga **1 I 1 1111I 1111111 Lawrencia viridigrisea **1 1 1 11111 1111111 Podolepis gardneri ***1 1 1 11111 11 11111 Zygophyllum compressum **1 1 1 11111 1111111 Halosarcia indica **1 1 1 1*1111 1111111 Halosarcia halocnemoides *** 1 1 1 11111 1111111 Swainsona kingii ** 1 1 1 I1111 1111111 Sondottia glabrata *** *1 1 1 11111 1111111 Didymanthus roei ***1 1 1 11111 1111111 Sclerolaena eurotioides ** *1 1* 1 11111 1111111 ------+------+-----+------+--+----+--+---+------+----+----+---+-+---+_.,+--- Species Assemblage P 1 1 1 1 1 1 1 1 1 1 1 1 Acacia roycei 1 1 1 * 1 1 1 1**** 1 1 1 1 1 Eucalyptus mannensis 1 1 1 * 1 1 1 1 *** 1 1 1 * 1 1 Calandrinia corrigioloides 1 1 1 * * 1 1 1 1 *** 1 1 1 1 1 * Bursaria occidentalis 1 1 1 * 1 1 1 1 ** 1 1 1** 1 1 Thysanotus manglesianus 1 1 1 ** 1 * 1 1 1**** 1 1 1 1 1 * Calandrinia lehmannii 1 1 * 1 *** 1 1 1 1** *1 1*1 1 1 Leucochrysum fitzgibbonii 1 1 1***** 1 1 1 1 *** 1 1 1 1 1 Wurmbea densiflora 1 1 *1 * 1 1 1 1****1 1*1 1 1 Callitris glaucophylla 1 1 1 1 1 1 1 ** * 1 1 1 1 1 Gilberta tenuifolia 1 1 1 1 1 1 1 *** 1 1 1 1 1 Lysiana casuarinae 1 1 1 1 1 1 1 * * 1 1 1 1 1 Trachymene ornata 1 1 1 * 1 1 1 1 *** 1 1 1 1 1 Grevillea paradoxa 1 1 1 1 1 1 1 * 1 1*1 1 1 Plantago aff. hispidula 1 1 1* 1 1 1 1 1 ** 1 1 1 1 1 Centrolepis drummondiana 1 1 1 1 1 1 1 1* * 1 1 1 1 1 Comesperma integerrimum 1 1 1 1 1 1 1 1* * 1 1 1 1 1 Gnephosis eriocephala 1 1 1 1 1 1 1 1** * 1 1 1 1 1 Poranthera microphylla 1 1 1 1 1 1 1 1** 1 1 1 1 1 Goodenia occidentalis 1 1 1 * 1 1 1 1* 1****1 1 1 1 1 Trachymene cyanopetala 1 1 1 1 1 1 1*1***1 11 1 1 Eremophila oldfieldii 1 1 1 * 1 1 1 1 1* 1 1 1 1 1 ptilotus stirlingii 1 1 1 * 1 1 1 1 1* 1 1 1 1 1 Brunonia australis 1 1 * 1 1 * * 1* *1 ****1****1 Monachather paradoxus 1 1 * 1 1 ** Velleia hispida 1 1 1 1 * 1 *1 ptilotus drummondii 1 1 1 1 ** 1 ***1 Synaptantha tillaeacea 1 1 1 1 * 1* *1 Gnephosis tenuissima 1 1 1 *1 1* *1 -+--+----+--+---+------+----+----+---+-+---+--+---

Species Assemblage Q 1 1 1 Acacia chartacea 1 1 **1 Diplopeltis intermedia 1 1 **1 Goodenia triodiophila 1 1 **1 Scaevola thesioides 1 1 **1 Acacia coriacea subsp. coriacea 1 1 * *** 1 Haloragis gossei 1 1 ***1 Triodia pungens 1 1 ****1 Daviesia hakeoides 1 1 ** 1 Brachysema macrocarpum 1 1 * **1* Scaevola sericophylla 1 1 ***1 * * Triodia basedowii 1 1 ****1 Dampiera incana 1 1 * **1 * Hibiscus sturtii 1 1 * 1 Indigofera brevidens 1 1 ****1 * Zygophyllum eremaeum 1 1 ** *1 Stipa ni tida 1 1 ** 1 * Olearia dampieri subsp. dampieri ms 1 1 *1 * Stackhousia muricata 1 1 * 1 -+---- -+- -+--+----+--+---+------+----+----+- -+-+---+--+-

Species Assemblage R 1 1 1 1 Acacia ligulata 1 1 * 1* *1* Trichanthodium skirrophorum 1 * *1 **1* *1 Beyeria cinerea 1 1 * 1 * 1 Eragrostis barrelieri 1 1 * 1 * 1 Erodium cicutarium 1 1 *1** 1 Triodia plurinervata 1 1 * 1** 1 Urospermum picroides 1 1 1** 1 Eremophila glabra 1 1 **1* 1 Wurmbea inframediana 1 1 * * 1 * 1 Lepidium puberulum 1 1 ** 1 1 Olearia aff. axillaris (GJK/NG 1025) 1 1 ** 1 1 Poa drummondiana 1 1 ** 1 1 Triglochin trichophorum 1 1 ** 1 1 Podotheca gnaphalioides 1 1 * ** 1 1 Exocarpos aphyllus 1 1 ** * **1***1 Melaleuca cardiophylla 1 1 * ** **1* *1 * Podotheca angustifolia 1 1 ** **1 *1 Thryptomene baeckeacea 1 1 *** **1***1 * Stipa crini ta 1 1 ** * 1 1 Waitzia suaveolens 1 1 *** **1 I Olearia axillaris 1 1 1 * 1 Trachymene elachocarpa 1 1 ** 1 1 * Table 3 (cant.)

Site Groups 1 2a 2b 2c 3a 3b 3c 3d 4a 4b 5 6 7 8 9a 9b

bbnbcplbbmbbmmlbbwwwlmmwwnplbclccccleelppplgkkggggmlkmmmlnnnnlkmklnlnnnlzzlzzz boabuelbbdooddlooooolddooaelbuluuuulllleeeljeejjjjrlerrrleeeelerelelaaaluuluuu 121311124345121133541451224156123451121253113423451152541134511321213451121453 1 1 1 11111 1111111 ------+------+-----+------+--+----+--+---+------+----+----+---+-+---+--+--- Atriplex cinerea 1 1 1 1 *1 1 1 * 1 1 1 1 scaevola crassifolia 1 1 1 1**1 1 1 1 1 1 1 Frankenia pauciflora * 1 1 1 1** 1 1**1 1 1 1 1 Rhodanthe humboldtiana 1 1 1 1 *1 1 *1 1 1 1 1 Zygophyllum ammophilum 1 * 1 1 1**1 * 1**1 1 1 1 1 Atriplex paludosa subsp. moquiniana * 1 1 1 1* 1* 1 1 1 1 1 1 Senecio lautus * 1 * 1* 1 1**1 *1 1 1 1 1 1 Cyperus bulbosus * 1 * 1 1 1 1 1 *1 1 1 1 1 Millotia myosotidifolia * 1 1 1 1 1 1 *1 * I 1 I I Bromus arenarius * 1* * 1 1 *1* 1 I *1 * I 1 I I Daucus glochidiatus * 1 * I 1 I 1 * I 1** * 1 1 1 I Cryptandra leuophracta 1 1 1 1 1 1 1 * I I *1 I Danthonia caespitosa 1 I 1 1 1 I 1* *1 1 1 1 1 Solanum oldfieldii 1 1 1 1 1 1 1* *1 1* 1 1 1 1 Dysphania plantaginella *1 1 1 11*I1*1 1 1 1 1 1 ------+------+-----+------+--+----+--+---+------+----+----+---+-+---+--+--- Species Assemblage S 1 1 1 1 1 1 1 1 I I Acacia blakelyi 1 I 1 1 1 IIII 1** Actinostrobus arenarius 1 1 1 1 I I 1 1 I 1** Allocasuarina campestris 1 I 1 I 1 I I 1 I 1** Calytrix strigosa 1 1 1 1 1 1 1 1 1 1** Leucopogon cucullatus 1 I I 1 1 1 I 1 I 1** Lyperanthus nigricans II I 1 1 I I 1 I 1** Tetraria microcarpa II I 1 I I I I I 1** Thryptomene denticulata II 1 1 I 1 1 I 1 1** Acacia cavealis fiS 1 1 1 1 1 1 1 I 1 *1* Drosera stolonifera subsp. stolonifera 1 1 1 1 1 1 1 1 1 *I* Boronia coerulescens II I 1 I 1 1 I I *I * Brachysema aphyllum I 1 1 I 1 I I 1 1 *I * Monotaxis lurida 1 1 I 1 1 I I 1 I *I * Petrophile semifurcata 1 1 1 I 1 1 I 1 1 *I * Scaevola canescens 1 1 1 I I I I 1 I *1 * Calothamnus blepharospermus 1 1 1 I 1 I I 1 1** I** Eremaea ebracteata 1 1 1 I 1 1 1 1 1** 1** Stylidium macrocarpum 1 1 1 1 1 1 1 I 1**1** Stylidium repens 1 1 1 1 1 1 I 1 1** I** Leucopogon cordifolius 1 1 1 1 1 1 I 1 1** I* Lysinema ciliatum 1 1 1 1 I 1 I 1 1** I* Neurachne alopecuroidea 1 1 1 1 1 I 1 1 1** 1 * Petrophile brevi folia 1 1 1 1 1 1 1 1 1**1 * ** * ~ Schoenus clandestinus 0 * ~ Melaleuca conothamnoides ~ * Melaleuca scabra ~ * ~ Mesomelaena preissii ~ ** * Opercularia spermacocea ~ * *** ~ Conospermum microflorum ms ~ *** ~ Lepidobolus preissianus * [ Phymatocarpus porphyrocephalus * *** o' Drosera stolonifera subsp. humilis ** *** ~ Malleostemon sp Cooloomia (Hopper 1353) * ** Mesomelaena pseudostygia *** Hibbertia racemosa Hibbertia subvaginata * Melaleuca acerosa Conostylis stylidioides ** Hibbertia pungens Melaleuca aff. leiopyxis (GJK/NG 1708) ** Ecdeiocolea monostachya ** ** ** Hibbertia conspicua ** ** Persoonia acicularis * ** ** Banksia ashbyi * ** * Banksia sceptrum ** Geleznowia verrucosa * ** Calytrix brevi folia ** * ** ** Grevillea annulifera * Plectrachne danthonioides * Scholtzia sp. Folly Hill (ME Trudgen 12097) * * Cassytha aurea * ** * ------+------+-----+-- ---+--+- -+--+---+------+----+----+---+-+---+--+--

Species Assemblage T I I II I I I I ** Amphipogon turbinatus I I II I I I I ** Boronia purdieana I I I I I I I I ** Cassytha racemosa I I II II I I ** Conostylis aculeata I I I I I I I I ** Dryandra borealis subsp. borealis I I I I I I I I ** Gompholobium tomentosum I I II I I I I ** Grevillea preissii ms I I I I I I I I ** Hibbertia spicata subsp. spicata I I I I I I I I ** Patersonia occidentalis I I I I I I I I ** Pimelea leucantha I I II I I I I ** Pityrodia oldfieldii I I I I II I I qr,0;~;um ** elongatum I I II II I I 96 G.}. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge

Figure 5 Differences in predicted climatic parameters between the nine site groups defined by the vascular flora. (L = lowest value, U =highest value, I =first quartile, M =mean, D =median).

Mean annual temperature (OC): (Kruskal-Wallis 49.9, df = 8, p < 0.0001)

20.80 21.52 22.25 22.98 23.70 +------+------+------+------+------

1 L1======DM======3---U 2 L------1======MD======3---U 3 L------1======M=D======* 4 L-1=* 5 L1M* 6 *=D=M===3U 7 * 8 * 9 *M3U

Diurnal temperature range (OC): (Kruskal-Wallis 37.5, df = 8, p < 0.0001)

12.00 12.88 13.75 14.63 15.50 +------+------+------+------+ 1 *======D====M======3------U 2 L------1======M======D==U 3 3 L---1=====D==M======3--U 4 L-----1======M==D======* 5 L1*=* 6 L1===MD=* 7 * 8 L1=*3U 9 *=M3-U

Annual temperature range (OC): (Kruskal-Wal1is 38.3, df = 8, p < 0.0001)

23.90 25.50 27.10 28.70 30.30 +------+------+------+------+ 1 *======D===M======3------U 2 L------1======M======D===U 3 3 L----1===D====M======3--U 4 L------1======M=====D===* 5 L1MD* 6 L---1======M====DU 3 7 8 L1=MD* 9 L-1=MD*

Precipitation wettest quarter (mm): (Kruskal-Wallis 30.2, df = 8, p 0.0002)

29.00 44.00 59.00 74.00 89.00 +------+------+------+------+ 1 L-1======D==M======3------U 2 L------1======D=M======3------U 3 L------1======D===M======3-U 4 *======D===M======3----U 5 L-1=MDU3 6 1 LD====M======3--U 7 * 8 L1=MD* 9 *DM3U Flora and vegetation 97

Figure 5 (cont.)

Precipitation coldest quarter (mm): (Kruska1-Wa11is 51.8, df 8, p < 0.0001)

69.00 101. 0 133.0 165.0 197.0 +------+- -+------+------+

1 *===D===M======3------U 2 *=====D==M======3------U 3 1LD======M======3------U 4 L1=*=3U 5 *DM=3-U 6 **3U 7 8 *=DM==3U 9 L1==MD=*

Annual precipitation (mm); (Kruskal-Wa11is = 43.5, df 8, P < 0.0001)

204.0 236.3 268.5 300.8 333.0 +------+------+------+------+

1 L-l=====*=====3---U 2 *=D======M======3------U 3 L------1======D=M======3 ------U 4 L-l=====M===D==* 5 *DM=3U 6 Ll=MD=* 7 * 8 Ll==*==3U 9 Ll===DM===*

/ plot. The four subgroups show strong assemblage. Assemblages Band C are largely geographical and edaphic separation with faithful to site group 4 and differentiate between subgroup 3a occurring on white dunes at Bush Bay two subgroups, 4a which tends to have shallower and Cuvier, subgroup 3b occurring on red sands at stony soils with significant calcrete in the upper Cuvier, subgroup 3c occurring on white sands at layer and 4b on deeper sandy soils (Wyrwoll, Edel Land and subgroup 3d occurring on red sands Stoneman, Elliott and Sandercock, 2000). Beard on Peron Peninsular. An separate analysis of sites (1990) states that Mulga does not normally occur in in site group 3 confirmed highly faithful and the Camarvon region but this community type was constant species assemblages for subgroup 3b (i.e. widespread in the north east quadrant of the study Triodia basedowii, Triodia pungens, Indigofera area. brevidens, Eragrostis lanata, Acacia rostellifera); 3c (i.e. Site group 5 consists of eucalypt - Callitris or Acanthocarpus robustus, Lepidium puberulum, Olearia eucalypt - Acacia woodlands on red sand on the sp. Edel, Poa drummondiana, Podolepis gnaphalioides, Toolonga plateau and site group 6 are sites of the Triglochin tridlOphorum); and 3d (i.e. Acanthocarpus red dunes on the top of the Kennedy Range (KEl, aff. robustus, Solanum orbiculare, Eragrostis dielsii, KE2) and tall dunes at Mardathuna (MR3). Both Dianella revoluta, Austrostipa elegantissima) while site group 5 and 6 differ from the other Camarvon subgroup 3a lacked species assemblages typical of groups (site groups 1 to 4) in lacking most species the other subgroups. from species assemblage K, which are generally Site group 4 consists of the inland Acacia ubiquitous (although site group 4 is under shrublands of the Mardathuna - Kennedy Range represented in some species). Species present from footslopes - Gascoyne Junction area (Figure 4). Most this assemblage differ between site groups 5 and 6 sites are dominated by A. aneura, two have a (Table 3). Species assemblage P is typical (ie eucalypt overstorey while one site (GP) is showing both high degrees of constancy and dominated by Senna artemisioides ssp helmsii. fidelity) of site group 5 and species assemblage L Average species richness is still high (39.6 taxa/ is shared with other Camarvon groups. Site group plot) but lower than site groups 2 and 3. Species 6 is largely defined by the lack of assemblages that assemblage A is typical of this site type with a much delimit the other Carnarvon groups, average higher constancy than site type 2 which shares this species richness is low for site group 6 (21.7 / plot) 98 G.]. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge while the red sands of the Toolonga plateau (site limestone. Species in species assemblage Tare group 5) are much richer with an average species southwestern taxa generally typical of massive richness of 38.3 species/plot and similar to site limestones. Average species richness for site group 9 groups 2 to 4. was 39.4 taxa/plot. The affinities of site group 7, which consists of a single site NE2, are not clear. This site is a Banksia Environmental correlates ashbyi-B. sceptrum woodland over heath on There is a strong correlation between the site yellowish - red sands near the top of a dune. It was groups and most of the major climatic gradients of an unusual site in that no annuals were recorded on the study area. The temperature and rainfall the plot in either year despite a significant annual parameters derived from ANUCLIM show a very flora being recorded at NEl (only some 3 km to the high degree of inter-correlation and it was not NW) in 1996. It is likely that this site is transitional possible from the available data to determine to between the limestone heaths of site group 9 and which aspect of these parameters the vascular flora the eucalypt woodlands of site group 5. With the is responding. Most parameters showed significant complete lack of annual taxa and only 12 perennial differences between site groups (Figure 5). taxa recorded from this plot, its strongest Patterns in mean annual temperature reflected relationship in the present classification is with the geographical spread of site groups with the highest species poor site group 6. mean temperatures predicted for site groups 4 and The remaining two site groups are typical of the 6 which are found in the north east of the study vegetation types of the Irwin (Geraldton Sandplains) area and the lowest for site group 9 in the south district. Site group 8 are eucalypt woodlands and west of the study area (Figures 4 and 5). The tree heaths on red dunes immediately south of Shark Widespread site groups (1, 2 and 3) had the widest Bay while site group 9 are limestone heaths or range of predicted mean temperatures. The inland woodlands over limestone heaths between Kalbarri site groups had the highest diurnal temperature and Shark Bay (Figure 4). Species assemblage N is range and the highest annual temperature range most typical of site group 8, largely comprising taxa (Figure 5). of sandy habitats with a southwestern distribution. Most of the study area had highest rainfall in Average species richness for site group 8 was 30.0 winter, with the exception of sites from the taxa/plot. Species assemblages S and T are both Kennedy Range (site groups 6 and 4 in part) which highly faithful to site group 9. Both these showed significant rainfall during the warmest assemblages are largely southwestern taxa common quarter (Figure 5). Annual precipitation was on calcareous sands. This site group can be divided considerably higher in the area between Kalbarri into two subgroups - 9a occurring on massive and Shark Bay (site group 9) than in any other part limestones and 9b occurring on deeper sands over of the study area, with the Acacia shrublands (site

