Research and Reviews in , 54 (3): 135-205 (1994) Published by A.P.E. © 1994 Asociaci6n de Parasit61ogos Espafioles (A.P.E) Printed in Barcelona, Spain

NOMENCLATURE OFTHE

Y.J. GOLVAN Parasitologie, Pathologie et Epidemiologic Parasitaires, U.R.A. 1344 du C.N.R.S., Faculte de Medecine Saint Antolne, rue Chaligny 27, 75571 Paris Cedex 12,

Received 7 April 1993; accepted 20 April 1994

REFERENCE:GOLVAN(Y.1.), 1994.- Nomenclature of the Acanthocephala. Research and Reviews in Parasitology, 54 (3): 135-205. SUMMARYTh: e whole Acanthocephala is reviewed from the point of view of the nomenclature. All the genera and described so far are alphabetically listed and their validity established and/or discussed. Synonyms and geographical distribution are noted for each species. All the references concerning these genera and species are included. The following new taxa are proposed: Arhythmorhynchus turbidus (Van Cleave, 1937) novocomb. (=Corynosoma turbidum Van Cleave, 1937); Arhythmorhynchus villoti nom. novo(= longicollis Villot, 1875); Cen- trorhynchus hartwichi nom. nov. (=Centrorhynchus conspectus sensu Hartwich, 1956); Centrorhynchus latai nom. novo[=Centrorhynchus indicus (Gupta et Lata, 1966) Gupta et Fatma, 1983]; Cleaveius inglisi (Gupta et Fatma, 1987) novocomb. (=Mehrarhynchus inglisi Gupta et Fatma, 1987); Cleaveius singhai (Gupta et Fatma, 1987) novocomb. (=Mehrarhynchus singhai Gupta et Fatma, 1987); Echinorhynchus (Metechinorhynchus) muraenolepisi (Rodyuk, 1984) novocomb. t=Metechinorhynchus muraenolepisi Rodyuk, 1984); dauai nom. novo(=Neoechi- norhynchus rutili sensu Datta, 1936 nec O.F. MUller, 1780); Neoechinorhynchus edmondsi nom. novo(=Neoechinorhynchus rutili sensu Edmonds, 1982); Neoechinorhynchus hartwichi nom. novo (=Neoechinorhynchus australis sensu Hartwich, 1956); Nephridiacanthus thapari (Sen et Chauhan, 1972) novocomb. (=Nephridiorhynchus thapari Sen et Chauhan, 1972); Plagiorhynchus pupa (von Linstow, 1905) novocomb. (=Echi- norhynchus pupa von Linstow, 1905); Porrorchis brevicanthus (Das, 1949) nov. comb. [=Centrorhynchus brevicanthus Das, (1949) 1950]; Profili- collis chasmagnathi (Holcman-Spector, Mane-Garzon et Dei-Cas, 1977) novocomb. [=Falsifilicollis chasmagnathi Holcman-Spector, Mane-Gar- zon et Dei-Cas, 1977; =Polymorphus (Profilicollis) chasmagnathi (Holcman-Spector, Mane-Garzon et Dei-Cas, 1977) Amin, 1985]; Prosthorhynchus asturi (Gupta et Lata, 1967) novocomb. (=Rhadinorhynchus asturi Gupta et Lata, 1967); Prosthorhynchus bullocki (Schmidt et Kuntz, 1966) novocomb. (=Plagiorhynchus bullocki Schmidt et Kuntz, 1966); Prosthorhynchus golvani (Schmidt et Kuntz, 1966) novocomb. i=Plagiorhynchus golvani Schmidt et Kuntz, 1966); Prosthorhynchus ogatai (Fukui et Morisita, 1936) nov. comb. [=Porrorchis ogatai Fukui et Morisita, 1936; =Plagiorhynchus (Prosthorhynchus) ogatai (Fukui et Morisita, 1936) Schmidt et Kuntz, 1966]; Prosthorhynchus russelli (Tadros, 1970) novocomb. (=Plagiorhynchus russelli Tadros, 1970); Prosthorhynchus schmidti nom. novo(=Prosthorhynchus rectus Sprehn, 1942 nee Lin- ton, 1892); Prosthorhynchus taiwanensis (Schmidt et Kuntz, 1966) novocomb. (=Plagiorhynchus taiwanensis Schmidt et Kuntz, 1966); Southwelli- na ardeae (Belopolskaja, 1958) novocomb. i=Polymorphus ardeae Belopolskaja, 1958); Southwellina pontica (Petrotschenko et Smogorjevskaja, 1962) nov. comb. (=Hemiechinosoma ponticum Petrotschenko et Smogorjevskaja, 1962); Sphaerirostris dollfusi nom. nov. (=Centrorhynchus picae sensu Dollfus, 1953); Sphaerirostris globuli (Nama et Rathore, 1984) nov comb. (=Centrorhynchus globuli Nama et Rathore, 1984); Sphaeri- rostris lesiniformis novocomb. [=Centrorhynchus lesiniformis (Molin, 1859) Meyer, 1932 (nec Bock, 1935)]; Sphaerirostris robustus (Datta, 1928) novocomb. (=Echinorhynchus robustus Datta, 1928); Sphaerirostris saxicoloides (Nama et Rathore, 1984) novocomb. (=Centrorhynchus saxicoloi- des Nama et Rathore, 1984); Sphaerirostris splendi (Gupta et Gupta, 1971) nov. comb. (=Centrorhynchus splendi Gupta et Gupta, 1971).

KEy WORDS:Acanthocephala, nomenclature, genera, species, synonyms, geographical distribution, review.

V.I. PETROTSCHENKO (1956 and 1958), Satyu YAMAGU- Nomina enim si pereunt, perit et rerum cognitio (Systema TI (1961), and more recently, Omar AMIN (1985), who Naturae, Xth edition). Filum ariadneum est Systema, sine quo Chaos est Historia presented a new classification of the Acanthocephala in naturalis (Philos. Botan.). the Biology of the Acanthocephala, which also contains a Carl von LINNE comprehensive index to information about the species mentioned in the book. I began to study the Acanthocephala in 1954. During INTRODUCTION the past 40 years, my only ambition was to make my own contribution to a very collective work. In this body The Linnean nomenclature of the Acanthocephala is of information and in the many taxonomic papers in more than 200 years old, So it should be no wonder that which the species have been described, many errors of many mistakes have been made during all these years. nomenclature were undoubtedly made. The present work Systematics is a very difficult science but it remains the is an attempt to correct errors for the benefit of Helmin- base for biological progress. In the field of the modern thology in the future. classification of the acanthocephalan , after the great precursors, such as Max LUHE (1911) and Anton MEYER (1932/1933), the status of a separate Phylum was GENERA AND SPECIES clearly established by the works of Harley J. VAN CLEA- VE (1948), who did much to promote the system of clas- In the following, the whole Phylum Acanthocephala is sification which is used today by the zoologists who reviewed from the point of view of the nomenclature. All study these parasites. the genera and species described so far are alphabetically After V AN CLEAVE, the work was carried on by many listed and their validity established and/or discussed. helminthologists. Especially noteworthy are the works of Synonyms and geographical distribution are noted for 136 Y.J.GOLVAN each species. Those considered valid taxa are noted in -A. aeutulus Van Cleave, 1931 (, USA) bold. All the references concerning these genera and spe- -A. (Termitorhynehus) aenigma Reichensperger, 1922: cies are included at the end. may be an Oligaeanthorhynehus sensu lato (fide This list takes into account the decision of the Interna- MEYER, 1932) tional Commission for Zoological Nomenclature (Lon- -A. alabamensis Amin et Williams, 1983: syn. of Aean- don, 1962) that the name of a which ends with thoeephalus etowani Williams, 1974 «rhynchus» is masculine and so the specific name which -A. amurensis Kostylew, 1941: syn. of Paraeantho- follows must also be masculine. eephalus eurtus (A. et D.-A.) Yamaguti, 1963 (nee amuriensis) ACANTHOCEPHALOIDES Meyer, 1932 -A. anguillae (O.F. Muller, 1780) Liihe, 1911 (type spe- -A. ehabanaudi Dollfus, 1951 (nee ehabaunadi, nee cies) (all Europe) ehabaudi): syn. of Paraeanthoeephaloides ehabanau- -A. anguillae Belous, 1952: syn. of Paraeanthoeephalus di (D.) Golvan, 1969 eurtus Achmerov et Dombrowskaja-Achmerova, 1941 -A. claviformis Araki et Machida, 1987 (Japan) -A. anthuris (Dujardin, 1845) Luhe, 1911 (West Euro- -A. eyrusi Bray, Spencer Jones et Lewis, 1988 (St. Lucia pe) Lake, Natal) -A. artatus Van Cleave, 1925 (nee aratus) [prov. of Shi- -A. distinetus Golvan, 1969 (Senegal) nano, Japan, also found in India (doubtful)]: may be a -A. genetieus de Buron, Renaud et Euzet, 1986 (Medite- syn. of Aeanthoeephalus lucidus (fide HARADA,1935) rranean -shore of France) -A. balkanieus Batcharov et Combes, 1974 () -A. iehiharai Araki et Machida, 1987 (North of Japan) -A. bigueti Houin, Golvan et Brygoo, 1965: syn. of -A. inerassatus (Molin, 1858) Meyer, 1932 (Adriatic Pseudoaeanthoeephalus bigueti (H., G. et B.) Golvan, and ) 1969 -A.japonieus (Fukui et Morisita, 1936) Yamaguti, 1939: -A. breviprostatus Kennedy, 1982 (East Asia) syn. of Aeanthoeephalus japonieus (F. et M.) Petrots- -A. bufonieola Kostylew, 1941: syn. of Pseudoaeantho- chenko, 1956 eephalus bufonieola (K.) Petrotschenko, 1956 -A. kostylewi Meyer, 1932 (Black Sea) -A. bufonis (Shipley, 1903) Southwell et Macfie, 1925: -A. neobythitis Yamaguti, 1939: syn. of Yamagutisentis syn. of Pseudoaeanthoeephalus bufonis (S.)Petrots- neobythitis (Y.) Golvan, 1969 chenko, 1956 -A. propinquus (Dujardin, 1845) Meyer, 1933 (type -A. bufonis sensu Southwell et Macfie, 1925 is another species) (Atlantic Ocean, ) (nee species (vide KENNEDY,1982) propinguus) -A. bufonis sensu Petrotschenko, 1953: syn. of Aeantho- -A. propinquus sensu Golvan, 1956: syn. of Aeantho- eephalus breviprostatus Kennedy, 1982 eephaloides distinetus Golvan, 1969 -A. caucasicus Petrotschenko, 1953: syn. of Pseudoa- -A. rhinoplagusiae Yamaguti, 1935: syn. of Yamaguti- eanthoeephalus eaueasieus (P.) Petrotschenko, 1956 sentis rhinoplagusiae (Y.) Golvan, 1969 (nee rhyno- -A. clavula (Dujardin, 1845) Grabda-Kazubska et plagusiae) Chubb, 1968 (nee Hamann, 1892) (all the south wes- -A. soleae (Porta, 1905) Meyer, 1932 in Petrotschenko, tern Europe) 1956: syn. of Solearhynehus soleae (P.) de Buron et -A. eorrealimai Machado Filho, 1970 (Parana, ) Maillard, 1985 -A. eriniae Snow, 1970 (Australia) -A. eurtus (Achmerov et Dombrowskaja-Achmerova, ACANTHOCEPHALORHYNCHOIDES Kostylew, 1941) Yamaguti, 1963 (eastern Siberia) 1941: may be a syn. of Pallisentis Van Cleave, 1928 -A. dirus (Van Cleave, 1931) Van Cleave et Townsend, -A. eholodkowskyi (Kostylew, 1928) Williams, Gibson 1936 (Mississippi, Illinois, USA) et Sadighian, 1980 (Armenia, ) -A. domerguei Golvan, Brygoo et Gassmann, 1972 -A. intermedius (Achmerov et Dombrowskaja-Achme- (Madagascar) rova, 1941) Petrotschenko, 1956: syn. of Aeantho- -A. eehigoensis Fujita, 1920 (Japan) (fide YAMAGUTI, eephalorhynehoides ussuriensis Kostylew, 1941 1963, the species Aeanthoeephalus aeerbus, Aeantho- -A. ussuriensis Kostylew, 1941 (type species) (Basin of eephalus aeuleatus and Aeanthoeephalus oneorhyn- the Amur River) chi are synonyms of it) -A. elongatus Van Cleave, 1937 (Hunan, ) ACANTHOCEPHALUS Koelreuther, 1771 -A. etowani Williams, 1974: may be a syn. of Aeantho- -A. aeerbus Van Cleave, 1931: (prov. of Shinano, eephalus alabamensis Amin et Williams, 1983 (fide Japan): may be a syn. of Aeanthoeephalus eehigoensis AMIN & WILLIAMS,1983) Fujita, 1920 (fide HARADA,1935) -A. faleatus (Frolich, 1789) Ltihe, 1911 (mountains of -A. aeuleatus Van Cleave, 1931 (pro v . of Shinano, the southern Europe, Alps, Carpathian mountains) Japan): may be a syn. of Aeanthoeephalus eehigoensis (vide GRABDA-KAZUBSKA1962), Fujita, 1920 (fide HARADA,1935) -A. fluviatilis Paperna, 1964 (nee fluviatilus) (Mount -A. aeutispinis Machado Filho, 1968 (nee aeutispinus) Carmel Israel) (Rio de Janeiro, Brazil) -A. galaxii Hine, 1978 (New Zealand) Nomenclature of the Acanthocephala 137 -A. goaensis Jain et Gupta, 1981 (India) -A. ranae sensu Van Cleave, 1915 (USA) is another spe- -A. gotoi Van Cleave, 1925 (market of Tokyo, Biwa and cies Suwa Lakes, Japan) -A. rauschi Golvan, 1969 (Alaska) (nee Schmidt) -A. graeilaeanthus Meyer, 1932 (East of , -A. serendibensis Crusz et Mills, 1970 (Sri Lanka) Poland): may be a syn. of Paracanthocephalus graci- -A. sinensis Van Cleave, 1937 (Hunan, Chikiang and lacanthus (M.) Grabda et Grabda-Kazubska, 1967 Hupei, China and Sulawesi) (a good species fide -A. haranti Golvan et Oliver in Golvan, 1969 (Fontes- YAMAGUTI,1963 nee 1954) tramer in the reappearance of a subterranean river on -A. srilankensis Crusz et Ching, 1975 (Sri Lanka) the sea shore of Languedoc, France) -A. tahlequahensis Oetinger et Buckner, 1976 (Oklaho- -A. hastae Baylis, 1944 (south of Australia) ma, USA) -A. inerassatus (Molin, 1858) Kostylew, 1916: syn. of -A. tenuirostris (Achmerov et Dombrowskaja-Achme- Acanthocephaloides incrassatus (M.) Meyer, 1932 rova, 1941) Yamaguti, 1963 (north east of Siberia) -A. jaeksoni Bullock, 1962: syn. of Acanthocephalus -A. tigrinae (Shipley, 1903) Yamaguti, 1963 (Thailand) dirus (VC.)Van Cleave et Townsend, 1936 -A. tumescens (von Linstow, 1896) Porta, 1905 (Tierra -A. japonicus (Fukui et Morisita, 1936) Petrotschenko, del Fuego) (vide HARTWICH,1956) 1956 (Japan) -A. vancleavei Hughes et Moore, 1943: syn. of Fessi- -A. kabulensis Datta et Soota, 1954 (Afghanistan) sentis vancleavei (H. et M.) Nickol, 1972 -A. kashmirensis Datta, 1936 (Laddak, north ofIndia) ACANTHODELTA Diaz-Ungrfa et Gracia-Rodrigo, -A. lateralis (Leidy, 1851) Petrotschenko, 1956: syn. of 1958 (nee Aeanthrodelta) Echinorhynchus (Metechinorhynchus) lateralis -A. scorzai (Dfaz-Ungrfa et Gracia-Rodrigo, 1957) Leidy, 1851 Diaz-Ungrfa et Gracia-Rodrigo, 1958 (type species) -A. lavareti Koelreuther, 1771: syn. of Pomphorhyn- (Orinoco river, ) (nee scorziai: may be a ch us laevis (Z. in O.F. M.) Van Cleave, 1924 (??) (vide DiAZ-UNGRIA& FARO- -A. lizus Li-Minmin, 1984 (Japan) OQI, 1989) -A. lucidus Van Cleave, 1925 [prov. of Shimosa, Japan, found also in India and Indochina (doubtful)]: may be a ACANTHOGYRUS Thapar, 1927 syn. of Acanthocephalus artatus (fide YAMAGUTI, -A. acanthogyrus Thapar, 1927 (type species) (India) 1939) (vide FAROOQI,1989) -A. lucii (O.F. Miiller, 1776) Liihe, 1911 (all Europe) -A. aeanthuri (Cable et Quick, 1954) Dollfus et Golvan, -A. lucii sensu Meyer, 1931: the arctic species is diffe- 1956: vide Acanthosentis rent from Acanthocephalus lucii (O.F.M.) Liihe, 1911 -A. anguillae (Wang, 1981) Amin, 1985: vide Acantho- sensu stricto, parasite of fresh water of Europe sentis (fide YAMAGUTI,1963) -A. antspinus (sic) (Verma et Datta, 1929) DoJlfus et -A. lutzi (Hamann, 1891) Meyer, 1932 (nee von Lins- Golvan, 1956 (nee antspinis) (India, Burma): vide tow, 1896) (vide HARTWICH, 1956) (Brazil): is not an Acanthosentis Acanthocephalus (fide GOLvAN, 1960 and 1969) -A. arii (Bilqees, 1971) Amin, 1985: vide Acanthosentis -A. madagascariensis Golvan, 1965 (Madagascar) -A. baeailai (Verma, 1973) Amin, 1985: vide Acantho- -A. minor Yamaguti, 1935 (Hukui, Hokkaido, Japan) sentis -A. nanus Van Cleave, 1925 (Kyoto, prov. of Omi, -A. eameroni (Gupta et Kajaji, 1969) Amin, 1985: vide Japan) Acanthosentis -A. oncorhynchi Fujita, 1921: may be a syn. of Acan- -A. cholodkowskyi (Kostylew, 1928) Golvan, 1959: thocephalus echigoensis Fujita, 1920 (fide YAMAGUTI, may be an Acanthocephalorhynchoides (fide KOSTY- 1963) LEW,1941) -A. opsariichthydis Yamaguti, 1935 (Suwa Lake, Naga- -A. dattai (Poddar, 1938) Dollfus et Golvan, 1956: vide no, Japan) (nee opsaliehtydis, nee opsaliehthydis) (vide Acanthosentis YAMAGUTI,1939: page 321) -A. giuris (Soota et Sen, 1956) Amin, 1985: vide Acan- -A. parallelotestis Achmerov et Dombrowskaja-Achme- thosentis rova, 1941 (Eastern Siberia) -A. golvani (Gupta et Jain, 1980) Amin, 1985: vide -A. parksidei Amin, 1975: syn. of Acanthocephalus Acanthosentis dirus (VC.) Van Cleave et Townsend, 1936 -A. guptai Gupta et Verma, (1976) 1977 (India): may be -A. paronai (Condorelli, 1897) Meyer, 1932 (): a syn. of A. acanthogyrus (fide FAROOQI,1989) may be a syn. of Acanthocephalus anguillae (OFM.) -A. holospinus (Sen, 1938) Dollfus et Golvan, 1956: Liihe, 1911 (fide GOLV AN, 1960) vide Acanthosentis -A. pesteri Tadros, 1966 (unknown geographical origin) -A. indicus (Tripathi, 1959) Chubb, 1982 (India) -A. propinquus sensu Kostylew, 1926: syn. of Acantho- -A. intermedius (Achmerov et Dombrowskaja-Achme- cephaloides kostylewi Meyer, 1931 rova, 1941) Dollfus et Golvan, 1956: vide Acantho- -A. ranae (Schrank, 1788) Liihe, 1911 [all Europe, but sentis also found in Maryland USA(doubtful)] (vide V AN -A. maroeeanus (Dollfus, 1951) Dollfus et Golvan, CLEAVE,1915) 1956: vide Acanthosentis 138 Y.J.GOLvAN -A. multispinus Wang, 1966: vide Acanthosentis -A. golvani Gupta et Jain, 1980 (Chandigarh, India) -A. nigeriensis Dollfus et Golvan, 1956: vide Acantho- -A. heterospinus Aly Khan et Bilqees, 1990 (Kalri sentis Lake, Sind-Pakistan) -A. oligospinus (Anantaraman, 1980) Amin, 1985: vide -A. hilsai Pal, 1963 (Uttar Pradesh, India): may be a syn. Acanthosentis of A. indicus Tripathi, 1959 (fide GUPTA&JAIN, 1980) -A. papilio Troncy et Vassiliades, 1974: vide Acantho- -A. holospinus Sen, 1938 (Bengal, India) sentis -A. indica (sic) Tripathi, 1959 (Bihar, Chilka Lake, -A. partispinus Furtado, 1963: vide Acanthosentis Matla and Ganga Rivers, India) -A. paucispinus Wang, 1966: vide Acanthosentis -A. intermedius (Achmerov et Dombrowskaja-Achme- -A. periophthalmi (Wang, 1980) Amin, 1985: vide rova, 1941) Golvan, 1959: may be a syn. of Acantho- Acanthosentis cephalorhynchoides intermedius (A. et D.-A.) (fide -A. putitorae Chowan, Gupta et Khera, 1988: vide PETROTSCHENKO1956, ) Acanthosentis -A. lizae Wang, 1986 (China, Fujian) -A. seomberomori (Wang, 1980) Amin, 1985: vide -A. maroccanus Dollfus, 1951 () Acanthosentis -A. multispinus Wang, 1966 (China) -A. shashiensis (Tso, Chen, Chien, Wu et Deng, 1974) -A. nigeriensis Dollfus et Golvan, 1956 (Basin of the Amin, 1985: vide Acanthosentis Niger River) -A. siamensis Farooqi et Sirikanchana, 1987: vide -A. oligospinus Anantaraman, 1980 (nee 1969) (India) Acanthosentis (vide GUPTA & JAIN, 1980): may be a syn. of A. antes- -A. similis (Wang, 1980) Amin, 1985: vide Acantho- pinus (fide CHOWAN,GUPTA& KHERA, 1987) sentis -A. papilio Troncy et Vassiliades, 1974 (West ) -A. sireari (Poddar, 1941) Dollfus et Golvan, 1956: vide -A. partispinus Furtado, 1963 (Malaysia) Acanthosentis -A. paucispinus Wang Pu Qin, 1966 (China) -A. thapari (Prasad, Sahay et Shambhunath, 1969) -A. periophthalmi Wang Pu Qin, 1980 (Fujian, China) Amin, 1985: vide Acanthosentis -A. putitorae Chowan, Gupta et Khera, 1988 (north of -A. tilapiae (Baylis, 1947) Dollfus et Golvan, 1956: vide India) Acanthosentis -A. scomberomori Wang Pu Qin, 1980 (Fujian, China) -A. tripathii Rai, 1967 (India) (nee tripathi): may be a -A. seenghalae Chowan, Gupta et Khera, 1988 (north of syn. of A. acanthogyrus (fide FAROOQI,1989) India) -A. vittatusi (Verma, 1973) Amin, 1985: vide Acantho- -A. shashiensis Tso, Chen, Chien, Wu et Deng, 1974 sentis (China) -A. siamensis Farooqi et Sirikanchana, 1987 (Thailand) ACANTHORHYNCHOIDES Achmerov, 1959: syn. of -A. similis Wang Pu Qin, 1980 (Fujian, China) Acanthocephalorhynchoides Kostylew, 1941 -A. sircari Podder, 1941 (India) -A. thapari Prasad, Sahay et Nath, 1969 (India) ACANTHOSENTHIS: errat. pro Acanthosentis -A. tilapiae Baylis, 1947 (Tanzania, , ) (nee 1958) ACANTHOSENTIS Verma et Datta, 1929: may be -A. vancleavei Gupta et Fatma, 1986 (India) admitted as a valid genus and not as a subgenus of -A. vittatusi Verma, 1973 (India) Acanthogyrus -A. acanthuri Cable et Quick, 1954 (Puerto Rico, Toba- Aeanthrus sipuneuloides Acharius, 1780, invalidated: go) syn. of Echinorhynchus salmonis O.F. Miiller, 1784 -A. anguillae Wang, 1981 (China) -A. antespinus Verma et Datta, 1929 (market of Calcut- ALLORHADINORHYNCHUS Yamaguti, 1959: the ta) (the original spelling "antspinus" is wrong and was synonymy of this genus with Micracanthorhynchina corrected by MEYER, 1932) (nee antespinis) Strand, 1936, proposed by ARAKI & MACHIDA(1987) -A. arii Bilqees, 1971 (Pakistan) cannot be admitted -A. bacailai Verma, 1973 (India) -A. segmentatus Yamaguti, 1959 (type species) (Japa- -A. betwai Tripathi, 1959 (nee 1956)(Bhopal, India): nese Inland Sea) valid species (fide GUPTA& JAIN, 1980 and ALY KHAN & BrLQEES,1990) ANDRACANTHA Schmidt, 1975 -A. bilaspurensis Chowan, Gupta et Khera, 1987 (India) -A. baylisi (Zdzitowiecki, 1986) Zdzitowiecki, 1989 -A. eameroni Amin, 1985: errat. pro A. commersoni (South Georgia, Antarctic) Gupta et Kajaji, 1969 -A. clavata (Goss, 1940) Zdzitowiecki, 1989 (Australia, -A. commersoni Gupta et Kajaji, 1969 (nee eammer- Antarctic) soni) (India) -A. gravida (Alegret, 1941) Schmidt, 1975 (types spe- -A. dattai Podder, 1938 (nee 1933) (Bengal, India) cies) (Cuba, Florida, , Gulf of Mexico) -A. giuris Soota et Sen, 1956 (India) -A. mergi (Lundstrom, 1941) Schmidt, 1975 (Scandina- -A. gobindi Chowan, Gupta et Khera, 1987 (India) via, found also from Alaska to , USA, Nomenclature of the Acanthocephala 139 but it is probably another species because the speci- -A. frontospinosus (Tubangui, 1935) Yamaguti, 1963 mens are very different) (Philippine Islands) -A. phalacrocoracis (Yamaguti, 1939) Schmidt, -A. fuseus Harada, 1929: syn. of Southwellina hispida 1975 (Takusima, Shikoku, Japan; east of Siberia; rvc» Witenberg, 1932 Alaska) -A. hispidus Van Cleave, 1925: syn. of Southwellina -A. tunitae (Weiss, 1914) Zdzitowiecki, 1989 [Tunisia, hispida (Ve.) Witenberg, 1932 found also in Alaska (doubtful)] -A. invaginabilis (von Linstow, 1902) Liihe, 1912 (West of Eurasia) (nee invaginavilis): may be a syn. of Shipley, 1899 Arhythmorhynchus longicollis (V.) (fide GOLVAN, -A. aculeatus Meyer, 1931 (Brazil) (nee aeuleatum) 1956) or of Arhythmorhynchus anser Florescu, 1941 -A. amphistomi Byrd and Denton, 1949 (Nearctic (fide KHOKHLOVA,1975) region) -A. invaginabilis sueeieus Lundstrorn, 1942: vide -A. bivolucrus Das, 1952 (India) (nee bivoluerus) (fide Arhythmorhynchus suecicus L., 1942 YAMAGUTI,1963 it is a Strigeid Trematode) -A.jeffreyi Schmidt, 1973 (Nearctic region) -A. chauhani Sen, 1975 (India) -A.johnstoni Golvan, 1960 (Queensland, Australia) -A. hemignathi (Shipley, 1896) Shipley, 1899 (type spe- -A. limosae Edmonds, 1971 (Queensland, Australia) cies) (Hawai) (nee 1897) -A. longicollis (Villot, 1875) Luhe, 1912: may be a syn. -A. paulonucleatus Khokhlova et Tsimbalyuk, 1966 (or of Arhythmorhynchus anser Florescu, 1941 (fide GOL- Cimbaluk) (nee 1971) (west of Siberia) VAN, 1956) or of Arhythmorhynchus invaginabilis -A. silesiacus Okulewicz et Maruszewski, 1980 (Poland) (vL.)Luhe, 1912 ifide KHOKHLOVA,1975 and in this case A. invaginabilis becomes the syn. of A. longico- ARHYNCHUS Shipley, 1896: preoccupied, syn. of Apo- llis because posteriorly proposed (law of priority)] rorhynchus Shipley, 1899 -A. macracanthus H.L. Ward et Winter, 1952 (Califor- -A. hemignathi Shipley, 1896: syn. of Apororhynchus nia) (may be a Southwellina) hemignathi (S.) Shipley, 1899 -A. maerourus (Bremser in Westrumb, 1821): may be a ARHYTHMACANTHUS Yamaguti, 1935 (nee Aryth- syn. of Arhythmorhynchus frassoni (M.) Luhe, 1911 maeanthus): the synonymy with Heterosentis Van Cle- (fide MEYER, 1932 it is a species inquirenda; fide ave, 1931, proposed by AMIN (1985) is not admitted KHOKHLOVA,1975 it is a nomen nudum) -A.fusiformis Yamaguti, 1935 (Japan)(type species) -A. petrotschenkoi (Schmidt, 1969) Atrashkevi tch, 1979 -A. overstreeti Schmidt et Paperna, 1978 (Elat, Israel) (SSSR) (nee petrosehenkoi) -A. paraplagusiarum Nickol, 1972 [Queensland, Aus- -A. plicatus (von Linstow, 1883) Meyer, 1932 (Central tralia, also found in India (doubtful)] Asia) -A. plotosi (Yamaguti, 1935) Schmidt et Paperna, 1978 -A. pumilirostris Van Cleave, 1916 (nee plumirostris, -A. septacanthus Sita, 1949 (India, China) nee pumiliorostris) (Washington D.e., USA, Nearctic -A. thapari Gupta et Fatma,(1979) 1980 (Tamil Nadu, region) India) -A. roseus (Molin, 1858) Meyer, 1932: valid species (fide KHOKHLOVA,1975), syn. of Arhythmorhynchus ARHYTHMORHYNCHUS Luhe, 1911 longicollis (V.) or Arhythmorhynchusfrassoni (M.) -A. anser Florescu, 1941 (Roumania): may be a syn. of Lilhe, 1911(fide GOLVAN, 1956 and PETROTSCHENKO, Arhythmorhynchus invaginabilis (vL.)Luhe, 1912 1958) (Holarctic) (fide GOLVAN,1956 and KHOKHLOVA,1975) -A. rubicundus (Molin, 1859) Meyer, 1932 (South -A. brevis Van Cleave, 1916 (Baltimore, USA, Nearctic Europe): good species (fide KHOKHLOVA, 1975) or region) juvenile of Arhythmorhynchus frassoni (M.) Luhe, -A. capellae (Yamaguti, 1935) Van Cleave et Rausch, 1911 (fide GOLVAN,1956) 1950 (Japan, also found in Colorado, USA) -A. sachalinensis Krotov et Petrotschenko, 1958 (East -A. eapellae sensu Schmidt, 1963: syn. of Arhyth- of Siberia): syn. of Arhythmorhynchus teres Van Cle- morhynchus jeffreyi Schmidt, 1973 ave, 1920 (fide KHOKHLOVA,1975) -A. comptus Van Cleave et Rausch, 1950 (Alaska, East -A. siluricola Dollfus, 1929: it is a Echinorhynchus of Siberia) sensu lata (fide GOLVAN,1969) -A. distinctus Baer, 1956 (Greenland) -A. suecicus Lundstrom, 1942 (Scandinavia) -A. duocinctus Chandler, 1935 (Texas, USA): may be a -A. teres Van Cleave, 1921 (Chili) syn. of Southwellina hispida (Ve.) Witenberg, 1932 -A. tigrinus Moghe et Das, 1953 (India) -A. eroliae (Yamaguti, 1939) Van Cleave et Rausch, -A. trichocephalus (Leuckart, 1876 in Kaiser, 1893) 1950 (Sizuoka, Japan, East of Siberia) (nee Schmidt, Liihe, 1911 (Florida, USA) 1973) -A. tringi Gubanov, 1952 (SSSR) -A. frassoni (Molin, 1858) Luhe, 1911 (nee 1958) (type -A. turbidus (Van Cleave, 1937) novo comb.: syn. of species) (Eurasia) Corynosoma turbidum Van Cleave, 1937 -A.frassoni sensu Johnston et Edmonds, 1951: syn. of -A. uncinatus (Leuckart, 1876 in Kaiser, 1893) Luhe, Arhythmorhynchusjohnstoni Golvan, 1960 1912 (USA, Florida) 140 Y.J.GOLvAN -A. villoti nom. nov.: vide Echinorhynchus longicollis -B. sajori Belous, 1952: syn. of Micracanthorhynchina Villot, 1875 sajori (B.) Golvan, 1969 -A. xeni Atrashkevitch, 1978 (SSSR) BOLBOSOMA Porta, 1908 ASPERSENTIS Van Cleave, 1929 -8. aurantiacum (Risso, 1826) Porta, 1909: may be a -A. austrinus Van Cleave, 1929 (Southern Shetlands, syn. of Bolbosoma vasculosum (R.)Meyer, 1932 Antarctic) -8. australis Skrjabin, 1972 (Antarctic) -A. heteracanthus (von Linstow, 1896) Golvan, 1969: -8. balaenae (Gmelin, 1790) Porta, 1908 (type species) syn. of Heterosentis heteracanthus (vL.) Van Cleave, (vide VAN CLEAVE, 1953) 1931 -B. balanae Van Cleave, 1953: errat. (?) pro 8. balae- -A.johni (Baylis, 1929) Chandler, 1934 (Falklands) nae (G.) Meyer, 1932 -A. megarhynchus (von Linstow, 1892) Golvan, 1960 -8. bobrovoi Krotov et Delyamure, 1952 (North Pacific) (type species) (South Georgia) (nec bobrovi, nee Delamure, nee Delamare) -A. wheeleri (Baylis, 1929) Chandler, 1934 (Antarctic): -B. brevicolle (Malm, 1867) Porta, 1908 (South Africa) may be a syn. of A. megarhynchus (vL.) Golvan, 1969 -8. caenoforme (Heitz, 1920) Meyer, 1932 (nec Heitz, 1917) (North Pacific): may be a Corynosoma (fide ATACTORHYNCHUS Chandler, 1935 GOLVAN,1960 and YAMAGUTT1, 963) -A. mugilis Machado Filho, 1951 (Rio de Janeiro, Bra- -8. capitatum (von Linstow, 1880) Porta, 1908 (Antarc- zil): may be a Floridosentis (fide BULLOCK,1962) tic, Cape Verde Islands) -A. verecundus Chandler, 1935 (type species) (Texas, -B. hamiltoni Baylis, 1929 (South Georgia, Antarctic) USA) -B. heteracanthe (Heitz, 1920) Meyer, 1932 ( of ATRACTORHYNCHUS in Dollfus et Golvan, 1956: errat. western Europe, found also in the Rhine River) pro Atactorhynchus Chandler, 1935 -8. nipponicum Yamaguti, 1939 (Japan,Kuril Islands) -8. physeteris Gubanov, (1952) 1955 (North Pacific) AUSTRALIFORMIS Schmidt et Edmonds, 1989 -B. porrigens (Rudolphi, 1819) Porta, 1908: syn. of Bol- -A. semoni (von Linstow, 1898) Schmidt et Edmonds, bosoma balaenae (G.) Meyer, 1932 1989 (type species) (Australia, New Guinea) -8. scomberomori Wang Pu Qin, 1980 (Fujian, China) -B. serpenticola (Fukui, 1929) Meyer, 1932: syn. of AUSTRALORHYNCHUS Lebedev, 1967 Diplospinifer serpenticola Fukui, 1929 -A. tetramorphacanthus Lebedev, 1967 (type species) -8. thunni Harada, 1935 (Japan): may be a syn. of Bol- (Tasmania, South of Australia) bosoma vasculosum (R.) Meyer, 1932 (fide PETROTS- BOLBORHYNCHOIDES Achmerov, 1959 CHENKO,1958) -8. exiguus (Achmerov et Dombrowskaja-Achmerova, -8. tuberculata Skrjabin, 1970 (South Atlantic; Indian 1941) Achmerov, 1959 (type species) (basin of Amur Ocean) River, East Siberia) -B. turbinella (Diesing, 1851) Porta, 1908 [South Geor- gia, North of the Atlantic (??)] BOLBORHYNCHUS Porta, 1906: preoccupied, syn. of -B. turbinella sensu Leiper et Atkinson, 1915 (New Zea- Bolbosoma Porta, 1908 land) is another species (fide MEYER, 1932) -B. turbinella australis Skrjabin, 1972: syn. of Bolboso- BOLBORHYNCHUS Achmerov et Dombrowskaja-Ach- ma australis Skrjabin, 1972 merova, 1941: preoccupied, syn. of Bolborhynchoides -8. vasculosum (Rudolphi, 1819) Porta, 1908 (North of Achmerov, 1959 the Atlantic, Mediterranean Sea) -B. brevicollis (Maim, 1867) Porta, 1906: syn. of Bolbo- soma brevicollis (M.) Porta, 1908 BREIZACANTHUS Golvan, 1969 -B. capitatus (von Linstow, 1880) Porta, 1906: syn. of -8. chabaudi Golvan, 1969 (type species) (North west Bolbosoma capitatum (vL.) Porta, 1908 of the Atlantic Ocean) -B. exiguus Achmerov et Dombrowskaja-Achmerova, -B. golvani Gaevskaja et Shukhgalter, 1984 (North of 1941: syn. of Bolborhynchoides exiguus (A. et D.-A.) the Atlantic Ocean) Achmerov,1959 -8. irenae Golvan, 1969 (Mediterranean Sea) -B. pellucidus (Leuckart, 1828) Porta, 1906: syn. of Bol- -8. ligur Paggi, Orecchia et Della Seta, 1975 (Ligur Sea, bosoma vasculosum (L.) Meyer, 1932 Italy) -8. porrigens (Rudolphi, 1819) Porta, 1906: syn. of Bol- bosoma balaenae (Gmelin, 1790) Meyer, 1932 BRENTISENTIS Leotta, Schmidt et Kuntz, 1982 -B. turbinella (Diesing, 1851) Porta, 1906: syn. of Bol- -8. uncinus Leotta, Schmidt et Kuntz, 1982 (type spe- bosoma turbinella (D.) Porta, 1908 cies) (Taiwan)

BOLBOSENTlS Belous, 1952: syn. of Micracan- BULLOCKRHYNCHUS Chandra, Rao et Shyamasun- thorhynchina Strand, 1936 dari, 1985 -B. hyporhamphi Belous, 1952: syn. of Micracan- -8. indicus Chandra, Rao et Shyamasundari, 1985 (type thorhynchina hemirhamphi (B.) H.L. Ward, 1951 species) (India) Nomenclature of the Acanthocephala 141

CABALLERORHYNCHUS Salgado-Maldonado, 1977 -CO (Malgacanthus) brumpti Golvan, 1965 (Madagas- -CO lamothei Salgado-Maldonado, 1977 (type species) car) (Vera Cruz, Mexico) -CO (Malgacanthus) brygooi, Golvan, 1965 (Madagas- car) CAPR/V/SENTIS van Dyk, Swanepoel et Saayman, -C. bubonis Yamaguti, 1939 [Kyushyu, Japan and India 1990 (??)] -CO hepseti van Dyk, Swanepoel et Saayman, 1990 (type -C. buckleyi Gupta et Fatma, 1981 (Uttar Pradesh, species) (Liambezi Lake, Namibia) India) -CO buteonis (Goeze, 1782 in Schrank, 1788) Kostylew, CATHAYACANTHUS Golvan, 1969 1914 [Eurasia, found also in India and in Australia -CO exilis (Van Cleave, 1928) Golvan, 1969 (type spe- (doubtful)] cies) (Pekin, China) -CO californicus Millzner, 1924 (Pacific Coast of USA) -CO chabaudi Golvan, 1958 (Western Africa) CA V/SOMA Van Cleave, 1931 -C cinctus (Rudolphi, 1819) Meyer, 193"2: syn. of -CO chromitidis Cable et Quick, 1954: syn. of Pseudoca- Sphaerirostris cinctus (R.) Golvan, 1956 visoma chromitidis (c. et Q.) Golvan et Houin, 1964 -CO clitorideus (Meyer, 1931) Golvan, 1958 (nee clitori- -CO magnum (Southwell, 1927) Van Cleave, 1931 (type deum) [Europe, Egypt, Israel, found also in India species) (Sri Lanka) (doubtful)] -CO conspectus Van Cleave et Pratt, 1940 [New York, CENTRORHYNCHUS Liihe, 1911 North Carolina, Nearctic region, found also in India -CO acanthotrias (von Linstow, 1883) Petrotschenko, (doubtful)] 1958 -C conspectus sensu Hartwich, 1956: syn. of Cen- -CO albense Rengaraju et Das, 1975 (India) (nee alben- trorhynchus hartwichi nom. novo sis) -C corvi Fukui, 1929: syn. of Sphaerirostris corvi (F.) -CO albidus Meyer, 1932 (Nicaragua, , , Golvan, 1956 [may be a syn. of Sphaerirostris skrjabi- Neotropical region) ni (P.) Golvan, 1956 (vide PETROTSCHENKO1949, )] -CO aluconis (O.F. MUller, 1780) Luhe, 1911 (type spe- -CO crocidurus Das, 1950 (Central provinces of India): cies) [Europe, found also in Taiwan (doubtful)] may be a juvenile of Centrorhynchus itatsinis Fukui, -CO amphibius Das, 1950 (Berar, India and Taiwan ?) 1929 (fide YAMAGUTI,1963) -CO andamanensis Soota et Kansal, 1972 (Andamans -CO crotophagicola Schmidt et Neiland, 1966 (Nicara- Islands, Nicobar): may be a syn. of Centrorhynchus gua) spilornae Schmidt et Kuntz, 1969 (fide ZAFAR & -C cylindraceus (Goeze, 1782 in Schrank, 1788) Kosty- FAROOQI,1981) lew, 1914: syn. of Prosthorhynchus cylindraceus (G.) -CO appendiculatus (Westrumb, 1821) Joyeux et Baer, Golvan, 1956 (may be erroneous) 1937 (nee appendieulatum) (Europe): is not a syn. of -CO dimorphocephalus (Westrumb, 1821) Meyer, 1932 Centrorhynchus aluconis (O.F.M.) Liihe, 1911 as it (South and Central Europe) was proposed by GOLVAN(1956) -CO dipsadis (von Linstow, 1888) Golvan, 1956 (Came- -C areolatus (Rudolphi, 1819) Petrotschenko, 1958: roon) syn. of Sphaerirostris areolatus (R.) Golvan, 1956 -CO elongatus Yamaguti, 1935 (Japan, Taiwan) (nee aerolatus) -C embae Cholodkowski et Kostylew, 1916: syn. of -CO asturinus (Johnston, 1912) Johnston, 1918 (Queens- Sphaerirostris embeae (c. et K.) Golvan, 1956 land, Australia) -C erraticus Chandler, 1925: syn. of Sphaerirostris -CO (Malgacanthus) atheni Gupta et Fatma, 1983 (Uttar erratic us (C.) Golvan, 1956 Pradesh, North of India) -C.falconis (Johnston et Best, 1943) Golvan, 1956 -C. bancrofti (Johnston et Best, 1943) Golvan, 1958 (Central Australia, Taiwan, Palawan) (Queensland, Australia) -C faleonis Das, (1949) 1950, preoccupied: syn. of -C. batrachus Das, 1952 (India) Centrorhynchus indicus Golvan, 1956 -C. bazaleticus Kurashvili, 1955 (SSSR, Georgia) -Ci fasciatus (Westrumb, 1821) Travassos, 1926 (Euro- -CO bengalensis Datta et Soota, (1954) 1955 (India): pe ?):fide MARVAL(1905) it is a syn. of Prostorhyn- may be a (fide NAMA & RATHORE, chus cylindraceus (G.) Golvan, 1956 and fide MEYER 1984) (1932) it is a Mediorhynchus -CO bethaniae George et Nadakal, 1987 (Kerala, India) -Co freundi (Hartwich, 1953) Golvan, 1956 (Kenya) -C bipartitus Solojew, 1912: syn. of Sphaerirostris (nee Hartwick) bipartitus (S.) Golvan, 1956 -C.fukiensis Wang Pu Qin, 1966 (Fukien, China and -C. brama Rengaraju et Das, 1980 (India) (nee bramae) India) -CO brevicanthus Das, (1949) 1950 (India): may be a -C. galliardi Golvan, 1956 (North Africa) Porrorchis fide GOLVAN (1956); syn. of Porrorchis -C. gendrei (Golvan, 1957) Golvan, 1960 (West Africa) brevicanthus (Das, 1949) novo comb. -Co giganteus Travassos, 1920/21 (nee 1919)(Brazil, -C. brevicaudatus Das, (1949) 1950 (India) Panama) 142 Y.J.GOLvAN -C. glaucidii Wang Pu Qin, 1966 (Fukien,China) -C maryasis sensu Gupta et Lata, 1967 is a Centrorhyn- -co globocaudatus (Zeder, 1800) Luhe, 1911 (Europe, ch us sensu stricto; a new specific name must be given Israel, Egypt, Central Asia) -Co merulae Dollfus et Golvan, 1961 -C globuli Nama et Rathore, 1984: syn. of Sphaeriros- -CO microcephalus (Bravo-Hollis, 1947) Golvan, 1956 tris globuli (N. et R.) novo comb. [Mexico, also found in India (doubtful)] -C. golvani Anantaraman et Anantaraman, 1969 (South -CO microcervicanthus Das, 1950 (Berar, India): may be of India) ajuvenile of Centrorhynchus tumidulus (R.) Neiva da -C.(Malgacanthus) grassei Golvan, 1965 (Madagascar) Cunha et Travassos, 1914 (fide YAMAGUTI,1963) (nec -C. guptai Golvan, 1969 (India) mic roce rviacanthus) -C. hagiangensis (Petrotschenko et Phan The Viet, -CO microrchis Fukui, 1929 (Japan, Penjab): may be a 1969) Amin, 1985 (Viet Nam) syn. of Centrorhynchus magnus Fukui, 1929 (fide -CO hargisi Gupta et Fatma,1983 (India): may be a syn. YAMAGUTI,1939) of Sphaerirostris corvi (F.) Golvan, 1958 -C. migrans Zuberi et Farooqi, 1974 (India) (nec Faro- -CO hartwichi nom. novo (Chili): syn. of Centrorhyn- oq) chus conspectus sensu Hartwich, 1956 -CO milvus H.L. Ward, 1956 [Egypt, Israel, India, Sene- -C. horridus (von Linstow, 1897) Meyer, 1932 (Bis- gal (doubtful)] marck Archipelago, New Caledonia, Queensland, Aus- -CO mysentri Gupta et Fatma, 1983 (Lucknow, India) tralia) -CO narcissae Florescu, 1942 (Romania) -C hylae (Johnston, 1914) Schmidt et Kuntz, 1967: syn. -CO nicaraguensis Schmidt et Neiland, 1966 (Nicara- of Porrorchis hylae (J.) Schmidt et Kuntz, 1967 gua) -C. indicus Golvan, 1956 (India) -co ninnii (Stossich, 1891) Meyer, 1932 (nec ninni) -C indicus (Gupta et Lata, 1966) Gupta et Fatma, 1983 (Italy, South of Europe) (preoccupied): syn. of Centrorhynchus latai nom. -C. olssoni Lundstrom, 1942 (Scandinavia) novo -CO opimus Travassos, 1919 [Sao Paulo, Brazil, also -CO insularis Tubangui, 1933 (Luzon, Leyte, Philippine found in India (doubtful)] Islands) -CO petrotschenkoi Kurashvili, 1955 (SSSR, Georgia) -CO itatsinis Fukui, 1929 (Japan, found also in Central -C physocoracis (Porta, 1913): syn. of Sphaerirostris Asia and Afghanistan) physocoracis (P.) Golvan, 1956 -C.javanicus Rengaraju et Das, 1975 (India) (necjava- -C picae (Rudolphi, 1819) Meyer, 1932: syn. of Sphae- nicans) rirostris picae (R.) Golvan, 1958 -CO knowlesi Datta et Soota, (1954) 1955 (India) -C picae sensu Dollfus, 1953: syn. of Sphaerirostris -CO kuntzi Schmidt et Neiland, 1966 (Galapagos, Nica- dollfusi nom. novo ragua, Louisiana, Florida) -C pinguis Van Cleave, 1918: syn. of Sphaerirostris -C lancea (Westrumb, 1821) Skrjabin, 1913: syn. of pinguis (VC.) Golvan, 1956 Sphaerirostris lancea (W.) Golvan, 1956 -CO polemaeti Troncy, 1970 (Chad) -C lanceoides Petrotschenko, 1949: syn. of Sphaeriros- -CO polyacanthus (Creplin, 1825): may be a syn. of Cen- tris lanceoides (P.) Golvan, 1956 trorhynchus buteonis (S.) Kostylew, 1914 (fide MAR- -C. latai nom. novo (Penjab, India): syn. of Centrorhyn- VAL, 1905) ch us indicus (Gupta et Lata, 1966) Gupta et Fatma, -CO polymorphus Travassos, 1926 (Brazil, Neotropical 1983 region) -C leguminosus Solovjew, 1912: syn. of Sphaerirostris -C polymorphus sensu Hartwich, 1956 (Uruguay) is leguminosus (S.) Golvan, 1956 probably another species -CO leptorhynchus Meyer, 1932 (Europe, Egypt) -C. ptyasus Gupta, 1950 (Lucknow, India) -C lesiniformis (Molin, 1859) Meyer, 1932 (Europe) -CO pycnonoti Wang, 1986 (China, Fujian) (nec Bock, 1935): syn. of Sphaerirostris lesiniformis -CO renardi (Lindemann, 1865) Van Cleave, 1924 novo comb. -C rep tans Bhalerao, 1931: syn. of Sphaerirostris rep- -C.lobianchii (Monticelli, 1887) Meyer, 1932 (Europe): tans (B.) Golvan, 1956 may be a Sphaerirostris -C saxicoloides Nama et Rathore, 1984: syn. of Sphae- -Co longicephalus Das, 1950 (Berar, India) rirostris saxicoloides (N. et R.) novo comb. -C. lucknowensis Gupta et Fatma, 1983 (Lucknow, -C scanensis Lundstrorn, 1942: syn. of Sphaerirostris North of India) scanensis (L.) Golvan, 1956; may be a syn. of Sphaeri- -C. mabuiae (von Linstow, 1908) Golvan, 1956 (Java) rostris lancea (W.) Golvan, 1956 (fide VANCLEAVE& -C. macrorchis Das, (1949) 1950 (India) WILLIAMS,1951) -CO madagascariensis (Golvan, 1957) Golvan, 1960 -C serpenticola (von Linstow, 1908): syn. of Sphaeri- (Madagascar) rostris serpenticola (vL.) Golvan, 1956 -Co magnus Fukui, 1929 [Japan, Taiwan, Bengladesh -CO sharmai (Gupta et Lata, 1966) Gupta et Fatma, 1983 and also Azerbaidjan (doubtful)] (India) -C maryasis Datta, 1932: syn. of Sphaerirostris marya- -CO sholapurensis Rengaraju et Das, 1975 (India) sis (D.) Golvan, 1956 -CO simplex Meyer, 1932 (Brazil) Nomenclature of the Acanthocephala 143 -C. sinicus Wang Pu Qin, 1966 (Fukien, China) -C zosteropis Porta, 1913: syn. of Mediorhynchus zos- -C skrjabini Petrotschenko, 1949: syn. of Sphaeriros- teropis (P.) Meyer, 1932 tris skrjabini (P.) Golvan, 1956 (nee skrjarbini) -CO spilornae Schmidt et Kuntz, 1969 (China, Taiwan, CLEAVE/US Subrahmanian, 1927 Bengladesh) -CO circumspinifer Subrahmanian, 1927 (type species) -C spinosus (Kaiser, 1893) Van Cleave, 1924 [Louisia- (Burma) (nee eireulspinifer) na, Florida and Galapagos (VAN CLEAVE, 1940)]; this -CO inglisi (Gupta et Fatma, 1987) novo comb. (Kerala, species was also found in India and Central Asia but India): syn. of Mehrarhynehus inglisi Gupta et Fatma, this ascription is very doubtful 1987 -C spinosus Van Cleave, 1916 is the same species as C. -CO leiognathi lain et Gupta, 1979 (Andamans Islands) spinosus (K.) Van Cleave, 1924 (vide VAN CLEAVE, -CO mysti (Sahay et Sinha, 1971) Amin, 1985 (Bihar, 1924) India) -C splendi Gupta et Gupta, 1971: syn. of Sphaerirostris -CO portblairensis lain et Gupta, 1979 (Andamans splendi (G. et G.) novo comb.; may be a syn. of Cen- Islands) trorhynchus batrachus Das, 1952 (fide lAIN, 1980) but -CO prashadi (Datta, 1940) Golvan, 1969 (Calcutta, East it seems erroneous ofIndia) -C tenuieaudatus (Marotel, 1899) Liihe, 1911: syn. of -CO secundus (Tripathi, 1959) Golvan, 1969 (Chilka Sphaerirostris tenuicaudatus (M.) Amin, 1985 Lake, Port Canning, India) -C teres (Westrumb, 1821) Travassos, 1926: syn. of -CO singhai (Gupta et Fatma, 1987) novo comb. (Kerala, Sphaerirostris teres (W.) Golvan, 1956 India): syn. of Mehrarhynehus singhai Gupta et Fatma, -CO tumidulus (Rudolphi, 1819) Neiva da Cunha et Tra- 1987 vassos, 1914 (nee tumidulum) (Brazil, Venezuela, Uru- guay, , Panama; also found in Penjab but it is CORYNOSOMA Luhe, 1904 (vide VAN CLEAVE,1945) certainly erroneous) -CO alaskensis Golvan, 1959 (Alaska) (nee alaskaensis) -C turdi Yamaguti, 1939: syn. of Sphaerirostris turdi -CO ambispinigerum Harada, 1935 (nee ambispige- (Y.) Golvan, 1956 rinum) (North Pacific): may be a syn. of Corynosoma -CO tyotensis Rengaraju et Das, 1975 (India) osmeri Fujita, 1921 (fide HARADA,1935) -CO undulatus (Nitzsch in Giebel, 1886) Dollfus, 1951 -CO anatarium Van Cleave, 1945 (nee anatarum) [USA, (Morocco) Texas, found also in Siberia (doubtful)] -CO wardae Holloway, 1958 (U.S.A.) -CO antarcticum (Rennie, 1906) Johnston et Best, 1937 -C. wedli (Sonsino, 1896) Porta, 1909: may be a syn. of (nee Leiper et Atkinson, 1914): may be a syn. of Cory- Centrorhynchus buteonis (G. in S.) Kostylew, 1914 (fide nosoma hamanni (vL.) Railliet et Henry, 1907 PORTA,1909) but for DOLLFUS(1951) it is a valid species -CO arctocephali Zdzitowiecki, 1984 (Antarctic) -C zosteropis Porta, 1913: syn. of Mediorhynchus zos- -CO australe Johnston, 1937 (Australia) teropis (P.) Meyer, 1932 -CO baylisi Zdzitowiecki, 1986 (Antarctic) -Centrosoma buteonis in Porta, 1910: vide Centrorhyn- -CO bipapillum Schmidt, 1965 (Alaska): may be a syn. ch us buteonis of Corynosoma constrictum Van Cleave, 1918) -C. bullosum (von Linstow, 1892) Railliet et Henry, CHENTRORHYNCHUS Neiva da Cunha et Travassos, 1907 [South Georgia, Antarctic, found also in Califor- 1914: errat. pro Centrorhynchus Liihe, 1911 nia (fide SCHMIDT& DAILEY, 1971)(doubtful) (vide ZDZITOWIECKI,1984)] CHENTROSOMA Monticelli, 1905 nee Porta, 1906 et -CO cameroni Van Cleave, 1953 (Quebec, Canada) 1908: composite genus, syn. of Centrorhynchus Luhe, -CO caspicum Golvan et Mokhayer, 1973 (Caspian Sea) 1911 pro parte, and of Mediorhynchus Van Cleave, -CO cetaceum Johnston et Best, 1942 (South Australia): 1916 pro parte and of Corynosoma Liihe, 1904 pro may be a Polymorphus (fide SCHMIDT & DAILEY, parte 1971) but it seems very doubtful because the worms they studied were from Stenodelphis blainvillei (Ger- CHENTROSOMA Porta, 1908: syn. of Centrorhynchus vais, 1844) [=Pontoporia b. (G.) Gray, 1846] endemic Liihe, 1911 in the Rio de la Plata -C alueonis (O.F. Miiller, 1780) Porta, 1909: syn. of -C clavatum Goss, 1940: syn. of Andracantha clavata Centrorhynchus aluconis (O.F.M.) Liihe, 1911 (G.) Zdzitowiecki, 1989 -C alueonis sensu Porta, 1913 (New Caledonia) is ano- -C clementei Giovannoni et Fernandes, 1965 in Amin, ther species 1985 (nee clementi): syn. of C. seropedicus Machado -C buteonis in Porta, 1908: syn. of Centrorhynchus Filho, 1965 (vide NORONHA,1988) buteonis (G. in S.) Kostylew, 1914 -co constrictum Van Cleave, 1918 (Wyoming, Illinois, -C horridum (von Linstow, 1897) Porta, 1913: syn. of Ohio, USA; Nearctic) Centrorhynchus horridus (vL.) Meyer, 1932 -CO curilensis Gubanov, 1952 (nee kurilense, nee euri- -C ninni (Stossich, 1891) in Porta, 1908: syn. of Cen- liens is) (Kurile Islands) trorhynchus ninnii (S.) Meyer, 1932 -CO enhydri Morosov, 1940 (North Pacific) 144 Y.J. GOLVAN -c. enhydris Afanas' ev, 1941: syn. of Corynosoma (the spelling piriforme is etymologically more correct): enhydri~orosov, 1940 may be a Polymorphus (fide ~ACHALSKA, 1981) -C. enrietti Molfi et de Freitas-Fernandes, 1953 (Parana, -Co rauschi Golvan, 1959 (Central Pacific, Midways) Brazil) -CO reductum (von Linstow, 1905) Railliet et Henry, -c. enrietti Molfi et de Freitas-Fernandes, 1953: syn. 1907 (Arctic) pro parte of C. molfifernandesi Machado Filho, 1962 -Co semerme (Forssell, 1904) Liihe, 1905 (nee Frossell) -CO eperlani (von Linstow, 1884) Petrotschenko, 1958 (Arctic) -CO evae Zdzitowiecki, 1984 (Antarctic) -c. semerme sensu Cordero, 1933 (Rio de la Plata): -C.falcatum Van Cleave, 1953 (Alaska) may be a syn. of Corynosoma (Polymorphus T) ceta- -c. gibber (Olsson, 1894) Liihe, 1911: syn. of Coryno- ceum Johnston et Best, 1942 (fide SCHMIDT& DAI- soma semerme (F.) Liihe, 1905 LEY, 1971) -Co gibsoni Zdzitowiecki, 1986 (Falklands) -Co semerme sensu Fukui, 1929 =?? -c. gravida Alegret, 1941: syn. of Andracantha gravida -c. semerme sensu Lyster, 1940: syn. of Corynosoma (A.) Schmidt, 1975 hadweni Van Cleave, 1953 pro parte and of Corynoso- -Co hadweni Van Cleave, 1953 (Alaska) (nee hadveni): ma validum Van Cleave, 1953 pro parte (vide V AN may be a syn. of Corynosoma wegeneri Heinze, 1934 CLEAVE,1953) (fide ~ARGOLlS, 1955) -CO septentrionalis Treschev, 1966 (Behring Sea) -C. hamanni (von Linstow, 1892) Railliet et Henry, -CO seropedicus Machado Filho, 1970 (Rio de Janeiro, 1907 (nee Leiper et Atkinson, 1915)(Booth- Wan del Brazil) Islands, South Orkneys, Antarctic) -C. shackletoni Zdzitowiecki, 1978 (South Shetlands, -Co hannae Zdzitowiecki, 1984 (Antarctic) Antarctic) -c. hystrix tunitae Weiss, 1914: syn. of Corynosoma -Co similis Neiland, 1962 (Alaska) tunitae Weiss, 1914 -c. singulare sensu Zdzitowiecki, 1978 pro parte: syn. -Co iheringi Machado Filho, 1961 (Sao Paulo, Brazil) of Corynosoma evae Zdzitowiecki, 1984 -c. kurilensis Gubanov, 1952: vide Corynosoma curi- -CO singularis Skrjabin et Nikolski, 1971 (Ross Sea, lensis Antarctic): may be a syn. of Corynosoma bullosum -Co longilemniscatus Machado Filho, 1961 (Brazil) (vL.) Railliet et Henry, 1907 pro parte and of Coryno- -Co macrosomum Neiland, 1962 (Alaska, North Paci- soma arctocephali Zdzitowiecki, 1984 pro parte (vide fic): may be a syn. of Corynosoma enhydri M. (fide ZOZITOWIECKI,1984) or may be a syn. of Corynosoma JELLlSON& NEILANO,1965) bullosum (fide ZOZITOWIECKI,1986) -C. magdaleni Montreuil, 1958 (nee magdaleiniy (Cana- -c. sipho Railliet et Henry, 1907: syn. of Corynosoma da, Gulf of the Saint Laurent) hamanni (vL.) Railliet et Henry, 1907 -Co mandarin ea Oschmarin, 1963 (East Siberia, Nearc- -CO stanleyi Smales, 1986 (Tasmania, Australia) tic) -co strumosum (Rudolphi, 1802) Liihe, 1904 (type spe- -C. marchii Porta, 1910 (Firenze, Italy): may be a Poly- cies) (Holarctic Region) morphus (fide ~EYER, 1932) -c. strumosum sensu Dollfus, 1962 is another species, -c. mergi Lundstrom, 1941: syn. of Andracantha mergi may be an Andracantha (Morocco) (L.) Schmidt, 1975 -c. strumosum sensu Dogiel et Bychowsky, 1938 is -Co mirabilis Skrjabin, 1966: may be a syn. of Coryno- another species (Caspian Sea) soma bullosum (fide ZOZITOWIECKI,1986) -c. strumosum sensu Hartwich, 1956 is another species -Co molfifernandesi Machado Filho, 1962 (Brazil) (Uruguay) -CO obtuscens Lincicome, 1943 (USA, California and -c. strumosum sensu Lyster, 1940: syn. of Corynosoma East Siberia) cameroni Van Cleave, 1953 -C. osmeri Fujita, 1921 (Japan, East Siberia) -C. sudsuche Belopolskaja, 1958 (East Siberia) -C. otariae Morini et Boero, 1960 (Argentina): may be a -c. tunitae Weiss, 1914: syn. of Andracantha tunitae syn. of Corynosoma australe Johnston, 1937 (fide (W.) Zdzitowiecki, 1989 ZDZITOWIECKI1989), -CO turbidum Van Cleave, 1937 (Dyer Island, South -CO pacifica Nikolski, 1974 (Ross and Wed dell Seas, Africa): may be a syn. of Arhythmorhynchus turbidus Antarctic): may be a syn. of Corynosoma hamanni novo comb. (vL.) Railliet et Henry, 1907 -Co validum Van Cleave, 1953 (Alaska) -CO peposacae (Porta, 1914) Travassos, 1924 (Ar- -Co ventronudum Skrjabin, 1959 in Amin, 1985 gentina) -CO villosum Van Cleave, 1953 (Alaska, Japan) (nee -Co peposaeae sensu Travassos, 1925: syn. of Coryno- viliosum) soma longilemniscatus ~.F. -Co wegeneri Heinze, 1934 (Greenland) -c. phalaeroeoraeis Yamaguti, 1939: syn. of Andra- cantha phalacrocoracis (Y.) Schmidt, 1975 DELTACANTHUS Diaz-Ungria et Gracia-Rodrigo, 1958: -CO pseudohamanni Zdzitowiecki, 1984 (South Shet- preoccupied, syn. of Acanthodelta D.-U. et G.-R., 1958 lands, Antarctic) -D. scorzai (Dfaz-Ungrfa et Gracia-Rodrigo, 1957) -C. pyriforme (Bremser in Rudolphi, 1819) Meyer, 1932 Diaz-Ungrta et Gracia-Rodrigo, 1958: syn. of Acan- Nomenclature of the Acanthocephala 145 thodelta scorzai (D.-U. et G.-R.) Dfaz-Ungria et Gra- -E. sagittifer (Linton, 1889) Monticelli, 1905: syn. of cia-Rodrigo, 1958 Serrasentis sagittifer (L.) Van Cleave, 1923

