Tulipa Sprengeri: the Long Lost Tulip
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Roozengaarde's Bulb Planting Guide
15867 Beaver Marsh Road Mount Vernon, WA 98273 Frequently 360-424-8531 or 866-4TULIPS Asked www.Tulips.com / [email protected] Questions WILL BULBS GROW ANYWHERE? Yes, bulbs will grow in many different climates. Warmer climates are more challenging and may require pre-cooling each fall prior to plant- ing. The year we ship your bulbs, we do any necessary pre-cooling here on our farm prior to shipment—your bulbs will be ready to plant when you receive them! DO BULBS NEED TO BE WATERED? Yes, if Mother Nature does not take care of it for you, then water during dry spells. Be sure not to oversoak the planting area. Water just enough so that it absorbs quickly. Water after planting, during growth, and even after topping if the soil is dry. WHAT IS THE BEST TYPE OF SOIL AND FERTILIZER? Soil needs to provide adequate drainage and oxygen for the bulbs. We recommend using a slow release fertilizer specifically designed for flower bulbs - usually a N - P - K formula. DO BULBS NEED TO BE DUG IN THE SUMMER? We recommend digging tulip bulbs each year after the foliage has died down naturally. They are more prone to disease and rot if left in the ground through the summer months. Other bulbs like daffodils can typically be dug on a 3-5 year schedule. If you experience diminished results in the spring, you should dig your bulbs that summer. WHAT IS THE BEST WAY TO STORE MY BULBS IF I DIG THEM? After digging, make sure to dry bulbs thoroughly. -
Liliaceae S.L. (Lily Family)
Liliaceae s.l. (Lily family) Photo: Ben Legler Photo: Hannah Marx Photo: Hannah Marx Lilium columbianum Xerophyllum tenax Trillium ovatum Liliaceae s.l. (Lily family) Photo: Yaowu Yuan Fritillaria lanceolata Ref.1 Textbook DVD KRR&DLN Erythronium americanum Allium vineale Liliaceae s.l. (Lily family) Herbs; Ref.2 Stems often modified as underground rhizomes, corms, or bulbs; Flowers actinomorphic; 3 sepals and 3 petals or 6 tepals, 6 stamens, 3 carpels, ovary superior (or inferior). Tulipa gesneriana Liliaceae s.l. (Lily family) “Liliaceae” s.l. (sensu lato: “in the broad sense”) - Lily family; 288 genera/4950 species, including Lilium, Allium, Trillium, Tulipa; This family is treated in a very broad sense in this class, as in the Flora of the Pacific Northwest. The “Liliaceae” s.l. taught in this class is not monophyletic. It is apparent now that the family should be treated in a narrower sense and some of the members should form their own families. Judd et al. recognize 15+ families: Agavaceae, Alliaceae, Amarylidaceae, Asparagaceae, Asphodelaceae, Colchicaceae, Dracaenaceae (Nolinaceae), Hyacinthaceae, Liliaceae, Melanthiaceae, Ruscaceae, Smilacaceae, Themidaceae, Trilliaceae, Uvulariaceae and more!!! (see web reading “Consider the Lilies”) Iridaceae (Iris family) Photo: Hannah Marx Photo: Hannah Marx Iris pseudacorus Iridaceae (Iris family) Photo: Yaowu Yuan Photo: Yaowu Yuan Sisyrinchium douglasii Sisyrinchium sp. Iridaceae (Iris family) Iridaceae - 78 genera/1750 species, Including Iris, Gladiolus, Sisyrinchium. Herbs, aquatic or terrestrial; Underground stems as rhizomes, bulbs, or corms; Leaves alternate, 2-ranked and equitant Ref.3 (oriented edgewise to the stem; Gladiolus italicus Flowers actinomorphic or zygomorphic; 3 sepals and 3 petals or 6 tepals; Stamens 3; Ovary of 3 fused carpels, inferior. -
Bulb Dormancy in Vitro—Fritillaria Meleagris: Initiation, Release and Physiological Parameters
plants Review Bulb Dormancy In Vitro—Fritillaria meleagris: Initiation, Release and Physiological Parameters Marija Markovi´c*, Milana Trifunovi´cMomˇcilov , Branka Uzelac , Sladana¯ Jevremovi´c and Angelina Suboti´c Department of Plant Physiology, Institute for Biological Research “Siniša Stankovi´c“—NationalInstitute of the Republic of Serbia, University of Belgrade, Bulevar Despota Stefana 142, 11060 Belgrade, Serbia; [email protected] (M.T.M.); [email protected] (B.U.); [email protected] (S.J.); [email protected] (A.S.) * Correspondence: [email protected] Abstract: In ornamental geophytes, conventional vegetative propagation is not economically feasible due to very slow development and ineffective methods. It can take several years until a new plant is formed and commercial profitability is achieved. Therefore, micropropagation techniques have been developed to increase the multiplication