Hymenoptera, Braconidae) of Finland, Part 2

@EntomoldgicaFennica. 15 October 1999

A survey of Opiinae (Hymenoptera, Braconidae) of Finland, part 2

Maximilian Fischer & Martti Koponen

Fischer, M. & Koponen, M. 1999: A survey of Opiinae (Hymenoptera, Braconidae) of Finland, part 2. - Entomol. Fennica 10: 129-160.

Faunistic records of 150 species of Opiinae are given mainly for Finland. 78 species are reported as new to Finland. The gross distributions and known hosts are mentioned. Psyttalia cariiiata (Thomson), Biosteres (Chilotrichia) bicolor (Wesmael) and B. (C.) sylvaticiis (Haliday) are deleted from the fauna of Fin- land. Paraphytomyza similis (Brischke) is a new host for Opiiis (Odontopoea) connivens Thomson, and Eziphranta connexa (Fabricius) (Diptera, Tphritidae) for Psyttalia concolor (Szépligeti).

Maximilian Fischel; Natural History MLrseum Vienna, 2. Zoological Department, Biirgring 7, A-1014 .Eenna, Airstria Martti Koponen, Department ofApplied Zoology, ?? O. Box 27, FIN-O0014 Uni- versiíy of Helsinki .

Received 1 Febrcrary 1998, accepted 2 Jdy 1999

Subgenus Odontopoea Fischer Host. in Finland reared from Diptera, Agro- myzidae: Paraphytomyza similis (Briscke) by E. Odontopoea Fischer, 1986: 610. Type species Thuneberg (new host). Opizrs epcilatirs Papp. Marerial examinen. 1d and 2 P from Finland (DAZH, MZH), N: Vantaa, 22.9.1979 (M. Koponen); Sa: Joutseno (E. Thuneberg); Obb: Ranua, 19.7.1980 (M. Koponen leg.), and Opius (Odontopoea) connivens Thomson 1 0 from Estonia, Laanemaa: Puhtu, 29.5.1990 (M. Kopo- nen leg.). Opiirs (Nosopaeus) connivens Thomson, 1895: 2190,?. Opius (Odontopoea) eprilatiis Pupp Opiirs coiznivens; Fischer 1959g: 70; 197 1a: 56. Opiiis (Nosopoea) connivetis; Fischer 1972: 292 Opiirs eprilatcrs Papp, 19SOa: 50, 53. 55; 1981b: (key), 306 (redescription); Papp 1979: 75. 261, C. . OpiT1c (Ai!ocp!iy)~-J:;,~;*PT;?: T~hii,s 19P6: 79 -. 9piií.b íGJc)ii;~)ji~~.!.(i)~piL:~i/~í~; i i>Liici 1986: GiC Opiiis ( Ociontopoeci) coriviiiieris;Fischer 1956: (key). 626 (redescription). 610. Distribution. Finland (map 72), Hungary. Opiirsfoveola Fischer, 1957b: 2 14. Synonymised A new species to Finland. by Fischer 197 1a. Host. Unknown. Distribcitiorz. Denmark, Estonia, Finland (map Material esrzniined. 1 d frorn Finland íDAZH), N: Hel- 7 l), Hungaq, Russia, Sweden. sinki, Viikki, 27.6.19S2 (Y.Zhongqi leg.). A new species t:, Estonia 2nd Finland. 130 - ' Fischer & Koponen: A survey of Opiinae ENTOMOL. FENNCA Vol. 10

Opius (Odontopoea) paranivens Fischer Subgenus Tolbiu Cameron

Opius (Odontopoea) paranivens Fischer, 1990: Tolbia Cameron, 1907: 85. Type species: Tolbia 223, Q. scaevolae Cameron. Distribution. the Canary Islands (Tenerife), Finland (map 73), Hungary. Opius (Tolbia) caesus Haliday A new species to the Canary Islands and Fin- land. Opius caesus Haliday, 1837: 215, 8; Marshall Host. Unknown. 1891: 45, O 8;Fischer 1958b: 192; Hellén 1959: Material examined. 2 O from Finland (DAZH), N: Hel- 21; Fischer 1967b: 140; 1971a: 49. sinki, 11.7.1978, NurmijarVi, 15.9.1984 (M. Koponen); and Opius (Tolbia) caesus; Fischer 1972: 347 (rede- 18,40 from ihe Canary Islands (DAZH), Tenerife: Sta Ur- scription); Papp 1982: 243; Tobias 1986: 73 (key, sula, La Quinta, 8.-11.12.1993 (M. &ponen leg.). distribution), Opius (Allotypus)punctiventris Thomson, 1895: Subgenus Stornosem¿z Fischer 2189, ? d. Synonymised by Fischer 1971a: 347. Opius subtilis Szépligeti, 1898: 400. Synonymised Stomosema Fischer, 1972: 330. vpe species: by Fischer 1958b: 200 with punctiventris. Opius mutus Fischer. Opius hydrelliae Rimsky-Korsakow, 1925: 69. Synonymised by Fischer 1971 a: 347. Opius (Stomosema) gyoerfii Fischer Opius hydrelliae Muesebeck, 1933: 197, ? . Pre- occupied. Opius gyOrfiiFischer 1958b: 190, O ;Hellén 1959: 22. Opius aquaticus Muesebeck 1967: 48 (new name Opius gyoerfii; Fischer 1971 a: 7 1. for hydrelliae Muesebeck). Synonymised by Opius (Stomosema) gyoerfii; Fischer 1972: 330 Fischer 1987: 31. (redescription); Tobias 1986: 70. Opius hydrellianus Fischer, 1971a: 74 (new name Distribution. Finland (map 74), Hungary. for hydrelliae Muesebeck). Host. Unknown. Distribution. Austria, Bulgka, former Czech- Material examined. 2 O from Finland 0,N: Deger6 oslovakia, Denmark, England, Finland (map 76), (A. Saarinen), Ekeniis, Notholm (Hakan Lindberg). France, Germany, Hungary, Ireland, Romania, Russia, Sweden, Caucasus, Nepal, Syria, Opius (Stomosema) mutus Fischer Tadzhikistan, North America. Hosts. Diptera, Agromyzidae: Phytomyza Opius mutus Fischer, 1964b: 142; Hellén 1966: primulae Robineau-Desvoidy; Ephydridae: Hy- 85; Fischer 1967a: 340; 1971a: 90. drellia cochleariae Haliday, Hydrellia griseola Opius (Stomosema) mutus; Fischer 1972: 330 Fallén on Holcus mollis; Drosophilidae: Scapto- (key), 332 (redescription); Tobias 1986: 70; Ko- myza graminum Fallén. ponen 1989: 29. Material examined. 1 O from Finland (MZH), St: Re- Distribution. Austria, Denmark, Finland (map posaan, 30.9.1950 (V. Lauro leg.). 75), Germany, Russia (Leningrad region, Hellén 1966). Opius (Tolbia) rudiformis Fischer Hosts. Diptera, Agromyzidae: Agromyza hen- deli Griffiths on Phragmites australis, Anthomy- Opius rudifonnis Fischer, 1958b: 201, 9 ; 1964a: iidae: Delia quadripila (Stein) on Honckenya 200,395 (8 nov.); 1971a: 106. peploides. Opius (Tolbia) rudiformis; Fischer 1972: 347 Murerial examined. 1 d,30 from Finland @AZH, MZH), (key), 349 (redescription); Tobias 1986: 73. N: Hanko, 20.8.1978, 17.6.1980 (M. Koponen), Helsinge = Opius (Gastrosema)rudifonnis; Papp 1981b: 269. Helsinki rural municipality W. Helién leg.. paratype); St: Opius rotundifonnis; Hellén 1959: 22 (mis- Sakyla, 13.6.1982 (M. Koponen); and 1 ? from Russia (MZH, spelling). paratype), Leningrad region: Tenjoki (W. Hellén leg.). . EhTT0MOL.r FENNICA Vol. 10 0 Fischer & Koponen: A survey of Opiinae 131

L 3 Distribiition. Austria, the Canary Islands, Opiiis (Lissosema)parvungula; Fischer 1972: 359 former Czechoslovakia, Denmark, England, Fin- (key), 361 (redescription); Papp 1980b: 205. land (map 77), France, Italy, Russia, eastern Can- Opius (Tolbia) parvungiiliis; Tobias 1986: 76. ada. Opiris docilis Fischer 1958b: 157 (partim) nec A new species to the Canary Islands. Haliday, 1837. Synonymised by Fischer 1971a: Host. Unknown. 361. Marerial examined. 3 O frorn Finland (DAZH, MZH), Distribution. Austria, former Czechoslovakia, Al: Lernland, 12.24.8.1954 (Hakan Lindberg); Kb: Pielis, Denmark, England, Estonia, Finland (map 79), (Gronvik); Lkoc: Kolari, (M. Ko- 29.7.-23.8.1865 9.7.1989 France, Germany, Hungary, Ireland, Poland, Swe- .ponen leg.); and ld frorn the Canary Islands (MZH), Lan- den; North Korea, Russian Far East (Vorosilov). zarote: Penhas del Chacha, 13.-22.3.1963 (Lindberg leg.). A new species to Estonia and Finland. Hosts. Diptera, Agromyzidae: Agromyza al- Subgenus Gastrosema Fischer bipennis Meigen on Phalaris arundinacea, Agro- myza graminicola Hendel on Phragmites austra- Gastrosema Fischer, 1972: 353. Type species: lis, Agromyza lucida Hendel on Deschampsia Opius pumilio Wesmael. caespitosa, Agromyza nigrella Rondani on Triti- Opius (Gastrosema) docilis Haliday cirm sp., Agromyza ambigua Fallén = niveipen- nis Zetterstedt on Secale cereale, Agromyza 6 reptans Fallén on Urtica dioica, Cerodontha Opius docilis Haliday, 1837: 2 11, @; Hellén 1946: 133; Fischer 1958b: 197; 1971a: 62. (Cerodontha) denticornis (Panzer) on Holcus Opius (Therobolus) docilis; Thomson 1895: 21 83, Eanatus. Marerial examined. 5 d ,5 O frorn Finland (DAZH, MZH), 9 d. Ab: Pojo, Aminne, 14.7.1965 (M. Meinander), Viiti, Siika- Opiiis (Gastrosema) docilis; Fischer 1972: 353 jarvi, 9.8.1964 (V. J. Karvonen); N: Helsinki, 11.7.1978, (key), 355 (redescription); Papp 1979: 77. 13.9.1979,Nurmijmi, 6.9.1986,Vantaa, 24.8.1979 (M.Ko- Opius (Tolbia) docilis; Tobias 1986: 76 (key, dis- ponen); St: Reposaari, 24.8.1954 (V.Lauro); Sa: Mikkelin mllc., 16.8.1976,11.8.1987, Pertunrnaa, 22.6.1983 (M. Ko- tribution). ponen); and ld frorn Estonia (DAZH), Saarernaa: Kaali, Distribrition. Austria, England, Finland (map 30.5.1990 (M. Koponen leg.). 78), France, Germany, Hungary, Poland, Russia, Sweden, Switzerland; Georgia, Kirgiziya, Russian Subgenus Opirrs Wesmael, s. str. Far East. Host. Diptera, Agromyzidae: Agromyzaphrag- Opius Wesmael, 1835: 115. Type species: Opius initidis Hendel on Phragmites australis. pallipes Wesmael. Material e-ramined.3 d,4 O frorn Finland (DAZH, MZH), Ab: Nystad = Uusikaupunki, 1923 (W.Hellén), Vihti, Siika- jarvi, 18.7.1961 (V. J. Karvonen); N: Helsinge, 6.8.-21.9.1915 Desmatophorus Thomson, 1895: 2 194. Type spe- íW’. Hellén). Helsinki (R. Frey), 24.7.1979 (M. Koponen); cies: Bracon pygtnaeator Nees. Sa: Mikkelin mlk., 24.8.1975 (M. Koponen); Kb: Joensiiii, 28.8.1924 (E.Thuneberg leg). Opiiis (Opius) agromyzicola Fischer p6 Subgenus Lissosema Fischer Opiiis agromyzicola Fischer, 1967a: 320, E@; 1971a: 40: 1974: 82. Lissosemcr Fischer: 1972: 359. Type species: Opiirs Opiirs (Opirrs) crgromyzicoli;; Fischer 1972: 366 pal-i)ungriln Thomson. (key), 367 (redescription); Tobias 1986: 78. Distribrition. Austria, England, Finland (map Opiris (Lissosema)parvririgrila Thomson SO), Sweden. A new species to Finland and Sweden. Opius (Therobolus)paniingrrla Thomson, 1895: Host. Diptera, Agromyzidae: Agromyza poten- 2183, 9 d. tillae fKaltenbach) = spiraeae Kaltenbach on Opius panwgula; Fischer 197 1a: 96; Zaykov Filipencliila i.*ulgaris. 1983: 36. Material examined. 1 d, 1 O frorn Finland (DAZH), N: 1 ‘132 *’ Fischer & Koponen: A srirvey of Opiirrne 0 ENTOMOL. FENNICA Vol. 10

Nurmijarvi, 7.8.1980 (M. Koponen); Ks: Kuusamo hdateiial exaniined. 4a, 39 from Finland (DAZIi), N: (7353:609), 3.7.1979 (M. Koponen); and Id from Sweden Nurmijib-vi, 30.5.1983,25.5.1988,5.8.1994, Sipoo, 3.8.1982; (DAZH), Nb: OvertomeA, 27.6.1979 (M. Koponen leg.). Kb: Liperi, 2.-3.7.1994;Om: Vimpeli, 8.7.1985 (al1 M. Ko- ponen); and 18 from Greece (DAZH), Corfu: Benitses, Opius (Opius) breviscapus Thomson 27.5.1981 (M. Koponen leg.). Opius (Opius) circinus Papp Opius (Desmatophorus) breviscapirs Thomson, 1895: 2197, ? d. Opius circiniis Papp, 1979b: 347, 9 1980a: Opius breviscapus; Fischer 1966: 9; 1967b: 139; 8; 104; 1981a: 128. 1971a: 48; 1974: 82.. Opius (Opius)circinus; Tobias 1986: 80; Fischer Opius (Opius) breviscapus; Fischer 1972: 366 1986: 620 (redescription). (key), 368 (redescription); Papp 1979: 73; Tobias Distribution. Finland (map 83), Hungary. 1986: 80. A new species to Finland. Distribution. Austria, Finland (map 81), Es- tonia, Hungary, Ireland, Russia, Sweden, Mon- Host. Unknown. Material examined. 1 O from Finland (DAZH), N: Nur- golia. mijarVi, 7.8.1980 (M. Koponen leg.). A new species to Estonia and Finland. Hosrs. Diptera, Agromyzidae: ? Phytomyzn Opius (Opius) exilis Haliday afinis Fallén on Euphrasia sp. Material e.rnrnined. 3 d , 209 from Finland (DAZH, MZH), Al: Brando, 11.7.1921 , Jomala, Kungso, 9.6.1939 Opiiis exilis Haliday, 1837: 208, ? ;Fischer 1960: (M. Hellén), Saltvik, 3.7.1984 (M. Koponen); Ab: Gustavs = 59, 3 d (redescription, figures); 1971a: 64. Kustavi, 1921 (W. Hellén), Pargas (Ingelius), Pojo, hnne. Opius (Pendopius) exilis; Fischer 1972: 410; Papp 14.7.1965 (M. Meinander); N: Nunnijiirvi, 28.7.1994,Tam- 1979: 78; 1981~:148. misaari, 14.8.1986 (M. Koponen); S€: Rauma, 1.-4.7.1986 (M. Koponen); Sa: Mikkelin mlk., 24.8.1975 (M. Koponen); Opius (Phaedrotoma) exilis; Tobias 1986: 85. IU: Parikkala, 15.7.1940 (W. Hellén); Ok: Kajaani, 16.8.1982 Opius (Opius) exilis; Fischer 1992: 48. (M. Koponen); Oba: Vaala, 14.8.1982 (M. Koponen); Lkoc: Opius (Desmatophonrs) piceus Thomson, 1895: Kittila 17.8.1983, 8.7.1989 (M. Koponen), and 1E from Es- 2198, ?. Synonymised by Fischer 1991b: 48. tonia (DAZH), Paniumaa: Massiaru, 4.6.1990(M. Koponen Opius piceus; Marshall 1898: 313, 8; Hellén leg.). 1959: 22; Fischer 1960: 51 (figs.); 1971a: 99. Opius (Opius) caricivorae Fischer Opius (Opius) piceus; Fischer 1972: 366 (key), 381 (redescription); Papp 1979b: 347; 1981c: 141; Fischer 1984a: 130; 1986: 656; Tobias 1986: 78 Opicrs caricivorae Fischer, 1964d: 1, ? d ; 1967b: 140; 1971a: 51. (key, distribution). Opius (Opius)caricivorae; Fischer 1972: 366 Opius (Pseudopius) extusus Papp, 198 lc: 140, (key), 370 (redescription); Papp 1981c: 139; Za- 148. Synonymised with piceiis by Fischer 1984a: ykov & Fischer 1983: 46; Tobias 1986: 80 (key, 130. distnbution). Opius (Pendopius)cisromensis Papp, 1982b: í 11. Distribiition. Finland (map 821, Germany, Synonymised with piceus by Fischer 1986: 656. Greece, Poland, Russia, Ukraine, North Korea. Disrribution. North, west and central Europe, A new species to Finland and Greece. including Finland (map 84), Italy, Balkans, Rus- si,, G:o$a. Kazakhstan. Norih Knrea. eostern Hosts. Diptera, Agromyzidae: Agromyza nigrescens Hendel, Cerodontha (Butomyza) angu- Canada, northwestern U.S.A. lata (Loew), Cerodontha (Dizygornyza) caricico- Hosts. Diptera, Agromyzidae: Lirioriiyza pli- la (Hering) and Cerodontha (Butomyza) eucari- si0 (Meigen) on Tragopogon prntensis, Ophio- cis Nowakowski on Carex hirta, Dizygornyza (Bu- niyia niaura (Meigen) on Lnthyrus torreyi; Dro- tomyza) caricivora (Groschke), Napomyza salvi- sophílidae: Scaptomyza grnminum Fallén on An- ae Hering on Salvia verticillata, Phytomyza lap- thyllis vulneraria. Material examinen. 298, from Finland (DAZH, pae Goureau on Arctium sp., Phytomyza sp. on 429 MZH), Ab: Karjalohja, 19.6.1993(M. Koponen), Pargas (ln- Caltha palustris, Drosophilidae: Scaptomyza api- gelius), Nystad = Uusikaupunki, 5.8.-17.9.1914(W. Hellén), calis Hardy on Amaranthus blitum. Vichtis = Vihti (R. Frey); N: Hanko, 17.6.1980, Helsinki, ENTOMOi. FENNICA Vol. 10 0 Fischer & Koponen: A survey of Opiinae 133