DELTANIA Diaz-Ungria et Gracia-Rodrigo, 1957: pre- ECHINOPARDALIS Travassos, 1918: it is valid genus, occupied, syn. of Acanthodelta D.-U. et G.-R., 1958 not a syn. of Oligacanthorhynchus Travassos, 1915 -D. scorzai Dfaz-Ungria et Gracia-Rodrigo, 1957: syn. nor of Oncicola Travassos, 1916 (fide NrCKOL & of Acanthodelta scorzai (D.-U. et G.-R.) Dfaz-Ungria DUNAGAN, 1989); the main distinctive characters are et Gracia-Rodrigo, 1958 the morphology of the trunk, the male genitalia and the structure of the protonephridia; moreover there are 36 DENDRONUCLEATA Sokolowskaja, 1962 hooks on the ; they are all parasitic in Carni- -D. dogieli Sokolowskaja, 1962 (type species) (Amur vora (Canidae, Viverridae, Felidae) River basin) -E. atrata Meyer, 1931 (sea shore of the Red Sea, -D. petruschewskii Sokolowskaja, 1962 (Amur River Egypt) basin) -E. bangalorensis Pujatti, 1951: may be a syn. of Oliga- canthorhynchus bangalorensis (P.) Schmidt, 1972. DENTITRUNCUS Sinzar, 1955 (nee Dentitrunkus) The original description was based on the examination -D. truttae Sinzar, 1955 (type species) (Jugoslavia, of a juvenile found in the South of India encysted North of Italy) in the neck of a pheasant -E. decrescens Meyer, 1931 (Brazil) DEVENDROSENTIS Sahay, Sinha et Ghosh, 1971 -E. lamasi de Freitas et Costa, 1964: syn. of Oligacan- -D. garuai Sahay, Sinha et Ghosh, 1971 (type species) thorhynchus lamasi (dF. et C.) Amato, Nickol et (Patna, India): may be a syn. of Pallisentis garuai (fide Froes, 1979 JAIN & GUPTA, 1979 and GUPTA& FATMA, 1986) -E. lerouxi Bisseru, 1956 (North ): may be a syn. of lerouxi (B.) Baer, 1959 DIPLOSENTIS Tubangui et Masilungan, 1937 -E. macrurae Meyer, 1931 (page 102) (nee macrou- -D. amphacanthi Tubangui et Masilungan, 1937 (nee rae in Meyer, 1931, page 64) ( and amphaeanthuri) (type species) (Mindanao, Philippine California, USA): is not a syn. of Oncicola macru- Islands) rae (M.) Witenberg, 1938 (vide NrCKOL & DUNA- -D. manteri Gupta et Fatma, 1979 (Tamil Nadu, India) GAN, 1989) -E. mariemilyi Tadros, 1969: syn. of Oligacanthorhyn- DIPLOSPINIFER Fukui, 1929 ch us mariemilyi (T.)Amin, 1985 (vide NrCKOL & -D. serpenticola Fukui, 1929 (type species) (Japan) DUNAGAN,1989) DISPIRON Bilqees, 1970 -E. pachyacanthus (Sonsino, 1889) Neveu-Lernaire, -D. catlai Aly Khan et Bilqees, 1987 (Kalri Lake, Pakis- 1936: may be a (fide TRAVASSOS, tan) 1917); may be a good species, different from E. atrata -D. heteroacanthus Aly Khan et Bilqees, 1985 (Pakis- (fide NrCKOL& DUNAGAN,1989) tan) -E. pardalis (Westrumb, 1821) Travassos, 1918 (type -D. mugilis Bilqees, 1970 (type species) (Pakistan) (nee species) [Brazil; also found in Chandigarh (India) mugili) (doubtful)J -E. pardalis sensu Travassos, 1917 pro parte: syn. of DISTEGANIUS Lehmann, 1953: nomen nudum, syn. of Oncicola campanulata (D.) Meyer, 1931 Mediorhynchus Van Cleave, 1916 -D. eolini Lehmann, 1953: nomen nudum, syn. of Me- ECHINORHYNCHOIDES Achmerov et Dombrowska- diorhynchus colini (W.) Schmidt et Kuntz, 1977 ja-Achmerova,1941 -E. dogieli Achmerov et Dornbrowskaja-Achrnerova, DOLLFUSENTIS Golvan, 1969 1941 (type species) (East Siberia) -D. bravoae Salgado-Maldonado, 1976 (Vera Cruz, -E. meyeri Gupta et Naqvi, 1984 (India) Mexico) -E. nudus (Harada, 1938) Petrotschenko, 1956: syn. of -D. chandleri Golvan, 1969 (type species) (gulf of Neorhadinorhynchus nudus (H.) Yamaguti, 1939 Mexico) -D. ctenorhynchus (Cable et Linderoth, 1963) Golvan, ECHINORHYNCHUS Zoega in O.F. Mi.iller, 1776 1969 (Jamaica) -E. (Echinorhynchus) abyssicola Dollfus, 1931 (North -D. heteracanthus (Cable et Linderoth, 1963) Golvan, of the Atlantic Ocean) 1969 (Curacao) -E. aeanthosoma Westrumb, 1821: syn. of Telosentis -D. longispinus (Cable et Linderoth, 1963) Golvan, molini VC.,1923 1969 (Jamaica): is not a syn. of Dollfusentis chandleri -E. aeanthotrias von Linstow, 1883: impossible to iden- Golvan, 1968 (fide SALGADO-MALDONADO,1978) tify [may be a Centrorhynchus (fide MARVAL, 1905), ECHINOGASTER Monticelli, 1905: preoccupied, syn. or Mediorhynchus (fide MEYER, 1932) or Cen- of Serrasentis Van Cleave, 1923 trorhynchinae (fide PETROTSCHENKO1958), J 146 Y.J. GOLVAN -E. acipenseris rutheni Rudolphi, 1819: syn. of Lep- -E. annulatus Molin, 1858: syn. of Bolbosoma vasculo- torhynchoides plagicephalus (W.) Kostylew, 1924 sum (R.) Meyer, 1932 -E. acus Rudolphi, 1802: syn. of Echinorhynchus gadi -E. antarcticus Rennie, 1906 (nec antarcticum): syn. of Zoega, 1776 (vide KOSTYLEW,1925) Corynosoma hamanni (vL.) Railliet et Henry, 1907 -E. acus sensu Leidy, 1856: syn. of Echinorhynchus -E. anthuris Dujardin, 1845: syn. of Acanthocephalus (E.) vancleavei Golvan, 1969 anthuris (D.) Liihe, 1911 -E. aequalis Zeder, 1803: syn. of Centrorhynchus alu- -E. appendiculatus Westrumb, 1821: syn. of Monilifor- conis (O.F.M.) Liihe, 1911 mis sp. (fide MEYER, 1932) or of Centrorhynchus alu- -E. affinis Gmelin, 1791: syn. of Neoechinorhynchus conis (O.F.M.) Luhe, 1911 (fide GOLVAN,1956) rutili (O.F.M.) Hamann, 1892 -E. arcticus von Linstow, 1901: syn. of Echinorhyn- -E. affinis Rudolphi, 1802: syn. of Acanthocephalus chus gadi O.F. Muller, 1776 (vide KOSTYLEW,1925) lucii (O.F.M.) Liihe, 1911 -E. arcuatus Diesing, 1851: syn. of Polyacanthorhyn- -E. agilis Rudolphi, 1819: syn. of Neoechinorhynchus ch us macrorhynchus (D.) Travassos, 1920 agilis (R.) Van Cleave, 1916 -E. ardeae Gmelin, 1789: syn. of Polymorphus striatus -E. agilis sensu Linton, 1889 et 1901: syn. of Neoechi- (G.) Liihe, 1911 norhynchus cylindratus (VC.) Van Cleave, 1916 -E. ardeae albeae Rudolphi, 1809: may be a syn. of -E. alaudae Rudolphi, 1819: syn. of Mediorhynchus Echinorhynchus garzeae Zeder, 1800 which is not an micracanthus (R.) Van Cleave, 1927 acanthocephalan (fide MEYER, 1932) -E. alcae Gmelin in O.F. Muller, 1780: syn. (?) of Fili- -E. ardeae purpureae Rudolphi, 1819: syn. of Arhyth- collis anatis (S.) Luhe, 1911 morhynchusfrassoni (M.) Liihe, 1911 -E. alcedinis Westrumb, 1821: species delenda (fide -E. areolatus Rudolphi, 1819: syn. of Sphaerirostris MEYER, 1932) areolatus (R.) Golvan, 1956 -E. alosae Hermann, 1782: syn. of Rhadinorhynchus -E. areolatus sensu de Marval, 1905: may be a syn. of alosae (H.) Meyer, 1932 Mediorhynchus oswaldocruzi Travassos, 1923 -E. (Metechinorhynchus) alpinus von Linstow, 1901 -E. argentinae Gmelin, 1789 is not an acanthocephalan (Central Asia, China): is not a syn. of Echinorhynchus (fide MEYER, 1932) salmonis O.F. Muller, 1784 -E. arioli in Rudolphi, 1819 may be a Mediorhynchus -E. aluconis O.F. Muller, 1780: syn. of Centrorhynchus (fide TRAVASSOS,1920/21) (vide Echinorhynchus orio- aluconis (O.F.M.) Luhe, 1911 li cristati) -E. amphipachus Westrumb, 1821 (nec amphipacus) -E. (Echinorhynchus) armoricanus Golvan, 1969 may be an (?) (fide MEYER, (Atlantic Coast of France) 1932) or a Nephridiacanthus or a Macracanthorhyn- -E. astaci fluviatilis Diesing, 1851: impossible to identify chus (fide GOLVAN,1962) -E. atherinae Rudolphi, 1819: syn. of Telosentis molini -E. anatarum Rudolphi, 1819: syn. of Polymorphus Van Cleave, 1923 minutus (G.) Luhe, 1911 pro parte and of Polymor- -E. (Echinorhynchus) attenuatus Linton, 1888 (nec ph us boschadis (S.) Railliet, 1919 pro parte 1890 et 1891) (North Atlantic coast of the USA) -E. anatis Schrank, 1788: syn. of Filicollis anatis (S.) -E. attenuatus O.F. Mtiller, 1779: syn. (?) pro parte of Ltihe,1911 Pomphorhynchus tereticollis (R.)Engelbrecht, 1957 -E. anatis sensu Gmelin, 1791: syn. of Polymorphus -E. attenuatus sensu Gracia-Rodrigo, 1960 is probably minutus (G.) Luhe, 1911 another species -E. anatis sensu Frolich, 1802: syn. of Polymorph us -E. aurantiacus Risso, 1826: syn. of Bolbosoma vascu- boschadis (S.) Railliet, 1919 losum (R.) Meyer, 1932 -E. anatis mollissimae Rudolphi, 1809: syn. of Polymor- -E. aurantiacus sensu Monticelli, 1887: syn. of Solea- phus phippsi Kosty lew, 1922 rhynchus soleae (P.) de Buron et Maillard, 1985 -E. anatarum Rudolphi, 1819: syn. of Polymorphus -E. bacillaris Zeder, 1803: syn. of Centrorhynchus alu- minutus (G.) Luhe, 1911 pro parte and of Polymor- conis (O.F.M.) Liihe, 1911 or of Centrorhynchus ph us boschadis (S.) Railliet, 1919 pro parte buteonis (G.) Kostylew, 1914 -E. anguillae O.F. Mtiller, 1780: syn. of Acanthocepha- -E. bacillaris sensu von Linstow, 1901 is not a Centro- Ius anguillae (O.F.M.) Liihe, 1911 rhynchus (fide KOSTYLEW,1925) -E. angustatus Rudolphi, 1802: syn. of Acanthocepha- -E. (Pseudoechinorhynchus) baeri Kostylew, 1928 Ius lucii (O.F.M.) Luhe, 1911 (Armenia) -E. angustatus sensu Leidy, 1856: syn. of Echinorhyn- -E. balaenae Gmelin, 1790: syn. of Bolbosoma balae- chus gadi O.F.M. (fide VAN CLEAVE, 1924) or of nae (G.) Meyer, 1932 Echinorhynchus (E.) vancleavei Golvan, 1969 -E. balaenocephalus Owen, 1803: syn. of Bolbosoma -E. annularis Gmelin, 1791 in Rudolphi, 1809 and in turbinella (D.) Porta, 1908 Westrumb, 1821: syn. of Pomphorhynchus laevis (Z.) -E. barbi Schrank, 1792: syn. of Pomphorhynchus lae- Van Cleave, 1924 vis (Z.) Van Cleave, 1924 -E. annulatus Gmelin, 1791: syn. of Pomphorhynchus -E. bazae Southwell et Macfie, 1925: may be a syn. of laevis (Z.) Van Cleave, 1924 Porrorchis bazae (S. et M.) Schmidt et Kuntz, 1967 Nomenclature of the Acanthocephala 147 -E. belgieus Railliet, 1918: syn. of Moniliformis monili- liformis moniliformis (B.) Travassos, 1915 formis (B.) Travassos, 1915 (fide MEYER, 1932) -E. canyonensis Huffman et Kliever, 1977 (nec Kleiver) -E. bifaseiatus Ltihe, 1904: syn. of Bolbosoma vasculo- (West USA) sum (R.) Meyer, 1932 -E. capitatus von Linstow, 1880 (nec capitatum): syn. of -E. bipennis Kaiser, 1893 (Argentina) is impossible to Bolbosoma capitatum (vL.) Porta, 1908 identify -E. carassii Rudo1phi, 1819: syn. of Neoechinorhyn- -E. blennii Rudolphi, 1810: syn. of Acanthocephalus ehus rutili (O.F.M.) Hamann, 1892 lucii (O.F.M.)Luhe, 1911 -E. carchariae Linton, 1888: species delenda -E. borealis Gmelin, 1791: syn. of Polymorphus phippsi -E. carehariae Linton, 1891: may be a syn. of Coryno- Kostylew, 1922 soma strumosum (R.)Luhe, 1904 (vide VAN CLEAVE, -E. borealis von Linstow, 1901: preoccupied, syn. of 1953) Echinorhynchus (Pseudoechinorhynchus) cinctulus -E. earpionis Gmelin, 1790: may be a syn. of Acantho- Porta, 1905 cephalus anguillae (O.F.M.) Luhe, 1911 (fide LUHE, -E. bosehadis Schrank, 1788: syn. of Polymorphus bos- 1911 and MEYER, 1932), or of Neoechinorhynchus chadis (S.) Railliet, 1919 rutili (O.F.M.) Hamann, 1892 (fide RUDOLPHI,1809) -E. bosehadis Gmelin, l791: syn. (?) of Filicollis anatis -E. carruccioi Condorelli, 1897: may be a syn. of (S.) Luhe, 1911 Mediorhynchus micracanthus (R.) Meyer, 1932 -E. bosehatis Frolich, 1802: syn. of Polymorphus minu- -E. caudatus Zeder, 1803: syn. of Centrorhynchus tus (G.) Luhe, 1911 buteonis (G. in S.) Kosty1ew, 1914 -E. bothinensis Zdzitowiecki et Valtonen, 1987 (Gulf of -E. caudatus sensu Rudolphi, 1819: syn. of Centrorhyn- Bothnia) eh us tumidulus (R.) Neiva da Cunha et Travassos, -E. bramae Gmelin, 1790: syn. of Pomphorhynchus 1914 laevis (Z.) Van Cleave, 1924 -E. caudatus sensu Westrumb, 1821: syn. of Cen- -E. brevieollis Malm, 1867: syn. of Bolbosoma brevico- trorhynchus aluconis (O.F.M.) Ltihe, 1911 pro parte lie (M.) Porta, 1908 and of Centrorhynchus tumidulus (R.) Neiva da Cunha -E. brevieollis sensu von Linstow, 1901 and 1908: may et Travassos, 1914 pro parte be a syn. of Echinorhynchus gadi Zoega (fide KOSTY- -E. caudatus sensu Diesing, 1851: syn. of Centrorhyn- LEW, 1925) ch us giganteus Travassos, 1919 -E. (Metechinorhynchus) briconi Machado Filho, 1959 -E. caudatus sensu Leidy, 1887: syn. of Centrorhyn- (Sao Paulo, Brazil) ch us spinosus (K.) Van Cleave, 1916 -E. brumpti Blanc et Cauchemez, 1911: syn. of Pros- -E. centropi Porta, 1910: syn. of Porrorchis centropi torhynchus cylindraceus (G.) Golvan, 1956 (P.) Schmidt et Kuntz, 1967 -E. bufonis Shipley, 1903: syn. of Pseudoacanthoce- -E. centropusi Tubangui, 1933: syn. of Porrorchis hylae phalus bufonis (S.) Petrotschenko, 1956 (1.) Schmidt et Kuntz, 1967 -E. bulboeaudatus Southwell et Macfie, 1925: syn. of -E. cernuae Gmelin, l791: syn. of Pomphorhynchus Porrorchis hylae (1.) Schmidt et Kuntz, 1967 laevis (Z.) Van Cleave, 1924 -E. bulbosus von Linstow, 1892: syn. of Corynosoma -E. cestodicola von Linstow, 1905: very doubtful species bullosum (vL.) Railliet et Henry, 1907 -E. cestodiformis von Linstow, 1904: syn. of Monilifor- -E. bullosus von Linstow, 1892: syn. of Corynosoma mis cestodiformis (vL.) Travassos, 1917 bullosum (vL.) Railliet et Henry, 1907 -E. charadrii pluvialis Rudolphi, 1819: nomen nudum, -E. buteonis Goeze, 1782 in Schrank, 1788: syn. of Cen- may be a syn. of Oligacanthorhynchus ricinoides (R.) trorhynchus buteonis (G.) Kostylew, 1914 Travassos, 1915 (fide MEYER, 1932) -E. eaenoformis Heitz, 1920: syn. of Bolbosoma caeno- -E. charcharine Linton, 1891: may be a syn. of Coryno- forme (H.) Meyer, 1932 soma strumosum (R.)Luhe, 1904 -E. (Echinorhynchus) calloti Golvan, 1969 (Atlantic -E. cherchiae Monticelli, 1889 (nee chierchiae) (Carib- coast of France) bean Sea) -E. eampanulatus Diesing, 1851 et 1856: syn. of Onci- -E. (Pseudoechinorhynchus ) cinctulus Porta, 1905 cola campanulata (D.) Meyer, 1931 (Volga River Basin, North East of Europe and Siberia): -E. eampanulatus sensu Leidy, 1856: syn. of Echino- may be a syn. of Echinorhynchus borealis von Lins- pardalis pardalis (W.)Travassos, 1918 tow, 1901 [fide GRABDA-KAZUBSKA & EJSYMONT -E. eampbelli Leiper et Atkinson, 1914: syn. of Meta- (1969) but this name is preoccupied by Echinorhyn- canthocephalus campbelli (L. et A.) Golvan, 1969 chus borealis Gme1in, 1791 which is probably a Poly- -E. campylurus Nitzsch in Giebel, 1857: syn. of Oligote- morphus phippsi Kostylew, 1922] rorhynchus campylurus (N.) Monticelli, 1905 -E. einetus Rudolphi, 1819: syn. of Centrorhynchus -E. candidus Zoega in O.F. Miiller, 1776: syn. of Echi- leptorhynchus Meyer, 1932 pro parte and of Sphaeri- norhynchus gadi Zoega, 1776 rostris lancea (W.) Go1van, 1956 pro parte -E. canis Kaupp, 1909: syn. of Oncicola canis (K.) Hall -E. circumflexus Molin, 1858: syn. of Oligacanthorhyn- et Wigdor, 1918 eh us circumflexus (M.) Meyer, 1932 -E. canis sensu Porta, 1914: may be a syn. (?) of Moni- -E. citilli Rudo1phi, 1806: syn. of Oligacanthorhynchus 148 Y.J. GOLVAN citilli (R.) Kostylew et Zemeev, 1939 -E. (Echinorhynchus) cotti Yamaguti, 1935 (Biwa -E. clavaeceps Zeder, 1800: syn. of Neoechinorhyn- Lake, Japan) ch us rutili (O.F.M.) Hamann, 1892 -E. crassicollis Villot, 1875: syn. of Plagiorhynchus -E. clavaeceps sensu Linton, 1913 in Sumner, Osborn et crassicollis (V.)Luhe, 1911 Cole, 1913: syn. of Neoechinorhynchus cylindratus -E. croaticus Stossich, 1889: syn. of Centrorhynchus (VC.) Van Cleave, 1921 aluconis (O.F.M.) Liihe, 1911 -E. clavaesceps in Rudolphi, 1810: errat. pro E. clavae- -E. (Metechinorhynchus) cryophilus (Sokolowskaja, ceps Zeder, 1800 1962) Amin, 1985 (Amur River Basin) -E. claviceps in Lunel, 1874: errat. pro E. clavaeceps -E. cuniculi Bellingham, 1844: species delenda Zeder, 1800 -E. curvatus von Linstow, 1897: syn. of Neoncicola -E. clavoeus in Lunel, 1879/1880: errat. pro E. clavae- curvata (vL.) Schrnidt, 1972 ceps Zeder, 1800 -E. cylindraceus Goeze, 1782 in Schrank, 1788: syn. of -E. clavula Dujardin, 1845: syn. of Acanthocephalus Prosthorhynchus cylindraceus (S.) Golvan, 1956 clavula (D.) Grabda-Kazubska et Chubb, 1968 -E. cylindraceus sensu Porta, 1913 from New Caledonia -E. clavula sensu Hamann, 1891 et 1892: syn. of Echi- is another species norhynchus (Metechinorhynchus) truttae Schrank, -E. cyprini idbari Rudolphi, 1819: syn. of Pomphorhyn- 1788 chus laevis (Z.) Van Cleave, 1924 -E. clavula sensu Ltihe, 1911: syn. of Echinorhynchus -E. cyprini rutili RudoJphi, 1819: syn. of Acanthoce- (Pseudoechinorhynchus) cinctulus Porta, 1905 phalus anguillae (O.F.M.) Luhe, 1911 -E. clavula sensu Southwell et Macfie, 1925: syn. of -E. debenhami Leiper et Atkinson, 1914: syn. of Meta- Acanthocephalus hastae Baylis, 1944 canthocephalus rennicki (L. et A.) Zdzitowiecki, 1983 -E. clavulus sensu Johnston et Deland, 1929, from Que- -E. decipiens Dujardin, 1845: incertae sedis, may be a ensland, is different from the species named by SOUTH- Prosthorhynchidae WELL& MACFIE(1925) -E. depressus Nitzsch, 1866 is an Oligacanthorhynchi- -E. cobitidis Gmelin, 1791: syn. of Neoechinorhynchus dae (fide MEYER, 1932) rutili (O.F.M.) Hamann, 1892 -E. dendrocopi Westrumb, 1821 (nec dentrocopi): spe- -E. cobitidis barbatulae Goeze, 1782: syn. of Neoechi- cies delenda norhynchus rutili (O.F.M.) Hamann, 1892 -E. devisiani Molin, 1858 (nec devisiana): syn. of Acan- -E. cobitinus Schrank, 1788: syn. of Neoechinorhyn- thocephaloides incrassatus (M.) Meyer, 1932 ch us rutili (O.F.M.) Hamann, 1892 -E. diffluens Zenker, J832: syn. of Polymorphus minu- -E. collaris Schrank, 1792: syn. of Polymorphus minu- tus (G.) Luhe, 1911 tus (G.) Liihe, 1911 pro parte and of Polymorphus -E. dimorphocephalus Westrumb, 1821: syn. of Centro- boschadis (S.) Railliet, 1919 pro parte rhynchus dimorphocephalus (W.) Meyer, 1932 -E. collurionis Rudolphi, 1819: nomen nudum, may be a -E. dipsadis von Linstow, 1888: syn. of Centrorhyn- syn. of Polymorphus contortus (B.) Travassos, 1926 chus dipsadis (vL.) Golvan, 1956 (fide MEYER, 1932) -E. dirus Van Cleave, 1931: syn. of Acanthocephalus -E. compressus Rudolphi, 1802: syn. of Oligacan- dirus (VC.) Van Cleave et Townsend, 1936 thorhynchus compressus (R.) Meyer, 1932 -E. (Echinorhynchus) dissimilis Yamaguti, 1939 -E. conicus Zed er, 1803 is not an acanthocephalan (fide (Sizuoka, Japan) MEYER, 1932) -E. dobulae Schrank, 1790 et 1792: syn. of Pompho- -E. constrictus Zeder, 1800: syn. of Filicollis anatis (S.) rhynchus laevis (Z.) Van Cleave, 1924 pro parte and Liihe, 1911 of Pomphorhynchus tereticollis (R.) Engelbrecht, -E. contortus Bremser in Westrumb, 1821: syn. of Poly- 1957 pro parte morphus contortus (B.) Travassos, 1926 -E. echinodiscus Diesi ng, 1851: syn. of Gigantorhyn- -E. contortus Molin, 1861: may be a syn. of Centro- ch us echinodiscus (D.) Hamann, 1892 rhynchus aluconis (O.F.M.) Luhe, 1911 -Echinorhynchus (Echinorhynchus) ekbaumi Golvan, -E. coraciae Rudolphi,1819 in Westrumb, 1821: syn. of 1969 (Pacific coast of North America) (nec eckbaumi): Oligacanthorhynchus ricinoides (R.) Travassos, 1915 is not a syn. of Echinorhynchus gadi Zoega, 1776 -E. (Metechinorhynchus) coregoni Linkins in Van Cle- -E. elegans Diesing, 1851: syn. of Prosthenorchis ele- ave, 1919 (Great Lakes): is not a syn. of Echinorhyn- gans (D.) Travassos, 1915 pro parte and of Oncicola chus salmonis O.F. Miiller, 1784 macrurae Meyer, 1931 pro parte -E. cornicis Rudolphi, 1819: syn. of Oligacanthorhyn- -E. embeae Cholodkowsky et Kostylew, 1916: syn. of ch us compressus (R.) Meyer, 1932 Sphaerirostris embeae (C. et K.) Golvan, 1956 -E. (sensu lato) corrugatus Sars, 1885 (expedition -E. emberizae Rudolphi, 1819: syn. of Mediorhynchus «Challenger» ) emberizae (R.) Travassos, 1924 (nec Van Cleave, 1916) -E. corrugatus sensu Monticelli, 1900: syn. of Solea- -E. emydis Leidy, 1851: syn. of Neoechinorhynchus rhynchus soleae (P.) de Buron et Maillard, 1985 emydis (L.) Van Cleave, 1919 -E. coryphaenae Zeder in Rudolphi, 1899 is not an -E. eperlani Rudolphi, 1809 may be a Polymorphinae acanthocephalan (fide MEYER, 1932) (fide MEYER, 1932) Nomenclature of the Acanthocephala 149 -E. eperlani sensu von Linstow, 1884: must be deleted -E. georgianus Rodyuk, 1986 (Western part of the because it is a juvenile impossible to identify and the Atlantic Ocean): may be a syn. of E.(Metechi- geographical origin is unknown norhynchus) petrotschenkoi (R) (fide ZOZITOWIECKl, -E. (Heruca) erinacei Rudolphi, 1795: syn. of Oliga- 1989) canthorhynchus erinacei (R.) Meyer, 1932 -E. gibber Olsson, 1894: syn of Corynosoma semerme -E. erinacei abdominalis Rudolphi, 1819: species inqui- (F.) Luhe, 1904 pro parte and of Corynosoma strumo- renda, may be an Oligacanthorhynchidae (fide sum (R.) Liihe, 1904 pro parte MEYER, 1932) -E. gibbosus Rudolphi, 1809: syn. of Corynosoma stru- -E. erinacei subcutaneus Rudolphi, 1819: may be a syn. mosum (R.) Luhe, 1904 pro parte of Oligacanthorhynchus (sensu lata) amphipachus -E. gigas (Goeze) Bloch, 1782: syn. of Macracantho- (W.) Meyer, 1932 rhynchus hirudinaceus (P.) Travassos, 1917 (nec -E. exiguus von Linstow, 1901: syn. of Telosentis exi- Block) guus (vL.) Van Cleave, 1923 -E. globicollis Creplin, 1829: syn. of Arhythmorhyn- -E. fabri Rudolphi, 1819: syn. of Acanthocephaloides chusfrassoni (M.) Luhe, 1911 propinquus (D.) Meyer, 1932 -E. globocaudatus Zeder, 1800: syn. of Centrorhynchus -E. falcatus Frolich, 1789: syn. of Acanthocephalus fal- globocaudatus (Z.) Luhe, 1911 catus (F.) Luhe, 1911 -E. globulosus Rudolphi, 1809: syn. of Acanthocepha- -E. falconis cyanei Rudolphi, 1819: syn. of Oligacan- Ius anguillae (O.F.M.) Luhe, 1911 thorhynchus lagenaeformis (W.) Travassos, 1917 -E. globulosus sensu Rudolphi, 1819: syn. of Acantho- -E. farionis Frol ich, 1802: syn. of Echinorhynchus cephaloides propinquus (D.) Meyer, 1932 (Metechinorhynchus) truttae Schrank, 1788 -E. globulosus sensu Leidy, 1856 syn. of Echinorhyn- -E. fasciatus Westrumb, 1821: species incertae sedis, chus (Metechinorhynchus) lateralis Leidy, 1851 may be a syn. of Centrorhynchus fasciatus (fide TRA- -E. globulosus sensu Linton, 1901: syn. of Paratenui- VASSOS, 1926) or of Mediorhynchus fasciatus (fide sentis ambiguus (VC.) Bullock et Samuel, 1975 MEYER, 1932) or of Prosthorhynchus cylindraceus -E. gobii Viborg, 1795: syn. of Pomphorhynchus laevis (G.) (fide GOLVAN, 1956) (O.F.M.) Van Cleave, 1924 -E. filicollis Rudolphi, 1809: syn. of Filicollis anatis -E. (Metechinorhynchus) gomesi Machado Filho, 1948 (S.) Luhe, 1911 (Brazil) -E. flavus Molin, 1861 (nec 1858): syn. of Acanthoce- -E. gracilis Rudolphi, 1819: syn. of Moniliformis graci- phaloides incrassatus (M.) Meyer, 1932 lis (R.) Meyer, 1931 -E. frassoni Molin, 1858: syn. of Arhythmorhynchus -E. gracilis sensu van Beneden, 1871: syn. of Neoechi- frassoni (M.) Liihe, 1911 norhynchus agilis (R.) Van Cleave, 1916 -E. fusiformis Zeder, 1803 (nec fusaeformis): syn. of -E. gracilis Machado Filho, 1948: preoccupied, syn. of Echinorhynchus (Metechinorhynchus) truttae Sch- Echinorhynchus (Metechinorhynchus) briconi Ma- rank, 1788 chado Filho, 1959 -E. gaboes McCallum, 1918: syn. of Pallisentis gaboes -E. grassii Deffke, 1891 (): syn. (?) of Monilifor- (MC.) Van Cleave, 1928 mis moniliformis (B.) Travassos, 1915 (fide TRAVAS- -E. (Echinorhynchus) gadi Zoega in O.F. Mtiller, 1776 sos, 1917) (type species) [North of the Atlantic Ocean but found -E. gruis Rudolphi, 1819: species delenda also in the Pacific Ocean (doubtful)] -E. gymnocyprii Liu Liqing, Wang Baodue et Yang -E. gadi sensu Van Cleave, 1924: syn. of Echinorhyn- Tao, 1981 (China) chus (Echinorhynchus) vancleavei Golvan, 1969 -E. haematopodis Rudolphi, 1819: syn. of Profilicollis -E. gadi sensu South well et Macfie, 1925 from Queens- sphaerocephalus (B.) Van Cleave, 1947 land is certainly another species -E. haeruca (Pall as, 1776) Rudolphi, 1802: syn. of -E. gadi sensu Ekbaum, 1938: syn. of Echinorhynchus Acanthocephalus ranae (S.) Luhe, 1911 pro parte (Echinorhynchus) ekbaumi Golvan, 1969 and of Acanthocephalusfalcatus (F.) Luhe, 1911 pro -E. gadi sensu Yamaguti, 1935: syn. of Echinorhyn- parte chus (Echinorhynchus) yamagutii Golvan, 1969 -E. haeruca de Lamarck, 1801 is not an acanthocephalan -E. gadi callariae Viborg, 1795: syn. of Echinorhyn- -E. haliaeti Rudolphi, 1819: syn. of Centrorhynchus chus (Echinorhynchus) gadi Zoega in O.F.M. 1776 buteonis (S.) Kostylew, 1914 -E. gadi virentis Rathke, 1799: syn. of Echinorhynchus -E. hamanni von Linstow, 1892: syn. of Corynosoma (Echinorhynchus) gadi Zoega in O.F.M. 1776 hamanni (vL.) Railliet et Henry, 1907 pro parte and of -E. galbulae Diesing, 1851: pro Echinorhynchus alce- Corynosoma antarcticum (R.) Johnston et Best, 1937 dinis galbulae Westrumb, 1821 and Echinorhynchus pro parte alcedinis Westrumb, 1821; species delenda -E. hamatus von Linstow, 1897: syn. (?) of Nephridia- -E. garzae (Gmelin, 1790) Zed er, 1803 is not an acan- canthus hamatus or of Oligacanthorhynchus hama- thocephalan (fide MEYER, 1932) tus (vL.) Schmidt, 1972 -E. gazae Gmelin, 1790 is not an acanthocephalan (fide -E. hamulatus Leidy, 1856: syn. of Neoechinorhynchus MEYER, 1932) emydis (L.) Van Cleave, 1919 150 Y.J. GOLVAN -E. hepaticola von Linstow, 1901: syn. of Echinorhyn- morhynchus longicollis (V.)Liihe, 1912 chus (Echinorhynchus) gadi O.F MUller, 1776 -E. (Metechinorhynchus)jucundus Travassos, 1923 -E. hepaticus Molin, 1858 et 1861: syn. of Sphaeriros- (Mato Grosso, Brazil) tris picae (R.) Go1van, 1956 -E. kerkoideus Westrumb, 1821: may be a syn. of Oliga- -E. heteracanthis Heitz, 1920: syn. of Bolbosoma hete- canthorhynchus erinacei (R.) Meyer, 1932 racanthis (H.) Meyer, 1932 -E. kostylewi Meyer, 1932: syn. of Acanthocephaloides -E. heteracanthus von Linstow, 1896: syn. of Hetero- propinquus (D.) Meyer, 1932 sentis heteracanthus (vL.) Van Cleave, 1931 -E. (Metechinorhynchus) kushiroensis Fujita, 1921 -E. heterorhynchus Parona, 1894: may be a Centrorhyn- (Japan) ch us sp. (fide MEYER, 1932) -E. labri Rudolphi, 1819: species delenda -E. hexacanthus Dujardin, 1845: may be a Polymorphi- -E. lacustris Zoega in O.F. MUller, 1776 is not an acan- nae (fide MEYER, 1932) thocephalan -E. hexagrammi Baeva, 1965 (East Siberia) -E. laevis Zoega in O.FMUller, 1776: syn. of Pompho- -E. hippuris Zeder, 1802 is not an acanthocephalan rhynchus laevis (Z.) Van Cleave, 1924 -E. hirudinaceus (pallas, 1781) O.F MUller, 1776: syn. -E. laevis sensu von Linstow, 1905: syn. of Filicollis of hirudinaceus (P.) Travas- anatis (S.) Luhe, 1911 sos, 1917 -E. lagaenaeformis Westrumb, 1821: syn. of Oligacan- -E. hirundinum Rudolphi, 1819: species delanda thorhynchus lagaenaeformis (W.) Travassos, 1917 -E. hominis Lambl, 1859 (nec Leuckart, 1876) is an Oli- pro parte and of Oligacanthorhynchus iheringi Tra- gacanthorhynchidae impossible to identify (fide vassos, 1916 pro parte MEYER, 1932) (may be Macracanthorhynchus) -E. lagenaeformis sensu Diesing, 1851: syn. of Oliga- -E. horridus von Linstow, 1897: syn. of Centrorhyn- canthorhynchus iheringi Travassos, 1917 ch us horridus (vL.) Meyer, 1932 -E. (Metechinorhynchus) lageniformis Ekbaum, 1938 -E. husonis Rudolphi, 1819: syn. of Leptorhynchoides (Pacific coast of North America) plagicephalus (W.) Kosty1ew, 1924 -E. lamelliger Diesing, 1854 (nec lamellifer): syn. of -E. hylae Johnston, 1912: syn. of Porrorchis hylae (1.) Serrasentis lamelliger (D.) Meyer, 1932 Schmidt et Kuntz, 1967 -E. lancea Westrumb, 1821: syn. of Sphaerirostris lan- -E. hystrix Bremser, 1824: syn. of Corynosoma semer- cea (W.) Golvan, 1956 me (F) Liihe, 1905 pro parte and of Corynosoma stru- -E. lanceolatus von Linstow, 1876: syn. of Plagiorhyn- mosum (R.) Luhe, 1904 pro parte ch us crassicollis (V.) Luhe, 1911 -E. hystrix sensu Leidy, 1887: syn. of Centrorhynchus -E. lari Rudolphi, 1819: syn. of Profilicollis sphaero- spinosus (K.) Van Cleave, 1916 pro parte and of Poly- cephalus (B.) Khokhlova, 1974 morphus obtusus Van Cleave, 1918 pro parte -E. (Metechinorhynchus) lateralis Leidy, 1851 (East of -E. idbari Gmelin, 1790: syn. of Pomphorhynchus lae- Canada) vis (Z.) Van Cleave, 1924 -E. lateralis Molin, 1858: preoccupied, syn. of Telosen- -E. impudicus Diesing, 1851: syn. of Paracavisoma tis molini Van Cleave, 1925 impudica (D.) von Kritscher, 1957 -E. lateralis sensu Leidy, 1851: species inquirenda -E. inaequalis Rudolphi, 1808 (nec inequalis): syn. of -E. (Echinorhynchus) laurentianus Ronald, 1957 (Gulf Centrorhynchus aluconis (O.FM.) Liihe, 1911 of St. Laurent River, Canada) -E. incrassatus Molin, 1858: syn. of Acanthocephaloi- -E. lavareti Rudolphi, 1809: syn. of Pomphorhynchus des incrassatus (M.) Meyer, 1932 laevis (O.F.M.) Van Cleave, 1924 -E. incrassatus sensu Linton, 1888: syn. of Dollfusentis -E. leguminosus Solowjew, 1912: syn. of Sphaerirostris chandleri Golvan, 1969 or of a Corynosoma sp. leguminosus (S.) Golvan, 1956 -E. incrassatus sensu Linton, 1891 is a larva of Coryno- -E. (Metechinorhynchus) leidyi Van Cleave, 1924 soma sp. (vide VAN CLEAVE,1953) (Great Lakes) -E. indicus Chandra, Rekha, Rao et Shyamasundari, -E. lendix (Phipps, 1774) Luhe, 1911: syn. of Polymor- 1982 (India) ph us phippsi Kostylew, 1922 -E. inflatus Rudolphi, 1809: syn. of Echinorhynchus -E. (Pseudoechinorhynchus) lenok Achmerov et Dom- (Metechinorhynchus) salmonis O.F MUlier, 1784 browskaja-Achmerova, 1941 (Eastern Siberia): may be -E. inflatus Creplin, 1829: preoccupied, syn. of Pla- a syn. of Echinorhynchus parasiluri Fukui, 1929 (fide giorhynchus crassicollis (V.) Luhe, 1911 GRABDA-KAZUBSKA& EJSYMONT,1969) -E. inflexus Cobbold, 1861: species delenda -E. lesiniformis Molin, 1859: syn. of Centrorhynchus -E. ingens von Linstow, 1879: syn. of Macracantho- lesiniformis (M.) Meyer, 1932 rhynchus ingens (vL.) Meyer, 1932 -E. linearis Westrumb, 1821 (nec 1891): syn. of Pla- -E. inscriptus Westrumb, 1821: syn. of Lueheia inscrip- giorhynchus linearis (W.) Golvan, 1956 ta (W.) Travassos, 1926 -E. lineolatus O.F. MUller, 1777: syn. of Echinorhyn- -E. invaginabilis von Linstow, 1902: syn. of Arhyth- chus (Echinorhynchus) gadi O.F MUller, 1776 morhynchus invaginabilis (vL.)Luhe, 1912 -E. linstowi Hamann, 1891: syn. of Acanthocephalus -E. invaginabile de Marval, 1905: syn. of Arhyth- anguillae (O.FM.) Liihe, 1911 Nomenclature of the Acanthocephala 151 -E. lobianchii Monticelli, 1887: syn. of Centrorhynchus -E. microcephalus Rudolphi, 1819: syn. of Hamannie- lobianchii (M.) Meyer, 1932 or of Sphaerirostris lla microcephala (R.) Travassos, 1915 lobianchii (M.) Golvan, 1958 -E. microcephalus sensu Leidy, 1856: syn. of Haman- -Ei longicollis (Pallas, 1781) Goeze, 1782: syn. of niella tortuosa (L.) Van Cleave, 1924 Pomphorhynchus tereticollis (R.) Meyer, 1932 -E. miliarius Zeder, 1832: syn. of Polymorphus minu- -E. longicollis Villot, 1875: syn. of Arhythmorhynchus tus (G.)Ltihe, 1911 or of Polymorphus boschadis (S.) longicollis (V.) Liihe, 1912 [may be considered as pre- Railliet, 1919 occupied by Echinorhynchus longicollis (P.) Goeze, -E. miniatus von Linstow, 1896: syn. of Polymorphus 1782 and, in this case, must be a syn of Arhythmorhyn- miniatus (vL.) Travassos, 1926 ch us villoti nom. nov.] -E. minutus Goeze, 1782: syn. of Polymorphus minutus -E. longicollis sensu Baylis, 1922 is another species (G.) LUhe, 1911 -E. longiproboscis Rodyuk, 1986 (Southwestern Atlan- -E. minutus sensu Zeder, 1800: syn. of Polymorphus tic Ocean, Falklands) boschadis (S.) Railliet,1919 -E. lophii Gmelin, 1791: syn. of Echinorhynchus -E. mirabilis Polonio, 1859: syn. of Centrorhynchus (Echinorhynchus) gadi O.F. MUlier, 1776 aluconis (O.FM.)Luhe, 1911 -E. (Echinorhynchus) lotellae Yamaguti, 1939 (Sizuo- -E. monechinus: errat. pro Echinorhynchus monoechi- ka, Japan) nus von Linstow, 1902 -E. lucii O.F. MUller, 1776: syn. of Acanthocephalus -E. moniliformis Bremser, 1811: syn. of Moniliformis lucii (O.F .M.)Liihe, 1911 moniliformis (B.) Travassos, 1915 -E. lutzi von Linstow, 1891: syn. of Acanthocephalus -E. moniliformis sensu Leidy collect.: syn. of Monilifor- lutzi (vL.) Meyer, 1932 mis clarki (W.) Van cleave, 1924 -E. mabuiae von Linstow, 1908: syn. of Centrorhyn- -E. monoechinus von Linstow, 1902: syn. of Monilifor- chus mabuiae (vL.) Golvan, 1956 mis monoechinus (vL.) Petrotschenko, 1958 -E. macracanthus Bremser in Westrumb, 1821: syn. of -E. monticellii Porta, 1904 (nec monticelli): species Oligacanthorhynchus ricinoides (R.) Travassos, 1915 impossible to be determined, may be a syn. of Echi- -E. macracanthus sensu de Marval, 1902: syn. of Oliga- norhynchus (Pseudoechinorhynchus) monticellii canthorhynchus compress us (R.) Meyer, 1932 Petrotschenko, 1956 (fide PETROTSCHENKO1956), -E. macrorhynchus Diesing, 1851 et 1856: syn. of Poly- -E. morinelli Rudolphi, 1819: nomen nudum, may be a acanthorhynchus macrorhynchus (D.)Travassos, syn. of Sphaerirostris lancea (W.) Golvan, 1956 1920 -E. motacillae atricapillae Rudolphi, 1819: may be a -E. macrourus Bremser in Westrumb, 1821: species syn. of Centrorhynchus fasciatus (W.) Travassos, inquirenda, probably a syn. of Arhythmorhynchus 1926 frassoni (M.) Luhe, 1911 or of Arhythmorhynchus -E. muraenae Bosc, 1802: may be a syn. of Echi- longicollis (V.)Liihe, 1912 norhynchus (Metechinorhynchus) salmonis O.F -E. magretti Parona, 1885 may be an Oligacanthorhyn- MUller, 1784 (fide MEYER, 1932), certainly not (fide chidae (fide MEYER, 1932) GOLVAN,1950) -E. major Bremser, 1811: syn. of Nephridiacanthus -E. (Metechinorhynchus) muraenolepisi (Rodyuk, major (B.) Golvan, 1962 1984) novo comb. (South Georgia): syn. of Metechi- -E. manifestus Leidy, 1851: syn. of Oligacanthorhyn- norhynchus muraenolepisi Rodyuk, 1984 ch us manifestus (L.) Van Cleave, 1924 -E. muris O.F MUller, 1787 is not an acanthocephalan -E. maraenae Gmelin, 1791: syn. of Echinorhynchus -E. muris Schrank, 1792 is not an acanthocephalan (Metechinorhynchus) salmonis O.F MUller, 1784 -E. muscicapae Rudolphi, 1819: syn. of Prosthorhyn- -E. medius Linton, 1908: syn. of Gorgorhynchus chus cylindraceus (G., 1782 in S., 1788) Golvan, 1956 medius (L.)Chandler, 1934 pro parte and of Centrorhynchus dimorphocephalus -E. megacephalus Westrumb, 1821 (nec megalocepha- (W.) Meyer, 1932 pro parte Ius): syn. of Centrorhynchus tumidulus (R.) Neiva da -E. mustelae Rudolphi, 1819: syn. of Oligacanthorhyn- Cunha et Travassos, 1914 ch us erinacei (R.) Meyer, 1932 -E. megarhynchus von Linstow, 1892: syn. of Aspersen- -E. mutabilis Rudolphi, 1819: syn. of Hexaglandula tis megarhynchus (vL.) Golvan, 1960 mutabilis (R.) Petrotschenko, 1950 pro parte and of -E. (Echinorhynchus) melanoglaeae Dollfus, 1960 Polymorphus striatus (G.) Liihe, 1911 pro parte (Senegal) -E. myoxi Galli-Valerio, 1929: syn. of Moniliformis -E. menurae Johnston, 1913: syn. of Plagiorhynchus myoxi (G.-V.) Meyer, 1932 (??) menurae (1.) Golvan, 1956 -E. napaeformis Rudolphi, 1802: syn. of Oligacan- -E. mergi Bloch, 1782 in Schrank, 1788: syn. of Cen- thorhynchus erinacei (R.) Meyer, 1932 trorhynchus aluconis (O.FM.) Liihe, 1911 -E. nardoi Molin, 1859 may be a Rhadinorhynchidae -E. merulae Gmelin, 1789 et 1791: syn. of Sphaeriros- (fide MEYER, 1932) tris lancea (W.) Golvan, 1956 -E. ninnii Stossich, 1891: syn. of Centrorhynchus nin- -E. micracanthus Rudolphi, 1819: syn. of Mediorhyn- nii (S.) Meyer, 1932 ch us micracanthus (R.) Van Cleave, 1927 -E. nitzschi Giebel, 1866: species incertae sedis 152 Y.J. GOLVAN -E. nodulosus Schrank, 1790 et 1792: syn. of Pompho- -E. pachyacanthus Sonsino, 1889 (nec 1899): syn. of rhynchus laevis (O.F.M.) Van Cleave, 1924 Prosthenorchis pachyacanthus (S.) Travassos, 1917 -E. nototheniae Zdzitowiecki, 1986 (South Shetlands, -E. pachysomus Creplin, 1829: syn. of Echinorhynchus Antarctic): may be a syn. of E. (Metechinorhynchus) (Metechinorhynchus) salmonis O.F. MUller, 1784 petrotschenkoi (R) (jide ZDZITOWIECKI,1989) -Echinorhynchus (Metechinorhynchus) paranensis -E. novellae Parona, 1890 (nec novellai): syn. of Neon- Machado Filho, 1959 (Mato Grosso, Brazil) cicola novellae (P.) Schmidt, 1972 -Echinorhynchus (Pseudoechinorhynchus) parasiluri -E. nudus (Harada, 1938) Petrotschenko, 1956: syn. of Fukui, 1929 (Japan) Neorhadinorhynchus nudus (H.) Yamaguti, 1939 -E. pardalis Westrumb, 1821: syn. of Oligacanthorhyn- -E. nycteae Schrank, 1788: syn. of Centrorhynchus alu- chus pardalis (W.) Schmidt, 1972 conis (O.F.M.) Luhe, 1911 or of Centrorhynchus bute- -E. pardalis sensu Travassos, 1917: syn. of Oncicola onis (S.) Kostylew, 1914 macrurae Meyer, 1931 -E. obesus von Linstow, 1902: syn. of Mediorhynchus -E. pardi Huxley in von Ihering, 1902: species delenda emberizae (R.) Travassos, 1924 -E. pari Rudolphi, 1819: species delenda -E. obliquus Dujardin, 1845: syn. of Prosthorhynchus -E. paronai Condorelli, 1897: syn. of Acanthocephalus cylindraceus (G. in S.) Golvan, 1956 paronai (C.) Meyer, 1932 -E. oblitus Golvan, 1969 (Japan) -E. parvus Furhmann in Wolffhiigel, 1900: nomen -E. obovatus Olsson, 1893: syn.(?) of Corynosoma nudum, syn. of Prosthorhynchus cylindraceus (G. in semerme (F.) Liihe, 1905 S.) Golvan, 1956 -E. obovatus Rudolphi in litteris: syn. of Polymorphus -E. patani Shipley, 1903: may be an Oligacanthorhyn- piriforme (B.) Machalska, 1981 chus (jide YAMAGUTI,1963) -E. odontaspidis Linton, 1891: nomen nudum -E. paucihamatus Leidy, 1891 (nec paucihamatum): -E. oedicnemi Rudolphi, 1819: syn. of Sphaerirostris may be a syn. of Neoechinorhynchus cylindratus lancea (W.) Golvan, 1956; often considered as a nomen (VC.) Van Cleave, 1919 nudum -E. peleci Grimm, 1870: Acanthocephala incertae sedis -E. oedicnemonis Rudolphi, 1819: vide E. oedicnemi -E. pellucidus Leuckart, 1828: syn. of Bolbosoma vas- -E. oligacanthoides Rudolphi, 1819: syn. of Oligacan- culosum (R.) Meyer, 1932 (vide VANCLEAVE,1953) thorhynchus spira (D.)Travassos, 1915 pro parte and -E. peposacae Porta, 1914: syn. of Corynosoma peposa- of Oligacanthorhynchus taenioides (D.) Travassos, cae (P.) Travassos, 1926 1917 pro parte -E. percae Gemlin, 1791: syn. of Acanthocephalus lucii -E. oligacanthus Rudolphi, 1819: syn. of Oligacan- (O.F.M.) Liihe, 1911 thorhynchus oligacanthus (R.) Meyer, 1932 -E. perforator von Linstow, 1908: syn. of Pomphorhyn- -E. oncicola von Ihering, 1902: syn. of Oncicola onci- chus perforator (vL.) Meyer, 1932 cola (vI.) Travassos, 1916 -E. (Metechinorhynchus) petrotschenkoi (Rodyuk, -E. (sensu lato) orestiae Neveu-Lemaire, 1905 (Titicaca 1984)Zdzitowiecki, 1989 (Antarctic) Lake, ) -E. phocae Gmelin, 1791 is not an acanthocephalan -E. oricola von Linstow, 1901: syn. of Leptorhynchoi- -E. phoenix Schneider, 1903: syn. of Echinorhynchus des thecatus (vL.) Kostylew, 1924 (Metechinorhynchus) salmonis O.F. Miiller, 1784 -E. (Echinorhynchus) orientalis Kaw, 1951 (Cachernire) -E. physocoracis Porta, 1913: syn. of Sphaerirostris -E. orioli Rudolphi, 1819: syn. of Sphaerirostris areola- physocoracis (P.) Golvan, 1956 tus (R.) Golvan, 1956 -E. picae Rudolphi, 1819: syn. of Sphaerirostris teres -E. orioli Westrumb, 1821: syn. of Mediorhynchus (W.) Golvan, 1956 or of Sphaerirostris picae (R.) Go1- emberizae (R.) Travassos, 1924 van, 1956 -E. orioli cristati Rudolphi, 1819: syn. of Mediorhyn- -E. pici Gmelin, 1791: syn. of Prosthorhynchus cylin- ch us emberizae (R.) Travassos, 1924 draceus (G. in S.) Golvan, 1956 -E. otidis Schrank, 1788: syn. of Centrorhynchus alu- -E. pici collaris Leidy, 1850: syn. of Oligacanthorhyn- conis (O.F.M.) Luhe, 1911 ch us manifestus (L.) Van Cleave, 1924 -E. otidis Miescher, 1841 in de Marval, 1904: syn. of -E. pigmentatus Fuhrmann in Wolffhiigel, 1900: nomen Mediorhynchus otidis (M.) Van Cleave, 1947 nudum -E. otidis houbarae Miescher, 1841 in Diesing, 1851: -E. pigmentatus de Marval, 1902: syn. of Prosthorhyn- syn. of Mediorhynchus taeniatus (vL.) Dollfus, 1936 chus cylindraceus (G. in S.) Golvan, 1956 or of Pros- -E. ovatus Zeder, 1800: syn. of Pomphorhynchus laevis thorhynchus pigmentatus (dM.) Meyer, 1932 (fide (O.F.M.) Van Cleave, 1924 MEYER, 1932) -E. ovatus sensu Leidy, 1850: syn. of Oncicola campa- -E. piriformis Bremser, 1811 in Rudolphi, 1819: syn. of nulata (D.) Meyer, 1931 or of Echinopardalis pardalis Polymorphus piriformis (B.) Machalska, 1981 (W.) Travassos, 1918 -E. piscinus Zeder, 1800: syn. of Pomphorhynchus -E. ovicola: errat. pro E. oricola von Linstow, 1901 tereticollis (R.) Meyer, 1932 -E. ovocristatus von Linstow, 1897: syn. of Promonili- -E. plagicephalus Westrumb, 1821: syn. of Leptorhyn- formis ovocristatus (vL.) Dollfus et Golvan, 1963 choides plagicephalus (W.) Kostylew, 1924 Nomenclature of the Acanthocephala 153 -E. platessae Rudolphi, 1809: syn. of Acanthocephalus -E. pupa von Linstow, 1905: syn. of Polymorphus pupa lucii (O.F.M.) Liihe, 1911 (vL.) (fide KOSTYLEW, 1922), syn. of Filicollis pupa -E. platessoida Bosc, 1820: is not an acanthocephalan (vL.) (fide TRAYASSOS, 1926); syn. of Prosthorhyn- -E. platessoidae Gmelin, 1791: is not an acanthocepha- chus pupa (vL.) (fide MEYER, 1931)(in this case it lan may rather be a Plagiorhynchus pupa novo comb. -E. platessoides Gmelin, 1790 (from Greenland): impos- because the life cycle is probably aquatic); it is impos- sible to identify sible to identify it (fide VAN CLEAYE& RAUSCH,1951) -E. platessoides maximi Rudolphi, 1809: impossible to -E. putorii Molin, 1858 is an Oligacanthorhynchus identify sensu lato -E. pleuronectis Gmelin, 1791: impossible to identify -E. pyriformis Bremser, 1811 in Westrumb, 1821: vide -E. pleuronectis maximi Rudolphi, 1809: impossible to Polymorphus piriformis (B.) Machalska, 1981 identify -E. quadrirostris Goeze, 1782 is a Cestode Tetrarhyncha -E. pleuronectis platessoides Viborg, 1795 is not an -E. ranae Schrank, 1788: syn. of Acanthocephalus acanthocephalan ranae (S.) Liihe, 1911 -E. plicatus von Linstow, 1883: syn. of Arhythmorhyn- -E. ranae sensu Rudolphi, 1803: syn. of Acanthocepha- ch us plicatus (vL.) Meyer, 1932 Ius lucii (O.F.M.) Luhe, 1911 -E. poliacanthoides in Condorelli, 1892: vide E. polya- -E. ranae bombynae Schrank, 1793: may be a syn. of canthoides Acanthocephalus ranae (S.) Luhe, 1911 -E. polyacanthoides Creplin, 1825: syn. of Centrorhyn- -E. raphaelis Sonsino, 1896: species delenda chus aluconis (O.F.M.) Luhe, 1911 -E. rectus Linton, 1892: syn. of Plagiorhynchus rectus -E. polyacanthus Creplin, 1825: syn. of Centrorhyn- (L.) Van Cleave, 1918 chus buteonis (G. in S.) Kostylew, 1914 -E. reductus von Linstow, 1905: syn. of Corynosoma -E. polymorphus Bremser, 1824: syn. of Polymorphus reductum (vL.) Railliet et Henry, 1907 minutus (G.) Luhe, 1911 pro parte and of Filicollis -E. rennicki Leiper et Atkinson, 1914: syn. of Metacan- anatis (S.) Liihe, 1911 pro parte thocephalus rennicki (L. et A.) Zdzitowiecki, 1983 -E. pomatostomi Johnston et Cleland, 1911: syn. of -E. reticulatus Westrumb, 1821: syn. of Plagiorhynchus Oncicola pomatostomi (1. et C.) Schmidt, 1983 reticulatus (W.) Golvan, 1956 -E. porrigens Rudolphi, 1814: syn. of Bolbosoma balae- -E. rheae Fuhrmann in Wolffhligel, 1900: nomen nudum nae (G.) Meyer, 1932 -E. rheae de Marval, 1902: syn. of Prosthorhynchus -E. porrigens sensu·Kaiser, 1893: syn. of Bolbosoma rheae (W.) Golvan,1966 brevicolle (M.) Porta, 1908 -Echinorhynchus (Metechinorhynchus) rhenanus -E. praetextus Molin, 1858: may be a syn. of Acantho- (Golvan, 1969) Amin, 1985 (Rhine River Basin) cephalus ranae (S.) Liihe, 1911 (fide PORTA, 1908) -E. rhopalorhynchus Diesing, 1851: syn. of Polyacan- -E. pristis Rudolphi, 1802: syn. of Rhadinorhynchus thorhynchus rhopalorhynchus (D.) Travassos, 1920 pristis (R.) Liihe, 1911 -E. rhytidodes Monticelli, 1905: syn. of Solearhynchus -E. pristis sensu Linton, 1891: syn. of Dollfusentis soleae (P.) de Buron et Maillard, 1985 chandleri Golvan, 1969 -E. ricinoides Rudolphi, 1808: syn. of Oligacantho- -E. pristis sensu von Linstow, 1901 is not a Rhadino- rhynchus ricinoides (R.) Travassos, 1915 rhynchus but a Echinorhynchus gadi O.F. Miiller, -E. robustus Datta, 1928: syn. of Sphaerirostris robus- 1776 (fide KOSTYLEW,1925) tus (D.) novo comb. -E. pristis tenuicornis Linton, 1905 et 1908: syn. of -E. rosai Porta, 1910: syn. of Prosthorhynchus cylin- Dollfusentis chandleri Golvan, 1969 draceus (G. in S.) Golvan, 1956 -E. propinquus Dujardin, 1845: syn. of Acanthocepha- -E. roseus Molin, 1858: syn. of Arhythmorhynchus loides propinquus (D.) Meyer, 1932 roseus (M.) Meyer, 1932 -E. propinquus sensu Miihling, 1898: syn. of Acantho- -E. roseus Van Beneden, 1871: species delenda cephalus anguillae (O.F.M.) Liihe, 1911 -E. rostratus Fuhrmann in Wolffhiigel, 1900: nomen -E. propinquus sensu Kostylew, 1926: syn. of Acantho- nudum cephaloides kostylewi Meyer, 1932 -E. rostratus de Marval, 1902: syn. of Prosthorhynchus -E. proteus Westrumb, 1821: syn. of Pomphorhynchus cylindraceus (G. in S.) Golvan, 1956 laevis (O.F.M.) Van Cleave, 1924 -E. rotundatus von Linstow, 1897: syn. of Porrorchis -E. proteus sensu Linton, 1851, 1856, 1888 et 1904: syn. rotundatus (vL.) Schmidt et Kuntz, 1967 of Pomphorhynchus bulbocolli Linkins, 1919 -E. rotundocapitatus Johnston, 1912: syn. of Sphaere- -E. proteus sensu Porta, 1905: syn. of Acanthocephalus chinorhynchus rotundocapitatus (1.) Johnston et anguillae (O.F.M.) Liihe, 1911 Deland, 1929 -E. pseudosegmentatus Knlipffer, 1888: syn. of Monili- -E. ruber Collett, 1886: syn. of Bolbosoma turbinella formis (sensu lato) pseudosegmentatus (K.) Meyer, (D.) Porta, 1908 1932 -E. rubetrae Rudolphi, 1819: syn. of Centrorhynchus -E. pumilio Rudolphi, 1819: syn. of Acanthocephaloi- fasciatus (W.) Meyer, 1932 (fide TRAYASSOS,1926) or des propinquus (D.) Meyer, 1932 of Mediorhynchusfasciatus (W.) (fide MEYER, 1932) 154 Y.J. GOLVAN -E. rubieundus Molin, 1859 in Stossich, 1890: syn. of -E. soeialis Leidy, 1856 nee 1851: syn. of Echinorhyn- Arhythmorhynchus rubicundus (M.) Meyer, 1932 chus (Echinorhynchus) vancleavei Golvan, 1969 -E. rutheni Rudolphi in Westrumb, 1821: syn. of Lep- -E. soleae Porta, 1905: syn. of Solearhynchus soleae torhynchoides plagicephalus (W.) Kostylew, 1924 (P.) de Buron et Maillard, 1985 -E. rutili O.F. Miiller, 1780: syn. of Neoechinorhyn- -E. solitarius Molin, 1858 (nee solitarium): syn. of Poly- ch us rutili (O.F.M.) Hamann, 1892 morphus sp. (fide MEYER, 1932) -E. rutili sensu Zeder, 1800: syn. of Acanthocephalus -E. sorieis Rudolphi, 1819: syn. of Centrorhynchus alu- anguillae (O.F.M.) Liihe, 1911 conis (O.F.M.) Liihe, 1911 -E. rythidodes Monticelli, 1905: vide E. rhytidodes -E. sp. Perroncito, 1882: syn. of Centrorhynchus ninnii -E. sagittifer Linton, 1889: syn. of Serrasentis sagittifer (S.) Meyer, 1932 (L.) Linton, 1932 -E. sp. Katoaka et Momma, 1937: syn. of Rhadinorhyn- -E. (Metechinorhynchus) salmonis O.F.Miiller, 1784 ch us oblitus Golvan, 1969 (North and West of Europe) -E. sp. Kagei et Watanuki, 1975: may be a syn. of E. -E. salmon is sensu Wang, 1980 may be another species (Metechinorhynchus) petrotschenkoi (R) (fide ZDZI- (China) TOWIECKJ,1989) -E. salmonorum Rudolphi, 1819: syn. of Pomphorhyn- -E. sphaerieus Rudolphi, 1810: syn. of Pomphorhyn- chus laevis (O.F.M.) Van Cleave, 1924 or of Pompho- ch us laevis (O.F.M.) Van Cleave, 1924 rhynchus tereticollis (R.) Meyer, 1932 -E. sphaeroeephalus Bremser, 1811 in Rudolphi, 1819: -E. salmonum Rudolphi, 1819: vide E. salmonorum syn. of Profilicollis sphaerocephalus (B.) Khokhlova, -E. (Metechinorhynchus) salobrensis Machado Filho, 1974 1948 (Mato Grosso, Brazil) -E. sphyrnae Rudolphi, 1809 is not an acanthocephalan -E. salvelini Schrank, 1788: syn. of Pomphorhynchus -E. spinosus Kaiser, 1893: syn. of Centrorhynchus spi- laevis (O.F.M.) Van Cleave, 1924 nosus (K.) Van Cleave, 1916 -E. salvelini Linkins in H.B. Ward et Whipple, 1918: -E. spira Diesing, 1851: syn. of Oligacanthorhynchus syn. of E. (Metechinorhynchus) leidyi Van Cleave, spira (D.) Travassos, 1915 1924 -E. spiralis Rudolphi, 1809: syn. of Plagiorhynchus spi- -E. seanensis Lundstrom, 1942: probably a syn. of ralis (R.) Golvan, 1956 Sphaerirostris lancea (W.) Golvan, 1956 -E. spirula von Olfers in Rudolphi, 1819: syn. of Onci- -E. seiaenae Rudolphi, 1819: syn. of Pomphorhynchus cola spirula (R.) Schmidt, 1972 pro parte and of Pros- tereticollis (R.) Meyer, 1932 thenorchis luehei Travassos, 1916 pro parte -E. seopis Gmelin, 1790: syn. of Centrorhynchus alu- -E. spirula sensu von Linstow, 1878 et 1879: syn. of conis (O.F.M.) Liihe, 1911 or of Centrorhynchus bute- Oncicola luehei (T.) Schrnidt, 1972 onis (S.) Kostylew, 1914 -E. stellaris Molin, 1858: syn. of Filicollis anatis (S.) -E. seorpaenae Rudolphi, 1819: syn. of Acanthocepha- Liihe, 1911 Ius anguillae (O.F.M.) Luhe, 1911 or of Acanthoce- -E. sternae Rudolphi, 1819: nomen nudum; may be a phaloides propinquus (D.) Meyer, 1932 syn. of Plagiorhynchus linearis (W.) Golvan, 1956 -E. sebastolobi Kovalenko, 1986 (Okhotsk Sea) -E. striatus Goeze 1782: syn. of Polymorphus striatus -E. segmentatus de Marval, 1902: is not a syn. of (G.) Liihe, 1911 Mediorhynchus taeniatus (vL.) (vide DOLLFUS,1936) -E. striatus Villot, 1875: syn. of Corynosoma strumo- -E. semermis Forssell, 1904: syn. of Corynosoma sum (R.)Liihe, 1904 semerme (F.)Liihe, 1904 -E. striatus sensu Linton, 1892: syn. of Corynosoma -E. semoni von Linstow, 1898: syn. of Australiformis constrictum Van Cleave, 1918 semoni (vL.) Schrnidt et Edmonds, 1989 -E. striatus sensu Leidy, 1856: syn. of Polymorphus -E. serpentieola von Linstow, 1908: syn. of Sphaeriros- crassus Van Cleave, 1924 pro parte and of Polymor- tris serpenticola (vL.) Golvan, 1956 ph us obesus Van Cleave, 1918 pro parte -E. serpentulus Grimm, 1870: impossible to indentify -E. stridulae Goeze, 1782: syn. of Centrorhynchus alu- -E. serrani Linton, 1888 et 1891: juvenile, may be a syn. conis (O.F.M.) Liihe, 1911 or of Centrorhynchus bute- of Bolbosoma vasculosum (R.) Meyer, 1932 (vide onis (G. in S.) Kostylew, 1914 VAN CLEAVE,1953) -E. strigis Gmelin, 1782: syn. of Centrorhynchus alu- -E. (Metechinorhynchus) sevani Dinnik, 1933 (nee conis (O.F.M.) Luhe, 1911 pro parte and of Centro- sevangi) (Armenia) rhynchus buteonis (G. in S.) Kostylew, 1914 pro parte -E. sigmoideus Westrumb, 1821: syn. of Sphaerirostris -E. strigis auricularis Goeze, 1782: syn. of Centrorhyn- areolatus (R.) Golvan, 1956 ch us aluconis (O.F.M.) Luhe, 1911 pro parte and of -E. simplex Rudo1phi, 1810: syn. of Euzetacanthus sim- Centrorhynchus buteonis (G. in S.) Kostylew, 1914 plex (R.) Golvan et Houin, 1964 pro parte -E. sipuneulus Schrank, 1788: syn. of Echinorhynchus -E. strumosus Rudolphi, 1802: syn. of Corynosoma (Metechinorhynchus) salmonis O.F. Miiller, 1784 strumosum (R.) Liihe, 1904 pro parte and of Coryno- -E. soeialis Leidy, 1851: syn. of Serrasentis sagittifer soma semerme (F.)Liihe, 1904 pro parte (L.) Linton, 1932 -E. sturionis Goeze, 1782: impossible to identify Nomenclature of the Acanthocephala 155 -E. subglobatus Zed er, 1800: syn. of Pomphorhynchus -E. triglae gurnardi Rathke, 1799: syn. Euzetacanthus laevis (O.F.M.) Van Cleave, 1924 simplex (R.) Golvan et Houin, 1964 -E. sublobatus Gmelin, 1782: syn. of Pomphorhynchus -E. tritonis Westrumb, 1821: syn. of Acanthocephalus laevis (O.F.M.) Van Cleave, 1924 falcatus (F.) Liihe, 1911 -E. subulatus Zed er, 1803: syn. of Rhadinorhynchus -E. trutta (sic) Southwell et Macfie, 1925: syn. of alosae (H.) Meyer, 1932 pro parte and of Rhadi- Acanthocephalus hastae Baylis, 1944 norhynchus pristis (R.) Liihe, 1911 pro parte -E. (Metechinorhynchus) truttae Schrank, 1788 (Euro- -E. sylviarum Rudolphi, 1819: syn. of Echinorhynchus pe) (sensu lato)fasciatus Westrumb, 1821 (Mediorhyn- -E. truttae sensu Zhukov, 1960, from Kurile Islands, ehus ??) may be another species -E. taeniaeformis von Linstow, 1890 (nee taeniaefor- -E. tuba Rudolphi, 1802: syn. of Centrorhynchus alu- me): may be a Polyacanthorhynchus (fide TRA- conis (O.F.M.) Liihe , 1911 pro parte and of Cen- VASSOS, 1920) or a Rhadinorhynchus (fide MEYER, trorhynchus globocaudatus (Z.) Liihe, 1911 pro parte 1932) -E. tuba sensu Creplin, 1848: syn. of Centrorhynchus -E. taeniatus von Linstow, 1901: syn. of Mediorhyn- aluconis (O.F.M.) Liihe, 1911 chus taeniatus (vL.) Dollfus, 1936 -E. tuberosus Zeder, 1800 and 1803: syn. of Neoechi- -E. taeniforme von Linstow, 1890: may be a Polyacan- norhynchus rutili (O.F.M.) Hamann, 1892 thorhynchus (fide YAMAGUTI,1963) -E. tuberosus sensu Linton, 1893: may be a syn. of Neo- -E. taenioides Diesing, 1851: syn. of Oligacanthorhyn- echinorhynchus crassus Van Cleave, 1919 (fide eh us taenioides (D.) Travassos, 1915 LYNCH, 1936) -E. tanagrae Rudolphi, 1819: species delenda -E. tuberosus sensu Leidy collect.: syn. of Neoechi- -E. tardae Rudolphi, 1809: nomen nudum (fide DOLL- norhynchus cylindratus (VC.) Van Cleave, 1919 FUS, 1936 is a species delenda) -E. tubifer Gmelin, 1782 is not an acanthocephalan -E. tenuieaudatus Marotel, 1889: syn. of Sphaerirostris -E. tumeseens von Linstow, 1896: syn. of Acanthoce- tenuicaudatus (M.) Amin, 1985 phalus (?) tumescens (vL.) Porta, 1905 -E. tenuieollis Frol ich, 1802: syn. of Polymorphus -E. tumidulus Rudolphi, 1819: syn. of Centrorhynchus minutus (G.) Luhe, 1911 pro parte and of Filicollis sp. tumidulus (R.) Neiva da Cunha et Travassos, 1914 pro parte (fide MEYER, 1932) -E. turbinella Diesing, 1851: syn. of Bolbosoma turbi- -E. tenuieollis Westrumb, 1821: syn. of Pomphorhyn- nella (D.) Porta, 1908 chus laevis (O.F.M.) Van Cleave, 1924 -E. uneinatus Kaiser, 1893: syn. of Arhythmorhynchus -E. tenuieornis Linton, 1891: syn. of Telosentis tenui- uncinatus (K.) Luhe, 1911 cornis (L.) Van Cleave, 1947 -E. undulata Giebel, 1866: vide E. undulatus -E. terebra Rudolphi, 1819: syn. of Raorhynchus tere- -E. undulatus Nitzsch in Giebel, 1857: syn. of Cen- bra (R.) Tripathi, 1959 trorhynchus undulatus (N.) Dollfus, 1951 -E. teres Westrumb, 1821: syn. of Sphaerirostris teres -E. ungriai Gracia-Rodrigo, 1960: syn. of Megapriapus (R.) Golvan, 1956 or of Sphaerirostris picae (R.) Gol- ungriai (GR.) Golvan, Gracia-Rodrigo et Dfaz-Ungrfa, van, 1956 1964 -E. teretieollis Rudolphi, 1810: syn. of Pomphorhyn- -E. upupae L6pez-Neyra, 1946: syn. of Prosthorhyn- ch us tereticollis (R.) Meyer, 1932 chus cylindraceus (G. in S.) Golvan, 1956 -E. theeatus Linton, 1891: syn. of Leptorhynchoides -E. urniger Dujardin, 1945: species ineertae sedis thecatus (L.) Kostylew, 1924 -E. urniger sensu Porta, 1905: syn. of Euzetacanthus -E. thymalli Schrank, 1803: syn. of Pomphorhynchus simplex (R.) Golvan et Houin, 1964 laevis (O.F.M.) Van Cleave, 1924 -E. uromastieis Fraipont, 1882 (nee uromastieus): syn. -E. tigrinae Shipley, 1903: syn. of Acanthocephalus (?) of Oligacanthorhynchus spira (D.) Travassos, 1915 tigrinae (S.) Yamaguti, 1963 -E. vaginatus Diesing, 1851 (nee vaginatum): syn. of -E. torquatus Frolich, 1802: syn. of Filicollis anatis (S.) Mediorhynchus vaginatus (D.) Meyer, 1932 Liihe, 1911 -E. (Echinorhynchus) vancleavei Golvan, 1969 (nor- -E. tortuosus Leidy, 1850 (nee tortuosa): syn. of Ha- thern of the Atlantic coast of USA): is not a syn. of manniella tortuosa (L.) Van Cleave, 1924 Echinorhynchus gadi Zoega, 1776 -E. totani Porta, 1910: syn. of Plagiorhynchus totani -E. vanelli Goeze, 1782: nomen nudum (P.) Golvan, 1956 -E. vanelli Gmelin, 1790 et 1791: syn. of Sphaerirostris -E. transversus Rudolphi, 1819: syn. of Prosthorhyn- lancea (W.) Golvan, 1956 ch us cylindraceus (G. in S.) Golvan, 1956 -E. variabilis Diesing, 1851: syn. of Octospinifer varia- -E. trichiuri Holton, 1802 is not an acanthocephalan bi/is (D.) von Kritscher, 1976 -E. triehoeephalus Leuckart, 1876 in Kaiser, 1893: syn. -E. variabilis Diesing, 1856: syn. of Gracilisentis varia- of Arhythmorhynchus trichocephalus (L.) Liihe, 1912 bi/is (D.) Travassos, Artigas et Pereira, 1928 (nee -E. triehuris Blumenbach, 1779 is not an acanthocephalan Petrotschenko, 1956) -E. triglae Rudolphi, 1819: syn. Euzetacanthus simplex -E. vaseulosus Rudolphi, 1819 (nee vaseulosum): syn. of (R.) Golvan et Houin, 1964 Bolbosoma vasculosum (R.)Meyer, 1932 156 YJ. GOLVAN -E. veli George et Nadakal, 1981 (nee 1978) (Veli Lake, EMPODIUS Travassos, 1916 (nee 1915): syn. of Medio- Trivandrum, India) rhynchus Van Cleave, 1916 (vide DOLLFUS, 1936 and -E. ventrieosus Rudolphi, 1809: syn. of Corynosoma VAN CLEAVE,1947) strumosum (R.)Luhe, 1904 -E. aleeturae Johnston et Edmonds, 1947: syn. of Me- -E. versieolor Rudolphi, 1819: syn. of Polymorphus diorhynchus alecturae (J. et E.) Golvan, 1962 boschadis (S.) Railliet, 1919 pro parte and of Poly- -E. empodius (Skrjabin, 1913) Travassos, 1916: syn. of morphus minutus (G.) Luhe, 1911 pro parte Mediorhynchus empodius (S.) Meyer, 1932 -E. vesieulosus Schrank, 1790: syn. of Filicollis anatis -E. gallinarum (Bhalerao, 1937) Petrotschenko, 1958: (S.) Luhe, 1911 syn. of Mediorhynchus gallinarum (B.) Golvan, 1962 -E. de vesianii Molin, 1856: syn. of Acanthocephaloi- -E. giganteus (Meyer, 1931) Meyer, 1932: syn. of Me- des incrassatus (M.) Meyer, 1932 diorhynchus giganteus Meyer, 1931 -E. waehniae Rudo1phi, 1819: syn. of Echinorhynchus -E. grandis (Van Cleave, 1916) Travassos 1923/24: syn. (Echinorhynchus) gadi O.F. Miiller, 1776 of Mediorhynchus grandis Van Cleave, 1916 -E. wedli Sonsino, 1896: syn. of Centrorhynchus buteo- -E. invaginatus Travassos, 1917: errat. pro E. vaginatus nis (S.) Kostylew, 1914 -E. mirabilis (de Marval, 1905) Travassos, 1917: syn. of -E. xenopeltidis Shipley, 1903: syn. of Pseudoacantho- Mediorhynchus mirabilis (dM.) Meyer, 1932 cephalus xenopeltidis (S.) Golvan, 1969 -E. numidae (Baer, 1925) Kasimov, 1956: syn. of -E. xiphiae Gmelin, 1791 is not an acanthocephalan Mediorhynchus numidae (B.) Meyer, 1932 -E.(Echinorhynchus) yamagutii Golvan, 1969 (Japa- -E. otidis (Miescher, 1841) Travassos, 1917: syn. of nese Sea) Mediorhynchus otidis (M.) Van Cleave, 1947 -E. (sensu lato) zanclorhynchi Johnston et Best, 1937 -E. segmentatus South well et Macfie, 1925 is probably a (Macquarie Island) (nee zanehlorhynehus) Mediorhynchus different from Echinorhynchus seg- -E. zenis Rudolphi, 1819: syn. of Acanthocephaloides mentatus de Marval, 1902 propinquus (D.) Meyer, 1932 -E. taeniatus (von Linstow, 1901) Travassos, 1917: syn. of Mediorhynchus taeniatus (vL.) Dollfus, 1936 ECHINOSOMA Porta, 1907: preoccupied; composite -E. turnixena Tubangui, 1933 (nee 1931): syn. of genus, syn. of Corynosoma Luhe, 1904 pro parte, of Mediorhynchus turnixena (T.) Webster, 1948 Rhadinorhynchus Luhe, 1911 pro parte and of Serra- -E. vaginatus (Diesing, 1851) Travassos, 1917: syn. of sentis Van Cleave, 1923 pro parte Mediorhynchus vaginatus (D.) Meyer, 1932 -E. gibber Olsson in Porta, 1907: syn. of Corynosoma strumosum (R.) Ltihe, 1904 EOCOLLIS Van Cleave, 1947 -E. miliarius (Zeder) Porta, 1907: syn. of Polymorphus -E. arcanus Van Cleave, 1947 (type species) (Illinois, minutus (G.) Luhe, 1911 or of Polymorphus boschadis Ohio, North America) (S.) Railliet, 1919 -E. Catostomi Bucker, 1992 (Alabama, USA) -E. peposaeae Porta, 1907: syn. of Corynosoma longi- -E. harengulae Wang, 1981 (China) lemniscatus Machado Filho, 1961 EORHYNCHUS Hamann, 1892: preoccupied, syn. of -E. pristis (Rudolphi) Porta, 1907: syn. of Rhadi- Neoechinorhynchus Hamann, 1892 norhynchus pristis (R.)Luhe, 1911 -E. agilis (Rudolphi, 1819) Van Cleave, 1914: syn. of -E. roseus (Molin) Porta, 1907: syn. of Arhythmorhyn- Neoechinorhynchus agilis (R.) Van Cleave, 1916 ch us roseus (M.) Meyer, 1932 -E. eylindratus (Van Cleave, 1913) Van Cleave, 1914: -E. (Eehinogaster) sagittifer (Linton) Porta, 1907: syn. syn. of Neoechinorhynchus cylindratus (VC.) Van of Serrasentis sagittifer (L.) Van Cleave, 1923 Cleave, 1919 -E. vaseulosus (Rudolphi) Porta, 1907: syn. of Bolboso- -E. emydis (Leidy, 1851) Van Cleave, 1914: syn. of ma vasculosum (R.) Meyer, 1932 Neoechinorhynchus emydis (L.) Van Cleave, 1919 -E. graeilisentis (Van Cleave, 1913) Van Cleave, 1914: EMPODISMA Yamaguti, 1963: syn. of Mediorhynchus syn. of Gracilisentis gracilisentis (VC.) Van Cleave, Van Cleave, 1916 1919 -E. aleeturae (Johnston et Edmonds, 1947) Yamaguti, -E. longirostris (Van Cleave, 1913) Van Cleave, 1914: 1963: syn. of Mediorhynchus alecturae (J. et E.) Gol- syn. of Tanaorhamphus longirostris (VC.) H.B. Ward, van, 1962 1918 -E. giganteum (Meyer, 1931) Yamaguti, 1963 (nee gi- -E. rutili (O.F. Muller, 1784) Van Cleave, 1914: syn. of ganteus): syn. of Mediorhynchus giganteus Meyer, Neoechinorhynchus rutili (O.F.M.) Hamann, 1892 1931 -E. tenellus (Van Cleave, 1913) Van Cleave, 1914: syn. -E. numidae (Baer, 1925) Yamaguti, 1963: syn. of of Neoechinorhynchus tenellus (VC.) Van Cleave, Mediorhynchus numidae (B.) Meyer, 1932 1919 -E. otidis (Miescher, 1841) Yamaguti, 1963: syn. of Mediorhynchus otidis (M.) Van Cleave, 1947 EOSENTIS Van Cleave, 1928 (nee 1914): may be a syn. -E. turnixena (Tubangui, 1933) Yamaguti, 1963: syn. of (or subgenus ??) of Neoechinorhynchus Hamann, Mediorhynchus turnixena (T.) Webster, 1948 1892 (fide YAMAGUTl,1963) Nomenclature of the Acanthocephala 157