Table 4 Mean values of soil parameters of nine groups derived by classifying sites according to their vascular flora. Values were derived from a bulked soil sample taken from 0-10 cm layer. (S.D. indicates level of significance in difference between the means based on Kruskal - Wallis non parametric one way analysis of variance" = P< 0.05, .... = P

Site Groups 1 2 3 4 5 6 7 8 9 S.D.

EC (mS/m) 651.0 3.7 9.9 12.7 1.3 1.0 1.0 8.0 1.6 ...... Cl (%) 0.84 0.0 0.0 0.01 0.0 0.0 0.0 0.0 0.0 ...... pH (HP) 8.68 7.48 9.15 6.88 5.53 6.50 5.80 8.50 6.56 ...... Sand (%) 84.8 92.7 93.7 90.2 91.5 95.5 94.0 95.7 95.4 .... Silt (%) 6.3 1.5 1.5 3.0 1.0 0.7 0.5 0.5 0.6 ...... Clay (%) 8.9 5.8 4.7 6.8 7.5 3.8 5.5 3.8 4.0 .. C (%) 0.44 0.23 0.45 0.20 0.36 0.13 0.32 0.41 0.45 ...... Total N (%) 0.04 0.02 0.04 0.02 0.02 0.01 0.01 0.02 0.02 ...... Total P (ppm) 185.0 129.6 222.5 135.4 65.0 60.7 51.0 35.3 34.0 ...... P (HCO) (ppm) 15.0 8.3 10.5 6.3 1.8 2.0 1.0 2.0 1.2 ...... ) ...... K (HC03 (ppm) 345.0 135.8 57.5 167.3 20.0 39.3 14.0 29.0 23.4 ) ...... Ca (HC03 (ppm) 15.8 0.6 25.3 0.2 0.0 0.0 0.0 1.0 0.0 CEC (me%) 4.3 2.7 3.2 3.0 1.0 1.3 1.0 3.7 2.0 .... ex Ca (me%) 4.08 1.36 2.32 1.31 0.33 0.57 0.28 2.14 1.24 ...... ex Na (me%) 5.46 0.06 0.09 0.16 0.02 0.01 0.02 0.04 0.02 ...... ex K (me%) 0.59 0.28 0.11 0.34 0.04 0.08 0.03 0.04 0.05 ...... ex Mg (me%) 2.49 0.44 0.32 0.82 0.10 0.15 0.07 0.26 0.21 ...... Flora and vegetation 99 groups 2 and 4) having the lowest average rainfall Table 5 Difference in soil chemistry and soil (Figure 5). mechanical analysis between the subgroups in The nine site groups differed significantly in site group 2. Values were derived from a bulked soil sample taken from 0-10 cm layer. terms of soil chemistry and soil mechanical analysis. (S.D. indicates level of significance in Soils from site group 1 were the most markedly difference between the means based on different soils with very high levels of electrical Kruskal - Wallis non parametric one way conductivity, Cl, K, exchangeable Na, and analysis of variance ns not significant, * = P< exchangeable Mg (Table 4). Soils of site group 1 0.05, ** P100 cm) (Wyrwoll, ex Na (me%) 0.24 0.02 Stoneman, Elliott and Sandercock, 2000). Site group ex K (me%) 0.41 0.18 ** ex Mg (me%) 1.09 0.27 9 has a much smaller geographical spread than site 100 G.]. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge groups 2, 3 or 4 and while the subgroups showed survey, saltbush communities were only sampled at no significance difference in soil chemistry or soil Bush Bay (BB1) and Boolathana (B01). In both cases mechanical analysis, subgroup 9a occurred on these sites classified most similar to a vegetation shallow soil over limestone or in soils with unit found nearby based on shared saline species in significant limestone outcropping while subgroup one case (species assemblage 0 - BB1) and a suite of 9b occurred on deeper sands (Wyrwoll, Stoneman, annuals (species assemblage G - B01) in the other. Elliott and Sandercock, 2000). More sampling of this vegetation unit may have increased resolution of the current analysis. Due to the limited number of sites it was also DISCUSSION difficult to separate floristic variation due to regional environmental gradients from that Flora associated with local edaphic factors. This is Although the flora of the southern Carnarvon complicated in the present analysis by the Basin is still imperfectly recorded, the current list of correlation in regional climatic variation with 2133 taxa is a marked improvement on the previous distribution of major soil types. Within some site list for the entire Carnarvon Basin (an area of 91 046 groups for which it was possible to distinguish 2 km ) of 1348 taxa (Keighery and Gibson, 1993). This subgroups, it was found that geographical improvement occurred at all levels; for example segregation (and hence correlation with regional Kalbarri National Park had 206 taxa recorded by environmental gradients) was still apparent; Bums (1984), 717 taxa for the area by Bellairs and nonetheless correlations with edaphic factors were Bellairs (1992) and we recorded 1071 taxa from clearly demonstrated within at least three site within the boundaries of the National Park. groups. The study area has a predominantly arid flora of The climate of this region is unpredictable from southern affinities with an overlay of temperate and year to year in terms of both precipitation and tropical taxa at their range ends. The area is, timing of rainfall events. During the two years of however, species rich because it encompasses the the study, at least one good season was experienced change-over zone from the temperate to the arid at all plots except NE2. For this reason the annual zone. flora was included in the current analysis. However Species level conservation of the vascular flora little is known of the patterns of accumulation of shows that the reserve network in the area is annuals at sites over longer time frames or the heavily biased towards the southern and coastal influence of time of year of rainfall events on the margin of the zone. The wetland [40% of the 265 annual accumulation curves. Work of Zahran (1986) wetland taxa listed by Gibson, Keighery and Lyons and Loneragan et al. (1991) provide some evidence (2000) are not known from reserves in the Basin] of longer term climatic response by the herbaceous and red sandplain floras are particularly lacking flora. Long term regular sampling at permanently from the reserve network. located plots is needed quantify these processes in terms of changes in cover, changes in species Vegetation patterning occurrence and patterns of species accumulation In a regional survey of this type the degree of through time. Annuals form a very significant discrimination within and between site groups is proportion of the total flora of the region and limited by the number of plots established. In our therefore could be expected to be important in

Table 7 Percentage of taxa in each lifeform class by site group. Only taxa present in two or more sites are included. Percentage is based on total flora list recorded for each site group.

Life form Site Group 1 2 3 4 5 6 7 8 9

Annual herbs / geophytes 60 44 40 42 58 18 17 14 16 Annual graminoides 5 5 3 8 1 8 0 0 0

Perennial herbs 9 7 5 4 4 4 0 2 8 Graminoides 6 5 10 10 4 10 8 5 12 Shrub 19 37 42 35 27 55 58 70 60 Trees 0 2 0 1 5 4 17 9 3

Total number of species 78 227 158 153 73 49 12 56 98 Total number of quadrats 6 18 11 12 4 3 1 3 5 Flora and vegetation 101 defining the major cornpositional gradients across study area to be the most floristically complex with the region. seven of the nine site groups (78'X,) being recorded The analysis of the 63 primary sites shows an from the 17 southern most sites (28%), reflecting the essentially similar result to the pattern discerned major change-over zone between the southwestern from our subjective appraisal of the biogeographic flora and the flora of more arid areas. This affiliations of the vascular flora. The vegetation difference between the floras of the Geraldton patterning is closely correlated to major climatic Sandplain [Beard's (1990) Irwin district] and the gradients, as well as the distribution of different soil Carnarvon can also be seen in an analysis of the types across the study area. At a local scale edaphic proportional representation in lifeform class factors were correlated with vegetation patterning between the major classificatory groups (Table 7). but for the more geographically widespread groups Species rich shrublands are typical of the it was not possible to separate effects of regional southwestern flora and site groups 8 and 9 show climatic differences and regional soil differences the expected high percentage of shrubs compared from local edaphic factors. to annual herbs. Most of the Carnarvon groups (site The analysis showed the southern part of the groups 1 to 5) show the opposite trend with

Table 8 Comparison of nine major floristic site types identified in the present study with the land system and Pasture Types defined by Payne et al. (1987). Divisions in the table reflect Payne et al. (1987) higher vegetation - soil type groupings.

Pasture Type/ Site Groups Land System 1 2 3 4 5 6 7 8 9

Acacia and eucalypt short grass forb Nerren 4 Heath Nanga 3 3 Zuytdorp 2

Acacia sandplain Brown 4 Lyons 3 1 Peron 3 Sandplain 5 Yalbalgo 3 Yaringa Coastal dune shrub Coast 1 Edel 2 Lyell 1 Hard spinifex Kennedy 2 Soft spinfex Cardabia 4

Acacia mixed shrub Mary Toolonga 2 Acacia short grass forb Windalia 2 Stony short grass forb Billy

Mixed saltbush and bluebush Delta 1 Sable I Wrarroora 1 2 Samphire Birrida 1 McLeod 2

Not mapped 5 102 G.]. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge moderate to low percentage of shrubs and high The resulting mapping recognised 89 land systems percentage of herbs. Site group 6, the species poor in 17 vegetation alliances termed Pasture Types. dune vegetation of the top of the Kennedy Range Emphasis was given to perennial taxa important to and Mardathuna, do not follow this trend with a stock production in the definition of Pasture Types much higher than expected representation of (Payne et al., 1987). These Pasture Types were shrubs. Further sampling following good rains classified into five broad groupings based on would be needed to confirm this finding. affinity of species present (Southwestern vs. Site group 7 consisted of only a single site (NE2) Eremaean) and broad soil type. Our plots covered from which only 12 perennial taxa were recorded 23 of these land systems in 11 Pasture Types in four over the two major sampling periods. At site NE1 of the broad vegetation - soil groupings (Table 8). only 3 km distant large numbers of annuals were At the land system level there was only moderate recorded in 1996 as with the other Nerren Nerren correlation between the 23 land systems and the sites located some distance to the east. The soil nine major floristic site groups. The nine site groups chemistry, soil mechanical analysis and soil profile were represented by between one and seven land data (Wyrwoll, Stoneman, Elliott and Sandercock, systems. Better correlation was evident between the 2000) were very similar between all Nerren Nerren floristic site groups and the major Pasture Types, sites. It is not clear if NE2 represents a local with the nine site groups represented by between vegetation type that is naturally poor in annual one and four Pasture Types. The correlation did not species or had not received the same degree of improve at the broadest vegetation - soil groupings rainfall as nearby sites. (Table 8). This lack of correlation between the two The floristic classification shows broad general studies relates to different aims and methodology agreement with the biogeographical boundaries of used. The rangelands study used perennial taxa to Beard (1990) with the tree heath on red sandplain help define mapped land units, while the present south of Shark Bay and the coastal limestone study used analysis of all taxa to define major woodlands and heaths (site groups 8 and 9) falling floristic units across the study area. A reanalysis of within the Irwin district and the saline group, the the 63 sites using only perennial taxa could be Acacia shrublands, the coastal Acacia and Melaleuca expected to be more closely correlated with the shrublands, the Mulga shrublands, and the red sand rangeland classification. dune vegetation groups of the Toolonga Plateau and the Kennedy Range falling in the Camarvon district. The present classification does not support ACKNOWLEDGEMENTS the inclusion of Edel Land as part of the Irwin Funding was provided by the Commonwealth district (renamed Geraldton Sandplain) as recently through the National Reserves System Co-operative suggested by Thackway and Cresswell (1995). Program of the Australian Nature Conservation Our analysis clearly shows the relationships Agency (now Environment Australia), together between Edel Land and Peron Peninsular with the with State funds provided by the Western rest of the coastal belt of the Camarvon district in Australian Department of Conservation and Land site group 3. The Edel Land sites are separated as a Management. Two referees provided valuable subgroup, with this subgroup being most similar to comment on an earlier draft of this paper. the Peron subgroup. Our present data are not detailed enough to assess if Beard's phytogeo­ graphic boundaries are optimally placed (but see Gibson, Burbidge, Keighery and Lyons, 2000). REFERENCES We found no strong correlation between Alford J.J., Keighery, G.J. and Trudgen, M.E. (2000). vegetation patteming and the broadly mapped Floras of the 29 islands of the Freycinet Estuary, surficial geology (Hocking, 1985). All sites occurred Shark Bay World Heritage Area, Western Australia. CALMScience (in press). on Pliocene to Holocene aged alluvial or colluvial deposits or aeolian sands except the lower slopes of Anonymous (1997). Shark Bay Regional Strategy. Western Australian Planning Commission, Perth. the Kennedy Range (KE3-5 which occurred on early Austin, M.P. and Belbin, L. (1982). A new approach to Permian to early Cretaceous sediments) and one site the species classification problems in floristic on lower slopes of the Pell Range (GJ1 which analysis. Australian Journal of Ecology 7: 75-89. occurred on Permian sediments). Beard, J5. (1975). The Vegetation of the Pilbara Area, 1; The only consistent land system mapping that has 1,000,000 map and explanatory notes. University of been undertaken across the study area is that Western Australia Press, Nedlands. produced by the Department of Agriculture Beard, J.5. (l976a). The Vegetation of the Murchison area, 1; rangelands survey group (Payne et al., 1987). This 1,000,000 map and explanatory notes. University of classification was based on 669 inventory sites Western Australia Press, Nedlands. sampled for perennial taxa and soil parameters in Beard, J5. (1976b). The Vegetation of the Ajana Area, conjunction with aerial photograph interpretation. Western Australia. Vegmap Publications, Perth. Flora and vegetation 103

