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Burrower Bugs of the Old World – a Catalogue (Hemiptera: Heteroptera

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Burrower Bugs of the Old World – a Catalogue (Hemiptera: Heteroptera Genus Vol. 10 (2): 165-249 Wroc³aw, 30 VI 1999 Burrower bugs of the Old World a catalogue (Hemiptera: Heteroptera: Cydnidae) JERZY A. LIS Department of Zoology, University of Opole, Oleska 22, 45-052 Opole, Poland e-mail: [email protected] ABSTRACT. The catalogue of all known Old World species of the family Cydnidae is presented; it includes 73 genera, 526 species and 4 subspecies. Following new synonymies are proposed: Cephalocteinae MULSANT et REY, 1866 (=Scaptocorinae FROESCHNER, 1960), Cephalocteus DUFOUR, 1834 (=Amblyottus AMYOT et SERVILLE, 1843), Byrsinocoris MONTANDON, 1900 (=Dallonia BERGEVIN, 1936), Cephalocteus scarabaeoides (FABRICIUS, 1803) (=Amblyottus dufouri AMYOT et SERVILLE, 1843), Byrsinocoris nigroscutellatus MONTANDON, 1900 (=Dallonia pangaeoides BERGEVIN, 1936), Byrsinus varians (FABRICIUS, 1803) (=Aethus riedeli JOSIFOV et KERZHNER, 1978), Geotomus punctulatus (COSTA, 1847) (=Aethus platysomoides WALKER, 1867); Canthophorus dubius var. brevipennis SIGNORET, 1884 is treated as a probable synonym of Canthophorus impressus HORVÁTH, 1881. The following new combinations are proposed: Aethus obscurus (WESTWOOD, 1837), Aethus hirsutus (HESSE, 1925), Byrsinus fuscipes (MULSANT et REY, 1866), Byrsinus pauculus (SIGNORET, 1882), Fromundus barbarae (LIS, 1991), Fromundus scutellopunctatus (SIGNORET, 1883), Microporus bimaculatus (HESSE, 1935), Microporus cruralis (STÅL, 1856), Microporus lepidus (STÅL, 1854), Microporus striipennis (LINNAVUORI, 1986), Microscytus babaeculus (LINNAVUORI, 1977), Raunoloma longiceps (LINNAVUORI, 1977), Lalervis alticola (LINNAVUORI, 1993), Lalervis tibialis (STÅL, 1854), Legnotus quadrilineus (WALKER, 1867). Parachilocoris minutus japonicus LIS, 1994 is raised to species rank (P. japo- nicus LIS, 1994). Raunoloma is proposed as a new name for the preoccupied Aloma LINNAVUORI, 1993. Lectotypes are designated for Cydnus australis ERICHSON, Cydnus flavicornis var. subinermis REY, Aethus nigropiceus SCOTT, Aethus nitens KIRBY, Choerocydnus coleopteroides BERGROTH, Hemixesta cribripennis BERGROTH, Macroscytus annulipes HORVÁTH, and Sehirus rotundipennis DOHRN. Key words: Entomology, taxonomy, catalogue, Old World, Hemiptera, Heteroptera, Cydnidae. 166 JERZY A. LIS CONTENTS I. Introduction .................................................................................................. 166. II. Catalogue ...................................................................................................... 169. 1. Subfamily Garsauriinae ........................................................................... 169. 2. Subfamily Cephalocteinae ....................................................................... 170. 3. Subfamily Cydninae ................................................................................. 172. 4. Subfamily Sehirinae ................................................................................. 218. III. References .................................................................................................... 229. IV. Index of Latin names ................................................................................... 242. I. INTRODUCTION The Cydnidae are one of the most speciose families (after Pentatomidae) within the superfamily Pentatomoidea. Its representatives are usually known under the common name of burrower bugs (or burrowing bugs) due to their specific way of life (many of them live in a soil and feed on roots). This catalogue attempts to assemble the taxonomic literature dealing with the Old World Cydnidae from 1758 to 1997, inclusively. All the names, years of publications, page numbers, distribution data and data on type materials have been checked with the original literature. Scientific names have been cited exactly as they appeared in the original publication, including typographical errors, misspelling and the use of foreign letters (except for the capitalization of the first letter in specific names, and hyphenated or separate original spelling). The type species with the means of designation are given for each genus. Where explanatory or additional information is appropriate it is placed after the taxonomic entry as a Note. The status of the type material is indicated in the section Type data. The following categories of types are included: holotype (either by monotypy, when there is clear from the original description or other sources that the species was based on a single specimen, or for one specimen originally designated as such), lectotype (previous or present designation), syntypes (for more than one speci- men), syntype(s) (if there are no indications of the number of specimens). Sex of the type material