-E. devdevi Datta, 1936 (India): may be a syn. of Neo- FILICOLLIS Liihe, 1911 echinorhynchus devdevi (D.) Kaw, 1951 -F. altmani Perry, 1942: syn. of Profilicollis altmani -E. formosanus Harada, 1938 (Taiwan): may be a syn. (P.) Khokhlova, 1974 of Neoechinorhynchusformosanus (H.) Kaw, 1951 -F. anatis (Schrank, 1788) Luhe, 1911 (type species) -E. rigidus Van Cleave, 1928 (Seistan, India)(type spe- (Holarctic region) cies): may be a syn. of Neoechinorhynchus rigidus -F. arcticus Van Cleave, 1920: syn. of Profilicollis arc- (VC) Kaw, 1951 ticus (VC.) Meyer, 1931 -E. yalei Datta, 1936 (India): may be a syn. of Neoechi- -F. botulus Van Cleave, 1916: syn. of Profilicollis botu- norhynchus yalei (D.) Kaw, 1951 Ius (VC.) Meyer, 1931 -F. kenti Van Cleave, 1947: syn. of Profilicollis kenti EUZETACANTHUS Golvan et Houin, 1964 (VC.) Khokhlova, 1974 -E. chorinemuri Gupta et Naqvi, 1984 (India) -F. major Lundstrorn, 1942: syn. of Profilicollis major -E. golvani Gupta et Fatma, 1985 (Tamil Nadu, India) (L.) Khokhlova, 1974 -E. simplex (Rudolphi, 1810) Golvan et Houin, 1964 -F. sphaeroeephalus (Bremser in Rudolphi, 1819) Tra- (type species) (coasts of France) vassos, 1926: syn. of Profilicollis sphaerocephalus (B. in R.) Khokhlova, 1974 FALSlFILlCOLLlS Webster, 1948: proposed as a subge- -F. sphaeroeephalus sensu Harrington et Pillbury, 1938: nus of Polymorphus Luhe, 1911 and syn. of Profilico- syn. of Profilicollis kenti (VC) Khokhlova;1974 llis Meyer, 1931 -F. trophimenkoi Atrashkevitch, 1982 (Siberia) -F. chasmagnathi Holcman-Spector, Mane-Garzon et Dei-Cas, 1977: syn. of Profilicollis chasmagnathi (H.- FILISOMA Van Cleave, 1928 S., M.-G. et D.-C) novo comb.: may be a Polymorphus -F. bucerium Van Cleave, 1940 (Pacific coast of USA (fide VIZCAINO,1988) and Mexico) (nee bueerinum) -F. kenti (Van Cleave, 1947) Yamaguti, 1963: syn. of -F.fulum Van Cleave et Manter, 1947 (USA, Florida) Profilicollis kenti (VC) Khokhlova, 1974 -F. hoogliensis Datta et Soota, 1962 (Calcutta, India): -F. major (Lundstrom, 1942) Yamaguti, 1963: syn. of may be a syn. of Filisoma indicum Van Cleave, 1928 Profilicollis major (L.) Khokhlova, 1974 (fide GUPTA& JAIN, 1979) -F. sphaeroeephalus (Bremser in Rudolphi, 1819) -F. indicum Van Cleave, 1928 (type species) (Chilka Yamaguti, 1963: syn. of Profilicollis sphaerocephalus Lake, India; found also in Sulawesi) (B. in R.) Khokhlova, 1974 -F. inglisi Gupta et Naqvi, 1984 (India) -F. texensis (Webster, 1948) Yamaguti, 1963: syn. of -F. japonicum Fukui et Morisita, 1936: syn. of Acantho- Profilicollis texensis (W.) Khokhlova, 1974 cephalusjaponicus (F. et M.) Petrotschenko, 1956 FARZANDIA Thapar, 1931: subgenus of Pallisentis -F. microcanthi Harada, 1938 (Taiwan) (nee micracan- Van Cleave, 1928 thi, nee maeroeanthi) -F. nagpurensis Bhalerao, 1931: syn. of Pallisentis -F. rizalinum Tubangui et Masilungan, 1946 (Luzon, (Farzandia) nagpurensis (B.) Baylis, 1933 Philippine Islands) -F. ophiocephali Thapar, 1931: syn. of Pallisentis (Neo- -F. scatophagusi Datta et Soota, 1960 (India) sentis) ophiocephali (T.) Baylis, 1933 FISSISENTlS: errat. pro Fessisentis FEMOGIBBOSUS Parukhin, 1973 -F. assi Parukin, 1973 (type species)(Indian Ocean, FLORIDOSENTIS H.L. Ward, 1953 region of Oman) -F. elongatus H.L. Ward, 1953 (type species) (Florida, Mexico, Puerto Rico) FESSISENTIS Van Cleave, 1931 -F. mugilis (Mach ado Filho, 1951) Bullock, 1962 (Bra- -F.fessus Van Cleave, 1931 (type species) (Mississippi, zil) Louisiana, Illinois, USA): the validity of this genus is -F. pacifica Bravo-Hollis, 1969 (Pacific coast of Me- not admitted by AMIN & REOLlN(1980) and by HUFF- xico) MAN(1973); these authors considered it to be an Echi- norhynchus sensu stricto; I think it must be used, at least, as a subgenus of Echinorhynchus FRESNYARHYNCHUS Golvan, 1960: syn. of Bol- -F. friedi Nickol, 1972 (Ohio, Louisiana, Mississippi, borhynchoides Achmerov, 1959 New Hampshire, USA) -F. exiguus (Achmerov et Dombrowskaja-Achmerova, -F. necturorum Nickol, 1967 (Louisiane, Georgie, 1941) Golvan, 1960: syn. of Bolborhynchoides exi- USA) guus (A. et D.-A.) Achmerov, 1959 -F. tichiganensis Amin, 1980 (Michigan, USA) -F. vancleavei (Hughes et Moore, 1943) Nickol, 1972 FUR CA TA Werby, 1938: syn. of Lueheia Travassos, (, USA) (nee vancleavi) 1919 -F. vancleavei sensu Haley et Bullock, 1953: syn. of -F. adlueheia Werby, 1938: syn. of Lueheia adlueheia Fessisentis friedi Nickol, 1972 (W.) Van Cleave, 1942 158 Y.1. GOLVAN Hamann, 1892 (nec Gigan- -G. spira (Diesing, 1851) Hamann, 1892: syn. ofOliga- thorhynchus) canthorhynchus spira (D.) Travassos, 1915 -G. asturinus Johnston, 1912: syn. of Centrorhynchus -G. spirula sensu Porta, 1909: syn. of Oncicola spirula asturinus (1.) Johnston, 1918 (0.) Schmidt, 1972 pro parte and of Oncicola luehei -G. aurae Travassos, (1912) 1913: syn. ofOligacan- (T.) Schrnidt, 1972 pro parte thorhynchus spira (D.) Travassos, 1915 (nec aurea) -G. spirula sensu Porta, 1912: syn. of Oligacanthorhyn- -G. canis (Kaupp, 1909) Hall et Wigdor, 1918: syn. of chus erinacei (R.) Meyer, 1932 Oncicola canis (K.) Van Cleave, 1921 -G. taenioides (Diesing, 1851) Hamann, 1892: syn. of -G. cestodiformis (von Linstow, 1904) Porta, 1909: syn. Oligacanthorhynchus taenioides (D.) Travassos, 1915 of Moniliformis cestodiformis (vL.) Travassos, 1917 -G. ungriai Antonio, 1958 (Venezuela) (nec ungariai) -G. circumflexus (Molin, 1858) de Marval, 1908 et 1909: syn. of Oligacanthorhynchus circumflexus (M.) GOACANTHUS Gupta et Jain, 1980 Meyer,1932 -G. panajiensis Gupta et Jain, 1980 (type species) (Goa, -G. compressus (Rudolphi, 1802) de Marval, 1905: syn. South of India) of Oligacanthorhynchus spira (D.) Travassos, 1915 pro parte, of Oligacanthorhynchus compressus (R.) GOLVANACANTHUS Paggi et Orecchia, 1972 Meyer, 1932 pro parte and of Oligacanthorhynchus -G. blennii Paggi et Orecchia, 1972 (type species) (Italy, iheringi Travassos, 1916 pro parte North of Sinai, Mediterranean Sea) -G. echinodiscus (Diesing, 1851) Hamann, 1892 (type -G. problematic us Mordinova et Parukin, 1978 (Medite- species) (Neotropical region) rranean Sea, Black Sea) (nec Mordvinova) -G. elegans Porta, 1909: syn. of Prosthenorchis elegans (D.) Travassos, 1915 -G. empodius Skrjabin, 1913: syn. of Mediorhynchus GOLVANORHYNCHUS Noronha, de Fabio et Pinto, empodius (S.) Meyer, 1932 1978 -G. gigas Bloch, 1782: syn. of Macracanthorhynchus -G. golvani Noronha, de Fabio et Pinto, 1978 (type spe- hirudinaceus (P.) Travassos, 1917 cies) (Brazil) -G. hamatus (von Linstow, 1897) Porta, 1908: syn. of Oligacanthorhynchus (Nephridiacanthus ?) hamatus GORD10RHYNCHUS Meyer, 1931: syn. of Cen- (vL.) Schmidt, 1972 pro parte and of Prosthenorchis trorhynchus Liihe, 1911 luehei Travassos, 1916 pro parte -G. bancrofti Johnston et Best, 1943: syn. of Cen- -G. hirudinaceus (Pall as, 1781) Hamann, 1892: syn. of trorhynchus bancrofti (1. and B.) Golvan, 1956 Macracanthorhynchus hirudinaceus (P.) Travassos, -G. californicus (Millzner, 1924) Golvan, 1957: syn. of 1917 Centrorhynchus californicus Millzner, 1924 -G. hirudinaceus sensu Porta, 1908 pro parte: syn. of -G. clitorideus Meyer, 1931: syn. of Centrorhynchus Oncicola spirula (0.) Schmidt, 1972 and of Oncicola clitorideus (M.) Golvan, 1956 luehei (T.) Schmidt, 1972 -G. crocidurus (Das, 1950) Golvan, 1957: syn. of Cen- -G. lopezneyrai Diaz-Ungrfa, 1958 (Venezuela) (nec trorhynchus crocidurus Das, 1950 lopezneirae) -G. falconis Johnston et Best, 1943: syn. of Centrorhyn- -G. lutzi Machado Filho, 1941 (Para, Brazil) chusfalconis (1. et B.) Golvan, 1956 -G. major (Bremser, 1811) Porta, 1908: syn. of Nephri- -G. freundi Hartwich, 1953: syn. of Centrorhynchus diacanthus major (B.) Golvan, 1962 freundi (H.) Golvan, 1956 -G. micracanthus (Rudolphi, 1819) Cholodkowski et -G. gendrei Golvan, 1957: syn. of Centrorhynchus gen- Kostylew, 1916: syn. of Mediorhynchus micracanthus drei (G.) Golvan, 1960 (R.) Van Cleave, 1927 -G. hagiangensis Petrotschenko et Phan, 1969: syn. of -G. microcephalus (Rudolphi, 1819) Porta, 1909: syn. of Centrorhynchus hagiangensis (P. et P.) Arnin, 1985 Hamanniella microcephala (R.) Travassos, 1915 -G. hylae (1ohnston, 1914) Johnston et Edmonds, 1948: -G. mirabilis de Marval, 1904: syn. of Mediorhynchus syn. of Porrorchis hylae (1.) Schmidt et Kuntz, 1967 mirabilis (dM.) Meyer, 1932 -G. insularis (Tubangui, 1933) Golvan, 1957: syn. of -G. moniliformis (Bremser, 1811) Hamann, 1892: syn. Centrorhynchus insularis Tubangui, 1933 of Moniliformis moniliformis (B.) Travassos, 1915 -G. itatsinis (Fukui, 1929) Golvan, 1957: syn. of Cen- -G. ortizi Sarmiento, 1954 (Colombia, Peru) trorhynchus itatsinis Fukui, 1929 -G. otidis (Miescher, 1841) Kostylew, 1914: syn. of -G. madag ascariensis Golvan, 1957: syn. of Cen- Mediorhynchus otidis (M.) Golvan, 1962 trorhynchus madagascariensis (G.) Golvan, 1960 -G. pesteri Tadros, 1966 (Zambia, Zimbabwe) -G. magnus (Fukui, 1929) Golvan, 1957: syn. of Cen- -G. rhopalorhynchus (Diesing, 1851) Porta, 1909: syn. trorhynchus magnus Fukui, 1929 of Polyacanthorhynchus rhopalorhynchus (D.) Tra- -G. microcephalus Bravo-Hollis, 1947: syn. of Cen- vassos, 1920 trorhynchus microcephalus (BH.) Golvan, 1956 -G. semoni (von Linstow, 1898) Porta, 1908: syn. of Aus- -G. turdi (Yamaguti, 1939) Kamegai, 1963: syn. of traliformis semoni (vL.) Schmidt et Edmonds, 1989 Sphaerirostris turdi (Y.) Golvan, 1956 Nomenclature of the Acanthocephala 159 GORGORHYNCHOIDES Cable et Linderoth, 1963 HAMANIA Neiva da Cunha, 1915: syn. of Hamanniella -G. bullocki Cable et Mafarachisi, 1970 (Floride, USA; Travassos, 1915 Bay Chetumal, Mexico) -H. microcephala Travassos, 1915: vide Hamanniella -G. elongatus Cable et Linderoth, 1963 (type species) microcephala (R.) Travassos, 1915 (Curacao) -G. epinepheli Wang, 1986 (China, Fujian) HAMANIELLA Travassos, 1915: errat. pro Hamannie- -G. lintoni Cable et Mafarachisi, 1970 (Northern part of lla Travassos, 1915 the Atlantic coast of USA) -G. orientalis Wang, 1986 (China, Fujian) HAMANNIA Travassos, 1915: preoccupied, syn. of -G. satoi (Morisita, 1937) Wang Pu Qin, 1966: syn. of Hamanniella Travassos, 1915 Gorgorhynchus satoi (M.) Yamaguti, 1963 -H. microcephala (Rudolphi, 1819) Travassos, 1915: syn. of Hamanniella microcephala (R.) Travassos, GORGORHYNCHUS Chandler, 1934 1915 -G. cablei Golvan, 1969 (Jamaica) -G. celebensis (Yamaguti, 1954) Golvan, 1969 (Sula- HAMANNIELLA Travassos, 1915 wesi) -H. carinii Travassos, 1916 (Brazil) -G. clavatus Van Cleave, 1940 (Galapagos) -H. eati Gupta et Lata, 1967: syn. of Oligacanthorhyn- -G. epinepheli (Yamaguti, 1939) Golvan, 1960 (Japan) chus cati (G. et L.) Schmidt, 1972 (nee epinephalii: syn. of Pseudauchen epinepheli (Y.) -H. microcephala (Rudolphi, 1819) Travassos, 1915 Yamaguti,1963 (type species) (Brazil, Colombia, Panama; Neotropical -G. gibber Chandler, 1934 (Texas, USA): may be a syn. region) of Gorgorhynchus medius (L.)Chandler, 1934 which -H. tortuosa (Leidy, 1850) Van Cleave, 1924 (Oklaho- becomes the type species (vide VANCLEAVE,1940) ma, North America) -G. lepidus Van Cleave, 1940 (Galapagos) -H. tumida (Van Cleave, 1947) Van Cleave, 1953 -G. medius (Linton, 1908) Chandler, 1934 (type spe- (USA) cies) (Vera Cruz, gulf of Mexico, Bermuda Islands) -G. nemipteri Parukhin, 1973 (Indian Ocean) HANUMANTHARAORHYNCHUS Chandra, 1983 -G. ophiocephali Furtado et Chau-Lan, 1971 (Malaysia) -H. hemirhamphi Chandra, 1983 (type species) (Ben- -G. polymixiae Kovalenko, 1981 (Philippine Islands) gal) -G. robertdollfusi Golvan, 1956 (Mauritanie) (may be a larva of Bolbosoma sp.) HEBESOMA Van Cleave, 1928 -G. satoi (Morisita, 1937) Yamaguti, 1963 (China, Tai- -H. derosum (?) in Amin 1992 wan) -H. violentum Van Cleave, 1928 (Wuchang, Changsha, -G. trachinotus Noronha, Vicente, Pinto et de Fabio, Shoahsing, China) (type species): may be a Neoechi- 1986 (Brazil) norhynchus (fide SALGADO-MALDONADO,1979) (vide AMIN,1992) GORYTOCEPHALUS Nickol et Thatcher, 1971 -G. elongorchis Thatcher, 1979 (Amazonia, Brazil) HEMIECHINOSOMA Petrotschenko et Smogorjevska- -G. plecostomorum Nickol et Thatcher, 1971 (type spe- ja, 1962: may be a syn. of Southwellina Witenberg, cies) (Panama, Peru) 1932 -G. spectabilis (Machado Filho, 1959) Nickol et That- -H. ardeae (Belopolskaja, 1958) Petrotschenko et Smo- cher, 1971 (Brazil) gorjevskaja, 1962 (SSSR) -G. talaensis Vizcaino et Lunaschi, 1987 (Buenos Aires, -H. mergi (Lundstrorn, 1941) Petrotschenko et Smogor- Argentina) jevskaja, 1962: syn. of Andracantha mergi (L.) Sch- midt, 1975 GRACILISENTIS Van Cleave, 1919 -H. ponticum Petrotschenko et Smogorjevskaja, 1962 -G. gracilisentis (Van Cleave, 1913) Van Cleave, 1919 (type species) (SSSR, Crimea): may be a syn. of South- (type species) (Mississippi, Illinois, USA) wellina pontica (P. et S.) novo comb. -G. mugilus Gupta et Lata, 1967 (Chandigarh, India) (nee mugilis) (= Gracilisentis mugilus sharmai G. et L.) HEMIGYRUS Achmerov et Dombrowskaja-Achmerova, -G. variabilis (Diesing, 1856) Travassos, Artigas et 1941 (nee Hemigurus): syn. of Acanthocephalorhyn- Pereira, 1928 (Brazil) choides Kostylew, 1941 -H. intermedius Achmerov et Dombrowskaja-Achmero- HAERUCA Rudolphi, 1793 va, 1941: syn. of Acanthocephalorhynchoides ussu- -H. erinacei Rudolphi, 1793: syn. of Oligacanthorhyn- riensis Kostylew, 1941 chus erinacei (R.) Meyer, 1932 HEMIRHADINORHYNCHUS Krotov et Petrotschenko, HAERUCULA Pallas, 1760: syn. of Acanthocephalus 1956: syn. of Achmerov et Koelreuther, 1771 Dombrowskaja-Achmerova, 1941 160 Y.J. GOLVAN -H. leueiseus Krotov et Petrotschenko, 1956: syn. of syn. of Arhythmacanthus overstreeti Schmidt et Pseudorhadinorhynchus leuciscus (K. et P.) Golvan, Paperna, 1978 1969 -H. paraplagusiarum (Nickol, 1972) Amin, 1985 -H. plotosi Yamaguti, 1935: syn. of Paracanthocepha- HEPTAMEGACANTHUS Spencer Jones, 1990 loides plotosi (Y.) Golvan, 1969 -H. niekerki Spencer Jones, 1990 (type species) (South -H. plotosi sensu Tripathi, 1959: syn. of Paracantho- Africa, Transkei) cephaloides tripathii Golvan, 1969 -H. rhinoplagusiae (Yamaguti, 1935) Tripathi, 1959: HETERACANTHOCEPHALUS Petrotschenko, 1956 syn. of Yamagutisentis rhinoplagusiae (Y.) Golvan, -H. hureaui Dollfus, 1964: syn. of Heterosentis mage- 1969 llanicus (S.) (fide ZOZITOWIECKI,1990) -H. septaeanthus (Sita, 1949) Amin, 1985: syn. of -H. peltorhamphi (Baylis, 1944) Petrotschenko, 1956 Arhythmacanthus septacanthus Sita, 1949 (type species) (South of the Pacific Ocean) -H. thapari (Gupta et Fatma, 1969) Amin, 1985: syn. of Arhythmacanthus thapari Gupta et Fatma, 1969 HETERACANTHORHYNCHUS Lundstrom, 1942: syn. of Mediorhynchus Van Cleave, 1916 HEXAGLANDULA Petrotschenko, 1950 -H. eehinopsi Horchner, 1962: syn. of Promoniliformis -H. ariusis Bilqees, 1971 (Pakistan) ovocristatus (vL.) Dollfus et Golvan, 1964 -H. corynosoma (Travassos, 1915) Petrotschenko, 1958 -H. vancleavei Lundstrom, 1942: syn. of Mediorhyn- (Brazil) ch us vancleavei (L.) Go1van, 1962 -H. inermis (Travassos, 1923) Petrotschenko, 1958 (Brazil) HETEROPHUS: errat. pro Heteroplus Kostylew, 1914 -H. karachiensis Bilqees, 1971 (Pakistan) -H. mutabilis (Rudolphi, 1819) Petrotschenko, 1950 HETEROPLUS Kostylew, 1914: preoccupied, syn. of (type species) (Brazil, Uruguay) Mediorhynchus Van Cleave, 1916 -H. paucihamatus (Heinze, 1936) Petrotschenko, 1958 -H. empodius (Skrjabin, 1913) Kostylew, 1915: syn. of (East of Europe) Mediorhynchus empodius (S.) Van Cleave, 1924 (?) -H. grandis (Van Cleave, 1916) Van Cleave, 1918: syn. HEXASPIRON Dollfus et Golvan, 1956 of Mediorhynchus grandis Van Cleave, 1916 -H. nigericum Dollfus et Golvan, 1956 (type species) -H. mirabilis (de Marval, 1905) Kostylew, 1914: syn. of (Basin of the Niger River) (nee nigeriensis) Mediorhynchus mirabilis (dM.) Meyer, 1932 -H. numidae Baer, 1925: syn. of Mediorhynchus numi- HORMORHYNCHUS H.B. Ward, 1917: syn. of Monili- dae (B.) Golvan, 1962 formis Travassos, 1917 pro parte and of Macracant- -H. otidis (Miescher, 1841 in de Marval, 1904) Kosty- horhynchus Travassos, 1917 pro parte lew, 1914: syn. of Mediorhynchus otidis (M. in dM.) -H. clarki H.B. Ward, 1917: syn. of Moniliformis clarki Van Cleave, 1947 (H.B.W.) Chandler, 1921 -H. taeniatus (von Linstow, 1901) in Baer, 1925: syn. of -H. gigas (Bloch, 1782) Johnston, 1918: syn. of Macra- Mediorhynchus taeniatus (vL.) Dollfus, 1936 canthorhynchus hirudinaceus (P.) Travassos, 1917 -H. vaginatus (Diesing, 1851) in Baer, 1925: syn. of -H. moniliformis (Bremser, 1811) H.B. Ward, 1917: Mediorhynchus vaginatus (D.) Meyer, 1932 syn. of Moniliformis moniliformis (B.) Travassos, 1917 HETEROSENTIS Van Cleave, 1931 -H. caballeroi Gupta et Fatma, 1985 (Tamil Nadu, HYPOECHINORHYNCHUS Yamaguti, 1939 India) -H. alaeopis Yamaguti, 1939 (type species) (Sizuoka, -H. fusiformis (Yamaguti, 1935) Tripathi, 1959: syn. of Japan, South Australia) Arhythmacanthus fusiformis Yamaguti, 1935 -H. golvani Gupta et Kumar, 1987 (Chilka Lake, Orissa, -H. heteracanthus (von Linstow, 1896) Van Cleave, India) 1931 (nee hetraeanthus) (type species) (Tierra del -H. magellanicus Szidat, 1950 (South coast of Argenti- Fuego) na) -H. magellanieus (Szidat, 1950) Zdzitowiecki, 1990: -H. thermaceri de Buron, 1988 (Eastern Pacific Dor- syn. of Heterosentis magellanicus (S.)Zdzitowiecki, sale) 1990 -H. magellanieus sensu Szidat, 1965 is another species ILLIOSENTIS Van Cleave et Lincicome, 1939 (fide ZOZITOVIECKI,1990) -I. africanus Golvan, 1955 (Senegal) -H. magellanieus sensu Szidat et Graefe, 1967 is another -I. cetratus Van Cleave, 1945 (nee eentratus) (USA, species (fide ZOZITOVIECKI,1990) California) -H. neobythitis (Yamaguti, 1939) Tripathi, 1959: syn. of -I. etenorhynehus Cable et Linderoth, 1963: syn. of Yamagutisentis neobythitis (Y.) Golvan, 1969 Dollfusentis ctenorhynchus (c. et L.) Go1van, 1969 -H. overstreeti (Schmidt et Paperna, 1978) Amin, 1985: -I. edmondsi Golvan, 1960 (Australia) Nomenclature of the Acanthocephala 161 -I.furcatus Van Cleave et Lincicome, 1939 (type spe- -L. minor Fukui et Morisita, 1936: syn. of Longicollum cies) (Louisiana, USA): may be a Tegorhynchus (fide alemniscus (H.) Fukui et Morisita, 1937 BUCKNER& MATEO, 1978); this synonymy cannot be -L. noellae Golvan, 1969 (Atlantic coast of Morocco) accepted because in the genus Illiosentis there are 2 -L. pagrosomi Yamaguti, 1935 (type species) (Japan) «lips» surrounding the female genital pore and those (nee pagrosoma) structures are absent in the genus Tegorhynchus (vide -L. pagrosomi sensu Johnston et Edmonds, 1951: syn. of LEOTTA,SCHMIDT& KUNTZ, 1982) Longicollum edmondsi Golvan, 1969 -I. furcatus sensu Edmonds, 1957: syn. of Illiosentis -L. psettodai Gupta et Gupta, 1979 (India) (nee psettod- edmondsi Golvan, 1969 sai) -I. heteraeanthus Cable et Linderoth, 1963: syn. of Doll- -L. quiloni Gupta et Naqvi, 1984 (India) fusentis heteracanthus (C. et L.) Golvan, 1969 -L. riouxi Golvan, 1969 (Senegal) -I. longispinus Cable et Linderoth, 1963: syn. of Dollfu- -L. sergenti (Choquette et Gayot, 1952) Golvan, 1969: sentis longispinus (C. et L.) Golvan, 1969 syn. of Paralongicollum sergenti (C. et G.) Amin, -I. multacanthus Mamaev, 1970 (Gulf of Tonkin) Bauer et Sidorov, 1991