Beard, J5. (1976c). The Vegetation of the Shark Bay and Edel BR. Wilson (eds), Research in Shark Bay, pp. 61-88. Areas, Western Australia. Vegmap Publications, Perth. Western Australian Museum, Perth. Beard, J5. (1990). Plant Life of Western Australia. Kangaroo Keighery, G.J. (1995). How many weeds? An annotated Press, Kenthurst. list of the naturalised vascular plants of Western Bellairs, D.R. and Bellairs, B. (1992). Checklist of Plants for Australia. In G. Burke (ed), Invasive Weeds and Kalbarri and the Murclllson Sandplalll, (with supplement). Regenerating Ecosystems in Western Australia: Kalflora, Kalbarri. Proceedings cif' the Conference held at Murdoch University, Burbidge, A.A, Fuller, P.J. and Cashin, K. (1980). The July 1994. Institute for Science and Technology Policy, wildlife of the proposed Toolonga Nature Reserve, Murdoch University, Perth. Shark Bay Shire, Western Australia. Dept. Fisheries and Keighery, G.J. (1996). Phytogeography, biology and Wildlife Report 39. conservation of Western Australian Epacridaceae. Burbidge, A.A and George, AS. (1978). The Flora and Annals of Botany 77: 347-355. fauna of Dirk Hartog Island, Western Australia. Keighery, G.J. and Gibson, N. (1993). Biogeography and Journal of the Royal Society of Western Australia 60: 71­ Composition of the flora of Cape Range peninsula, 90. Western Australia. Records of the Western Australian Burbidge, AI-I., McKenzie, N.L. and Harvey, MS. (2000). Museum 45: 51-85. A biogeographic survey of the southern Camarvon Kenneally, K.F. (1978). Notes on the vegetation and flora Basin, Western Australia: background and methods. of Rocky Pool, Gascoyne River. Western Australian Records of the Western Australian Museum Supplement Herbarium Research Notes 1: 29-39. No. 61: 1-12. Loneragan, W.A., Holm, A.McR, Watson, I.W. (1991) Bums, AC. (1984). List of Plants: Kalbarri National Park. Assessing long-term changes in native rangeland National Parks Authority, Perth. vegetation using functional species groups and CALM (1998). Declared Rare and Priority flora list - 21/10/ multivariate analysis. Proceedings IVth International 1998. Unpublished Report, CALM, Perth. Rangeland Congress. April 22-26, 1991, Montpellier. Carolin, RC., Rajput, M.J.M. and Morrison, D. (1992). McArthur, W.M. (1991). Reference Soils of South-western Goodeniaceae. In AS. George (ed) Flora of Australia, Australia. Department of Agriculture, Perth. Volume 35. Bnllloniaceae, Goodeniaceae, pp. 4-334. McMahon, J.P., Hutchinson, M.F., Nix, H.A. and Ord, Bureau of Flora and Fauna and Australian K.D. (1995). ANUCLIM Users Guide, Version 1. Government Publishing Service, Canberra. Centre for Resource and Environmental Studies, George, AS (1999). William Dampier in New Holland: Australian National University, Canberra, Australia. Australia's First Natural Historian. Bloomings Books, Newbey, K.R. and Newbey, B.J. (1992). Notes on the Hawthorn, Victoria. Vegetation, Flora and Birds ofParts ofthe Kennedy Range, Gibson, N., Keighery, G.J. and Keighery, B.J. (1997). Gascoyne District, Western Australia. Unpublished Contributions of N.H. Speck to the biogeography of report to CALM, Perth. Proteaceae in Western Australia. Journal of the Royal Payne, A.L., Curry, P.J. and Spencer, G.F. (1987). An Society ofWestern Australia 80: 73-77. inventory and condition survey of rangelands in the Gibson, N., Burbidge, AH., Keighery, G.J. and Lyons, Carnarvon Basin, Western Australia. Department of M.N. (2000). The temperate to arid transition of the Agriculture Western Australia Technical Bulletin No. 73. Irwin - Carnarvon phytogeographic boundary, Royce, RD. (1962). Botany .In A.J. Fraser (ed), The results Western Australia. Records of the Western Australian of an expedition to Bernier and Dorre Islands, Shark Museum Supplement No. 61: 155-173. Bay, Western Australia in July 1959. Fauna Bulletin of Gibson, N. Keighery, G.J. and Lyons, M. (2000). The flora Fisheries Department, Western Australia 2: 31-53. and vegetation of the seasonal and perennial Siege!, S. (1956). Non Parametric Statistics for Behavioural wetlands of the southern Carnarvon Basin, Western Sciences. McGraw-Hill, New York. Australia. Records of the Western Australian MusCllm Sneath, P.H.A. and Sokal, R.R. (1973). Numerical Supplement No. 61: 175-199. Taxonomy: The Principles and Practice of Numerical Green, J.W. (1985). Census of the Vascular Plants of Western Classification. Freeman, San Francisco. Australia. Department of Agriculture, Perth. Thackway, R. and Cresswell, I.D. (1995). (eds) An interim Hocking, RM. (1985). Geology of the Carnarvon Basin. biogeographical regionalisation for Australia: a framework Geological Survey of Western Australia Bulletin No. 133. for establishing the natIOnal system of reserves, version Hopper, S.D. and Maslin, B.R (1978). Phytogeography of 4.0. Australian Nature Conservation Agency, Acacia in Western Australia. Australian Journal of Canberra. Botany 26: 63-78. Trudgen, M.E. and Keighery, G.J. (1995). Flora of the Shark Hussey, B.M.J., Keighery, G.J, Cousens, RD, Dodd, J. Bay World Heritage Area and Environs. Report prepared and Uoyd, S.G. (1997). Western Weeds: a Guide to the by Western Australian Department of Conservation Weeds of Western Australia. Plant Protection Society of and Land Management for the Australian Heritage Western Australia, Victoria Park, W.A Commission. Keighery, GJ. (1984). Phytogeography of the Tyler, J.P. (1988). Vegetation surveys near Lake MacLeod. Monocotyledons of Western Australia. Kings Park KlIIgia 1: 49-74. Research Notes 8: 39-67. Wilson, P.G. (1984). Chenopodiaceae. In A.5. George (ed) Keighery, G.J. (1990). Vegetation and flora of Shark Bay, Flora of Australia, Volume 4. Phytoloccaceae to Western Australia. In P.F. Berry, S.D. Bradshaw and Chenopodlcaeae, pp. 81-317. Bureau of Flora and Fauna 104 G.}. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge

and Australian Government Publishing Service, Zahran, B.H.A. (1986) Forage composition and production of Canberra. annual and biennial species on contrasting soils in Wyrwoll, K.-H., Courtney, J., and Sandercock, P. (2000) rangeland grazed by sheep near Carnarvon, Western The climatic environment of the Carnarvon Basin, Australia. MSc (Agriculture) Thesis, University of Western Australia. Records of the Western Australian Western Australia. Museum Supplement No. 61: 13-27. Wyrwoll, K.-H., Stoneman, T., Elliott, G. and Sandercock, P. (2000) The geo-ecological setting of the Carnarvon Manuscript received 4 February 2000; accepted 14 April Basin, Western Australia: geology, geomorphology 2000. and soils of selected sites. Records of the Western Australian Museum Supplement No. 61: 29-75. Flora and vegetation 105

APPENDIX 1

Vascular plants recorded for study area, their CALM 3. Priority Three Poorly Known Taxa conservation codes and occurrence in major conservation Taxa which are known from several populations, at reserves. least some of which are not believed to be under Explanation of codes: immediate threat (ie. not currently endangered). Conservation code [as defined by the Western Australian Such taxa are under consideration for declaration as Department of Conservation and land Management 'rare flora', but are in need of further survey. (1998)]. 4. Priority Four - Poorly Known Taxa R. Declared Rare Flora - Extant taxa (= Threatened Taxa which are considered to have been adequately Flora Endangered + Vulnerable) surveyed and which, whilst being rare (in Taxa which have been adequately searched for, and Australia), are not currently threatened by are deemed to be in the wild either rare, in danger identifiable factors. These taxa require monitoring of extinction, or otherwise in need of special every 5 -10 years. protection, and have been gazetted as such, following approval by the Minister for the Environment, after recommendation by the State's Distribution Endangered Flora Consultative Committee. Distributions refer to named places, pastoral leases, and conservation reserves (see below and Figure 1). Range 1. Priority 1 - Poorly Known Taxa limits refer to populations occurring at the limit of their known geographic range at or within the named place, Taxa which are known from one or a few (generally pastoral lease or conservation reserve. < 5) populations which are under threat, either due to small population size, or being on lands under VCl Vacant Crown land immediate threat, eg. road verges, urban areas, WHA Shark Bay World Heritage Area farmland, active mineral leases, etc., or the plants are under threat from disease, grazing by feral Conservation Reserve Codes animals etc. May include taxa with threatened BINR Bemier Island Nature Reserve populations on protected lands. Such taxa are under DINR Dorre Island Nature Reserve consideration for declaration as 'rare flora', but are CNR Cooloomia Nature Reserve in urgent need for further survey. FPNP Francois Peron National Park 2. Priority Two - Poorly Known Taxa KNP Kalbarri National Park KRNP Kennedy Range National Park Taxa which are currently known from one or a few TNR Toolonga Nature Reserve (generally < 5) populations, at least some of which ZNP Zuytdorp National Park are believed to be under immediate threat (ie. not currently endangered). Such taxa are under consideration for declaration as 'rare flora', but are in urgent need of further survey. Familyrraxon Conservation Distribution Conservation Reserve Code KNP TNR ZNP CNR FPNP BINR DlNR KRNP

Acanthaceae Dipteracanthus australasicus subsp. australasicus Harnieria kempeana Hypoestes floribunda

Adiantaceae Anogramma leptophylla Cheilanthes austrotenuijo/ia Chei/anthes lasiophylla Cheilanthes sieberi

Agavaceae * Agave americana

Aizoaceae * Carpobrotus aequilaterus Carpobrotus sp. (GJK & NG 1707) Carpobrotus virescens Disphyma crassifo/ium subsp. clavellarum Gunniopsis rodwayi Gunniopsis septifraga * Mesembryanthemum crystallinum Sesuvium portulacastrum 5 limit FPNP Tetragonia coronaw Endemic WHA Tetragonia cristata * Tetragonia decumbens Tetragonia diptera Tetragonia implexicoma Tetragonia tetragonoides Trialllhema oxycalyptra var. oxycalyptra 5 limit Nerren Nerren 5tn. Trialllhema pi/osa Trianthema triquetra Trianthema turgidifo/ia Zaleya galericulara

Amaranthaceae Alternanthera nodi/lora * Alternanthera pungens Amaranthus clementii Amaranthus mitchellii Al1Ulrllllthus pallidi/lorus Gomphrena canescens Gomphrena cunninghamii Hemichroa diandra Pti/otus aervoides N limitZNP Pti/otus aff. chamaecladus Pti/otus aff. grandi/lorus Ptilotus al,'xandri 2 Endemic DHI Ptilotus arthrolasius Ptilotus astrolasills Ptilotus atnp!icifil!ius Ptilotus heardli Ptilotus culostachvus var calostuchyus Ptilotus caPitlllus Ptilotus ciwmaecladus Ptilotus divaricatus vaL diVilncatus Ptilotus divariclllus vaT. rubescens Endemic WHA (Edel Land - Tamala Stn.) Ptilotus drummondli var drummondii Ptilotus drummondii var drummondii N limitZNP Plilows drurnmondii var muwr N limit FPNP Ptilotus erwtrichus Ptilotus cxaltatus var exaltatus Ptilotus Rardneri Ptilotus RlUidiciwudli vaT. Raudichaudii Ptilotus Raudichaudii var. parViflorus N limit FPNP Ptilotus JilWliiiflorus var RYimdiflorus N limit WHA Ptilotus Rraruliflorus vaT. lepidus Endemic Peran Peninsular Ptilotus helipteroi/ies Ptilotus humilis subsp. humilis Ptilotus latifiJiius Ptilotus macrocephalus Ptilotus flwnRlesii Ptilotus murrayi Ptilotus ohovlllus var. lancifiJiius Endemic WHA Ptilotus obOVIlIUS VaT. obovatus Ptilotus petiolatus Ptilotus polystachyus forma polyslachyus Ptilotus roei Ptilotus rotundifi!!ius Ptilotus schwartm var ReorRei Plilo[us sc!n-vartzii var sc}nvartzii Ptilotus spathulatus Ptilotus StipilatuS Ptilotus stirlinRh vaT. pumilus Endemic WHA Ptilotus stirlinRii var. stir!ingh Ptilotus villosif/orus

Amaryllidaceae Crinum flaceidum S limit Lake MaCleod

Anthericaceae Amocrinum drummondii 3 N limit ZNP Borya scirpoidea Borya spiwerocephala N limit KNP Caesia alfimlii ms N limit KNP Chamaeseilla ,·orymhoS([ Corynotheca mierantha vaT. acunthoclada N limit Carrang Stn. ... Familyffaxon Conservation Distribution Conservation Reserve o Code KNP TNR ZNP CNR FPNP BINR DINR KRNP <»

Corynotheca micrantha var. micrantha Corynotheca pungens S limil Tamala SIn. Dichopogon preissii N limil KNP Dichopogon tyleri Endemic to Sludy Area Laxmannia sessilif/ora subsp. sessi!iflora N limitCNR Murchisonia fragrans 2 N range limit KNPrrNR Murchisonia volubilis Sowerbaea laxiflora Thysanotus arenarius N limit Tamala SIn. Thysanotus brittanii ms Endemic KNP Thysanotus kalbarriensis ms 2 Thysanotus manglesianus N limit Nanga Stn. Thysanotus patersonii Thysanotus ramulosus Thysanotus sparteus N limit Carrang SIn. Thysanotus speckii Thysanotus teretifo!ius Nlimit KNP Thysanotus thyrsoideus Tricoryne aff. corynothecoides (GJK & NG 1274) Tricoryne arenicola ms Tricoryne corynothecoides Tricoryne elatior N limit Tamala Stn.

Apiaceae Actinotus leucocephalus Apium annuum NlimitKNP Apium prostratum * Cyclospermum leptophyllum Daucus glochidiatus Hydrocotyle pi!ifera var. glabrata Hydrocotyle rugulosa Hydrocotyle scutellifera Neosciadium glochidiatum N limit FPNP Platysace commutata Platysace juncea N limit KNP Platysace sp. Kalbarri (0.& B.Bellairs 1383) 2 Endemic KNP Platysace sp. Kennedy (P.G.Wilson 8450) 2 Endemic KRNP Platysace xerophila N limitKNP Trachymene aff. oleracea (GJK & NG 719) Confined KRNP Trachymene coerulea var. leucopetala NlimitKNP Trachymene cyanopetala Trachymene elachocurpa Trachymene glaucifolia Trachymene ornata N limit Cobum Stn. Trachymene pi!osa N limitWHA Uldinia ceratocarpa N limit FPNP Apocynaceae Alyxw hlln/olw N limit Nanga Stn * CUl/wraf/thus roseus

Arecaceae * PhOelll,1 doctylI/em

Asclepiadaceae C'ytlotlchllm aff /lonlJUndllm (KRN 11847) CVtlatlChll1ll flilrlhllndllm Gymnanfhera cUflninghamii S limit from (Cape Range) G.Ymnema grtmiriul rns N limit FPNP lWarsdenia australis A1arsdenia granttfco/a MlIrsdellw vlndt/lom Rh,"clwrrhe"a IIllearls Sarcostemnw vlminale subsp austrule

Asphodelaceae * ASfihodcllls (ISlllloslIs BlIllnlle ,'emdJilrbof(/

Aspleniateae

Pleurosorus rut~I(}lius

Asteraceae Acttn()!J()!e COf1dervd.ltum N limit DHUFPNP Actifl()/)(Jle drurnnumdiana S limit Hamelin Stn, Actlnobolc IIhgilwslIm Angiaflthus acrohva!inus S limit Hamelin Stn, Angianthus ('unninglwfnfi Anglalllhus mlcrI!cepfwllls 2 AnUiu!1[hus mifnei Angianthus prelSSUlflUS Angwnlhus pusWus Angwlllhlls sp, (A,S,George 11362) Endemic WHA Angwlllhus sp, (A,S,George 11471) Endemic WHA Angian!hus {ornenlosus N limit DHI * Arelolhcca calendlllo ArClolheca pOplllU/ilIo Aster suhulolllS ASlerldeu aslermdes N limit KNP Belllda gram/nco * Bldells hlplmulIo Blenfl{}-"'{J0ra drurnnI{Jndii Brachvscome chelll!I'orp" Brachysi'()me (";ffaris Brach)'s('olne ciliocorpa Brachyscomc fwllJphtla Brachvscorne iberidi(o/w S limit ZNP .... Farnilyrraxon Conservation Distribution Conservation Reserve .... o Code KNP TNR ZNP CNR FPNP BINR DINR KRNP