was given if it was available from the original description or other sources. The known distribution of each species is also provided, using current political units. The name of the former Yugoslavia is used only in the case when it was not possible to ascertain the exact locality from the original paper. Each doubtful country record is accompanied by a question mark; new records are marked with asterisks. The arrangement of subfamilies and tribes follows FROESCHNER (1960a) and LIS (1994a). CATALOGUE OF BURROWER BUGS OF THE OLD WORLD 167 Abbreviations for depositories of the type material: AMNH - American Museum of Natural History, New York, USA; BAU - Beijing Agricultural University, Beijing, China; BMNH - Natural History Museum, London, Great Britain; BNHM - Beijing Natural History Museum, Beijing, China; CMNH - Carnegie Museum of Natural History, Pittsburgh, USA; DEIC - Deutsche Entomologisches Institut, Eberswalde, Germany; HNHM - Hungarian Natural History Museum, Budapest, Hungary; IMTU - Inner Mongolia Teachers University, Department of Biology, Huhehot, Inner Mongolia, China; ISNB - Institut Royal des Sciences Naturelles de Belgique, Brussels, Belgium; IZAS - Institute of Zoology, Academia Sinica, Beijing, China; JEC - J.E. EGER collection, Tampa, Florida, USA; JM - J.M. TUSÁK collection, Prague, Czech Republic; LSUK - Linnean Society, London, Great Britain; MCSN - Museo Civico di Storia Naturale Giacomo Doria, Genova, Italy; MCZC - Museum of Comparative Zoology, Harvard University, Cambridge, Massa- chusetts, USA; MDLIB - Museo do Dundo, Laboratório de Investigações Biológicas, Dundo, Angola; MGAB - Muzeul de Istoria Naturala Grigore Antipa, Bucharest, Romania; MNHN - Muséum National dHistoire Naturelle, Paris, France; MNMS - Museo Nacional de Ciencas Naturales, Madrid, Spain; MRAC - Musée Royal de lAfrique Centrale, Tervuren, Belgium; MRSN - Museo Regionale di Scienze Naturali, Torino, Italy; MSNT - Museo Civico di Storia Naturale, Trieste, Italy; MZBS - Museo de Zoologia, Barcelona, Spain; MZHF - Zoological Museum, University of Helsinki, Finland; MZLU - Museum of Zoology, Lund University, Lund, Sweden; MZUF - Museo Zoologica, Università di Firenze, Florence, Italy; NHMB - Naturhistorisches Museum Basel, Switzerland; NHMW - Naturhistorisches Museum Wien, Vienna, Austria; NHMUK - Natural History Museum, Karachi University, Pakistan; NHRS - Naturhistoriska Riksmuseet, Stockholm, Sweden; NKUM - Nankai University, Department of Biology, Tianjin, China; NMV - National Museum of Victoria, Melbourne, Australia; NSMT - National Science Museum, Tokyo, Japan; NTM - Museums and Art Galleries of the Northern Territory, Darwin, Australia; NZAC - New Zealand Arthropod Collection, Auckland, New Zealand; OXUM - Oxford University Museum, Hope Entomological Collections, Oxford, Great Britain; QMB - Queensland Museum, Brisbane, Australia; 168 JERZY A. LIS REL - R.E. LINNAVUORI collection, Raisio, Finland (belongs to AMNH); RNMH - Nationaal Natuurhistorisch Museum, Leiden, the Netherlands; SAM - South Australian Museum, Adelaide, Australia; SMFD - Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt am Main, Germany; SMNS - Staatliches Museum für Naturkunde, Stuttgart, Germany; SMTD - Staatliches Museum für Tierkunde, Dresden, Germany; TMP - Transvaal Museum, Pretoria, South Africa; UMCA - Universidad de Madrid, Cátedra de Arthrópodos, Spain; UMIZ - Universität Mainz, Institut für Zoologie, Germany; UODZ - University of Opole, Department of Zoology, Poland; UQIC - Queensland University Insects Collection, Brisbane, Australia; USNM - United States National Museum of Natural History, Washington, USA; UWNM - Wroc³aw University, Natural History Museum, Wroc³aw, Poland; UZMT - Zoological Museum, University of Turku, Finland; ZIUN - Museum of Zoology, University of Naples, Italy; ZMAN - Zoölogisch Museum, University of Amsterdam, the Netherlands; ZMAS - Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia; ZMHB - Zoologisches Museum, Humboldt Universität, Berlin, Germany; ZMMU - Zoological Museum, Moscow University, Moscow, Russia; ZMPA - Zoological Institute, Polish Academy of Sciences, Warsaw, Poland; ZMUC - Zoologisk Museum, University of Copenhagen, Denmark; ZMUH - Zoologisches Institut und Museum, Universität Hamburg, Germany; ZSMC - Zoologische Staatssammlung München, Germany. Acknowledgements. I would like to express my sincere thanks to Dr. Masaaki TOMOKUNI (Tokyo, Japan) for his help is resolving the problem of the authorship of Neostibaropus formosanus, Dr. Dominique PLUOT-SIGWALT (Paris, France) and Dr. Tamás VÁSÁRHELYI (Budapest, Hungary) for their hospitality and help during my visits to Paris and Budapest respectively, Dr. LE-YI ZHENG (Tianjin, China) for information on the distibution of some Chinese Cydnidae, Dr. Andrej GOGALA (Ljubljana, Slovenia) for his help in verification of records
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