INDORHYNCHUS Golvan, 1969 LONGIROSTRIS Golvan, 1956: syn. of Centrorhynchus -I. indicus (Tripathi, 1959) Golvan, 1969 (type species) Liihe,1911 (Puri, Chilka Lake, India) LUEHEIA Travassos, 1920 KALEOPS or KOLEOPS Lockwood, 1872: syn. of -L. adlueheia (Werby, 1938) Van Cleave, 1942 (USA, Acanthocephalus Koelreuther, 1771 Washington) -L. boreotis Van Cleave et Williams, 1951: syn. of LEIPERACANTHUS Bhalerao, 1937: syn. of Me- Pseudolueheia boreotis (VC. et W.) Schrnidt et Kuntz, diorhynchus Van Cleave, 1916 1967 -L. gallinarum Bhalerao, 1937: syn. of Mediorhynchus -L. cajabambensis Machado Filho et Ibanez, 1967 gallinarum (B.) Golvan, 1962 (Peru) -L. inscripta (Westrumb, 1821) Travassos, 1920 (nee LEPIDOSOMA Porta, 1908: preoccupied, syn. of Serra- inseipta) (Neotropical region: Brazil, Panama, Nicara- sentis Van Cleave, 1923 gua, Puerto Rico) -L. lamelliger (Diesing, 1854) Porta, 1908: syn. of -L. lueheia Travassos, 1920 (type species) (South Ame- Serrasentis lamelliger (D.) Meyer, 1932 rica) (nee luehei)

LEPTORHYNCHOIDES Kostylew, 1924 MACRACANTHOCEPHALUS Baylis, 1944: errat. pro -L. aphredoderi Buckner et Buckner, 1976 (USA, Loui- Micracanthocephalus Harada, 1938 siana) -L. eampbelli (Leiper et Atkinson, 1914) sensu Johnston MACRACANTHORHYNCHUS Travassos, 1917 (nee et Best, 1937: syn. of Metacanthocephalus johnstoni 1916) Zdzitowiecki, 1983 -M. catulinus Kostylew, 1927 (Turkestan, Central Asia) -L. debenhami (Leiper et Atkinson, 1914) Johnston et -M. (?) erinacei Dollfus, 1953 (North Africa) Best, 1937: syn. of Metacanthocephalus rennicki (L. -M. hirudinaceus (Pallas, 1781) Travassos, (1916) 1917 et A.) Zdzitowiecki, 1983 (nee hirundinaeeus) (type species) (doubtless cosmo- -L. ovieephalus Zhukov, 1960: syn. of Metacanthoce- politan) phalus ovicephalus (Z.) Golvan, 1969 -M. ingens (von Linstow, 1879) Meyer, 1932 (Nearctic -L. plagicephalus (Westrumb, 1821) Kostylew, 1924 region) (type species) (Volga River Basin, Caspian Sea, Black -M. major Machado Filho, 1963: syn. of Oligacantho- Sea, Mediterranean Sea) rhynchus major (MF.) Schrnidt, 1972 -L. thecatus (Linton, 1891) Kostylew, 1924 (Nearctic region) MALGACANTHUS Golvan, 1965: subgenus of Cen- trorhynchus Luhe, 1911 LONGICOLLUM Yamaguti, 1935 -L. alemniscus (Harada, 1935) Fukui et Morisita, 1937 MAMANNIELLA Travassos, 1917: errat. pro Haman- (Taiwan) niella Travassos, 1915 -L. cadenati Gupta et Naqvi, 1984 (India) -L. chabanaudi Dollfus et Golvan, 1963 (South Africa) MEDIORHYNCHUS Van Cleave, 1916 -L. edmondsi Golvan, 1969 (Queensland, Australia) -M. alecturae (J ohnston et Edmonds, 1947) Gol van, -L. engraulisi Gupta et Fatma,(1983) 1985 (Kerala, 1962 (Australia) India) -M. armenicus Petrotschenko, 1958 (nee armenieum) -L. indicum Gupta et Gupta, 1970 (India) [Central Asia, also found in Brazil (very doubtful)]: -L. lutiani Jain et Gupta, 1980 (Goa, India) (nee lutjani) may be a syn. of Mediorhynchus micracanthus (R.) 162 Y.J. GOLVAN Van Cleave, 1927 (fide SCHMIDT& KUNTZ, 1977) -M. najasthanensis Gupta, 1976 (India) -M. bakeri Byrd et Kellogg, 1971 (Florida, USA): may -M. numidae (Baer, 1925) Golvan, 1962 (South and be a syn. of Mediorhynchus papillosus Van Cleave, West Africa) 1916 (fide SCHMIDT& KUNTZ, 1977) -M. orientalis Belopolskaya, 1953 (nee orientale) [Cen- -M. bullocki Gupta et Jain, 1973 (India): may be a syn. tral Asia, India, also found in Indonesia, Taiwan, Bor- of Mediorhynchus orientalis Belopolskaja, 1953 (fide neo, Pescadores and Hawai (doubtful)] SCHMIDT& KUNTZ, 1977) -M. oswaldocruzi Travassos, 1923 (Brazil) -M. cambellensis Soota, Srivastava et Ghosh, 1969 -M. otidis (Miescher, 1841 in de Marval, 1904) Van (India): this species is doubtful and cannot be identified Cleave, 1947 (Europe, Central Asia) (fide SCHMIDT& KUNTZ, 1977) -M. papillosus Van Cleave, 1916 (nee papillosum) (type -M. centurorum Nickol, 1969 (Louisiana, USA) species) (Holarctic region, also found in Taiwan, Pes- -M. channapettae George et N adakal, 1984 (Kerala, cadores Islands) India) -M. passeris Das, 1951 (India) -M. colini Webster, 1948 (Texas, USA): may be a syn. -M. pauciuncinatus Dollfus, 1959 (Peru) of Mediorhynchus papillosus Van Cleave, 1916 (fide -M. petrotschenkoi Gvozdev et Soboleva, 1966 (Central SCHMIDT& KUNTZ, 1977) Asia) (nee petroehenkoi) -M. conirostris H.L. Ward, 1966 (Egypt, Israel) -M. pintoi Travassos, 1923 (Brazil): the original des- -M. corcoracis Johnston et Edmonds, 1951 (Queens- cription was based on a broken female and impossible land, South Australia) to identify (fide SCHMIDT & KUNTZ, 1977): species -M. edmondsi Schmidt et Kuntz, 1977 (Palawan, Philip- dubia pine Islands) -M. rajasthanensis Gupta, 1976 (India) -M. emberizae (Rudolphi, 1819) Travassos, 1924 (nee -M. robustus Van Cleave, 1916 (nee robustum) (Nearc- Van Cleave, 1916) (Brazil, Puerto Rico) tic region) -M. empodius (Skrjabin, 1913) Van Cleave, 1924 (nee -M. rodensis Diaz-Cosin, 1971 (Spain) Meyer, 1932) (Turkestan, found also in Belgium ?) -M. segmentatus (de Marval, 1902) Dollfus, 1936 -M. fasciatus (Westrumb, 1821) Meyer, 1932: may be a (Sudan) syn. of Prosthorhynchus cylindraceus (fide MARVAL, -M. selengensis Harris, 1973 (nee selenensis) (Kenya): 1905) is not a syn. of Mediorhynchus gallinarum (B.) Van -M. gallinarum (Bhalerao, 1937) Van Cleave, 1947 Cleave, 1947 as suggested by SCHMIDT & KUNTZ (Africa, found also in the Philippine Islands, in New (1977) Guinea and Sulawesi, etc.) -M. sharmai Gupta et Lata, 1967 (India): this species -M. garruli Yamaguti, 1939 (Toyohasi, Japan): may be was found in a (Mugil sp.) and,fide SCHMIDT& a syn. of Mediorhynchus robustus Van Cleave, 1916 KUNTZ(1977) it is a Centrorhynchus sharmai swallo- (fide SCHMIDT & KUNTZ, 1977) but it seems to me wed with the food of this abnormal questionable -M. sipocotensis Tubangui, 1935 (nee sipposotensis, nee -M. giganteus Meyer, 1931 (nee giganteum) (East Africa) sipoeotense) (Luzon, Philippine Islands) -M. grandis Van Cleave, 1916 (nee grande) (USA) -M. sipocotensis sensu Baker and Hamon, 1968 may be -M. grande (sic) sensu Kelly and Finnie, 1972 (Austra- a syn. of Mediorhynchus robustus Van Cleave, 1916 lia) may be another species (fide SCHMIDT& KUNTZ, 1977) -M. indicus George, Nadakal, Vijayakumaran et Rajen- -M. taeniatus (von Linstow, 1901) Dollfus, 1936 (nee dran, 1981 (Kerala, India) taeniatis, nee taeniatum) (Palaearctic region, Ural, -M. kuntzi H.L. Ward, 1960 (Egypt, Israel) Egypt) -M. lagodekhiensis Kuraschvili, 1955 (nee lagodek- -M. tanagrae (Rudo1phi, 1819) Travassos, 1921: a spe- hiense) (Central Asia) cies which cannot be identified (fide SCHMIDT & -M. leptis H.L. Ward, 1966 (Egypt, Israel) KUNTZ, 1977) -M. lophurae Wang Pu Qin, 1966 (nee lophura) -M. tenuis Meyer, 1931 (nee tenue) (Egypt) (Fukien, China) -M. textori Barus, Six! et Majumdar, 1978 (Ghana, West -M. mariae George et Nadakal, 1984 (Kerala, India) Africa) -M. mattei Marchand et Vassiliades, 1982 (Senegal, -M. turnixena (Tubangui, 1933) Golvan, 1962 (Philip- West Africa) pine Islands) -M. meiringi Bisseru, 1960 (South West of Africa) -M. vaginatus (Diesing, 1851) Meyer, 1932 (nee vagi- -M. micracanthus (Rudolphi, 1819) Meyer, 1932 (nee natum) (Neotropical region): a species which cannot be mieroeanthus) (Europe, found also in Brazil but this identified (fide SCHMIDT& KUNTZ, 1977) identification seems erroneous ?) -M. vancleavei (Lundstrom, 1942) Golvan, 1962 (Scan- -M. mirabilis (de Marval, 1905) Travassos, 1925 (nee dinavia) mirabile) (probably from the Neotropical region becau- -M. wardi Schmidt et Canaris, 1967 (Kenya) se parasitic in a Vultur sp.) -M. zosteropis (Porta, 1913) Meyer, 1932 (New Cale- -M. murtensis Lundstrom, 1942 (nee muritensis, nee donia): a species which cannot be identified (fide murtense) (West of Russia, Scandinavia) SCHMIDT& KUNTZ, 1977) Nomenclature of the Acanthocephala 163

MEGAPRIAPUS Golvan, Gracia-Rodrigo et Dfaz- -M. coregoni (Linkins in Van Cleave, 1919) Golvan, Ungria, 1964 1969: vide Echinorhynchus (M.) coregoni Linkins in -M. ungriai (Gracia-Rodrigo, 1960) Golvan, Gracia-Ro- Van Cleave, 1919 drigo et Diaz-Ungrfa, 1964 (type species) (Venezuela) -M. eryophilus Sokolowskaja, 1962: vide Echinorhyn- chus (M.) cryophilus (Sokolowskaja, 1962) Amin, MEGISTACANTHA Golvan, 1960 1985 -M. horridum (Liihe, 1912) Golvan, 1960 (type species) -M. gomesi (Mach ado Filho, 1948) Golvan, 1969: vide (Nile River Basin, Egypt) Echinorhynchus (M.) gomesi Machado Filho, 1948 -M. jueundus (Travassos, 1923) Golvan, 1969 (nee MEHRARHYNCHUS Datta, 1940: syn. of Cleaveius jueundum): vide Echinorhynchus (M.)jucundus Tra- Subrahmanian, 1927 vassos, 1923) -M. inglisi Gupta et Fatma, 1987: syn. of Cleaveius -M. kushiroensis (Fujita, 1921) Golvan, 1969: vide inglisi (G. et F. ) novo comb. Echinorhynchus (M.) kushiroensis Fujita, 1921 -M. mysti Sahay et Sinha, 1971: syn. of Cleaveius mysti -M. lageniformis (Ekbaum, 1938) Petrotschenko, 1956: (S. et S.) Amin, 1985 vide Echinorhynchus (M.) lageniformis Ekbaum, -M. prashadi Datta, 1940: syn. of Cleaveius prashadi 1938 (D.) Golvan, 1969 -M. lateralis (Leidy, 1851) Golvan, 1969: vide Echi- -M. seeundus Tripathi, 1959: syn. of Cleaveius secun- norhynchus (M.) lateralis Leidy 1851 dus (T.) Golvan, 1969 -M. leidyi (Van Cleave, 1924) Golvan, 1969: vide Echi- -M. singhai Gupta et Fatma, 1987: syn. of Cleaveius norhynchus (M.) leidyi Van Cleave, 1924 singhai (G. et F.) novo comb. -M. malacocephali Parukin, 1985 (South of the Atlantic Ocean, Capetown) METACANTHOCEPHALOIDES Yamaguti, 1959 -M. muraenolepisi Rodyuk, 1984 (Antarctic): syn. of -M. zebrini Yamaguti, 1959 (type species) (Japanese Echinorhynchus (Metechinorhynchus) muraenolepi- Inland Sea) si (Rodyuk, 1984) novo comb. -M. paranensis (Mach ado Filho, 1959) Golvan, 1969 METACANTHOCEPHALUS Yamaguti, 1959 (nee paranensis): vide Echinorhynchus (M.) paranen- -M. campbelli (Leiper et Atkinson, 1914) Golvan, 1969 sis Machado Filho, 1959 (Antarctic) -M. petrotsehenkoi Rodyuk, 1984: syn. of E. (Metechi- -M. dalmori Zdzitowiecki, 1983 (Antarctic) norhynchus) petrotschenkoi (R.)Zdzitowiecki, 1989 -M. johnstoni Zdzitowiecki, 1983 (Antarctic) -M. rhenanus Golvan, 1969: vide Echinorhynchus (M.) -M. ovicephalus (Zhukov, 1960) Golvan, 1969 (Kurile rhenanus (G.) Islands) -M. salmonis (O.F. Muller, 1780) Petrotschenko, 1956: -M. pleuronichthydis Yamaguti, 1959 (type species) vide Echinorhynchus (M.) salmon is O.F. Muller, 1780 (Japanese Inland Sea) -M. salobrensis (Machado Filho, 1948) Golvan, 1969: -M. rennicki (Leiper et Atkinson, 1914) Zdzitowiecki, vide Echinorhynchus (M.) salobrensis Machado 1983 (Antarctic) Filho,1948 -M. thapari Gupta et Gupta, 1975 (India) -M. sevani (Dinnik, 1933) Golvan, 1969: vide Echi- norhynchus (M.) sevani Dinnik, 1932 METARHADINORHYNCHUS Yamaguti, 1959 -M. truttae (Schrank, 1788) Petrotschenko, 1956: vide -M. arii (???)Wang, 1986 (China, Fujian) Echinorhynchus (M.) truttae Schrank, 1788 -M. cyprini (???) Wang, 1986 (China, Fujian) -M. lateolabracis Yamaguti, 1959 (type species) (Japa- MICRACANTHOCEPHALUS Harada, 1938: preoccu- nese Inland Sea) pied, syn. of Micracanthorhynchina Strand, 1936 -M. valiyathurae Nadakal, John et Jacob, 1990 (Kerala, -M. dakusuiensis Harada, 1938: syn. of Micracan- India) thorhynchina dakusuiensis (H.) H.L. Ward, 1951 -M. hemirhamphi Baylis, 1944 in Johnston et Edmonds, METECHINORHYNCHUS Petrotschenko, 1956: sub- 1952: syn. of Micracanthorhynchina hemirhamphi genus of Echinorhynchus O.F. Muller, 1776 (B.)H.L. Ward, 1951 -M. alpinus (von Linstow, 1901) Petrotschenko, 1956: -M. motomurai (Harada, 1935) Harada, 1938: syn. of vide Echinorhynchus (M.) alpinus von Linstow, 1901 Micracanthorhynchina motomurai (H.) H.L. Ward, -M. baeri (Kostylew, 1928) Petrotschenko, 1956: vide 1951 Echinorhynchus (Pseudoechinorhynchus) baeri Kostylew,1928 MICRACANTHORHYNCHINA Strand, 1936 -M. brieoni (Machado Filho, 1959) Golvan, 1969: vide -M. cynoglossi Wang Pu Qin, 1980 (Fujian, China) Echinorhynchus (M.) briconi Machado Filho, 1959 -M. dakusuiensis (Harada, 1938) H.L. Ward, 1951 (Tai- -M. eampbelli (Leiper et Atkinson, 1914) Petrotschenko, wan) 1956: syn. of Metacanthocephalus campbelli (L. et -M. hemiculturus Demshin, 1965 (nee hemieultrus) A.) Golvan, 1969 (SSSR) 164 Y.J. GOLVAN -u. hemirhamphi (Baylis, 1944) H.L. Ward, 1951 -M. moniliformis aegyptieus Meyer, 1932: vide Monili- (New Zealand, South Australia, Tasmania) formis aegypticus -M. indica Farooqi, 1980 (India) (nee indieus) -M. moniliformis sieiliensis Meyer, 1932: vide Monili- -M. lateolabracis Wang Pu Qin, 1980 (Fujian, China) formis siciliensis -Mo motomurai (Harada, 1935) H.L. Ward, 1951 (type -M. monoechinus (von Linstow, 1902) Petrotschenko, species) (Korea, Taiwan) 1958 (Neotropical region) (nee moneehinus) -M. sajori (Belous, 1952) Golvan, 1969 (South Pacific) -M. myoxi (Galli- Valerio, 1929): species inquirenda -M. segmentata (Yamaguti, 1959) Araki et Machida, (Europe) may be closely related if not identical to 1987: syn. of Allorhadinorhynchus segmentatus Moniliformis siciliensis Yamaguti,1959 -M. ovoeristatus (von Linstow, 1897) Petrotschenko, 1958: syn. of Promoniliformis ovocristatus (vL.) Doll- MICRACANTHORHYNCHUS Travassos, 1918: syn. of fus et Golvan, 1963 Mediorhynchus Van Cleave, 1916 -M. pseudosegmentatus (Kntippfer, 1888) Meyer, 1932: -M. emberizae (Rudolphi, 1819) Travassos, 1918: syn. species inquirenda (SSSR) may be closely related if of Mediorhynchus emberizae (R.) Van Cleave, 1916 not identical to Moniliformis siciliensis -M. invaginatus (Diesing, 1851) Travassos, 1918: syn. -M. semoni (von Linstow, 1898) Johnston et Edmonds, of Mediorhynchus vaginatus (D.) Meyer, 1932 1952: syn. of Australiformis semoni (vL.) Schmidt et -M. mieraeanthus (Rudolphi, 1819) Travassos, 19l7: Edmonds, 1989 syn. of Mediorhynchus micracathus (R.) Van Cleave, -M, siciliensis Meyer, 1932 (South of Italy) 1927 -M. spiradentatus Mac Leod, 1933 (nee spiradentatis, nee spirodentatus): may be a syn. of Moniliformis MICRACANTHORHYNCHUS Harada, 1935: preoccu- clarki (H.B.W.) Chandler, 1921 (fide CHANDLER, pied, syn. of Micracanthorhynchina Strand, 1936 1924) or of Moniliformis dubius Meyer, 1932 (fide -M. motomurai Harada, 1935: syn. of Micracant- VANCLEAVE,1946) horhynchina motomurai (H.) H.L. Ward, 1951 -M. spiralis Subrahmanian, 1927 (Burma) -Mo tarsii Deveaux, Schmidt et Krishnasamy, 1988 MICROSENTIS Martin et Multani, 1966 (East Malaysia, National Park of Poring, Sabah) -Mo wardae Martin et Multani, 1966 (type species) -Mo travassosi Meyer, 1932 (Brazil): may be a syn. of (Mexico, Baja California) Moniliformis moniliformis (fide MACHADO FILHO, 1946 and VAN CLEAVE, 1953) MONILIFORMIS Travassos, 1915 -M. acomysi H.L. Ward et Nelson, 1967 (Egypt, Israel) NEOACANTHOCEPHALOIDES Cable et Quick, -M. aegypticus Meyer, 1932 (Egypt) 1954: may be a syn. of Pseudorhadinorhynchus Ach- -M. (sensu lato) appendieulatus (Westrumb, 1821): may merov et Dombrowskaja-Achmerova, 1941 (fide be a syn. of Centrorhynchus aluconis (O.F.M.) Liihe, GUPTA& FATMA, 1981) 1911 -N. neobythitis (Yamaguti, 1939) Cable et Quick, 1954: -M. cestodiformis (von Linstow, 1904) Travassos, 1917 syn. of Yamagutisentis neobythitis (Y.) Golvan, 1969 (Central Africa) -N. rhinoplagusiae (Yamaguti, 1939) Cable et Quick, -u. clarki (H.B. Ward, 1917) Chandler, 1921 (nee Van 1954: syn. of Yamagutisentis rhinoplagusiae (Y.) Gol- Cleave, 1924 or 1925)(North America) van, 1969 -M. convolutus Meyer, 1932 (Brazil) -No spinicaudatus Cable et Quick, 1954 (type species) -M. dubius Meyer, 1932 [North America, also found in (Puerto Rico) Eastern Asia (doubtful)]: may be a syn. of Monilifor- mis moniliformis (B.) T., 1915 (vide VAN CLEAVE, NEOACANTHORHYNCHUS Morisita, 1937: syn. of 1953) Gorgorhynchus Chandler, 1934 -M. echinosorexi Deveaux, Schmidt et Krishnasamy, -N. satoi Morisita, 1937: syn. of Gorgorhynchus satoi 1988 (Malay Peninsula, Ulu Gombak, Selangor) (M.) Yamaguti, 1963 -M. erinaeei Southwell et Macfie, 1925: syn. of Monili- formis cestodiformis (vL.) Travassos, 1917 NEOECHINORHYNCHUS Hamann, 1892 in Stiles et -Mo gracilis (Rudolphi, 1819) Meyer, 1931 (Europe) Hassal, 1905 -M, kalahariensis Meyer, 1931 (South West of Africa) -No acanthuri Farooqi, 1981 (India) -Mo merionis Golvan et Theodorides, 1960 (Iranian -No afghanus Moravec et Arnin, 1978 (Afghanistan) Kurdistan) -No africanus Troncy, 1969 (Chad, Central Africa) -M, moniliformis (Bremser, 1811) Travassos, 1915 -No agilis (Rudolphi, 1819) Van Cleave, 1916 (Japan, (type species)(cosmopolite) Holarctic region) (nee agile) -M. moniliformis sensu Travassos, 19l7: syn. of Monili- -No aldrichettae Edmonds, 1971 (South Australia, New formis travassosi Meyer, 1932 Zealand) -M. moniliformis sensu Sandground, 1926: syn. of -No argentatus Chandra, Rao et Shyamasundari, 1987 Moniliformis dubius Meyer, 1932 (Bay of Bengale) Nomenclature of the Acanthocephala 165 -N. armenicus Mikhailov, 1975 (Armenia) -N.formosanus (Harada, 1938) Kaw, 1951 (Taiwan) -N. asymetricus Belous, 1952 (Eastern Siberia): may be (neeformosanum, neeformosans) a syn. of Neoechinorhynchus tylosuri Yamaguti, 1939 -N. gilbesoni Aly Khan et Bilqees, 1989 (Kalri Lake, (fide YAMAGUTI,1963) (nee asymmetrieus) Sind, Pakistan) -N. australis Van Cleave, 1931 (nee australe) (Missis- -N. glyptosternumi Fotedar et Dhar, 1977 (Cachemire, sippi, Nearctic region) India) (published as a new species in FOTEDAR& KHA- -N. australis Van Cleave, 1931: syn. pro parte of Neo- Roo,1985) echinorhynchus distractus Van Cleave, 1949 -N. golvani Salgado-Maldonado, 1978 (Brazil, Tabasco, -N. australis sensu Hartwich, 1956 (Uruguay, Neotropi- Vera Cruz, Mexico) cal region) is a syn. of Neoechinorhynchus hartwichi -N. graeilisentis (Van Cleave, 1913) Van Cleave, 1916: nom. novo syn. of Gracilisentis gracilisentis (VC.) Van Cleave, -N. bangoni Tripathi, 1959 (Calcutta, India) 1919 -N. buttnerae Golvan, 1956 (Brazilian Amazonia, -N. graptemydis Cable et Hopp, 1954 (USA): it is a Colombia) nomen nudum or a syn. of Neoechinorhynchus emydis -N. carassii Roitman, 1961 (nee Rotman) (Central (L.) Van Cleave, 1919 (fide YAMAGUTI,1963) Asia): may be a syn. of Neoechinorhynchus rutili -N. hartwichi nom. novo (Uruguay, Neotropical region): (O.F.M.) Stiles et Hassal, 1905 (fide SKRJABINA,1979) syn. of Neoeehinorhynehus australis sensu Hartwich, but it needs a new examination 1956 -N. carpiodi Dechtiar, 1968 (Erie Lake) -N. hutchinsoni Datta, 1936 (Cachemire, India) -N. chelonos Schmidt, Esch et Gibbons, 1970 (USA, -N. ichthyobori Saoud, El N affar et Abu Sinna, 1974 South Carolina) (Egypt) -N. chilkaensis Podder, 1937 (India) (nee ehilkaense) -N. idahoensis Amin et Heckman, 1992 (Idaho, USA) (vide GUPTA& JAIN, 1983) -N. johnii Yamaguti, 1939 (East of the China Sea, -N. chrysemydis Cable et Hopp, 1954 (Indiana, USA) Japan, also found in Goa in India) (vide GUPTA& JAIN, -N. cirrhinae Gupta et Jain, 1979 (Chandigarh, India) 1983) (nee johni) -N. coiliae Yamaguti, 1939 (Kiou Shiou, Japan) -N. karachiensis Bilqees, 1972 (Pakistan) -N. constrictus Little and Hopkins, 1968 (USA, Louisia- -N. kashmirensis Fotedar et Dhar, 1977 (Cachemire, na and Texas): may be a syn. of Neoechinorhynchus India) pseudemydis Cable et Hopp, 1954 (fide SCHMIDT, -N. limi Muzzal et Buckner, 1982 (Michigan, USA) ESCH & GIBBONS,1970) -N. lingulatus Nickol et Ernst, 1987 (Florida, USA) -N. crassus Van Cleave, 1919 (Douglas Lake, Michi- -N. longilemniscus Yamaguti, 1954 (Sulawesi) gan, USA) (nee erassum) -N. longirostris (Van Cleave, 1913) Van Cleave, 1916: -N. cristatus Lynch, 1936 (Washington, Pacific Coast of syn. of Tanaorhamphus longirostris (VC.)H.B. Ward, USA) (nee eristatum) 1918 -N. curemai Noronha, 1973 (Para, Brazilian Amazonia, -N. longissimus Farooqi, 1980 (India) Rio de Janeiro) (vide NORONHA,1984) -N. macronucleatus Machado Filho, 1954 (Espirito -N. cyanophlyctis Kaw, 1951 (nee eyanophyetis) (Ca- Santo, Brazil) (nee maeronucleatum) chemire, India) -N. magnapapillatus Johnson, 1969 (USA, Alabama, -N. cylindratus (Van Cleave, 1913) Van Cleave, 1919 North Carolina) (Nearctic region) (nee eylindratum) -N. magnus Southwell et Macfie, 1925 (Queensland, -N. dattai nom. novo (India): syn. of Neoeehinorhyn- Australia) (nee magnum) ehus rutili sensu Datta, 1936 nee O.F. Muller, 1780 -N. manasbalensis Kaw, 1951 (Manasbal Lake, Cache- -N. devdevi (Datta, 1936) Kaw, 1951 (India) mire, India) (nee manasbalense) -N. distractus Van Cleave, 1949 (Nearctic region) (nee -N. nematalosi Tripathi, 1959 (Chilka Lake, India) distraetum) -N. nigeriensis Farooqi, (1980) 1981 (Lagos, , -N. doryphorus Van Cleave et Bangham, 1949 (Florida, basin of River Niger) USA) (nee doryphorum) -N. notemigoni Dechtiar, 1967 (USA, Ontario Lake, -N. edmondsi nom. novo (Heron Island, Queensland): basin of River Mississippi) syn. of Neoeehinorhynehus rutili sensu Edmonds, 1982 -N. octonucleatus Tubangui, 1933 (Luzon, Philippine -N. elongatus Tripathi, 1959 (Chilka Lake, Madras, Islands) (nee oetonecleatum) India): may be a syn. of Neoechinorhynchus chilkaen- -N. oreini Fotedar, 1968 (India) sis Podder, 1937 (fide CHANDRA, REKHA, RAO & -N. ovalis Tripathi, 1959 (Chilka Lake, India) (nee SHYAMASUNDARI,1982) (nee elongatum) ovale) -N. emydis (Leidy, 1851) Van Cleave, 1919 (nee 1916) -N. paraguayensis Machado Filho, 1959 (Rio de Janei- (USA) ro, Rio Paraguay, Brazil) (nee paraguayense) -N. emydis sensu Bravo-Hollis, 1946 (nee Leidy, 1851): -N. paucihamatus (Leidy, 1891) Petrotschenko, 1956: syn. of Neoechinorhynchus emyditoides Fisher, 1960 species impossible to identify (fide V AN CLEAVE, -N. emyditoides Fisher, 1960 (South of the USA, Loui- 1924) (nee paueihamatum) siana, Mexico) -N. prochilodorum Nickol et Thatcher, 1971 (Colombia) 166 Y.J. GOLVAN

-N. prolixoides Bullock, 1963 (USA) NEOGORGORHYNCHUS Golvan, 1960: syn. of Neo- -N. prolixus Van Cleave et Tirnmons, 1952 (Oklahoma, rhadinorhynchus Yamaguti, 1939 USA) (nee prolixum) -N. aspinosus (Fukui et Morisita, 1937) Golvan, 1960: -N. pseudemydis Cable et Hopp, 1954 (Indiana, USA) syn. of Neorhadinorhynchus aspinosus (F. et M.) -N.pterodoridis Thatcher, 1981 (Brazilian Amazonia) Yamaguti, 1939 -N. pungitius Dechtiar, 1971 (Huron Lake, Ontario) -N. nudus (Harada, 1938) Golvan, 1960: syn. of Neo- -N. quinghaiensis Liu Liqing, Wang Baodue et Yang rhadinorhynchus nudus (H.) Yamaguti, 1939 Tao, 1980 (China) -N. robustus Edmonds, 1964: syn. of Sclerocollum -N. rigidus (Van Cleave, 1928) Kaw, 1951 (India)) (nee robustum (E.) Schmidt et Papema, 1978 rigidum, nee rigidis) -N. robertbaueri Amin, 1985 (USA, ) NEONCICOLA Schmidt, 1972 -N. roonwali Datta et Soota, 1963 (India) -N. avicola (Travassos, 1917) Schmidt, 1972 (nee avicu- -N. roseum (sic) Salgado-Maldonado, 1978 (Mexico, la) (Sao Paulo, Brazil) Sinaloa) (nee Salgado et Maldonado) -N. bursata (Meyer, 1931) Schmidt, 1972 (type species) -N. rutili (O.F. Muller, 1780) Hamann, 1892 in Stiles et (Malaysia) Hassal, 1905 (type species) (Holarctic region) -N. curvata (von Linstow, 1897) Schmidt, 1972 [the -N. rutili sensu Datta, 1936 nee O.F. Muller, 1780: syn. geographical origin is certainly erroneous because of Neoechinorhynchus dattai nom. novo there is no lizard of the genus Eumeees in Madagascar, -N. rutili sensu Edmonds, 1982: syn. of Neoechinorhyn- but only in North Africa and Middle East (vide DOLL- ch us edmondsi nom. novo FUS, 1938)] -N. saginatus Van Cleave et Bangham, 1949 (USA) -N. novellae (Parona, 1890) Schmidt, 1972 (nee nove- (nee saginatum) llai) [Puerto Rico, also found in Central Asia (doubt- -N. salmonis Lei Ching, 1984 (British Columbia) ful)]. This species is very poorly described and figured -N. satoi Morisita, 1937: syn. of Gorgorhynchus satoi -N. pintoi (Machado Filho, 1950) Schmidt, 1972 (Rio (M.) Yamaguti, 1963 Grande do SuI, Brazil) -N. simansularis Roitman, 1961 (SSSR): may be a syn. -N. potosi (Machado Filho, 1950) Schmidt, 1972 (Para, of Neoechinorhynchus rutili (O.F.M.) Hamann, 1892 Brazil and Venezuela) (fide SKRJABfNA,1979); it needs a new examination -N. sinensis Schmidt et Dunn, 1974 (China) -N. sinicus Wang Pu Qin, 1966 (Fukien, China) -N. skrjabini (Morosow, 1951) Schmidt, 1972 (Siberia) -N. speetabilis Machado Filho, 1959: syn. of Gorytoce- phalus spectabilis (MF.) Nickol et Thatcher, 1971(fide NEORHADINORHYNCHUS Yamaguti, 1939 NfCKOL& THATCHER,1971) -N. aspinosus (Fukui et Morisita, 1937) Yamaguti, 1939 -N. strigosus Van Cleave, 1949 (basin of the Mississippi (type species) (Sizuoka, Japan) (nee aspinosum) River, Tennessee, Wisconsin, also found in China) (nee -N. atlanticus Gaevskaja et Nigmatullin, 1977 (North of strigosum) the Atlantic Ocean) -N. stunkardi Cable et Fisher, 1961 (USA, Illinois, -N. dogieli (Achmerov et Dombrowskaja-Achmerova, Louisiana) (vide ACHOLONU,1969) 1941) Yamaguti, 1963: syn. of Echinorhynchoides -N. tenellus (Van Cleave, 1913) Van Cleave, 1919 dogieli Achmerov et Dombrowskaja-Achmerova, 1941 (USA and Canada) (nee tenellum) -N. madagascariensis Golvan, 1969 (Madagascar) -N. topseyi Podder, 1937 (Port Canning, Goa, India) -N. myctophumi Mordinova, 1988 (Central Eastern (vide GUPTA& JAfN, 1983) Atlantic) -N. tsintaoensis Morisita, 1937 (Tsintao, China) (nee -N. nudus (Harada, 1938) Yamaguti, 1939 (Taiwan) tsintaoense) (nee nudum) -N. tumidus Van Cleave et Bangham, 1949 (Wisconsin, -N. robustus (Edmonds, 1964) Golvan, 1969: syn. of USA and Saskatchewan, Canada, Siberia) (nee tumidum) Sclerocollum robustum (E.) Schmidt et Paperna, 1978 -N. tylosuri Yamaguti, 1939 [Hamana Lake (Koti), Japan, North Pacific] NEORHADINOTHYNCHOIDES Tripathi, 1959: lapsus -N. variabilis (Diesing, 1856) Meyer, 1932: syn. of Gra- calami pro Neorhadinorhynchus Yamaguti, 1939 cilisentis variabilis (D.) Travassos, Artigas et Pereira, 1928 NEORHYNCHUS Hamann, 1892: preoccupied, syn. of -N. venustus Lynch, 1936 (Washington Lake, Washing- Neoechinorhynchus Hamann in Stiles et Hassal, 1905 ton, USA) (nee venustum) -N. agilis (Rudolphi, 1819) Hamann, 1892: syn. of Neo- -N. wuyiensis Wang, 1981 (China) echinorhynchus agilis (R.) Van Cleave, 1916 -N. yalei (Datta, 1936) Kaw, 1951 (Cachemire, India) -N. eylindratus Van Cleave, 1913: syn. of Neoechi- -N. zacconis Yamaguti, 1935 (Kobata Lake, Japan) norhynchus cylindratus (VC.) Van Cleave, 1919 -N. emydis (Leidy, 1851) Van Cleave, 1913: syn. of NEOGORGORHYNCHOIDES Gupta et Fatma, 1987 Neoechinorhynchus emydis (L.) Van Cleave, 1919 -N. cablei Gupta et Fatma, 1987 (Mandapam; Tamil -N. graeilisentis Van Cleave, 1913: syn. of Gracilisentis Nadu, India) gracilisentis (VC.) Van Cleave, 1919 Nomenclature of the Acanthocephala 167 -N. longirostris Van Cleave, 1913: syn. of Tanaorham- -N. maroeeanus Dollfus, 1951: syn. of Nephridiacan- ph us longirostris (VC.) H.B. Ward, 1918 thus maroccanus (D.) Golvan, 1962 -N. rutili (O.F. Muller, 1780) Hamann, 1892: syn. of -N. palawanensis Tubangui et Masilungan, 1938: syn. of Neoechinorhynchus rutili (O.F.M.) Hamann, 1892 in Nephridiacanthus palawanensis (T. et M.) Golvan, Stiles et Hassal, 1905 1962 -N. tenellus Van Cleave, 1913: syn. of Neoechinorhyn- -N. thapari Sen et Chauhan, 1972: syn. of Nephridia- chus tenellus (VC.) Van Cleave, 1919 canthus thapari (S. et C.) novo comb.