Brachyscome latisquamea Brachyscome lineariloba Brachyscome perpusilla Calocephalus aervoides 3 Calocephalus francisii Calocephalus knappii Calocephalus multiflorus Calocephalus sp. Pilbara (MET 11454) Calocephalus sp. Wiltenoom (A.S.George 1082) Caloti.1 hispidula Caloti.1 multicaulis * Centaurea melitensis Celllipeda cunninghamii Centipeda minima Centipeda thespidioides Cephalipterum drummondii Ceratogyne obionoides Chondropyxis halophila Chrysocephalum apiculatum ChrysoceplUllum semicalvum Chthonocephlllus muellerianus 2 Chthonocephalus oldfieldianus I Endemic Carnarvon to Shark Bay ChtlulIlocephalus pseudevax Chthonocephalus spathulatus I Chthonocepmllus tomelllellus 2 * ColUla bipinnata ColUla coronopifolia N limit KNP Cotula cotuloides N limitDHI ColUla sp. Kalbarri (GJK & NG 2044) Cratystylis subspinescens Decazesia hecatocephnla Erymophyllum compactum Erymophyllum ramosum Erymophyllum ramosum subsp. involucratum Erymophyllum ramosum subsp. ramosum Erymophyllum tenellum Gilberta tenuif{!lia N limit Coburn Stn. Gilruthia osbornei * Gnaphalium polycaulon S limit D1NR Gnephosis acicularis Gnephosis angianthoides Gnephosis arachnoidea Gnephosis brevifolia Gnephosis eriocephala Gnephosis gynotricha Gnephosis macrocephala Gnephosis sp. Billabong (Nordenstam/Anderberg 203) Endemic study area Gnep/wsis lenuissima Haplolriclllon COIWJllli Endemic southern Carnarvon Basin Haplolrichion conicum Hedypnois rha[{w.iio/ilides * Helianlhus Wlnuus Heliplerum cmspcdlOides Hyalosperma co[ula Hyalospemw demlssum Hvalosperma [{IUllflOSum subsp [{IUllllOSum Hyal()spernw glutin{}slurl subsp venus!um N limit Tamala Stn • H\lwclwens [{Iabm Isoelopsi.\' gmmlfll!olw * I..,,(JCfUCl1 serrio/a Lagem/era hue[{elll L(Hvrencella d(rvenportii N limit Hamelin Stn. Lawrence/la rrJsca N limit Tamala Stn. LeuuJcht)'sum fitzgihb(mii Leucophyla broWflll Millolia depauper(J{a Mlllotia jaCkWlIll 2 Millolia myosolidtjillw Ivfinuria integerrinw Mlfluna leplophvlla lvlyriocepfwlu.\' gascoyncnsis Ins Endemic Carnarvon Basin Myrioceplwlu.\' guenllae Afynocephalus nudus Myriocephalus pyw,weus Myrioccphalus sp. (Demarz 12152) Endemic Carnarvon Basin MyrlOccp/wlus sp. (Nordenstam &Anderberg 188) Endemic WHA Olearia adenolasia Olearia affaxillaris (GJKJNG 1025) Endemic WHA ()fearia axillaris Olearia dampien subsp. dampieri ms Olearia dampieri subsp. eremicola ms Olearia humitis Olearia imbricula Olearia mue/leri O/earia occidentissinw 2 Endemic WHA Olearia plme/eoides Olearia p/ucheacca Oleana revoluw Olearia rudis Olearia sWarlii * Ostcospennurn co[cndu!aceum P/uchea rubelll!7ora P/uchea s'luarmsa Pluchea lelmnlhera S limit Wooramel R. Podolepis aunculaw S limit Wooramel R. Podolepis wnescells N limit Giralia Podolepi.\' wpillans .... Familyrraxon Conservation Distribution Conservation Reserve .... Code KNP TNR ZNP CNR FPNP BINR DINR KRNP N

Podolepis gardneri Podolepis lessonii Podolepis microcephala S limit Tamala Stn. Podotheca angustifolia N limit DHI Podotheca gnaphalioides N limit Woodleigh Stn. Podotheca pollackii Podotheca pygmaea Pogonolepis muelleriana Pogonolepis stricta * Pseudognaphalium luteoalbum Pterocaulon sphacelatum Pterodweta paniculata * Reichardia tingitana Rhodanthe charsleyae Rhodanthe chloroceplwla subsp. splendida Rhodanthe citrina Rhodanthe condensata N limit KRNP Rhodanthe corymbosa Rhodanthe cremea Endemic WHA Rhodanthe humboldtiana Rhodanthe laevis Rhodanthe "umglesii Rhodanthe margarethae Rhodanthe maryonii Rhodanthe oppositifolia subsp. oppositifolia NlimitDHI Rhodanthe oppositifolia subsp. ornata 2 Endemic WHA Rhodanthe polycephala Rhodanthe psammophila Rhodanthe rubella Rhodanthe sp. Overlander (P.S.Short 2096) Endemic WHA Rhodanthe spicata Rhodanthe sterilescens Rhodanthe stricta Schoenia ayersii Sclwenia cassiniana Schoenia jilifolia subsp. arenicola Senecio glossanthus Senecio gregorii Senecio lautus subsp. dissectifolius Senecio lautus subsp. maritimus * Sonchus oleraceus * Sonchus tenerrimus Sondottia glabrata 2 Endemic WHA Streptoglossa cylindriceps Streptoglossa liatroides Streptoglossa tenuiflora Trichanthodium scarlettianum ...------Triehantlll}(hlll/l skirrophorum '* Urosperrnum picroides * Ursinia (Jf/lhemoides * Vel!crcopll)'1on dcalbarllm Vittudinia cervicularis var cervicularis N limitDHI Vifflldil11ll CC/Tlellians var. old/ieldii N limit Kalbarri area Vittadinia erel11aea Waltzia ucurnina!u var. ucununala

~V(1itziaacuminata var albicans N limit Hamelin Stn. WWIZUl corvmbosll Wllmill /und" N Iil11itZNP Wwtzw podolcplS

~Vaif::.iusuav('o!ens

Aviccnniaccac .Avicennia matlna

Boragina('cac CynoghJSsum australe * Echlwn plalllllgil/cum Hlllglll/w afT. Ilrgyrophyl!a Halgal/w afT Iworalts Hlllgal/ia al/agal!oides var. wwglll!oides I11S N limit KNP Ha{gal/ia al/aglll!oides var preiSSlww ms Ha{gal/ia argvrophvl!a Ha{gal/la be!Jrww N limit Tamala Stn. Ha!t(unia C}'llIU'(J HIl{gal/w cyal/ca var {Illisepalll ms S limit FPNP Ha{gwlia guslatvel/it var. compactus I11S Ha{gwua {,fforalts Ha{ganla senu/{ora Ha/gonia VlSCOS(i Helio[ropiulIl curassaVlCUfrI He{wtrol'ium heleral/thum Heliotropium panicu{alUm Heltotrol'ium andu{alum Omp{w{,J{apl'u{a conCIlHl P{llgiobothrys p{urisepa{us Disjunct from Mt Jackson Trichodesma zey{wllcum var. zey{anicum

Brassicaccac Brassica juncca Brassica {ournefortii CakUe maritima C"roI/OpUS didym/ls D'l'{otaxis muralis Dip{otaxis tenutfldia Hymel/o{o!J/lS procumbens N limit DINR Lel'ldium !Jiplicat/lm Lel'idium jlJlios/lm Familyffaxon Conservation Distribution Conservation Reserve Code KNP TNR ZNP CNR FPNP BINR DINR KRNP

Lepidium linitiJlium Lepidium lyratogynum Lepidium oxytrichum Lepidium phlebopetalum Lepidium platypetalum Lepidium pseudoruderale Lepidium puberulum 4 N limit D1NR Lepidium rotundum Lepidium scandens Menkea australis Menkea villosula Phlegmatospermum drummondii 3 Range extension from Wongan Hills * Raphanus raphanistrum * Raphanus sativus * Sisymbrium erysimoides * Sisymbrium irio * Sisymbrium orientale Stenopetalum anfractum Stenopetalum filifolium N limit Cobum Stn. Stenopetalum linea re Stenopetalum pedicel/are Stenopetalum robustum Stenopetalum sphaerocarpum NlimitDHI

Caesalpiniaceae Labichea cassioides Labichea eremaea Labichea lanceolata subsp. lanceolata Labichea teretijiJlia subsp. teretifolia N limit Cobum Stn. Petalostylis cassioides Petalostylis labicheoides Senna artemisioides subsp. filitiJlia Senna artemisioides subsp. helmsii Senna artemisioides subsp. x sturtii Senna glutinosa subsp. charlesiana Senna glutinosa subsp. chatelainiana Senna glutinosa subsp. x luerssenii Senna notabilis Senna pleurocarpa var. pleurocarpa Senna sp. Austin (AStrid 20210) Senna sp. BilIabong (J.D.Alonzo 721) Senna sp. Meekatharra (E.Bailey 1-26) Senna sp. Tamala (M.ETrudgen 7334A) Endemic WHA

Callitrichaceae * Callitriche stagnalis Campanulaccac WahlcnberlilU capensls

~V(Jhlef1herRi(JcornmuntS WahlenberlilU limnlelllll N limit DHI Wahlenberliw limcilts Wahlcnberlilll mullluw!L, \Vdhlenbergi(l prelssll Wahlenberli/a III mIIJ!frucia

Capparaceae Cappans laslIlnlllU Cappdris Splt/flStl var numfnu!arlcl Cleome unlllem ('{come viscosa

Caryophyllaceac Cerasfiurn g/ornCUl1um C'orrigiolu Iitora!Js Gvpsophila auslmlis Petrorhagiu velutilw Polycarpaea ("or.ym!}()S(J Pul)'l'arpull tetmphvllum Saginu opera/a SlIene lia/hea var RaWca

..')'i/enc !l{)Cfurna SpClJ

Casuarinaceae Al/uC(Jsuamw acullm/v/s subsp aeullva!\,/s N limitZNP Allocasuar/rl(l campestns AI/(!C(Js/lI1riml cam/I'u/ala N limit Nerren Nerren Stn, Al/oCl/suarilla dielsiarw Al/oUlsuarina helmsi/ N limit DHI Al/oCl/suarinll huelic/iww Al/oCl!suarirw humihs

Alloc(lsuarif1l1 lehnwnniana N limit Tamala St~, Casuarint1 {fhesa

Celastraceae PSallUn()tf/(}.va (~h()relr()ides N limit KNP PsanlnUJllUJya ephednJides

Centrolepidaccae Aphelia nuluns N limit KNP Cenfrolepis arisfuI(J N limit KNP Centro!epis ceplwlolormi:." subsp. rnurrayi N limit Kalbani Area Familyffaxon Conservation Distribution Conservation Reserve Code KNP TNR ZNP CNR FPNP BINR DINR KRNP

Centrolepis drummondiana Centrolepis eremica Centrolepis glabra N limit KNP Centrolepis humillima NlimitDHI Cemrolepis polygyna

Chenopodiaceae Atriplex amnicola Atriplex bunburyana Atriplex cinerea Atriplex codonocarpa Atriplex IlOlocarpa Atriplex isatidea Atriplex lindleyi subsp. in/lata Atriplex nummularia subsp. spathulata Atriplex paludosa subsp. baudinii Atriplex paludosa subsp. moquiniana Atriplex semilunaris Atriplex spongiosa Atriplex vesicaria subsp. incompta Atriplex vesicaria subsp. variabilis * Chenopodium ambrosioides Chenopodium auricomum Chenopodium cristatum Chenopodium curvispicatum Chenopodium desertorum Chenopodium gaudichaudianum Chenopodium melanocarpum forma leucocarpum Chenopodium melanocarpum forma melanocarpum * Chenopodium murale Chenopodium pumilio Didymanthus roei N limit FPNP Dissocarpus paradoxus Dysplumia glandulosa Dysplumia glomuli{era subsp. eremaea Dysplumia kalpari Dysph(mia plamaginella Dysphimia platycarpa Dysphania rhadinostachya subsp. in/lata Dysplumia rhadinostachya subsp. rhadinostachya Dysph(mia sphaerosperma S limit Tamala Stn. Einadia nutans Enehylaena tomentosa var. tomentosa Eremophea aggregata Eriochiton sc!erolaenoides Halosarcia aurieulata Halosarcia doleijormis Ha/omrcia /w/ocnemoides subsp. /llI/ocnemoides Ha/osarcia ha/ocnemoides subsp. lenuls S limit Nanga Stn. Ha/omnia ""ilca subsp. hldens Ha/osarcia Indu(i subsp /eiostachya Ha/osarcw /eploc/ada Ha/osw'C/a pe/llIw Ha/osarcla per/iWmililla subsp. per/iWnu/ala Ha!osarcia prulflosa Ha/osarcia pteryg()spernw subsp den/feu/ora S limit Nanga Stn Halosarcia pteryguspenna suhsp pter.vgospernw N limit Nanga Stn lv/aireat/a (illU!(:'fW Mwreana aphylla /vlaircaf/a appressa :\4aircatla afkin.\'iarw Nlalreana cunl0Sd A-1airca!la CO!li-'exa Mairca!la gcorget Maircana lfltegra !rfaircat1a lanosa Afillreww /o/n/low lv!airca!1a me/a!locoma Malreww OP!Wslll/olla AJuireana pen[0J.,;(J!1{1 MiIlreana p/am/o/w Malreana p/alvcar!'a A1aircollo P()!)'Ptcr)\t;iil Maircana sc!eroplera Mlllreana sll/)f[ala S limit Tamala Stn. Ataircat1l1 thesioides Mairea!la tomentos'u lVfaireana tric!l(Jptera Afmreana trlpfera Alaireoflo rillosa Neohassia asfroc'arpa S limit BINR Ostco(arpum acropterurn var. acrn!Jterum Rha/iodia hdccald Rha/iodw crassl/o/w Rhag(hiia drurnm(}fuhi Rhagodia erenwea R/w/iodl([ /all/o/w subsp /all/o/I([ R/w/iodw /all/olld subsp recto R/wgodlil preissit subsp. obo\'([[(J Sa/so/a ka/I 5'arcocorniu bllu'kitlfW Sarcllcllmla qllll/que/lout Sclem/ael/a blcusl'is Disjunct from South Australia Sc!errJ!ocfw o}stota Sc/emlaefla cUI/ea la Sc/em/aerw derw/lout Sclemlaena desertlclI/a .... Familyrraxon Conservation Distribution Conservation Reserve .... QC Code KNP TNR ZNP CNR FPNP BINR DINR KRNP

Sclerolaena diacantha Sclerolaena eriacantha Sclerolaena eurotioides Sclerolaena j(Jrrestiana Sclerolaena limbata Sclerolaena medicaginoides Sclerolaena obliquicuspis Sclerolaena recurvicuspis Sclerolaena stylosa Sclerolaena tridens Sclerolaena uniflora Sclerostegia disarticulata Suaeda australis Tecticornia arborea Tecticornia verrucosa Threlkeldia diffusa

Chloanthaceae Cyanostegia angustifolia Dicrastylis fulva N limit Nerren Nerren Stn. Dicrastylis linearijiJlia 3 N limit Meadow Stn. Dicrastylis micrantha N limit Nanga Stn. Dicrastylis sp. Peron Peninsula (M.E.Trudgen 7373) Endemic WHA Dicrastylis sp. Shark Bay (J.J.Alford 1318) Lachnostachys eriobotrya Lachnostachys ferruginea Physopsis chrysophy/la 3 Endemic WHA & S to Murchison R. Pityrodia atriplicina NlimitDHI Pityrodia axillaris Pityrodia cuneata N limit Peron Peninsular / DHI Pityrodia glabra Endemic WHA Pityrodia glutinosa 2 Endemic Nerren Nerren Stn. area Pityrodia hemigenioides N limit Eurardy Stn. Pityrodia loxocarpa Pityrodia oldfieldii N limit Tamala Stn. Pityrodia paniculata S limit Cobum Stn. Pityrodia teckiana Pityrodia verbascina Spartothamne/la teucriif/ora

Clusiaceae Hypericum gramineum Hypericum japonicum

Colchicaceae Burchardia multiflora Burchilrdia rosea Endemic Kalbarri Area Wunnbea cernua lVurmhea dellS/flom N limit KNP ~Vurrnbcl1,Jp ",'rrn,l" \Vurmhea dilo{(if(1 N limit CNR lVurmhea mframedwl/iI lVurmbea mm/wullll N limit DINR H/urmhca mun'hisunidna 4 \Vurmhca otlorafa lVurmbea lVurmbea sp (KRN 11582) Wurmbea N limit Tamala Stn.