.) 28.6.. 9.-24.8.1976, 26.6.1980, 15.9.1983. Kirkkonunimi, si&,Spain, Sweden, Switzerland, Israel, Mongo- 24.7.1991,Nurmijarvi, 20.7.1975, 1.8.1976, 14.7.,2.9.1979, lia. 7.-24.8.1980,17.7.1981, 28.7.. 25.8., 14.9.1987, 16.6.1991, 17.7.,5.8.1994, Porvoon mik.. 16.6.1979, Vantaa, 3.8.1975, Hosts. Diptera, Agromyzidae: Liriomyza con- 24.8.1979 (M. Koponen); Ka: Ylhaa, 13.8.1987 (M. Kopo- gesta (Becker) on Vicia faba, Liriomyta piisilia nen); Ta: Janakkala, 24.8.1980 (M. Koponen), Juupajoki, Meigen on Hieracium pilosella, Ophiomyia sp. 15.6.1982 (Y. Zhongqi), Jamsa, 26.8.1991,Lahti, 15.8.1980 on Medicago sativa, Phytomyza atricornis Mei- (M. Koponen); Sa: Anttola, 15.8.1976,Joutsa, 22.9.1978, Mikkelin mik., 2.9.1974, 22.8.1975,24.7.1976, 26.8.1978, gen on Tarnxacum ofcinale. Weber; Cecidomyi- 22.6.1979,21.6., 29.7.1980, 23.6.1983, 10.8.1987,23.7., idae: Asphondylia verbasci Vallot. 4.9.1989, 27.6.1992, Ristiina, 23.6., 28.7.1979, 22.6., Material examined. 940 279 from Finland (DAZH, 31.7.1980,23.7.1985, 19.7.1987 (M. Koponen); Oa: Narpes, MZH), Al: VArdo, 8.7.1984(M. Koponen); Ab: Lohjan kun- 31.7.1950 (G. Nordstrom),Ylih&rma, 13.7.1980;Om: Himan- ta, 29.5.1992, Vihti, 30.5.1979 (M. Koponen); N: Espoo, ka, 22.7.1980; Ok: Kuhmo, 8.7.1980; Obb: Tornio, 6.7.1982 29.5.1986 (M. Koponen), Hango = Hanko, 1931 (W. Hel- (M. Koponen); Ks: Kuusamo, 19.8.1965 (M. Meinander), En), 20.8.1978, 19.5.1984 (M. Koponen), Helsinki, 22.5, 4.7.1979 (M. Koponen); Lkoc: Kittila, 8.7.1989 (M. Kopo- 7.9.1976,6.6.1979,15.-30.5.1983,15.5.-7.6.1984,4.8.,25.8., nen leg.); 18 from Estonia (DAZH), Laanemaa: Puhtu, 14.9.1987,2. -16.6. 1991, 14.-31.5.1992,5.8.1994, Sipoo, 29.5.1990 (M. Koponen); 1 O from Georgia (MZH), pr Fi. 3.6.1982, 15.6.1983, Tammisaari, 20.5.1984, Vantaa, Hrami, 22.4.1958 (Lindberg); 18 1 O from Russia (MZH), 10.8.1980(M. Koponen); Ta: Hattula, 11.6.1987, Hameen- Karelia: Gumbaritza (S. Platonoff); 18 from Sweden linna, 22.5.1982,Janakkala, 26.5.1982,26.5.1984 (M. Ko- (DAZH), Vb: Brbnland, 14.7.1980 (M. Koponen leg.); 1 8 ponen); Sa: Hirvensalmi, 5.6.1982,Mikkelin mik., 31.5.1980, from Germany (DAZH), Mecklenburg, 2.9.1895 (Fr. W. 4.6.1983,31.5.1988,3.6.1994 (M. Koponen); Tb: Rautalam- Konow); 1 O from Poland (DAZH), Pomorze: Pietrzykowo, pi, 19.7.1983 (M. Koponen); Oba: ii, 20.7. 1980 (M.Kopo- 13.7.1988 (M. Koponen); and 18 from Greece (DAZH), nen); 78, 49 from Estonia (DAZH), Harjumaa: Lahemaa Corfu: Perama, 29.5.1978 (M. Koponen leg.). andvihasoo, 25.5. 1991;Paniumaa: Nigula, 3.6.1990;Saare- maa: Asuka and Kuressaare, 2.6.1990(M. Koponen); 1 O from Opius (Opius) gigapiceus Fischer Sweden (MZH), Gotland (A. Nordman); and 18 from Swit- zerland (MZH), Münster Tal, Santa Maria, 19.-22.6.1953 (Hakan Lindberg). Opius (Opius)gigapiceus Fischer, 1990: 2 11, 3 d. Distribution. Czechia, Finland (map 85), Hun- Opius (Opius)meracus Fischer gary. A new species to Finland. Opius meraciis Fischer, 1960: 38, 3 ;Hellén 1959: Host. Unknown. 22; Fischer 1962b: 97 (d nov.); 1971a: 87. Material examined. 1 O from Finland (DAZH), Kb: Kesalah- Opius (Opius)meracus; Fischer 1972: 366 (key), ti, 17.7.1982 (M. Koponen leg.). 375 (redescription); Tobias 1986: 76 (key); Ko- Opius (Opius) lugens Haliday ponen 1989: 29. Distribution. Denmark, Estonia, Finland (map Opicis lugens Haliday, 1837: 206, 3 ;Hellén 1959: 87), Germany, Hungary, Italy, Russia (Karelia, 22; Fischer 1960: 36 (figs.); 1971a: 83. Murmansk region, Krasnodar region), Slovenia. Opius (Opius) lugens; Fischer 1972: 366 (key), Hosts. Diptera, Agromyzidae: Napomyza ni- 373 (redescription); Tobias 1986: 78, 92; Papp gritrila (Zetterstedt) = cineracea (Hendel) on Ra- 1988: 58 (distribution). nunciilus acris, Nnpomyza ? evanescens (Hendel) Opius (Desmatophorus) abscissiis Thomson, on Raniinciiliis lanuginosus. Material e.rarnined. 6d,6 9 from Finland (DAZH, MZH), 1895: 2 196, 9 d.Synonyrnised by Fischer 1971 a: N: Espoo, 29.5.1986 (M. Koponen), Hango = Hanko, 373. 14m31 (Y ~m~~),1~1.~3,w s.19~ (M ~.:oporicn), üpius nbscissiis; Fischer 1960: 41 (redescription). Kyrkslatt = KirAkonunimi (R. Frey), Numiij&vi, 7.6.i9S4 Opius obscurus Szépligeti, 1901: 152, 8 ;Fischer (M. Koponen), Tvarminne, 20.6.1966 (V. Varis); Ta: Lammi, 7.7.1976 (M. Koponen); Kl: Parikkala, 7.-14.6.1940(S. Hel- 9). 1960: 47 (redescription, Synonymised by En); Ok: Ristijbi, 3.7.1991 (M. Koponen), Suomussalmi, Fischer 1971a: 373. 16.6.-3.7.1917 (W. Hellén); Li: Utsjoki (R. Frey); 10 from Opius adveniens Fischer 1960: 43, d. Syno- Estonia (MZH), “St. RAgo” = Suur-Pakri island, 5.6.1933 (H. nymised by Fischer 1971a: 373. Backlund); 28 frorn Russia (MZH), Karelia: Sortavala, 4.6.1928 (Rantalainen), Murmansk region: Ylaluostari, 1930 Distribution. Austria, Denmark, England, Es- (M. Hellén); and 1 Ofrom Slovenia (MZH), Laibach = Ljublja- tonja, Finland (map 86), Germany, Ireland, Rus- na, 1870 (J. A. Palmén). , 134 ~ 1' Fischer & Koponen: A sriwey of Opiinae ENTOMOL. FENNICA Vol. 10

o Opius (Opius)pallipes Wesmael (Zetterstedt); Drosophilidae: Scaptomyza apica- lis Hardy on Amaranthiis blitum, Scaptomyza Opicis pallipes Wesmael, 1835: 118, 9 8 ; Hellén graminiim Fallén on Anthyllis vulnerarin;Tephriti- 1959: 22; Fischer 1960: 49 (figs.); 1971a: 95; dae: Myiolia caesio Hanis, Philophylln heraclei Wharton 1987: 66. (Linnaeus), Vidalia spinifrons Schroder. Hypolabis pallipes; Foerster 1862: 260. Material examined. 64 6,74 O from Finland (DAZH, MZH), Al: Foglo, 9.7.1984,Geta, 29.6.1983 (M. Koponen), Opiiis pallidipes Marshall, 1891: 27. 9 8. Jomala, 9.6.1939 (W. Hellén); Ab: Pojo, Aminne, 14.7.1965 Opiiis (Opiiis)pnllipes; Fischer 1972: 366 (key), (M. Meinander); N: Espoo, 10.8.1980 (M. Koponen), Hels- 377 (redescription); Tobias 1986: 78 (key, distri- ingin pit., 18.6.1920 (Y. Hukkinen), Helsinki, 28.6.,20.7., bution); Papp 1982b: 106. 9.8.1976, 18.6.1979, 2.8.1982,Kirkkonummi, 23.7.1991, Distribution. North, west and central Europe, 24.7.1994,Mantsaia, 24.8.1975,Nurmijarvi, 7.8.1976, 24.8.1980. 1.6.,22.6., 28.8.1982,5.-21.6., 18.8.1984, 16.6., including Finland (map 88), Italy, Balkans, Rus- 26.6.,6.-13.7., 8.8.1985, 25.7.1986, 22.7.,4. -14.9.1987, sia, Ukraine, central Urals, Caucasus, Kazakhstan, 27.6.1988, 22.6., .13.9.1989, 14.7.1991,13.6., 5.7., 18.7., Mongolia, North Korea, Siberia, westem North 9.8.1992,3. -5.8.1994,Pyhtaa, 14.8.1981 (M. Koponen), Sib- Amenca. bo, Nevas, 7.-13.7.1986(A. Albrecht), Sipoo, 13.8.1977,1.6., 15.6.1983,Tammisaari, 13.-14.8.1986,Vantaa, 13.8.1977, Hosts. Fischer & Koponen: A survey of Opii- 8.9.1978, 17.8.1979 (M. Koponen); Ka: Kotka, 31.7.1977 nae . Diptera, Apromyzidae: Agromyza nigrescens (M. Koponen); St: Euta, 7.7.1981, 16.6.1982, Huittinen, Hendel, Agromyza spiraeae on Filipendula vul- 29.5.1993,Kokemaki, 30.5.1993,Loimaan mlk., 30.5.1993, garis, Cerodontha (Butomyza) angzrlata (Loew) Rauma, 1.7.1986,Vammala, 2.7.1993,YlZne,17.6.1982 (M. Koponen); Ta: Hartola, 13.6.1987,Heinolan mik., 20.7.1983, or Cerodonta (Dizygomyza) niorosa (Meigen) on Janakkala, 24.8.1980,26.5.1982, 12.6.1983,Korpilahti, Carex hirta, Amaiiromyza (Trilobomyza) labia- 20.8.1986,Lammi, 6.4.7.1975,4. -15.7.1976,Loppi, 12. tariim (Hendel), Amaiiromyza (Trilobomyza) ver- 7.1991, Nastola, 25.6.1994,Palkane, 23.7.1980 (M. Kopo- basci (Bouché), Liriomyzn amnrellae Hering on nen); Sa: Imatra, 8.-11.6.1956(Hakan Lindberg), Mikkelin mlk., 24.7.1976,7.7.1978, 29.7., 16.8.1950,4.-24.6.1983. Gentiana gerníanica, Liriornyzn bryoniae (Kalten- 29.7.1985, 10.9. 1994, Pertunmaa, 22.6.1983, Ristiina, bach) on Po~c~~c~sberolinensis, Liriornyza centau- 28.7.1979, 4.7.1980, 1.8.1981, 5.6., 20.8.1983, 21.6.,25.7. reae Hering, Lirioinyza eupatoriana Spencer, Lir- 1985, 10.7.1993,22.7.1994 (M.Koponen); K1: Parikkala, iomyzn sp. on Crepis praemorsn, Liriomyza stri- 2.7.1980, 27.-28.6.1990 (M. Koponen), Uukuniemi, 20.7.1983 (M. Pulkkinen); Sb: Temo, 18.7.1983 (M. Kopo- gata (Meigen) on Hyoscyamus niger and Senecio nen); Kb: Kitee, 22.7.1982,Liperi, 2.-5.7.1994,Tohmajirvi, nemorensisfiichsii, ?Liriomyza strigata (T ileigen) 18.-23.7.1982(M. Koponen); Om: Pietarsaan, 30.6.1978 (M. on Hypochoeris aetnensis, Liriornyza taraxaci Koponen); Ok: Puolanka, 29.6.1989 (M. Koponen); Ks: Ku- Hering, Liriomyza wachtli Hendel = veratri Gro- usamo, 4.7.1979 (M. Koponen), Kuusamo, Juuma, 22.7.1934 (W. Hellén); Lkor: Sodankyla, 4.7.1989 (M. Koponen); 2 O schke; Paraphytomyza tremiilae (Hering), Phyto- from Estonia (DAZH), Saaremaa: Maebe, 31.5.1990;Vaipa- rnyza nngelicne Kaltenbach on Arigelicn sylves- maa: Koikküla, 5.6.1990 (M. Koponen); Id from Sweden tris, Phytornyzn fnllnciosn Bnschke, Phytomyza (MZH), Og: Rystad, 15.-17.6.1933(Hakan Lindberg); 48, feriilae Hering on Biiriiiim creticiim, Phytoinyza 3 Q from Poland (DAZH); 1 d from Russia (MZH, Murmansk region); 1 O from Slovenia (MZH), Laibach = Ljubljana, 1870 h?fliisnrin Hering, Phq'toinyza leiicnntheini Her- (J. A. Palmén leg.); and 39 from Greece (DAZH), Corfu: ing on Leiicaiitheiniim mnxirniirn, Phytoinyzn piil- Acharavi, 21.4.1994,Benitses, 23.5.19s1 (M. Koponen 1eg.j. liila Zetterstedt = rnatricarine Hendel, Phytornyzn ininusciiln Goureau on Thnlictriim rninus, Phyto- Opius (Opius)peterseni Fischer niyzn obsciira Hendel on Peiicednniim oreo- ~~ii!j;~ii~i,Plij [o,/ij..li y'iii1ilot.it.i I-Ici-iLi 011 ,411- Upiiisyerer-setii Fischer, 1964b: 242. Y d' ; 197 1 a: theiii is tinctorin, P1i';tomyzn ran ii~cu li (Schrank) 98. on Rnnuricriliis acris, Phgtomyza sp. on Arrernone Opiiis (Opiiis)peterseiii; Fischer 1972: 366 (key), sylvestris, Phytornyzn sphonclyiii Robineau-Des- 380 (redescription); Papp 1979: 86; Tobias 1986: voidy, Phytomjzn tannceti Hendei, Phytornyzn 80. tlíysselirii Hering on Peiicednnrrin palustre, Para- Distribution. Austria, Denmark, Finland (map phytorriyzn hencleliaiia (Hering), Phytoinyza per- 89), Hungary. iclymeríii de Meijere; Anthomyiidae: Anthornyia Host. Unknown. sp., Delia echirintn Séguy, Pegornya bicolor (Hoff- Material exainined. 2 O from Finland (DAZH), Sa: mannsegg in Wiedemann), Pegoinq'n nigritcirsis Mikkelin mlk., 5.6.1983 (M. Koponen); Oba: Ii, 20.7.1980

--. EN?IOM.OL. FENNICA Vol. 10 0 Fischer & Koponen: A siirvey of Opiinne 135

(M. Koponen leg.). 387 (key), 389; Papp 1982a: 24;; Tobias 1986: 80 (key, distribution). Opius (Opius)pygmaeator (Nees) Opiiis semiaciciilatcis Stelfox, 1949: 308, ? d. Synonymised by Fischer 1957a: 5 1. Bracon pygmaeator Nees, 1834: 52, ? d. Distribution. North and central Europe, includ- Opiiispygmaeator; Haliday 1837: 205, ? 0;Hel- ing Finland (map 92), Armenia, Georgia, Kirgiz- lén 1959: 22; Fischer 1960: 54; 1971a: 103. iya. Reported from Finland by Hellén (1959). Opiiis (Desmatophoriis) pygmaeator; Thomson Hosts. Diptera, Agromyzidae: Cerodontha 1895: 2195, 9 d. (Cerodonta) denticomis (Panzer) on Holccis lana- Opius (Opicts) pygmaeator; Fischer 1972: 366 tcis, Ophiomyia sp. on Medicago sativcim, Phyto- (key), 383 (redescription); Tobias 1986: 80. rnyza grifithsi Spencer on Plantago media, Phy- Distribcition. Austria, Belgium, former Czech- tomyza plantaginis Robineau-Desvoidy on Plan- oslovakia, England, Finland (map 90), Germany, tago lanceolata, Phytomyza ramosa Hendel, Russia, Sweden, Georgia. Phytomyza siiccisae Hering. Host. Uncertain. Material aamined. 2 d, 3 9 from Finland (DAZH, MZH), Material examined. 58, 27 9 from Finland (DAZH, Al: Lemiand, Flaka, 12.-24.8. 1954 (HAkan Lindberg); Ab: MZH), Al: Geta, Hockbole, 16.6.1939, Kokar, 17.7.1936, Houtskar, 5.7.1936 (W. Hellén); N: Nurmijarvi, 29.5.1988, Kokar, Ido, 18.7.1936 (W. Hellén); Ab: Nystad = Uusikau- Porvoo, 1.8.1982 (M. Koponen); Ta: Palkane, 23.6.1921 (M. punki, 1920,1923 (W. Hellén); N: Hango = Hanko, 14.6.1931 Hellén); 1G from Estonia (DAZH), Laanemaa: Puhtu, (W. Hellén), HyvinkXa, 8.8.1976 (M. Koponen), Tvarminne, 29.5.1990 (M. Koponen leg.); and 1E from Holland (MZH). 13.-30.7.1914 (W. Hellén); Ka: Hamina, 30.7.1953 (E. Thuneberg); St: Karkku, 12.-20.6.1914 (W. Hellén); Ta: Opiris (Phaedrotoma) diversus Szépligeti Ruovesi, 25.6.193 1 (V. Saarinen); Sa: Hirvensalmi, 31.7.1983, Mikkelin mlk., 1.-2.8.1983, Ristiina, 23.7.1994 (M. Koponen); K1: Parikkala, 16.-27.6.1940 (S. Hellén), Opicis diversus Szépligeti, 1898: 308, 400, ? d; 21.7.1942 (W. Hellén), UuEruniemi, 25.7.1940 (W. Hellén); Fischer 1960: 102 (redescription). Sb: Kuopio, 27.6.1989 (M. Koponen); Kb: KontioiakdLibe- Opicis (Phaedrotoma) diversus; Papp 1979: 77. lits, 1 1.-25.6.1865 (Woldstedt); Obb: TorneA = Tornio, 4.7.1924 (HAkan Lindberg); Lkoc: Kittila, 8.7.1989 (M. Ko- Opiiis diversipes; Hellén 1959: 2 1 (misspelling). ponen); Lkor: Sodankyla, 9.8.1983 (M. Koponen); and 28, Treated as synonym of Opiiis exigciiis Wesmael 4 O from Russia (MZH), Leningrad and Murmansk regions. by Fischer (1971a and 1972) and Tobias (1986). Fischer (1990) is still uncertain about the inde- Opiiis (Opius) subpallipes Tobias pency of this species. Distribrition. Austria, Bulgaria, the Canary Is- Opicis (Opiiis) scibpallipes Tobias, 1986: 80. - lands, former Czechoslovakia, England, Estonia, Fischer 199 1a: 19 1 (redescr.). Finland (map 93), Germany, Greece, Hungary, Distribiition. Finland (map 9 1), Moldova. Italy, Sweden, Uzbekistan. A new species to Finland. A new species to the Canary Islands and Host. Unknown. Greece. Material exarnined. 1 9 from Finland (DAZH), Ta: Jana- Host records mixed with those of Opiiis e.xig- kkala, 2.8.1975 (M. Koponen leg.). 1111s. Material examined. 22 8, 22 9 f from Finland (DAZH, Subgenus Phaedrotoma Foerster MZH), Al: Lemland, 6.7.1984, Sund, 1.7.1983 (M. Kopo- nen); Ab: Pargas (Ingelius), Pernio, 17.9.1994 (M. Koponen); Ii. Ii~i~idi,71.7.i97.í, 12.S.1977, 2-+.’,.iS7&,iO.S.I9SC, Phaedr~toninFoerster, 1 562: 260. Type species: 4.8.1952, 16.6.1983, 23.8.1981, Nurmijiirvi, 25.7.1975, Phaedrotoma depeciilntor Foers ter. 15.7.1978, 14.7.1979, 14.7.1952, 20.5., 19.6.. 6.8.1983, 24.8.1985, 12.7.,25.8.1987, 14.8.1989,16.6.1991.19.6.1993, Opiiis (Pltaedrotoma)depeculator (Foerster) 5.8.1994, Porvoon mlk., 30.7.1978, Vantaa, 6.8.1950 (M. Ko- ponen); St: Rauma. 1.7.1986, Vammala, i .7.1993 (M. Kopo- nen); Ta: Forssa, 11.8.1992, Hartola, 10.6.1987, Heinolan Phnedrotoma depeciilntor Foerster, 1862: 260. rnlk., 21.8.1983, Korpilahti, 20.8.1986 (M. Koponen); Sa: Opiiis depeciilator; Hellén 1959: 22; Fischer Mikkelin mlk., 19.8. 1994, Ristiina, 9.9.1990, Ruokolahti, 1957a: 51; 1971a: 60. 2.7.1975 (hl. Koponen); 18, 29 from Estonia (DAZH), Saaremaa: Montu park 31.5.1990, Asuka and Lihulinna, Opiits (Phnedrotoma) depeciilator; Fischer 1972: 2.6.1990 (M. Koponen); 2d, 49 from the Canary Islands ~ r36 ,’ Fischer & Koponen: A suwey of Opiinae ENTOMOL. FENNICA Vol. 10