NEORHYNCHUS de Marval, 1905: preoccupied, syn. of NIPPORHYNCHUS Chandler, 1934: syn. of Rhadi- Apororhynchus Shipley, 1899 norhynchus Luhe, 1911 -N. hemignathi (Shipley, 1896) de Marval, 1905: syn. of -N. africanus Golvan, Houin et Deltour, 1963: syn. of Apororhynchus hemignathi (S.)Shipley, 1899 Rhadinorhynchus africanus (G., H. et D.) Golvan, 1969 NEOSENTIS Van Cleave, 1928: subgenus of Pallisen- -N. aspinosus (Fukui et Morisita, 1937) H.L. Ward, tis Van Cleave, 1928 1951: syn. of Neorhadinorhynchus aspinosus (F. et -N. eelatus Van Cleave, 1928 (nee eellatus): syn. of M.) Yamaguti, 1939 Pallisentis (Neosentis) celatus (VC.) Baylis, 1933 -N. atheri Farooqi, (1980) 1981: syn. of Rhadinorhyn- (vide HARADA,1935) ch us atheri (F.)Amin, 1985 -N. cadenati Golvan et Houin, 1964: syn. of Rhadi- NEPHRIDIA CANTHUS Meyer, 1931: fide SCHMIDT norhynchus cadenati (G. et H.) Golvan, 1969 (1972) this genus is a perfect synonym of Oligacan- -N. carangis (Yamaguti, 1939) H.L. Ward, 1951: syn. of thorhynehus, but for BAER (1959) and GOLv AN(1962) Rhadinorhynchus carangis Yamaguti, 1939 such a synonymy cannot be accepted because those -N. ditrematis (Yamaguti, 1939) H.L. Ward, 1951: syn. species are parasitic in entomophagous mammals, pla- of Rhadinorhynchus ditrematis Yamaguti, 1939 ying in the Old World the role played by the genus -N. erumei Gupta et Fatma, 1981 (nee erumeii, nee Fati- Gigantorhynchus in the New World; Nephridiorhyn- ma): syn. of Rhadinorhynchus erumei (G. et F.) Amin, ehus Meyer, 1931 is a synonym 1985 -N. gerberi Baer, 1959 (Zaire) -N. katsuwonis (Harada, 1928) Chandler, 1934: syn. of -N. hamatus (von Linstow, 1897) Meyer, 1932: syn. of Rhadinorhynchus katsuwonis (H.) Van Cleave et Lin- Oligacanthorhynchus hamatus (vL.) Schmidt, 1972 cicome, 1939 -N. kamerunensis Meyer, 1931 (type species) (Came- -N. nudus (Harada, 1938) Van Cleave et Lincicome, run) 1940: syn. of Neorhadinorhynchus nudus (H.) Yama- -N. longissimus Golvan, 1962 (Zaire, Kenya, Central guti, 1939 Africa) -N. ornatus (Van Cleave, 1918) Chandler, 1934: syn. of -N. major (Bremser, 1811 in Westrumb, 1821) Golvan, Rhadinorhynchus ornatus Van Cleave, 1918 (nee 1962 (Europe, Egypt, , Cyprus, Israel, ) ornatum) -N. manisensis Meyer, 1931 (Camerun) -N. ornatus sensu Van Cleave, 1940: syn. of Rhadi- -N. maroccanus Dollfus, 1951 (Morocco) norhynchus vancleavei Golvan, 1969 -N. palawanensis (Tubangui et Masilungan, 1938) Gol- -N. seriolae (Fukui et Morisita, 1937) Yamaguti, 1963: van, 1962 (Philippine Islands) syn. of Rhadinorhynchus seriolae (Y.) Golvan, 1969 -N. shillongensis Sen et Chauhan, 1972: syn. of Oliga- -N. trachuri (Harada, 1935) Van Cleave et Lincicome, canthorhynchus shillongensis (S. et C.)Amin, 1985 1940: syn. of Rhadinorhynchus trachuri Harada, 1935 -N. thapari (Sen et Chauhan, 1972) novo comb. (India): syn. of Nephridiorhynehus thapari Sen et Chauhan, OCTOSPlNIFER Van Cleave, 1919 1972 -0. chandleri (Bullock, 1957) Yamaguti, 1963: syn. of Octospiniferoides chandleri Bullock, 1957 NEPHRIDIORHYNCHUS Meyer, 1931: syn. of Nephri- -0. macilentus Van Cleave, 1919 (type species)(North diacanthus Meyer, 1931 America) (nee maeilentis) -N. gerberi (Baer, 1959): syn. of Nephridiacanthus ger- -0. rohitae Zuberi et Farooqi, 1976 (India) (nee rohitau, beri Baer, 1959 nee Farooq) -N. kamerunensis (Meyer, 1931) Schmidt, 1972: syn. of -0. torosus Van Cleave et Haderlie, 1950 (California, Nephridiacanthus kamerunensis Meyer, 1931 North America) -N. longissimus (Golvan, 1962) Schmidt, 1972: syn. of -0. variabilis (Diesing, 1851) von Kritscher, 1976 Nephridiacanthus longissimus Golvan, 1962 -N. major (Bremser, 1811 in Westrumb, 1821) Meyer, OCTOSPlNIFEROIDES Bullock, 1957 1932: syn. of Nephradiacanthus major (B.) Golvan, -0. australis Schrnidt et Hugghins, 1973 (Colombia) 1962 -0. chandleri Bullock, 1957 (type species) (Texas, -N. manisensis (Meyer, 1931) Schmidt, 1972: syn. of USA) Nephridiacanthus manisensis Meyer, 1931 -0. incognita Schmidt et Hugghins, 1973 (Bolivia) 168 Y.J. GOLVAN OLIGACANTHORHYNCHUS Travassos, 1915: the -0. oti Machado Filho, 1964 (Mato Grosso and Rio de transfer of some species of this genus to Oligacan- Janeiro, Brazil) thorhynchus proposed by SCHMIDT (1972) seems to -0. pardalis (Westrumb, 1821) Schmidt, 1972: syn. of me abusive; the ecology of the intermediate or encyst- Echinopardalis pardalis (W.)Travassos, 1918 ment hosts, and the trophic behaviour of the definitive -0. pomatostomi (Johnston et Cleland, 1911) Tubangui, hosts are very different; we must say that the morpho- 1933: syn. of Oncicola pomatostomi (1. et C.) Schmidt, logical criteria are not sufficient to make an easy dis- 1983 tinction between these closely related genera; the biolo- -0. ricinoides (Rudolphi, 1808) Travassos, 1915 (Euro- gical characters, when they are available, are often of a pe and Middle East) better quality for that purpose (vide BURON, 1986) -0. shillongensis (Sen et Chauhan, 1972) Amin, 1985 -0. (Termitorhynchus) aenigma (Reichensperger, (India ?) 1922) Meyer, 1932 (Brazil) -0. spira (Diesing, 1851) Travassos, 1915 (type species) -0. (sensu lato) amphipachus (Westrumb, 1821) (Neotropical region) Meyer, 1932: may be a syn. of Nephridiacanthus -0. taenioides (Diesing, 1851) Travassos, 1913 (Brazil) major (B.) Golvan, 1962 -0. thumbi Diesing in von Haffner, 1939 (Haiti) -0. atratus (Meyer, 1931) Schmidt, 1972: syn. of Echi- -0. tortuosus (Leidy, 1850) Schmidt, 1972: syn. of nopardalis atrata Meyer, 1931 Hamanniella tortuosa (Leidy, 1850) Van Cleave, 1924 -0. bangalorensis (Pujatti, 1951) Schmidt, 1972: syn. of -0. tumidus (Van Cleave, 1947) Schmidt, 1972: syn. of Echinopardalis bangalorensis Pujatti, 1951 Hamanniella tumida (VC.) Van Cleave, 1953 -0. carinii (Travassos, 1916) Schmidt, 1972: syn. of Hamanniella carinii Travassos, 1916 OLIGOTERORHYNCHUS Monticelli, 1914: may be a -0. cati (Gupta et Lata, 1967) Schmidt, 1972 (Chandi- syn. of Ca vis om a Van Cleave, 1931 (fide PETROTS- garh, India) CHENKO,1956) -0. (sensu lata) circumflexus (Molin, 1858) Meyer, -0. campylurus (Nitzsch, 1857) Monticelli, 1914 (type 1932 (West Europe) species): species dubia (Europe) -0. citilli (Rudolphi, 1806) Kostylev et Zmeew, 1939 -0. magnus Southwell, 1927: syn. of Cavisoma mag- (SSSR) num (S.) Van Cleave, 1931 -0. (sensu lato) compress us (Rudolphi, 1802) Travas- -0. malayensis Tubangui, 1935 (Luzon, Philippine sos, 1917 (Europe) Islands) may be a Prosthorhynchus -0. decrescens (Meyer, 1931) Schmidt, 1972: syn. of Echinopardalis decrescens Meyer, 1931 ONCICOLA Travassos, 1917 -0. erinacei (Rudolphi, 1793) Meyer, 1932 (Europe) -0. angolensis Golvan, 1957: syn. of Pachysentis ango- -0. gerberi (Baer, 1959) Schmidt, 1972: syn. of Nephri- lensis (G.) Baer, 1959 diacanthus gerberi Baer, 1959 -0. atrata (Meyer, 1931) Golvan, 1962: syn. of Echino- -0. hamatus (von Linstow, 1897) Schmidt, 1972 pardalis atrata Meyer, 1931 (Madagascar) -0. bangalorensis (Pujatti, 1951) Golvan, 1962: syn. of -0. iheringi Travassos, 1917 (Brazil, Panama) Echinopardalis bangalorensis Pujatti, 1951 [may be a -0. kamerunensis (Meyer, 1931) Schmidt, 1972: syn. of syn. of Oligacanthorhynchus bangalorensis (P.) fide Nephridiacanthus kamerunensis Meyer, 1931 SCHMIDT(1972)] -0. kamtschaticus Khokhlova, 1966 (East of Siberia) -0. bursata Meyer, 1931: syn. of Neoncicola bursata -0. lagenaeformis (Westrumb, 1821) Travassos, 1917 (M.) Schmidt, 1972 (Europe) -0. campanulata (Diesing, 1851) Meyer, 1931 (Neotro- -0. lamasi (de Freitas et Costa, 1964) Amato, Nicko1 et pical region) Frees, 1979 (Brazil) -0. canis (Kaupp, 1909) Van Cleave, 1921 (Central and -0. lerouxi (Bisseru, 1956) Schrnidt, 1972 (Africa) Northern America, Panama) -0. longissimus (Golvan, 1962) Schmidt, 1972: syn. of -0. chibigouzouensis Machado Fi Iho, 1963 (Mato Nephridiacanthus longissimus Golvan, 1962 Grosso, Brazil) -0. major (Mach ado FiLho, 1963) Schmidt, 1972 (Brazil) -0. confusa (Machado Filho, 1950) Schmidt, 1972 -0. manifestus (Leidy, 1851) Travassos, 1917 (North (Minas Gerais, Brazil) (nee confusus) America) -0. decrescens (Meyer, 1931) Golvan, 1962: syn. of -0. manisensis (Meyer, 1931) Schmidt, 1972: syn. of Echinopardalis decrescens Meyer, 1931 Nephridiacanthus manisensis Meyer, 1931 -0. dimorpha Meyer, 1931 (Africa, West Camerun) -0. mariemily (Tadros, 1969) Arnin, 1985 -0. fraterna Baer, 1959: may be a syn. of Prosthenor- -0. microcephalus (Rudolphi, 1819) Schmidt, 1972: chisfraternus (B.) (fide SCHMIDT,1972) syn. of Hamanniella microcephala (R.) Travassos, -Oi freitasi (Mach ado Filho, 1950) Schmidt, 1972 (Espi- 1917 (nee microcephala) rito Santo, Brazil) -0. minor Machado Filho, 1964 (Minas Gerais, Brazil) -0. gigas Meyer, 1931 (Indonesia) -0. oligacanthus (Rudolphi, 1819) Meyer, 1932 (South -0. juxtatesticularis (Machado Filho, 1950) Schmidt, of Europe) 1972 (Espirito Santo, Brazil) Nomenclature of the Acanthocephala 169 -0. lamasi (de Freitas et Costa, 1964) Schmidt, 1977: -Po ehrenbergi Meyer, 1931 (Egypt) syn. of Oligacanthorhynchus lamasi (dF. et C.) -Po gethi (Machado Filho, 1950) Schmidt, 1972 (Para, Amato, Nickol et Froes, 1979 (fide AMATO,NICKOL& Rio de Janeiro, Brazil; Guerrero, Mexico) FROES, 1979) -Po lenti (Mach ado Filho, 1950) Schmidt, 1972 (Espirito -Oneieola lerouxi (Bisseru, 1956) Golvan, 1957: syn. of Santo, Brazil) Echinopardalis lerouxi Bisseru, 1956 -Po (Oligaeanthorhynehus ?) lerouxi (Bisseru, 1956) -0. luehei (Travassos, 1917) Schmidt, 1972 (Sao Paulo, Baer, 1959: syn. of Echinopardalis lerouxi Bisseru, Para, Minas Gerais, Mato Grosso, Brazil; Panama) 1956 -0. machadoi Schmidt, 1972 (Espirito San to, Brazil) -Po procumbens Meyer, 1931 (Egypt) -0. macrurae Meyer, 1931 (maerurae, page 101, -r. procyonis (Machado Filho, 1950) Schmidt, 1972 maerourae, page 64)(Brazil, observed in (Rio de Janeiro, Brazil; Panama) the Zoological Gardens in U.S.A.) -Po rugosus (Machado Filho, 1950) Schmidt, 1972 (Rio -0. magalhaesi Machado Filho, 1962 (Sao Paulo, Bra- de Janeiro, Brazil) zil) -Po septemserialis (Mach ado Filho, 1950) Schmidt, -0. malayana Toumanoff, 1947 (Indochina) (nee mala- 1972 (Para, Brazil) yanus) -0. martini Schmidt, 1977 (Paraguay) PALLIOLISENTIS Machado Filho, 1960 -0. michaelseni Meyer, 1931 [Indonesia (doutbful)]: -Po ornatus Machado Filho, 1960 (Brazil, Paraguay) may be a Echinopardalis (fide MEYER, 1932) or a syn. -Po polyonca Schmidt et Hugghins, 1973 (Colombia) of Oncicola gigas (fide WITENBERG,1938) -Po quinqueungulis Machado Filho, 1960 (type species) -0. micracantha Machado Filho, 1949 (Rio Grande do (Mato Grosso, Brazil) SuI, Brazil) -0. oncicola (von Ihering, 1902) Travassos, 1917 (type PALLISENTIS Van Cleave, 1928 species) (Brazil) -Po (P.) allahabadi Agarwal, 1958 (Haryana and Allaha- -0. paracampanulata Machado Filho, 1963 (Sao Paulo, bad, India): may be a syn. of P. ophiocephali (T.) Bay- Brazil) lis, 1933 (fide SOOTA& BHATTACHARYA,1982) (fide -0. pardalis (Westrumb, 1821) Travassos, 1917: syn. of CHOWHAN,GUPTA& KHERA, 1987, it is good species) Echinopardalis pardalis (W.) Travassos, 1918 (nee allahabadii) -0. pomatostomi (Johnston et Cleland, 1911) Schmidt, -Po (P.) basiri Farooqi, 1958 (India): may be a syn. of P. 1983 (Australia, Philippine Islands, Borneo, Malaya) colisai Sarkar, 1956 (fide SOOTA & BHATTACHARYA, -0. schacheri Schmidt, 1972 (Lebanon, Israel) 1982) -0. sigmoides (Meyer, 1932) Witenberg, 1938 (Rio de -Po (P.) buckleyi Tadros, 1966 (Aligarh, India): may be Janeiro, Brazil;Surinam) a syn. of P. colisai Sarkar, 1956 (fide SOOTA& BHAT- -0. skrjabini Morosow, 1951: syn. of Neoncicola skrja- TACHARYA,1982) or of P. allahabadi Agarwal, 1958 bini (M.) Schmidt, 1972 (fide SCHMIDT,1972) (fide JAIN & GUPTA, 1979) -0. spirula (von Olfers, 1816 in Rudolphi, 1819) Sch- -Po (P.) cavasii Gupta et Verma, 1980 (Lucknow, North midt, 1972 (Rio de Janeiro, Brazil) India) -0. travassosi Witenberg, 1938 (Israel) -Po (Neosentis) celatus (Van Cleave, 1928) Baylis, 1933 -0. venezuelensis Marteau, 1977 (Venezuela) (type species of the subgenus Neosentis) (Japan, China) (nee cleatus) OWILFORDIA Schmidt et Kuntz, 1967 -Po cholodkowskyi (Kostylew, 1928) Amin, 1985 -0. olseni Schmidt et Kuntz, 1967 (type species) (Tai- -Po (P.) colisai Sarkar, 1954 (India) wan) -Po (P.)fasciati Gupta et Verma, 1980 (Lucknow, North -0. schmidti Gupta et Fatma, 1988 (Uttar Pradesh, India) (nee faseiata) Tamil Nadu, India) -Po (Farzandia) gaboes (McCallum, 1918) Van Cleave, -0. teliger (Van Cleave, 1949) Schmidt et Kuntz, 1967 1928 (type species of the sub genus Farzandia) (Bor- (Java, Indonesia) neo, Sulawesi) (vide YAMAGUTI,1954) -Po garuai (Sahay, Sinha et Ghosh, 1971) Jain et Gupta, PACHICENTIS: errat. pro Pachysentis 1979 (nee Gupta et Fatma, 1986) (Patna, India) -Po (P.) golvani Troncy et Vassiliades, 1973 (Chad, PACHYSENTIS Meyer, 1931 Senegal) -Po angolensis (Golvan, 1957) Baer, 1959 (Angola) -Po (P.) gomtii Gupta et Verma, 1980 (Lucknow, North -Po canicola Meyer, 1931 (type species) (designated by India) VAN CLEAVE, 1953) (Brazil) -Po (P.) guntei Sahay, Nath et Sinha, 1967 (Bihar): may -Po dollfusi (Machado Filho, 1950) Schmidt, 1972 (Bra- be a syn. of P. colisai Sarkar, 1956 (fide SOOTA & zil, but the captive , mainly the Primates in the BHATTACHARYA,1982) Zoological Gardens of the whole world, are often con- -Po guptai Gupta et Fatma, 1986 (India, Uttar Pradesh) taminated and the original geographical origin of this -Po kalriai Aly Khan et Bilqees, 1985 (Pakistan) now cosmopolite species is probably not Madagascar) -Po (P.) magnum Saeed et Bilqees, 1971 (India): may be 170 Y.J. GOLVAN a syn. of P. ophiocephali (Thapar, 1931) Baylis, 1933 PARA CA VISOMA von Kritscher, 1957 (fide SOOTA& BHATIACHARYA,1982) -Po impudica (Diesing, 1851) von Kristcher, 1957 (type -Po mehrai Gupta et Fatma, 1986 (India, Uttar Pra- species) (Mato Grosso, Brazil; Peru) (nee impudi- desh) cum) -Po (Farzandia) nagpurensis (Bhalerao, 1931) Baylis, 1933 (Penjab, India, Burma, Sri Lanka) PARADOXITES Lindemann, 1865: preoccupied, syn. of -Po (P.) nandai Sarkar, 1953: may be a syn. of P. ophio- Centrorhynchus Luhe, 1911 cephali (T.) Baylis, 1933 (fide SOOTA & BHATTA- -Po renardi Lindemann, 1865: syn. of Centrorhynchus CHARYA, 1982) renardi (L.) Van Cleave, 1923 -Po (Neosentis) ophiocephali (Thapar, 1931) Baylis, 1933 (India) PARAECHINORHYNCHUS Bilqees et Aly Khan, -Po (P.) panadei Rai, 1967 (Mathura, India) (pandei in 1983 GEORGE& NADAKAL,1973): may be a syn. of P. coli- -Po kalriai Bilqees et Aly Khan, 1983 (type species) sai Sarkar, 1956 (fide SOOTA & BHATTACHARYA, (Ka1ri Lake, Sind, Pakistan) 1982) -Po sindensis Aly Khan et Bilqees, 1987 (Kalri Lake, PARAFlLICOLLIS Petrotschenko, 1956: syn. of Profili- Pakistan) collis Meyer, 1931 -Po (P.) tetraodontae Troncy, 1970 (Chad) -Po altmani (Perry, 1942) Petrotschenko, 1958: syn. of -Po (P.) umbellatus Van Cleave, 1928 (type species of Profilicollis altmani (P.) Khokhlova, 1974 the genus Pallisentis) (Pekin, Wuchang, China) -Po kenti (Van Cleave, 1947) Petrotschenko, 1958: syn. -Po (?) ussuriensis (Kostylew, 1941) Golvan, 1959: syn. of Profilicollis kenti (VC.) Khokhlova, 1974 of Acanthocephalorhynchoides ussuriensis Kostylew, -Po major (Lundstrom, 1942) Petrotschenko, 1958: syn. 1941 of Profilicollis major (L.) Khokhlova, 1974 -Po sphaeroeephalus (Bremser in Rudolphi, 1819) PANDOSENTIS Van Cleave, 1920 (nee Pandoneentis) Petrotschenko, 1958: syn. of Profilicollis sphaeroce- -Po iracundus Van Cleave, 1920 (type species) (Mara- phalus (B. in R.) Khokhlova, 1974 cay, Venezuela) PARAGORGORHYNCHUS Golvan, 1957 -Po albertianus Golvan, 1957 (type species) (Zaire) (nee PARACANTHOCEPHALOIDES Golvan, 1969 albertianum) -Po chabanaudi (Dollfus, 1951) Golvan, 1969 (type spe- -Po chariensis Troncy, 1969 (basin of the Chari River, cies) (Zanzibar) Chad) -Po golvani Chandra, Hanumantha Rao et Shyamasun- dari, 1984 (Bengale) PARALONGICOLLUM Amin, Bauer et Sidorov, 1991 -Po plotosi (Yamaguti, 1935) Golvan, 1969 (Mie, Japan -Po nemacheili Amin, Bauer et Sidorov, 1991 (type spe- et Chilka Lake, Port Canning, India) cies) (Balkhash Lake, Kazakhstan) -Po soleae (Porta, 1905) Paggi et Orecchia, 1983: syn. of -Po sergenti (Choquette et Gayot, 1952) Amin, Bauer et Solearhynchus soleae (Porta, 1905) de Buron et Mai- Sidorov, 1991 (North Africa) lIard, 1985 -Po tripathii Go1van, 1969 (India) PARARHADINORHYNCHUS Johnston et Edmonds, 1947 PARACANTHOCEPHALUS Achmerov et Dombrows- -Po aspinosus (Fukui et Morisita, 1937) Petrotschenko, kaja-Achmerova, 1941: syn. of Acanthocephalus 1956: syn. of Neorhadinorhynchus aspinosus (F. et Koelreuther, 1771 M.) Yamaguti, 1939 -Po eurtus Achmerov et Dombrowskaja-Achmerova, -Po coorongensis Edmonds, 1973 (Australia) 1941: syn. of Acanthocephalus curtus (A. et D.-A.) -Po mugilis Johnston et Edmonds, 1947 (type species) Yamaguti, 1963 (South Australia) -Po graeitaeanthus (Meyer, 1932) Grabda et Grabda- Kazubska, 1967: syn. of Acanthocephalus gracila- PARATENUISENTIS Bullock et Samuel, 1975 canthus Meyer, 1932 -Po ambiguus (Van Cleave, 1921) Bullock et Samuel, -Po rausehi (Go1van, 1969) Schmidt, 1969: syn. of 1975 (type species) (Maryland, East of USA) Acanthocephalus rauschi Golvan, 1969 -Po tenuirostris Achmerov et Dombrowskaja-Achmero- PARDALIS Travassos, 1917: preoccupied, syn. of Echi- va, 1941: syn. of Acanthocephalus tenuirostris (A. et nopardalis Travassos, 1918 D.-A.) Yamaguti, 1963 -Po pardalis (Westrumb, 1821) Travassos, 1917: syn. of Oncicola campanulata (D.) Meyer, 1931 or of Oliga- PARACANTHORHYNCHUS Edmonds, 1967 canthorhynchus pardalis (W.) Schmidt, 1972 -Po galaxiasus Edmonds, 1967 (type species) (South Australia) PARIDOSENTlS: errat. pro Pandosentis Nomenclature of the Acanthocephala 171 PAR V/CA V/SOMA: errat. pro Paracavisoma -Po lemnisalis Belopolskaja, 1959 (SSSR) (nee lemnis- calis) PAULISENTlS Van Cleave et Bangham, 1949 -Po limnobaeni (Tubangui, 1933) Golvan, 1956 (Luzon, -P.fractus Van Cleave et Bangham, 1949 (type species) Philippine Islands) (Ohio, Indiana, USA) -Po linearis (Westrumb, 1821) Golvan, 1956 (Europe) -Po missouriensis Keppner, 1974 (basin of the (nec lineare) River, Nebraska, USA) -Po longirostris (Travassos, 1926)Amin, 1985: syn. of Prosthorhynchus longirostris T. PILUM Williams, 1976 -Po (Prosthorhynehus) malayensis (Tubangui, 1935) -Po pilum Williams, 1976 (type species) (Alabama, Schmidt et Kuntz, 1966 (nee malayense): syn. of USA) Prosthorhynchus malayensis Tubangui, 1935 -Po menurae (Johnston, 1912) Golvan, 1956 (New PLAGIORHYNCHUS Lilhe, 1911: the synonymy of the South Wales, Australia) genera Plagiorhynchus and Prosthorhynchus propo- -Po nicobarensis (Soota et Kansal, 1972) Zafar et Faroo- sed by SCHMIDT&KUNTZ (1966) cannot be admitted qi, 1981: syn. of Prosthorhynchus nicobarensis Soota because the first genus contains acanthocephalans para- et Kansal, 1972 sitic in aquatic arthropods and which become adults in -Po odhneri Lundstrom, 1942 (Scandinavia, SSSR) limicole , whereas the acanthocephalans of the -Po (Prosthorhynchus) ogatai (Fukui et Morisita, 1936) second genus develop in terrestrial arthropods and are Schmidt et Kuntz, 1966: syn. of Prosthorhynchus oga- adults in «terrestrial» birds; this important ecological tai (F. et M.) novo comb. difference is supported by the egg morphology; Pla- -Po paulus Van Cleave et Williams, 1951(Alaska) (nec giorhynchus has fusiform eggs with polar prolonga- paulum) tions, whereas the species of the «terrestrial» genus -Po (Prosthorhynchus) pittarum (Tubangui, 1935) Sch- Prosthorhynchus present «short oval shaped» eggs midt et Kuntz, 1966: syn. of Prosthorhynchus pitta- always lacking polar prolongations rum Tubangui, 1935 -Po (Prosthorhynehus) angrense (Travassos, 1926) Sch- -Po pupa (von Linstow, 1905) novo comb. (West of midt et Kuntz, 1966: syn. of Prosthorhynchus angren- Tairnyr Peninsula): syn. of Echinorhynchus pupa von sis Travassos, 1926 Linstow, 1905 -Po bullocki Schmidt et Kuntz, 1966: syn. of Prostho- -Po rectus (Linton, 1892) Van Cleave, 1918 (nec rhynchus bullocki (S. et K.) novo comb. Sprehn, 1942) (Mexico) -Po charadrii (Yamaguti, 1939) Van Cleave, 1951 (nee -Po reticulatus (Westrumb, 1821) Golvan, 1956 (nec Golvan, 1956)(Taiwan, Japan, Pescadores Islands, retieulatum) (Neotropical region) Oceania) -Po rheae (de Marval, 1902) Schmidt et Kuntz, 1966: -Po charadriicola (Dollfus, 1953) Golvan, 1956 (North syn. of Prosthorhynchus rheae (dM.) Travassos, 1926 Africa) -Po rossicus (Kostylew, 1915) Schmidt et Kuntz, 1966: -Po crassicollis (Villot, 1875) Luhe, 1911 (type species) syn. of Prosthorhynchus rossicus Kostylew, 1915 (North Europe) (nee erassieolle) -Po rostratus (de Marval, 1902): species incertae sedis -Po (Prosthorhynehus) cylindraceus (Goeze, 1782) Sch- (nec rostratum) midt et Kuntz, 1966: syn. of Prosthorhynchus cylin- -Po russelli Tadros, 1970: syn. of Prosthorhynchus rus- draceus (G.) Golvan, 1956 selli (T.) novo comb. -Po formosus Van Cleave, ] 918: syn. of Prosthorhyn- -Po scolopacidis (Kostylew, 1915) Schmidt et Kuntz, chus formosus (VC.) Travassos, 1926 (may be a syn. 1966: syn. of Prosthorhynchus scolopacidis Kostylew, of P. cylindraceusfide SCHMIDT,1981) 1915 -Po freitasi Vicente, 1977 (Neotropical region) -Po spiralis (Rudolphi, 1809) Golvan, 1956 (Europe) -Po (Prosthorhynehus) gallinagi (Schachtachtinskaja, -Po taiwanensis Schmidt et Kuntz, 1966: syn. of Pros- 1953) Schmidt et Kuntz, 1966: syn. of Prosthorhyn- thorhynchus taiwanensis (Schmidt et Kuntz, 1966) ch us gallinagi Schachtachtinskaja, 1953 novo comb.; may be a syn. of Prosthorhynchus (Pla- -Po genitopapillatus (Lundstrom, 1942) Amin, 1985: syn. giorhynchus ???) cylindraceus (Goeze, 1782) (fide of Prosthorhynchus genitopapillatus Lundstrorn, 1942 SCHMIDT,1981, but it is very doubtful) -Po golvani Schmidt et Kuntz, 1966: syn. of Prostho- -Po totani (Porta, 1910) Golvan, 1956 (Italy) rhynchus golvani (S. et K.) novo comb. -Po urichi (Cameron, 1936) Golvan, 1956 (Trinidad): -Po gracilis (Petrotschenko, ] 958) Schmidt et Kuntz, species incertae sedis 1966: syn. of Prosthorhynchus gracilis Petrotschenko, 1958 PLEURORHYNCHUS Nau , 1787: preoccupied, syn. of -Po kuntzi Gupta et Fatma, 1988 (Uttar Pradesh, Tamil Leptorhynchoides Kostylew, 1924 Nadu, India) -Po laneeolatus (von Linstow, 1876) Liihe, 1911: may be POLYACANTHORHYNCHUS Travassos, (1919) 1920 a syn. of Plagiorhynchus crassicollis (Villot, 1875) (nec 1926) Luhe, 1911 -Po arcuatum (Diesing, 1851) Travassos, 1920/21: syn. 172 Y.J. GOLVAN of Polyacanthorhynchus macrorhynchus (D.) Travas- -Po (Subcorynosoma) corynoides Skrjabin, 1913 (Cen- sos, 1920 tral Asia) -Po caballeroi Diaz-Ungrfa et Gracia-Rodrigo, 1960 -Po eorynosoma Travassos, 1915: syn. of Hexaglandula (Venezuelan Amazonia) corynosoma (T.) Petrotschenko, 1958 -Po kenyensis Schmidt et Canaris, 1967 (Naivasha Lake, -Po (Polymorphus) crassus Van Cleave, 1924 (Nearctic Kenya) region) -Po macrorhynchus (Diesing, 1851) Travassos, (1919) -Po (Polymorphus) cucullatus Van Cleave et Starrett, 1920 (nee 1856, nee Baylis, 1927)(type species) (Bra- 1940 (Illinois, USA) zilian Amazonia) -Po (Polymorphus) diploinflatus Lundstrorn, 1942 -Po megaeephalus (Westrumb, 1821) Travassos, (Scandinavia, SSSR) 1920121: syn. of Centrorhynchus tumidulus (R.) -Po eroliae Yamaguti, 1939: syn. of Arhythmorhynchus Neiva da Cunha et Travassos, 1914 eroliae (Y.) Schmidt, 1973 -Po megaloeephalus (Westrumb, 1821) Travassos, -Po (Profilieollis)formosus Schmidt et Kuntz, 1967: syn. 1920121: vide P. megacephalus of Profilicollis formosus (S. et K.) Khokhlova, 1974 -Po rhopalorhynchus (Diesing, 1851) Travassos, 1920 (?) (Mato Grosso, Amazonia) -Po frontospinosus Tubangui, 1935: syn. of Arhyth- morhynchusfrontospinosus (T.) Yamaguti, 1963 POLYMORPHUS Luhe, 1911 -Po (Subfilicollis) gavii Khokhlova, 1965 (SSSR) -Po (Polymorphus) actuganensis Petrotschenko, 1949 -Po inermis Travassos, 1923 (nee inerme): syn. of Hexa- (SSSR) glandula inermis (T.) Petrotschenko, 1958 -Po (Polymorphus) acutus Van Cleave et Starrett, 1940 -Po kenti Van Cleave, 1947: syn. of Profilicollis kenti (nee aeutis) (Illinois, Nearctic region, also found in (VC.) Khokhlova, 1974 SSSR) -Po (Polymorphus) kostylewi Petrotschenko, 1949 -Po altmani (Perry, 1942) Van Cleave, 1947: syn. of (SSSR) Profilicollis altmani (P.) Khokhlova, 1974 -P. (Polymorphus) magnus Skrjabin, 1913 (Europe) -Po aretieus (Van Cleave, 1920) Van Cleave, 1937: syn. .cp major Lundstrorn, 1942: syn. of Profilicollis major of Profilicollis arctic us (VC) Meyer, 1931 (L.) Khokhlova, 1974 -Po arctocephali Smales, 1986 (nee aretoeephalus) -Po (Polymorphus) marchii (Porta, 1910) Meyer, 1932 (Victoria, Australia; Philippine Islands) (Italy) -Po ardeae Belopolskaja, 1958: may be a syn. of South- -Po (Polymorphus) marilis Van Cleave, 1939 (Oklaho- wellina ardeae (B.) novo comb. ma, Illinois, Nearctic region) -Po (Polymorphus) biziurae Johnston et Edmonds, 1948 -Po (Polymorphus) mathevossianae Petrotschenko, (Murray riv., Australia) 1949 (nee mathevossiaenae) (SSSR) -Po boschadis (Schrank, 1788) Railliet, 1919 (it is the -Po (Polymorphus) meyeri Lundstrorn, 1942 (Scandina- type species fide YAMAGUTI,1963) (North Eurasia) via) -Po botulus (Van Cleave, 1916) Van Cleave, 1937 (vide -Po (Polymorphus) miniatus (von Linstow, 1896) Tra- VAN CLEAVE, 1939): syn. of Profilicollis botulus vassos, 1926 (Tierra del Fuego) (VC) Meyer, 1932 -Po (Polymorphus) minutus (Goeze, 1782) Luhe, 1911 -Po brevis (Van Cleave, 1916) Travassos, 1926: syn. of (type species) (North of Eurasia) Arhythmorhynchus brevis Van Cleave, 1916 -Po mutabilis (Rudolphi, 1819) Travassos, 1926: syn. of -Po eapellae Yamaguti, 1935: syn. of Arhythmorhyn- Hexaglandula mutabilis (R.) Petrotschenko, 1950 ch us capellae (Y.) Van Cleave et Rausch, 1950 (nee -Po (Polymorphus) obtusus Van Cleave, 1918 (USA, Schrnidt, 1973) Florida) -Po caryosoma Travassos, 1915 (Brazil) -Po (Polymorphus) paradoxus Connell et Corner, 1957 -Po cetaceum sensu Schmidt et Dailey, 1971: probably (Alberta, Canada) is not a true synonym of Corynosoma cetaceum Johns- -Po paueihamatus Heinze, 1936 (nee paueihamata): syn. ton et Best, 1942 because the acanthocephalans exami- of Hexaglandula paucihamatus (H.) Petrotschenko, ned by SCHMIDT& DAILEY (1971) were from the Rio 1958 de la Plata (Uruguay) and the C. cetaceum sensu stricto -Po petrotchenkoi (sic) Schmidt, 1969 (Alaska): syn. of of JOHNSTON& BEST(1942) were from South Australia! Arhythmorhynchus petrotchenkoi (Schrnidt, 1969) -Po (Profilieollis) ehasmagnathi (Hoicman-Spector, Atrashkevitch, 1979 Mane-Garzon et Dei-Cas, 1977) Amin, 1985: syn. of -Po (Subfilicollis) phippsi Kostylew, 1922 (Alaska, Profilicollis chasmagnathi (HS., MG. et DC) novo Strait of Behring, Tshouktschis, Mourman): may be a comb. syn. of Polymorphus boschadis Railliet, 1919 (fide -Po (Polymorphus) cincli Belopolskaja, 1958 (East VANCLEAVE& RAUSCH,1951) Siberia) -Po piriformis (Bremser, 1811 in Rudolphi, 1819) -Po contortus (Bremser in Westrumb,1821) Travassos, Machalska, 1981 (Europe) 1926 (Europe) (probably this species does not belong -Po (Subfilicollis) pupa (von Linstow, 1905) (nee to the genus Polymorphus) Kostylew, 1922) (North Europe): this species cannot be Nomenclature of the Acanthocephala 173 identified and must become a species delendafide VAN -Po tereticollis (Rudo1phi, 1809) Meyer, 1932 (Nearctic CLEAVE&RAUSCH(1951) region, found also in Ireland) (nee teretieolle) -Po sphaeroeephalus (Bremser, 1811 in Rudolphi 1819) -Po tori Fotedar et Dhar, 1974 (Cachernire, India) Van Cleave, 1947: syn. of Profilicollis sphaerocepha- -Po turbinella (Diesing) in Leiper et Atkinson, 1915 is a Ius (B.) Khokhlova, 1974 Bolbosoma different from Bolbosoma turbinella (D.) -Po spindlatus Arnin et Heckmann, 1991 (Peru) Porta, 1908 (fide MEYER, 1933) -Po (Polymorphus) striatus (Goeze, 1782) Luhe, 1911 -Po yamagutii Schrnidt et Hugghins, 1973 (Chili) (North of Eurasia) -Po yunnanensis Wang, 1981 (China) -Po (Subcorynosoma) strumosoides Lundstrom, 1942 (Scandinavia, SSSR) PORRORCHIS Fukui, 1929 -Po (Subcorynosoma) swartzi Schmidt, 1965 (Alaska) -Po bazae (Southwell et Macfie, 1925) Schmidt et -Po texensis Webster, 1948: syn. of Profilicollis texensis Kuntz, 1967 (Queensland, Australia) (W.) Khokhlova, 1974 -Po brevicanthus (Das, 1949) novo comb. (India): syn. -Po trochus Van Cleave, 1945 (Ohio, New York, Illi- of Centrorhynehus brevieanthus Das, (1949) 1950 nois, Nearctic region) -Po centropi (Porta, 1910) Schmidt et Kuntz, 1967 (Gui- nea Bissau, West Africa) POMPHORHYNCHUS Monticelli, 1905 -Po centropusi (Tubangui, 1933) Petrotschenko, 1958: -Po bosniacus Kiskaroly et Cankovic, 1969 (South of may be a syn. of Porrorchis hylae (1.) (fide SCHMIDT& Europe) KUNTZ, 1967) -Po bufonis Fotedar, Duda et Raina, 1970 (Srinagar, -Po chauhani Gupta et Fatman, 1986 (India, Lucknow) Cachemire) -Po crocidurai Gupta et Fatma, 1986 (India, Lucknow) -Po bulbocolli Linkins in Van Cleave 1919 (Nearctic -Po elongatus Fukui, 1929 (type species) (Japan, Tai- region; Campeche, Mexico) wan, Philippine Islands) -Po cylindrica Wang Pu Qin et Guo Qizhi, 1983 (Yun- -Po houdemeri (Joyeux et Baer, 1935) Schmidt et Kuntz, nan, China) (nee cylinderica: the spelling cylinderica 1967 (Indochina) used on page 99 for the explanation of the drawing and -Po hydromuris (Edmonds, 1957) Schmidt et Kuntz, on page 102 and the spelling cylinderius which appears 1967 (Australia) only on page 98 must be deleted; the correct name -Po hylae (Johnston, 1914) Schmidt et Kuntz, 1967 which must be adopted for this species is eylindriea) (Queensland, Australia, Philippine Islands, Taiwan; -Po dubious Kaw, 1941 (India) (nee dubium) also found in India) -Po francoisae Gol van, 1969 (Morocco) -Po indicus (Das, 1957) Schmidt et Kuntz, 1967 (Chan- -Po indicus Gupta et Lata, 1967 is a syn. of Centrorhyn- digarh, India) chus guptai Golvan, 1969 -Po keralensis George et Nadakal, 1984 (Kerala, India) -Po intermedius Engelbrecht, 1957 (Central Europe) -Po leibyi Schmidt et Kuntz, 1967 (Taiwan, Palawan, -Pi jammuensis Fotedar et Dhar, 1974 (Jammu, India) Philippine Islands) -Po kashmirensis Kaw, 1941 (Cachemire, India) (nee -Po maxvachoni (Golvan et Brygoo, 1965) Schrnidt et kashmirense) Kuntz, 1967 (Madagascar) -Po kawi Fotedar, Duda et Raina, 1970 (Cachemire, -Po ogatai Fukui et Morisita, 1936: syn. of Prosthorhyn- India) chus ogatai (F. et M.) novo comb. -Po kostylewi Petrotschenko, 1956 (Armenia) -Po oti Yamaguti, 1939 (nee goti) (Japan) -Po laevis (Zoega in O.F. Miiller, 1776) Van Cleave, -Po rotundatus (von Linstow, 1897) Schmidt and Kuntz, 1924 (nee laeve) (type species) (Palaearctic region) 1967 (Madagascar) -Po lucyi Williams et Rogers, 1984 (Floride, South of the -Po teliger Van Cleave, 1949: syn. of Owilfordia teliger USA) (VC.) Schmidt et Kuntz, 1967 -Po megacanthus Fotedar et Dhar, 1974 (Cachemire, India) -Po oreini Fotedar et Dhar, 1974 (India) PROFILICOLLIS Meyer, 1931: it is a valid genus para- -Po orientalis Fotedar et Dhar, 1974 (Cachernire, India) sitic in the marsh birds with Decapod as -Po patagonicus Ortubay, Ubeda, Semenas et Kennedy, intermediate hosts and not a synonym of Polymorphus 1991 (Patagonia, Argentina) Liihe, 1911 (fide KHOKHLOVA,1974) -Po perforator (von Linstow, 1908) Meyer, 1932 (Tur- -Po altmani (Perry, 1942) Khokhlova, 1974 (USA, Cali- kestan) fornia) -Po proteus (Westrumb, 1821) Porta, 1905: syn. of -Po antarcticus Zdzitowiecki, 1985 (South Shetlands) Pomphorhynchus laevis (Z.) Van Cleave, 1924 pro -Po arcticus (Van Cleave, 1920) Meyer, 1932 (North parte and of Pomphorhynchus tereticollis (R.) Meyer, West Territory, Canada) 1932 pro parte -Po botulus (Van Cleave, 1916) Meyer, 1932 (type spe- -Po rocci Cordonnier et H.L. Ward, 1967 (USA, North cies) (Maine, USA, Spitzberg Islands, North of Europe) Carolina) -Po chasmagnathi (Holcman-Spector, Mane-Garzon et -Po sebastichthydis Yamaguti, 1939 (Japan) Dei-Cas, 1977) novo comb. (Rio de la Plata): syn. of 174 Y.J. GOLVAN Falsifilicollis chasmagnathi Holcman-Spector, Mane- -Po juxtatesticularis Machado Filho, 1950: syn. of Onci- Garzon et Dei-Cas, 1977 and of Polymorphus (Profili- colajuxtatesticularis (MF.) Schmidt, 1972 collis) chasmagnathi (Holcman-Spector, Mane-Garzon -Po lemuri Machado Filho, 1950 (these worms were ori- et Dei-Cas, 1977) Amin, 1985 ginally found in the Zoological Garden in Paris and not -Po formosus (Schmidt et Kuntz, 1967) Khokhlova, in Madagascar as was wrongly written. Now, it seems 1974 (Taiwan) present in all the zoological parks around the world, -Po kenti (Van Cleave, 1947) Khokhlova, 1974 (New where it causes severe enzootia, especially among cap- Brunswick, Canada) tive Primates) -Po major (Lundstrom, 1942) Khokhlova, 1974: (Scan- -Po lenti Machado Filho, 1950: syn. of Pachysentis lenti dinavia, SSSR) (MF.) Schmidt, 1972 -Po sphaerocephalus (Bremserin Rudolphi, 1819) -Po luehei Travassos, 1916: syn. of Oncicola luehei (T.) Khokhlova, 1974 (Australia, South America) Schmidt, 1972 -Po texensis (Webster, 1948) Khokhlova, 1974 (South of -Po (sensu lato) novellae (Parona, 1890) Travassos, the USA) 1917: syn. of Neoncicola novellae (P.) Schmidt, 1972 -Po pachyacanthus (Sonsino, 1889) Travassos, 1917 PROMONILIFORMIS Dollfus et Golvan, 1964 (Egypt): species inquirenda -Po ovocristatus (von Linstow, 1897) Dollfus et Golvan, -Po pardalis Southwell et Macfie, 1925 (Sierra Leone): 1964 (type species) (Madagascar) species dubia -Po pintoi Machado Filho, 1950: syn. of Neoncicola PROSTENORCHIS: errat. pro Prosthenorchis pintoi (MF.) Schmidt, 1972 (vide SCHMIDT& DUNN, 1974) PROSTHENORCHIS Travassos, 1915 -Po potosi Machado Filho, 1950 (Brazil, Para): may be a -Po avicola Travassos, 1917 (nec avicula): syn. of Neon- syn. of Neoncicola potosi (MF.) Schmidt, 1972 cicola avicola (T.) Schmidt, 1972 -Po procyonis Machado Filho, 1950: syn. of Pachysentis -Po (sensu lato) circumflexus (Molin, 1858) Travassos, procyonis (MF.) Schmidt, 1972 1915: syn. of Oligacanthorhynchus circumflexus (M.) -Po rugosus Machado Filho, 1950: syn. of Pachysentis Meyer, 1932 rugosus (MF.) Schmidt, 1972 -Po confusus Machado Filho, 1950: syn. of Oncicola -Po (sensu lato) semoni (van Linstow, 1898) Travassos, confusa (MF.) Schmidt, 1972 1917: syn. of Australiformis semoni (vL.) Schmidt et -Po curvatus (van Linstow, 1897) Travassos, 1917: syn. Edmonds, 1989 of Neoncicola curvata (vL.) Schmidt, 1972 -Po septemserialis Machado Filho, 1950: syn. of Pachy- -Po dollfusi Machado Filho, 1950: syn. of Pachysentis sentis septemserialis (MF.) Schmidt, 1972 dollfusi (MF.) Schmidt, 1972 -Po sigmoides Meyer, 1932: syn. of Oncicola sigmoides -Po elegans (Diesing, 1851) Travassos, 1915 (type spe- (M.) Schmidt, 1972 cies) (Rio de Janeiro, Para, Brazil; Panama) (vide SCH- -Po sinicus Hu, 1990 (China) MIDT,1972) -Po spirula (van Olfers, 1816 in Rudolphi, 1819) Travas- -Po elegans sensu Travassos, 1917: syn. of Pachysentis sos, 1917: syn. of Oncicola spirula (0.) Schmidt, 1972 gethi (MF.) Schmidt, 1972 pro parte, of Pachysentis (vide DOLLFUS,1938) lenti (MF.) Schmidt, 1972 pro parte, of Neoncicola -Po spirula sensu Travassos, 1917 pro parte: syn. of Onci- pintoi (MF.) Schmidt, 1972 pro parte and of Oncicola cola confusa (Machado Filho, 1950) Schmidt, 1972 sigmoides (M.) Schmidt, 1972 pro parte -Po spirula sensu Porta, 1909 is another species found in -Po elegans sensu Stiles et Baker, 1935: syn. of Neonci- Egypt cola pintoi (MF.) Schmidt, 1972 -P. spirula sensu Dollfus, 1938: syn. of Pachysentis -Po elegans sensu Dollfus, 1938: syn. of Prosthenorchis dollfusi (Machado Filho, 1950) Schmidt, 1972 lemuri Machado Filho, 1950 -Po travassosi Machado Filho, 1950: syn. of Oncicola -Po erinacei (Rudolphi, 1793) Travassos, 1917: syn. of machadoi Schmidt, 1972 Oligacanthorhynchus erinacei (R.) Meyer, 1932 -Po fraterna (Baer, 1959) Schmidt, 1972 (geographical PROSTHORHYNCHUS Kostylew, 1915 origin unknown ?) -Po angrensis Travassos, 1926 (nec angrense) (Neotro- -Po freitasi Machado Filho, 1950: syn. of Oncicola frei- pical region) tasi (MF.) Schmidt, 1972 -Po asturi (Gupta et Lata, 1967) novo comb. (Penjab, -Po gethi Machado Filho, 1950: syn. of Pachysentis India): syn. of Rhadinorhynchus asturi Gupta et Lata, gethi (MF.) Schmidt, 1972 1967 -Po (sensu lato) hamatus (von Linstow, 1897) Travassos, -Po asymetricus Belopolskaja, 1983 (coasts of the Baltic 1917: syn. of Nephridiacanthus hamatus (vL.) Meyer, Sea) 1932 -Po avicola Travassos, 1917: syn. of Neoncicola avicola -Po (sensu lato) ingens (van Linstow, 1897) Travassos, (T.) Schmidt, 1972 1917: syn. of Macracanthorhynchus ingens (vL.) -Po bazae (Southwell et Macfie, 1925) Travassos, 1926: Meyer, 1932 syn. of Porrorchis bazae (S. et M.) Schmidt et Kuntz, Nomenclature of the Acanthocephala 175 1966 (fide SCHMIDT & KUNTZ, 1966 it is a species -Po pittarum Tubangui, 1935 (Luzon, Philippine incertae sedis) Islands) -Po bulboeaudatus (Southwell et Macfie, 1925) Travas- -Po pupa (von Linstow, 1905) Meyer, 1932 (Taimyr): sos, 1926: syn. of Porrorchis hylae (1.) Schmidt et this species cannot be identified (fide VANCLEAVE& Kuntz,1967 RAUSCH,1951), but (fide KOSTYLEW,1922) it may be a -Po bullocki (Schmidt et Kuntz, 1966) novo comb.(Tai- Polymorphus wan): syn. of Plagiorhynchus bullocki Schmidt et -Po rectus (Linton, 1892) Travassos, 1926: syn. of Pla- Kuntz,1966 giorhynchus rectus (Linton, 1892) Van Cleave, 1918 -Po charadrii Yamaguti, 1939: syn. of Plagiorhynchus (vide VAN CLEAVE, 1931); species incertae sedis (fide charadrii (Y.) Van Cleave, 1951 SCHMIDT& KUNTZ, 1966) -Po charadrii sensu Johnston et Edmonds, 1947, from -Po rectus Sprehn, 1942 nee Linton, 1892: syn. of Pros- Queensland, is probably another species thorhynchus schmidti nom. novo -Po charadriicola Dollfus, 1953: syn. of Plagiorhyn- -Po reticulatus (Westrumb, 1821) Travassos, 1926 (nee ch us charadriicola (D.) Golvan, 1956 retieulatum): syn. of Plagiorhynchus recticulatus -Po cylindraceus (Goeze, 1782) Golvan, 1956 (type spe- (Westrumb, 1821) Golvan, 1956 cies) (Eurasia) -Po rheae (de Marval, 1902) Travassos, 1926 (Neotropi- -Po dimorphocephalus (Westrumb, 1821) Travassos, cal region) 1926: syn. of Centrorhynchus dimorphocephalus (W.) -Po rosai (Porta, 1910) Meyer, 1932: syn. of Pros- Meyer,1932 thorhynchus cylindraceus (G.) Golvan, 1956 -Po erinacei (Rudolphi, 1893) Stiles et Stanley, 1932: -Po rossicus Kostylew, 1915 (SSSR) syn. of Oligacanthorhynchus erinacei (R.) Meyer, -Po rostratus (de Marval, 1902) Meyer, 1932 (Dniepo- 1932 petrovsk, East Europe) -Po formosus (Van Cleave, 1918) Travassos, 1926 (New -Po russelli (Tadros, 1969) novo comb. (United King- York, Nearctic region) (vide VA CLEAVE, 1931): may dom): syn. of Plagiorhynchus russelli Tadros, 1970 be a syn. of Prosthorhynchus cylindraceus (G.) Gol- -Po schmidti nom. novo (Germany): syn. of Prost- van, 1956 (fide SCHMIDT,1981) horhynehus rectus Sprehn, 1942 nee Linton, 1892 -Po gallinagi Schachtachtinskaja, 1953 (Azerbaidjan, -Po scolopacidis Kostylew, 1915 (type species) (Europe SSSR) and Central Asia) -Po genitopapillatus Lundstrorn, 1942 (Scandinavia, -Po scolopacidis sensu Lundstrorn, 1942: may not be a Central Asia) syn. of Prosthrorhynchus spiralis (R.) Golvan, 1956 -Po golvani (Schmidt et Kuntz, 1966) novo comb. (Tai- nor of Prosthorhynchus totani (R.) Golvan, 1956 (fide wan and Pescadores Islands): syn. of Plagiorhynchus GOLVAN,1960) golvani Schmidt et Kuntz, 1966 -Po spiralis (Rudolphi, 1809) Meyer, 1932: syn. of Pla- -Po gracilis Petrotschenko, 1958 (Armenia, SSSR) giorhynchus spiralis (R.) Golvan, 1956 or species -Po horridus (von Linstow, 1897) Travassos, 1926: syn. incertae sedis (fide SCHMIDT& KUNTZ, 1966) of Centrorhynchus horridus (vL.) Meyer, 1932 -Po taiwanensis (Schmidt et Kuntz, 1966) nov. comb. -Po limnobaeni Tubangui, 1933 (description based (Taiwan): syn. of Plagiorhynchus taiwanensis Schmidt solely on a male specimen): syn. of Plagiorhynchus et Kuntz, 1966 limnobaeni (T.) Golvan, 1956 -Po totani (Porta, 1910) Meyer, 1932: syn. of Pla- -Po linearis (Westrumb, 1821) Meyer, 1932: syn. of Pla- giorhynchus totani (P.) Golvan, 1956 giorhynchus linearis (W.) Golvan, 1956 -Po transversus (Rudolphi, 1819) Travassos, 1926 (Eu- -Po longirostris Travassos, 1926 (Neotropical region) rasia) -P.luehei Travassos, 1916 (Brazil) -Po upupae L6pez-Neyra, 1946: syn. of Prosthorhyn- -Po malayensis Tubangui, 1935 (nee malayense) ch us cylindraceus (G.) Golvan, 1956 (Luzon, Philippine Islands; Taiwan) -Po urichi Cameron, 1936: species incertae sedis (fide -Po menurae (Johnston, 1912) Meyer, 1932: syn. of Pla- GOLVA ,1956 and SCHMIDT& KUNTZ, 1966) becau- giorhynchus menurae (J.) Golvan, 1956 se the description of this species is too poor; it may -Po nicobarensis Soota et Kansal, 1972 (Nicobar be a Plagiorhynchus spiralis (C.) Golvan, 1956 ins- Islands) tead -Po odhneri (Lundstrom, 1942): syn. of Plagiorhynchus -Po varispinus Wang Pu Qin, 1966 (Fukien, China) odhneri Lundstrom, 1942 -Po ogatai (Fukui et Morisita, 1936) novo comb. (Japan): PROTORHADINORHYNCHUS Petrotschenko, 1956: syn. of Porrorchis ogatai Fukui et Morisita, 1936 and syn. of Rhadinorhynchus Luhe, 1911 of Plagiorhynchus (Prosthorhynehus) ogatai (Fukui et -Po ea rang is (Yamaguti, 1939) Petrotschenko, 1956: Morisita, 1936) Schmidt et Kuntz, 1966 syn. of Rhadinorhynchus carangis Yamaguti, 1939 -Po paulus (Van Cleave et Williams, 1951) Petrotschen- (nee earrangis) ko, 1958: syn. of Plagiorhynchus paulus Van Cleave -Po ditrematis (Yamaguti, 1939) Petrotschenko, 1956 et Williams, 1951 (nee H.L. Ward, 1951): syn. of Rhadinorhynchus -Po pigmentatus (de Marval, 1902) Meyer, 1932 (Europe) ditrematis Yamaguti, 1939 176 Y.J. GOLVAN PSEUDAUCHEN Yamaguti, 1963 of Echinorhynchus (Pseudoechinorhynchus) parasi- -P. epinepheli (Yamaguti, 1939) Yamaguti, 1963 luri Fukui, 1929 (type species) (Japanese Inland Sea) (nee epine- PSEUDOGORDIORHYNCHUS Golvan, 1957 phali) -P. antonmeyeri Golvan, 1957 (type species) (West of Africa) PSEUDOACANTHOCEPHALUS Petrotschenko, 1956 -P. betsileo Golvan, Houin et Brygoo, 1969 (Madagas- PSEUDOLEPTORHYNCHOIDES Salgado-Maldona- car) do, 1976 -P. bigueti (Houin, Golvan et Brygoo, 1965) Golvan, -P. lamothei Salgado-Maldonado, 1976 (type species) 1969 (Madagascar) (Sinaloa, Mexico) -P. bufonicola (Kostylew, 1941) Petrotschenko, 1958 (Tadjikistan, Central Asia, Eastern Europe) PSEUDOLUEHEIA Schmidt et Kuntz, 1967 -P. bufonis (Shipley, 1903) Petrotschenko, 1958 (type -P. boreotis (Van Cleave et Williams, 1951) Schmidt et species) (Hong Kong, Thailand) Kuntz, 1967 (Alaska, Nearctic region) -P. caucasicus (Petrotschenko, 1953) Petrotschenko, -P. korathai Gupta et Fatma, 1988 (Uttar Pradesh, 1958 (Central and Eastern Europe) Tamil Nadu, India) -P. elongatus (Van Cleave, 1937) Petrotschenko, 1958 -P. pittae Schmidt et Kuntz, 1967 (type species) (Pala- (Hunan, China) wan, Philippine Islands) -P. perthensis Edmonds, 1971 (Perth, Australia) -P. rauschi Gupta et Fatma, 1986 (Lucknow, India) PSEUDOPORRORCHIS Joyeux et Baer, 1935: syn. of -P. xenopeltidis (Shipley, 1903) Golvan, 1969 (Thai- Porrorchis Fukui, 1929 land) -Po bazae (Southwell et Macfie, 1925) Petrotschenko, 1958: syn. of Porrorchis bazae (S. et M.) Schmidt et Kuntz, 1967 PSEUDOCAVISOMA Go1van et Houin, 1964 -Po bulboeaudatus (Southwell et Macfie, 1925) Joyeux -P. chromitidis (Cable et Quick, 1954) Golvan et Houin, et Baer, 1935: syn. of Porrorchis hylae (1.) Schmidt et 1964 (type species) [Puerto Rico, also found in India Kuntz, 1967 (doubtful)] -Po centropi (Porta, 1910) Joyeux et Baer, 1935: syn. of Porrorchis centropi (P.) Schmidt et Kuntz, 1967 PSEUDOECHINORHYNCHUS Petrotschenko, 1956: -Po centropusi (Tubangui, 1933) Joyeux et Baer, 1935: the validity of this genus is contested by AMIN &RED- syn. of Porrorchis centropusi (T.) Schmidt et Kuntz, UN (1980) and by HUFFMAN(1973) who considered it 1967 a perfect synonym of Echinorhynchus sensu stricto; in -Po elongatus (Fukui, 1929) Joyeux et Baer, 1935: syn. my opinion it may be used as a sub genus of Echi- of Porrorchis elongatus Fukui, 1929 norhynchus -Po houdemeri Joyeux et Baer, 1935: syn. of Porrorchis -Po baeri (Kostylew, 1928) Golvan, 1969: syn. of Echi- houdemeri (1. et B.) Schmidt et Kuntz, 1967 norhynchus (Pseudoechinorhynchus) baeri Kostylew, -Po hydromuris Edmonds, 1957: syn. of Porrorchis 1928 hydromuris (E.) Schmidt et Kuntz, 1967 -Po cinctulus (Porta, 1905) Petrotschenko, 1956: syn. of -Po hylae (Johnston, 1912) Edmonds, 1957: syn. of Echinorhynchus (Pseudoechinorhynchus) cinctulus Porrorchis hylae (J.) Schmidt et Kuntz, 1967 ) Porta, 1905 -Po indicus Das, 1957: syn. of Porrorchis indicus (D.) -P. clavula (Dujardin, 1845) Petrotschenko, 1956: syn. Schmidt et Kuntz, 1967 of Acanthocephalus clavula (D.) Grabda-Kazubska et -Po maxv achoni Golvan et Brygoo, 1965: syn. of Chubb, 1968 Porrorchis maxvachoni (G. et B.) Schmidt et Kuntz, -Po clavula sensu Petrotschenko, 1956 is probably a syn. 1967 pro parte of Echinorhynchus (P.) cinctulus Porta, .cp rotundatus (von Linstow, 1897) Joyeux et Baer, 1905 1935: syn. of Porrorchis rotundatus (vL.) Schmidt et -Po dirus (Van Cleave, 1931) Petrotschenko, 1956: syn. Kuntz, 1967 of Acanthocephalus dirus (VC.) Van Cleave et Town- -Po teliger Van Cleave, 1949: syn. of Owilfordia teliger send, 1936 (VC.) Schmidt et Kuntz, 1967 -Po lenok (Achmerov et Dombrowskaja-Achmerova, 1941) Petrotschenko, 1956 (nee lenoki): syn. of Echi- PSEUDORHADINORHYNCHUS Achmerov et Dom- norhynchus (Pseudoechinorhynchus) lenok Achme- browskaja-Achmerova, 1941 rov et Dombrowskaja-Achmerova, 1941 -P. cinereus Gupta et Naqvi,(1981) 1983 (Karachi, -Po monticellii (Porta, 1904) Petrotschenko, 1956 (nee Pakistan) montieelli): syn. of Echinorhynchus (Pseudoechi- -P. cochinensis Gupta et Naqvi,(1981) 1983 (Cochin, norhynchus) monticellii Porta, 1904 (in fact this spe- Kerala, South of India) cies cannot be identified) -P. dussamicitatum Gupta et Gupta, 1971 (India) -Po parasiluri (Fukui, 1929) Petrotschenko, 1956: syn. -P. ernakulensis Gupta et Gupta, 1971 (India) Nomenclature of the Acanthocephala 177 -Po leueiseus (Krotov et Petrotschenko, 1956) Golvan, RHADINORHYNCHOIDES Fukui et Morisita, 1937 1969 (Shakaline, Japan) -R. ehromitidis (Cable et Quick, 1954) Yamaguti, 1963: -Po markewitsehi Achmerov et Dombrowskaja-Achme- syn. of Pseudoeavisoma ehromitidis (c. et Q.) Golvan rova, 1941 (type species) (East of Siberia, Japan) (nee et Houin, 1964 markewitehi) -R. ehromitidis sensu Gupta et Lata, 1967 is another spe- -Pomujibi Gupta et Naqvi,(1981) 1983 (Karachi, Pakistan) cies collected in India -Po neobythitis (Yamaguti, 1939) Petrotschenko, 1956: -R. miyagawai Fukui et Morisita, 1937 (type species) syn. of Yamagutisentis neobythitis (Y.) Golvan, (Japan) 1969 -Po orissai Gupta et Fatma, (1983) 1985 (Puri, Orissa, RHADINORHYNCHUS Liihe, 1911 India) -R. afrieanus (Golvan, Houin et Deltour, 1963) Golvan, -Popseudaspii Achmerov et Dombrowskaja-Achmero- 1969 (Gabon) va, 1941 (Basin of the Amur River) -R. alosae (Hermann, 1782) Meyer, 1932 (Basin of the -Po rhinoplagusiae (Yamaguti, 1935) Petrotschenko, Rhine River): is not a syn. of Rhadinorhynehus pristis 1956: syn. of Yamagutisentis rhinoplagusiae (Y.) Gol- -R. arii Wang, 1966: syn. of Metarhadinorhynehus van, 1969 eyprini (W.) Wang, 1986 -Po samegaiensis Nakajima et Egusa, 1975 (nee same- -R. aspinosus Fukui et Morisita, 1937: syn. of gainensis) (Japan) Neorhadinorhynehus aspinosus (F. et M.) Yamagu- -Po spinieaudatum (Cable et Quick, 1954) Yamaguti, ti,1939 1963: syn. of Neoacanthoeephaloides spinieaudatus -R. asturi Gupta et Lata, 1967: syn. of Prosthorhynehus Cable et Quick, 1954) asturi (G. et L.) novo comb. -Po srivastavai Gupta et Fatma, (1983) 1985 (Puri, Oris- -R. atheri (Farooqi, 1981)Amin, 1985 (Lagos, Nigeria) sa, India) -R. bieireumspinis Hooper, 1983 (New South Wales, Australia) QUADRlGYRUS Van Cleave, 1920 -R. eadenati (Golvan et Houin, 1964) Golvan, 1969 -Q. brasiliensis Machado Filho, 1941 (Mato Grosso, (Senegal, Nigeria) Brazil) -R. eamerounensis Golvan, 1969 (Camerun) -Q. ehinensis Mao, 1979 (North China) -R. eapensis Bray, 1974 (South Africa) -Q. eholodkowskyi Kostylew, 1928 (Armenia): may be a -R. earangis Yamaguti, 1939 (Japanese Inland Sea) syn. of Aeanthoeephalorhynehoides eholodkowskyi -R. eelebensis Yamaguti, 1954: syn. of Gorgorhynehus (K.) Williams, Gibbons et Sadighian, 1980 (jide SCH- celebensis (Y.) Golvan, 1969 (nee eelebesense) MlDT & HUGGINS,1973) -R. ehongmingnensis Huang, Zheng, Deng, Fan et Ni, -Q. maehadoi Fabio, 1983 (Rio de Janeiro, Brazil) 1988 (Chongming, China) -Q. niekoli Schmidt et Hugghins, 1973 (Central Colom- -R. ehromitidis (Cable et Quick, 1954) Yamaguti, 1963 bia) (vide VIZCAINO,1988) -R. eololabis Laurs et McCauley, 1964 (Oregon, USA) -Q. polyspinosus Li Minmin, 1984 (China) -R. eyprini (???): syn. of Metarhadinorhynehus eyprini -Q. rhodei Wang Pu Qin, 1980 (Fujian, China) (??) Wang, 1986 -Q. simhai Gupta et Fatma, 1986 (India, Uttar Pradesh) -R. deeapteri Parukhin et Kovalenko, 1976 (Hawai) -Q. torquatus Van Cleave, 1920 (type species) (Colom- -R. ditrematis Yamaguti, 1939 (Japanese Inland Sea) bia, Venezuela, Mato Grosso, Brazil) -R. dollfusi Gupta et Fatma, 1987 (Kerala, India) -Q. torquatus sensu Ortlepp, 1924 may be a different -R. dujardini Golvan, 1969 (Curacao, Brazil) species from Surinam (fide MEYER, 1932) -R. epinepheli Yamaguti, 1939 (nee epinephaliy: syn. of Gorgorhynchus epinepheli (Y.) Golvan, 1969 RAORHYNCHUS Tripathi, 1959 -R. erumei (Gupta et Fatma, 1981) Amin, 1985 (India) -R. guptai Gupta et Kumar, 1987 (Orissa, Bay of Ben- (nee erumeii, nee Fatima) gal) -R. exilis Van Cleave, 1928: syn. of Cathayacanthus -R. inexspeetatus Golvan, 1969 (Antilles, Atlantic exilis (VC.) Golvan, 1969 (nee exile) Ocean) (nee inexpeetatus) -R. ganapatii Chandra, Rao et Shyamasundari, 1985 (India) -R. meyeri (Heinze, 1934) Golvan, 1969 (Indian Ocean) -R. horridus Liihe, 1912: syn. of Megistaeantha horri- -R. polynemi Tripathi, 1959 (Puri, India) dum (L.) Golvan, 1969 (nee horridum) -R. sehmidti George et Nadakal, 1978 (Kerala, India) -R. indicus Tripathi, 1959: syn. of Indorhynehus indi- -R. terebra (Rudolphi, 1819) Tripathi, 1959 (type spe- eus (T.) Golvan, 1969 cies) (Hawai, India) -R. japonieus Fukui, 1920 (Japan) (nee japonieum): -R. thapari Gupta et Fatma, 1981 (India) may be a syn. of Rhadinorhynchus seriolae (Y.) Gol- van, 1969 (jide ICHIHARAet al., 1968) RAOSENTIS Datta, 1946 -R. johni Baylis, 1929: syn. of Aspersentis johni (B.) -R. dattai Gupta et Fatma, 1986 (India, Uttar Pradesh) Chandler, 1934 -R. podderi Datta, 1946 (type species) (Calcutta, India) -R.johnstoni Golvan, 1969 (South Australia) -R. thapari Rai, 1967 (India) -R. katsuwonis Harada, 1928 (Hatijosima, Japan): may 178 Y.J. GOLVAN be a syn. of Rhadinorhynehus ornatus (fide V AN -R. wheeleri Baylis, 1929: syn. of Aspersentis me- CLEAYE,1940) garhynehus (vL. ) Golvan, 1960 -R. keralensis Gupta et Fatma, 1987 (Kerala, India) -R. zhukovi Golvan, 1969 (Kourile Islands) -R. /intoni Cable et Linderoth, 1963 (Atlantic Ocean, Mediterranean) SACCOSENTIS Tadros, 1966: may be a syn. of Palli- -R. medius (Linton, 1908) Van Cleave, 1918: syn. of sentis (fide CHOWANGUPT, A& KHERA, 1987) Gorgorhynehus medius (L.)Chandler, 1934 -So ophioeepha/i Jain et Gupta, 1979 (India) -R. meyeri Heinze, 1934: syn. of Raorhynehus meyeri -So pesteri Tadros, 1966 (type species) (Far East ?) (H.) Golvan, 1969 -R. nilotieus Meyer, 1932: syn. of Tenuisentis ni/otieus SACHALINORHYNCHUS Krotov et Petrotschenko, (M.) Van Cleave, 1936 1956 -R. nudus Harada, 1938: syn. of Neorhadinorhynehus -So skrjabini Krotov et Petrotschenko, 1956 (type spe- nudus (H.) Yamaguti, 1939 cies) (Shakaline Island) -R. ornatus Van Cleave, 1918 [Atlantic coast of the USA, also found in Japan (doubtful)] SCLEROCOLLUM Schmidt et Paperna, 1978 -R. peltorhamphi Baylis, 1944: syn. of Heteraeantho- -So robustum (Edmonds, 1964) Schmidt et Paperna, eephalus peitorhamphi (B.) Petrotschenko, 1956 1978 (Heron Island, Queensland, Australia, New Cale- -R. plagioscionis Thatcher,1980 (Amazonia) donia) -R. plotosi Parukhin, 1985 (Oman Gulf) -So rubrimaris Schmidt et Paperna, 1978 (type species) -R. polynemi Gupta et Lata, 1967 (India) (Israel, Red Sea) (nee rubimaris) -R. pristis (Rudolphi, 1802) Liihe, 1911 (type species) (North of the Atlantic Ocean) SERRASENTIS Van Cleave, 1923 (nee Serraeentis) -R. pristis sensu Linton, 1891: syn. of Rhadinorhyn- -So ehauhani Datta, 1953 (India) eh us ornatus Van Cleave -S.fotedari Gupta et Fatma, (1979) 1980 (Tamil Nadu, -R. pristis sensu Liihe, 1911 pro parte: syn. of Rhadi- India) norhynehus /intoni Cable et Linderoth, 1963 -So gigantieus Bilqees, 1972 (India): may be a syn. of S. -R. pristis sensu Fukui et Morisita, 1937: syn. of Rhadi- sagittifer (fide GUPTA& JAIN, 1977) norhynehus seriolae (Yamaguti, 1963) Golvan, 1969 -So golvani Gupta et Kumar, 1987 (Orissa, Bay of Ben- -R. pristis sensu Chandler, 1943: syn. of Rhadinorhyn- gale) eh us selkirki Van Cleave, 1921 -So lamelliger (Diesing, 1854) Meyer, 1932 (may be the -R. pristis sensu Johnston et Edmonds, 1947: syn. of generotype) Rhadinorhynehusjohnstoni Golvan, 1969 -So longa Tripathi, 1959 (Puri, India): may be a syn. of -R. pristis sensu Zhukov, 1960: syn. of Rhadinorhyn- S. sagittifer (fide GUPTA& JAIN, 1977) (nee longus) eh us zhukovi Golvan, 1969 -So longiformis Bilqees, 1971 (Pakistan): may be a syn. -R. pristis sensu Cable et Linderoth, 1963: syn. of Rha- of S. sagittifer (fide GUPTA& JAIN, 1977) dinorhynehus dujardini Golvan, 1969 -So mujibi Bilqees, 1972 (Pakistan) -R. pristis sensu Solonchenko, 1968 and Kovaleva, 1970 -So nadakali George et Nadakal, 1978 (Kerala, India) (from South Africa) is another species -So sagittifer (Linton, 1889) Linton, 1932 (type species) -R. saltatrix Troncy et Vassiliades, 1973 (West Africa) (Tropical Atlantic) (vide VAN CLEAYE & LINCICOME, -R. selkirki Van Cleave, 1920 (Juan Femandez Island) 1940) (is not a syn. of Rhadinorhynehus pristis) -So sciaenus Bilqees, 1972 (India) (nee seianis) -R. seriolae (Yamaguti, 1963) Golvan, 1969 (Japan) -So seomberomori Wang, 1981 (China): may be a syn. -R. stunkardi Gupta et Fatma, 1987 (Orissa, India) (nee of S. sagittifer fide AMIN et al. (1984) stunkardii) -So sidaroszakaio Tadros, Iskander et Wassef, 1979 -R. tenuieornis (Linton, 1891) sensu Van Cleave, 1947: (Red Sea, Egypt) (sadros sakaio ???) may be a syn. of Dollfusentis longispinus (C. et L.) -So socialis (Leidy, 1851 nee 1856) Van Cleave, 1924: Golvan, 1969 syn. of Serrasentis sagittifer (Linton, 1889) (fide LI - -R. tenuieornis (Linton, 1905) sensu Van Cleave, 1947 TON, 1932) pro parte: syn. of Dollfusentis ehandleri Golvan, 1969 -R. tenuieornis sensu Chandler, 1934: syn. of Dollfusen- SERRASENTOIDES Parukhin, 1971 tis ehandleri Golvan, 1969 -So fistulariae Parukhin, 1971 (type species) (unknown -R. tenuirostris Van Cleave, 1918: errat. pro R. tenuicornis origin ??) -R. terebra (Rudolphi, 1819) Liihe, 1911: syn. of Rao- rhynehus terebra (R.) Tripathi, 1959 SIPUNCULUS Phipps, 1774: preoccupied and composi- -R. traehuri Harada, 1935 (market of Tokyo, Japan): te, syn. of Bolbosoma Porta, 1908 pro parte and of may be a syn. of Rhadinorhynehus japonieus Fujita, Corynsosoma Liihe, 1904 pro parte 1920 and of Rhadinorhynehus selkirki Van Cleave, -So lendix Phipps, 1774: syn. of Bolbosoma balaenae 1921 (fide YAMAGUTI,1939) (G.) Meyer, 1932 pro parte and of Polymorphus -R. vancleavei Golvan, 1969 (Galapagos) phippsi Kostylew, 1922 pro parte Nomenclature of the Acanthocephala 179 SKRJABINORHYNCHUS Petrotschenko, 1956: may be -s. maryasis (Datta, 1933) Golvan, 1956 (India) a syn. of Arhythmorhynchus Luhe, 1911 -S. opimus (Travassos, 1919) Golvan, 1956: syn. of -So capellae (Yamaguti, 1935) Petrotschenko, 1958: syn. Centrorhynchus opimus Travassos, 1919 of Arhythmorhynchus capellae (Y.) Van Cleave et -So orientalis (?): may be a syn. of Gorgorhynchoides Rausch, 1950 orientalis (fide WANG, 1986) -So eroliae (Yamaguti, 1939) Petrotschenko, 1958: syn. -So physocoracis (Porta, 1913) Golvan, 1956 (New of Arhythmorhynchus eroliae (Y.) Van Cleave et Caledonia) Rausch, 1950 (nec erolie) -S. picae (Rudolphi, 1819) Golvan, 1956 (type species) (Europe) SOLEARHYNCHUS de Buron et Maillard, 1985 -S.pinguis (Van Cleave, 1918) Golvan, 1956 (Nankin, -S. soleae (Porta, 1905) de Buron et Maillard, 1985 China; Taiwan) (type species) (West of the Mediterranean Sea) -So reptans (Bhalerao, 1931) Golvan, 1956 (India) -So robustus (Datta, 1928) novo comb. (India): syn. of SOUTHWELLINA Witenberg, 1932 Echinorhynchus robustus Datta, 1928 -So ardeae (Belopolskaja, 1958) novo comb. (SSSR): -S. saxicoloides (Nama et Rathore, 1984) novo comb. syn. of Polymorphus ardeae Belopolskaja, 1958 (Rajasthan): syn. of Centrorhynchus saxicoloides -So dimorpha Schmidt, 1973 (USA, Louisiana) Nama et Rathore, 1984 -S. duocincta Chandler, 1935 (Texas, USA) -So scanensis (Lundstrom, 1941) Golvan, 1956 (Scandi- -So hispida (Van Cleave, 1925) Witenberg, 1932 (type navia, Alaska) species)[Japan, Taiwan, also found in Florida (doubt- -So serpenticola (van Linstow, 1908) Golvan, 1956 ful)] (Java) -So macracantha (H.L. Ward et Winter, 1952) Schmidt, -So skrjabini (Petrotschenko, 1949) Golvan, 1956 (Cen- 1973 (USA) tral Asia) -So pontica (Petrotschenko et Smogorjevskaja, 1962) -So splendi (Gupta et Gupta, 1970) novo comb. (Chandi- novo comb.: vide Hemiechinosoma ponticum Petrots- garh, India): syn. of Centrorhynchus splendi Gupta et chenko et Smogorjevskaja, 1962 Gupta,1971 -So tenuicaudatus (Marotel, 1889) Amin, 1985 (Euro- SPHAERECHINORHYNCHUS Johnston et Deland, pe) 1929 -So teres (Westrumb, 1821) Golvan, 1956 (Europe, -So rotundocapitatus (Johnston, 1912) Johnston et Israel, Egypt) Deland, 1929 (type species) (New South Wales, Que- -So turdi (Yamaguti, 1939) Golvan, 1956 (Japan, Tai- ensland, Australia) wan) -S. serpenticola Schmidt et Kuntz, 1966 (North of Boneo) -So wertheimae Schmidt, 1975 (Israel)