rate and thus shorten the multiplication and regeneration period. The majority of these techniques rely on the formation of new bulbs and their sprouting. Dormancy is one of the main limiting factors to speed up multiplication in vitro. Bulbous species have a period of bulb dormancy which enables them to survive unfavorable natural conditions. Bulbs grown in vitro also exhibit dormancy, which has to be overcome in order to allow sprouting of bulbs in the next vegetation period. During the period of dormancy, numerous physiological processes occur, many of which have not been elucidated yet. Understanding the process of dormancy will allow us to speed up and improve breeding of geophytes and thereby achieve economic profitability, which is very important for horticulture. This review focuses on recent findings in the area of Citation: Markovi´c,M.; Momˇcilov, bulb dormancy initiation and release in fritillaries, with particular emphasis on the effect of plant M.T.; Uzelac, B.; Jevremovi´c,S.; growth regulators and low-temperature pretreatment on dormancy release in relation to induction of Suboti´c,A. -
Guide to the Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- LILIACEAE
Guide to the Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- LILIACEAE LILIACEAE de Jussieu 1789 (Lily Family) (also see AGAVACEAE, ALLIACEAE, ALSTROEMERIACEAE, AMARYLLIDACEAE, ASPARAGACEAE, COLCHICACEAE, HEMEROCALLIDACEAE, HOSTACEAE, HYACINTHACEAE, HYPOXIDACEAE, MELANTHIACEAE, NARTHECIACEAE, RUSCACEAE, SMILACACEAE, THEMIDACEAE, TOFIELDIACEAE) As here interpreted narrowly, the Liliaceae constitutes about 11 genera and 550 species, of the Northern Hemisphere. There has been much recent investigation and re-interpretation of evidence regarding the upper-level taxonomy of the Liliales, with strong suggestions that the broad Liliaceae recognized by Cronquist (1981) is artificial and polyphyletic. Cronquist (1993) himself concurs, at least to a degree: "we still await a comprehensive reorganization of the lilies into several families more comparable to other recognized families of angiosperms." Dahlgren & Clifford (1982) and Dahlgren, Clifford, & Yeo (1985) synthesized an early phase in the modern revolution of monocot taxonomy. Since then, additional research, especially molecular (Duvall et al. 1993, Chase et al. 1993, Bogler & Simpson 1995, and many others), has strongly validated the general lines (and many details) of Dahlgren's arrangement. The most recent synthesis (Kubitzki 1998a) is followed as the basis for familial and generic taxonomy of the lilies and their relatives (see summary below). References: Angiosperm Phylogeny Group (1998, 2003); Tamura in Kubitzki (1998a). Our “liliaceous” genera (members of orders placed in the Lilianae) are therefore divided as shown below, largely following Kubitzki (1998a) and some more recent molecular analyses. ALISMATALES TOFIELDIACEAE: Pleea, Tofieldia. LILIALES ALSTROEMERIACEAE: Alstroemeria COLCHICACEAE: Colchicum, Uvularia. LILIACEAE: Clintonia, Erythronium, Lilium, Medeola, Prosartes, Streptopus, Tricyrtis, Tulipa. MELANTHIACEAE: Amianthium, Anticlea, Chamaelirium, Helonias, Melanthium, Schoenocaulon, Stenanthium, Veratrum, Toxicoscordion, Trillium, Xerophyllum, Zigadenus. -
Hill View Rare Plants, Summer Catalogue 2011, Australia
Summer 2011/12 Hill View Rare Plants Calochortus luteus Calochortus superbus Susan Jarick Calochortus albidus var. rubellus 400 Huon Road South Hobart Tas 7004 Ph 03 6224 0770 Summer 2011/12 400 Huon Road South Hobart Tasmania, 7004 400 Huon Road South Hobart Tasmania, 7004 Summer 2011/12 Hill View Rare Plants Ph 03 6224 0770 Ph 03 6224 0770 Hill View Rare Plants Marcus Harvey’s Hill View Rare Plants 400 Huon Road South Hobart Tasmania, 7004 Welcome to our 2011/2012 summer catalogue. We have never had so many problems in fitting the range of plants we have “on our books” into the available space! We always try and keep our lists “democratic” and balanced although at times our prejudices show and one or two groups rise to the top. This year we are offering an unprecedented range of calochortus in a multiplicity of sizes, colours and flower shapes from the charming fairy lanterns of C. albidus through to the spectacular, later-flowering mariposas with upward-facing bowl-shaped flowers in a rich tapestry of shades from canary-yellow through to lilac, lavender and purple. Counterpoised to these flashy dandies we are offering an assortment of choice muscari whose quiet charm, softer colours and Tulipa vvedenskyi Tecophilaea cyanocrocus Violacea persistent flowering make them no less effective in the winter and spring garden. Standouts among this group are the deliciously scented duo, M. muscarimi and M. macrocarpum and the striking and little known tassel-hyacith, M. weissii. While it has its devotees, many gardeners are unaware of the qualities of the large and diverse tribe of “onions”, known as alliums. -