O (DAZH), Tenerife: Anaga, 4.12.1992, Sta Ursula, 8,- 1986: 82. 11.12.1993 (M. Koponen); Greece (DAZH), Corfu: Perarna, Opicis (Opiothorax) affectcis Papp, 1981 c: 139, 28.5.1977, Acharavi, 24.4.1994 Koponen); 1 frorn Ita- (M., d 144, O d. Synonymised by Fischer (1 984a). ly (MZH), Tneste, Tirnavo, 1870 (J. A. Pairnén); and 1 O frorn Georgia (DAZH), Batumi, 9.7.1977 (M. Koponen lep.). Distribution. Austria, former Czechoslovakia, England, Finland (map 93, France, Germany, Opius (Phaedrotoma) exiguus Wesmael Poland, Russia, Switzerland, Kazakhstan, North Korea. Opius exiguus Wesmael, 1835: 123, E; Hellén A new species to Finland. 1959: 22; Fischer 1960: 104 (redescription); Hosts. Diptera, Agromyzidae: Amauromyza 1971a: 64. (Trilobomyza) verbasci (Bouché) on Verbascum Opius (Phaedrotoma)exiguus; Fischer 1972: 387 thapsiforme, Phytomyzn agromyzina Meigen, (key), 392; Papp 1981c: 140; Tobias 1986: 82; Phytol iriomyza hila rella (Zetters ted t) on Pterid- Papp 1981c: 140; 1982c: 184,191; 1988: 58 (dis- ium aquilinzirn. In DAZH there is 10 from Fin- tribution) . land, reared at the Agricultura1 Research Centre, Distribution. North, west and central Europe, Department of Pest Investigation, from a sample including Finland (map 94), Greece, Italy, Rus- of Hordeiim leaves, with larvae and puparia of sia, Caucasus, Israel, Kazakhstan, North Korea, Phytomyza sp. Egypt, Ethiopia, South Africa. Material emrnined. 2 9 frorn Finland (DAZH), N: Nur- mijarvi, 28.9.1980 (M. Koponen); St: Kokemaki, 1960, Host records mixed with those of Opius di- reared. versus. Material examined. 226. 599 frorn Finland (DAZH, Opirrs (Phaedrotoma) variegatus Szépligeti MZH), Al: Geta, 29.6.1983, Hammarland, 1.7.1984 (M. Ko- ponen); Ab: Nauvo, Seili, 17.-18.7.1978, Pernio, 6.8.1982 (M. Koponen); N: Hanko, 17.6.1980, 20.5.1989, Helsinki, Opiiis variegatus Szépligeti, 1896: 3 15,381, EG; 11.9. 1977, 28.7.1980, 25.8.1982, 16.6.1983, 30.5.1984, Hellén 1959: 22; Fischer 1960: 107; 1967a: 347; Kirkkonurnmi, 23.5.1991, Nurrnijarvi, 14.7.1979, 10.- 1971a: 121. 29.7.1982, 14.6.-7.7.1983, 7.6.1984, 11.6., 6.9.1986, 23.6., 14.9.1987, 17.7.. 7.6.1991,25.7., 16.8.1992,20.5.,4.7.1993, Opicis (Phaedrotorna) variegatus; Fischer 1972: 5.8. 1994, Siuntio, 20.8.1978, Tarnrnisaari, 20.5.1984, 387 (key), 401 (redescription); Tobias 1986: 82 12.8.1986 (M. Koponen); Ta: Hattula, 1.8.1992, Janakkala, (key , distribution) . 12.6.1988, Jarnsa, 7.7.1982, Loppi, 1.8.1992, Palkane, Distribution. Austria, former Czechoslovakia, 1 1.7.1985, Ruovesi, 1 1.7.1985 (M. Koponen); Sa: Mikkelin mlk., 23.6., 31.7. 1983, Rautjmi, 3.7.1990, Ristiina, 27.6.- Denmark, England, Estonia, Finland (map 96), 1.7., 9.9. 1990 (M. Koponen); Tb: Rautalampi, 15.7.1983 France, Germany, Hungary, Poland, Russia, cen- (M. Koponen); Sb: Nilsia, 17.6.1988, Pieksarnaen rnlk., tral Urals, Sweden, Kazakhstan, Mongolia. 14.7.1983 (M. Koponen); Kb: Liperi, 24.6.1982, 5.7.1994, Hosts. Diptera, Agromyzidae: Agromyza al- Tohrnajarvi, 21.7.1982 (M. Koponen); Ok: Puolanka, 29.6.1989 (M. Koponen); Obb: Kemi, 6.7.1982 (M. Kopo- bitarsis Meigen, ?Agromyza apfelbecki Strobl on nen); Ks: Taivalkoski, 11.8.1983 (M. Koponen); Id from Ci rs ii im e r ioph 3 riiin , Ag romyza ph rag m itidis Russia (MZH), Leningrad region: Tytasaari, 16.7. 1935 (Vi’. Hendel, Cerodontlia (Poemyza) descharpsiae Hellén): 58, 69 frorn Estonia (DAZH), Laanernaa: Puhtu, (Spencer) on Descliampsia caespitosa, Cero- 28.-29.5.1990, Saaremaa: Kaali, Kübassaare and Maebe, 30.- 3 1.5.1990 (M. Koponen); 12, 1 ? from Poland (DAZH), Po- dontha (Poeinyzci) iricisa (Meigen) on Calama- rnorze: StaraBrda, il.-i2.7.1988(M. Koponen); ld,30 from grostis canescens, Calarnagrostis epigeios, Mel- Grrece, Corfu: Benitses, 25-31.5. 1951. Perarna, 27.5.1951 ica nritnns and Phlerrrii pmterrse, Ceroclorztlia (Po- (A: Kui,di.?iL), c.2iJ i fid,iii C,oiLi.i (iI!tZIl), ELitlxiili, erii?*,íci)pJgriicieíi (Meigen) on tlrncli>pocliiinrsyí- 11.7.1977 (M. Koponen lee.). saticiiiti, Calcitnciyrostis nriindinacea, Dactylis glomeratn. and Deschanipsia caespitosa, Cero- Opius (PIIaedrotoma)prrlchriventris Fischer dorithci (Britornom~zi)scirpi (Karl) on Scirpris Opiiis prilchriventris Fischer, 1958c: 301, O 0; sylvaticiis, Dizygornyza sp. on Brachypodiiiin syl- 1960: 106 (figs.); 197 1a: 102. vaticurti, Amaiirornyci (Trilobomyza) Pavifrons Opiiis (Phaedrotoma) piilchriiyentris; Fischer (Meigen) o11 Dianthiis barbntiis, Dianthiis cary- 1972: 3S7 (key), 397 (redescnption); Papp 198 Ic: ophylliis, Lychnis coronarici, Melandriiiin album, 141; 1982b: 107; Fischer 1984a: 131; Tobias Melandriiirn riibriitn, Saponaria oficinalis and ENTOMQL'. FENNICA Vol. 10 O Fischer & Koponen: A survey. of Opiinne 137

o Stellnrin sp., Amauromyza (Trilobomyza) verbasci 1!#4 (M.Koponen); St: Vammala, 30.6.1993 (M. Koponen); (Bouché) on Scrophcilaria nodosa, Verbascum Ta: Lammi, 6.7.1976 (M. Koponen); Kb: Polvijiirvi, 5.7.1994 (M. Koponen); Om: Reisjwi, 11.7.1980 (M. Koponen); Ok: nigrum., Verbascumphlomoides, Verbascum thnp- Kajaani, 5.7.1991 (M. Koponen); Id, 19 from Estonia siforme Schrad. and Verbascum thapsus; An- (DAZH), Saaremaa: Kaali 30.5.1990, LOpollu, 31.5.1990 íM. thomyiidae: Pegomyn hyoscyami (Panzer) on Koponen), and 1 d, 1 9 from Greece (DAZH), Corfu: Achar- Hyoscyamus niger. avi, 23.4.1994, Benitses, 24.5.1981 (M. Koponen leg.). Material examined. 66, 159 from Finland (DAZH, hfZH, TUU), Al: Foglo, 9.7.1984 (M. Koponen); Ab: Korp- Subgenus Allophlebus Fischer poo, 4.7.1958, Nauvo. 8.7.1948 (E. K. Lahtinen); N: Hango = Hanko, 1931 (W. Hellén), Helsinki, 18.9.1975 (M. Vii- tasaan),24.8.1976, 12.8.1979,26.6.1980,8.9.,30.9.1982(M. Allophlebus Fischer, 1972: 416. Type species: Koponen), Nurmijarvi, 20.6.1975, 15.7., 31.8.1979, 9.6., Opicis victus = singularis sensu Fischer unti11992 16.7.1994, Sipoo, 15.6. 1983 (M. Koponen), Tvarminne, (see Fischer 1995: 238). 14.8.1955 (Hakan Lindberg); Ta: Loppi, 29.8.1993 (M. Ko- ponen), Nokia, 5.8.1940 (W. Hellén); Sa: Mikkelin mlk., 73.6.1983 (M. Koponen); Kb: Lipen, 4.8.1993 (M. Kopo- Opius (Alloph1ebus)frrscipennis Wesmael nen); 2d from Estonia (DAZH), Saaremaa: Salme, 31.5.1990, Viidumagi, Sihi, 1.6.1990 (M. Koponen); ld from Poland Opiusfiiscipennis Wesmael, 1835: 123, 9 ;Fischer (DAZH), Pomorze: Stara Brda, 12.7.1988 (M. Koponen); and 1960: 72 (figs.); 1971a: 68; Papp 1979c: 185; 1 9 from Russia (MZH), Leningrad region: Terijoki, 1982c: 190. 12.8.1927 (W. Hellén leg.). Opiiis (Allophlebus) firscipennis; Fischer 1972: Subgenus Pendopius Fischer 416 (key), 420 (redescription). Opicis (Phnedrotoma)jiiscipennis; Tobias 1986: Pendopius Fischer, 1972: 409. Type species: Opius 87. pendulus Haliday. Distribution. Austria, Belgium, former Czech- Subgenus Pendopius; Fischer 1991 b: 47 (new oslovakia, England, Finland (map 98), Greece, definition). Hungary, Moidova. A new species to Finland. Opirrs (Pendopius) pendulus Haliday Host. Diptera, Agromyzidae: Agromyzn nigro- ciliata Hendel on Arrhenatherzim elnticis. Material examined. 4 8,69 from Finland (DAZH, MZH), Opicis penduliis Haliday, 1836: 205, O O ; Hellén Ab: Nauvo, Sando, 19.7.1978; N: Nurmijiirvi, 13.6.1992, 1959: 22; Fischer 1960: 66 (redescription); 1964a: 3.8.1994; Sa: Mikkelin mlk., 22.8.1975, 18.7.1987, Ristii- 200; 1971a: 97; van Achterberg 1997: 64. na, 23.6.1979, 19.7.1987; Sb: Tervo, 18.7.1983; Kb: Tohma- Opiiis (Pendopius) pendulus; Fischer 1972: 4 1O jarvi, 25.7.1982 (al1 M. Koponen leg.). (key), 414 (redescription); 1992: 47; Papp 1979: 85. Opiris (Allophlebus)padidalis Fischer Opiiis (Phnedrotonia)pendulus; Tobias 1986: 85 (key. distribution). Opiirs (Allophlebus)padidnlis Fischer, 1989: 96, Disti-ibution. Austria, England, Estonia, Fin- o. land (map 97), France, Germany, Greece, Hun- Distribution. Finland (map 99), Slovakia. gary, Ireland, Russia, Sweden, west Caucasus, A new species to Finland. Canada (Ont., Que.), U.S.A. (N.Y.). Host. Unknown. Alnrc.rirr1 rrorniiird 1 d froni Finlnnd (DAZH).Sa. Ris- A new species to Estonia and Greece. tiina, G.S.1977 (hl. Koponen leg.). Hosts. Diptera. Agromyzidae: Lirionivza con- gesta (Becker) on Vicia sepiiim, Napoiiiym ern- Opirrs (Allophlebus)pitlcherrini us Fischer nescens (Hendel) on Raniiiicrilus lnniiginosirs, Naponryzn nigritiila (Zetterstedt) = Phytomyzcl Opiiis piilcherrimiis Fischer, 1958f 176, O d; cineracea Hendel on Ranunculus acris and R. 1971a: 102. bulbosils. Opius (Allophlebus)pulcherrinius; Fischer 1972: hfmrrial examined. 46. 6 O from Finland (DAZH, MZH), N: Helsinki, 28.6.1976, 8.6.1979, Inkoo, 21.5.1989 (M. Ko- 416 (key), 422 (redescription). ponen), Kyrkslatt = Kirkkonummi (R. Frey), Nudjiirvi, 5.8. Opius (Phnedrotoma)prilcherriinus; Tobias 1986: - 138 ,. Fischer & Koponen: A survey of Opiinae 0 ENTOMOL. FENNICA Vol. 10 * 89 (key, distribution). Opiiis (Allophlebus) victus: Fischer 1995: 238. Distribution. Austria, Finland (map lOO), Ger- Opiiis singiilaris (sensu Fischer, nec Wesmael): many, Hungary, Moldova, Russia, Ukraine (Cri- Hellén 1959: 22; Fischer 1960: 76, 9 d (figs.); mea) . 1962d: 89; 1971a: 111. A new species to Finland. Opiiis (Allophlebus) singcilaris (sensu Fischer, nec Host. Unknown. Wesmael); Fischer 1972: 416 (key), 424 (rede- Material examined. 1 O frorn Finland (DAZH), Sa: scnption); papp 1982a: 245. Mikkelin mlk., 6.9.1981 (M. Koponen leg.). Opiirs (Allophlebus) tarni Papp, 1982: 249. . Atomorlis victus; van Achterberg 1997: 87 (new Opiiis (Allophlebus) pulchriceps Szépligeti status & new combination, not applied here). Treated as synonym for Opiiis (Allopitlebiis) OPitls PulchricePs SzéPligeti, 1898: 4009 5' singcilaris Wesmael, 1835 by e.g. Fischer (1972) Fischer 1960: 74 (redescription); 1971a: 102. and Tobias (1986). Opiiis (Allophlebus) pulchriceps; Fischer 1972: Distribiction. Austria, Belgium, Bulgaria, 416 (keY), 423 (redescnption); PaPP 1981~141. former Czechoslovakia, Denmark, England, Fin- OPiLlS (PhaedrOtoma) PulchricePs; Tobias 1986: land (map 103), Germany, Hungary, Italy, Poland, 89. Romania, Sweden, Switzerland, Armenia, Geor- Distribution. Austria, Finland (map lOl), Ger- gia, Mongolia. many, Hungary, North Korea. Hosts. Diptera, Agromyzidae: Agrornyza abi- Host. Unknown. ens Zetterstedt, Agromyza alnibetiilae Hendel on Material examined. 2 O frornFinland (DAZH), Ab: Siirk- ~l~~~~ incana, A~~~~~~~anthracina ~~i~~~, isalo, 2 1.9.1994; N: Nurmij&vi, 16.7.1994 ( both M.Kopo - nen leg.). Agrornyza arrinci Hering on Ariincirs viilgaris, Agromyza nigripes Meigen on Agrotis canina, Opiiis (Allophlebiis) staryi Fischer Agromyza reptans Fallén, Agromyza urficneNow- akowski on Urtica dioica, Cerodontha pygmaea Opilcs staryi Fischer, 1958~:303, 3 ; 1971a: 113; Meigen On DnCtyliS gfomerü?B,Liri0m)'za amoe- Papp 1979c: 185. na Meigen, Liriomyza eupatorii Kaltenbach on Opiirs (Allophlebus) staui; Fischer 1972: 41 6 Gdeopsis tetrahit, Phytorriyza actaeae Hendel on (key), 427 (redescription); Papp 197%: 348,352; Actaea spicata, Phytomyta angelicastri Henng on 1981c: 141. Angelica sylvestris, Phytoinyza anthrisci Hendel Opiirs (Phaedrotoma) staryi; Tobias 1986: 87. on Anthrisccis sylvestris, Phytomyza heracleana Opiirs (Allophlebus) sitagnis papp, 1982b: 117, Henng on Peucedanlim ostrllthilim, Rilbiomyza O. Synonymised by Fischer 1986: 656. hendeliana Henng on Lonicera pericljmenium, Distribrition. former Czechoslov&a, England, Rerbiornyza periclyrnenii Meijere on Lonicera per- Finland (map 102), Hungary, Sweden, Noflh Ko- icl~iizenicri?i,Trilobomyzn labiatanirn Hendel on rea. Stachys sjlvatica. Unknown hosts on Heracleiim A new species to Finland and Sweden. sp. and Holccis lanatus. Hosts. Diptera, Agromyzidae: Paraphytornyza Mnterial examined. 238, 41 O from Finland (DAZH, MZH). Al: Saltvik, 30.6.1983 (M. Koponen); Ab: Karjalo- treirritkne(Hering) onPopuzirstrerTiuki, Phytom)?za hja, 19.6.1992 (M. Koponen), Lohjan kunta, 21.6.1991, aql(ilfQ?iaeHardY on Aqllilegia sp., Ph)'to~?~)'Zfl19.6.1992 íM Koponen), Pojn, Aniinne, 14.7 1965 (A4 Mein- niiniisclclCi Goureau on Aqcli/egia sp. and 7yzalic- iinder). Abo = ?'urku (K. i-rey), Vichtis = Vihti (K. Frey) trirrii sp. Vastanfjard, 6.8.1983 (M. Koponen); N: H:fors = Helsinki Frey), 19.8.1975, 21.9.1976. 27.7.. 4.-15.9.1978 (19 Af

L 1979, 17.8.1980, 11.-12.8. 1987, 10.9.1994, Rstiina, 14.6.1986 (A. Albrecht), Tacktom, 9.6.1955 (V. Nystrom), 25.7.1985,4.7.1987 (M. Koponen);Tb: Atinekoski, 21.8.1986 and 1 O from Estonia (DAZH), Laanemaa: Puhtu, 29.5.1990 (M. Koponen); Kb: “Carel. Bor.” (E W. Woldstedt); Ok: Kuh- (M. Koponen leg.). mo, 10.7.1980, Suomussalmi, 29.6.1991 (M. Koponen); Obb: Ranua, 19.7.1980 (M. Koponen); and 28 from Estonia (DAZH), Paniumaa: Nigula, 3.6.1990, Saaremaa: Viiduma- Opius (Hypocynodus) mongolaltaiensis Fischer gi, Sihi, 1.6. 1990 (M. Koponen leg.); and 2 O from Poland (DAZH), Bialowieza, 8.7.1988, Pomorze: Stara Brda, Opiiis (Hypocynodus) mongolaltaiensis Fischer, 12.7.1988 (M. Koponen leg.). 1991: 169, O. Distribution. Finland (map 106), Mongolia. Subgenus Hypocynodus Foerster A new species to Finland. Host. Unknown. Hypocynodiis Foerster, 1862: 260. Type species: Material examined. 2 O from Finland (DAZH), N: Nur- Opius crassipes Wesmael. mijaNi, 21.5., 14.6.1983 (M. Koponen leg.).

Opius (Hypocynodus) crassipes Wesmael Subgenus Adontopius Fischer

Opius crassipes Wesmael, 1835: 127, O ; Fischer Adontopius Fischer, 1984a: 93. Type species: 1958c: 306; 1971a: 58; Wharton 1987: 66. Opiiis (Opiothorax) adentatiis Fischer. Hypocynodus crassipes Foerster, 1862: 260. Opius (Hypocynodus) crassipes; Fischer 1972: Opius (Adontopius) adentatus Fischer 428 (key), 430 (redescription); Papp 1979b: 353; 1981c: 154; 1982a: 243. Opiiis (Opiothorax)adentatus Fischer, 1980: 202, Opiiis (Phaedroctoma)crassipes; Tobias 1986: 89. O. Distribcition. Austria, Belgium, Denmark, Opiiis (Adontopius)adentatiis Fischer, 1984a: 93. England, Finland (map 104), Germany, Greece. Opiiis (Phaedrotoma)adentatiis; Tobias 1986: 92. Hungary, Italy, Romania, Russia, Switzerland,Ar- Distribution. Austria, Finland (map 107). menia. A new species to Finland. A new species to Finland and Greece. Host. Unknown. Host. Unknown. Material examined. 1 6 (aliotype) from Finiand (DAZH), Material examined. 18, 3 O from Finland (DAZH), Al: K: Nurmijmi, 5.8.1993 (M. Koponen leg.). Hammarland, 7.7.1984, Jomaia, 29.6.1984; Ab: Nauvo, Sei- li, 20.7.1978, Pernio, 6.8.1982 (M. Koponen), and 28, 1 O Subgenus Opiothorax Fischer from Greecr (DAZH), Corfu: Aghii Deka, 24.5.1982, Pera- ma, 3.6.1977, 24.5.1978 (M. Koponen leg.). Opiothorax Fischer, 1972: 441. Type species: Opiiis levis Wesmael. Opirts (Hypocynodus) latipes Fischer Opius (Opiothorax) abditiformis Fischer Opiirs latipes Fischer, 1958c: 308, O d; 1971a: 80; Hellén 1959: 22. Opiirs (Opiothorax) abditifomis Fischer, 1984a: Opiiis (Hypocynodus) latipes; Fischer 1972: 429 94 (key), 96, 1995: 220 (key). (key), 434 (redescription). Distribiition. Finland (map 1OS), Slovakia. Opiirs (Phnedrotomn) latipes; Tobias 1 986: 92 A new speri?. to Finland. (Ley, dis tri buti on). Host. Unknown. Distribzition. former Czechoslovakia, Den- kfaterial e.\.ni?ziried.4s from Finland (DAZH), N: Nur- mark, England, Estonia, Finland (map los),Ger- mijarvi, 7.7.1983; Sa: Ristiina, 19.7.1987; KI: Parikkala, many, Hungary, Kazakhstan, Moldova, Turkey. 1.7.1990; Ks: Kuusamo, 2.7.1979 (M. Koponen leg.). A new species to Estonia. Host. Diptera, Agromyzidae: Napomyza nig- Opirts (Opiothorax) attribrttus Fischer ritiila (Zetterstedt) = cineracea (Hendel) on Ra- ncinciiliis ncris. Opiiis attribiitiis Fischer, 1962e: 23, d. Material e.rnmirted. 3 O from Finland (DAZH, MZH), Opiiis (Opiothorax)attributiis; Fischer 1972: 441 N: Nurniijiii, 20.5.1993 (M. Koponen). Sibbo, Nevas, 8.- 1