SPHAERIROSTRIS Golvan, 1956 SPIRACANTHORHYNCHUS Harada, 1938: syn. of -So areolatus (Rudolphi, 1819) Golvan, 1956 (Europe) Longicollum Yamaguti, 1935 -So bipartitus (Solowjew, 1912) Golvan, 1956 (Turkes- -So alemniscus (Harada, 1935) Harada, 1938: syn. of tan, Kazakstan) Longicollum alemniscus (H.) Fukui et Morisita, 1937 -s. cinctus (Rudolphi, 1819) Go1van, 1956 (Italy) -S. corvi (Fukui, 1929) Golvan, 1956 (Siga, Japan) SPIRORHYNCHODES Strand, 1942: syn. of Longico- -So dollfusi nom. novo (Morocco): syn. of Centrorhyn- llum Yamaguti, 1935 chus picae sensu Dollfus, 1953 -So alemniscus (Harada, 1935) Strand, 1942: syn. of -So embae (Kolodkowsky et Kostylew, 1916) Golvan, Longicollum alemniscus (H.)Fukui et Morisita, 1937 1956: may be a syn. of Sphaerirostris lancea (West- rumb, 1821) Golvan, 1956 SPIRORHYNCHUS Harada, 1935: preoccupied, syn. of -So erraticus (Chandler, 1925) Golvan, 1956 (India) Longicollum Yamaguti, 1935 -So globuli (Nama et Rathore, 1984) novo comb. -So alemniscus Harada, 1935: syn. of Longicollum (Rajasthan): syn. of Centrorhynchus globuli Nama et alemniscus (H.)Fukui et Morisita, 1937 Rathore, 1984 -So lancea (Westrumb, 1821) Golvan, 1956 (Europe, SUBCORYNOSOMA Khokhlova, 1967: subgenus of also found in Taiwan) Polymorphus Liihe, 1911 -So lanceoides (Petrotschenko, 1949) Golvan, 1956 (Central Asia) SUBFILICOLLIS Khokhlova, 1967: subgenus of Poly- -s. leguminosus (Solowjew, 1912) Golvan, 1956 (Cen- morphus Liihe, 1911 tral Asia) -So lesiniformis (Molin, 1859) novo comb. (Saratov, Taenia haeruca Pallas, 1766: syn. pro parte of Macra- valley of the Volga River): syn. of Centrorhynchus canthorhynchus hirudinaceus (P.) Travassos, 1917 lesiniformis (Molin, 1859) Meyer, 1932 (nec Back, Taenia hirudinacea Pallas, 1781: syn. of Macracan- 1935) thorhynchus hirudinaceus (P.) Travassos, 1917 180 Y.J. GOLVAN TANAORHAMPHUS H.B. Ward, 1918 -T. carinii (Travassos, 1916) Meyer, 1932: may be a -T. ambiguus Van Cleave, 1921: syn. of Paratenuisen- syn. of Oligacanthorhynchus carinii (T.) Schmidt, tis ambiguus (VC.) Bullock et Samuel, 1975 1972 or of Hamanniella carinii Travassos, 1916 -T. longirostris (Van Cleave, 1913) H.B. Ward, 1918 -T. tumida Van Cleave, 1947: syn. of Oligacanthorhyn- (type species) (Nearctic region) chus tumidus (VC.) Schrnidt, 1972 or of Hamanniella tumida (VC.) Van Cleave, 1953 TCHADORHYNCHUS Troncy, 1970 -T. quentini Troncy, 1970 (type species) (Chad, Lebanon) TRA VASSOSINA Witenberg, 1932: syn. of Centrorhyn- chus Luhe, 1911 pro parte and of Sphaerirostris Gol- TEGORHYNCHUS Van Cleave, 1921: the synonymy van, 1956 pro parte of Tegorhynchus and Illiosentis proposed by -T. eorvi (Fukui, 1929) Witenberg, 1932: syn. of Sphae- BULLOCK& MATEO(1970) and BUCKNER,OVERSTRE- rirostris corvi (F.) Golvan, 1956 ET &HEARD(1978) is not accepted here (vide LEOTTA, -T. pinguis (Fukui, 1929) Witenberg, 1932 SCHMIDT& KUNTZ, 1982) -T. afrieanus (Golvan, 1956) Amin, 1985: syn. of Illio- WOLFFHUGELIA Mane-Garzon et Dei-Cas, 1974 sentis africanus Golvan, 1956 -Wo matercula Mane-Garzon et Dei-Cas, 1974 (type -T. brevis Van Cleave, 1921 (type species) (Juan Fer- species) (Uruguay) nandez Islands) (nee breve) -T. eetratus (Van Cleave, 1945) Bullock et Mateo, 1970: YAMAGUTISENTIS Golvan, 1969: the synonymy of syn. of Illiosentis cetratus Van Cleave, 1945 this genus with Acanthocephaloides Meyer, 1932, pro- -T. edmondsi (Golvan,1960) Amin, 1985: syn. of Illio- posed by ARAKI& MACHIDA(1987), cannot be accepted sentis edmondsi Golvan, 1960 -Y. neobythitis (Yamaguti, 1939) Golvan, 1969 (type -T.fureatus (Van Cleave et Lincicome, 1939) Bullock et species) (Hukui, Japan) Mateo, 1970: syn. of Illiosentisfurcatus Van Cleave et - Y. rhinoplagusiae (Yamaguti, 1935) Gol van, 1969 Lincicome, 1939 (Japanese Inland Sea) - T. pectinarius Van Cleave, 1940 (Puerto Rico) (nee peetinarium) ZEYLANECHINORHYNCHUS Fernando et Furtado, TELOSENTIS Van Cleave, 1923 1963 -T. australiensis Edmonds, 1964 (Australia) -Z. longinuchalis Fernando et Furtado, 1963 (type spe- -T. exiguus (von Linstow, 1901) Van Cleave, 1923 cies) (Sri Lanka) (Black Sea) -T. mizellei Gupta et Fatma, 1988 (Uttar Pradesh, India) ZEYLONECHINORHYNCHUS: errat. pro Zeylanechi- -T. molini Van Cleave, 1923 (type species) (Sicily, norhynchus and Mediterranean Sea) -T. tenuieornis (Linton, 1892) Van Cleave, 1947: syn. of Dollfusentis longispinus (C. et L.) Golvan, 1969 REFERENCES -T. tenuieornis sensu Linton, 1905 in Van Cleave, 1947 ACHMEROV (A.K.), 1959.- [Acanthocephales des poissons du fleu- pro parte: syn. of Dollfusentis chandleri Golvan, 1969 ye Amour]. Trudy Gelminthologicheskoi Laboratorii, Akademii Nauk S.S.S.R., 9: 23-44. (in Russian) TENUIPROBOSCIS Yamaguti, 1935 ACHMEROV (A.K.) & DOMBROWSKAJA-AcHMEROVA (O.S.D), -T. guptai Gupta et Sinha (1989) 1990 (Calcutta, West 1941.- [Nouveaux Acanthocephales des poissons du fleuve Bengal, India) Amour]. Dokladi Akademii Nauk S.S.S.R., 31: 517- 520.(in -T. misgurni Yamaguti, 1935 (type species) (Japan) Russian) -T. sergenti Choquette et Gayot, 1952: syn. of Paralongi- ACHOLONU (A.D.), 1969.- Acanthocephala of Louisiana turtles collum sergenti (C. et G.) Amin, Bauer et Sidorov, 1991 with a redescription of Neoechinorhynchus stunkardi Cable et Fisher, 1961. Proceedings of the Helminthological Society of TENUISENTIS Van Cleave, 1936 Washington, 36: 177-183. AFANAS'EV (V.P.), 1941.- [Faune parasitaire des marnmiferes -T. niloticus (Meyer, 1932) Van Cleave, 1936 (type spe- dinteret commercial des iles du Commandeur]. Uchenye Zapis- cies) (White Nile, Africa) ki Leningradskogo Gosudarstvennogo Universiteta, Seriya Bio- logicheskikh Nauk, 18 (74): 93-117.(in Russian) TERMITORHYNCHUS Reichensperger, 1922: subgenus of AGARWAL (S.C.), 1958. - A new species of the genus Pallisentis Acanthocephalus (???) or of Oligacanthorhynchus (???) (Acanthocephala). Current Science, Bangalore, 27 (3): 107. ALEGRET (MJ.), 1941.- Contribucion al conocimiento de los ver- TRA VASSOSIA Meyer, 1931: nomen nudum mes parasites de Phalacrocorax auritus floridanus. Tesis en Ciencias Naturales, Facultad de Ciencias, Universidad de la Habana, La Habana, 32 pp. TRA VASSOSIA Meyer, 1932: syn. of Hamanniella Tra- ALY KHAN & BILQEES (F.M.), 1985.- Dispiron heteroacanthus, vassos, 1915; this genus is strictly restricted to America new species (Acanthocephala: ) from the (vide VANCLEAVE,1953); may be a syn. of Oligacan- fish Labeo rohita (Ham.) of Kalri Lake, Sind, Pakistan. Pakistan thorhynchus (fide SCHMIDT,1972) Journal of Zoology, 17: 11-l3. Nomenclature of the Acanthocephala 181

ALY KHAN & BILQEES(F.M.), 1985.- Pallisentis kalriai, new spe- ANANTARAMAN (S.) & ANA TARAMAN (M.), 1969.- On Cen- cies (Acanthocephala: ) from the fish Labeo rohi- trorhynchus ( Centrorhynchus} golvani, a new species of an ta of Kalri Lake, Sind, Pakistan. Philippine Journal of Science, Acanthocephalan from the booted eagle, Hieraetus pennatus in 114(1/2): 101-111. Madras. Annales de Parasitologie Humaine et Comparee, 44: ALY KHAN & BILQEES (F.M.), 1987.-Two new Acanthocephala 5-12. species from freshwater fishes of Kalri Lake. Pakistan Journal A TO 10 (H.), 1958.- Descripci6n de una nueva especie del gene- of Zoology, 19: 263-271. ro Gigantorhynchus Hamann, 1892 (Acanthocephala). Acta Bio- ALY KHA & BILQEES (F.M.), 1989.- On a new acanthocephala, logica veneruelica, 2 (24): 291-298. Neoechinorhynchus gibsoni, new species from a freshwater fish, ARAKI (J.) & MACHIDA(M.), 1987.- Some acanthocephalans from Labeo rohita (Ham.). 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(in (Acanthocephala: Neoechinorhynchidae) from two species of Russian) suckers (Catostornidae) in Alabama, with comments on Eocollis BHALERAO (G.D.), 1931.- On a new species of Acanthocephala arcanus. Proceedings of the Helminthological Society of Was- from Ophiocephalus striatus. Annals and Magazine of Natural hington, 59: 159-162. History, Ser. 10, 7: 569-573. BUCKNER(R.L.) & BUCKNER (S.c.), 1976.- A new species of Lep- BHALERAO(G.D.), 1937- On a remarkable Acanthocephala from a torhynchoides Kostylew, 1924 (Acanthocephala) from the pirate fowl in India. Proceedings of the Zoological Society of , perch, Aphredoderus sayanus (Gilliams). Journal of Parasito- 107, ser. B (2): 199-203. logy, 62: 955-958. BILQEES (F.M.), 1970.- Parasites of grey mullets (Mugil buchana- BUCKNER (R.L.) & NICKOL (B.B.), 1978.- Redescription of Fessi- ni) with description of Dispiron mugili n. gen. and sp., a fifth sentis vancleavei (Hughes and Moore, 1943) Nickol, 1972 genus of Neoechinorhynchinae (Acanthocephala). Agriculture (Acanthocephala: Fessisentidae). Journal of Parasitology, 64: Pakistan, 21 (1): 103-109. 635-637. BILQEES (F.M.), 1971.- Parasites of the cat fish Arius serratus. I. BUCKNER (R.L.), OVERSTREET (R.M.) & HEARD(R.W.), 1978.- Acanthocephala. Pakistan Journal of Zoology, 3: 57-65. Intermediate hosts for Tegorhynchus furcatus and Dollfusentis BILQEES (F.M.), 1972.- Descriptions of two Acanthocephala, chandleri. Proceedings of the Helminthological Society of Was- including a new species Neoechinorhynchus karachiensis (Neo- hington,45: 195-20 I. echinorhynchidea: Neoechinorhynchidae), from marine fishes BULLOCK (W.L.), 1957.- Octospiniferoides chandleri n. gen., n. of Karachi. Sind University Research Journal, Science ser., 6: sp., a neoechinorhynchid acanthocephalan from Fundulus gran- 93-100. dis Baird and Girard on the Texas coast. Journal of Parasito- BILQEES (F.M.), 1972.- Serrasentis giganticus n. sp.(Acanthocep- logy, 43: 97-100. hala, Rhadinorhynchidae : Serrasentinae), from fishes of Kara- BULLOCK (W.L.), 1960.- Some acanthocephalan parasites of Flori- chi coast. In: Helminth parasites of some chiefly da fishes. Bulletin of Marine Science of the Gulf and Caribbean, from fishes of West Pakistan (F.M. Bilquees, R. Saeed, R. Reha- 10 (4): 481-484. na, A. Khatoon & S.H. Kaikabad edit.), Agricultural Research BULLOCK(W.L.), 1962.- A new species of Acanthocephalus from Council, Government of Pakistan, Karachi, Pakistan: 60-65. ew England fishes with observations on variability. Journal of BILQEES (F.M.) & MY KHAN, 1983.- Paraechinorhynchus kalriai Parasitology, 48: 442-445. n. g., n. sp.(Neoechinorhynchidea: Neoechinorhynchinae) from BULLOCK(W.L.), 1962.- The status of the acanthocephalan genus Labeao rohita (Ham.). Pakistan Journal of Agricultural Floridosentis Ward, 1953 and Atactorhynchus Chandler, 1935. Science, 20 (3/4): 116-120. Proceedings of the Helminthological Society of Washington, 29: BISSERU (B.), 1956.- On a new Acanthocephalan, Echinopardalis 217-218. lerouxi n. sp., from a jackal (Canis adustus) in Central Africa. BULLOCK (W.L.), 1963.- Neoechinorhynchus prolixoides n. sp. Journal of Helminthology, 30: 41-50. (Acanthocephala) from North American fishes. Proceedings of BISSERU(B.), 1960.- The parasites of Heterocorax capensis from the Helminthological Society of Washington, 30: 92-96. the Eastern Province. Fort Hare Papers, 2 (2): 1-86. BULLOCK (W.L.), 1966.- A redescription of Octospiniferoides BLANC(G.R.) & CAUCHEMEZ(L.St.), 1911.- Sur un echinorhynque chandleri Bullock, 1957. Journal of Parasitology, 52: 735-738. nouveau (Echinorhynchus brumpti novo sp.) parasite du heris- BULLOCK (W.) & MATEO (E.), 1970.- A reconsideration of the son. Comptes Rendus des Sceances de la Societe de Biologie, acanthocephalan genera Telosentis, Illiosentis and Tegorhyn- 71: 120-121. chus on the basis of the type specimens. Proceedings of the 11 BLOCH (M.E.), 1782.- Abhandlung von der Erzeugung der Einge- International Congress of Parasitology (Washington), Journal weidewiirmer und den Miueln wider deselben. Berlin, pp. 26-28. of Parasitology, 56 (4), Sect. 2, Part I: 41-42. Bosc (L.A.G.), 1802.- Histoire naturelle des vers, contenant leurs BULLOCK (W.L.) & SAMUEL (G.), 1975.- Paratenuisentis gen. n. descriptions et leurs moeurs avec des figures dessinees d' apres for Tanaorhamphus ambiguus Van Cleave, 1921 (Acanthoce- nature. n. 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Jr.), 1957.- A fourth species of CHOQUETTE (L.P.E.) & GAYOT (G.), 1952.- Tenuiproboscis ser- Neoechinorhynchus (Acanthocephala) in turtles of the United genti nov. sp., acanthocephale trouve chez le chien a Alger. States. Journal of Parasitology, 43, Sect. 2 (5): 29. Archives de l'Institut Pasteur d'Algerie, 30: 51-54. CABLE (R.M.) & FISHER (F.M. Jr.), 1961.- A fifth species of Neoe- CHOWAN (J.S.), GUPTA (N.K.) & KHERA (S.), 1987.- On two spe- chinorhynchus (Acanthocephala) in turtles. Journal of Parasito- cies of the genus Pallisentis Van Cleave, 1928 from fishes of logy, 47: 666-668. Gobindsagar Lake and synonymy of the genus Saccosentis CABLE (R.M.) & Hors (W.B.), 1954.- Acanthocephalan parasites Tadros, 1966. Research Bulletin of the Panjab University, of the genus Neoechinorhynchus in North American turtles with Science, 38 (1/2): 21-26. the descriptions of two new species. 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Sobolev], history of a new species of the acanthocephalan genus Arhyth- Dalnevostoschnii Gosudarstevennii Universitet, Vladivostok: morhynchus from the frog Rana tigrina (Daud.) from India. 81-83.(in Russian) Records of the Indian Museum, 51 (I): 39-50. DEVEAUX (T.P.), SCHMIDT (G.D.) & KRISHNASAMY(M.), 1988.- DAS (E.N.), 1957.- On juvenile and adult forms of Pseudoporror- Two new species of Moniliformis (Acanthocephala: Monilifor- chis indicus, a new species of Acanthocephala from India. Jour- midae) from Malaysia. Journal of Parasitology, 74: 322-325. nal of Parasitology, 43: 659-663. DHAR(R.L.) & KHAROO (V.K.), 1984.- A new species of Acantho- DATIA (M.N.), 1928.- On Echinorhynchus robustus n. sp. from the cephala, Neoechinorhynchus glyptosternumi n.sp. from the common crows Corvus corax and Corvus splendens of Allaha- intestine of a Kashmir fish, Glyptosternum sp. Indian Journal of bad. The Allahabad University Studies, 4: 41-61. Helminthology, 36: 36-39. 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On a new DfAZ-U GR(A(C.) & GRACIA-RoDRIGO (A.), 1957.- Revisi6n de la genus of acanthocephalan parasite of the family Quadrigyridae, familia Quadrigyridae (Acanthocephala) con descripci6n de from a Calcutta fish, Mystus cavasius (Ham.). Records of the Deltania scorzai gen. n.; sp. n. Novedades Cientificas, Contribu- Indian Museum, 44 (4): 363-367. ciones ocasionales del Museo de Historia Natural La Salle DATIA(M.N.), 1954.- On a new species of acanthocephalan para- (Caracas), Ser. Zoologia, 22: 3-19. site from fishes of Bombay. Records of the Indian Museum, 51 DfAZ-UNGRfA (C.) & GRACIA-RoDRIGO (A.), 1958.- Un cambio de (I): 51-55. nombre por preocupaci6n. Novedades Cientificas, Contribucio- DATIA (M.N.) & SOOTA(T.D.), (1954) 1955.- Two new species of nes ocasionales del Museo de Historia Natural La Salle (Cara- acanthocephalan parasites of Birds. Records of the Indian cas), Ser. Zoologia, 23: 42. Museum, 52 (I): 15-20. DiAz-U GRfA(C.) & GRAC!A-RODRIGO (A.), 1960.- Revisi6n del DATIA (M.N.) & SOOTA (T.D.), (1954) 1956.- On a new species of genero Polyacanthorhynchus Travassos, 1920 (Acanthocephala) the genus Acanthocephalus from Rana sp. and a new host record con descripci6n de una especie neuva. In: Libro Homenaje al of Centrorhynchus cinctus (Rudolphi). Records of the Indian Dr. Eduardo Caballero y Caballero, Mexico: 429-433. Museum, 52: 185-188. DIESING (K.M.), 1851.- Systema Helminthum. 1 and 2. Vindobo- DATIA (M .. ) & SOOTA (T.D.), (1960) 1962.- Two new Acantho- nae, W. Braurnuller, 588 pp. + Corrigenda 3 pp. Nomenclature of the Acanthocephala 185

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GEORGE (P.V.) & NADAKAL (A.M.), 1982.- Histopathological FOTEDAR(D.N.) & DHAR(R.L.), 1977.- On five new species of the changes in the intestine of the fish Synaptura orientalis (BI. et genus Pomphorhynchus Monticelli, 1905 from fresh-water fi- Sch.) parasitized by an acanthocephalid worm, Echinorhynchus shes of Jammu and Kashmir, India. Indian Journal of Zoology, veli (George et Nadakal, 1978). Japanese Journal of Parasito- 5: 15-24. logy, 31: 99-103. FOTEDAR(D.N.), DUDA(P.L.) & RAINA (M.K.), 1970.- On a new GEORGE (P.V.) & NADAKAL (A.M.), 1984.- Three new species of species of Pomphorhynchus Monticelli, 1905 from a fresh-water Acanthocephala (Gigantorhynchidae) from birds of KeraIa. Acta fish in Kashmir. In: Proceedings of the Indian Science Congress Parasitologica Polonica, 29 (9/19): 97-106. Association, Abstracts, 57 (3): 455-456. GEORGE (P.V.) & NADAKAL (A.M.), 1987.- Centrorhynchus be- FOTEDAR(D.N.), DUDA (P.L.) & RAINA (M.K.), 1970.- A new spe- thaniae sp. n.(Acanthocephala:Gigantorhynchidae) from bird, cies of the genus Pomphorhynchus Monticelli, 1905 from Bufo Accipiter badius (Gmelin) of Kerala, India. Japanese Journal of viridis in Kashmir. Indian Journal of Helminthology, 22 (1): Parasitology, 36: 9-12. 41-45. GEORGE (P.V.), NADAKAL (A.M.), VIJAYAKUMARA (N.K.) & Nomenclature of the Acanthocephala 187