Bio 308-Course Guide
COURSE GUIDE BIO 308 BIOGEOGRAPHY Course Team Dr. Kelechi L. Njoku (Course Developer/Writer) Professor A. Adebanjo (Programme Leader)- NOUN Abiodun E. Adams (Course Coordinator)-NOUN NATIONAL OPEN UNIVERSITY OF NIGERIA BIO 308 COURSE GUIDE National Open University of Nigeria Headquarters 14/16 Ahmadu Bello Way Victoria Island Lagos Abuja Office No. 5 Dar es Salaam Street Off Aminu Kano Crescent Wuse II, Abuja e-mail: [email protected] URL: www.nou.edu.ng Published by National Open University of Nigeria Printed 2013 ISBN: 978-058-434-X All Rights Reserved Printed by: ii BIO 308 COURSE GUIDE CONTENTS PAGE Introduction ……………………………………......................... iv What you will Learn from this Course …………………............ iv Course Aims ……………………………………………............ iv Course Objectives …………………………………………....... iv Working through this Course …………………………….......... v Course Materials ………………………………………….......... v Study Units ………………………………………………......... v Textbooks and References ………………………………........... vi Assessment ……………………………………………….......... vi End of Course Examination and Grading..................................... vi Course Marking Scheme................................................................ vii Presentation Schedule.................................................................... vii Tutor-Marked Assignment ……………………………….......... vii Tutors and Tutorials....................................................................... viii iii BIO 308 COURSE GUIDE INTRODUCTION BIO 308: Biogeography is a one-semester, 2 credit- hour course in Biology. It is a 300 level, second semester undergraduate course offered to students admitted in the School of Science and Technology, School of Education who are offering Biology or related programmes. The course guide tells you briefly what the course is all about, what course materials you will be using and how you can work your way through these materials. It gives you some guidance on your Tutor- Marked Assignments. There are Self-Assessment Exercises within the body of a unit and/or at the end of each unit. -
Thermonastic TROPISM
THERMOnastic TROPISM Eleni Katrini | Ruchie Kothari | Mugdha Mokashi Bio_Logic Lab School of Architecture, Carnegie Mellon University ABSTRACT Motivation: To develop dynamic elements which respond to changes in temperature Approach: The thermonastic movements of the Tulip flower were looked at for inspiration to develop conceptual systems based on movement and heat. The tulip flower opens and closes its petals based on the external temperature. The petals open up when the temperature is high and close when the temperature is low. This movement of the petals is facilitated by movement of water through the plant. Experiments were conducted using paper which is a fibrous material exhibiting properties similar to that of a flower petal. Water at different temperatures was studied as a conducting medium. Findings: Based on analysis, different forms of flowers were created. This includes flowers with varying surface areas, flowers made using composite materials, laminated and chiseled flowers, flowers with varying edge exposure etc. The study was focused on exploring the effects of capillary action on fibrous materials. The observations were focused on- - Range of movement of petals - Duration for capillary action - Effect of hot and cold temperature It was seen that the range of movement was the same for most of the prototypes. All the flowers other than the flowers for which the range of movement was restricted opened up to 180o. The time taken for the water to rise depended on the density and the surface area of the paper in contact with the water. Increase in density of the paper in contact with water, increases the time taken for the water to rise up. -
GENOME EVOLUTION in MONOCOTS a Dissertation