9,140 : Fischer & Koponen: A survey of Opiinae ENTOMOL. FENNICA Vol. 10

O (key), 443 (redescription); Papp 1981c: 154; Opius (Opiothorax)funebris Wesmael Fischer 1984a: 94, 1995: 220 (key). Opius (Phaerdrotoma) attributus;Tobias 1986: 92 Opiusfiinebris Wesmael, 1835: 124, O d ;Fischer (key, distribution). 1960: 34 (redescription); 1971a: 68; 1974: 83. Distribiition. Austria, former Czechoslovakia, Opius (Opiothorax)jiinebris; Fischer 1972: 441 Finland (map 109), Hungary, Lithuania, Ukraine. (Key), 447 (redescription); Papp 1979b: 349; A new species to Finland. Fischer 1984a: 94 (key); 1995: 220 (key). Host. unknown. Opiiis (Phaedrotomn)fiinebris; Tobias 1986: 94 Marerial examined. 1 d 2 O from Finland @AZH, MZH), (key, distribu tion) . N: NurmijWi, 12.6.1986 (M. Koponen), Sibbo, Nevas, 29.6.- Opius rriminnns 'Fischer, 1957d: 9, d. Syno- 4.7.1986 (A. Albrecht); Sa: Ristiina, 28.7.1986 (M. Kopo- nymised by Fischer 1971a: 68. nen leg.). Opius dilatatus Fischer, 1960: 56, 8 ; 197 1a: 6 1. Opius (Opiothorax) dureseaui Fischer Synonymised by Fischer 1972: 447. Distribzition. Austria, Belgium, former Czech- Opiiis (Opiothorax)dureseaui Fischer, 1975: 53; oslovakia, Denmark, England, Estonia, Finland 1984a: 94 (key); 1995: 221 (key). (map 112), France, Germany, Hungary, Lithua- Opiiis (Phaedrotoma)dureseaui; Tobias 1986: 94 nia, Romania, Russia. (key, distribution) . A new species to Estonia and Finland. Distribution. Finland (map 1lo), France, Hun- Hosts. Diptera, Agromyzidae: Napomyza nigritiila (Zetterstedt) = cineracea (Hendel) on Ra- gary. A new species to Finland. ntinczilus acris, Napomyza evanescens (Hen- del) on Rancinculus lnnuginosus. Host. Diptera, Agromyzidae: Agromyza fron- Material exnrnined. 1 1d, 14 O from Finland (DAZH), telfn Rondani. Al: Saltvik, 28.6.1983; N: Inkoo, 24.5.1989, Nurmijarvi, Material examined. 1d frorn Finland (DAZH),N: Nur- 16.6.1986, 2.6.1989; Ta: Hameenlinna, 22.5.1982, Tamme- mijmi, 21.6.1987 (M. Koponen leg.). la, 20.6.1982; Sa: Mikkelin mik., 11.6.1977, 20.6.1981, Ris- tiina, 5.6.1983; Sb: Nilsia, 17.6.1988; Orn: Kaustinen, Opiiis (0piothorax)Jilicornis Thomson 27.6.1978, Pyhajoki, 29.6.1978, Ruukki, 29.6.1978; Ok: Paltamo, 30.6.-6.7.1991, Ristijarvi, 3.7.1991: Le: Enontekio, Hetta (7592:361), 4.7.1982 (al1 M. Koponen); and 1 O from OpiusJilicomis Thomson, 1895: 2 196,E; Fischer Estonia (DAZH), Saaremaa: Kübassaare, 30.5.1990 (M. Ko- 1960: 70, O d; 1971a: 65. ponen leg.). Opius (0piothorm)filicomis;Fischer 1972: 442 (key), 446 (redescription); 1984a: 95 (key); 1995: Opius (Opiothorax) inflammatus Fischer 221 (key). Opiris (Phaedrotomn)Jilicornis;Tobias 1986: 96. Opiiis inflammatus Fischer, 1962a: 38, O ; 197 1a: Distribution. Austria, the Canary Islands (Ten- 76. erife), Denmark, Finland (map 11 1), Germany, Opiiis (Opiothornx) injlatnmatus; Fischer 1972: Itaiy, Russia, Sweden, Kazzkhstan. 441 (key), 450 (redescription); 19842: 94; 1995: A new species to tbe Canary Islands. 220 (key). Host. Diptera, Agromyzidae: Phjtomyza scoti- Opiiis (Phnedrotomn)itiflammntus; Tobias 1986: iicl Hendel on SnIvin prntensis. 94 (key). .1' ,icí~ /'L(! ~i~t!t~ítf.'~!.:L-, C T f,,~,..Fiii:,.t~d (D=',ZII), A!J Uisit ibiiliuti. kiisti-;a, FiriiaIid tii~api 131, h'au\o, Seili, 20.7.1978 (M. Koponen); N: Helsinki, Greece, Hungary. 2j.7.1978, Nurmijarvi, 6.6., 25.5.1987, 17.7.1993, Porvoo, 1.8.1952 (M. Koponen); Ta. Heinolan mlk., 23.6.1985, A new species to Finland and Greece. Hamernlinna, 26.5.19S2, Janakkala, 26.5.1982, Lahti, Host. Unknown. 9.8.1984 (M. Koponen); Sa: MiAkelin rnlk., 19.6.1981 (M. Mnterinl examined. 15 O from Finland (DAZH, MZH), Koponen); 1 O from Russia (MZH), Leningrad re,'oion: Ab: Pojo, Aminne, 14.7.1965 (M. Meinander); N: Nurmi- Raisala, 22.6.1920 (W. Hellén); and 26, 1 P from the Ca- jarvi, 5.8.1979, Tammisaari, 17.6.1980 (M. Koponen), nary Islands (DAZH), Tenerife: Sta Ursula, La Quinta, 8.- Tvarminne, 1.5.-31.7.1964 (MZH); St: Rauma, 1.7.1986 (M. 11.12.1993 (M. Koponen leg.). Koponen); Sa: Ristiina, 23.6.1984.23.7.1994 (M. Koponen); ENiOMQk. FENNICA Vol. 10 0 Fischer & Koponen: A siin’ey of Opiinae 141

o Sb: Pieksarnaen mlk., 14.7.1983 (M. Koponen); Kb: Lipen, tio, 30.8.198 1, Tammisaari, 13.8.1986, Tuucu!u, 23.6.1975 5.7.1994, TohrnajWi, 23.7.1982 (M. Koponen); Ok: Kuh- (M. Koponen). Tvarrninne, 22.8.1948 (U. v. Udvardy), mo, 7.7.1980 (M. Koponen); and 1 O frorn Greece, Kha- 19.8.1951 (P. Palrngren),Vantaa, 5.9., 14.10.1978,24.8.1979 lkidike: Kassandra, Pallini, 5.5.1976 (M. Koponen leg.). (M. Koponen): Ka: Miehikkala, 30.-3 1.8.1977 (M. Koponen); St: Huittinen, 29.-30.5.1993, Raurna, 1.7.1986, Rauman mlk., Opius (Opiothorax) levis Wesmael 2.7.1986 (M. Koponen), Reposaan, 24.8.1950, 4.6.1956 (V. Lauro), Varnmala, 28.6.-1.7.1993 (M. Koponen); Ta: Hattu- la, 11.7.1976, Heinolan mlk., 25.7.1982 (M. Koponen), Ju- Opius levis Wesmael, 1835: 56, ? d ;Hellén 1959: upajoki, 13.-14.6.1982 (Y. Zhongqi), Jarnsa, 26.-29.8.1991 22; Fischer 1960: 86 (figs.); 1971a: 82; Hellén (M. Koponen), Kangasala, 20.6.1938 (A. Saarinen), Korpi- 1959: 22; Wharton 198.7: 66. lahti, 203.1986, Kuhrnoinen, 25.8.1991, Lahti, 15.8.1980, 17.7.1987, Larnrni, 6.-13.7.1976, 27.8.1984, Loppi, Opiiis (Opiothorax) levis; Fischer 1972: 442 12.7.1991, 1.8.1992, 29.8.1993 (M. Koponen), Vanaja, (Key), 453 (redescription); Papp 1982b: 106; 18.6.1921 (Y. Hukkinen); Sa: Lappeenranta, 26.6.1990, 1982c: 184; Fischer 1984a: 95; 1995: 221 (key). Mikkelin mlk., 15.6., 22.8.1975, 24.7., 14.8.1976, 5.8., Opicis (Phaedrotoma)levis; Tobias 1986: 96 (key); 26.8.1978, 22.6.1979, 5.7., 30.7.1980, 19.6., 5.9.1981, 23.6.1983, 12.8.1984, 31.7.1986, 5.-18.7.1987, 23.7.1989, Papp 1988: 58 (distribution). 27.6.1992, 6.8.1993, Mantyharju, 5.9.1981, 4.6.1982, Per- Opiiis varipes Szépligeti, 1898: 389, 401, 0 d. tunrnaa, 22.6.1983, Ristiina, 23.7.1977, 22.6. 1979, 13.7., Synonymised by Fischer 1971 a: 82. 5.9.1981, 4.-5.6.1983, 23.6.1984, 30.8., 5.10.1985, Distribution. the Canary Islands, Europe, Al- 29.7.1986, 4.-19.7.1987, 23.7.1989, 27.6.1992, Suornennie- mi, 23.7.1978, Valkeala, 28.7.1977 (M. Koponen); K1: Pa- geria, Central Asia, Isreal, North Korea. This is nkkala, 27.6.-2.7.1990 (M. Koponen), Uukuniemi, 4.7.1983 the most common species of Opiinae in Finland (M. Pulkkinen); Tb: Rautalampi, 18.-19.7.1983, Vesanto, (map 114). 28.6.1991 (M. Koponen),Viitasaari, 30.6.-3.7.1947 (W. Hack- A new species to the Canary Islands. man); Kb: Kesalahti, 17.7.1982, Kitee, 22.7.1982, Liperi, 24.6.1982, 3.-5.7.1994, Tohmajarvi, 19.-23.7. 1982 (M. Ko- Hosts. Diptera, Agromyzidae: Liriomyza con- ponen); Om:Virnpeli, 8.-10.7.1955 (M. Koponen); Ok: Ris- gesta (Becker), Liriomyza j7aveola (Fallén) on tijWi, 10.7.1989, Sotkamo, 15.8.1982 (M. Koponen); Oba: Gramineae, Liriomyza strigata (Meigen) on Cam- Pudasjarvi, 20.7.1980, Vaala, 20.7.1980 (M. Koponen); Obb: panda sp., Phytomyza glechomae Kaltenbach. Rovaniemen mlk., 9.8.1983, Simo, 17.7.1980, Tornio, 6.7.1982, Ylitornio, 12.8.1982 (M. Koponen); Lkoc: Kittila, Material examined. 263 d, 129 O frorn Finland (DAZH, Sirlcka, 7.7.1950 (J. Jalava & R. Vaisanen); Lkor: Sodankyla, MZH), Al: Foglo, 9.7.1984, Lernland, 6.-8.7.1984 (M. Ko- 10.-11.8.1983 (M. Koponen); Li: Inari, 6.7.1989 (M. Kopo- ponen), Lemland, Flaka, 12.-3.-12.6.1954 (Waselius), nen); 1 0 frorn Sweden (DAZH); 11 d, 2 ? frorn Estonia 24.8.1954 (Hakan Lindberg), Mariehamn, 11.6.1976, Vikdo, (DAZH); 7 d , 1 0 frorn Poland (DAZH), 2 ,90 frorn Russia 8.7.1984 (M. Koponen); Ab: Kajalohja, 19.6.1992 (M. Ko- d (MZH) (Leningad region, Karelia, Moscow, Taganrog), 1 O ponen), Lojo = Lohja (R. Frey), Nauvo, Sando, 19.7.1978, from Greece, Corfu: Acharavi, 28.4.1994 (M. Koponen); and Nauvo, Seili, 18.-20.7.1978, Paimio, 7.8.1982, Pernio, 50d,34? frorn the Canary Islands (DAZH), Tenenfe: Porte- 6.8.1982, 19.-20.9.1994 (M. Koponen), Pojo, Arninne, la Baja, 4.4.1988, Puerto de la 11.-15. 12.1986, 28.3.- 14.7.1965 (M. Meinander), Finby = Sarkisalo (W. Hellén), Cruz, 9.4.1988, Sta Ursula, 11.12.1993 (M. Koponen leg.). Turku, 2.10.1976, 18.-19.9.1951 (M. Koponen), Nystad = Uusikaupunki, 1919, 20.7.1924 (M. Hellén), Vichtis = Vihti (R. Frey), 18.7.1964,6.-8.6.1967 (V. J. Kanonen), 11.7.1982 Opiiis (Opiothorax) longicornis Thomson (M. Koponen), Vastanfjard, 15.8.1993 (M. Koponen); N: Espoo, 27.8.1982, 25.-26.7.1953 (hl. Koponen), Hango = Opiiis longicoriiis Thomson, 1895: 2 197, 0 ; Hanko, 23.7.1 969 (W. Hellén), 20.5.1989 (M. Koponen), H:fors = Helsinki (R. Frey), 2.9 1965 (E. Karvonen), 26.8.- Fischer 1960: 63; 1971a: 82. 4.9.1975, 26.7.-11.9.1976, 20.7., 23.8.-8.9.1977, 7.- Opiiis (Pendopiiis)longicornis; Fischer 1972: 41 O 24.9.1978, 28.7., 2.-6.9.1980, 22.6., 22.7.. 15.-22.9.1981, (key), 41 3 (redescription). 31.S., 2 -4 8., 23 9.1982, 16.6.1983, 19.9.1935 (M. KOPO- 0,r~iri~(Opiotlmrqi-) 1onyicnlai.v: Ficcher 19843: nen). 27.6.. 30.7.. 29.8.. 12.9.1983 (Y. Zhorigqi), Hyvinküa, 9.8.1955, Kirkkonumnii, 30.8.19S1, 23.-21.7.1991, Nurnii- 95; 1995: 220 (key). jarvi, 21.6., 20.7., 13.8. 1975, 16.7.1978. 14.7.1979, 23.- Opiiis (Phaedrotonin)longicorriis; Tobias 1986: 24.8.1980, 11.7., 30.7.1981, 14.-19.6., 6.8., 27.8.1983, 2.- 85. 7.6., 8.8.1984, 14.8., 24.8., 14.-19.9.1985. 23.6., 28.8., Opiiis ccuidifer Fischer, 195Sc: 295, 9 8 ; Hellén 6.9.1986, 1.8.1987,20.6., 20.8., 6.9.1989, 16.6., 9.-17.7.1991, 20.6., 18.-24.7.1992, 19.6.1993, 6.6., 9.7., 5.-13.8.1994 (M. 1959: 21; Fischer 1967b: 140; 1971a: 52. Syno- Koponen), PernA = Pernaja, 12.8.1915 (A. Nordstrom), Por- nymised by Fischer 1981 b: 23. vom rnlk., 30.7.1978, 1.8.1952 (M. Koponen), Sibbo = Sipoo, Opiiis (Opiothorax) calidver; Fischer 1972: 441 27.8.1975, 3.6., 3.8.1952, 1.-15.6.1983. 26.9.1985 (hl. KO- (key), 444 (redescription); Papp 1982~:189. ponen), 19.-31.8. 1985, 15.6.-4.7.1986 (A. Albrecht), Siun- - ,152 >* Fischer & Koponen: A survey of Opiinae ENTOMOL. FENNICA Vol. 10

Ii Opius (Phaedrotoma) caudifer; Tobias 1986: 94 MZH), Ab: Nauvo, Seiii, 17.7.1978 (M. Koponen); N: Es- (key, distribu tion). poo, 26.7.1983, Helsinki, 10.8.1976, 20.7.1977, Nurmijarvi, 27.5.1984,20.6.1987,16.6.1991,Porvoonmlk.,3.6.1982(M. Distribution. Austria, the Canary Islands, Koponen); St: Vammala, 28.6.1993 (M. Koponen); Ta: Kan- former Czechoslovakia, Finland (map 1 15), gasala (R. Frey); Sa: Ristiina, 10.7.1993 (M. Koponen); Tb: France, Greece, Hungary, Romania, Russia, Slov- "Tav. bor." (Woldstedt); Om: Vimpeli, 7.7.1985 (M. Kopo- enia, Sweden, forrner Yugoslavia. nen); Ok: Paltamo, 5.7.1991, Ristijbi, 4.7.1991 (M. Kopo- nen); Obb: Tomea = Tornio, 4.9.1924 (Hakan Lindberg); A new species to the Canary Islands and Lkoc: Kittila, Sirkka, Katkatunturi, 7.-12.7.1980 (J. Jalava Greece. & R. Vaisanen); 1 O from Sweden (DAZH), Nb: Overtomea, Host. Diptera, Agromyzidae: Phytomyza isa- 27.6.1979 (M. Koponen); 7 O from Estonia (DAZH), Saare- is Hering on seeds of Odontites verna. maa: Kaali, 30.5.1990, Lihulinna, 2.6.1990, LOpollu and M&tu park, 31.5.1990, Pangapank, 2.6.1990 (M. Koponen); Material examined. 8 8, 82 O from Finland (MZH), Al: and 4 O from Poland (DAZH), Podlasie: Bialowieza, 7.7.1988, Finstrom, 5.7.1984, Foglo, 30.6.1984, Geta, 5.7.1984, Lem- land, 6.7.1984 (M. Koponen); Ab: Hitis (Reuter), Nauvo, Mielnik, 5.7.1988 (M. Koponen leg.). Seili, 20.7.1978, Pernio. 6.8.1982, Rymattyla, 27.6.1983 (M. Koponen); N: Helsinge, 1915 (W. Hellén), Espoo, 25.7.1983, Opius (Opiothorax) minuscuiue Fischer Helsinki, 17.7.1976, 10.-19.8.1976, 20.7., 12.8.1977, 2.8., 13.9.1982, Hyvinkaa, 19.7.1981, Kirkkonummi, 23.- 24.7.1991, Nurmijarvi, 4.9.1976, 16.7., 16.8.1978, 29.- Opicis minusculae Fischer, 1967a: 328, O ; 1971a: 3 1.7.1984, 1.8.1987,24.7.1989, 18.-24.7.1992.9.-16.7.1993. 88. Sipoo, 27.8.1975, 13.8.1977,26.7.1981,3.8.1982,24.7.1983 Opius (Opiothorar) minusculae; Fischer 1972: (Me Koponen), TvMnne, 13.-30.7-1914(w. Helh); Ka: 442 (key), 458 (redescription); 1984a: 95 (key); Ylamaa, 4.7.1990 (M. Koponen); St: Kokemiiki, 10.8.1992, Rauma, 1.7.1986, Rauman mlk., 2.7.1986 (M. Koponen). 1995: 221 (key); Peydro 1983: 240. Reposaari (V. Lauro); Ta: Hausjmi, 23.8.198 1, Jiakkala, Opicis (Phaedrotoma) minusculae; Tobias 1986: 9.7.1983, Loppi, 12.7.1991 (M. Koponen); Sa: Mikkelinmik., 96. 24.6.1983. 25.7.1985, Pertunmaa, 4.8.1983, Ristiina, Distribution. England, Finland (map 117), 23.6.1984, 19.7.1987 (M. Koponen); Tb: Rautalampi, 19.7.1983 (M. Koponen); Sb: Kuopio, 18.8.1983, Nilsia, Hungary* 17.6.1988, Suonenjoki, 15.8.1983 (M. Koponen); Kb: Kitee, A new species to Finland. 19.-22.7.1982, Liperi, 4.8.1993, 3.7.1994, Tohmajarvi, Host. Diptera, Agromyzidae: Phytomyza mi- 21.7.1982 (M. Koponen); Ok: Kuhmo, 10.7.1980 (M. Kopo- nclscula G~~~~~~. nen); Obb: Simo, 18.7.1980, Tervola, 13.8.1982 (M. Kopo- Material examined. 18 from Finland (DAZH), N: Es- nen); Lkor: Sodankyla (7486:507), 10.8.1983 (M. Koponen leg.); 1 O from Greece (DAZH), Corfu: Acharavi, 23.4.1994 POo3 25-7*1983 ('. Koponen leg.)* (M. Koponen); and 1 O from the Canary Islands (DAZH), Tenerife: Sta ürsuia, La Quinta, 11.12.1993 (M. Koponen). Opius (Opiothorax) mirabilis Fischer