RAJENDRAN (M.), 1981.- A new species of Acanthocephala, GOLVAN (YJ.), 1957.- Acanthocephales d'oiseaux. Septierne note. Mediorhynchus indicus n. sp., an intestinal parasite of domestic Gordiorhynchus (Gordiorhynchus) madagascariensis n. sp. fowl, Gallus gallus domesticus in Kerala. Japanese Journal of parasite d'un rapace, Polyboroides madagascariensis. Annates Parasitology, 30: 301-304. de Parasitologie Humaine et Comparee, 32: 200-207. GIEBEL (C.G.A.) & NITZSCH (C.L.), 1857.- Helminthologische Go LVAN (Y.J.), 1958.- Acanthocephales d' Afrique Occidentale Untersuchungen. Zeitschrift fur die Gesammten Naturwissens- Francaise recoltes par le Dr. Pierre Claude Morel. Bulletin de chaften, Berlin, 9 (4): 264-269. l'lnstitut Francais d'Afrique Noire, 20, ser. A (I): 59-72. GIEBEL (C.G.A.) & NITZSCH (C.L.), 1866.- Echinorhynchus nitzs- GOLVAN (Y.J.), 1959.- Le Phylum des Acanthocephala. Deuxierne chi n. sp. Zeitschrift fiir die Gesammten Naturwissenschaften, note. La Classe des Eoacanthocephala (Van Cleave, 1936). Berlin, 28: 87-88. Annales de Parasitologie Humaine et Comparee, 34: 5-52. GMELI (J.F.), 1790.- Systema naturae per regna tria naturae GOLVAN (Y.J.), 1959.- Acanthocephales du genre Corynosoma secundum classes, ordines, genera, species, cum caracteribus Liihe, 1904, parasites de rnamrniferes d' Alaska et de Midway. differentis, synonymis locis. No. 6 (Vermes): 3021-3910. Annales de Parasitologie Humaine et Comparee, 34: 288-321. GMELIN(J.F.), 1791.- Caroli a Linne Systema Naturae. 13th Edi- GOLVAN (YJ.), 1960.- Le Phylum des Acanthocephala. Troisierne tion, I. Pars VI. Lipsiae. note. La Classe des Palaeacanthocephala (Meyer, 1931). Anna- GOEZE (J.A.E.), 1782.- Versuch einer Naturgeschichte der Einge- les de Parasitologie Humaine et Comparee, 35: 138-165. weidewiirmer thierischer Korper. Blankenburg, 471 pp. GOLVA (Y.J.), 1960.- Le Phylum des Acanthocephala. Troisierne GOLVAN (Y J .), 1956.- Parasites des poissons de mer ouest-afri- note. La Classe des Palaeacanthocephala (Meyer, 1931) (suite). cains recoltes par J. Cadenat. VIII. Acanthocephales, Bulletin de Annales de Parasitologie Humaine et Comparee, 35: 350-386. l'lnstitut Francais d'Afrique Noire, 18, ser. A (2): 467-481. GOLVAN (YJ.), 1960.- Le Phylum des Acanthocephala. Troisierne GOLVAN (YJ.), 1956.- Acanthocephales d'oiseaux. Premiere note. note. La Classe des Palaeacanthocephala (Meyer, 1931) (suite). Description d'Arhythmorhynchus longicollis (Villot, 1875) et revi- Annales de Parasitologie Humaine et Comparee, 35: 573-593. sion du genre Arhythmorhynchus Liihe, 1911 (Acanthocephala). GOLVAN (Y.J.), 1960.- Le Phylum des Acanthocephala. Troisierne Annales de Parasitologie Humaine et Comparee, 31: 199-224. note. La Classe des Palaeacanthocephala (Meyer, 1931) (suite). GOLVAN (YJ.), 1956.- Acanthocephales d' Amazonie. Redescrip- Annales de Parasitologie Humaine et Comparee, 35: 713-723. tion d ' Oligacanthorhynchus iheringi Travassos, 1916 et des- GOLVAN (Y.J.), 1961.- Le Phylum des Acanthocephala. Troisierne cription de Neoechinorhynchus buttnerae n. sp. (Neoacantho- note. La Classe des Palaeacanthocephala (Meyer, 1931) (suite). cephala, eoechinorhynchidae). Annates de Parasitologie Annales de Parasitologie Humaine et Comparee, 36: 76-91. Humaine et Comparee, 31: 500-524. GOLVA (Y.J.), 1961.- Le Phylum des Acanthocephala. Troisierne GOLVAN (Y.J.), 1956.-Une espece et une var iete nouvelles note.La Classe des Palaeacanthocephala (Meyer, 1931) (suite). dacanthocephales parasites de poissons de mer des cotes du Annales de Parasitologie Humaine et Comparee, 36: 612-647. Senegal et redescription de Serrasentis socialis (Leidy, 1851) GOLVAN (Y.J.), 1961.- Le Phylum des Acanthocephala. Troisieme Van Cleave, 1924. Annales de Parasitologie Humaine et Com- note.La Classe des Palaeacanthocephala (Meyer, 1931) (suite et paree, 31: 225-241. fin). Annales de Parasitologie Humaine et Comparee, 36: GOLVA (Y.J.), I956.-Acanthocephales d'oiseaux. Troisierne 717-736. note. Revision des especes europeennes de la sous-famille des GOLVAN (Y.J.), 1962.- Le Phylum des Acanthocephala. Quatrierne Plagiorhynchinae A. Meyer, 1931 (Polymorphidae). Annales de note. La Classe des (A. Meyer, 1931). Parasitologie Humaine et Comparee, 31: 351-384. Annales de Parasitologie Humaine et Comparee, 37: 1-82. GOLVAN (Y.J.), 1956.- Acanthocephales d'oiseaux. Cinquierne GOLVAN (YJ.), 1962.- Acanthocephales parasites de mamrniferes. note. Les especes du genre Oligacanthorhynchus Travassos, Pare National de l'Upemba, Mission G.F. de Witte, 65 (3): 1915 (Archiacanthocephala - Oligacanthorhynchidae Meyer, 65-76. 1931) dans I'Ancien Monde. Bulletin de la Societe Zoologique GOLVA (YJ.), 1965.- Acanthocephales de Madagascar recoltes de France, 81: 344-353. par E.R. Brygoo. (Premiere note). Annales de Parasitologie GOLVAN (Y.J.), 1956.- Acanthocephales d'oiseaux. Note addition- Humaine et Comparee, 40: 303-316. nelle. Pseudoporrorchis rotundatus (0. von Linstow, 1897) GOLVAN (Y J .), 1969.- Systernatique des Acanthocephales (Acan- (Palaeacanthocephala, Polymorphidae), parasite d'un Cuculidae thocephala Rudolphi, 1801). L'ordre des Palaeacanthocephala Centropus madagascariensis (Bris.). Bulletin de la Societe 200- Meyer, 1931. I. La super-famille des Echinorhynchoidea (Cob- logique de France, 81: 339-344. bold, 1876) Golvan et Houin, 1963. Memoires du Museum GOLVA (Y.J.), 1956.- Le genre Centrorhynchus Luhe, 1911 National d'Histoire Naturelie, nouvelle serie, ser. A, Zoologie, (Acanthocephala, Polymorphidae). Revision des especes euro- 57 (fascicule unique): 1-373. peennnes et description d'une nouvelle espece africaine parasite GOLVAN (Y.J.) & BRYGOO (E.R.), 1965.- Acanthocephales de de rap ace diume. Bulletin de l'lnstitut Francais d'Afrique Noire, Madagascar. Deu x ierne note. Le genre Pseudoporrorchis 18, ser, A (3): 732-785. Joyeux et Baer, 1935. Annales de Parasitologie Humaine et GOLVAN (Y.J.), 1957.- Acanthocephales des poissons. Exploration Comparee, 40: 543-568. Hydrobiologique des Lacs Kivu, Edouard et Albert (1952- GOLVA (Y.J.) & Houi (R.), 1963.- Revision des Palaeacantho- 1954). Resultats Scientifiques. Institut Royal des Sciences Natu- cephala. Premiere note. La famille des Diplosentidae Tubangui relies de Belgique, 3 (2): 55-64. et Masilungan, 1937. Annales de Parasitologie Humaine et GOLVAN (Y.J.), 1957.- Acanthocephales de I'Angola. 1. Oncicola Comparee, 38: 807-822. angolensis n. sp.(Archiacanthocephala, Pachysentidae), parasite GOLVAN (Y.J.) & HOUIN(R.), 1964.- Euretacanthus simplex novo du chacal Canis adustus Sundevall. Publicacion Servicos Cultu- gen.(Rudolphi, 1810)(Palaeacanthocephala, Echinorhynchi- rais de la Companhia de Diamantes de Angola, Museo do dae) parasite du surmulet (Mullus surmuletus Linne, 1758) 11. Dundo, 34: 39-50. Banyuls-sur-Mer. Annales de Parasitologie Humaine et Compa- GOLVAN (Y.J.), 1957.- Acanthocephales d'oiseaux. Sixierne note. ree, 39: 61-68. Deux especes nouvelles parasites doiseaux d' Afrique Occiden- GOLVAN (YJ.) & HOUIN (R.), 1964.- Revision des Palaeacantho- tale Francaise, Gordiorhynchus (Gordiorhynchus) gendrei n. sp. cephala. Deuxierne note. La famille des Gorgorhynchidae Van et Pseudogordiorhynchus antonmeyeri n. gen et n. sp. Annales Cleave et Lincicome, 1940. Annales de Parasitologie Humaine de Parasitologie Humaine et Comparee, 32: 83-97. et Comparee, 39: 535-605. 188 Y.J. GOLVAN

GOLVAN (YL) & MOKHAYER(B.), 1973.- Acanthocephales des Nadu. Indian Journal of Helminthology, 31 (I): 45-53. esturgeons de la mer Caspienne. Annales de Parasitologie GUPTA(V.) & FATMA (5.), 1981.- On three acanthocephalan para- Humaine et Comparee, 48: 597-602. sites of fishes. Rivista di Parassitologia, 42: 443-449. GOLVAN (Y.J.) & THEODORIDES (1.),1960.- Cycle evolutif d'un GUPTA (V.) & FATMA (5.), (1981) 1983.- On five new species of acanthocephale parasite de Gerbi llides du genre Meriones en the genus Centrorhynchus Liihe, 1911 (Acanthocephala: Cen- Iran. Comptes Rendus des Seances de l'Academie des Sciences trorhynchidae Van Cleave, 1916) from avian and amphibian de Paris, 250: 224-225. hosts of Lucknow,Uttar Pradesh. Indian Journal of Helmintho- GOLVAN (YJ.), BRYGOO (E.R.) & GASSMANN (M.), 1972.-Acan- logy, 33 (2): 105-120. thocephalus domerguei n. sp., parasite d'un amphibien malga- GUPTA (V.) & FATMA (5.), (1983) 1985.- On some acanthocepha- che. Annales de Parasitologie Humaine et Comparee, 47: lan parasites of fishes. Indian Journal of Helminthology, 35 (2): 305-308. 137-154. GOLVA (Y.l.), GRACIA-RODRIGO (A.) & DfAZ-UNGRfA (C.), GUPTA (V.) & FATMA (5.), (1985) 1986.- Three acanthocephalan 1964.- Megapriapus ungriai (Gracia-Rodrigo, 1960) n. gen. parasites of amphibians and mammalian hosts from Lucknow. (Palaeacanthocephala) parasite d'une pastenague d'eau douce Indian Journal of Helminthology, 37 (2): 137-148. du Venezuela (Potamotrygon hystrix). Annales de Parasitologie GUPTA (V.) & FATMA (5.), (1985) 1986.- On some acanthocepha- Humaine et Comparee, 39: 53-59. lan parasites (family Quadrigyridae Van Cleave, 1920) from fis- GOLVAN (Y.J.), HOUlN(R.) & DELTOUR (F.), 1963.- Nipporhyn- hes of Uttar Pradesh and Tamil Nadu. Indian Journal of Hel- ch us africanus n. sp.(Palaeacanthocephala), parasite d'un minthology, 37 (2): 149-180. Carangidae du Gabon. Bulletin de la Societe Zoologique de GUPTA (V.) & FATMA (5.), 1987.- On some acanthocephalan para- France, 88: 71-85. sites (family Rhadinorhynchidae Travassos, 1923) from marine Goss (O.M.), 1941.- Platyhelminth and Acanthocephala parasites fishes of Kerala, Puri and Tamil Nadu. Indian Journal of Hel- of local shags. Journal of the Royal Society of Western Austra- minthology, 39 (1): 1-26. lia, 26: 1-13. GUPTA(V.) & FATMA (5.), (1987) 1988.- On four acanthocephalan GRABDA (1.) & GRABDA-KAZUBSKA (B.), 1967.- Redescription of parasites of vertebrates from Uttar Pradesh and Tamil Nadu. Paracanthocephalus graciliacanthus (Meyer, 1932) comb. nova Indian Journal of Helminthology, 39 (2): 128-142. (Acanthocephala). Acta Parasitologica Polonica, 15 (4): 35-41. GUPTA (B.V.) & SI HA(G.), (1989) 1990.- On a new acanthoce- GRABDA-KAZUBSKA(B.), 1962.- On the validity of the species phala Tenuip roboscis guptai sp. novo (Pomphorhynchidae Acanthocephalus falcatus (Frolich, 1789). Acta Parasitologica Yamaguti, 1939) from a marine fish Cerres setifer Ham. from Polonica, 10 (21/27): 377-394. Calcutta, West Bengal. Indian Journal of Helminthology, 41 (2): GRABDA-KAzuBSKA(B.) & CHUBB (l.C.), 1968.-Acanthocephalus 104-107. - the correct generic name for Echinorhynchus clavula Dujardin, GUPTA(B.V.) & SINHA(G.), (1991) 1992.- On some acanthoce- 1845 (Acanthocephala). Acta Parasitologica Polonica, 15 phalan parasites from marine fishes of Bay of Bengal, Puri (40/50): 305-312. Coast, Orissa. Indian Journal of Helminthology, 43 (2): 108-188. GRABDA-KAZUBSKA(B.) & EJSYMONT (L.), 1969.- Studies on the GUPTA (N.K.) & GUPTA (K.), (1970) 1971.- New species of the morphology, variability and systematic status of Echinorhyn- genus Centrorhynchus Luhe, 1911 (Acanthocephala) from chus borealis Linstow, 190 I (Acanthocephala, Echinorhynchi- crow, Corvus splendens.Research Bulletin of the Panjab Uni- dae). Acta Parasitologica Polonica, 17 (1/19): 65-87. versity of Science, 21 (112): 101-104. GRACIA-RoDRIGO (A.), 1960.- Contribuci6n al conocimiento de GUPTA (N.K.) & GUPTA(K.), (1970) 1971.- Longicollum indicum los acantocefalos parasites de los peces en Venezuela. Revista n. sp., an acanthocephalan parasite of a marine fish, Tylosurus Veterinaria Yeneeolana, 8 (42): 3-32. strongylurus from Emakulam (South India). Research Bulletin GRIMM (O.A.), 1870.- Ueber Echinorhynchus pachysomus und of the Panjab University of Science, 21 (3/4): 507-510. andere Rundwurrner. Beschreibung zweier neuer Rundwurrner GUPTA (N.K.) & GUPTA (K.), (1971) 1972.- Two new species of aus der Abteilung der Acanthocephala. Travaux de la Societe Pseudorhadinorhynchus Achmerow et Dombrovskaja-Achme- d'Histoire Naturelle de St. Petersbourg, 1 (3): 214-223. rova, 194 I (Acanthocephala, family Micracanthorhynchinidae GUBANOV (N.M.), 1955.- [Faune helminthologique des animaux Yamaguti, 1961) from marine food fishes at Emakulum. Re- dinteret econornique de la Mer d'Okhot et de l'Ocean Pacifi- search Bulletin of the Panjab University of Science, 22 (3/4): que]. Autoreference and dissertation held in 1952, In: Faune 345-351. helminthologique des mammiferes marins a la lumiere de leur GUPTA(N.K.) & GUPTA(K.), (1971) 1972.- Observations on two ecologie et de leur phylog enie (S.L. Delyamure), Akademii acanthocephalan parasites from marine fishes in India. Research Nauk S.S.S.R., Moskva, 5 I 7 pp. (in Russian) Bulletin of the Panjab University of Science, 22 (3/4): 417-420. GUPTA (P.V.), 1950.- On some stages in the development of the GUPTA (N.K.) & GUPTA(K.), 1972.- Observations on genus Cen- acanthocephalan genus Centrorhynchus. Indian Journal of Hel- trorhynchus Luhe, 1911 (Acanthocephala: Gigantorhynchidea). minthology, 2 (1): 41-48. Research Bulletin of the Panjab University of Science, 23: 1-12. GUPTA (P.D.), 1976.- A new species of the genus Mediorhynchus GUPTA (N.K.) & lAIN(M.), 1973.- On a new acanthocephalan (Acanthocephala, Gigantorhynchidae) from the great Indian parasite, Mediorhynchus bullocki, from a crow pheasant at bustard, Choriotis nigriceps (Vigors). Journal of the Bombay Chandigarh. Zoologischer Anzeiger, 191: 320-324. Natural History Society, 73 (I): 182-183. GUPTA (N.K.) & lAIN (M.), 1975.- On two already known species GUPTA (P.C.) & KUMAR (P.), (1983) 1987.- Hypoechinorhynchus of Acanthocephala of the genus Porrorchis Fukui, 1929 (Gigan- golvani sp. novo (Acanthocephala: Hypoechinorhynchidae) from torhynchidea, Prosthorhynchidae) from Chandigarh, India. Acta estuarine fish, Clarius (Clarias) batrachus of Chilka Lake, Oris- Parasitologica Polonica, 23 (34): 38\-387. sa. Kanpur University Research Journal, Science, 4: 85-89. GUPTA(N.K.) & lAIN(M.), 1977.- On Serrasentis socialis (Leidy, GUPTA (P.e.) & KUMAR (P.), (1984) 1987.- Two new species of 1851) Van Cleave, 1924 and discussion on synonymy of some acanthocephalan parasite (family Rhadinorhynchidae) of marine related species with it. In: All-India Symposium on Helmintho- fishes, from Bay of Bengal off Puri Coast, Orissa. Kanpur Uni- logy (Srinagar), Abstracts: 3-4. versity Research Journal, Science,S: 75-80. GUPTA (N.K.) & lAIN(M.), 1979.- On Neoechinorhynchus cirrhi- GUPTA (V.) & FATMA (5.), (1979) 1980.- On three new species of nae sp. n., an acanthocephalan parasite from a fresh-water fish at acanthocephalan parasites of marine fishes of Mandapam, Tamil Chandigarh. Helminthologia, 16: 147-151. Nomenclature of the Acanthocephala 189

GUPTA (N.K.) & JAIN(M.), 1979.-A note on Filisoma indicum Jenaisch Zeitschrift fiir Naturwissenschaften, 25: 113-231. Van Cleave, 1928. Indian Journal of Parasitology, 3: 135-137. HAMANN (0.), 1891.- Zur Kenntnis des Baues der Nemathelmin- GUPTA(N.K.) & JAIN (M.), 1980.- On Goacanthus panajiensis n. then. Sitzungberichte der Konigliche Preuss Akademie der Wis- g., n. sp. (Acanthocephala, ll1iosentidae Golvan, 1960). Rivista senschaften, Berlin: 57-61. di Parassitologia, 41: 47-59. HAMANN (0.), 1892.- Das System der Acanthocephalen. Zoologi- GUPTA(N.K.) & JAIN (M.), 1980.- On a new species of acantho- cher Anzeiger, 15 (392): 195-197. cephalan parasite, Acanthosentis golvani n. sp. from a feshwater HA RADA (1.), 1928.- A new species of Acanthocephala from the fish at Chandigarh. Revista Iberica de Parasitologia, 40: 149-160. Japanese bonito, Euthynnus vagans.Japanese Journal of Zoo- GUPTA (N.K.) & JAIN(M.), 1983.- On three already known species logy, Tokyo, 2 (I): 1-4. of the genus Neoechinorhynchus Hamann, 1892 (Acanthocepha- HARADA (I.), 1929.- Ueber eine neue Species der Acanthocepha- la) from fish of the Arabian Sea. Acta Parasitologica Polonica, len. Japanese Journal of Zoology, Tokyo, 2 (2 ): 195-198. 28 (38/42): 407-416. HARADA (1.), 1935.- Zur Acanthocephalenfauna von Japan. GUPTA (N.K.) & LATA (V.), 1967.- On the morphology, anatomy Memoirs of the Faculty of Science and Agriculture, Taihoku and histology of Echinopardalis pardalis, an acanthocephalan Imperial University of Formosa.Japan, 14 (2): 7-23. parasite of Felis domestica in India. Research Bulletin of the HA RADA (1.), 1938.- Acanthocephalen aus Formosa (1.). Dr. A. Panjab University of Science, 18 (3/4): 395-403. OKA Jub., Annotationes Zoologicae Japonenses, 17 (3/4): 419427. GUPTA (N.K.) & LATA (V.), 1967.- On six new species of Acan- HARRI GTON (R.W.) & PILSBURY (N.R. Jr.), 1938.- Preliminary thocephala from some hosts in India. Research Bulle- report on the internal parasites of the herring gull. Bulletin of tin of the Pan jab University of Science, 18 (3/4): 253-268. Bowdoin Collection (mimeographed), 4: 8-9. GUPTA(N.K.) & LATA (V.), 1967.- Observations on eight already HARRIS (M.T.), 1973.- A new acanthocephalan from an East Afri- known acanthocephalan parasites from vertebrate hosts. Research Bulle- can galliform bird. Bulletin of the British Museum (Natural His- tin of the Panjab University of Science, 18 (3/4): 325-341. tory), Zoology, 24 (9): 455-460. GUPTA (S.P.) & KAJAJI (V.L.), 1969.- A new acanthocephalan, HARTWICH(G.), (1952) 1953.- Gordiorhynchus freundi n. sp. Acanthosentis cameroni n. sp., from the intestine of a freshwater (Acanthocephala, Polymorphidae) aus dem Sekretar (Sagittarius fish, Rohtee cotio Skyes, 1839, from Lucknow. Journal of the serpentarius Miller). Wissenschaftliche Zeitschrift der Martin Fisheries Research Board of Canada, 26 (4): 965-974. Luther Universitdt, Ha lie, Wittenberg, 2 (II), Mathematisch- GUPTA (S.P.) & NAQvl (M.), (1981) 1983.- Acanthocephalan para- Naturwissenchaftliche Reihe, 6: 917-919. sites of fishes. On three new acanthocephalan parasites of the HARTWICH (G.), 1956.- Sudarnerikanische Acanthocephalen aus genus Pseudorhadinorhynchus Achmerow et Dombrowskaja- der Zoologischen Sammlung des Bayerischen Staats. Zoologi- Achmerova, 1941 from marine fishes of Arabian Sea. Indian cher Anzeiger, 156 (11/12): 299-308. Journal of Helminthology, 33 (2): 144-153. HEl ZE(K. VO ), 1934.- Wissenchaftliche Ergebnisse der Deuts- GUPTA (S.P.) & NAQvl (M.), (1982) 1984.- On some acanthoce- chen Groenland Expedition Alfred Wegener 1929 und 1930-31. phalan parasites of marine fishes from India and Pakistan. Anthropologie und Zoologie, 6: 185-189. Indian Journal of Helminthology, 34 (I): 61-84. HEI ZE (K. VON), 1934.- Ein neuer Acanthocephale (Rhadi- GUPTA(S.P.) & VERMA(S.L.), (1976) 1977.- On a new Acantho- norhynchus meyeri n. sp.) aus dem Zoologischen Museum zu cephala, Acanthogyrus guptai n. sp. from the intestine of a fresh- Greifswald. Zoologicher Anzeiger, 108 (9/I0): 225-256. water fish, Labeo rohita (Ham.) from Lucknow. Rivista di HEINZE (K. VON), 1936.- Ein neuer Acanthocephalen aus der Parassitoliogia, 37: 143-149. Polarrnowe, Zoologischer Anzeiger, 115: 139-142. GUPTA (S.P.) & VERMA (S.L.), 1980.- On three new Acanthoce- HEITZ (A.), (1918 ) 1920.- Salmo salar L., seine Parasitenfauna phala parasites of the genus Pallisentis Van Cleave, 1928 from und seine Ernahrung im Meer und im Susswasser. Archiv fur fresh water fishes of Lucknow. Helminthologia, 17: 269-282. Hydrobiologie und Planktonkunde, 12 (2): 311-372. GUPTA (R.C.) & GUPTA (S.P.), 1975.- On a new acanthocephalan, HEITZ(A.), (1919) 1920.- Salmo salar L., seine Parasitenfauna Metarhadinorhynchus thaparai n. sp. from the intestine of a und seine Ernahrung im Meer und im Susswasser. Archiv fur marine fish, Arius platystomus (Griffith) from Puri, Orissa. In: Hydrobiologie und Planktonkunde, 12 (3): 485-561. Proceedings of the 62th Indian Science Congress (Delhi), Cal- HERMANN (J.), 1782.- Helminthologische Beobachtungen. Halle, cutta Indian Science Congress Association, Part Ill, Gr. B: 215. 17: 171-182. GVOZDEV (E.V.) & SOBOLEVA (T.N.), 1966.- [Faune des acantho- HINE (P.M.), 1977.- Acanthocephalus galaxii n. sp. parasitic in cephales des oiseaux sauvages dans le Kazakhstan meridional]. Galaxias maculatus (Jenyns, 1842) in the Lema Stream, New Zealand. Materialy Nauchnykh Konerentsii Vsesoyuznogo Obshchestva Journal of the Royal Society of New Zealand, 7 (1): 51-57. Gelminthoogov, Part Ill: 84-89. (in Russian) HOLCMA -SPECTOR (B.), MANE-GARZON (F.) & DEI-CAS (E.), HAFFNER (K. VON), 1939.-Untersuchungen iiber einen bisher 1977.- Ciclo evolutivo y descripcion de Falsifilicollis chasmagnathi unbekannten Acanthocephalen aus Schlitzruesselrn (Solenodon n. sp. (Acanthocephala). Revista de Biologia del Uruguay, 5 (2): 77-91. paradoxus Brandt). Zeitschrift fur Wissenschaftliche Zoologie, HOLLOWAY (H.L. Jr.), 1956.- A new species of the acanthocepha- 152 (2): 277-304. lan genus Centrorhynchus from Spilogale putorius (Linn.). HALEY (AJ.) & BULLOCK (W.L.), 1953.- A new species of Acan- A.SA Bulletin, 3 (l): 10-1 1. thocephala from the sunfish Lepomis gibbosus (Linnaeus) with a HOLLOWAY(H.L. Jr.), 1958.- Notes on the helminths of mammals redescription of the family Fessisentidae Van Cleave, 1931. in the Mountain Lake Region. Part Ill. The genus Centrorhyn- American Midland Naturalist, 50: 202-205. chus in North America with the description of a new species. 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Ihre Entwicklungsgeschischte, Histogenie und Hu (J .0.), 1990.- [Une nouvelle espece de Prosthenorchis (Oli- Anatomie nebst Beitrligen zur Systematik und Biologie. gacanthorhynchida: Oligacanthorhynchidae)]. Acta Veterina- 190 Y.J. GOLVAN

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Annals of Tropical Medici- HUFFMAN(D.G.) & KLlEVER(R.G.), 1977.- Echinorhynchus can- ne and Parasitology, Liverpool, 8 (I): 106-112. yonensis sp. n. (Acanthocephala) from Maynea californica JOHNSTON(T.H.), 1916.- A census of the endoparasites recorded (Osteichthyes: Zoarcidae) from the Monterey Submarine Can- as occurring in Queensland, arranged under their host. Procee- yon, California. Proceedings of the Helminthological Society of dings of the Royal Society of Queensland, 28: 31. Washington, 44 (2): 171-176. JOHNSTON(T.H.), 1918.- Notes on miscellaneous endoparasites. HUGHES(R.c.) & Moo RE(G.A.), 1943.- Acanthocephalus vancle- Proceedings of the Royal Society of Queensland, 30: 215. avei: a new echinorhynchid worm from a salamander. American JOHNSTON(T.H.), 1937.- Entozoa from the Australian hair seal. Pro- Midland Naturalist, 29: 724-729. ceedings of the Linnean Society of New South Wales, 62: 9-16. iCHTHARA(A.), KATO(K.), KAMEGAI(S.) & MAcHlDA (M.), 1968.- JOHNSTON(T.H.), 1942.- Australian Acanthocephala. 3. Transac- On the parasites of fishes and shell-fishes in Sagami Bay. 4. tions of the Royal Society of South Australia, 66: 250-254. Parasitic helminths of the mackrel, Pneumatophorus japonicus JOHNSTON(T.H.) & BEST (E.W.), 1937.- Report on the Acantho- (Houttuyn). Research Bulletin of the Meguro Parasitological cephala. Scientific Reports of the Australian Antarctic Expedi- Museum, 2: 45-57. tion 1911-1914, ser. C, Zoolog., Botan., 10 (2): 5-20. IHERING(H. VON), 1902.- Die Helminthen als Hilfsmittel der zoo- JOHNSTON(T.H.) & BEST (E.W.), 1942.- Australian Acanthoce- geographischen Forschung. Zoologischer Anzeiger, 26 (686): phala. 3. Transactions of the Royal Society of South Australia, 42-51. 66: 250-254. JAIN (M.), 1980.- Centrorhynchus splendi Gupta et Gupta, 1970 JOHNSTON(T.H.) & BEST (E.W.), 1943.- Australian Acanthoce- (Acanthocephala) as the adult of C. batrachus Das, 1952. Indian phala. 4. Transactions of the Royal Society of South Australia, Journal of Parasitology, 4 (1): 25. 67: 226-230. JAIN (M.) & GUPTA (N.K.), 1979.- On two already known species JOHNSTON (T.H.) & CLELAND (B.1.), 1911.- Echinorhynchus of the genus Pallisentis Van Cleave, 1928 (Acanthocephala) and pomatostomi (n. sp.) a subcutaneous parasites of Australian discussion on the validity of Pallisentis buckleyi Tadros, 1966 birds. Journal and Proceedings of the Royal Society of New and genus Devendrosentis Sahay, Sinha et Ghosh, 1971. Hel- South Wales, 45: 111-114. minthologia, 16 (3): 173-183. JOHNSTON(T.H.) & DELAND(E.W.), 1929.- Australian Acanthoce- JAIN(M.) & GUPTA(N.K.), 1979.- On Saccosentis ophiocephali n. phala. I. Census of recorded hosts and parasites. Transactions of the sp. 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JAIN (M.) & GUPTA (N.K.), 1981.- On two species of the genus JOHNSTON(T.H.) & EDMONDS(S.J.), 1948.- Australian Acantho- Acanthocephalus Koelreuther, 1771 (Acanthocephala: Echi- cephala. 7. Transactions of the Royal Society of South Australia, norhynchidae) including the description of a new species, A. 72: 69-76. goaensis. Rivista di Parassitologia, 42: 163-176. JOHNSTON(T.H.) & EDMONDS(S.J.), 1951.- Australian Acantho- JAlN (M.) & GUPTA (N.K.), 1983.- On some already known spe- cephala. 8. Transactions of the Royal Society of South Australia, cies of the genus Centrorhynchus Luhe, 1911 (Acanthocephala) 74: 1-5. from varied vertebrate hosts in India. Research Bulletin of the JOHNSTON(T.H.) & EDMONDS(S.1.), 1952.- Australian Acantho- Panjab University of Science, 34 (1/2): 15-17. cephala. 9. Transactions of the Royal Society of South Australia, JAMES(P.M.), 1954.- On some helminths from British small mam- 75: 16-21. mals with a redescription of Echinorhynchus rosai. 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KAISER (J.E.), 1893.- Die Acanthocephalen und ihre Entwicklung. thologia, Bratislava, 15 (1/4): 803-811. (in Russian) Bibliotheca Zoologica Cassel, 7: 1-136. KHOKHLOVA(I.G.), 1975.- [Revision du genre Arhythmorhynchus KASIMOV(G.), 1956.- [Faune helminthologique des gallinaces sau- Luhe, 1911 (Acanthocephala: Polymorphidae)]. Trudy Gelmin- vages et domestiques]. Akademii Nauk S.S.S.R.: 488-554. (in thologicheskoi Laboratorii, Akademii Nauk S.S.S.R. (Issledova- Russian) nie po Sistematike Zhirnennym Tsiklam i Biokhimii Celmintov), KATAOKA (N.) & MOMMA(K), 1937.- Helminths from the sal mo- 25: 195-203. (in Russian) noid fish, Plecoglossus altivelis T. et S. Collected Papers from KHOKHLOVA(I.G.), 1978.- [Revue taxonomique des acanthoce- the Faculty of Medicine, Osaka Imperial University: 67-72. phales d'oiseaux de I'U.R.S.S]. Trudy Gelminthologicheskoi KAuPP (B.F.), 1909.- Echinorhynchus canis. Review of American Laboratorii, Akademii Nauk S.S.S.R. 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Acad. nis (Shipley, 1903) Southwell and Macfie, 1925 (Acanthocepha- Scientif. Imper. Petropolitanae, 15: 494-503. la: Echinorhynchidae) from the black-spotted toad, Bufo mela- KOELREUTHER (J.T.), 1771.- Descriptio Piscis e Coregonorum no stictus from Boyor (Indonesia). Canadian Journal of genere, russicae Sig vocari. Historico-anatomica. Nov. Com- Zoology, 60: 356-360. ment. Acad. Scientif. Imper. Petropolitanae, 15: 504-516. KEpPNER (EJ.), 1974.- The life history of Paulisentis missourien- KOSTYLEW( .N.), 1914.- Ueber die Stellung einiger Acanthocep- sis n. sp.(Acanthocephala, Neoechinorhynchidae) from the halen in System. Zoologicher Anzeiger, 44: 186-188. creek chubb Semotilus atromaculatus.Transactions of the Ame- KOSTYLEW(.. ), 1915.- Contributions a la faune des acantho- rican Microscopical Society, 93: 89-100. cephales de la Russie. Annuaire du Musee Zoologique de l'Aca- KHAN (A.): see MY KHAN. emie Imperiale des Sciences de Petrograd, 20: 389-394. 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N.), 1926.- Zur Kenntnis der Acanthocephalen der Fis- 1911 (Acanthocephala: Polymorphidae)]. (Voprosy Biologii, che des Schwarzen Meeres. Zoologischer Anzeiger, 67: 177-183. Fiziologii i Biokhimii Gelminthol. Zhivotnykh. i Rastenii. Trudy KOSTYLEW(N.N.), 1928.- Acanthocephalen der Fische der Gokts- Celminthologicheskoi Laboratorii, Akademii Nauk S.S.S.R., 21: cha-Sees. Centralblatt fur Bakteriologie, Parasitenkunde und 120-126. (in Russian) Infektionskrankheiten, Abteilung I. Orig., 108: 146-150. KHOKHLOVA (I.G.), 1971.- [Acanthocephales d'oiseaux dans la KOSTYLEW (N.N .), 1941.- [Contribution a la connaissance de la region de Yakutsk. (A.S.S.R.)]. Trudy Celminthologicheskoi faune helminthologique des poissons d'Extreme-Orienr]. Trudy Laboratorii, Akademii Nauk S.S.S.R. (Teoreticheskie Voprosy Voenno-Meditsinskoi Akademii Krasnoi Armii im S.M. Kirova, Obshchei Gelmintologii), 22: 215-223. (in Russian) 25: 336-340. 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Pakistan Jour- (2): 483-498. nal of Zoology, 24 (2): 143-144. LI STOW (0. VON), 1878.- Compendium der Helminthologie bis KURASHVILI(B.E.), 1955.- [Acanthocephales des oiseaux de Geor- rum. Jahre 1878. Ein Verzeichniss der bekannten Helminthen, gie]. Soobshchenie Akademii Nauk S.S.S.R., 16 (9): 723-730. (in uSW. Hahn Edit., Hannover, 382 pp. Russian) LI STOW (0. VO ), 1879.- Compendium der Helminthologie bis LAMBL (W.), 1859.- Mikroskopische Untersuchungen der Darm- zum Jahre /878. Ein Yerzeichniss der bekannten Helminthen, Excrete. Prager.Viertelj. Prakt.Heilkd., 61: 45. uSW.Nachtrag. Hahn Edit., Hannover, 151 pp. LAURS(R.M.) & Mc CAULEY (1.E.), 1964.- A new acanthocephalan LINSTOW (0. VON), 1879.- Helminthologische Untersuchungen. from the Pacific saury. Journal of Parasitology, 50: 469-471. Jahresheft des Vereins for vaterlandische Naturkunde in Wurt- LEBEDEV (B.I.), 1967.- [Australorhynchus tetramorphacanthus n. temberg, Stuttgart, 35: 313-342. g., n. sp.(Acanthocephala, Rhadinorhynchidae) recolte chez des LI STOW (0. VON), 1880.- Helminthologische Untersuchungen. poissons d' Australie et de Nouvelle Zelande]. Zoologicheskii Archiv fur Naturgeschichte, 46 (I): 49. Zhurnal, 46: 279-282. (in Russian) LINSTOW (0. VON), 1882.- Helminthologische Studien. Archiv fur LEHMANN (V.W.), 1953. - Bobwhite population fluctuations and Naturgeschichte, 48 (1): 1-25. vitamin A. Transactions of the North American Wildlife Confe- LINSTOW (0. VON), 1883.- Nematoden, Trematoden und Acantho- rence, 18: 199-246. cephalen gesammelt nach Prof. Fedtschenko in Turkestan. LEI CHING (H.), 1984.- Description of Neoechinorhynchus salmo- Archiv fur Naturgeschichte, 49: 304-305. nis sp. n. (Acanthocephala: Neoechinorhynchidae) from fresh- LINSTOW (0. VON), 1884.- Helminthologisches. Archiv fur Natur- water fishes of British Columbia. Journal of Parasitology, 70: geschichte,50: 125-145. 286-291. LINSTOW (0. VON), 1888.- Helminthologisches. Archiv fur Natur- LEIDY (1.), 1850.- Contributions to Helminthology. Proceedings of geschichte, 54 (I): 235-246. the Academy of Natural Science, Philadelphia, 5: 96-98. LINSTOW (0. VON), (1876) 1888.- Entozoa collected by H.M.S. LEIDY (1.), 1851.- Contributions to Helminthology. Proceedings of Challenger during the years 1873-1876. Report Challenger, 23 the Academy of Natural Science, Philadelphia, 5: 205-210. (71): 1-18. LEIDY (1.), 1851.- Descriptions of new species of Entozoa. Procee- LINSTOW (0. VON), 1890.- Beitrag zur Kenntnis der Vogeltaenien dings of the Academy of Natural Science, Philadelphia, 5: 155-156. nebst Bemerkungen an neuen und unbekannten Helminthen. LEIDY (1.), 1856.- A synopsis of Entozoa and some of their ecto- Archiv fur Naturgeschichte, 56 (I): 171-188. congeners observed by the author. Proceedings of the Academy LI STOW(0. VON), 1892.- Helminthen von Siidgeorgien. ach der of Natural Science, Philadelphia, 8: 42-58. Ausbeute der deutschen Station von 1882-1883. Jahrbuch der LEIDY (1.), 1887.- Notice on some parasitic worms. Proceedings of Hamburgischen Wissenschaftlichen Anstalten, 9 (2): 59-77. the Academy of Natural Science, Philadelphia, 39: 20-24. LI STOW(0. VON), 1894.- Helminthologischen Studien. Jenaische LEIDY (1.), 1888.- Parasites of the rock-fish. Proceedings of the Zeitschrift fiir Naturwissenchaften, 28: 328-342. Academy of Natural Science, Philadelphia, 40: 166-167. LINSTOW (0. VON), 1896.- Nemathelminthen. Hamburger Magal- LEtDY (1.), 1888.- Parasites of the striped bass. Proceedings of the haensische Sammelreise.L. Friedrischen und Mittarbeiter, Academy of Natural Science, Philadelphia, 40: 96-98. Hamburg, 22 pp. LEIDY (1.), 1888.- Entozoa of the Terrapin. Proceedings of the LINSTOW (0. VON), 1897 a.- Nemathelminthen grosstenteils in Academy of Natural Science, Philadelphia, 40: 127-128. Madagascar gesammelt. Archiv for Naturgeschichte, 63 (I): 27-34. LEIDY (1.), 1891.- Notices on Entozoa. Proceedings of the Aca- LINSTOW (0. VON), 1897 b.- Nemathelminthen, gesammelt von demy of Natural Science, Philadelphia, 43: 410-418. Herrn Prof. Dahl in Bisrnark-Achipel. Archiv fur Naturges- LELPER(R.T.) & ATKINSON(E.L.), 1914.- Helminths of the British chichte, 63 (1) 281-291. Nomenclature of the Acanthocephala 193

LiNSTOW (0. VON), 1898.- Nemathelminthen von Herrn Richard che Annalen, Zeitschrift Geschichte Zoologie Herausgegeben Semon in Australien gesammelt. R. Semon Zoologische Fors- von B. Braun, Wurzburg, 1: 139-250. chungsreisen in Australien Vo!. 5, Denkschrift der Medizinis- LOHE (M.), 1905.- Geschichte und Ergebnisse der Echinorhyn- chen Naturwissenchaftlichen Gesellschaft Jena, 8: 467-472. chen-Forschung bis auf Westrumb (1821) (Mit Bemerkungen LINSTOW (0. VON), 1901 a.- Helminthen von den Ufern des Nyas- iiber alte und neue Gattungen der Acanthocephalen). Zoologis- sa-Sees. Ein Beitrag zur Helminthenfauna von Siid-Africa. che Annalen, Zeitschrift Geschichie Zoologie Herausgegeben Jenaische Zeitschrift fur Naturwissenschaften, 35: 409-428. von B. Braun, wurzburg, 1: 139-353. LiNSTOW (0. VON), 1901 b.- Entozoa des Zoologischen Museums LOHE (M.), 1911.- Acanthocephalen. Register der Acanthocepha- der Kaiserlichen Akademie der Wissenschaften zu St. Peters- len und parasitischen Plattwiirmer geordnet nach ihren Wirten. burg. Bulletin de l' Academic Imperiale des Sciences de St. Pro! Dr. Brauer, Die Siisswasserfauna Deutschlands, Eine Petersbourg, 15 (3): 271-291. Exkursionsfauna, Jena, 16: 144 pp. LINSTOW (0. VON), 1902.- Beobachtungen an neuen und bekann- LOHE (M.), 1912.- Zur Kenntnis der Acanthocephalen. Zoologis- ten Nemathelminthen. Archiv fur Mikroskopische Anatomie, 60: che Jahrbiicher, Suppl. 15: 271-306. 217-232. LUNDSTROM (A.), 1941.- Corynosoma mergi n. sp., eine neue Art LINSTOW (0. VON), 1904.- Neue Helminthen aus Westafrika. Cen- der Acanthocephalen. Kungl. Fysiografiska Sallskapets I Lund tralblatt fur Bakteriologie, Parasitenkunde und Infektionskrank- Forhandlingar, 11 (11): 1-7. heiten, 36 (3): 380-383. LUNDSTROM (A.), 1942.- Die Acanthocephalen Schwedens mit LINSTOW (0. VON), 1905.- Helminthen der Russischen Polarexpe- Ausnahme der Fischacanthocephalen von Siisswasserstandor- dition (1900-1903). Memoires de l'Academie Imperiale des Scien- ten. Doctor Thesis in Sciences, University of Lund, Sweden, ces de St. Petersbourg, ser. 8, Class. Phys. Math., 18 (1): 17 pp. 238 pp. LINSTOW (0. VON), 1908 a.- Helminthen von Herrn Edward Jacob- LYNCH (I.E.), 1936.- New species of Neoechinorhynchus from the son in Java (Semarang) gesammelt. Notes from the Leiden Western sucker, Catostomus macrocheilus Girard. Transactions Museum, 29: 81. of the American Microscopical Society, 55 (1): 21-43. LINSTOW (0. VON), 1908 b.- Beitrage zur Kenntnis der Fauna Tur- LYSTER (L.L.), 1940.- Parasites of some Canadian sea mammals. kestans auf Grund des von D.D. Fedtschenko gesammelten Canadian Journal of Research, 18: 395-409. Materials. l. Nemathelminthen. Travaux de la Societe Imperiale MAC CALLUM (G.A.), 1914.- Notes on the genus Camallanus and des Naturalistes de. St. Petersbourg, 37: 1-6. other Nematodes from various hosts. Zoopathologica, I: 123-134. LiNSTOW (0. VON), 1908.- Helminthen, Nematoden und Acantho- MAcHADo FILHO (D.A.), (1940) 1941.- Pesquisas helrninthologi- cephalen. Denkschrift der Medizinischen Naturwissenchaftli- cas realizadas no Estado de Mato Grosso. Acanthocephala. chen Gesellschaft Jena, 13: 19-28. Memorias do lnstituto Oswaldo Cruz; 35 (3): 593-601. LiNTON (E.), 1889.- Notes on Entozoa of marine fishes of New MAcHADo FILHO (D.A.), 1941.- Sabre alguns acantocefalos prove- England with description of several new species. Report of the nientes do Estado do Mato Grosso. Revista Brasileira de Biolo- U.S. Commisioner of Fish and Fisheries for 1886,14: 453-511. gia, 1 (1): 57-61. LiNTC(N(E.), 1891.- Notes on Entozoa of marine fishes with des- MAcHADo FILHO (D.A.), 1941.- Sabre alguns acantocefalos do cription of new species. (Part 3). Report of the U.S. Commisio- Estado do Para. Revista Brasileira de Biologia, 1 (2): 223-226. ner of Fish and Fisheriesfor1888, 16: 523-542. MAcHADo FILHO (D.A.), 1947.- Revisao do genero Polyacant- LINTON (E.), 1892.- Notes on avian Entozoa. Proceedings of the horhynchus Travassos, 1920 (Acanthocephala, Rhadinorhynchi- U.S. National Museum of Washington, 15: 87-113. dae). Revista Brasileira de Biologia, 7 (2): 195-201. LINTON (E.), 1893.- On fish Entozoa from Yellostone National MAcHADo FILHO (D.A.), 1948.- Echinorhynchidae do Brasil. I. Park. Report of the U.S. Commisioner of Fish and Fisheries Tres especies novas de «Echinorhynchus» Zoega in Miiller, for1889-1891, 17: 545-564. 1776 e redescricao de «Echinorhynchus jucundus» Travassos, LlNTON (E.), 1901.- Parasites of fishes of the Woods Hole Region. 1923. Revista Brasileira de Biologia, 8 (2): 265-273. Bulletin of the U.S. Fisheries Commisioner for 1889, 19: 405-492. MACHADO FILHO (D.A.), 1949.- Uma nova especie do genero LINTON (E.), 1905.- Parasites of fishes of Beaufort, North Caroli- Oncicola Travassos, 1916 (Acanthocephala). Revista Brasileira na. Bulletin of the U.S. Bureau for Fisheries. 1904, 24: 323-428. de Biologia, 9 (4): 467-468. LINTON (E.), 1908.- Notes on parasites of Bermuda Fishes. Proce- MACHADOFTLHO(D.A.), 1950.- Revisao do genero Prosthenorchis edings of the U.S. National Museum of Washington, 23: 85-126. Travassos, 1915 (Acanthocephala). Memorias do lnstituto LINTON (E.), 1911.- Notes on the distribution of Entozoa of North Oswaldo Crur, 48: 495-544. American Marine Fishes. Proceedings of the 7th International MAcHADo FILHO (D.A.), 1951.- Uma nova especie do genero Zoological Congress: 686-696. Atactorhynchus Van Cleave, 1935 (Acanthocephala, Neoechi- LINTON (E.), 1932.- On the taxonomic position of Echinorhynchus norhynchidae). Revista Brasileira de Biologia, 11 (1): 29-31. sagittifer Linton. Science, 76: 193. MAcHADo FILHO (D.A.), 1954.- Uma nova especie do genero Neo- LITTLE (J.W.) & HOPKlNS (S.H.), 1968.- Neoechinorhynchus con- echinorhynchus (Hamann) (Neoechinorhynchidae, Acanthoce- trictus sp. n., an acanthocephalan from Texas turtles. Procee- phala). Revista Brasileira de Biologia, 14 (1): 55-57. dings of the Helminthological Society of Washington, 35: 46-49. MACHADO FILHO (D.A.), 1959.- Echinorhynchidae do Brasil. Il. LlU LIQING, WANG BAODUE & YANG TAO, 1981.- [Notes sur les Nova especie do genero Echinorhynchus Zoega in Miiller, 1776. parasites de la carpe nue de Qinhai Hu]. Journal of Fisheries of Memorias do Instituto Oswaldo Cruz: 57 (2): 195-198. China, 5 (4): 295-299. (in Chinese) MACHADO FILHO (D.A.), 1959.- Neoechinorhynchus spectabilis n. LOCKWOOD (S.), 1872.- A new Entozoan (Kaleops anguillae n. g. sp. (Neoechinorhynchidae, Acanthocephala). Revista Brasileira and n. sp.) from the eel (Anguilla acutirostris) with Woodcuts. de Biologia, 19 (2): 191-194. American Naturalist, 6: 449-454. MAcHADo FTLHO(D.A.), 1959.- Uma nova especie do genero Neo- LOPEZ-NEYRA (C.R.), 1946.- Compendio de Helminthologfa Iberi- echinorhynchus Hamann, 1892, parasita de «peixe-rnartir» do ea (Continuaci6n). Parte Il, Cap. IV. Clase Acanthocephala Paraguai (Neoechinorhynchidae, Archiacanthocephala). Revista Rudolphi, 1808. Revista Iberica de Parasitologia, 6: 3-50. Brasileira de Biologia, 19 (4): 379-381. LOHE (M.), 1904.- Geschichte und Ergebnisse der Echinorhyn- MACHADO FTLHO(D.A.), 1960.- Urn novo genero da familia «Qua- chen-Forschung bis auf Westrumb (1821) (Mit Bemerkungen drigyridae» Van Cleave, 1920 (Metacanthocephala, Palaecan- iiber alte und neue Gattungen der Acanthocephalen). Zoologis- thocephala). Revista Brasileira de Biologia, 20 (1): 79-84. 194 Y.J. GOLVAN