Commelinaceae

(~(1rr(J!l('nwphifydroides N limit KNP

Convolvulaceae Bonamia C'(J!1\'()!vu!us COllwlmlus Evn[vulus lllsinoides Ipomoea C(}S{(i[U Ipomoea muellen Ip(uw;ea peS-C{lprae subsp, brasiliensis S limit FPNP Polymerill sp. (RJ Cranfield 2563) Porana sericea \\/ilsonio hacklwL/s'ei N limit Carrang Stn. lVi/SOil ill hUflu"s N limit Salutation Isi.

Crassulaecac Crassu[a coloru[a acununufu N limit FPNP (,'russula co[orafa var co{oraw (.'rassula dcc'urnbcns dec'umbens Cmssulll * Crassula flOWn.\,' Crllssu!1I ped/(ellosiI CroSSU!iI pedul/cu!lIns

Cucurbitaceae A-1ukia nu.lderllspafana S limit Wooramel R

Cunoniaceac Aphwlllpelll!wl/ ,!el/wlldeum N limit DHI

Cupressaccac AClirwstrohus arefwnus N limit CNR Callitris g!aucoph ..,.{fa

Cuscutaccac Cuscula epuhvmum * C'uscuw p/anijlura Cuscuta rictoriana .... Familyrraxon Conservation Distribution Conservation Reserve N Code KNP TNR ZNP CNR FPNP BINR DINR KRNP o

Cymodoceaceae Amphibo/is un/urcticu Amphibolis griffithii Cymodoceu unguslala Cymodoceu serrulula Hulodule uninervis Syringodium isoeti/iJlium

Cyperaceae Buumeu ucutu N limit KNP Buwneu urticulala Baumea junceu N limit KNP Bulbostylis burbuf(l Cuustis dioica N limit KNP Chrysitrix distigmu!osu N limit CNR Cyperus aff. cunninglwmii (GJK & NG 592) Cyperus ulterni/lorus Cyperus bifax Cyperus blakeanus Cyperus bulbosus Cyperus castaneus Cyperus difformis S limit Gascoyne R. Cyperus gymnocuulos * Cyperus humulosus Cyperus ixiocarpus Cyperus rigidel/us Cyperus squurrosus Cyperus vuginutus Eleocharis acuta Eleocharis geniculuta Eleoc!wris pul/ens Guhniu lunigeru N IimitDHI Gahniu trifidu N limit KNP Isolepis cernuu N limit DHI Isolepis congruu Isolepis cyperoides Isolepis murginata lsolepis nodosa N limit Carrang Stn. Isolepis setiformis Lepidosperma angustatum N limit ZNP Lepidosperma rupestre 4 Endemic KNP Lepidospermu scabrum N limit KNP Lepidosperma tenue N limit KNP Lipocarpha microcephala Mesomeluena preissii N limit KNP Mesomelaena pseudostygia N limit Carrang Stn. ------Sc!1{!efl()pleCIUs dis.wclwfll!1us ScllOefl(lpleCfUS lalen/lorus S limit Coolcalalaya Stn. (from Hamersley Rg.) Sch(Jcf1(Jplectus li{(lra/is Sc!1oenople,}us suhulafus 5'clwcnus andrewsif N limit KNP 5'clwcnus drmeria N limit KNP Sclwcnus clandeslinus N limit ZNP Sc!1oenus curYI/oltus N limit KNP .Schocnus e!ega!ls Sclwenus glolll/er N limit KNP Sc!1ocnus glwu!J/lon/s N limit CNR Sc/wel/us gn/lil1lWluS N limit Murchison House Stn. Sc/weflus !1cxal/drus Schoeflus !1unults Se/wcl/us lalirafls N limit CNR Schoenus !/(lflUS N limit KNP Sclwenus (}dontocarpus ,S('h{)enus plel(},\-tefn(ineUS N limit KNP Sc!1oel/us sp (Keighery & Alford 1966) N limit Tamala Stn. Sc!1oeflus sp G Broad Sheath (KLWilson 2633) N limit Carrang Stn. Sclwel/us sp Kalbarn (K.R.Newhey 9352) 2 Endemic Kalbarri area Sc!1oCI/US sp Murchison (KLWilson 2647) Endemic KNP to TNR ,S'chocnus VtJrii('cflae N limit KNP Tt/raria microcarpa N limit CNR

Dasypogonaceae AWfllhocarpus afT. mhuslus (Hopper 1367) N limit FPNP AWfllhocarpus !1ufluhs S limit Quobba Stn, Acafll/wcarpus parVl/h!rus 2 Endemie to KNP Acarlt!l()("arpus prelssll Acunlfwcarpus UJhusfuS S limit ZNP Acafllh""alpus \'crllcillalus S limit Tamala Stn. (,'a/n>{usia ('\,(ltlC[J N limit KNP Calccra,na gwndt/!ow N limit KNP Chl1nll1c.xeros serra N limit KNP Lmllllfldw /lIlslJlis N limit KNP IAurwndra nUlritinw N limit Carrang Stn, Xero!irio!l di"'aricala

Dilleniaceae Hlhbertia acemsa N limit ZNP HiJlbertia uJ/upit'ua N hmit Nanga Stn Hibbertia desfIlophylla N limit KNP Hihhertia exasperala N limit KNP Hihherlia glofllcm.w N limit KNP Hibberlw huegelti N limit KNP Hihhertia hypericoides N limit KNP Hlhherlla POfCflfilll/!ow N limit KNP Hlbberflll racefllosa N limit Tamala Stn...... HII!berfia sp. Red Bluff (JRWhecler 2376) N limit CNR N ..... Familyrraxon .... Conservation Distribution Conservation Reserve N Code KNP TNR ZNP CNR FPNP BINR DINR KRNP N

Hibbertia spicata subsp. spicata Hibbertia stellaris N limit KNP Hibbertia subvaginata N IimitCNR

Dioscoreaceae Dioscorea hasti/i,!ia N limit Salutation Isl.

Droseraceae Drosera bu/bosa N limit KNP Drosera coo/amon ms N limit KNP Drosera ericksoniae N limit KNP Drosera g/andu/igera N limit KNP Drosera indica Drosera macrantha subsp. nwcrantha N limit KNP Drosera menziesii subsp. thysanosepa/a N limitZNP Drosera neesii subsp. borea!is N limitZNP Drosera radicans N limit KNP Drosera ramellosa N limit KNP Drosera sw/onitera subsp. humilis N limit Nanga Stn. Drosera st%nitera subsp. prostrata S limit KNP-Nanga Stn.

Elatinaceae Bergia auricu/ata 2 Bergia perennis subsp. obtusijo!ia E/atine grati%ides

Emblingiaceae Emb!ingia ca/ceo/if1ora

Epacridaceae Astr%ma g/aucescens N limit KNP Astr%ma microdonta N limit KNP Astr%ma serrati/i'/ium N limit KNP Astr%ma xerophyllum N limit KNP Brachy/oma sp. Kalbarri (A.H. Burbidge 4504) Conostephium preissii N limit CNR Leucopogon aff cordito!ius Leucopogon aff. strongy/ophyllus Leucopogon allitlii N limitKNP Leucopogon cOfwstephioides N IimitKNP Leucopogon cordito!ius N limit Carrang Stn. Leucopogon crassi/lorus N limit KNP Leucopogon hamu/osus N limit KNP Leucopogon hispidus N limit KNP Leucopogon leptanthus N limit Kalbarri Area Leucopogon marginatus R Presumed extinct Ajana area Leucopogon parvijlorus Leucopogon plafllf(,tius N limit KNP Leucopogon polymorphus Leucopoglm slrlmgrlophrl!us N limit KNP Leu('(J[}()gOf1 !aNunlJlt>/lSlS N limit KNP Lysincnw citialum

Euphorbiaceae Adriaf1i1 lomen[osa Beyeria Clnerea N limit BINR Beyeria (')'l1nes("cns Beyeria cygn()rUnl Beyena gardnen 4 Beyeria lepidopelala R BeVNw simitis Calycopcplus nwrginatus 3 Eupllllrhia afT alsll1fj/ora (ASW 1(547) Eupllllrhia alsifllj/ora S limit DINR Euphorhia alolo Euphorhia auslralis subsp auslratis Euplwrhia hoophlhona Euphorhw coghlanil Eupllllrhia drummondii * Euphorhia 11frta Euphorhia kimherlcyenns Euphorhia myrtlJldes EupllOrhia slwrkoensis S limit DHI Euphorhia tannctl.\'IS subsp eremophila Monolaxis graciliS Mono/iuis grmullj/om t'l1ono/axis lurida N limit ZNP Phyllanthus calycinus N limit Tamala Stn Phyllanthus juernmhrii S limit DHI Phyllanthus laeunel!us Phyllanlhus maderaspatcnsis Phyllanlhus maillandianus N limit Nanga Stn, Phyllanlhus sp, (ME Trudgen 7563) Endemic WHA Poranlhera miemphylla Pseudl1nthus intrico{us Ins Ricinocmpos glaueus Ricinocarp()s murico(Us N limit ZNP * Ricinus communis Sauropus emssif(JllUs S limit KNP Stachyslemon sp, (S, Van Leeuwen 210)

Frankeniaceae Fmnkenia amhita Frankenia one rca Frankenia conlusa 2 N limit KNP Frankenia lax/flora Frankenia magflljica I<'amilyrraxon Conservation Distribution Conservation Reserve Code KNP TNR ZNP CNR FPNP BINR DINR KRNP

Frankenia pauciflora Frankenia setosa

Gentianaceae * Centaurium erythraea Centaurium spicatum * Centaurium tenuiflorum

Geraniaceae Erodium angustilobum * Erodium aureum * Erodium cicutarium Erodium crinitum Erodium cygnorum subsp. cygnorum Pelargonium liUorale N limitZNP

Goodeniaceae Brunonia australis Dampiera altissima N limit Meadow Stn. Dampiera haematotricha subsp. dura Dampiera incana var. fuscescens Dampiera incana var. incana Dampiera krauseana 2 N limit KNP Dampiera lindleyi N limit Meadow Stn. Dampiera oligophylla N limit Murchison House Stn. Dampiera spicigera N limit Tamala Stn. Dampiera stenostachya N limit KNP Dampiera wellsiana Nlimit KNP Goodenia aff. havilandii (GJK & NG 1505) Goodenia berardiana Goodenia caerulea Goodenia corynocarpa S limit Murchison R. Goodenia drummondii N limit KNP Goodenia hassallii Goodenia havi!andii Goodenia maideniana Goodenia micrallllw Goodenia microptera Goodenia mimuloides Goodenia neogoodenia 4 Goodenia occidentalis N IimitKRNP Goodenia ochracea SlimitDHI Goodenia pinnatifida Goodenia pulchella NlimitKNP Goodenia salmoniana Type only Gascoyne R. Goodenia sericostachya 3 N limit Cobum Stn. Goodenia tenui!oba Goodenia Iriodioplllla G(}(Jdenia xunlh()spennu Lecherwullio chlomnlha R Endemic Kalbarri Area Leehenouilio /lorilJlmdo N limit Eurardy Stn, Lechenoulllo I{(rsula N limit CNR Lechelloultw lil/oT/mdes N limit DHllFPNP Lechenaultia macriiflr!za N limit Meadow Stn, Lechenoultio suhcwlwso S limit Carrang Stn Penla{Jri/o!l carey; N limit Coburn Stn, Scacvu!fl acacioides Scoel'olo anchusI/lllw N limit KNP ScaeV(J!a cancscens ScaeV(J/u ('hrys()!)(Jgon :> N limit WHA Sca('\'o[a crassif(J!iu N limit Coral Bay .)'('a('vola CUtltlillf.{hamii S limit Tamala Stn Scaevolo tlielsll N limit S of WHA SUlevola glal/dub/I'm Scoel'olo glohub/I'm N limit KNP Scal'volo humilloflil N limit KNP SUlel'olu kal/ophyl/a Endemic KNP Scaew){a !U!J('('()/o!a Scul'l'olu nilidu Scuevolu old/ieldil Type only Murchison R. ScaCl'o{a oxyc!O!la SUlevolo porViflllw subsp pilhome Scuevolu phle!Jopclolo N limit KNP Scaevo!tl {Jorocarya N limit VCL S of Tamala Stn SUlevolu pulchelto S limit near Carnarvon Scaevo[a repens var ereclll Ins Scaet'o[a resllucca SUlCl'olu sericophsl/u S limit Nerren Nerren Stn S'caevo/a spinesct'ns SUlevolo thesioldl's subsp, thesimdes N limit Wooramel R Scacvola [omentusa Seal'l'olu Virgutu N limit KNP Symphyohusis muuople,'tm Vel/eia C)-L!lO[')()lamica N limit KNP Vel/cia discopfwm Vel/cia glabmw Velteia hispida Verreau:ria reirHt'ardtii N limit KNP

Gyrostcmonaccac CodmwUlrpus ,'Ofinl/'lllus Gyrostemo!l facenuger N limit Coburn Stn G}'rostenwn fartlu!osus Gyrostemofl subnudus N limit Murchison House Stn Tenonia cyaffll/lon; N limit KNP Familyrraxon Conservation Distribution Conservation Reserve Code KNP TNR ZNP CNR FPNP BINR DINR KRNP

Haemodoraceae Anigownthos humilis subsp. humilis N limit KNP Anigowntlws kalbarriensis N limit Murchison House Stn. Anigownthos manglesii subsp. quadrans N limit Nanga Stn. Conostylis aculeata subsp. echinissima Endemic to Nerren Nerren Area Conosty!is aculeata subsp. rhipidion N limit KNP Conosty!is aculeata subsp. septentrionora Endemic KNP to Tamala Stn. Conosty!is androstemma N limit KNP Conosty!is aurea Conosty!is candicans subsp. flavifo!ia N limit Nanga Stn. Conosty!is prolifera N limit KNP Conostylis resinosa N limit KNP Conosty!is robusta N limit KNP Cmwsty!is stylidioides NlimitDHl Haemodorum brevisepalum N limitKNP Haemodorum discolor N limit KNP Haemodorum paniculatum N limit KNP Haemodorum simulans N limit CNR Haemodorum spicatum N IimitKNP Tribonanthes australis N IimitKNP Tribonanthes violacea Nlimit KNP

Haloragaceae Glischrocaryon aureum var. angustifolium G!ischrocaryon aureum var. aureum N limit Tamala Stn. G!ischrocaryon flavescens N limit KNP Gonocarpus conferti/iJ!ius Haloragis gossei Haloragis trigonocarpa Myriophyllum decussatum

Hypoxidaceae Hypoxis glabella var. leptantha N limitKNP Hypoxis occidenta!is var. occidenta!is N limit KNP

Iridaceae Orthrosantlws laxus var. laxus N limit KNP Patersonia drummondii subsp. borea!is ms Patersonia graminea N limit KNP Pater.wnia occidenta!is

Isoetaceae !soetes drummondii !soetes muellcri

Juncaceae Juncus aridicola * Juneus bufrJllws Juneus kmuss;l Juneus p{wufu{ws

Juncaginaceae Tng{uclllu aff m{Cllmpa (GJK 1(430) TriR,I(}chin ("a!('ilrapuflI Trig!(}('hin centro('urpurn Tng{uc!lln 111lnullSSlmUJIl Trigl(jchin mU(T(matum Trig{uchlll InclwphuruJll

Lamiaceae Herniandra pungens N limit KNP Hemialldm sp (Beard 7(59) Endemic WHA Hemiandm sp (Trudgen sn) Endemic WHA Hem;alldm sp Kalbarri (D Bellairs 15(5) 2 Endemic KNP Hemlgoua ,lip{alllhem N limit KNP Hell/fReniu mo('t(ult!ra N limit Tamala Sin. Hellugellw {"JIle{t/u{1II 3 Hnnigenfu scohra IYlicrocorys !oganil1cca Mif'rfN'()rys !t'fluU()liu 2 Proslall/hem allnflum ProslOll/hem eampln'{!ll Prostantheru '.,>'ilkituna * Salt'ia rerhenaca Weslrillgw aft' dampleri (GJK & NG 798) Wes/r;llgw damp/{'ri N limit DlNR VVeslringia rigid(J

Lauraceae Cassylha (Jurea var (lutea Cassylha a meemusa forma pilusa Cass.vtha facernosa fonna racemusa

Lentibulariaccae Ulricu!aria lellclla N limit KNP

Lobeliaceae !so[onw hypucralerU(Jrmis N limit KNP Lube{wa{ala N limit KNP

Lube/w glhhusa N limit KNP .... Lube!w helerophv!la "IN Farnilyrraxon .... Conservation Distribution Conservation Reserve N Code KNP TNR ZNP CNR FPNP BINR DINR KRNP 00

Lobelia rhytidosperma N limit KNP Lobelia winfridae * Monopsis debilis

Loganiaceae Logania afL litoralis (AH Burbidge 4406) Logania biloba N limit VCL W of Nerren Nerren Stn. Logania flaviflora Logania litoralis S limit KNP Logania nuda Logania spermacocea Logania vaginalis Phyllangium paradoxum

Loranthaceae Amyema benthllmii Amyema .fitzgeraldii Amyema gibberula var. gibberula Amyema hilliana Amyema linophylla Amyema maidenii subsp. maidenii Amyema melaleucae Amyema miquelii Amyema miraculosa subsp. miraculosa N limit NW Cape Amyema preissii Amyema sanguinea var. sanguinea Lysiana casuarinae Lysiana exocarpi Lysiana murrayi Lysiana subfalcata S limit Wooramel R. Nuytsia floribunda N limit ZNP

Lythraceae Ammannia baccifera S limit Carey Downs Stn. (from Barlee Range) Ammannia multi/lora Lythrum sp. Towrana (RJ Cranfield 2183) Rotala diandra S limit Carey Downs Stn. (from Rudall R.)