Opius (Opiothorax) magnicauda Fischer Opius mirabilis Fischer, 1958f 173, ? d ; 1971 a: 88. Opius magnicaiida Fischer, 1958: 298, 3 d; Opizis (Opiothorax) mirabilis; Fischer 1972: 441 1971a: 84; Hellén 1959: 22. (key), 459 (redescription); 1984a: 95; 1995: 221 Opir is (Opiotho rax) magri ica ir da ; Fischer 1972 : (key). 441 (Key), 456 (redescription); Papp 1979: 82; Opicis (Phaedrotoma) mircrhilis;Tobias 1986: 94. Fischer 1984a: 94; 1995: 220 (key). Distribiition. Austria, Finland (map 11S), Opiirs (Phaedrotorna)mngnicarrda; Tobias 1986: France, Germany, Italy, Poland, Russia, former 9 1 (?:q,?,ist:.ih:iticz?j. TIA U~UJ~V~LL, AI iliciíia, Ldli~~~ii, TLiiiisia. Distribrrtion. Austria, former Czechoslovakia, A new species to Finland and Poland. Denmark, Estonia, Finland (map 116), Hungary, Hosts. Diptera, Agromyzidae: Lirioiriyza bal- Italy, Poland, Russia, Sweden, Switzerland, Ka- canica S trobl, Liriomyza sp. on Eriphorbia zakhstan. nicaeensis All. A new species to Estonia, Poland and Swe- Material e.ramined. 1 a, 2 O from Finland (DAZH), N: den. Nurmijarvi, 14.7.1979, 11.6.1988; Sa: Mikkelin mlk., Host. Unknown. 25.7.1976 (M. Koponen); and 1 O from Poland (DAZH), Pod- lasie: Bialowieza, 9.7.1988 (M. Koponen leg.). Material e-raniined. 3 d, 18 O from Finland (DAZH, ENTOMOL. FENNICA Vol. 10 0 Fischer & Koponen: A siirvey of Opiinae 143

e oi Opius (Opiothorax) nigricoloratus Fischer nen); Kb: Liperi, 4.8.1983 (M. Koponen); Lkoc: Kittila, Sirk- ka, Katkatunturi, 7.7.1980 (J. Jalava & R. Vaisanen leg.). Opicis nigricoloratus Fischer, 1958f 175, d ; 1971a: 90; Hellén 1959: 22. Opius (Opiothorax) spretus Haliday Opius (Opiothorax)nigricoloratiis; Fischer 1972: 441 (Key), 462 (redescription); 1984a: 95; 1995: Opiiis spretcrs Haliday, 1837: 207, O d ; Fischer 221 (key); Papp 1979b: 349. 1960: 91 (figs.); 1967a: 347; 1971a: 113. Opiiis (Phaedrotoma) nigricoloratus; Tobias Opius (Opiothorax) spretiis; Fischer 1972: 441 1986: 94. (key), 469 (redescription); Papp 1981c: 141; Distribution. Denmark, Finland (map 119), Fischer 1984a: 95; 1995: 221 (key), 226, figs 7- Germany, Hungary, Italy ; Mongolia. 9. Host. Unknown. Opius (Phaedrotoma) spretus; Tobias 1986: 94 Material examined. 1 4 frorn Finland (MZH), N: Hoplax (key, distribution). (R. Frey). Opiiis clarus Haliday, 1837: 206, 9.Synonymised by Fischer 197 1a: 469. Opius (Opiothorax) opacus Fischer Opiiis arenosus Szépligeti, 1898: 389,401, 9 8; Hellén 1959: 21; Fischer 1960: 82. Synonymised Opiiis opacus Fischer, 1968: 107, 9 ; 1971a: 93. by Fischer 1971a: 469. Opicis (Opiothorax) opacus; Fischer 1972: 442 Opiirs spretiis; van Achterberg 1997: 77 (singu- (Key), 463 (redescription); 1984a: 95; 1995: 221 laris Wesmael as new synonym, but stated as val- (key); Papp 1982c: 184. id name on page 78. The question is left open Opicis (Phaedrotoma) opacus; Tobias 1986: 96. here). Distribufion. Finland (map 120), Greece, Hun- Distribzition.Austria, former Czechoslovakia,- gary, Moldova, Mongolia. Denmark, England, Finland (map 122), Hunga- A new species to Finland. ry, Ireland, Italy, Poland, Russia, Sweden, Swit- Host. Unknown. zerland, former Yugoslavia, North Korea. Material examined. 1 4 frorn Finland (DAZH), N: Nur- A new species to Sweden. mijarvi, 5.6.1986 (M. Koponen leg.). Hosts. Diptera, Agromyzidae: Agromyza abi- ens Zetterstedt, Agromyza ferrciginosa Wulp on Opius (Opiothorax) soenderrrpianus Fischer Sy mph y t ii m oficinale, Ag romy za my osotidis Kaltenbach and Agromyza pseudorujipes Nowa- Opiiis soenderiipianiis Fischer, 1967a: 33 1, Q ; kowski on Myosotis laxa, Agromyza reptans Fal- 1971a: 112. lén on Urtica dioica, Agromyza rclfipes Meigen Opiiis (Opiothorax) soerideriipianus; Fischer on Myosotis paliistris and Symphytum oficinale. 1972: 441 (Key), 467 (redescription); 1995: 220 Material e.raniined. 5 d, 11 4 frorn Finland (DAZH, (key). hlZH). N: Helsinki (R.Frey), 31.8.-1.9.1978, 16.8.-2.8.1979 (M. Koponen), Nurrnijarvi, 22.6.1989, Porvoon rnlk. Opiiis (Phaedrotorna) soenderiipiariiis; Tobias 26.8.1979, 10.8.1950; Ta: Hattula, 11.7.1976; Sa: Mikkeli, 1986: 94 (key, distribution). 5.9.1987, Mikkelin mlk., 10.8.1987, Ruokolahti, 2.7.1975; Distribiition. Finland (map 12 1), Germany, Kl: Rautjarvi, 29.6.1990; Kb: Tohrnajarvi, 2 1.7.1982; Obb: Russia. Kemi, 6.7.1952 (M. Koponen); 1d from Sweden (DAZH). Vb: Brannland, 14.7.1950 (M. Koponen); and 1 from Po- A new speries to Finlarirl. 0 idid (D;iZli,. Host. Diptera, Apromyzidae: Cerotlonrhn (Dizygomyza) caricicola (Hering) = soeizderiipi Opircs (Opiothorax) turcicus Fischer (Hering). Marerial e.rnmiried. 20 4 frorn Finland (DAZH, hlZH). Ab: Pernio, 6.8.1982 (M. Koponen); N: Espoo, 26.7.1983, Opiiis tirrciciisFischer, 1960: 93, O 0;1971a: 119. Helsinki, 3 1.7.1982, Hyvinkaa, 9.8.1985, Nurrnijarvi, Opiiis (Opiothorax) tiirciccis; Fischer 1972: 441 5.9.1984, Sipoo, 30.8.1983 (M. Koponen); Ka: Ylamaa, (key), 471 (redescription); 1984a: 95; 1995: 221 13.8.1987 (M. Koponen); Ta: Lamnii, 27.8.1964 (hl. Kopo- (key); Papp 1988: 59 (distribution). nen); Sa: Mikiielin rnlk., 3.8.1953, 25.7.1985, Ristiina, 25.7.1985, 19.7.1987 (hl. Koponen); Tb: Rautalampi, Opiiis (Phaedrotoma) tiircicirs; Tobias 1986: 96 14.7.1983 (M. Koponen); Sb: Tervo, 18.7.1983 (M. Kopo- (key, distribution). ,144 ***‘ Fischer & Koponen: A sziney of Opiinne ENTOMOL. FENNICA Vol. 10

5 Distribiition. Finland (map 123), France, Ger- Psyttalia rhagoleticola (Sachtleben) many, Hungary, Netherlands, Crimea, Turkey, Algeria, India, Iraq, Israel, Jordan, Mongolia. Opiiis rhagoleticoliis Sachtleben, 1934: 76, 3 d ; A new species to Finland. Fischer 1958b: 210. Host. Diptera, Agromyzidae: Agromyza lath- Opiiis rhncoleticola; Fischer 1965a: 18; 197 1a: yri Hendel on Pisirm sativum. 105. Material examined. 3 d from Finland (DAZH, MZH), Opizrs (Psyttalin) rhagoleticola;Fischer 1972: 334 Al: Lumparland, 30.6.1984 (M. Koponen); K1: Parikkala, (Key), 344 (redescription); Papp 1979: 88. 22.6.1940 (W. Hellén); Paltamo, 6.7.1991 (M. Koponen Ok: Opiiis (Psyttalin) rhngoleticolus; Tobias 1986: 73 leg.). . (key, distribution). Genus Psyttalia Walker Opiiis carinatus auct.; Hellén 1946: 132 (misidentified). Psyttalia Walker, 1860: 3 11. Type species: Psyt- Distribiition. Austria, former Czechoslovakia, talia testacea Walker. Finland (map 125), France, Germany, Russia, Synonym of Opiiis; Fischer 1971a: 38. Switzerland, former Yugoslavia, Armenia, Kaza- Opicis subgenus Psyttalia; Fischer 1972: 334. khstan, Kirgizhia, Siberia. Psyttalia; Wharton 1987: 63,64. A new species to Finland. Hosts. Diptera, Tephritidae: Rhagoletis cera- Psyttalia carinata (Thomson, 1895) si (Linnaeus) on Cerasiirm nvium and Lonicera SP- Marerial e.rar>iiiied.3 d ,6O from Finland (DAZH, MZH), This species was incorrecty reported from Fin- N: Helsinki, 7.8.1990 (on Lonicera sp.), Porvoon mlk., land (Hellén 1946). See Psyttalia rhngoleticola 10.9.1980, Vantaa, 21.8.1991 (M. Koponen); Ta: Hauho, (Sac h tleben). 16.8.1935 (hl. Hellén), Saaksmaki, 10.7.1937, 6.-8.8.1938 (E. Kivirikko). Psyttalia concolor (Szépligeti) Genus Ezirytenes Foerster Opiiis concolor Szépligeti, 1910: 244. Opiiisfuscitarsis Szépligeti, 1913: 605, d.Syno- Eiiiytenes Foerster, 1862: 259. Type species: Opi- nymised by Fischer 1963b: 679. LISnbnonnis Wesmael; Fischer 1972: 60,472 (use Opiiisperproximcis Silvestri, 1914: 103, 9 d.Syn- of Eiiytenes Foerster sensu auct. Olim, in new onymised by Fischer 1963b: 679. sense only Euytenes s. str.); Fischer 1987: 549 Opiiis (Psyttalia) concolor; Fischer 1972: 334 (two Old World species). (key), 337 (redescription); Tobias 1986: 73 (key, Eiirytenes, Fischer 1998: 21,22. New interpreta- distribution); Papp 1988: 58 (distribution). tion. Distribiition. Finland (map 124), Greece, Ita- ly, Spain, former Yugoslavia, Israel, Africa. Eir ryteri es abn orrn is (Wesmael) A new species to Finland. Hosts. Diptera, Tephritidae: Cnrpomyin iti- Opiiis abrionnis Wesmael, 1935: 117, 9 O. completa (Becker) on Zizyphzrs satirw, Cnppa- Eiinteries abnorinis; Foerster, 1ü62: 259; Fischer rirnjin scivastaiiii (Martelli), Cerntitis cnpitnta 1959b: 248 (figs.); 1971a: 123; 1972: 473 (rede- (Wiedsriiann); Dcrcirs oleae (Ginelirij on Oleci scirpticjiij; ‘lubias 1980: 96. europea. In Finland reared from Eiiplircrnta cotl- Distribzition. Austria, Belgium, Croatia, Fin- iiexa (Fabricius) on Cynnchum rincetoxicurn. This land (map 126), Germany, Poland, Russia, Cau- is a new host. casus, North America. Marerial e,ranziiied. 1 O from Finland (MZH), Al: Lem- A new species to Finland. land, Marso, 1918 (A. Nordman lep.). Hosts. Diptera, Agromyzn albitarsis Meigen; E"f0MOL. FENNICA Vol. 10 0 Fischer & Koponen: A siirvey of Opiinne 145

L Agromyza sp. on Holciis lanatiis, Amaiirornyza Impatiens noli-tangere, Phytomyza campanariae lnmii Kaltenbach on Lcrminstrum galeobdolon, Nowakowski on Pulsatilla pratensis. Cerodontha (Butomyza) angiilata (Loew) and Material eramined. 1 d and 1 ? from Finland (DAZH), Cerodontha (Biitoniyzn) eiicaricis Nowakowski N: Nurmijarvi, 1.9.1979 (M. Koponen); Le: Enontekio, KilpisjarVi, 7.7.1966 (Karvonen leg.); 1 d and 1 ? from Rus- on Carex hirta, Cerodonthn (Biitomyza) cnriciv- sia (MZH), Murmansk region: Gavrilova, 21.-27.7.1913, ora (Groschke) on Carex hirtn and Carex sp., Pummanki, 1930 (W. Hellén leg.), and 18 from the Canary Cerodontha (Diqgornyza)iraeos (Robineau-Des- Islands (DAZH), Tenerife: Sta Ursula, La Quinta, 8.12.1993 voidy) on Iris pseudacoriis , Cerodontha (Phyta- (M. Koponen leg.). gromyza)flavocingiilatn (Strobl) on Festuca pratensis and Holciis lanatus, Cerodontha (Poemyzn) Eiiryten es cu bitalis (Fischer) incisn (Meigen) on Alopeciirus pratensis and Ph- leiim pratense, Cerodontha (Poemyza) pygmaea Opius cubitalis Fischer, 19593: 7 1, O d ; 1971 a: (Meigen) on Dactylis glomerata and Deschamp- 58. sin caesiptosa, Liriomyza balcanica (Strobl), Lir- Opiiis (Nosopoen) cubitalis; Fischer 1972: 29 1 iomyza demeijerei Hering on Artemisia vulgaris, (key), 308 (redescription). Liriomyza eripntoriana Spencer on Eupntoriiim Opius (Allotypiis) cubitnlis; Tobias 1986: 65. cannabiniim, Liriornyza flnveola (Fallén) on Fes- Opius (Stigmntopoea) cubitalis; Fischer 1986: tuca pratensis, Liriomyza scorzonerne Rydén on 611. Scorzonera hiimilis, Phytoliriomyza variegnta Eiirytenes cubitalis; Fischer 1998: 25 (nov. (Meigen), Phytomyza abdorninalis Zetterstedt on comb.). Hepatica nobilis, Phytomyza albimargo Hering, Distribution. Austria, Bulgaria, Finland (map Phytomyza fnllaciosn Brischke on Rnniinculiis 128), Poland, Japan, North Korea. repens, Phytomyza obsciira Hendel on Clinopo- A new species to Finland. dium vulgare, Phytornyza piilmonariae Nowa- Host. Unknown. kowski on Piilmonaria angiistifolia, Phytomyzn Material examined. 1 d from Finland (MZH), St: Re- posaan, 26.8.1966 (V. Lauro leg.); and 1 d, 3 ? from Poland senecionis Kaltenbach on Senecio nemorensis, (DAZH), Bialowieza, 7.43.7.1988 (M. Koponen leg.). Amnuromyza (Trilobomyzn) lnbiatnriim (Hendel) on Galeopsis tetrahit, Antniiromyzn (Trilobomyzn) Eurytenes impatientis (Fischer) verbasci (Bouché). Material examined. 1 ? from Finland (DAZH), Ta: Lam- mi, 6.7.1976 (M. Koponen leg.). Opiiis impatientis Fischer, 1957c: 338, O ; 1971a: 74. Eiirytenes campanariae (Fischer) Opius (Cryptonastes) impatientis; Fischer 1972: 179 (key), 193 (redescription, O 0);Tobias 1986: Opiiis cnmpannrine Fischer, 1959i: 82, d. 45 (key, distribution). Opicis (Cr-yptorznstes)cnmpnariae; Fischer 1972: Opiiis (Jiicundopiiis) impntientis; Fischer 1984b: 179 (key), 183 (redescription); Tobias 1986: 43 34 (key). (key, distribution). Euvtenes impatientis: Fischer 1998: 25, nov. Opiiis (Jiiciindop iiis) campnria r ine ; Fi s c he r comb. 1984b: 34. Distribiition. Austria, the Canary Islands, FIII?~PV~Sff~7p7tlfli-iclP' Fiqcher. 1998: 2.5 (nov former Czechoslovakia, Finland (map 129), Ita- c0mb.j. ly, Poland, Russia, Amienia, Kazakhstan, Kirgi- Distrihrition. Austria, the Canary Islands (Ten- sia. erife), former Czechoslovakia, Finland (map 127), A new species to Finland and the Canary Is- Hungary, Poland, Russia, Armenia, Kazakhstan, iands. Mongolia. Hosts. Diptera, Agromyzidae: Agi-ornyza jo- A new species to Finland and the Canary Is- hannae de Meijere on Cytisus scopnriiis, Phyto- lands. liriornyza melampyrn (Loew) = impatientis (Br- Hosts. Diptera, kgromyzidae: Pliytolirioniyza ischke), Paraphytoniyzn luteosciitellntn (de Mei- melnmpyra (Loew) = inipatientis (Brischke) on jere) = xylostei (auct.). , , ..,,

146 1 Fischer & Koponen: A survey of Opiinae ENTOMOL. FENNICA Vol. 10

o Material examined. 1 9 from Finland (DAZH), N: Kirk- Opilis (Nosopoea)rnacrocerus; Fischer 1972: 291 konummi, 23.7.1991 (M. Koponen), 2 9 from Poland (key), 316 (redescription). (DAZH), Bialowieza, 7.7.1988 (M. Koponen); and 1 9 from the CanqIslands (DAZH), Tenenfe: Sta Ursula, La Quinta, OPiUs (StigmatoPoefl) ntflcroCerLls; Fischer 1986: 8.12.1993 (M. Koponen leg.). 611. Ecirytenes rnacrocerus;Wharton 1988: 357 (new Eurytenes leptostigma (Wesmael) combination); Fischer 1998. Opius hians Stelfox, 1949: 306, O 8. Syno- Opiiis leptostigma Wesmael, 1835: 138, 9; Hel- nymised by Fischer 1959g: 59. lén 1959: 22; Fischer 1971a: 81. Distribution. Austria, former Czechoslovakia, Opius leptostigmus, Marshall 1891: 22, ? d. Finland (map 131), France, Germany, Hungary, Opius (Allotypus) leptostigma, Thomson 1895: Ireland, Poland, Romania, Russia, Japan, U.S.A. 2 187. (Mich.). Opius (Opiostomcis) leptostigma; Fischer 1972: Hosts. Diptera, Agromyzidae: Agromyza pre- 75 (redescription); 1986: 613 (key); Zaykov & spana Spencer on Elymcis (=Agropyron) repens; Fischer 1983: 44; Tobias 1986: 23 (key, distribu- Tephritidae: Trypeta immnculatn Macquart. tion) . Material examined. loa, 159 from Finland (DAZH, MZH),Ab: Vichtis =Vihti (R. Frey), 30.6., 4.-9.8.1962,17.7., Eiirytenes leptostigma Wesmael; Fischer 1998,23, 26.8.1964,20.7.1965 (V. J. Karvonen); N: Esbo = Espoo (R. nov. comb. Frey), 7.9., 24.9.1978 (M. Koponen), Helsinki, 4.9.1978 (M. Distribution. Austria, former Czechoslovakia, Koponen), Kirkkonummi. 10.8.1963 (V. J. Karvonen), Nur- Denmark, Finland (map 130), France, Germany, mijarvi, 21.8.1987 (M. Koponen), PernA = Pemaja, 16.- 18.8.1944 (A. Nordstrom), Sibbo, Gesterby, 22.-28.8.1983 Hungary, Kazakhstan (Akmolinsk), Russia (Len- (A. Albrecht); Ta: Korpilahti, 20.8.1986 (M. Koponen); Sa: ingrad region), Sweden. Ristiina, 23.7.1977, 6.8.1978 (M. Koponen); Oa: Lappfjard, Host. Doubtful record of a fly, family Phon- 30.7.1959 (G. Nordstrom); Sb: Joroinen, 9.8.1982 (Y. Zhong- dae: “Phora tuberorum Goureau” (= ”Phora tu- qi leg.). berum”). Material examined. 158 33 O from Finland (DAZH, Eurytenes vockerothi (Fischer) MZH), Ab: Pargas = Parainen (Ingelius), Pojo, Aminne, 14.7.1965 (M. Meinander), AbO = Turku (IngehS), Nystad opillsvockerothi ~i~~h~~,1964~:348, ? d. = Uusikaupunki, 12.-22.6.1916, 6.9.1920 (M. Hellén); N: Hanko, 1931 (W. Hellén), 17.6.1980 (M. Koponen), Helsin- ‘PiUs (CryPtonastes) vockerothi; Fischer 972: ki, 28.6.1966 (1. Jalas), 19.8.1976, 15.8.1978, 15.6.1980 (M. 179 (key), 204 (redescription, syn.); Tobias 1986: Koponen), Nurmijikvi, 6.9.1986, 25.8.1987, Porvoon dk., 43. 26.8.1979, Sipoo, 31.8.1982 (M. Koponen), Ekenas = Tam- Opiiis (Jucundopius) vockerothi; Fischer 1984b: misaari (W. Hellén), Lappvik =Tammisaari, Lappohja (A. Berg), Vantaa, 24.8.1979 (M. Koponen); St: Vammala, 34. 28.6.1993 (M. Koponen); Ta: Janakkala, 24.8.1980, Jamsa, Eurytenes vockerothi:Fischer 1998: 25, new com- 7.7.1982 (M. Koponen), Kangasala, 22.6.1938 (A. Saarin- bination. en), Lahti, 9.6.1978, La&, 12.-14.7.1976 (M. Koponen); Opius bnrsceanus Fischer, 1967a: 325,Q.Syno - Sa: Mikkelin mlk., 15.6.1975, 14.-15.8.1976, 29.7.1980, 5.9.1981 (M. Koponen); Kl: Paikkala, 14.6.1930 (W.Hel- nymised by Fischer 1972: 204. En); Tb: Keuruu, 3.8.1928 (W. Hellin), Rautalampi, Distribution. Austria, Finland (mal 132), Ger- 17.7.1983 (M. Koponen); Kb: Kesalahti, 17.7.1982, Tohma- many, North Korea, Canada, U.S.A.. jkvi, 19.7.1982 (M. Koponen): Om: Pietarsaan 30.6.1978, A new species to Finland. Pyhanta, 16.8.1982 (M. Koponen); Ok: Kuhnio, 7.7.1980 (M. Kopoiien); Oba: Hailuoto, 27.6.190s (Wuorentaus), and 1 V H~SI.DiyIcia, Agl o~iy~iJíit’.Lit ioii,-\):u I)i 1i.b- from Russia, Leningrad region: Terijoki, 8.1927 (%’. Hellén). cae Hering on Eqiiisetiim arveiise. Marfrinfe.irarniried. 2 d 8 O from Finland (DAZH, MZH), Eirrytenes rnacrocerus (Thomson) N: Helsingfors = Helsinki, 10.7.1971 (\Y. Hellén), 26.7.1976, 8.6.1979 (M. Koponen), Nurmijarvi, 30.6.1957, 17.7.1993, Tamrnisaari, 17.6.1980 (M. Koponen); Oba: Kuivaniemi OpillS (Allotypcis) macrocenis Thomson, 1895: (7273:417), 18.7.1980 (M. Koponen); KS: Kuusamo, 21 86, ? 8 ; Tobias 1986: 63. 19.8.1965 (M. Meinander); Lkor: Sodankyla, 11.8.1983 (M. Opiris macrocenis; Fischer 19j98: 58; 1971a: 84; Koponen); Li: Ind. 6.7.1989 (M. KoPonen leg.). Hellén 1959: 22. I 'ENTOMOL. FENNICA Vol. 10 Fischer & Koponen: A siirvey ofopiinae 147

o Genus Biosteres Foerster sen), 10.6.1987 (M. Koponen), Hattula (L. von Essen), Hameenlinna. 7.6. 1980, Janakkala, 30.5.1982, Lammi, 7.6.1980 (M. Koponen), Saaksmaki 17.6.1921 (W. Hellén); Biosteres Foerster, 1862: 259. Type species: Bra- Sa: Mikkelin rnik.; 10.6.1978,5.6.1983 (M. Koponen), Ran- con carbonarius Nees. tasaimi, 1.7.1941 (1. Hellén), 15.7.1941 (S. Hellén), Ristii- Remark. van Achterberg (1 997) follows a dif- na, 1.6.1980,5.6.1982,4.-5.6. 1983.23.6.1985,3.6.1989 (M. ferent definition of the taxa Biosteres, Chilotrichia Koponen); KI: Pankkala, 7.-9.6.1940 (W. Hellén); Kb: Pyha- selka, Hammaslahti, 8.6.1927,3.6.1928 (P. Kont-kanen); Om: and Zetetes. We follow here the old conventional Pietarsaan, 2.7.1981 (M. Koponen); and 19 from Russia interpretation as used by Fischer. The same re- (MZH), Mumansk region. mark for the following Biosteres with author Hal- iday. Biosteres (Chilotrichia) borealis (Zet terst ed t)