GENOME EVOLUTION IN MONOCOTS A Dissertation Presented to The Faculty of the Graduate School At the University of Missouri In Partial Fulfillment Of the Requirements for the Degree Doctor of Philosophy By Kate L. Hertweck Dr. J. Chris Pires, Dissertation Advisor JULY 2011 The undersigned, appointed by the dean of the Graduate School, have examined the dissertation entitled GENOME EVOLUTION IN MONOCOTS Presented by Kate L. Hertweck A candidate for the degree of Doctor of Philosophy And hereby certify that, in their opinion, it is worthy of acceptance. Dr. J. Chris Pires Dr. Lori Eggert Dr. Candace Galen Dr. Rose‐Marie Muzika ACKNOWLEDGEMENTS I am indebted to many people for their assistance during the course of my graduate education. I would not have derived such a keen understanding of the learning process without the tutelage of Dr. Sandi Abell. Members of the Pires lab provided prolific support in improving lab techniques, computational analysis, greenhouse maintenance, and writing support. Team Monocot, including Dr. Mike Kinney, Dr. Roxi Steele, and Erica Wheeler were particularly helpful, but other lab members working on Brassicaceae (Dr. Zhiyong Xiong, Dr. Maqsood Rehman, Pat Edger, Tatiana Arias, Dustin Mayfield) all provided vital support as well. I am also grateful for the support of a high school student, Cady Anderson, and an undergraduate, Tori Docktor, for their assistance in laboratory procedures. Many people, scientist and otherwise, helped with field collections: Dr. Travis Columbus, Hester Bell, Doug and Judy McGoon, Julie Ketner, Katy Klymus, and William Alexander. Many thanks to Barb Sonderman for taking care of my greenhouse collection of many odd plants brought back from the field. -
Generic Classification of Amaryllidaceae Tribe Hippeastreae Nicolás García,1 Alan W
TAXON 2019 García & al. • Genera of Hippeastreae SYSTEMATICS AND PHYLOGENY Generic classification of Amaryllidaceae tribe Hippeastreae Nicolás García,1 Alan W. Meerow,2 Silvia Arroyo-Leuenberger,3 Renata S. Oliveira,4 Julie H. Dutilh,4 Pamela S. Soltis5 & Walter S. Judd5 1 Herbario EIF & Laboratorio de Sistemática y Evolución de Plantas, Facultad de Ciencias Forestales y de la Conservación de la Naturaleza, Universidad de Chile, Av. Santa Rosa 11315, La Pintana, Santiago, Chile 2 USDA-ARS-SHRS, National Germplasm Repository, 13601 Old Cutler Rd., Miami, Florida 33158, U.S.A. 3 Instituto de Botánica Darwinion, Labardén 200, CC 22, B1642HYD, San Isidro, Buenos Aires, Argentina 4 Departamento de Biologia Vegetal, Instituto de Biologia, Universidade Estadual de Campinas, Postal Code 6109, 13083-970 Campinas, SP, Brazil 5 Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611, U.S.A. Address for correspondence: Nicolás García, [email protected] DOI https://doi.org/10.1002/tax.12062 Abstract A robust generic classification for Amaryllidaceae has remained elusive mainly due to the lack of unequivocal diagnostic characters, a consequence of highly canalized variation and a deeply reticulated evolutionary history. A consensus classification is pro- posed here, based on recent molecular phylogenetic studies, morphological and cytogenetic variation, and accounting for secondary criteria of classification, such as nomenclatural stability. Using the latest sutribal classification of Hippeastreae (Hippeastrinae and Traubiinae) as a foundation, we propose the recognition of six genera, namely Eremolirion gen. nov., Hippeastrum, Phycella s.l., Rhodolirium s.str., Traubia, and Zephyranthes s.l. A subgeneric classification is suggested for Hippeastrum and Zephyranthes to denote putative subclades. -
Liliaceae Lily Family
Liliaceae lily family While there is much compelling evidence available to divide this polyphyletic family into as many as 25 families, the older classification sensu Cronquist is retained here. Page | 1222 Many are familiar as garden ornamentals and food plants such as onion, garlic, tulip and lily. The flowers are showy and mostly regular, three-merous and with a superior ovary. Key to genera A. Leaves mostly basal. B B. Flowers orange; 8–11cm long. Hemerocallis bb. Flowers not orange, much smaller. C C. Flowers solitary. Erythronium cc. Flowers several to many. D D. Leaves linear, or, absent at flowering time. E E. Flowers in an umbel, terminal, numerous; leaves Allium absent. ee. Flowers in an open cluster, or dense raceme. F F. Leaves with white stripe on midrib; flowers Ornithogalum white, 2–8 on long peduncles. ff. Leaves green; flowers greenish, in dense Triantha