Malvaceae Abutilon afL oxycarpum (Morat 8145) Aburilon amplum Abutilon cryptopetalum Abutilon exonemum S limit DINR Abutilon geranioides S limit Tamala Stn. Abutilon leucopetalum Abutilon otocarpum Abutilon oxycarpum S limit Tamala Stn. Abuti!on sp. all. oxyearpum (Morat 8145) Abull!on sp. Hamelin (A.M.Ashby 2196) 2 Abuti!on sp. Quobba (HDemarz 3858) 2 Endemic Carnarvon Basin A!yoRyne cunel!il/1lllS A!yoRyne iwkel/idw A!yogrne hueRelil var R!ossu!ariae!illtus ms A!yoRrne hueRe!1I var hueRelti illS N limit Carrang Stn A(voKyne huegelil var ~vrayaerns AlyoRyne pinonwna var lepwch!amrs N limit Tamala Stn. Alyogync pinrJ!1ia!la var. pit/on/ono Gossypiurn austru{e Hibiscus brachych!aenus HibIscus brachySlphonlus Hibiscus burtOfll1 Hibiscus coatesii Hibiscus drummondil Hihiscus gardnerii Ins Hibiscus leptocladus Hibiscus sturtil var RfWldl!lonls HibiSCUS sturtil vac tnlflcatus * Lavlltcra creriea Lavulera plebeil1 var. {ornenlosa N limit DHl LaH-'ren(·ja densUl()ra Lawrencia R!()!'nerata LLm'fencia sp Mulein StatIon (Seller 317) Lo\-vrencia spi('u[(/ l.-

Marsilcaccac lvfamlea anRustl/iJ!w Marsilea drummondli MalSilea exarata tvfalSi/ea hirsuw

Menyanthaccac Vil/alSw conResll!lora 3 N limit KNP Farnilyrraxon .... Conservation Distribution Conservation Reserve w Code KNP TNR ZNP CNR FPNP BINR DINR KRNP o

Mirnosaceae Acacia aciphylla N limit KNP Acacia acuaria N limitTNR Acacia acuminata Acacia amblyophylla Endemic WHA Acacia ampliceps Acacia anastema Acacia ancistrocarpa Acacia andrewsii N limit ZNP Acacia aneara Acacia ashbyae N limit KNP Acacia aalacophylla Acacia bidentata N limit KNP Acacia bivenosa Acacia blakelyi N limit Tarnala Stll. Acacia cavealis ms N limit Murchison House Stn. Acacia chartacea Acacia citrinoviridis Acacia cochlearis Acacia colletioides N limit KNP Acacia comans Acacia coolgardiensis subsp. coolgardiensis Acacia coolgardiensis subsp. effusa Acacia coriacea subsp. coriacea Acacia coriacea subsp. pendens Acacia caspidifolia S limit Wooramel R. Acacia cathbertsonii subsp. cuthbertsonii Acacia didyma 3 Endemic Shark Bay & DHI Acacia dielsii Acacia drepanophylla 3 Endemic Carbla Stn. area Acacia ericijiJlia N limit Murchison House Stn. Acacia erinacea N limit Nerren Nerren Stn. Acacia jilrnesiana Acacia galeata Acacia gelasina 2 Acacia grasbyi Acacia gregorii Acacia idiomorp}w Acacia inaeqailatera Acacia isoneura subsp. nimia ms 3 N limit Nerren Nerren Stn. Acacia kempeana Acacia lasiocarpa var. lasiocarpa N limitZNP Acacia latipes subsp. latipes ms Acacia leptospermoides subsp. leptospermoides NlimitDHI Acacia leptospermoides subsp. obovala 2 N limitZNP Acacia ligulata Acacia ligustrina Acacia !ineo/ata subsp. lineo!a/([ Amcia linophylla N limit Eurardy Stn. Acacia ltmgispinea Acacia microhofrya AC(J("ia micrt}('ulyx A("(1("10 tn()rri,Y(}nii Acacia multisfJic(J[(l Acacia mUrUI)'{UUJ Amcia neuml'hrlla subsp erugata N limitCNR AmCla o/dlie!dll N limit KNP Acacia oxyclada N limit KNP Awcia l'a/ustns Acacia fJdraneura Awcia p!autellu ms 3 Acacia pruiw)('arpa Acacia puncricu!a!a illS N limit KNP Awcia l'ynl(l!w Awcia quadrimargll1ea Awcia quadnsu!cata N limit ZNP A('acia romu/usa Acacia resliacca N limit KNP Awcia rlwdoph!oUl Awcia mstellt!('ra Acacia ro}'cei N lirmt Hamelin Stn Acacia sa!Ju/osa illS Acacia saliRlla Acacia sa:xotilis N limit KNP AC(J{'ia s('irpU()!itl N limit KNP Acacia sc!erospernw subsp r.:/ouces('ens 3 Awcw sclemspemw subsp sclemsperma N limit Yaringa Stn. Acacia sessilis N limit KNP Acacia slRfWfu N limit Eurardy Stn. Awclil sp. (KRN 11816) Amcia spathulil()lia Amcw sphenaphvlla ms N limitCNR Acacia stereophylla var. cylindrata 2 N limit CNR Acaclil stereophylla var stereophvlla N limit Nerren Nerren Stn. Acaclil suIJrigida N limit Tamala Stn. Af'licia subtessan)g(JrJo Acacia synchronicia AcaCIa tetragonol'hyl!a Acacia ulicifW N limit KNP Acacia vlctorwe Acacia Hianyu Acacia \i'ISeOI1(1 Acacia xanthina N lirmt Tamala Stn. Acacia xiplwphvlla Neptunia dimorp!wntha * ProSilpls palllda Familyrraxon Conservation Distribution Conservation Reserve Code KNP TNR ZNP CNR FPNP BINR DINR KRNP

Molluginaceae Glinus IOfoides Macarthuria ausfralis Macarfhuria georgeana I N limit TamaJa Stn. Macarfhuria infricata 3 N limit Nanga Stn.

Moraceae Ficus platypoda var. lachnocaulos S limit DINR Ficas platypoda var. minor

Myoporaceae Eremophila clarkei Eremophila compacta subsp. compacta Eremophila crenulata ms Eremophila cuneata ms Endemic WHA Eremophila cuneifolia Eremophila decipiens subsp. decipiens ms Eremophila deserfi Eremophila exilijiJlia Eremophila flaccida ms Eremophila jorresfii Eremophilajraseri subsp fraseri ms Eremophila jraseri subsp. parva ms Eremophila jreelingii Eremophila gibsonii Eremophila glabra Eremophila glabra subsp. Zuytdorp (GJK & NG 518) Eremophila glabra subsp. albicans Eremophila glabra subsp. carnosa ms Eremophila glabra subsp. psammophora ms 2 S limit DINR Eremophila glabra subsp. tomentosa ms Eremophila glufinosa Eremophila granitica Eremophila lanata ms Eremophila latrobei subsp. lafrobei ms Eremophila leucophyl/a Eremophila longijiJlia Eremophila mackinlayi subsp. mackinlayi ms Eremophila mackinlayi subsp. spathulafa ms Eremophila maculata subsp. brevifolia ms Eremophila mait/andii Eremophila mackinlayi subsp. mackinlayi ms Eremophila microfheca R Eremophila occidens ms 2 S limit Nanga Stn. Eremophila o/dfieldii subsp. oldfieldii Eremophila oppositifo!ia Eremophila pantonii N limit Wooramel R. Eremol'h!la l'hyllol'oda ms Eremol'hila l'hysocalyx ms N limit Meadow Stn. Eremol'llIla I'lalYcalyx Eremol'llIla I'lerocucl'a Eremol'llIla serrulma N limit Hamclin Stn. Eremol'htla selacea ms Eremol'h!la sl'lendells rns Endemic WHA Erem0l'hila sl'ur!a ms Eremol'hila slrlillgylol'hylla Eremol'llIla subf/oceosa Eremol'hila YOUIl;;!t subsp YOWI;;II rns A1}'(}!l(JrUnl u('umif1a[um llt!}'(J!}(Jrurn insulare lV(WJ[J(JrurN nl()!11anum

Myrtaccac Baeck,'a "Iackallit ms Ba('ckeu {lUllRurethllc Ins Baeckea me;;af/o«J ms Baeckea l'enta;;onanlha N limit Coburn Stn. Bankea robusla Baeckea sp (M.ETrudgen 7180) N limit Tamala Stn. Baeckea sp Murchison River (SD.Hoppcr 1428) N limit CNR Baeckea sp Mmgencw (M.E.Trudgen 12(29) Baeckea sp Nanga (AS.George 11346) N limit WHA Stn. Baeckea sulJCuneata 2 Beaujflrt!a daml'!en N limit DINR

Beal~l(jrti{jeleguns N limit KNP

Be{1l~l(Jrti(Jsquurrosu N limit KNP C'(lllistem()fl plu;eni("eus Calollwmnl1s blel'harosl'erml1s N limit Tamala Stn. Calolhamnus borealts Calolhamlll1s ('hrysalllherus Calollwmnl1s /fll7r!OSl1S subsp. formosus Endemic WHA Calolhamnl1s ;;Iabcr ms N limit Nanga Stn. Ca!lJlhamnl1s 11IimallJl'hyllus Calolhamnus Iwlbarr!ens!s N limit Tamala Stn. Calollwmnus oldj/cldi! N limit Eurardy Stn, Ca!lJllwmnus {hlchyslachyus 4 N limit KNP Calollwmnus quadnf/dus N limit Nanga Stn. ('ab)thamnus songulneus Calylrix hrenjolll1 N limit Tamala Stn Calylrix del'ressa Calylr!x desolata Calylr!x drummondtl 2 N limit KNP Calylrix eealycala Calylrix /fJrllwsa 2 Calylrix frasen N limit KNP Calylrix ;;lul"/Osa N limitKNP Calylrix ;;mCllts N limit KNP Familyrraxon Conservation Distribution Conservation Reserve Code KNP TNR ZNP CNR FPNP BINR DINR KRNP

Calytrix harvestiana 2 Calytrix leschenaultii Calytrix oldfieldii N limit CNR Calytrix paucicostata 2 Endemic Kalbarri Area Calytrix purpurea 2 Endemic Kalbarri Area Calytrix sapphirina Nlimit KNP Calytrix strigosa NlimitDHI Calytrix truncatifo/ia Chamelaucium brevifo/ium N limit KNP Chamelaucium conostigmum ms 3 Chamelaucium marchantii 2 Endemic Kalbarri Area Chamelaucium micranthum Chamelaucium oenanthum ms N limitZNP Chamelaucium paucif!orum subsp. pauci!lorum ms N limit KNP Chamelaucium sp. Coolcalalaya (A.H.Burbidge 4233) Chamelaucium uncinatum N limit Murchison House Stn. Chamelaucium vinosum ms Conothamnus trinervis N limit Tamala Stn. Darwinia capitel/ata N limit KNP Darwinia oldfieldii Endemic Kalbarri Area Darwinia paucif!ora N limit KNP Darwinia virescens N limit KNP Eremaea acuti!{)/ia 2 Eremaea dendroidea N limit Nanga Stn. Eremaea ebracteata Eremaea ebracteata var. ebracteata Eucalyptus aff. prominens (MIH Brooker 4569) S limit Tamala Stn. Eucalyptus aff. socia/is Eucalyptus beardiana R Eucalyptus camaldulensis var. obtuSll Eucalyptus decipiens Eucalyptus dichromophloia Eucalyptus diminuta ms 2 Eucalyptus do/ichocera ms Eucalyptus drummondii Eucalyptus erythrocorys N limit ZNP Eucalyptus eudesmioides subsp. eudesmioides Eucalyptus eudesmioides subsp. pallida ms N limit Towarana Stn. Eucalyptus eudesmioides subsp. sel/achiana ms Endemic WHA Eucalyptus j1ocktoniae Eucalyptus foecunda subsp. Coolimba (M.I.H.Brooker 9556)3 N limit KNP Eucalyptus fruticosa S limit KNP Eucalyptus gillinsii N limit Nanga Stn. Eucalyptus hypochlamydea subsp. hypochlamydea ms Eucalyptus jucunda N limit CNRlCoburn Stn. Eucalyptus kochii Eucalyptus lenziana Eucalyptus leptopoda Eucalyptus loxophleha subsp. supralaevis Eucalyptus mallllensls subsp. vespertllla Eucalsptus ohtusljlora Eucalyptus old/ieldu N Iimil Nerren Nerren Sin. Eucalyptus orllrio N limit BINR Eucalyptus pmllllllellS Eucalyptus psn/ormis N Iimil KNP Eucalyptus rif.(idula N IimitWHA Eucalyptus mln'l Endemic WHA Eucalvptus sheathiww Eucalyptus srna/lcalvx Eucalyptus suhallf.(usto sllbsp. pusilla Eucalsptus trallsomtilU'lltalis Eucalyptus vicrnx Horna!oca(v:r afT (lurc(J HomaloclJ!Yx aureus N Iimil KNP Hspocalymma IOIlf.(ljiilll1m R N limil KNP Lamarchea hakeljolw var. hreVl/olw N limit Nerren Nerren Sin. Lamarchea IwkCl/iilia vac hakeljiilia Endemic WHA Malleostemoll hurstluluset N limit Coburn Sin. :\}ullcosfertlof1 minil....,aensis Malleostemoll pedlllu'ulatus N limit Carrang Sin. Malleostemoll peltif.(er N limit Coburn Sin. Mal!costemon roseus Malle'ostemoll sp Nerren Nerren (Payne 360) Endemic Nerren Nerren Sin. Malleostemoll sp Meadow Station (A.H.Burbidge sile 69) Malleostemoll sp Rabbit Proof Fence (A.H.Burbidge sile 59) . Malleostemoll sp Cooloomia (S.D.Hopper 1353) 2 Endemic WHA Malleostemoll sp. Hardabull Rapids (DBellairs 1654A) 1 Malleostemoll sp. Kalbarri (LACraven 7083) 2 Malleostellwll sp Moonyoonooka (R.J.Cranfield 2947) 2 N limit KNP Malleostemoll tuherculatus Afelaleuca accm.w N limit Tamala Stn. Melaleuca adnato Melaleuca afL leiopyxis (GJK & NG 17(8) ,Helaleuca afL eremaea (GJK & NG 1332) Melaleuca aff Iwlosericca Melaleuca all. nesophila (AHB 4520 ) Melaleuca afT. old/ieldii/nesophylla (Beard 6768) Endemic WHA Melaleuca afL old/leldii (C. Gardner 13412) Endemic WHA Melaleuca calotlullrlIwides Endemic Kalbarri - Mllrchison House Stn. Melaleuca cardiophylla Afelaleuca cilwsa N limit KNP Melaleuca COil crI' to Endemic Kalbarri Area Me/aleuca CO!lo{!wmnoides Melaleuca cordata Melaleuca depressa N limit KNP Melaleuca eleutemswchya Melaleuca ererrwea Familyffaxon Conservation Distribution Conservation Reserve Code KNP TNR ZNP CNR FPNP BINR DlNR KRNP