Subgenus Chilotrichiu Foerster Bracon borealis Zetterstedt, 1838: 400. Opiiis borealis: Papp 1994: 304. Chilotrichia Foerster, 1862: 258. Type species: Opiiis brevipalpis Thomson, 1895: 2 191, d ;Hel- Opiiis blandus Haliday. lén 1959: 21. Synonymised by Papp 1994. Opiiis (Biosteres) brevipalpis; Fischer 1959a: 11 Biosteres (Chilotrichiu) bicolor (Wesmael, 1835) (redescription). Biosteres brevipalpis; Fischer 197 1a: 2 1. Specimens reported from Finland (Hellén 1946) Biosteres (Chilotrichia)brevipalpis; Fischer 1972: are misidentified, see B. magnicomis. This spe- 488 (key), 496 (redescription); Papp 1979: 91. cies is deleted from lists for Finland. Opiiis (Chilotrichia) brevipalpis; Tobias 1986: 14. Biosteres castaneiventris auct., Hellén 1946: 133 Biosteres (Chilotrichia) blandus (Haliday) (misidentified). Distribiition. Austria, former Czechoslovakia, Opius blandiis Haliday, 1836: 220, ? d ; Denmark, Finland (map 134), Germany, Lithua- Chilotrichia blandiis; Foerster 1862: 258. nia, Russia, Sweden. Biosteres blandiis; Marshall 1891: 54, ?; Hellén Host. Diptera, Anthomyiidae: Pegomya sp. es- 1946: 133; Fischer 1971a: 21. iiriens-group on Betu viilgaris. Opiiis (Biosteres) blandiis; Fischer 1959a (figs.). Material e.rarnined. Id from Finland (MZH), St: Re- Biosteres (Chilotrichia) blandiis; Fischer 1972: posa&, 11.9.1950 (V. Lauro); and 2d from Russia, Karelia (MZH). 487 (key), 494 (redescription): Papp 1979d: 231. Opius (Chilotrichia) blandiis; Tobias 1986: 14. Biosteres (Chilotrichia) clypealis (Thomson) Chilotnchia blanda; van Achterberg 1997: 19 (new combination, not regarded here). Opiiis clypealis Thomson, 1895: 2202, 0 ;Fischer Distribiition. Austria, former Czechoslovakia, 1964b: 258, new a). Denmark, England, Finland (map 133), France, Opiits (Biosteres) clypealis; Fischer 1959a: 14. Germany, Hungary, Ireland, Italy, Russia, Swe- Biosteres clypealis; Fischer 1969b: 83; 197 1a: 23. den, Switzerland, Mongolia. Biosteres (Chilotrichia) clypealis; Fischer 1972: Host. Diptera, Agromyzidae: Agromyza si. on 487 (key), 498 (redescription). Vicia sylvatica. 0pi::s (Chilotrichia) clypenlis; Tobias 1986: 1 1. Marerial exaniined. 35 8, 19 O from Finland (DAZH, Distribiition. Denmark, Finland (map 133, h1ZH). Ab: Lohjan kunta, 29.5.1992 (M. Koponen), Vihti, France, Germany, Latvia, Lithuania, Russia (Kare- 17.61952. 6.6.1964 íV. J. Karvonen); N: H:fors = Helsinki. .. 7.1941 (A. Nordnian), 28.6.1976, 6.6.1980, 18.5.1983 (hl. iiit), fii iiiciiia. Koponen), 2.6.1977 (Eila Karvonen), Helsinge, Hopisx, Host. Unknown species on Datiirn sp. 28.5.1916 (M. Hellén), Kirkkonummi, 30.6.1975 (V. J. Kar- Marerial e.umitzed. 2 d,4 O from Finland (DAZH, MZH), vonen), Munksnas (W. Hellén), Nurmijarvi, 25.5.1988, Ab: Vihti, 25.6.1966 (V. J. Karvonen); N: Hanpo = Hanko 19.5.1990, 31.5.1992, 15.5.1993 (M. Koponen), Sibbo, Ne- (A. Berp), Tvarminne (B.Poppius); St: Ylane, 6.7.1872 (J. vas, 1 .-7.6.1986 (A. Albrecht). Sipoo, 24.5.1983 (M. Kopo- Sahlberg); Ta: Hattula (L. v. Essen); Oa: Solf = Sulva, 4.7. nen), Tammisaari, 20.5.1981 (M. Koponen), Tikkurila, 1930 (Hakan Lindberg); and 18 from Russia (MZH), Kare- 4.6.1922 (J. Listo); Ka: Virolahti, 1.-6.7.1971 (Karvonen); lia: Soutjarvi, 4.10.1932, rcared from Diptera, Pegotnya sp. St: Ahlaiiien, 1.6.1959 (V. Lauro); Ta: Hartola (L. von Es- or Rhagoleris sp. (L. Tiensuu). Fischer & Koponen: A survej of Opiinae ENTOMOL. FENNICA Vol. 10 .

o Bwsteres (Chilotrichia) haemomhoeus (Haliday) France, Hungary, Moldova. A new species to Finland. Opius haemorrhoeirs Haliday, 1836: 2 19, ? d ; Host. Unknown. Biosteres haemorrhous; Marshall 1891: 53, 0 d ; Mureriul exumined. 1d, 2 P frorn Finland (DAZH), Ab: Hellén 1946: 133; Fischer 1971a: 25. Turku, 19.5.1979; N: Helsinki, 16.8.1979 (M. Koponen lec.). Opius (Biosteres) haemorrhoeus; Fischer 1959a: 20. Biosteres (Chilotrichia) riisticus (Haliday) Biosteres (Chilotrichia) haemorrhoeus; Fischer 1972: 488 (key), 500 (redescription). Opiirs rusticcrs Haliday, 1837: 2 18; Hellén 1959: Opius (Chilotrichia)haemorrhoeus; Tobias 1986: 22. 14 (key, distribution). Biosteres rusticus; Marshall 1891 : 54; Fischer Biosteres (Btetes)haemorrhoeus; van Achterberg 1971a: 30. 1997: 45. Opiirs (Biosteres) rusticus; Fischer 1959a: 23 (re- Opius castaneiventris Thomson, 1895: 2201, ? d ; description). Fischer 1959a: 13. Synonymised by Fischer Biosteres (Chilotrichia) rusticus; Fischer 1972: 1971a: 25. 488 (key), 508 (redescription). Distribirtion. Austria, Belgium, Corsica, Opiirs (Chilotrichia) rusticus; Tobias 1986: 15 former Czechoslovakia, Denmark, England, Fin- (key, distribution). land (map 136), France, Germany, Hungary, Ire- Biosteres (Zetetes) rusticus; van Achterberg 1997: land, Poland, Russia, former Yugoslavia, Geor- 75. gia, East of Lake Baikal, North America. Distribution. Austria, former Czechoslovakia, Host. Diptera, Anthomyiidae: Pegomya bi- Denmark, England, Finland (map 138), France, color (Hoffmannsegg in Wiedemann), Pegomya Germany, Ireland, Russia (European part, Altai, holosteae (Hering) on Stellaria holostea, Pego- Far East), Switzerland. mya nigritarsis (Zetterstedt) on Rrtmex sp. Un- Hosts. Diptera, Anthomyiidae: Delia echina- known host on Chenopodium polyspetmum. ta Séuguy on Dianthus barbatus, Pegomya sp.. Mureriul e,rumined.4 d. 1 O frorn Finland (DAZH, MZH), esirriens group, Phorbia sp. on Cerastium holoste- N: Helsinki, 29.6.. 6.9.1980 (M. Koponen), Tikkurila, oicies. 24.6.1924 (Agricultura1Research Centre, Departrnent of Pest Muteriul e.rurnined. 34d, 40 O frorn Finland (DAZH, Investigation); Ta:Tammela (Helenius leg.); 28, 1 O frorn MZH), Al: Lemiand, 13.8.1953, 12.6.-28.7.1954 (CVaselius); Russia (MZH), Leningrad region: Sakkola, 13.5.1920 (W. Ab: Brornarv, 11.-18.8.1953 (R.Gronblom), Nauvo, Sando, Hellén), Vyborg, 22.6.1920 (E. Thuneberg); Karelia: Vala- 19.7.1978 (M. Koponen), Vichtis = Vihti (R.Frey); N: Esbo mo (Woldstedt leg.); 1d frorn Poland (DAZH), Bialowieza, = Espoo (R.Frey), Helsinki, 14.8.1975,30.8.1976, 10.6., 11.- 8.7.1988 (M. Koponen); and 1 6 frorn Georgia (DAZH), Ba- 18.7.1979, 9.6.1980, 28.7.1981, 27.9.1983, 29.9.1987 (M. tumi, 9.7.1977 (M. Koponen leg.). Koponen), Kirkkonummi, 8.8.1954 (V. J. Karvonen), Num- ijarvi, 1.9.1975, 16.5., 14.7., 18.8., 1.-2.9.1979, 24.8.1980 Biosteres (Chilotnchia) pir nctiscii ta homson) (M. Koponen), Sibbo, 22.-28.8.1953, 19.-31.8.1955, 29.5.- (T 5.6.1989 (A. Albrecht), Tikkurila, 27.6.1920, 24.6.1922 (Y. Hukkinen), Tvarminne, 25.6.195 I (P. Palrngren), Vantaa. Opiiis piinctiscirta Thomson, 1895: 2204, O ; 2.9.1977, 5.9.1978, 7.9.1979 (hI. Koponen); St: Reposaari, Opiiis punctisciitis; Marshall 1898: 3 16, Y . 18.9.1950.9.7., 27.8.1951.9.8.-7.9.1954 (V.Lauro);Ta: Jana- Opiirs (Biosteres)punctiscuta; Fischer 1959a: 2 1. kkalri, 12.9.1982 (M. Koponen), Kangasala, 19.6.1938 (A. Saarinen). Lahti, 15.8.1980, Nastola, 18.9.1977, Sysrr,S, Biosteres punctiscutcr: Fischer 197 1a: 29; Papp 27.9.1978 (M. Koponen); Sa: Imatra, 8.-11.6.1956 (Hakan 1979a: 92. Lindberg), Mrintyharju, 5.9.1981, hlikkelin nilk., 27.8.1975, R iostc res ( CIi ilnt rich in 17 I I n c t i.r c i I fn : Fi s c h er IS.7.1987, 4.8.19S9 (hl. Koponen): and 2 0 from Ruscia 1972: 488 (key), 505 (redescription); Payp 1979: (MZH). Leiiiiigrd region: i<;uolciiiajar\ 1. 19.-1ü.7.1S9Y (1. 92; Fischer 1984a: 128. H.Jkrvi). 8.6.1930 (Ikaschnikoff lcg.). Opiirs (CI~ilotrichicr)pcrnctisciitcr;Tobias 1986: 16 (key) . Biosteres (Chilotrichia) scabricir liis (Wesmael) Opiiis nzoIdai~icirsJakimavicius, 1977: 86. Syno- nymised by Papp 1979a and Fischer 1984a: 128. Opiiis scabriciilirs Wesmael, 1835: 154; Hellén Distribirtion. Depmark, Finland (map 137), 1959: 22. ' 9 ENTQMOL. FENNICA Vol. 10 O Fischer & Koponen: A survey ofOpiinne 149

Biosteros scabriculus; Marshall 1891 : 55; FiscLer Bidsteres carbonariux; Hellén 1946: 133; Fischer 1971a: 30. 1974: 85. Opius (Biosteres) scabriculus; Fischer 1959a: 25. Opius Wesmaelii Haliday, 1837: 2 19; Fischer Biosteres (Chifotrichia)scabriculus; Fischer 1959a: 31. 1972: 487 (key), 510 (redescription); 1995: 236, Opius wesmaeli; Hellén 1959: 22. figs 28-3 1. Biosteres wesmaelii; Marshall 1891: 55, 9 O; Opius (Chilotrichia) scabriculus; Tobias 1986: 11. Fischer 1967b: 161; 1971a: 32. Distribution. Belgium, former Czechoslova- Biosteres (Chilotrichia) wesmaelii; Fischer 1972: kia, Finland (map 139), Germany, Hungary, Ire- 487 (key), 517 (redescription); 1981a: 129. land, Russia. Opius (Chilotrichia)wesmaelii; Tobias 1986: 13. Host. Unknown. Biosteres (Zetetes) wesmaelii; van Achterberg Material examined. 3 d,3 ? from Finiand (DAZH, MZH), 1997: 89,90. N: H:fors = Helsinki, 2.10.1961 (A. Nordman), NurmijWi, Opiiis jonaitisi Jakimavicius, 1977: 86. Syno- 22.9.1974,3.10.1982 (M. Koponen); St: Reposad, 6.10.1950 (V. Lauro); Sa: Mikkelin rnik., 6.8.1993 (M. Koponen); Ks: nymised with wesmaelii by Fischer 1981a: 129. Kuusamo, 3.7.1979 (M. Koponen leg.). Distribiition. North, west and central Europe, including Finland (map 141), North America. Biosteres (Chilotrichia) sylvaticus (Haliday, Hosts. Diptera, Anthomyiidae: Delin quadri- 1836) pila (Stein) on Honckenya peploicies; Pegomya betae (Curtis) and Pegomya hyoscyami (Panzer) Biosteres (Zetetets) sylvaticus; van Achterberg on Befa vulgaris, Pegomya hyoscyami or Musci- 1997:79. na stabulans Fabricius on Hyoscyamiis niger, This species was reported by Hellén (1946) Pegomya nigrita rsis (Zetterstedt) ,Pegomya setar- from Finland, but the specimens in the MZH with ia (Meigen). In Finland reared from Pegomya this name were misidentified. This species is de- hyoscyami by J. Listo. leted from the Finnish list. Material examined. 42d, 459 from Finland (DAZH, MZH), Al: Lerniand, 8.8.1953 (W. Hellén), Lemiand, Haka, 13.8.1953 (Hakan Lindberg); Ab: Pojo, hnne, 14.7.1965 Biosteres (Chilotric hiu) u ltor (Foers t e r) (M. Meinander), Runsala = Turku, Ruissalo (Ingelius), Nys- tad = Uusikaupunki. 5.8.- 17.9.1914,13.- 15.8.1920,1.7.192 1, Zetetes ultor Foerster, 1862: 258. Vichtis = Vihti (R.Frey), 1929 (J. Listo); N: Esbo = Espoo. Opius (Biosteres) ultor; Fischer 1957a: 52, O; 1.6.1916 (M. Hellén). Hanko, 17.6.1980 (M. Koponen), Hels- inge, 16.6.-31.7.1915 (W. Hellén), H:fors = Helsinki (W. 1959a: 31. Nylander, W. Hellén), 24.8.1977, 10.6.1979, 26.6., 26.7., Biosteres ultor; Fischer 1971a: 32. 6.9.1980 (M. Koponen), Munksnas = Munkkkiniemi (R. Biosteres (Chilotrichia) ultor; Fischer 1972 :487 Frey), Nurmijarvi, 16.5.1979, 23.8.1980, 17.7.1981, 14.- (key), 5 16 (redescription); 1983: 36. 17.8.1985,14.8.,21.9.1989,26.7.1992(M.Koponen),Strom- fors, 8.6.1974 (A. Albrecht), Tikkurila, 24.6.1922 (Y. Huk- Opizis (Chilotrichia) ultor; Tobias 1986: 11. kinen), Tvarminne (A. Nordman), 6.-10.7.1914 (W. Hellén), Distribiition. Austria, Belgium, former Czech- 17.6.1966 (V. Varis), Vantaa, 13.8.1977, 5.9.1978, 6.8.1980 oclcvakia, Denmark, England, Finland (map 140), (M. Koponen); Ka: Kymi. 1929 (J. Listo), Sippola, 1 1.6.1978 Germany. Hungary, Ireland. íM. Viitasaan); St: Eurajoki, 17.8.1920 (E. Kalervo), Re- posaari. 24.7.1950 (V. Lauro); Ta: Asikkala, 1925 (Nurmin- Host. Unknown. In Finland reared from sam- en), Hartola, 11.6.1987 (bl. Koponen), Hattula (L. von Es- ple of Riimex sp. by E. O. Peltonen. sen), Janakkala, 13.8.1979, iz=5rkC!Z. 15.8.1974 (hl. Kopo- Mriferid e.rrimined. 2 d ,2 0 from Finland (DAZH, MZH), nen), Tamiela (Helenius), Tammerfors = Tampere (Woldst- N: Helsinki, reared 19.12.1977 (E. O. Peltonen), Nunnijibvi, edt), Ylojarvi, 1.8.1937 (S. Saarinen); Sa: Luumaki, 2.7.1975, I 1 .h. 19S3 (M. Koponen); Ta: Tarnniela (Hslenius); Kb: Li- Mikkelin rnlk.. 12.6.1977, 15.8.1951, 24.6.1958, 3.6.1989, peri, 3.7.1994 (M. Koponen 1eg.k Kistiiiia, 20.6. ¡%O (M. Kopurien): Oh;i: ii, 2u.7. i 9S0 (M. Koponen), Oulu (Wuorentaus): Le: Enontekio, Kilpisjarvi, 8.7.1919 (Gronvall,A. Nordman), Enontekio. Saana (T. Gron- Biosteres (Chilotrichia) wesrnaelii (Haliday) blom); 2 0 from Sweden (MZH); 2 d,4 0 froni Russia (MZH), Leningrad and Murmansk regions; 3 6,2 O from Estonia Opius cnrbonarius Wesmoel, 1835: 152 (not Bra- (DAZH); and 1 0 from Poland (DAZH). con carbonariris Nees, 1834); Hellén 1959: 21. 1$Q Fischer & Koponen: A siirvej of Opiinae 0 ENTOMOL. FENNICA Vol. 10