racemes on very short peduncles. dd. Leaves oval to elliptic, present at flowering. G G. Flowers in an umbel, 3–6, yellow. Clintonia gg. Flowers in a one-sided raceme, white. Convallaria aa. Leaves mostly cauline. H H. Leaves in one or more whorls. I I. Leaves in numerous whorls; flowers >4cm in diameter. Lilium ii. Leaves in 1–2 whorls; flowers much smaller. J J. Leaves 3 in a single whorl; flowers white or purple. Trillium jj. Leaves in 2 whorls, or 5–9 leaves; flowers yellow, small. Medeola hh. Leaves alternate. K K. Flowers numerous in a terminal inflorescence. L L. Plants delicate, glabrous; leaves 1–2 petiolate. Maianthemum ll. Plant coarse, robust; stems pubescent; leaves many, clasping Veratrum stem. -
Tecophilaeaceae 429 Tecophilaeaceae M.G
Tecophilaeaceae 429 Tecophilaeaceae M.G. SIMPSONand P.J. RUDALL Tecophilaeaceae Leyb., Bonplandia JO: 370 (1862), nom . cons . Cyanastraceae Engler (1900). Erect, perennial, terrestrial herbs. Roots fibrous. Subterranean stem a globose to ellipsoid corm, 1- 4 cm in diameter, in some genera with a membra nous to fibrous tunic consisting of persistent sheathing leaves or fibrovascular bundles . Leaves basal to subbasal, or cauline in Walleria, spiral; base sheathing or non-sheathing, blades narrowly linear to lanceolate -ovate, or more or less petiolate in Cyanastrum and Kabuyea; entire, glabrous, flat, or marginally undulate; venation parallel with a major central vein. Flowers terminal and either Fig. 122A-F. Tecophilaeaceae. Cyanastrum cordifolium . A Flowering plant. B Tepals with sta mens. C Stamens. D Pistil. E solitary (or in small groups) and a panicle or (in Ovary, longitudinal section. F Capsule. (Takh tajan 1982) Walleria) solitary in the axils of cauline leaves. Bracts and bracteoles (prophylls) often present on pedicel. Flowers 1- 3 cm long, pedicellate, bisexual , trimero us. Perianth variable in color, zygomor fibrous scale leaves or leaf bases or the reticulate phic or actinomorphic, homochlamydeous, ba fibrovascular remains of these scale leaves (Fig. sally syntepalous; perianth lobes 6, imbricate in 2 123). The tunic often continues above the corm, in whorls, the outer median tepal positioned anteri some cases forming an apical tuft. Corms of orly; minute corona appendages present between Walleria, Cyanastrum, and Kabuyea lack a corm adjacent stamens in some taxa. Androecium aris tunic (Fig. 122). ing at mouth of perianth tube, opposite the tepals Leaves are bifacial and spirally arranged. -
Networks in a Large-Scale Phylogenetic Analysis: Reconstructing Evolutionary History of Asparagales (Lilianae) Based on Four Plastid Genes
Networks in a Large-Scale Phylogenetic Analysis: Reconstructing Evolutionary History of Asparagales (Lilianae) Based on Four Plastid Genes Shichao Chen1., Dong-Kap Kim2., Mark W. Chase3, Joo-Hwan Kim4* 1 College of Life Science and Technology, Tongji University, Shanghai, China, 2 Division of Forest Resource Conservation, Korea National Arboretum, Pocheon, Gyeonggi- do, Korea, 3 Jodrell Laboratory, Royal Botanic Gardens, Kew, Richmond, United Kingdom, 4 Department of Life Science, Gachon University, Seongnam, Gyeonggi-do, Korea Abstract Phylogenetic analysis aims to produce a bifurcating tree, which disregards conflicting signals and displays only those that are present in a large proportion of the data. However, any character (or tree) conflict in a dataset allows the exploration of support for various evolutionary hypotheses. Although data-display network approaches exist, biologists cannot easily and routinely use them to compute rooted phylogenetic networks on real datasets containing hundreds of taxa. Here, we constructed an original neighbour-net for a large dataset of Asparagales to highlight the aspects of the resulting network that will be important for interpreting phylogeny. The analyses were largely conducted with new data collected for the same loci as in previous studies, but from different species accessions and greater sampling in many cases than in published analyses. The network tree summarised the majority data pattern in the characters of plastid sequences before tree building, which largely confirmed the currently recognised phylogenetic relationships. Most conflicting signals are at the base of each group along the Asparagales backbone, which helps us to establish the expectancy and advance our understanding of some difficult taxa relationships and their phylogeny.