Melaleuca julgens Melaleuca glomerata Melaleuca holosericea NlimitDHI Melaleuca huegelii subsp. prislicensis 2 Endemic WHA Melaleuca lanceolata subsp. occidenlalis NlimitDHI Melaleuca lasiandra Melaleuca lalerijlora subsp. aculifolia ms Melaleuca lalerilia N limit KNP Melaleuca laxif/ora Melaleuca leiopyxis N limit Nanga Stn. Melaleuca leucadendra Melaleuca linophyl/a Melaleuca longislaminea subsp. longistaminea ms N limit KNP Melaleuca megacephala Melaleuca nematophyl/a Melaleuca oldfieldii Melaleuca psammophila Melaleuca radula Melaleuca rhaphiophyl/a N limit KNP Melaleuca scabra sen. lat. Melaleuca sp. (Trudgen sn) Endemic WHA Melaleuca sublrigona Melaleuca Irichophyl/a Melaleuca uncinata Melaleuca undulala Melaleuca urceolaris Melaleuca viminea subsp. viminea N limit Carrang Stn. Micromyrlus racemosa Micromyrtus sulphurea Phymatocarpus porphyrocephalus Pileanlhus aff. filifolius (Bellairs 1684) Pileanthus aff. peduncularis (Hopper 1385) Pileanthus auranlicus ms Endemic Murchison House Stn. Pileanlhus bel/us ms N limit ZNP Pileanlhus limacis S limit Hamelin Stn. Pileanlhus peduncularis Pileanthus seplenlrionalis (ML 2492) Pileanlhus vernicosus N limit Cobum Stn. Schollzia cordala ms Scho/tzia leplanllw Schollzia sp. Cookalalaya (A.H.Burbidge sn) . Schollzia sp. Ajana (T.A.Halliday 137) 3 Schollzia sp. East Yuna (A.C.Bums 6) 2 Schollzia sp. Eurardy (J.S.Beard 6886) I Schollzia sp. Folly Hill (M.E.Trudgen 12097) I Schollzia sp. Galena (W.E.Blackall 4728) 2 Schollzia sp. Kalbarri (N.Hoyle 623) Endemic Kalbarri Area S,!loll:w sp Murchlson River (A.S.George 7098) Sc!loll;w NCH1hampton (AStrid 2(714) Sc!loll;w sp Red Bluff (A Gunness 2373) Sc!loll;w sp Ross Graham lookout (S.Maley 6) 2 Sc!loll;w sp Shark Bay (MET 7429) N limit Nanga Stn. Sc!lo!r;w Z-Bend (Belhurs-Kalnora 912a) 2 Sclwllz/{/ ,pawlala Sclwlwa "ben/lora Sc!zo/t::)(J umhelli!£'Tu T!lrs!'loIl1Cnc aspera ssp glabra MS Thr.\'pffflUe!lC hacI'kC(J('Cll Thryptomene dc('ussart[ N limit Cohurn Stn Thn'!,IOll1ene denllclllala Thryptomenc mw'ronu/a!o Thrypfomene ninglwnensis Ins Thnploll1enc "rolt/era N limit CNR Thryp[()fuene racef1lulosa N limit Cohurn Stn. ThnpwlI1"ne sp Kalharri (A Gmness 2378) ThrsplOll1ene sp Wandana (AJI.Burhidge site I.') ThrsplOlI1ene sp Carrarang(M.ETrudgen 7420) I Endemic WHA Thnploll1"ne sp Eagle Gorge(A.G.Gunness 23(0) 2 Endemic KNP to ZNP ThrsplOll1ene sp Eurardy(D.& BBellairs) 2 ThrvplOlI1cnc sp Junga Dam(AGGunness 2383A) 2 Thnploll1ene sp Steep Pomt(M.ETrudgen 7421) Endemic WHA Thnploll1ene sp Tamala(METrudgen 7384) ThrvpwlI1ene slenophsl/a 2 Thrsploll1ene slrongvlophslla N IimitCNR Thryploll1ene slrongslop!lsl/a suhsp Tamala (METrudgen 7384) Endemic WHA Verllcordia ((!flll/ans 4 Verticordul chr.\'santha VCftl('(Jrdiu t'/,f-YS'{)S'lu('hys var. ('!zrysf)s[(J('h.vs Verticordia c()o!oomia 3 Endemic WHA Verlicordw dasyslylls suhsp. kallwrnensls 2 Verlicordia dell.H/lora var roseoslel/a 3 Verllcordia dens/flora var. slel/aligera \ferricurdio. dichroma vaT dichroma 3 N limit CohurnlMeadow Stn.

Verticordia dichro!lla vac Srtlfonw :3 N limit CNR VertiuJrdia cri(}ceplwla Verticordia ethe!iatla var eth(J!iana N limit VCl S of Tamala Stn Verticordiu cthe!iaf/u var j(JntlOSa Verlicordw /orresllI Verlicordia galeala 2 Verticordia 1lIlertons VerI/cordia lepldophvlla var lepu}ophyl/a Verlicordia lepidophyl/a var ,!uamula Verl/cordw irlonadelpha var. call1lric!la N limit Kalharri Area Verlicordia nobilis Verticordia oculata Verticordia penflIRcUl Verli"ordia pluJlu}op!lyl/a Familyrraxon Conservation Distribution Conservation Reserve Code KNP TNR ZNP CNR FPNP BINR DINR KRNP

Verticordia picta Verticordia polytriclUl 4 Verticordia spicata subsp. spicata N limitCNR

Najadaceae Najas marina

Nyctaginaceae Boerhavia burbid[?eana S limit Carnarvon Boerhavia coccinea Boerhavia [?ardneri Boerhavia repanda Boerhnvia schombur[?kiana Commicarpus australis

Olacaceae Olax aurantia

Oleaceae Jasminum calcarium S limit KNP Jasminum sp. (B. Turner I) S limit Carrang Sin.

Ophioglossaceae Ophio[?lossum [?ramineum Ophio[?lossum lusitanicum e") -:- ~ Orchidaceae /'l) Caladenia bical/iata N limit KNP ciQ" ::r Caladenia bryceana subsp. cracens ms R /'l) Caladenia discoidell N limit KNP ~ Caladenia drummondii ?- Caladenia .flava subsp. maculatll ms e") Caladenia hirtll subsp. rosea ms N limit KNP ;: Caladenia hoffmanii ms R '"0 Call1denia incensa ms .? Call1denill latifolia N limit KNP ~ N limitKNP Caladenia lon[?icauda subsp. borealis ms ~ Call1denill pachychilll ms r-' '< ClIladenia reptans subsp. impensa ms Endemic KNP 0 Caladenia reptans subsp. reptllns ms N limit KNP :::l !!' ClIladenia roei Caladenia varians ms > ;:z: Call1denia vllrians subsp. nobi/is ms N limit KNP Caladenia vllrians subsp. varians ms O'.l:: ... Caladenia wanosa R <:r Cyanicula amplexans ms N limit Kalbarri Area Q.; Oll Cyaniculll dej{mnis ms /'l) Cyunicu/o gernrnata Ins ()'rtoS!Ylis huegelil N limit KNP Dwris afT IiUlflora rAP Brown 32) N limit Kalbam Area Dwns !anflora N limit KNP

Diuris m{/,~ni/ic(J N limitZNP Diun, 4 N limit KNP Drakoca CO!ico!or ms R DrakonorchlS harbarel!a ms R Ei\lhranlheru bnlf10nls N limit KNP E!Ylhranlheru emarglnaw N limit KNP Enoclll!us dllawlus subsp dl!alalus ms N limit Tamala Stn Enoclll!us dllalalus subsp undu!alus ms N limit KNP L"pore//a flmhnala Microlls medw subsp. medw PuraCU!CiUIU IYOllsii Ins Endemic TNR Nerren Nerren Stn. Paracu/cana terminalis ms P/asophyl!um co/nc% ms N limit KNP Prasophy/!um cyphochl!um N limit KNP Prasophy/!um e/olulII Prasophyl!um glgonleum Prasophyl!um gruclle Prasophyllum nngens Prasophyllum sargentll Plerostvlis all /wno (GJK & NG 1867) Plerostvlis aspera N limit KNP Pleroslrlis /m/bow N limit KNP Pterostylis nano Pleroslylis plc/a N limit KNP Pterostvlts vangUlUC(J. Pterosty!is scuhra Pyrorchis nigricans N limit Tamala Stn. Splcu/aea n/ww N limit KNP The/YIIII/ra anlenmfera N limit KNP The/ymllra campanu!ala N limit KNP The/ymilra jlexuosa N limit KNP The/ymllra sargenlll The!ymllra spiralis N limit KNP

Orobanchaccac Orobonche {ninor var. oustraliano

Oxalidaccac Oxalis perennans * Oxalis pes caprae

Papavcraccac * ,Aruenwne ochro!euca

Papilionaeeac Aolus phylic(}{des N limitZNP Familyrraxon Conservation Distribution Conservation Reserve Code KNP TNR ZNP CNR FPNP BINR DlNR KRNP

Bossiaea eriocarpa N IimitZNP Bossiaea spinescens N limit KNP Bossiaea walkeri Brachysema aphyllum N limit ZNP Brachysema macrocarpum Chorizema ericiji/!ium Chorizema racemosum Clianthus formosus Crotalaria cunninghamii Cullen cinereum Cullen leucanthum ms Cullen patens Daviesia divaricata N limit Murchison House Sw. Daviesia hakeoides subsp. hakeoides Daviesia in/ricata Daviesia nudiflora subsp. nudi/lora Daviesia peduncula/a N limit KNP Daviesia podophylla N limit Murchison House Stn. Daviesia ramosissima Gastrolobium bennellsianum N limit KNP Gastrolobium oxylobioides N limit KNP Glycine canescens Glycine c!andestina Glycine tabacina Glycyrrhiza acanthocarpa Gompholobium sp. Kalbarri (M.D. Crisp 6300) Endemic Kalbarri Area Gompholobium Iomenlosum N limit Tamala Stn. Indigofera australis Indigofera boviperda Indiw!fera brevidens Indigofera colutea Indigofera georgei Indigofera !inifolia Indig(ifera linnaei Indigofera monophylla Indigo/era occidenta!is ms Indigo/era sp. (KRN 11615) Isotropis cuneifolia N limit Nanga Stn. Jacksonia aff. spinoS(l (RJ. Cranfield 2576) Endemic Carnarvon Area Jacksonia angulata N IimitTNR Jacksonia arenicola ms N limit Nerren Nerren Stn. Jacksonia arida ms Jacksonia calcicola ms Jacksonia cupulifera N limit KNP Jacksonia folio.1'il N limit KNP Jacksonia h

Tephrosia flammea Tephrosia gardneri Tephrosia rosea Tephrosia sp. (K.R. Newbey 11636) * Trifolium campestre Trigonella suavissima Viminaria juncea N limit KNP

Phormiaceae Dianella revoluta Stypandra glauca N limit KNP

Pittosporaceae Billardiera bicolor var. lineata N limit Tamala Stn. Billardiera erubescefls Billardiera ringens Bursaria occidefltalis N limit Nanga Stn. Cheiranthera filif{}lia Cheirallthera preissiana var. preissiana Piuosporum phylliraeoides var. phylliraeoides

Plantaginaceae Plalltago aff. hispidula (A.H.Burbidge 4517) * Plalltago coronopus Plantago debilis Plantago drummondii Plantago sp. Mt. Magnet (A.S. George 6793)

Plumbaginaceae Muellerolinum salicomiaceum

Poaceae * Aira caryophyllea Amphipogofl caricinus Amphipogon strictu.I· N limit TNR Amphipogon turbinatus N limitZNP Aristida aflthoxallthoides Aristida cOfltorta Aristida holathera var. holathem Astrebla elymoides Astrebla pectinata Austrostipa compressa Austrostipa crinita Austrostipa elegantissima Austrostipa macalpinei Austrostipa nitida Austrostipa tenaifolia Auslroslq){l Inchophrllll AWNrosripa fLIcker! * Avellmill nllche!" * A vena barba!a * At'cna Iowa * A vena stcrili,\ Bothri(fChloa eH'artuuw Brachyo(>hne pnJsfra{(1 * Bri::a ma.yima '" Bri;a minor Bro!llus oreflarlUS * Bromus diandrus BrofHus hordeaceus *' Bro!nIL) japonicus * Bromus madritensis Bromus riRidus * Cenchrus ciftans * ('enchrlls echinarus '" C'enchrus setigerus Ch!oris !obala Ch/oris peUi,tala Ch!oris {'umifw Ch !on\' virgll!a Chr\'Sopogon Jallox Chrssopogon {'aU/(/us Chryso{'ogon sp, Zuytc!orp (GlK & NG 9(6) Endemic to ZNP Csmbopogon wn!Jlgllus C'ynlbopogr)JJ !NJlnbyr'inus C'ymbo{Jogon ohtecfuS ('ynodofl dacty!o!l '" [)act.v!ocleniurn australe Daus!ouenium radu!ans f)anfh()nia caf!spi!osa N limit FPNP [)anlhonia rucemosa Dichanlhium senCtum suhsp, hum/tius Dig/tar/a brown/i Digi[orla ctefwnlha Dip!achne /US((I Dq,!achne mueUen Dlp!achne parn/lora * Ehr!wrta bren/il!/(! * Ehrhllrlll ca!YC/tllI * Ehrhllrtu !ongl/lora * Eleus/ne ind/co Enneap()g()!l cacrules{'ens Enneapogon po!vphsllus Enteropogofl acicularis Eragrostis austra!asica * Eragrostis barrelieri Eragrostis hasedrHvii Farnilyrraxon Conservation Distribution Conservation Reserve .... Code KNP TNR ZNP CNR FPNP BINR DlNR KRNP t

Eragrostis brown;; Eragrostis cumingii • Eragrostis curvula Eragrostis dielsii Eragrostis elongata Eragrostis eriopoda Eragrostis falcuta Eragrostis lanipes Eragrostis leptocurpa Eragrostis pergracilis Eragrostis setifolia Eragrostis tenellula Eragrostis xerophila Eriachne aristidea Eriachne avenacea Eriachne beflthllmii Eriachne flaccida Eriachne gardneri Eriachne helmsii Eriachne lanata Eriachne mucronata Eriachne obtusa Eriachne ovata Eriachne pulchella subsp. dominii Eriachne pulchella subsp. pulchella Eriochloa australiensis Eriochloa procera Eriochloa pseudoacrotricha Eulalia aurea Eulalia fulvu • Hordeum geniculatum • Hordeum leporinum Iseilema eremaeum Iseilema membranaceum Iseilema vaginiflorum • Lamarckia aurea Leptochloa digitata • Lolium perenne • Lolium rigidum • Lophoch/oa pumila Monachather paradoxus Neurachne alopecuroidea Panicum australiense Panicum decompositum Panicum eUusum Panicum laevinode Paractaenum novae-hollandiae Paraneurachne mue//eri * Parapho!is incurvu Pospolidlllm IWSlclodllm Pus!w/idium (,I('mentii Puspa/idium (Dflstn('lWn POSPOlldlll1ll gmClle POSpolldlllm fllhtflorum Pospolidllllll re/lexlIm * Pentaschistis airoides Perolis rani * Phularis tlzinor Ploginselllm refmcllIlll Plenrac!me hnul{(Jides 4 N limit Tamala Stn Pleclrm1111c dalltlumi()ides N limit FPNP Pleclrachlle dlelsll Ple('tru('hne drUtllnl(}ndii N limit Nanga Stn Pleuraelme meil'lllel Pleurac/me rigidissi!1la P/('('fru('hne P(Nl annua Poa dranzmondiuna N limit Nanga Stn, * Po{ypogOtl mOflspehensis Psammagrostis H'iseafw * Roslraria crislala

:;: Roslraria pumilu * S'chismus ara}ncus Sehisllllls borlmllls SelOrtil dlelsll Setaria surgens * S'etaria vertici/ltJlo

~)'(Jrghump{Utlz(Jsum Spilllfl'x IOllgltollllS S!}(Jr(Jh(J/us (l(,tin(}('/adus Spombollls blokel Spombollls romll Spombollls Illllrhl'lIll S'poroho/us l'tr){lfllCl/S Themeda [riat/dra Thyrtdolepis lll11ehelllilf/{/ Tragus auslralionus Triodia aff !JasedoH,'ii

Triodia basedo~\'ii Triodia epaC[Ul Triodia trnfans Trtodio Imllgem Triodia plurinervalu Triodia pungens Triodia scariosu Triodia lriticoides Tnpogoll lolilt/mms Familyffaxon Conservation Distribution Conservation Reserve Code KNP TNR ZNP CNR FPNP BINR D1NR KRNP

Triraphis mollis * Trisetaria cristata Urochloa gilesii Urochloa piligera * Vulpia myuros Yakirra australiensis

Polygalaceae Comesperma acerosum 3 N limit KNP Comesperma calymega N limit KNP Comesperma integerrimum N limit ZNP Comesperma polygaloides Comesperma scoparium N limit Tamala Stn. Polygala isingii