Subgen& Biosteres Foerster, s. str. Biosteres (Biosteres) impressur;Fischer 1972: 522 (key ), 53 O (redescription) . Biosteres Foerster, 1862: 259. Distribution. England, Finland (map 143), Ire- Rhabdospilus Foerster, 1862: 259. Type species: land, Central Europe, former Yugoslavia, Russia Opitis placidiis Haliday. (European part, Kanichatka), Japan, North Ko- rea, North America. Biosteres (Biosteres) breviszilcus (Thomson) Hosts. Diptera, Anthomyiidae: Pegomya betae (Curtis) and Pegomya hyoscyami (Panzer) on Opius brevisulcus Thomson, 1895: 2203; Hellén Beta vulgaris, Pegomya nigritarsis (Zetterstedt) 1959: 21: on Rumex acetosa, Pegomya nigrisquama (Stein) Opius (Biosteres) brevisulcus;Fischer 1959d: 520; on Solidago virgaurea, Pegomya steini Hendel, Tobias 1986: 19 (key, distribution). Delia antiqua (Meigen), Delia brassicae (Hoff- Biosteres brevisulcus; Fischer 1969: 49 (figure); mannsegg in Wiedemann). 1971a: 22; 1974: 85. Material examined. 53d, 68 O from Finland (DAZH, Biosteres (Biosteres) brevisulcus; Fischer 1972: MZH), Al: Lemland, 13.8.-18.9.1953 (Wegelius), Mariehamn, 9.7.1968 (W. Hellén); Ab: Nagu, 10.1953 (Th. Clayhills), 523 (redescription); Papp 1979a: 91; 1979d: 234. Vichtis =Vihti (R. Frey); N: Hango = Hanko, 1931,20.7.1969 Opius melanosoma Szépliegti, 1896: 3 18, 384, (W. Hellén), 20.8.1978, 17.6.1980 (M. Koponen), 14.- 3 d. Synonymised by Papp 1979a. 26.6.1989, 9.-20.8.1990 (A. Albrecht), Helsingin pit., 1951 Distribution. Austria, Finland (map 142), Rus- (1. Jalas), H:fors = Helsinki (R. Frey), 23.9.1915, 18.7.1918 (W. Hellén), 11.9. 1976, 15-18.9.1978, 11.-12.7., 8.10.1979, sia (Leningrad region, Hellén 1959), Sweden, 14.-26. 6.1980, 15.6.1981 (M.Koponen), Inkoo, 5.7.1941 former Yugoslavia, Russian Far East (Kamchat- (A. Saarinen), 24.5.1989 (M. Koponen), Nurmijarvi, ka) . 25.6.1978, 16.5, 14.7.1979,7.8.1950, 11.6.1983, 11.6.1986, Host. Unknown. 19.6.1987, 2.6.1992, Fredriksb. = Helsinki, Pasila (R.Frey), PernS = Pernaja, 15.6.1960 (A. Nordstrorn), Sibbo = Sipoo, Material examined. 48,4 O from Finiand (DAZH, MZH), 13.8.1977 (M. Koponen), 23.6.-31.8.1985, 22.6.-13.7.1986, Ab: Nauvo, Seili, 20.7.1978 (M. Koponen), Abo = Turku, 14.-22.9.1989 (A. Albrecht), Ekenk =Tammisaari, 29.6.1948 10.6.1919 (W. Hellén); N: Elimaki, 16.6.1984 (M. Koponen); (Hakan Lindberg), Tikkurila, 24.6.1922 Hukkinen), Van- St: Suoniemi, 23.6.1950 (A. Saarinen); Ta: Loppi, 12.7.1991 (Y. taa, 22.8.1978, 6.8.1980 (M. Koponen); Ka: Vehkalahti, (M. Koponen); Ks: Kuusamo, 2.7.1979 (M. Koponen); and 14.8.1953 (Fagerstrom); St: Karkku, 6.6.-3.7.1914 (W. Hel- Russia (MZH): Karnchatka, 20.7.1917 (Y. Wuorentaus). En), Menkarvia, 26.6.1978 (M. Koponen), Reposaari, 7.8., 18.9.1950.27.7.1954 (V. Lauro);Ta: Aitolahti, 21.6.1932 (A. Biosteres (Biosteres) carbonarius (Nees) Saarinen), Hartola, 11.6.1987 (M. Koponen), Hattula (L.von Essen), Harneenlinna, 4.6.192 1 (Y. Hukkinen), Jokioinen, 20.6.1987, Lammi, 12.-15.7.1976 (M. Koponen), Messuby Bracon carbonarius Nees, 1834: 58, 3 d. = Tampere, Messukyla (R. Frey), Ruovesi, 22.6.1931 (V. Biosteres carbonarius; Foerster 1862: 259; Saarinen), Saaksrnaki, 17.6.193 1 (E. Kivirikko); Sa: Irnatra, Fischer 197 1a: 22. 11.6.1970 (E. Thuneberg), 16.8.1954 (O. Nybom), Mikkelin Opius carbonnrius; Hellén 1959: 2 1. mlk., 22.8.1975, klantyharju, 2.6.1989 (M. Koponen), Ran- tasalmi, 15.7.1941 (S. Hellén),Ristiina,23.6.1979,21.6.1985, Opius (Biosteres) carbonarius; Fischer 1959d: 4.7.1987, 3.6.1989 (M. Koponen); Kl: Pankkala, 15.6.1940 523 (redescription). (LV. HellCn); Tb: Laukaa (Woldstedt); Sb: Nilsia, 17.6.1985 Biosteres (Biosteres) carbonarius; Fischer 1972: (M. Koponen); Kb: Lipen, 17.6.1988, 2.7.1994 (M. Kopo- 523 (key), 525 (redescription); van Achterberg nen); Ok: Paltamo, 6.7.199 1, Ristijarvi, 3.7.199 1 (M.Kopo - nen), Suomussalmi (Sorsakoski), 29.6.1991 (M. Koponen); 1975: 175: Papp 1981c: 139. Obb: Simo, 17.7.1980 (M. Koponeii); 83 3? from Russia Opius (Biosteres) carbonarius; Tobias 1986: 17. (MZH), Lenigrad region, Karelia. Murrnansk region; 1 O from Opius proceriis Wesmael, 1835: 156, d; Syno- Estonia, Saarenua: Viidurnapi, Tarnmiku, 1.6.1990 (M. Ko- njmised !y’ ?;r.chr~ 1959c!: 52.3. \Vnrtor?. 19S7: ponen): 3 Q from Nonrny (FLIZH). Sarek íR. Poppiusl Dovre. ll.7.i953 (W. HeilEri): 1 d 2 9 tron! Polarid (DAZH): aiid 67. 1 5 froni Austria (MZH). Nassfeld. 1570 (J. A. Palnign leg.). Biosteres procerus; Hellén 1946: 133. Opius irnpressrrs Wesmael, 1835: 157, d. Syno- Biosteres (Biosteres) cuiidutiilzis (Thomson) nymised with carbonarius by van Achterberg 1975. Opiiis caudatuliis Thomson, 1895: 2204, 9 ;Hel- Biosteres inipressus; Marshall 1891: 52, 3 d ; lén 1959: 21. Fischer 1971a: 25; 1974: 85. ' ENTOMOL. FENNICA Vol. 10 0 Fischer & Koponen: A survey ofopiinae 151

c Biosteres caudatuliis; Hellén 1946: 134; Fischer 1972: 522 (key), 528 (redescription); Papp 1979: 1967a: 334; 1971a: 22. 92. Opiiis (Biosteres)cniidatulus; Fischer 1959d: 526; Opiiis (Biosteres) colorativentris;Tobias 1986: 17. Tobias 1986: 20. Distribution. Austria. Finland (map 145), Ger- Biosteres (Biosteres) caudatulus; Fischer 1972: many, Hungary, Sweden. 523 (key), 527 (redescription); 1995: 219 (fig), A new species to Finland. 232. Hosts. Diptera, Anthomyiidae: Chirosia his- Biosteres longicauda; Hellén 1946: 133, misiden- tricinn Rondani on Pteridium aquilinum; Scat- tified. ophagidae: Chjlizosotna beckeri Séguy Polygo- Distribution. former Czechoslovakia, Den- natrtm miiltijloriri~i. mark, Finland (map 144), Russia, Sweden, Swit- Marerial examiried. 7 O from Finland (DAZH, MZH), zerland. Ab: Pernio, 18.9.1991(M. Koponen); N: Helsinki, 21.8.1978, 30.8., 15.9., 14.10.1980 (M. Koponen), PemA = Pemaja, Hosts. Diptera, Anthomyiidae: Pegomya hyo- 19.7.1945 (A. Nordstrorn); Sa: Ruokolahti, 4.7.1975 (M. scyami (Panzer) on Chenopodiiim boncrs-henri- Koponen). CliS. Material examined. 48 (uncertain identification), 131 O Biosteres (Biosteres) magnicornis (Wesmael) from Finland (DAZH, MZH), Al: Eckero. 2.-7.7.1984, Sund, 4.7.1984 (M. Koponen); Ab: Houtskar, 6.6.1936 (W. Hel- En); Ab: Korppoo, Avensor, 20.7.1978, Nauvo, Sando, Opius magnicornis Wesmael, 1835: 155, 9. 19.7.1978, Nauvo, Seili, 17.-20.7.1978 (M. Koponen); N: Biosteres magtiicornis; Marshall 1895: 348; Hango = Hanko, 1931, 19.7.1970, 15.7.1972 (W. Hellén), Fischer 1971a: 27. 29.7.-3.8.1981 (Kock & Perttula), Helsinki, 19.7.-25.8.1976, 23.8.1977,24.-28.7.1978,4.8.1980 (M. Koponen), 30.7.1982 Opius (Biosteres) magnicornis; Fischer 1959d: (Y. Zhongqi), Sveaborg = Helsinki, Suomenlinna (R. Frey), 532; 1963: 15, d. Nurmijarvi, 10.-29.7.1982, 7.7.1983, 21.7.1984, 13.7.- Biosteres (Biosteres) magnicornis; Fischer 1972: 17.8.1985, 29.7.1987, 8.7.1990, 17.7.1994 (M. Koponen), 522 (key), 532 (redescription). Tvarminne, 29.7.1923, 15.-31.7.1964 (R. Frey), Vantaa, 11.8.1979 (M. Koponen); Ka: Ylamaa, 13.8.1987 (M. Kopo- Opius (Biosteres) magniconiis; Tobias 1986: 17. nen); St: Rauman mik., 2.7.1986 (M. Koponen), Reposaari, Biosteres bicolor; Hellén 1946: 133, misidenti- 27.-29.7.1954 (V. Lauro); Suodenniemi, 22.7.1933 (E. fied. Thuneberg); Ta: Hattula, 11.7.1976, Heinolan mik., 20.7.1983 Distribiition. Austria, Belpium, Denmark, Fin- (M. Koponen), Hameenlinna, 20.8.196s (E. Valkeila), Ko- rpilahti, 20.8.1986, Lammi, 17.7.1975, Loppi, 12.7.1991, land (map 146), Hungary, Russia. 1.8.1992 (M. Koponen); Sa: Anttola, 15.8.1976, Mikkelin A new species to Finland and Russia. mlk., 24.7.1976, 11.8.1987, 4.8.1989, Ristiina, 6.8.1977, Host. Unknown. 31.7.1950, 29.7.1983, 11.8.1987, 23.7.1989, 23.7.1994 (M. Marerial e.ramined. 1d,2 O from Finland (DAZH, MZH), Koponen), Vakeala, 8.7.1974 (M. Viitasaari); Kl: Rautjarvi, N: Helsinki, 18.9.1975 (M. Viitasaari); Ta: Lammi, 11.7.1975 4.7.1990 (M. Koponen), Uukuniemi, 25.7.1940 (W.Hellén); (M. Koponen), Saaksmaki, 15.6.1921 (W. Hellén); and Oa: Yliharma, 13.7.1980 (M. Koponen); Tb: Keuruu, lEfrom Russia, Leningrad region: St. Andrae = Antrea, 26.7.1928 Hellén), Rautalampi, 18.-19.7.1953 (M. Ko- (W. 1.7.1920 (W. HellCn). ponen), Ahtari (R. Frey); Sb: Varkaus, 15.-20.7.1963 (E. Pul- liainen): Kb: Ilomantsi, 20.6.1865 (Vi. Woldstedt), Polvijarvi, 2.8.1865 (Gronvik), Tohmajarvi, 21.-22.7.1952 (M. Kopo- Biosteres (Biosteres)placidirs (Haliday) nen): Ok: Kajaani, 16.8.1952, Paltamo. 10.7.1959 (M. Ko- ponen); Oba: Ii, 20.7.1950, Utajarvi, 14.8.1983 (M. Kopo- Opiiis placidirs Haliday, 1836: 149, ? . nen); Lkor: Savukoski, 10.8.1983 (M. Koponen); and 3 O from Rtissia (MZH). Leningrad region and Chita region. 1842 (R. Rhabdospilirs placidirs; Foerster 1862: 259. E Sahlbeg). Biosteres placidirs; Marshall 1891: 56; Fischer 1971a- 3Q. Biosteres (Biosteres) colorativeiitris (Fischerj Qpiiis (Diachasnicr)placidiis; Fischer 1458a: 57. Opiiis (Biosteres) placidiis; Fischer 1959d: 536; Opiiis (Biosteres)colorativentris Fischer, 1957b: Tobias 1986: 17. 211, 0. Biosteres (Biosteres)placidiis; Fischer 1972-S22 Biosteres colorativentris; Fischer 1967a: 334; (key), 535 (redescription). 1971a: 23. Biosteres (Biosteres) placodus; van Achterberg Biosteres (Biostcres) colorativentris; Fischer 1997: 67. 6

1 152 Fischerd Koponen: A survey of Opiinae ENTOMOL. FENNICA Vol. 1O

o Opius melanocerus Wesmael, 1838: 149, O. Syn- Bathystomus xanthopus Foerster. onymised by Fischer 1959d. Opius (Diachasma); Fischer 1959a: 57. Biosteres melanocerus; Marshalll895: 347; Hel- lén 1946: 133. Diachasma rugiventre (Thomson) Opius tarsator Thomson, 1895: 2193, O. Syno- nymised by Fischer 1959d. Opius rugiventris Thomson, 1895: 2178; Hellén Biosteres indotatus Viereck, 1905: 273, O. Syno- 1946: 132. nymised by Fischer 1965b: 152. Opicis (Diachasma) riigiventris; Fischer 1959e: Distribution. Austria, Belgium, Finland (map 158. 147), Ireland, Russia, Sweden,'Canada, U.S.A. Diachasma rugiventre; 1971a: 35; Fischer 1972: Host. Diptera, Anthomyiidae: Pegomya nigri- 543 (key), 556 (redescription). tarsis (Zetterstedt) on Rumex sp. Opius (Diachasma) rugiventris; Tobias 1986: 23. Material examined. 69 frorn Finland (MZH), N: PemA Distribution. Denmark, Finland (map 149), = Pemaja, 3.9.1944, Sibbo = Sipoo, 7.-15.7.1985 (A. Albre- Germany, Switzerland. cht), Tvarminne, 20.-26.9. 1989 (A. Albrecht). We have not seen the specimens frorn Ab: Abo = Turku and N: Helsinge Host. Unknown. reported by Hellén (1946). Marerial examined. 1 9 frorn Finland (MZH), Ta: Saaksmaki, 18.7.1940 (E. Kivirikko). Biosteres (Biosteres) spinaciue (Thomson) Tribus Desmiostomatini Opius spinaciae Thomson, 1895: 2201, d ; Opius (Biosteres)spinaciae; Fischer 1959d: 539. Genus Desmiostoma Foerster Biosteres spinaciae; Hellén 1946: 133; Fischer 1971a: 30. Desmiostoma Foerster, 1862: 260. Type species: Biosteres (Biosteres)spinaciae; Fischer 1972: 522 Opius parvulus Wesmael. (key), 538 (redescription); ipapp 1979: 93. Opius (Biosteres) spinaciae; Tobias 1986: 17. Desmiostoma parvulum (Wesmael) Opiuspegomyiae Gahan, 1917: 201, ? d. Opius (Chilotrichia) hyoscyamiellus Viereck, Opiiispawulcis Wesmael, 1835: 139, ? 8 ;Fischer 1958b: 189; 1971a: 96. 1925: 278, d. Distribution. Austria, former Czechoslovakia, Desmiostonta parvuliim; Fischer 1972: 567 (re- Denmark, Finland (map 148), Germany, Poland, description); Tobias 1986: 9. Russia, Sweden, Mongolia, Canada (Quebec), Opius nwdiscutum Fischer, 1964a: 373, O O. Syn- U.S.A. (from Alsk. to Calif.). onymised by Fischer 1970: 3 1 1. Hosts. Diptera, Anthomyiidae: Pegomya be- Distribiition. Belgium, Denmark, England, tne (Curtis) and Pegomya hyoscyami (Panzer) on Finland (map 150), Germany, Hungary, Switzer- Beta vulgaris and Rumex acetoselln, Pegornya ni- land, Canada, U.S.A. (Calif.), Bermuda. gritarsis (Zetterstedt). A new species to Finland. Material examined. 1 d frorn Finland (MZH), N: Hango Hosts. Diptera: Agromyzidae: Agromyza ni- = Hanko, 12.7.1914 (W. Hellén); and 3 8 from Russia (MZH), grelln Rondani on Hordeiim iwlgare, Liriomyza Leninpd region: Kivinebb = Kivennapa (K. Ehnbeg); Ros- fnsciola aiid Phytomy,-n sp. on Bellis perennis. tov region: Taganrog. 20.-28.5.1925 (C. Ahnger). Material e.ramined. 1 ¿ ,2O from Finland (DAZH, MZH), N: Degero, 26.6.1945 (A. Saarinen); Lkor: Sodankyla. 3n h 1980 chl Kopnncv): 1 i. Tnnri 12 8 1983 (M Knponcn 1C.g.).

Diachasma Foerster, 1862: 259. Type species: Acktio,cled~errzetzrs.\Ve are grateful to Anders Albrecht Opiiis fiilgidiis Haliday. for his help in producing the distribution rnaps and Mark R. Atoreiiteiis Foerster, 1862: 241. Type species: Shsw for his valuabie comrnents on the manuscript. Hans Atoreuteris striatiis Foerster. Silfverberg has helped to find notebooks of some of the col- lectors in MZH. We are indebted also to Lisa Heidorn for the Bathystomiis Foerster, 1862: 235. Type species: English language revision. l .<

, EWOMOL. FENNICA Vol. 10 0 Fischer & Koponen: A suwey of Opiinae 153

Riferences Wesrn. Teil VI. - Ann. Mus. Civ. Stor. Nat. Genova 71: 145-16 1. Fischer, M. 1959fi Die europaischen Arten der Gattung Opi- Achterberg, C. van 1975: About the identity of Biosteres us Wesrn. Neue Revision der fallax-Gmppe. - Ann. (Biosteres) impressus (Wesmael) (Braconidae, Opiinae). Hist.-nat. Mus. Natn. Hung. 51: 420-423. - Entomol. Ber. 35: 115-176. Fischer, M. 1959g: Die europaischen Arten der Gattung Opi- Achterberg, C. van 1997: Revision of the Haliday collection us Wesrn. Teil 11 c. - Opuscula Entomol. 24: 55-78. of Braconidae (Hymenoptera). - 2001. Verhandl. 3 14: Fischer, M. 1959h: Fünf neue Opius-Arten. - Entornol. 3-1 15. Nachrbl. Ost. Schweiz. Entomol. 11: 19-25. Achterberg, C. van & Salvo, A. 1997: Reared Opiinae (Hy- Fischer, M. 1959i: Neue Opius-Arten aus Polen. -Ann. 2001. menoptera: Braconidae (Hymenioptera) from Agenti- Warszawa 18: 81-87. na. - 2001. Med. Leiden 71: 189-209. Fischer, M. 1960 Die europaischen Arten der Gattung Opi- Ashmead, W. H. 1900 (1901): Classification of the Ichneu- us Wesm., Teil IVa. - Ann. Zool., Warsz. 19: 33-1 12. mon flies, or the superfamily Ichneumonoidea. - h-oc. Fischer, M. 1961: Erganzungen zur Kenntnis der palaark- U. S. natn. Mus., Washington 23: 1-220. tischen Opius-Arten. - Ann. Mus. Civ. Stor. Nat. Gen- Carneron, E. 1907: Percy Sladen Trust expedition. No. IV. ova, 72: 1-16. Hymenoptera. - Trans. Linn. Soc. London 12: 69-86. Fischer, M. 1962a: Die Opius-Arten des Burgenlandes. - Dalla Torre, C. G. 1898: Catalogus Hyrnenopterorum 4: Bra- Wiss. Arb. Burgenland, 29: 30-67. conidae. - VIII+323 pp.. Leipzig. Fischer, M. 1962b: Die Opiinae des Museo Civico di Stona Fischer, M. 1957a: Die Opiinen-Typen der Sammlung Foer- Naturale in Genua. - Ann. Mus. civ. Stor. nat. Genova ster aus dem Zoologischen Museurn in Berlin. - D. 73: 71-97. Entomol. Z., N. F. 4: 47-53. Fischer, M. 1962c: Neues über die palaarktische Opius-Fau- Fischer, M. 1957b: Neue Opius-Arten aus Schweden. - na. - Polskie Pismo Entomol. 32: 89-107. Opuscula Entornol. 22: 21 1-225. Fischer, M. 1962d: Beitrage zur Kenntnis der Wirte von Opi- Fischer, M. 1957c: Die europaischen Arten der Gattung Opi- us-Arten. - Entomophaga 7: 79-90. us Wesrn. Teil IIa. Sektion B. - D. Entomol. Z. (N. F.) Fischer, hl. 1962e: Zwei neue Opius-Arten aus Niederoster- 4: 332-358. reich. - Nachrichtenbl. bayer. Entomol. 11: 23-25. Fischer, M. 1957d: Beitrage zur Kenntnis der palaarktischen Fischer, M. 1963a: Die Opiinae Niederosterreichs (Fortset- Braconiden. - Mitt. Münch. Entomol. Ges.. 47: 1-21. zung). - Z. Arbeitsgern. Ost. Entomol. 15: 7-18. Fischer, M. 1958a: Die europaischen Arten der Gattung Opi- Fischer, M. 1963b: Die Opiinae der athiopischen Region. - us Wesm. Teil Ia. -Annali Mus. Civ. Stor. Nat. Giaco- Beitr. Entornol. 13 (1/2): 191-221, (516) 662-747. mo Dona 70 33-70. Fischer, M. 1964a: Die Opiinae der nearktischen Region. 1. Fischer, M. 1958b: Die europaischen Arten der Gattung Opi- Teil. - Polskie Pisrno Entomol. 34: 197-530. us Wesrn. Teil IiI. Opius s. str., Sektion C. - Beitr. En- Fischer, M. 1964b: Die Opiinae Danernarks. - Entomol. tornol. 8: 189-21 1. Medd. 32: 236-260. Fischer, M. 1958c: Die europaischen Arten der Gattung Opius Fischer, M. 1964c: Über Opiinae der westlichen Palaarktis Wesm. Teil IVb. -Acta Entomol. Mus. Nat. Pragae 32: (Hym., Braconidae). - Ann. Naturhist. Mus. Wien 67: 295-3 16. 409433. Fischer, M. 195Sd: Die europaischen Arten der Gattung Opius Fischer, M. 1964d: Die gezüchteten Opiinae der Sammlung Wesm. Teil IIb. -Ann. Hist.-nat. Mus. Natn. Hung. 50: Groschke. - Stuttgart. Beitr. Naturk. 136: 1-12. 241-260. Fischer, M. 1964e: Die gezüchteten Opiinae aus dem Zoolo- Fischer, M. 1958e: Die europaischen Arten der Gattung Opius gischen Museum der Humbolt-Universitat zu Berlin. - Wesm. Teil IId. - Ann. Naturhist. Mus. Wien 62: 210- Z. angew. Entomol. 55: 55-70. 2 19. Fischer, M. 19651: Die Braconidae des Steiermarkischen Fischer, M. 1958f Neue Braconiden aus dem Zoologischen Landesmuseums “Joanneurn” in Graz. -Mitt. Abt. 2001. Museum Berlin, Sarnmlung Foerster. -Mitt. Zool. Mus. Bot. Joanneurn 21: 1-29. Berlin 34: 173-1 8 1. Fischer, M. 1965b: Die Opiinae der nearktischen Region. 11. Fischer, M. 1958g: Die europaischen Arten der Gattung Opi- Teil. - Polskie Pisrno Entornol. 35: 3-212. us Wesm. Teil Ib. -Annali Miis. Civ. Stor. Nat. Giaco- Fischer, M. 1966: Opius breviscapus Thomson irn Burgen- rno Dona 70: 245-304. innd. - Nachrichtenbl. Bayer. Entornol. 15: 7-10. Fischer, M. 1959a: Die europaischen Arten der Gattung Opi- Fischer, M. 1967a: Über gezüchtete Opiinae aus Europa. - us Wesrn. Teil Va. - Mitt. Münch. Entornol. Ges. 49: I- Z. Angew. Entomol. 60: 3 18-350. 35. Fischer. M. 1967b: Die Opiinne der Steiermnrk. - Mitt Fischer, hl. 1Y59b: Die europiiischen Opiinae. -Acta Entu- Abt. Zool. Bot. Joaniieuni 36: 135-165. mol. Mus. Nat. Pragae 33: 241-363. Fischer, M. 1965: Ergebnisse der zoologischen Forschungen Fischer, M. 1959c: Ein neuer Beitrag zur Kenntnis der von Dr. Z. Kaszab in der Mongolei. 141. Opiinae. - palaarktischen Opius Wesni.-Arten. - Polskie Pisrno Reichenbachia 11: 97-1 14. Entomol. 29: 17-31. Fischer, M. 1969a: Opiinae aus dem Tiroler Hochgebirpe. Fischer. M. 1959d: Die europaischen &en der Gattung Opius - Ber. Naturw.-med. Ver. Innsbruck 57: 39-58. Wesrn. Teil V b. - Polskie Pisrno Entornol. 29: 519- Fischer, M. 1969b: Die von Dr. H. Buhr gezüchteten Opii- 542. nae. - Z. angew. 2001. 56: 65-88. Fischer, M. 1959,: Die europaischen Arten der Gattung Opius Fischer, M. 1970: Die Opiinae der University of Minnesota, l