Polygonaceae * Emex australis Muehlenbeckia adpressa N limitZNP Muehlenbeckia cunninghamii Rumex crystallinus 2 * Rumex vesicarius

Portulacaceae Calandrinia aff. pumila (Andeberg 206) Endemic Carnarvon Basin Calandrinia calyptrata Calandrinia corrigioloides N limit FPNP Calandrinia disperma Calandrinia eremaea Calandrinia granulifera Calandrinia lehmannii Calandrinia linif10ra Calandrinia papil/ata Calandrinia polyandra Calandrinia polypetala Calandrinia primulijlora Calandrinia ptychosperma Calandrinia pumila Calandrinia remota Calandrinia sp. Coolcalalaya (GJK & NG 698) Calandrinia sp. Shark Bay (J.J. Alford 1376) Endemic WHA Calandrinia sp. (GJK & NG 1495) Calandrinia stagnensis Portulaca oleracea

Posidoniaceae Posidonia angustijiJIia Posidonia australis ,..------Posidonia ('onacea Posidonia SffllUJsa

Potamogetonaceae P()!alll(}gelr)!) !J('("!lfl(l!US Po{unu}gewn tnCartfla!US Ruppia mcg(u"orpa RUflflw po!rcw/,a Ruppia ruhcrosa

Primulaccae * Arwgullis {lfTCnsis var an'CllSlS * Anagollls (UTCflSis va[ ('ocrulca Sa!1/()!us JUN('CUS Samolus rcpcns var. jlorihundus Samo!us repens var. paun'j"o!ius N limit FPNP Samolus sp Shark Bay (METrudgen 7410) S limit Hamelin Stn

Proteaceae Adcnwlf!los a(Wlf!l{)p!lrllus Endemic to WHA Adc!lanth(}s ('ygl/()rum N limit KNP Banksw ashh\'l BUflksia oftenualu N limit ZNP Banksia lepfoflhrlla var. melllfiCil N limit KNP Banksia lindlewlf/a N limit Tamala Stn, Banksia menZlesii N limit KNP Banksia pri(JfUJ{CS N limit Tamala Stn, Banksia sccpfrum N limit Nanga Stn, Banksia vicforia(' N limit ZNP ('oflospermum aceroswn subsp hir.\'ulUm N limit KNP ('ufjospernuun horca{e subsp. !Jurea!e Endemic Kalbarri Area ('{)f1(JSpennufll micnJjl(Jrurn N limit Nanga Stn CO!lospennum stocclwdis subsp_ sloechadis N limit Tamala Stn, Dryandm afl. ashh)'1 (GlK & NG 1571) Eodemic Murchison House Stn, to Zuytdorp Dryandm horealis subsp, borealls Dryandm jmsen Dryandra s('ssllis Grevillea afl. sfenohofrya (MET 7554) S limit Tamala Stn. Grevillea wlfluli{era 2 N limit WHA Grevillea argrl'Op!lylla N limit Tamala Stn Grevillea hltimms subsp, bJ/lmnis N limit KNP Grevillea bmchrsfac!lra N limit ZNP Grevillca Cilndelabl'Oujes N limit FPNP Grevillea calldlcwlS N limit KNP Grevillca ('(Jrnmutoto N limit Nerren Nerren Stn Grevillea coSlofo 2 Endemic lower Murchison R, Grel'illell dU!VlIlo/wfrYIl subsp, dU!VlIlo!Jofrya N limit Nerren NelTcn Stn, Gret'illca die/siand N limit CNR Grerillca erios[(1chya Grevi/lc(l gtJrd(}nitJtltJ Familyffaxon Conservation Distribution Conservation Reserve Code KNP TNR ZNP CNR FPNP BINR DINR KRNP

Grevillea hakeoides subsp. stellophylla NlimitDHI Grevillea illtricalll N limit KNP Grn'illea leptopoda 3 N limit KNP Grevillea leucoclada N IimitTNR Grevillea leucopteris N IimitZNP Grevillea levis N limit KNP Grevillea obliquistigma subsp. obliquistigma N limit KNP Grevillea paradoxa Grevillea petrophiloides N limit KNP Grevillea pillaster N limit ZNP Grevillea preissii N limitZNP Grevillea pterosperma Grevillea rogersonialla 2 Endemic Shark Bay - Tamala Stn. Grevillea sp. Shark Bay (MET 7553) Endemic to WHA Grevillea stellobotrya Grevillea stellomera 2 Endemic KNP to Tamala Grevillea stellostachya 3 N limit Cobum Stn. Grevillea trachytheca Endemic Kalbarri Area Grevillea tri/oba 2 N limit KNP Grevillea variiji,lia subsp. bUllderra ms S limit Lake McLeod Grevillea vestita subsp. isopogoides N limit KNP Hakea arida Hakea bucculelllll N limit CNRffNR Hakea cundollealla N limit KNP Hakea circumalalll N limit Tamala Stn. Hakea coriacea Hakea coSlll1ll N limit KNP Hakea illcrassata Hakea kippistialla Hakea lissocurpha N limit Murchison House Stn. Hakea orthorrhyllclw Hakea preissii Hakea pycllolleura N limit KNP Hakea recurva Hakea stenocarpa Hakea stellophylla Hakea suberea Hakea trij'urcalll N limitZNP !sopogon divergells Persoonia acicularis N limit Nanga Stn. Persoollia angustif/ora Persoollia biglalldulosa N limit CNR Persoonia bowgada N limit Nanga Stn. Persoollia brachystylis Persoollia hexagolla N limit CNR Persoonia papillosa N limit KNP Persoollia rufij10ra N limit KNP PerSlJOf1la saundersuuw N limit Nanga Stn Persoonw sp. (Craven 112) N limit Nanga Stn Persoonia \'trtc!a N limit KNP PelrophIie Inlenwlll N limit KNP Pelropll/!e hrerifo!1I1 N limit Tamala Stn. Pelro!,hIic (o/!lfera N limit KNP PetrtJ[Jhile nUUI(istm)lYo N limit Murchison House Stn PelropllI!e S((lhnUsiu!a N limit KNP Petr(Jphile \'cI/litur("(it(l N limit ZNP Perrophile \"cminwfa N limit KNP Pelrophl!e shull!eH(Jrlh,wlil N limit KNP SlIrlingw !allfolia N limit KNP Svnaphea rnlllTa N limit KNP 5'yl/aphell spinulosa subsp !Jor('u!is N limit Eurardy StnlCNR Xv!ome!um IIngllsllfo!wm N IimitCNR

Psilotaceae PSI!olum nlldum S limit KNP

Ranuncu!aceae C!emalls !mcllnfo!w N limIt DHI Ranuni'ulus pumlllo Runul/('ulus H'sslhilorus

Restionaceae Desnloc{adus US/h'r Ins N limit DHI Desmocludus hi/flrmis Ins N limit KNP l::cdeim'o!eu nUJ!l(JSluchru N limit Tumala Stn Lepidob(J!us c!wcl(Jicplw!lIs N limit CNR Lepidoho!lIs densus ms N limit DHI Lepidohollls prelsslIlnlls N limit KNP Lept()curpus lirisltllus Lo),."OCorYli [lorthenicu IllS N limit KNP LYf.{inia hur!Jata N limit KNP

Rharnnaceae Cnpwndra arbullllora var horeahs Cryptandru cOIU/ala Cnplandra gllllmflora 2 Cryptandru IcwJphruc!u Cry/Jlandru mUfi!u N lunit DINR (',)ptandra mvriantha Cnpwndra nl/dlf!ora N limit DHI ('I)ptandra nulans Cryplandra pllngens Cr)ptandra scopariu var. ,nicrocephu!£l 2 ('ryptundra s/Jyridi(Jides Srenunrhenlurn ("(Jfnplicotum N limit DHI Stef/anthernum dll'uricUlum 3 S limit DHI ...... Stenanthemum tnfncatum N limit KNP 'J:) ------

Familyrraxon Conservation Distribution Conservation Reserve Code KNP TNR ZNP CNR FPNP BINR DINR KRNP

Stenanthemum notiale N IimitZNP Stenanthemum pomaderroides N limit KNP

Rubiaceae Canthium attenuatum Canthium latifo/ium Canthium /ineare Oldenlandia galioides Opercularia aff. spermacocea (A.H.Burbidge 4502) Opercularia spermacocea Opercularia vaginata N limit CNR Synaptantha tillaeacea Vat. tillaeacea

Rutaceae Boronia coerulescens subsp. spinescens N IimitZNP Boronia crenulata vat. gracilis N limit Carrang Stn. Boronia cymosa N limit Hamelin Stn. Boronia purdieana N limit Tamala Stn. Boronia ramosa subsp. anethifolia N limit KNP Diplolaena geraldtonensis ms Diplolaena grandiflora Diplolaena mollis ms NlimitDHI Eriostemon brucei Eriostemon deserti Eriostemon sericeu.l' Eriostemon sp. Kalbarri (GJK & NG 2034) Eriostemon tomentellus GeleZflowia verrucosa 3 NlimitDHI Phebalium tuberculosum

Santalaceae Anthobolus jiJVeolalus N limit FPNP Chorelrum prilzelii N limit KNP Exowrpos aphyllus Exocarpo.l' sparteus Leplomeria preissiana N limit Cobum Stn. Leplomeria spinosa Santalum acuminalum Santalum lanceolatum Santalum spicalum Santalum spicalum subsp. novo Endemic Study atea

Sapindaceae Aleclryon oleifo/ius subsp. oleifolius Diplopellis eriocarpa Diplopellis huegelii Vat. subinlegra N limit Nanga Stn. Diplopellis intermedia Vat. incana S limit KNP Dip/ope/liS illlermedw var illlermedw Dip/ope/us pcuo/uns N limitZNP Dodolluco umb/yophs'l/u Dodolluco oplera N limit DHI Dodolluf'o bu (Sorii/olio Dodo!laca ('acs[Jl{osa

[)odoflllca i!l(J('4U~l(}!i(1 N limit FPNP Dodolloeu /urrcoides N limit KNP Dodolloco lobu/olo Dodo!lllCa pctio!aris DodofW('O plfll/O/W Dodmweu !'IUrllIICue/o/w [)odonaca Vlscosa subsp l1ngustlsSltJW N limit FPNP Dodollucu "ISCOSU subsp, spululillu

Scrophulariaceae * Dischisma arenarium E/ucholollw IlIImil Glossosligmu diulldrum G/ossosligmu drummolldii Mimulus graCilis Mimulus repells P('plidlum uirhochetlum Peplidium muel/en PeplidlUm sp, (N T Burbidge & A Kain 8152) Sremodw /lorulell((i Stemodia grossa 5'temodlO \'lSCOsa Srngu curn/lora

Solanaceae Antlulcercis genislllides Allr/wcerCls IliClj(llw subsp, caldurlllla N limitCNR Anthoce rCls llier/oli" subsp iliCl/illiu N limit KNP Anthocercis mlncala N limitWHA AlIlllllcercis Ii{{oreu AIII!rocercis sp, Shark Bay (T,E,H,Aplin 3335) Endemic Cape Cuvier Tamala Stn, Antholroche rtl)'oJ1oroides 2 N limit Meadow Stn AIlI!rorroche H'U/CO{{II 3 Cyp!rullrhera racemosu * Durura /eichlwrdlil Duboisiu IwpH'oodll Lycium uustra.!e * Lycium ferocissimum Nicofiana (,llvico!a

* Nico/iaf/a ~ltiUC(J NicoriilflU occidentu/IS subsp hespens Nicoliullu occidcllwlis subsp obliquu Nicot{afla ro(undilo{ia Nico[iana simu/at1s Familyrraxon Conservation Distribution Conservation Reserve Code KNP TNR ZNP CNR FPNP BINR D1NR KRNP

* Solanum americanum Solanum cleislogamum Solanum diversiflorum Solanum esuriale Solanum hesperium Solanum horridum Solanum lasiophyllum * Solanum nigrum Solanum oldfieldii N limit S of WHA Solanum orbiculatum subsp. orbiculalum Solanum phlomoides Solanum slurtianum Solanum symonii

Stackhousiaceae Slackhousia dielsii Stackhousia megaloplera Stackhousia mOf!ogyna Stackhousia muricala Slackhousia pubescens N limit just S of WHA Stackhousia viminea Triplerococcus brunonis

Sterculiaceae Brachychiton gregorii Commersonia crispa Commersonia gaudichaudii Commersonia melanopelala N limit KNP Commersonia microphylla N limit KNP Guichenolia all micranlha (Wittwer 18(9) N limit Tamala Stn. Guichenolia ledifo/ia NlimitDHI Guichenolia macrantha Guichenolia micrantha Hannafordia quadrivalvis Keraudrenia hermanniifolia Keraudrenia inlegrifolia Lasiopetalum anguslifo/ium Lasiopetalum oldfieldii 3 ulSiopellllum opposilifo/ium 3 N limit DlNR Ru/ingia craurophylia Rulingia cygnortlm Ru/ingia densiflora Rulingia kempeana Ru/ingia IUleij]ora Rulingia malvifolia var. borealis NlimitDHI Wallheria indica Stylidiaceae Lel'ellhookw !Cl'tallt/w Levenh(wkia (Jctoma('Ulala N limit KNP StylidiulIl brurwllwllulIl subsp. hrWWllllllllllll N limit KNP StvlidlUlIl ca/caratulIl N limit KNP Stv!ldlUlIl crasstfiJllUlIl N limit Tamala Stn. StvlidlUIIl dlc/wtolllUIIl N limit KNP StylidiulIl ,ltspermulIl N limit Nerren Nerren Stn.lCNR StylidiulIl nome N limit KNP Stvlidium !eptophvllulIl N limit KNP Stylidium !lIllhatum N limit CNR

S~tylidiurnfWUTocarpum N limit Tamala Stn StylidlUlIl ptll/erUIIl N limit KNP Stylidium pU/JI/ierulIl N limit KNP StylidlUIIl repells N limit Tamala Stn. StylidiulIl sp Kalbam (ACarr 145)

Stylidiurn ~·~.'i!r())

Surianaceae Sty!obaslllm austra!e N limit Murchison House Stn Sty!obasiwll spathll!atwll

Tamaricaceae * Tamanx aphylla

Thelypteridaceae Cyclosol'Us Illterrllptlls

Thymelaeaceae Pillle!ca wllllloc/wris Pime!ca all/iUsti/illia N limit KNP Pillle!ea jlonhwlIja Pime!ea /ii!/iialla N limit DHI Pime!ea !ellCl.lIItha N limit Tamala Stn. Pime!ea lIlicrocel'ha!a subsp microcep/w!a Pillle/ea sessills N limit Tamala Stn Pime/ea tric/wstachya

Tiliaceae Corclwl'Us Ct!IZ!ll'hun/iilills Corclwrus e!achocarl'us Corchorlls sp. Camarvon (A.M.Ashby 2321) Corc/wrus wa!coll" Triwll(et/a sp. T Kimberley Flora

Tremandraceae Tetratheca hirsuta

Typhaceae Typlw d()mingensis Farnilyffaxon Conservation Distribution Conservation Reserve Code KNP TNR ZNP CNR FPNP BINR DINR KRNP

Urticaceae Parietaria cardiostegia Parietaria debilis • Urtica urens

Violaceae Hybanthus calycinus N limit Murchison House Stn. Hybanthus .floribundus subsp. .floribundus N IimitZNP

Vitaceae Clematicissus angustissima N limitZNP

Xanthorrhoeaceae Xanthorrhoea aff drummondii N limit KNP Xanthorrhoea preissii

Zannichelliaceae Lepilaena preissii

Zygophyllaceae Nitraria billardierei Tribulus astrocarpus Tribulus cistoides S limit DH! Tribulus .!(Jrrestii Tribulus macrocarpus S limit Harnelin Stn. Tribulus occidentalis Tribulus platypterus Tribulus sp. (KRN 11569) Tribulus terrestris Zygophylium aff. simile (RD Royce 5997) Zygophylium ammophilum Zygophylium apicularum NlimitDHl Zygophylium aurantiacum Zygophylium biliardierei NlimitDHl Zygophylium compressum Zygophylium eremaeum Zygophylium fruticulosum Zygophylium glaucum Zygophylium iodocarpum Zygophylium kochii Zygophylium lobularum Zygophylium ovatum Zygophylium retivalve S limit Carrang Stn. Zygophylium simile Zygophylium sp. Karratha (J .S.Beard 3508) Zygophylium tesquorum