154 Fischer & Koponen: A swvey of Opiinne ENTOMOL. FENNICA Vol. 10 4,

e Departrnent of Entornology, in St. Pacl, U.S.A. - Acta der palaarktischen Opiinae. - Sitzungsber. Osterr. Ak. Zool. Cracoviensia 15: 299-3 13. Wiss., mn. Ki. 1, 197 (1-4): 75-1 18. Fischer, M. 1971a: Hyrn., Braconidae, World Opiinae. - In Fischer, M. 1990 Palaarktische Opiinae (Hyrnenoptera, Bra- Delucchi, V. & Rernaudiere (ed.) Index of entornopha- conidae): Neue Arten und neue Funde aus dern Ungar- gous insects 5, 189 pp. ischen Naturwissenschaftlichen Museurn in Budapest. Fischer, M. 1971b: Die athiopischen Opius-Arten der Sek- - Ann. Hist.-nat. Mus. natn. Hung. 81: 205-238. tion A, aufgeteilt auf die Untergattungen. - Ann. Fischer, M. 199 1a: Wiederbeschreibungen und Neubesch- Naturhist. Mus. Wien 75: 387433. reibungen von Opiinae aus der Alten Welt (Hyrnenop- Fischer, M. 1972: Hyrnenoptera, Braconidae (Opiinae 1). - tera: Braconidae). - Ann. Naturhist. Mus. Wien 92 B: Das Tierreich 91: 1-630. 139-203. Fischer, M. 1973 (1974): Studien an Alysiinen-Typen. - 2. Fischer, M. 1991 b: Erganzungen zur Taxonornie der Opiinae ArbGern. Osterr. Entornologen 35: 47-51. mit Neubeschreibungen (Hymenoptera, Braconidae). - Fischer, M. 1974: Opiinae aus dern Otztal (Osterreich) (Hy- 2. Arbeitsgern. Osterr. Entornol. 44: 4248. menoptera: Braconidae). - Beitr. Entornol. 24: 73-86. Fischer, M. 1995: Korrekturen und Erganzungen zur Taxon- Fischer, M. 1975: Opius dureseaui n.sp. (Hyrn., Braconidae, omie altweltlicher Opiinae und Neufassung eines Bes- Opiinae) gezogen aus Agromyza frontella Rondani tirnrnusbgsschlüssels für die palaarktischen Arten des (Dipt., Agrornyzidae). - 2. Arbeitsgem. Ost. Entornol. Subgenus Opiothorax Fischer, 1972 des Genus Opius 27: 53-55. Wesrnael, 1835 (Hyrnenoptera, Braconidae). - Ento- Fischer, M. 1977: Opiinae aus Kamten, gesammelt irn Jahre mofauna 16 (9): 217-244. 1973 in der Ungebung von Hüttenberg (Hyrnenoptera, Fischer, M. 1998: Neue taxonomische Untersuchungen über Braconidae). - Carinthia 11 167 (87): 351-366. Madenwespen der Alten Welt mit besonderer Berück- Fischer, M. 1978: Die Madenwespen (Opiinae und Alysiini) sichtigung der Gattungen Eurytenes Foerster, Aulono- der FischbacherAlpen aus den Ausbeuten 1973 und 1974 tus Ashrnead, Biosteres Foerster und der Untergattung (Ins., Hyrnenoptera, Braconidae). - Mitt. Abt. Zool. Gastroserna Fischer. (Hyrnenoptera, Braconidae: Opii- Landesmus. Joanneurn 7: 45-66. nae). - Linzer Biol. Beitr. 30 (1): 2 1-5 1. Fischer, M. 1980: Opiinen aus den osterreichischen Alpen- Foerster, A. 1862: Synopsis der Familien uiid Gattungen der Iandern und angrenzenden Gebieten, gesarnrnelt von Braconen. - Verh. Naturhist. Ver. Preuss. Rheinl. 19: Herrn Dr. E. Haeselbarth (München). - Ber. nat.-med. 225-288. Ver. Insbruck 67: 185-21 1. Fulrnek, L. 1962: Parasitinsekten der Blattrninier Europas. Fischer, M. 1981a: Zur Synonymie von Opiinae, eine Neu- - Verlag Dr. W. Junk, den Haag, 203 pp. beschreibung, Redescnptionen und Erganzungen von Gahan, A. B. 1917: Descriptions of sorne parasitic Hyrnenop- ' Beschreibungen. - Boll. 1st. Entornol. agr. Oss. Fitopat. tera. - Proc. U.S. nat. Mus. 53: 195-217. Palermo 10: 227-250. Gahan, A. B. 1919: Descriptions of seven new species of Fischer, M. 1981b: Catalogus Faunae Austriae. Teil 16 j: Opius (Hyrnenoptera, Braconidae). - Proc. Entornol. Hymenoptera Fam. Braconidae. Opiinae. - Verlag der Soc. Wash. 21: 161-170. Osterreichischen Akademie der Wissenschaften: 1-36. Haliday, A. H. 1833: An essay o11 the classification of the Fischer, M. 1982: Die palaarktischen Arten der Subgenera parasitic Hyrnenoptera of Bntain, which correspond with Misophthora Foerster und Agnopius n. des Opius the Ichneurnones minuti of Linnaeus. - Entornol. Mag. Wesrnael sowie über andere Opiinae (Hyrnenoptera, Bra- London 1: 259-276. conidae, Opiinae). - Folia Entornol. Hung. 43 (1): 21- Haliday, A. H. 1837: Essay on parasitic Hyrnenoptera. - 37. Entornol. Mag. London 4: 38-59, 92-106, 203-221. Fischer, M. 1983: Illustrierte Redeskriptionen von Opiinen Hellén, W. 1916: Für die fauna Finnlands neue Braconiden aus der UdSSR und neuer Bestimmungsschlüssel zu den (Hyrn.) 11. - Notulrie Entornol. 25: 13CL-137. palaarktischen Arten des Subgenus Utetes Foerster (Hy- Hellén, W. 1958: Zur Kenntnis der Braconiden (Hym.) rnenoptera, Braconidae). - Ann. Naturhist. Mus. Wien Finnlands 11. Subfamilie Helconinae (Part.). - Fauna 85 B: 13-37. Fennica 4: 1-37. Fischer, M.19S1a: Deskriptionen und Redeskriptionen von Hellén, W. 1959: Zur Kenntnis dcr Opiinen Finnlands (Hyrn., Opiinae (Hyrnenoptera, Braconidae) aus paliiarktischen Braconidae). - Notulae Entornol. 39: 21-22. Region. - Ann. Naturhist. Mus. Wien 86 B: 85-131. Jakmavicius, A. B. 1977: New species of subfamily Opiinae Fischer, M. 19S1b: Aufteilung des Forrnenkrrises um das (Hyrnenoptera, Braconidae). - In: Skarlato, O. A. (Ed.) Subgenus Cryptonastes Foerster des Genus Opius New and little known species of insects of the European Wesmael sowic Erganzungen zum Subgenus Tolbia Cam- part of the USSR. 113 pp. Sbornik Nauchnjkh Rabot. eron (Hynienoptern, Rraconidae. Opiinw.) - Z Arhe- Aknd Nii~ikSSSR. L.enincrad. 8S-F;8 íTn Rii\sian with itsgeni. Ost. Entoniol. 36: 3340. English sumrnq.) Fischer. hl. 1956: Keue Bestimmungsschlüssel fiir palaark- Koponen, hl. 19S9: Lists of the insect tjpes in the Zoologi- tische Opiinae, neue Subgenera, Redeskriptionen und cal Museurn, University of Helsinki. 13. Hymenoptera: eine neue Art (Hymenoptera, Braconidae). - Ann. Braconidae, excluding types of the taxa described by Naturhist. Mus. Wien 88/89 B: 607-662. Wolter Helltn. -Acta Entomol. Fennica 55: 27-31. Fischer, M. 1987: Hymenoptera, Opiinar JII - atiopische, Kcponen, M. Bi Tobias, V. 1. 19S9: Lists of the insect types in onentalische, australische und ozeanische Region. - Das the Zoological Museurn, University of Helsinki. 12. Tierreich 104: 1-734. Hymenoptera: Braconidae descnbed by Wolter Hellén. Fischer, M. 19S9 (1988/89): Ein neuer Beitrag zur Kenntnis - Acta Entornol. Fennica 55: 23-25. , EN'T,OMOL. FENNICA Vol. 10 0 Fischer & Koponen: A srirvey of Opiinne 9 155

* Krogerus, R. 1927: Parasitstekiar fdn Karelska nwts dynom- - Ann. Mus. Goulandns 5: 183-193. raden. - Mem. Soc. Fauna Flora Fennica 2: 44-48. Papp, J. 1985: Taxonomical and faunistical novelties of the Marshall, T. A. 1891: A monograph of British Braconidae. Opiinae from the Old World Tropics (Hymenoptera: Bra- Part IV. - Trans. Entomol. Soc. London 1891: 7-61. conidae). -Acta Zool. Hung. 31: 185-216. Marshall, T. A. 1895: Les Braconides. - In André, E.: Spe- Papp. J. 1988: A contribution to the braconid fauna of Israel cies des Hyménopteres d'Europe et d'Algerie. 5. Beaune, (Hymenoptera), 2. - Israel J. Entornol. 22: 45-59. 632 pp. Papp, J. 1994: A revicion of the Braconidae (Hyrnenoptera) Marshall, T. A. 1898: Les Braconides (Supplérnent). - In species described by J. W. Zetterstedt. -Entomol. Scand. André, E.: Species des Hyrnénoptkres d'Europe et 25: 303-310. d'Algerie. 5 bis: 374 + 72 pp. + 15 pl. Perepetchayenko, V. L. 1994: The species of braconids new Muesebeck, C. F. W. 1933: Seven new species of reared Bra- for the fauna of Ukraine. - Izvest. Khark. Entomol. conidae (Hymenoptera). - Proc. Entornol. Soc. Wash. Obshch. 2 (2): 36-39. (In Russian) 35: 193-200. Peydro, R. J. 1983: Opius (Opiothorax) longiradialis nov. Muesebeck, C. F. W. 1967: In Krombein, K. V. & Burks, B. sp. from Spain (Hymenoptera, Braconidae). - Nouv. D. Hymenoptera of America north of Mexico: synoptig Rev. Entomol. 13: 239-241. catalog. Second supplement. - Agric. Monogr. U. S. RimsS.-Korsakow. M. 1925: Über Wasserhymenopteren der Dep. Agnc. No. 2: 1-584. Umgegend vom See Seliger. - Russ. Hydrobiol. Z. 4: Nees ab Esenbeck, C. G. 1812 (1811): Ichneumonides adsci- 67-7 1. ti in genera et familias divisi. - Mag. Ges. Naturforsch. Ruschka, F. 1915: In: Ruschka, F. & Fulrnek, L.: Verzeichnis Fr. Berlin 5: 3-37. der an der K. K. Pflanzenschutz-Station in Wien erzo- Nees ab Esenbeck, C. G. 1818: Appendix ad J. L. C. Graven- genen parasitischen Hymenopteren. - Z. angew. Ento- horst conspectum generum et familianim Ichneurnoni- mol. 2: 395412. dum, genera et familias Ichneumonidurn adscitorum ex- Sachtleben, H. 1934: Deutsche Parasiten der Kir- hibens. - Nova Acta Leop.-Carol. 9: 299-310. schfruchtfiiege. - Arb. Morph. Taxon. Entomol. Berlin Nees ab Esenbeck, C. G. 1834: Hymenopterorum Ichneu- 1: 7682. monibus affinium monograph, genera Europaea et spe- Silvestn, F, 1914: Viaggio in Africa per cercare parassiti di cies illustrantes. - Stuttgart & Tübingen, 312 pp. mosche dei frutti. Descnzione e notizie biologiche dei Papp, J. 1978a: Braconidae (Hyrnenoptera) from Korea. 3. parassiti. - Boll. Lab. 2001. Gen. Agr. Portici, 8: 3- - Acta Zool. Acad. Sci. Hung. 24: 133-148. 164. Papp, J. 1978b: New Opius Wesm. species frorn Hungary Stelfox, A. W. 1949: Descriptions of four new species be- (Hymenoptera: Braconidae, Opiinae). 1. - Acta Zool. longing to the genus Opius (Hyrn., Braconidae), - En- Acad. Sci. Hung. 24: 363-370. tomol. Mon. Mag. 85: 306-308. Papp, J. 1979a: Contributions to the braconid fauna of Hun- Szépligeti, G. 1896: Adatok a Magyar fauna Braconidáinak gary, 1. Opiinae (Hymenoptera: Braconidae). - Folia ismeretéhez. -Termesztr. Füzet. 19: 165-186,285-321. Entomol. Hung, 32 (1): 71-95. Szépligeti, G.V. 1898: Beitrage zur Kenntnis der ungarischen Papp. J. 1979b: New Opius Wesrn. species frorn Hungary Braconiden. Dritter Teil. - Termesztr. Füzet. 21: 381- and Czechoslovakia (Hyrnenoptera: Braconidae, Opii- 408. nne). 2. - Acta Zool. Acad. Sci. Hung. 25: 347-356. Szépligeti, G. V. 1901: Hyrnenoptera - In: Mocsáry, A. & Papp. J. 1979c: Braconidae (Hymenoptera) frorn Tunisia. 1. Szépligeti, V.: Hymenopteren. In: Zoologische Ergeb- - Folia Entomol. Hung. 32 (2): 175-187. nisse der dritten Asiatischen Forschungsreise des Graf- Papp. J. 1979d: Two new Biosteres Forst. species from the en Eugen Zichy. 2: 121-169. Carpathian Basin (Central Europe) (Hymenoptera, Bra- Szépligeti, G. V. 1904: Hyrnenoptera Fam. Braconidae conidae: Opiinae). - Ann. Hist.-nat. hltis. Natn. Hung. (Premiere partie). - Genera Insect. 22 and 23: 1-253. 71: 235-250. Szépligeti, G. V. 1910, In: Marshall, P.: Sur un Braconide Papp. J. 19803: Braconidae III/A - Gyilkosfürkészek IIVA (Hym.) nouveau, parasite du Dacus oleae. - Bull. Soc. 13. aicsalád: Opiinae. - Fauna Hung. 144: 1-163. Entomol. France, 1910: 241. Papp. J. 19SOb: Braconidae (Hymenoptera) from Korea. IV. Szépligeti, G. V. 1913: Neue afrikanische Braconiden aus - Acta Zaol. Acad. Sci. Hung. 26: 197-210. der Samndung des ungarischen Nstional-Museums. - Papp. J. 19Sla: Contnbutions to the Braconid fauna of Hun- Ann. Hist.-nat. Mus. Hung. 11: 592405. gary, 111. Opiinae and Microgastrinae (Hymenoptera: Thomson, C. G. 1S95: LVII bidrag ti11 braconidemas kanne- Braconidae). - Foliri Entomol. Hung. 34 (2): 139-155. dom. - Opuscula Entomol. 20: 214 1-2339. Papp. J. 19Slb: A survey of the hraconid fauna of the Horto- Tobias. V. 1. 1977: Braconids of the genus Opius \Vesm. (Hy-

!Y::, 1. í11~7:~ii !$ :z, R I';ICO I: i c!:: 6: ) . - - 1: iol!: :,!, 95>./r.: Ii iJ ,: 55: P;2t. Hist. Nritn. Paiks Iíüng. 1: 255-774. 420430. (k Rtissiün) Papp. J. 19Slc: Braconidae (Hymenoptera) from Korea. \'. Tobias, V. 1. 1956: Opiinac: p. 7-100. In: Opredelitel naseko- -ActaZool. Acad. Sci. Hung. 37: 139-15s. mych Evropejskoi tchasti SSSR 3, Perepontchntokrylye Papp. J. 1983: Taxonomical and faunistical novelties of the 5. - Opredel. Faune SSSR 117: 1-30s. (In Russian) Opiinae in thr Arctogaea (Hymenoptera, Braconidae). Vappula, N. 1962: Suomen viljelykawien tuhoelainlajisto. - - Ann. Hist.-nat. hlus. Natn. Hiing. 74: 241-253. Ann. Ag. Fenniae 1, Ser. 5, Suppl. 1: 1-275. Papp. J. 1983b: Braconidae (Hyrnenoptera) from Korea, 6. Viereck, H. L. 1905: Notes and descriptions of Hyrnenoptera - Acta Zool. Acad. Sci. Hung. 28: 105-124. from the western United States, in the collection of the Pipp. J. l'-)S?c: Braconidae (Hyrnenoptern) frorn Greece, 1. LJniversity of Kansas. ~-Trans. Kansas Acad. Sci. 19: * 156 Fischer di Koponen: A survey of Opiinae ENTOMOL. FENNICA Vol. 10

L 264-326. cation of some opiine Braconidae (Hymenoptera). - Viereck, H. L. 1925: Two new Canadian Ichneumonoidea. Proc. Entomol. Soc. Wash. 89: 61-73. - Can. Entomol. 57: 278. Wharton, R. A. 1988: Classification of the braconid sub- Waker, E 1860: Characten of some apparentiy undescnbed family Opiinae (Hymenoptera). - Can. Entomol. 120: Ceylon insects. -Ann. Mag. nat. Hist. (3) 5: 304-3 1 1. 333-300. Wesmael, C. 1835: Monographie des Braconides de Belgique. Zaykov, A. N. 1983: New for Bulgarian fauna species of sub- - Nouv. Mém. Acad. R. Sci. Bruxelles 9: 1-252. family Opiinae (Hymenoptera, Braconidae). Faunistic Wesmael, C. 1838: Monographie des Braconides de and taxonomic contribution. 1. -Acta Zool. Bulg. No. Belgique. - Nouv. Mém. Acad. R. Sci. Bruxelles 11: 23: 33-42. 1-166. Zaykov, A. & Fischer, M. 1983: Über Opiinae aus Bulgarien Wharton, R. A. 1986: Opius Wesmael, 1835 (Insecta, Hy- (Hymenoptera, Braconidae). - Ann. Nat.-hist. Mus. menoptera): proposed designation of Opius pallipes Wien (Bot. Zool.) 85: 39-51. Wesmael, 1835 as type species. 2. N. (S.) 2561. - Bull. Zetterstedt, J. W. 1838: Insecta Lapponica. Braconidae. - 2001. Nom. 43 (4): 369-371. Lipsiae: 397407. Wharton, R. A. 1987: Changes in nomenclature and classifi- 1 -,

ENTOMOL. FENNICA Vol. 10 Fischer & Koponen: A srirvey of Opiinne 157 <- 8)

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0- - ? .- . >. . '\' - - bp-f-ci . ?' .. -_. '9- --Tr -*a/ .- -;,s - --- *- c 1 Euwenes - Eurytenes Biosteres Biosteres Biosteres macrocerus (Thornson, 1895) vociremthi (Fischer, 1964) blandus (Haliday. 1837) boredis (Zetterstedt.1838) dypealis (Thornson.1895)

Map 136 Map 137 Map 138 Map 139 Map 140

-_ . .'* \. --..,' ,9; .. . * -,~ --- -- . e 5 '- .J -.' .__ .-.,.* r . r w Biosteres Biosterei Biosteres üiosterei Biosteres haemonhoeus (Haliday. 1837) puncfiscuta (Thornson. 1895) ruslicus (Haliday, 1837) Scabriculus (wesrnaei, 1835) ubr(Fdrster, 1862)

Map 141 Map 142 Map 143 Map 144 Map 145 8 .- - ..., _. -7

.. - .-8 - .. .I~. . 0,-. .\-- 3 :8'- .O -8 e . e. .-,.z*. .-,.z*. 9 . .. . -r_-. . . ** 0. :-3 - -...-3.. üiosteres Biosteres Biosteres Biosteres üiosteres WemaelJi(Haliday, 1837) brevisulcus (Thornson,1895) carbonarius (Nees. 1834) CaudatUluS (Thornson. 1895) colorativentris (Fischer, 1957)

Map 146 Map ? 4: Map 148 Map 149 Map 150

. ..

Biosteres Biosteres Biosteres Diachasma Desmiostoma mwicom (Wesmael. 1835) placidus (Haliday. 1837) spinanae (Thomson. 1895) rugiventre (Thomson. 1895) parvulum (Wesmael, 1835)