MACHADOFILHO(D.A.), 1961.- Contribuicao para 0 conhecimento Dentitruncus Sinzar, 1955. Parassitologia, 18 (113): 71-78. do genero Corynosoma Luhe, 1904 (Metacanthocepha1a, Palae- MANTER (H. W .), 1928.- Notes on the eggs and larvae of the acanthocephala, Polymorphidae). Memorias do Instituto Oswal- thorny-headed worm of hogs. Transactions of the American do Cruz; 59 (2): 181-184. Microscopical Society, 47: 342-347. MACHADOFILHO(D.A.), 1961.-Nova contribuicao para 0 conheci- MAO (G.L.), 1979.- [Etudes sur les acanthocephales des poissons mento do genero "Corynosoma" Luhe, 1904 (Metacanthocepha- d'eau douce de Liaoning, Chine, avec la description d'une nou- la, Palaeacanthocephala, Polymorphidae). Revista Brasileira de velle espece]. Acta Zootaxonomica Sinica, 4 (4): 316-321. (in Biologia, 21 (3): 249-251. Chinese) MACHADOFiLHO(D.A.), 1962.- Sabre Corynosoma enrietti Molfi et MARCHAND (B) & VASSILIADES(G.), 1982.- Mediorhynchus mat- Fernandes, 1953 (Metacanthocephala, Palaeacanthocephala, tei sp. n. (Acanthocephala, Gigantorhynchidae) from Tockus Polymorphidae). Revista Brasileira de Biologia, 22 (2): 143-151. erythrorhynchus (Aves), the red-beaked in West Africa. MACHADOFILHO(D.A.), 1962.- Nova especie do genero Oncicola Journal of Parasitology, 68: 1142-1145. Travassos, 1916 (Metacanthocephala, Archiacanthocephala, MARGOLlS(L.), 1955.- Corynosoma hadweni Van Cleave, a pro- Pachysentidae). Revista Brasileira de Biologia, 22 (4): 367-369. bable synonym of C. wegeneri Heinze (Acanthocephala). Jour- MACHADOFILHO(D.A.), 1963.- Oncicola chibigouzouensis n. sp., nal of Parasitology, 41: 326-327. parasito de Felis chibigouzou (Griff.) (Archiacanthocephala, MAROTEL (G.), 1899.- Sur un type particulier dacanthocephale Pachysentidae). Revista Brasileira de Biologia, 23 (1): 15-17. (Echinorhynchus tenuicaudatus n. sp.). Note prelirninaire. MACHADOFILHO (D.A.), 1963.- Oncicola paracampanulata n. sp., Comptes-Rendus Hebdomadaires des Seances de la Societe de parasito de Herpailurus jaguarundi (Fish.) (Archiacanthocephala, Biologie de Paris, 10 (6): 226-228. Pachysentidae). Revista Brasileira de Biologia, 23 (2): 153-155. MAROTEL (G.), 1899.- Etude zoologique d' Echinorhynchus tenui- MACHADO FILHO(D.A.), 1964.- Contribuicao para 0 conhecimen- caudatus novo sp. Archives de Parasitologie, 2: 291-302. to do genero Oncicola Travassos, 1916 (Archiacanthocephala, MARTEAU (M.), 1977.- Oncicola venezuelensis n. sp. (Archiacan- Pachysentidae). Revista Brasileira de Biologia, 24 (1): 23-30. thocephala, Oligacanthorhynchidae), parasite de I' (Felis MACHADO FILHO (D.A.), 1964.- Sabre 0 genero «Oligacan- pardalis L.). Annales de Parasitologie Humaine et Comparee, thorhynchus» Travassos, 1915 (Archiacanthocephala, Oligacan- 52: 25-33. thorhynchidae). 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DE), 1905.- Monographie des Acanthocephales soma Luhe, 1904 (Acanthocephala, Echinorhynchidea). Revista d'oiseaux. Revue Suisse de Zoologie, 13: 195-387. Brasileira de Biologia, 30 (3): 377-379. MEYER (A.), 1928.- Die Furchung nebst Eibildung, Reifung und MACHADO FILHO(D.A.), 1970.- Nova especie do genero Acantho- Befruchtung des Gigantorhynchus gigas (Ein Beitrag zur Mor- cephalus Koelreuther, 1771 (Acanthocephala, Echinorhynchi- phologie der Acanthocephalen). Zoologische Jahrbiicher, Abtei- dae). Atas da Sociedad de Biologia do Rio de Janeiro, 13 (1/2): lung fur Anatomie und Ontogenie der Tiere, 50: 177-218. 53-54. MEYER (A.), 1931.- Neue Acanthocephalen aus dem Berliner MACHADO FILHO(D.A.) & IBANEZ (H.N.), 1967.- Fauna helminto- Museum. Begrundung eines neuen Acanthocephalensystems auf 16gica do Peru: Lueheia cajabambensis sp. n. (Acanthocephala, Griind einer Untersuchung der Berliner Sammlung. Zoologische Echinorhynchidae, Plagiorhynchinae). Nota previa. Atas da Jahrbucher,Abteilung fur Systematik, Oekologie und Geogra- Sociedad de Biologia do Rio de Janeiro, 10 (6): 157-158. phie der Tiere, 62 (112): 53-108. MACHALSKA(J.), 1981.- Helminth fauna of birds of the genus Tur- MEYER (A.), 1931.- Die Stellung des Genus Heterosentis Van Cle- dus examined during their spring and autum migration. 11. ave, 1931 in Acanthocephalensystem. Zoologischer Anzeiger, Acanthocephalans. Acta Parasitologica Polonica, 28 (12/24): 94 (9/10): 258-265. 171-177. MEYER (A.), 1931.- Gordiorhynchus, ein neues Acanthocephalen- MACHIDA(M.) & ARAKl (J.), 1982.- Redescription of Pseudorhadi- genus mit inner ovarialer Pseudosegmentierung. Zoologische norhynchus leuciscus (Krotov et Petrotschenko, 1956). Research Iarhbiicher, Abteilung fur Systematik, Oekologie und Geogra- Bulletin of the Meguro Parasitological Museum, 8: 49-51. phie der Tiere, 60: 457-470. MAC LEOD (J.A.), 1933.- A parasitological survey of the genus Cite- MEYER (A.), 1932.- Acanthocephala. Dr. H.G. Bronns, Klassen llus in Manitoba. Canadian Journal of Research, 9: 108-127. und Ordnungen des Tier-Reichs, Leipzig, Bd. 4, 2 Abt., 2 Buch, MALM (A.W.), 1867.- Monographie illustree du balenoptere trou- I Lief.: 1-332. ve le 29 Octobre 1865 sur la cote occidentale de la Suede. MEYER (A.), 1933.- Acanthocephala. Dr. H.G. Bronns, Klassen Stockholm, 95 pp. und Ordnungen des Tier-Reichs, Leipzig, Bd. 4, 2 Abt., 2 Buch, MAMAEV (Y.L.), 1970.- [Helminthes de quelques poissons 2 Lief.: 333-582. d'inreret econornique du Golfe du Tonkin]. In: [Helminthes de MEYER (A.), 1938.- Klasse Acanthocephalen, Kratzer. In: Die I'Asie du Sud-Est] (P.G. Oshmarin, Y.L. Mamaev & B.T. Lede- Tierwelt Mitteleuropas (P. Brohmer, P. Ehrmann & G. Ulmer), bev), Izadatel'stvo «Nauka», Moskva: 187-190. (in Russian) Leipzig, 1 (6): 40 pp MANE-GARZON (F.) & DEI-CAS (E.), 1974.- Un acanthocephale MIESCHER (F.), 1841 (???).- Ueber Echinorhynchus in Otis houba- nouveau Woif.fhugelia matercula n. gen., n. sp. de l'intestin de rae. Verhandlungen der Schweizer Naturforschung Gesells- Fitzroyia lineata (Jenyns) (poisson) en Uruguay. Annales de chaft, Zurich, 26: 75-76. Parasitologie Humaine et Comparee, 49: 83-89. MILLZNER (R.), 1924.- A larval acanthocephalid Centrorhynchus MANILLA (G), ORECCHIA(P.) & PAGGI (L.), 1976.- Parassitofauna californicus n. sp. from the mesentery of Hyla regilla. Univer- di Salmo trutta L. del Fiume Tirino. Nota 1. Ridescrizione di sity of California, Publications in Zoology, 26: 325-327. Dentitruncus truttae Sinzar, 1955 e considerazioni suI genere MOGHE (M.A.) & DAs (E.N.), 1953.- On a new species of Acan- Nomenclature of the Acanthocephala 195

thocephala of the genus Arhythmorhynchus (Luhe, 1911), from lium. Daniae et Norvegiae Indigenarum Characteres, Nomina et India. Thapar Commemoration Volume: 211-216. Synonyma imprimis popularium. Havniae, I: 214-215. MOLFI (A.) & FREITAS-FERNANDES (B. DE), 1953.- Corynosoma MULLER (O.F.), 1777.- Zoologiae Danicae seu Animalium Daniae et enrietti n. sp. parasita de patos e marrecos domesticos (Pal aea- Norvegiae rariorum ac minus notorum. !cones. Havniae, I (3): 4. canthocephala: Polymorphidae). Arquivos de Biologia e Tecno- MULLER (O.F.), 1778.- Von den Thieren in den Eingeweiden der logia, Curitiba, 8 (I): 3-6. Thiere, insonderheit vom Kratzer im Hecht. Der Naturfoscher, MOLl (R.), 1858.- Prospectus helminthum quae in parte seconda Halle, 12: 178-196. prodromi faunae helminthologicae venetae continentur. Sit- MULLER (O.F.), 1779.- Zoologia Danica seu Animalium Daniae et zungsbetrichte der konigliche Akademie der Wissenschaften zu Norvegiae rariorum ac minus notorum Descriptiones et Histo- Wien, Mathem=Naturwiss. Kl., 30: 141. ria. Havniae et Lipsiae, I: 103 pp. MOLlN(R.), 1859.- Cephalocotylea e Nematoidea. Parte seconda MULLER (O.F.), 1780 a.- Unterbrochene Bemiihungen bei den prodromi faunae helminthologicae venetae continentur. Sit- Intestinalwiirmern. Schriften der Berlinischen Gesellschaft zungsbetrichte der konigliche Akademie der Wissenschaften zu naturforschende Freunde, Berlin, I: 201-210. Wien, Mathem=Naturwiss.Kl., 38: 16. MULLER (O.F.), 1780 b.-!cones. Havniae, 2. MOLlN(R.), 1861.- Prodromus faunae helminthologicae venetae MULLER(O.F.), 1784.- Zoologia Danica seu Animalium Daniae et adjects disquisitionibus anatomicis et criticis. Denkschrift der Norvegiae rariorum ac minus notorum Descriptiones et Histo- konigliche Akademie der Wissenschaften zu Wien, Mathem> ria. Havniae et Lipsiae, 2. Naturwiss. Kl., 19: 189-338. MUZZAL (P.M.) & BUCKNER (R.L.), 1982.- Neoechinorhynchus MONTICELLl (F.S.), 1887.- Osservazioni intorno ad alcune specie limi sp. n.(Acanthocephala, Neoechinorhynchidae) from the di Acantocefali. Bolletino della Societa di Naturalisti in Napoli, central mudminnow, Umbra limi. Proceedings of the Helmin- ser. I (I): 1-11; 19-29. thological Society of Washington, 49: 231-234. MONTICELLl (F.S.), 1900.- Sui parassiti del Relalecus glesne . NADAKAL (A.M.), JOHN (K.O.) & JACOB (A.), 1990.- Metarhadi- Monitore Zoologico ltaliano, Firenre, 11 (Suppl.): 36-37. norhynchus valiyathurae sp. nov., an acanthocephalid worm MONTICELLl (F.S.), 1905.- Su di un Echinorinco dell a Collezione parasitic in the marine fish, Caranx melampygus (Cuv, and Val). del Museo Zoologico di Napoli (Echinorhynchus rhytitodes Zoologischer Anzeiger, 225 (5/6): 377-382. Montic.). Annuario del Museo Zoologico della Reale Universita NAKAJlMA(K.) & EGUSA(S.), 1975.- [Pseudorhadinorhynchus di Napoli, Nuova Serie, I (25): 1-13. samegaiensis n. sp., (Acanthocephala), parasite de la truite arc- MONTICELLl (F.S.), 1905.- Sull' Echinorhynchus aurantiacus en-ciel delevage, Salmo gairdneri, au Japon). Fish Pathology, Risso. Annuario del Museo Zoologico della Reale Universita di 10 (I): 58-68. (in Japanese) Napoli, Nuova Serie, I (30): 1-3. AMA(H.S.) & RATHORE (G.S.), (1982) 1984.- The acanthoce- Mo TICELLl (F.S.), 1905.- A proposito di una proposta classifica- phalan genus Centrorhynchus occurring in Indian birds with zione degli Acantocefali (per una rettifica). Bolletino della description of two new species from Rajasthan. Indian Journal Societa di Naturalisti in Napoli, 19: 217-218. of Helminthology, 34 (2): 143-150. MONTREUIL(P.L.), 1958.- Corynosoma magdaleni sp. nov. NEILAND (K.A.), 1962.- Alaskan species of acanthocephalan (Acanthocephala), a parasite of the grey seal in Eastern Canada. genus Corynosoma Luehe, 1904. Journal of Parasitology, 48: Canadian Journal of Zoology, 36: 205-215. 69-75. MORAVEC (F.) & AM IN(A.), 1978.- Some helminth parasites, NEIVA(A.Marques da Cunha) & TRAVAssos (L.), 1914.- Contri- excluding Monogenea, from fishes of Afghanistan. Acta Scien- buicoes Parasitologicas. 1. Memorias do Instituto Oswaldo Cruz, tiarum Naturalium Academiae Scientiarum Bohemoslovacae, 6: 180-191. Brno, 12 (6): 1-45. NEVEU-LEMAIRE(M.), 1905.- Sur un nouvel acanthocephale MORDINOVA (T.N.), 1988.- [Neorhadinorhynchus myctophumi (Echinorhynchus orestiae nov. sp.) parasite des poissons du (Acanthocephala) a new acanthocephalan species from Atlantic genre Orestias. Comptes Rendus des Seances de la Societe de myctophiid fish). Zoologicheski Zhurnal, 67 (9): 1411-1414. (in Biologie de Paris, 59: 31-32. Russian) NEVEU-LEMAIRE (M.), 1905.- Sur les parasites des poissons du MORDINOVA (T.N.) & PARUKHIN(A.M.), 1978.- [Golvanacanthus genre Orestias (Mission de Crequi-Montfort et de Senechal de problematicus n. sp., un nouvel acanthocephale de Gammarus la Grange). Bulletin de la Societe Philomatique de Paris: 1-5. olivii de la Mer Noire). Biologiya Morya, Kiev, (Parazitofauna NEVEU-LEMAlRE(M.), 1936.- Traite d'Helminthologie medicate et Zhivotnykh Yuehnykb Morei), 45: 42-44. (in Russian) veterinaire. Monographie, 1520 pp. MORINI (E.G.) & BOERo (1.J.), 1960.- Corynosoma otariae n. sp. NICKOL (B.B.), 1967.- Fessisentis necturorum sp. n.(Acanthoce- (Acanthocephala, Polymorphidae) parasite de un lobo marino phala: Fessisentidae), a parasite of the , (Otariaflavescens). In: Aetas y Trabajos dell Congreso Suda- Necturus beyeri. Journal of Parasitology, 53: 1292-1294. mericano de Zoologia (La Plata, 1959 J, 2: 229-234. NICKOL (B.B.), 1969.- Acanthocephala of Louisina Picidae with MORISITA (T.), 1937.- [Etudes des acanthocephales de Tsintao). description of a new species of Mediorhynchus.Journal of Zikken.lgaku Zasshi, 21 (12): 1909-1913. (in Japanese) Parasitology, 55: 324-328. MOROZOV (F .. ),1940.- [Nouvelles especes dacanthocephales de NICKOL (B.B.), 1972.- Fessisentis, a genus of acanthocephalan la loutre de mer (En hydra lutris L.)). Trudy Gor'koskogo parasitic in North American poikilotherms. Journal of Parasito- Gosundarstvennogo Pedagogicheskogo Intsitut im M. Gorkogo, logy, 58: 282-289. 5: 61-65. (in Russian) lCKOL(B.B.), 1972.- Two species of Acanthocephala from Aus- MOROZOV (F. .), 1951.- [Helminthes des loups de la Reserve tralian fishes with description of Arhythmacanthus paraplagu- Naturelle d'Etat de Mordovian). Trudy Gelminthologicheskoi siarum sp. n. Journal of Parasitology, 58: 778-780. Laboratorii, Akademii Nauk A.N. S.S.R., 5: 146-149. (in Rus- ICKOL(B.B.), 1983.- Centrorhynchus kuntzi from the U.S.A with sian) description of the male and redescription of C. spinosus (Acan- MUHLlNG(P.), 1898.- Die Helminthenfauna der Wirbeltiere Ost- thocephala, Centrorhynchidae). Journal of Parasitology, 69: preussens. Archiv fur Naturgeschichte, 64 (I): 1-118. 221-225. MUHLING (P.), 1898.- Studien aus Ostpreussens Helminthenfauna. NICKOL (B.B.) & DUNAGAN(T.T.), 1989.- Reconsideration of the Zoologischer Anzeiger, 21: 16-24. acanthocephalan genus Echinopardalis, with a description of MULLER (O.F.), 1776.- Zoologiae Danicae Prodromus seu Anima- adult E. atrata and a key to genera of the Oligacanthorhynchi- 196 Y.J. GOLVAN

dae. Proceedings of the Helminthological Society of Washing- gen. e n. sp.(Palaeacanthocephala: Rhadinorhynchidae Travas- ton, 56: 8-13. sos, 1923) parasita di Blennius pavo e proposta di una nuova sot- NICKOL (B.B.) & ERNST (CH.), 1987.- Neoechinorhynchus lingu- tofamiglia Golvanacanthinae novo subfarn. In: VII Congrezo latus sp. n. (Acanthocephala: Neoechinorhynchidae) from Pseu- Nazionale de la Societa Italiana di Parassitologia (Bologna), demys nelsoni (Reptilia: Emydidae) of Florida. Proceedings of Parassitologia, 14 (I): 175-18l. the Helminthological Society of Washington, 54: 146-149. PAGGI (L) & ORECCHIA (P.), 1983.- Paracanthocephaloides sole- NICKOL (B.B.) & HEARD (R.W.), 1970.- Arhythmorhynchus fras- ae (Porta, 1905) n. comb. (Arhythmacanthidae: Paracanthoce- soni from the clapper rail, Raltus longirostris, in North America. phaloidinae) acantocefalo parassita di Solea impar Benn. Paras- Journal of Parasitology, 56: 204-206. sitologia, 25 (I): 99-103. NICKOL (B.B.) & PADILHA(T.N.), 1979.- Neoechinorhynchus PAGGI (L.), ORECCHIA(P.) & DELLA SETA (G.), 1975.- Su di un paraguayensis (Acanthocephala: Neoechinorhynchidae) from nuovo acantocefalo Breizacanthus ligur sp. n. (Palaeacantho- Brazil. Journal of Parasitology, 65: 987-989. cephala: Arhythmacanthidae Yamaguti, 1935) parassita di alcu- NICKOL (B.B.) & THATCHER (V.E.), 1971.- Two new acanthoce- ne specie di pesci bentonici del Mar Ligure. Parassitologia, 17 phalans from neotropical fishes: Neoechinorhynchus prochilo- (1/3): 83-94. dorum sp. n. and Gorytocephalus plecostomorum gen. and sp. n. PAL (R.N.), 1963.- A new acanthocephalan parasite, Acanthosentis Journal of Parasitology, 57: 576-581. hilsai n. sp. from the Indian shad, Hilsa ilisha (Hamilton). NIKOLSKY(OK), 1974.- [Faune des acanthocephales des pinnipe- Indian Journal of Helminthology, 15 (2): 96-99. des de la partie marine Pacifique de l' Antarctique], lzvestiya PALLAS (P.S.), 1766.- Elenchus zoophytorum sistens generum Tikhookeanskogo Nauchno-Issledovatelskogo Instiuta Rybnogo adumbrationes generaliores et speciarum cognitarum, succinc- Khoryaistva i Okeanografii (T.I.NR.O.) (Pararity Morskikh tas descriptiones cum selectis auctorum synonymis . Hagae- Zhivotnykh), 88: 101-106. (in Russian) Comitum. NORONHA(D.), 1973.- Sobre Neoechinorhynchus curemai sp. n. PALLAS (P.S.), 1781.- Bemerkungen iiber die Bandwiirmer in (Acanthocephala: Neoechinorhynchidae). Atas da Sociedade de Menschen und Thieren. Neue nordische Beitrdge Physikal. und Biologia do Rio de Janeiro, 17 (I): 19-21. Geograph. Erd. Yolkbeschr. Naturg.Oekonom. (Petersburg und NORO HA(D.), 1984.- Remarks on Neoechinorhynchus curemai Leipzig), I (I): 39-112. oronha, 1973 (Eoacanthocephala: Neoechinorhynchidae). PAPERNA (I.), 1964.- Parasitic helminths of inland-water fishes in Memorias do Instituto Oswaldo Cruz, 79 (2): 271. Israel. Israel Journal of Zoology, 13: 1-20. NORONHA(D.), 1988.- Corynosoma clementei Giovannoni and PARONA (C), 1885.- Di alcuni elminti raccolti nel Sudan orientale Fernandes, 1965 and its synonym Corynosoma seropedicus da Beccari e P. Magretti. Annali del Museo Civico di Storia Machado Filho, 1956, an accidental parasite of Canis familiaris Naturale di Genova, 2 (2): 424-445. L, 1758. Arquivos Fluminenses de Medicina Yeterinaria, 3 (4): PARONA (C), 1890.- Di una nu ova species di Echinorhynchus (E. 119-120. novellae) parassita di un chirottero di Porto-Rico. Annali del NORONHA (D.), FABIO (S.P. DE) & PINTO (R.M.), 1987.- Golva- Museo Civico di Storia Naturale di Genova, sec. 2, 10 (30): 1-3. norhynchus golvani gen. n., sp. n. (Palaeacanthocephala, Allor- PARONA (C), 1893.- Sopra una straordinaria polielmintiasida echi- hadinorhynchinae). Atas da Sociedade de Biologia do Rio de norinco net Globicephalus svineval Flow., pescato net mare di Janeiro, 19 (I): 63-64. Genova. Alii delta Societa Ligustica di Scienze Naturali e Geo- NORO HA (D.), FABIO(S.P. DE) & ROLAS (FJoT), 1973.- Revisao graf,4: 1-11;314-324. da subfamilia Atactorhynchinae Petrotschenko, 1956 (Acantho- PARONA (C), 1894.- L'Elmintologia italiana da sui primi tempi cephala, Neoechinorhynchidae). Memorias do Instituto Oswaldo all'anno 1890. Storia sistematica, corologia e bibliographia. Atti Cruz; 71 (4): 369-382. delta Reale Universita di Genova, 13: 733 pp. NORONHA(D.), VINCENTE (U.), PINTO (R.M.) & FABIO (S.P. DE), PARUCHIN(AM.): see PARUKHIN(A.M.). 1986.- Gorgorhynchus trachinotus n. sp. and new hosts records PARUKHIN(A.M.), 1971.- [Une nouvelle espece dacanthocephale for Dollfusentis chandleri Golvan, 1969 (Palaeacanthocephala: de la famille des Rhadinorhynchidae, parasite d'un poisson de Echinorhynchoidea). Memorias do Instituto Oswaldo Cruz, 81 l'Ocean Indien]. Zoologicheskii Zhurnal, 50 (11): 1734-1737. (2): 145-148. (in Russian) OETINGER(D.F.) & BUCKNER (RL), 1976.- Acanthocephalus tah- PARUKHIN (AM.), 1973.- Femogibbosus assi novo gen. et spec., lequahensis sp. n. (Acanthocephala: Echinorhynchidae) from ein neuer Kratzer aus dem Indik. Angewandte Parasitologie, 14 the stippled darter, Etheostoma punctulatum (Agassiz), in north- (I): 18-23. easten Oklahoma. Journal of Parasitology, 62: 237-241. PARUKHIN(A.M.), 1973.- [Gorgorhynchus nemipteri n. sp., une OKULEWICZ (J.) & MARUSZEWSKI (W.), 1980.- Apororhynchus nouvelle espece dacanthocephale parasite d'un poisson de silesiacus sp. n. (Apororhynchidae, Acanthocephala), a parasite l'Ocean Indien]. Parazitologiya, 7 (I): 72-74. (in Russian) of birds (Passeriformes), Acta Parasitologica Poloni- PARUKHI (A.M.), 1980.- [Position systernatique de I'acanthoce- ca, 27 (46/60): 459-470. phale Serrasentoides fistulariae Parukhin, 1971]. Biologiya OLSSO (P.), 1894.- Bidrag till Skandinaviens Helrninthenfauna. Morya, Vladivostok, 1: 83-85. (in Russian) Kongl. Svenska Yetenskaps. Akad. Handlinger, Stockholm, 25, PARUKHIN(A.M.), 1982.- Uber die systematische Stellung von VoL 2. Serrasentoides fistulariae (Acanthocephala). Angewandte Para- ORTLEPP (RJ.), 1924.- On a collection of helminths from Dutch sitologie, 23 (2): 83-85. Guiana, Journal of Helminthology, 2: 15-40. PARUKHIN (A.M.), 1985.- [Nouvelles especes dacanthocephales ORTUBAY (S.G.), UBEDA (C), SEMENAS (LG.) & KENNEDY (C), de I'ordre des Palaeacanthocephala Meyer, 1931 chez les pois- 1991.- Pomphorhynchus patagonicus n. sp. (Acanthocephala: sons de lOcean Indien et de I'Atlantique Sud]. Ekologiya Pomphorhynchidae) from freshwater fishes from Patagonia, Morya, Kiev, 20: 26-29. (in Russian) Argentina. Journal of Parasitology, 77: 353-356. PARUKHlN(A.M.) & KOVALE KO(L.M.), 1976.- Rhadinorhynchus OSHMARlN(P.G.), 1968.- [Faune helminthologique des Cinclidae decapteri n. sp., parasite d'un poisson de I'Ocean Pacifique. en relation avec la question de la formation de la faune d'hel- Zoologicheskii Zhurnal, 55 (I): 137-138. minthes des animaux]. In: [Parasites des animaux et des PERRY (M.L), 1942.- A new species of the acanthocephalan genus plantes], Izdatelstvo «Nauka», Moskva, 4: 122-128. (in Russian) Filicollis. Journal of Parasitology, 28 (5): 385-388. PAGGI (L) & ORECCHIA(P.), 1972.- Golvanacanthus blennii novo PETROCHENKO(V.!.): see PETROTSCHENKO(V.!.). Nomenclature of the Acanthocephala 197

PETROTSCHENKO(V.I.), 1949.- [Nouvelles especes dacanthoce- PORTA(A.), 1909.- Gli acantocefali degli anfibi e dei rettili. Archi- phales d'oiseaux d' Asie Centrale]. Trudy Gelminthologicheskoi vio Zoologico, Napoli, 3: 225-259. Laboratorii, Akademii Nauk S.S.S.R., 2: 114-127.(in Russian) PORTA (A.), 1910.- Acantocefali nuovi 0 poco noti. Zoologischer PETROTSCHENKO(V.I.), 1950.- [Notes sur les acanthocephales des Anzeiger, 35 (22): 699-703. oiseaux de la Kirghizie du Sud]. Trudy Gelminthologicheskoi PORTA (A.), 1912.- SuI Gigantorhynchus spirula Olf. parassita Laboratorii, Akademii Nauk S.S.S.R., 4: 100-105. (in Russian) dell' Erinaceus algirus Duv. Zoologischer Anzeiger, 39 (5/6): PETROTSCHENKO(V.!.), 1950.- [Notes sur les acanthocephales des 233-235. oiseaux des lacs de Barbarin]. Trudy Gelminthologicheskoi PORTA (A.), 1913.- Acanthocefali dell a Nuova Caledonia e delle Laboratorii, Akademii Nauk S.S.S.R., 4: 106-107. (in Russian) isole Loyalty. F. Sarazin et 1. Roux. Nova Caledonia Zoologie, PETROTSCHENKO(V.I.), 1953.- [Acanthocephales des amphibiens I-Ill (4): 167-170. de I'U.R.S.S.]. In: [Contributions de Publications en Helmintho- PORTA(A.), 1914.- Acanthocephali nuovi e note sinonimiche. logie pour la Commemoration du 75eme Anniversaire de K.1. Zoologischer Anzeiger, 44: 483-485. Skrjabin]: 508-517. (in Russian) PRASAD (D.), SAHAY (U.) & SHAMBHUNATH,1969.- On a new spe- PETROTSCHENKO(V.I.), 1956.- [Acanthocephales des animaux cies of an acanthocephalan of the genus Acanthosentis Verma et domestiques et sauvages]. In: Editions de l'Academie des Scien- Datta, 1929 from a fresh water fish, Hilsa ilisha (Hamilton). ces de l'URSS, Societe des Helminthologistes de I'Union Sovieti- Zoologischer Anzeiger, 183 (3/4): 278-283. que, Moskva - English translation by Israel Programme of Scien- PUJATTI(D.), 1951.- Uma nuova specie del genere Echinopardalis tific Translations, Keter Press, Jerusalem (1971), I: 435 pp. Travassos, 1918 rinvenuta in India. Rivista di Parassitologia, 12 PETROTSCHENKO (V.I.), 1958.- [Acanthocephales des animaux (2): 119-128. domestiques et sauvages]. In: Editions de l'Academie des Scien- RAI (P.), 1967.- On four acanthocephalan genera parasitic in fresh- ces de l'URSS, Societe des Helminthologistes de I'Union Sovieti- water fishes with description of three new species. Indian Jour- que, Moskva - English translation by Israel Programme of Scien- nal of Helminthology, 19 (1): 27-44. tific Translations, Keter Press, Jerusalem (1971),2: 458 pp. RAILLlET (A.), 1918.- Echinorhynchus belgicus. Zoologischer PETROTSCHENKO(V.I.), 1959.- [Nouvelles especes dacanthocep- Anzeiger, 44: 483 hales d'oiseaux]. Rabot. Gelmintolog 80 Let. Skrjabin, Akad. RAfLLLET(A.), 1919.- Les acanthocephales des animaux dornesti- Selskohziaistv Nauk V.I. Lenina, I: 144-147. (in Russian) ques. Recueil de Medecine Wterinaire, 94 (7): 185-198. PETROTSCHE KO(V.I.) & PHAN THE VIET, 1969.- [Gordiorhyn- RAILLLET(A.) & HE RY(A.C.L.), 1907.- Nemathelminthes parasi- chus hagiangensis n. sp.(Acanthocephala, Gigantorhynchidae)]. tes. Expedition Antarctique Francaise 19031/905 (Dr. Jean Parazitologiya,3 (4): 375-377. (in Russian) Charcot), Sciences Naturelles, Documents Scientifiques: 11-15. PETROTSCHENKO (V.I.) & SMOGORZHEVSKAJA (L.A.), 1962.- RATHKE (1.), 1799.- lattagelser henhorende til Indvoldeormenes [Hemiechinosoma ponticum n. sp., n. gen. (Acanthocephala) de og Bloddyrenes Naturhistorie. Skrivt. a Naturhist. Selsk., Kjo- Phalacrocorax aristotelis, cormoran du littoral de la Mer benhavn,5 (1): 61-148. Noire]. Zoologicheskii Zhurnal, 41 (6): 936-939. (in Russian) REICHENSPERGER(A.), 1922.- Neue eigenartige Parasiten von Ter- PHIPPS (C.I.), 1774.- A voyage towards the North Pole. London. miten. Bulletin de la Societe Fribourgeoise,Sciences Nature- PODDER (T.N.), 1937.- A new species of Acanthocephala, N. top- lles, 26: 103-111. seyi n. sp. from a Calcutta fish, Polynemus heptadactylus (Cuv. RENGARAJU(V.) & DAS (E.N.), 1980.- A new species of the genus et Val.). Parasitology, 28: 365-369. Centrorhynchus Liihe, 1911 (Acanthocephala) from India. PODDER (T.N.), 1937.- On a new species of Neoechinorhynchus Record of the Zoological Survey of India, 76 (1/4): 83-85. parasitic on Mugil cephalus from Chilka Lake. Records of the RENGARAJU(V.) & DAS (E.N.), 1981.- On a new species of the Indian Museum, 39: 129-131. genus Oligacanthorhynchus Travassos, 1915 (Acanthocephala: PODDER (T.N .), 1938.- A new species of Acanthocephala, Acan- Oligacanthorhynchidae) from India. Record of the Zoological thosentis dattai n. sp. from a freshwater fish of Bengal, Barbus Survey of 1ndia, 78 (1/4): 37-40. ticto (Ham.-Buch.) and Barbus stigma (Cuv. et Val.). Parasito- RENNIE (1.), 1906.- «Scotia» collections. On Echinorhynchus logy, 30: 171-175. antarcticus n. sp. and its Allies. Proceedings of the Royal PODDER (T.N.), 1941.- On a new Acanthocephala, Acanthosentis Society of Edinburgh, 26: 437-446. sircari, from a Calcutta fish, Rasbora elonga (Ham.). Records of Risso (A.), 1826.- Histoire naturelle des principales productions the Indian Museum, 43: 137-142. d'Europe meridionale et principalement de celles des environs POLONIO (A.F.), 1859.- Prospectus helminthum qui in reptilibus et de Nice et des Alpes Maritimes. Paris, 5 (27): 402 pp. amphibiis faunae italicae continentur. Echinirhinchi. Padova, RODYUK (G.N.), 1984.- [Nouveaux representants du genre Mete- 20 pp.(reference in pp. 10-1 I). chinorhynchus (Acanthocephala), parasites des poissons dans la PORTA(A.), 1904.- Nota sugli Echinorinchi di pesci del Museo partie occidentale de I' Antarctique]. Zoologicheskii Zhurnal, 63 Zoologico di apoli. Annuario del Museo Zoologico della Reale (12): 1893-1896. (in Russian) Universita di Napoli, 1 (4): 14. RODYUK (G.N.), 1985.- [Faune parasitaire des poissons de la par- PORTA(A.), 1905.-Gli Echinorhynchi dei Pesci. Archivio Zoologi- tie Atlantique de I' Antarctique (archipel de la Georgie du Sud eo, Napoli, 2: 149-214. et des iles des Shetlands du Sud)]. In: [Parasitologie et Patho- PORTA (A.), 1906.- Ricerche sull' Echinorhynchus capitatus v. logie des Organismes Marins des Oceans du Monde] (WJ. Linstow e nota sulla sistematica degli Echinorinchi dei Cetacei. Hargis lr. edit.), NOAA, Technical Report, NMFS, 25: 31-32. Zoologischer Anzeiger, 30: 235-271. (in Russian) PORTA (A.), (1907) 1908.- Contributo allo studio degli acantocefa- RODYUK (G.N.), 1986.- [Nouvelles especes dacanthocephales du li dei pesci. Biologia, 1 (19): 377. genre Echinorhynchus (Echinorhynchidae) parasites de poissons PORTA (A.), 1908.- Gli acantocefali dei mammiferi. Nota preventi- de I' Atlantique du sud ouest]. Parazitologiya, 20 (3): 224-227. va. Archives de Parasitologie, Paris, 12 (2): 268-282. (in Russian) PORTA (A.), 1908.- Nota sugli acantocefali di mammiferi del ROITMAN (V.A.), 1961.- [Acanthocephales des poissons du bassin Museo Zoologico di Napoli. Annuario del Museo Zoologico de la Zeida]. Trudy Gelminthologicheskoi Laboratorii, Akade- della Reale Universita di Napoli, 2 (22): 1-6. mii Nauk S.S.S.R., 11: 194-207.(in Russian) PORTA (A.), 1909.- Gli acantocefa1i dei mammiferi. Archivio Zoo- RONALD (K.), 1957.-The metazoan parasites of the Heterosomata logico, Napoli, 4 (2): 239-285. of the Gulf of SI. Lawrence. I. Echinorhynchus laurentianus sp. 198 Y.J. GOLVA

novo (Acanthocephala, Echinorhynchidae). Canadian Journal of Universidad Nacional Aut6noma de Mexico: 493-50 I. Zoology, 35: 437-439. SALGADO-MALDONADO (G.), (1976) 1977.- Acantocefalos de RUDOLPHI(K.A.), 1793.- Observationes circa vermes intestinalis peces. H. Descripcion de un genero y especie nuevos (Acantho- quas consensu ampliss. facul. philos. in memoriam concilii cephala: Leptorhynchoididae) parasite de Centropomus robalito puriorum sacrorum in Suecia stabiliendorum gratia ante ducen- de la laguna de Caimanero, Sinaloa, Mexico. Anales del Institu- tos annos Upsaliae habiri praeside etc. Inaugural Dissertation, to de Biologia, Universidad Nacional Aut6noma de Mexico, Ser. Gryphiswaldiae, Pars I: 46 pp. Zoologia, 47 (I): 1-6. RUDOLPHI (K.A.), 1795.- Observationes circa vermes intestinalis SALGADO-MALDOADO (G.), (1976) 1978.- Acantocefalos de quas consensu ampliss. facul. philos. in memoriam concilii peces. Ill. Redescripcion de Dollfusentis chandleri Golvan, puriorum sacrorum in Suecia stabiliendorum gratia ante ducen- J 969 (Acanthocephala, ) y descripcion de una tos annos Upsaliae habiti praeside etc. Inaugural Dissertation, nueva especie del mismo genero. Anales del Instituto de Biolo- Gryphiswaldiae, Pars 2. gia, Universidad Nacional Aut6noma de Mexico, Ser. Zoologia, RUDOLPHI (K.A.), 1801.- Beobachtungen uber die Eingeweide- 47 (2): 19-34. wurmer. Wiedemann's Archiv Zoologie Zootomie, Braunsch- SALGADO-MALDO ADO (G.), (1978) 1979.- Acantocefalos de weig, 2 (1): 1-65. peces. IV. Descripcion de dos especies nuevas de Neoechi- RUDOLPHI (K.A.), 1802.- Forsetzung der Beobachtungen uber die norhynchus Hamann, 1892 (Acanthocephala, Neoechinorhyn- Eingeweidewurrner. Wiedmann's Archiv Zoologie Zootomie, chidae) y algunas consideraciones sobre este genero, Anales del Braunschweig, 2 (2): 1-67. Instituto de Biologia, Universidad Nacional Aut6noma de Mexi- RUDOLPHI(K.A.), 1808.- Entozoorum sive vermium intestinalium co, Ser. Zoologfa, 49 (I): 35-47. historia naturalis. Amsteldaemi-Treuttel et Wurtz Edit., I: 527 pp. SALGADO-MALDONADO (G.), (1978) 1979.- Acantocefalos de RUDOLPHI(K.A.), 1809.- Entozoorum sive vermium intestinalium peces. V. Redescripcion de cuatro especies de palaeacantocefa- historia natural is. Amsteldaemi-Treuttel et Wurtz Edit., Volume los parasites de peces de Mexico. Anales del Instituto de Biolo- 2, Part I: 457 pp. gia, Universidad Nacional Aut6noma de Mexico, Ser. Zoologfa, RUDOLPHI(K.A.), 1810.-Entozoorum sive vermium intestinalium 49 (I): 49-69. historia naturalis. Amsteldaemi-Treuttel et Wurtz Edit., Volume SALGADO-MALDOADO (G.), 1979.- Acantocefalos de peces. VI. 2, Part 2: 251-319. Hallazgo de Gorgorhynchoides bullocki Cable et Mafarachisi, RUDOLPHI(K.A.), 1819.- Entozoorum synopsis, cui accedunt man- 1970 (Acanthocephala, Arhythmacanthidae) y descripcion de tissa duplex et indices locupletissimi. A. Riicker Edit., Berolini, algunos de sus estadios juveniles. Anales del Instituto de Biolo- 811 pp. gia, Universidad Nacional Aut6noma de Mexico, Ser. Zoologfa, SAEED (R.) & BILQEES (F.M.), 1971.- Pallisentis magnum, new 50 (I): 35-50. species (Acanthocephala, Quadrigyridae), from the fish Wallago SALGADO-MALDONADO (G.), 1985.- Crecimiento alornetrico y attu of Kalri Lake, West Pakistan. Pakistan Journal of Zoology, consideracions taxonomicas sobre Neoechinorhynchus golvani 3 (2): 221-223. Salgado-Maldonado, 1978 (Acanthocephala: eoechinorhyn- SAEED (R.) & BILQEES (F.M.), 1972.- Acanthocephala of some fis- chidae) parasite de peces dulceacuicolas en Tabasco, Mexico. hes from the vicinity of Kalri Lake (Sind), West Pakistan. In: Universidad y Ciencia, 2 (3): 57-66. Helminth parasites of some vertebrates chiefly from fishes of SALGADO-MALDONADO (G.) & BARQUIN-ALVAREZ (N.P.), 1978.- West Pakistan (F.M. Bilquees, R. Saeed, R. Rehana, A. Khatoon Floridosentis elongatus Ward, 1953 y Contracaecum sp., para- & S.H. Kaikabad edit.), Agricultural Research Council, Govern- sitos de Mugil cephalus Linnaeus, 1758. Anales del Instituto de ment of Pakistan, Karachi, Pakistan: 82-86. Biologia, Universidad Nacional Aut6noma de Mexico, Ser. Zoo- SAHAY (U.) & SI HA(A.), 1969.- On a new host record of Cen- logia, 49 (I): 71-82. trorhynchus bengalensis Datta et Soota, 1954 (Centrorhynchi- SAOUD (M.F.A.),EI-NAFFAR (M.K.) & ABU-SI A(H.), 1974.- dae Van Cleave, 1916; Centrorhynchus Luhe, 1911). Zoologis- Neoechinorhynchus ichthyobori n. sp.(Acanthocephala: eo- cher Anzeiger, 183 (3/4): 292-298. echinorhynchidae) from a freshwater fish in the Sudan. Bulletin SAHAY (U.) & SINHA (A.), 1971.- On Mehrarhynchus mysti n. sp. of the Zoological Society of Egypt, 26: 89-93. (Echinorhynchidae South well et Macfie, 1925; Rhadinorhynchi- SARKAR (H.L.), 1953.- On a new acanthocephalan, Pallisentis dae Travassos, 1923, Rhadinorhynchinae Liihe, 1912) from the nandai n. sp. from the fish Nandus nandus (Ham.). In: Procee- intestine of a fresh water fish Mystus seenghala. Indian Journal dings of the 40th Indian Science Congress, Abstracts, Part Ill: of Helminthology, 23 (I): 66-70. 192-193. SAHAY(U.), NATH (S.) & SINHA(A.), 1967.- On an acanthocepha- SARKAR (H.L.), 1953.- On a new Acanthocephala Pallisentis nan- lan from a hill stream fish, Lepidocephalichthys guntea (Hamil- dai n. sp. from the fish Nandus nandus (Hamilton) with notes on ton). Zoologischer Anreiger, 178 (5/6): 348-353. the other species of the genus. Proceedings of the Zoological SAHAY (U .), SINHA(A.) & GHOSH (A.K.), 1971.- On Devendro- Society of Bengal, 6 (2): 139-147. sentis garuai gen. et sp. novo (Neoechinorhynchidae South well SARKAR (H.L.), 1956.- On a new Acanthocephala, Pallisentis coli- et Macfie, 1925; Quadrigyridae Van Cleave, 1920), Acantho- sai, from the fish Colisafasciatus (Bloch et Schneider) with a cephala from a fresh water Indian fish, Clupisoma g aruai note on Acanthogyrus acanthogyrus Thapar from the fish Labeo (Ham.). Annates de Parasitologie Humaine et Comparee, 46: rohita (Hamilton). Records of the Indian Museum, Year 1954, 69-80. 52 (2/4): 349-362. SAHAY (U.), SI HA(H.) & NARAYA (S.), 1967.- On an acantho- SARMIENTO(L.), 1954.- Gigantorhynchus ortizi n. sp., an acantho- cephalan of the genus Acanthosentis Verma et Datta, 1929 (Neo- cephalan from Metachirus nudicaudatus. Journal of Parasito- echinorhynchidea, Quadrigyridae) from a new host Mystus logy, 40: 448-452. seenghala (Sykes). Anales dellnstituto de Biologia, Universidad SCHINZ (R.), 1841.-«Miescher (Pr.)» 1841.In: Verhandlungen der Nacional Aut6noma de Mexico, Ser. Zoologra, 38 (I): 17-22. Schweizerischen Naturforschung Gesellschaft, 26 Versammlung SALGADO-MALDONADO (G.), 1977.- Acantocefalos de peces. I. (Zurich), Zoologische Sect.: 75-76. Descripcion de Caballerorhynchus lamothei gen. nov, sp. novo SCHMIDT (G.D.), 1963.-Arhythmorhynchus capellae sp. n. (Poly- (Acanthocephala: Fessisentidae) parasite de Diapterus olisthos- morphidae, Acanthocephala) a parasite of the common snipe tomus de Sontecomapan, Vera Cruz, Mexico. Excerta Parasito- Capella gallinago delicata. Journal of Parasitology, 40: 483-484. logica en Memoria del Doctor Eduardo Caballero y Caballero, SCHMIDT (G.D.), 1965.- Polymorphus swartzi sp. n., and other Nomenclature of the Acanthocephala 199

Acanthocephala of Alaskan ducks. Journal of Parasitology, 51 : SCHMIDT(G.D.) & HUGGINS (E.J .), 1973.- Acanthocephala of 809-813. South American fishes. Part 2. Palaeacanthocephala. Journal of SCHMIDT(G.D.), 1965.- Corynosoma bipapillum sp. n. from Bona- Parasitology, 59: 836-838. parte's gull Larus philadelphia in Alaska, with a note on C. SCHMIDT (G.D.) & Ku TZ (R.E.), 1966.- New and little-known const rict um. Van Cleave, 1918. Journal of Parasitology, 51: plagiorchid Acanthocephala from Taiwan and the Pescadores 814-816. Islands. Journal of Parasitology, 52: 520-527. SCHMIDT (G.D.), 1965.- A collection of Acanthocephala from fi- SCHMIDT (G.D.) & Ku TZ(R.E.), 1966.- Sphaerechinorhynchus shes of George Lake, Central Alaska. Canadian Journal of Zoo- serpenticola sp. n. (Acanthocephala: Sphaerechinorhynchinae), logy, 43: 651. a parasite of the Asian cobra, Naja naja (Canthor), in Borneo SCHMIDT(G.D.), 1969.- Polymorphus petroschenkoi sp. n. (Acan- (Malaysia). Journal of Parasitology, 52: 913-916. thocephala) from the red phalarope, Phalaropus fulicarius L., in SCHMIDT (G.D.) & Ku TZ(R.E.), 1967 a.- otes on the life cycle Alaska. Journal of Parasitology, 55: 335-336. of Polymorphus (Profilicollis) formosus sp. n. and records of SCHMIDT (G.D.), 1969.- Paracanthoceph alus rauschi sp. n. Arhythmorhynchus hispidus Van Cleave, 1925 (Acanthocepha- (Acanthocephala, Paracanthocephalidae) from grayling, Thyma- la) from Taiwan. Journal of Parasitology, 53: 805-809. lIus arcticus (Pallas), in Alaska. Canadian Journal of Zoology, SCHMIDT(G.D.) & KUNTZ (R.E.), 1967 b.- Revision of the Porror- 47: 383-385. chinae (Acanthocephala: Plagiorhynchidae) with description of SCHMIDT (G.D.), 1972 a.- Revision of the class Archiacanthoce- two new genera and three new species. Journal of Parasitology, phala Meyer, 1931 (Phylum Acanthocephala), with emphasis on 53: 130-141. Oligacanthorhynchidae Southwell et Macfie, 1925. Journal of SCHMIDT (G. D.) & KUNTZ (R.E.), 1969.- Centrorhynchus spilor- Parasitology, 58: 290-297. nae sp. n. (Acanthocephala) and other Centrorhynchidae from SCHMIDT (G.D.), 1972 b.- Oncicola schacheri sp. n. and other the Far East. Journal of Parasitology, 55: 329-334. Acanthocephala of Lebanese mammals. Journal of Parasito- SCHMIDT (G.D.) & KUNTZ (R.E.), 1977.- Revision of Mediorhyn- logy, 58: 279-281. chus Van Cleave, 1916 (Acanthocephala) with a key to species. SCHMIDT (G.D.), 1973.- Resurrection of Southwellina Witenberg, Journal of Parasitology, 63: 500-507. 1932, with a description of Southwellina dimorpha sp. n. and a SCHMIDT(G. D.) & NEILAND (K.A.), 1966.- Helminth fauna of key to genera in Polymorphidae (Acanthocephala). Journal of Nicaragua. III. Some Acanthocephala of birds, including three Parasitology, 59: 299-305. new species of Centrorhynchus. Journal of Parasitology, 52: SCHMIDT (G.D.), 1975.- Sphaerirostris wertheimae sp. n., and 739-745. other Acanthocephala from vetebrates of Israel. Journal of SCHMIDT (G.D.) & PAPERNA (I.), 1978.- Sclerocollum rubrimaris Parasitology, 61: 298-300. gen. et sp. n. (Rhadinorhynchidae: Gorgorhynchinae), and other SCHMIDT (G.D.), 1975.- Andracantha, a new genus of Acantho- Acanthocephala of marine fishes from Israel. Journal of Parasi- cephala (Polymorphidae) from fish-eating birds, with descrip- tology, 64: 846-850. tion of three species. Journal of Parasitology, 61: 615-620. SCHMIDT(G.D.), Escn (G.W.) & GIBBO S (J.W.), 1970.- Neoechi- SCHMIDT (G.D.), 1975.- Redrescription of Acanthosentis acanthu- norhynchus chelonos, a new species of acanthocephalan parasite ri Cable et Quick, 1954 (Acanthocephala: Quadrigyridae). Jour- of turtles. Proceedings of the Helminthological Society of Wa- nal of Parasitology, 61: 865-867. shington, 37: 172-174. SCHMIDT(G.D.), 1977.- Observations on the type specimens of SCH EIDER(G.), 1903.- Beitrage zur Kenntnis der Helrninthenfau- two species described by Lauro Travassos. Journal of Parasito- na des Finnischen Meerbusens. Acta Societatis pro Fauna et logy, 63 (2): 343. Flora Fennica, Helsingfors., 26 (3): 22-35. SCHMIDT(G.D.), 1977.- Oncicola martini sp. n., and other Archia- SCHRANK (F. VON P.), 1788.- Verteichnis der bischer hinliinglich. canthocephala of the Chaco Boreal, Paraguay. Journal of Para- bekannten Eingeweidewiirmer, nebst einer Abhandlung iiber sitology, 63: 508-510. ihre Anverwandtschaften. Miinchen, 116 pp. SCHMIDT (G.D.), 1981.- Plagiorhynchus formosus Van Cleave SCHRANK(F. VONP.), 1790.- Fortekning pa nagra hittils obeskrife- 1918, a synonym of Plagiorhynchus cylindraceus (Goeze, 1782) ne Intestinal krak. Kong/. Vetensk. Acad. Nya. Handlingar for ar Schmidt et Kuntz, 1966. Journal of Parasitology, 67: 597-598. XI,Stockholm: 118-126. SCHMIDT (G. D.), 1983.- What is Echinorhynchus pomatostomi SCHRANK(F. VONP.), 1792.- Verzeichnis einiger noch unbeschrei- Johnston et Cleland, 1912? Journal of Parasitology, 69: 397-399. bener Eingeweidewiirmer. Kg/. Schweden Akademie der Wis- SCHMIDT (G.D.) & CANARIS (A.G.), 1967.- Acanthocephala from senschaften, Neue Abhandlungen, IJ: 111-118. Kenya with description of two new species. Journal of Parasito- SEN (P.), 1938.- On a new species of Acanthocephala, Acantho- logy, 53: 634-637. sentis holospinus sp. novo from the fish Barbus stigma (Cuv. et SCHMIDT (G.D.) & DAILEY (M.D.), 1971.- Zoogeography and the Val.). Proceedings of the Indian Academy of Science, 7 (I, sect. generic status of Polymorphus (Polymorphus) cetaceum (Johns- B): 41-46. ton et Best, 1942) comb. n. (Acanthocephala). Proceedings of SEN (J.K.), 1975.- On a new species of Apororhynchus Shipley, the Helminthological Society of Washington, 38 (I): 137. 1899 (Apororhynchidea: Apororhynchidae) from India. Dr. B.S. SCHMIDT(G. D.) & DAILEY (M.D.), 1971.- A zoogeographic note on Chauhan Commemoration Volume 1975 (K.K. Tiwari & C.B. the acanthocephalan Corynosoma bullosum (Linstow, 1892). Tran- Srivastava edit.): 211-213. sactions of the American Microscopical Society, 90 (I): 94-95. SEN (J.K.) & CHAUHA(B.S.), (1972) 1973.- On some acantho- SCHMIDT(G.D.) & Du N (A.M.), 1974.- Neoncicola sinensis sp. n. cephalan parasites from Indian mammals with descriptions of (Acanthocephala: Oligacanthorhynchidae) from China. Journal two new species. Journal of the Zoological Society of India, 24 of Parasitology, 60: 626-627. (I): 39-45. SCHMIDT (G.D.) & EDMo DS(S.J.), 1989.- Australiformis semoni SHAKHTAKHTI SKAJA(E.M.), 1956.-[Vers parasites des oiseaux (Linstow, 1898) n. gen., n. comb. (Acanthocephala: Monilifor- domestiques et du gibier d'eau en Azerbaijan]. Trudy Gelminto- midae) from of Australia and ew Guinea. Journal logicheskoi Laboratorii, Akademii Nauk S.S.S.R., 8: 286-296. of Parasitology, 75: 215-217. (in Russian) SCHMIDT (G.D.) & HUGGI S (E.J.), 1973.- Acanthocephala of SHIPLEY (A.E.), 1897.- On Arhynchus hemignathi, a new genus of South American fishes. Part I. Eoacanthocephala. Journal of Acanthocephala. Quaterly Journal of Microscopical Science, Parasitology, 59: 829-835. new ser., 39 (2): 207-218. 200 Y.J. GOLVAN

SHIPLEY (A.E.), 1899 a.- Note: Arhynchus hemignathi. Quaterly Indian Science Congress Association, Abstracts, 57 (Ill): 450. Journal of Microscroscopical Science, new ser., 42 (3): 36l. SOOTA (T.D.) & KANSAL (K.C.), 1972.- The helminth fauna of SHIPLEY (A.E.), 1899 b.- Notes on the species of Echinorhynchus Andaman and Nicobar. Acanthocephala. Records of the Zoolo- parasitic in the Cetacea. Archives de Parasitologie, 2 (2): 262-269. gical Survey of India, 66 (1/4): 303-307. SHIPLEY (A.E.), 1903.- On the Entoparasites collected by the SOOTA (T.D.) & SEN (J.K.), 1956.-On a new species of Acantho- «Skeat Expedition» to Lower Siam and the Malay Peninsula in sentis Verma et Datta from Glossogobius giuris (Hamilton). the years 1899-1900. Proceedings of the Zoological. Society of Records of the Indian Museum Year 1954,52 (2/4): 363-365. London, 2: 145-156. SOUTHWELL (T.), 1927.- On a new species of Acanthocephala SINZAR (D.), 1955.- Porilog poznavanju entoparasita pastrmke (Oligoterorhynchus magnus) from a marine fish. Annals of Tro- Salmo trutta L. Prethodno. Saopstenje Glasnik Prir. Mus. Srps- pical Medicine and Parasitologie, 21: 165-169. ke., ser. B, Biolog., Nauke, 7 (4): 223-229. SOUTHWELL(T.) & MACFIE(J.W.S.), 1925.- On a collection of SITA, 1949.- Description d'Arhythmacanthus septacanthus. These Acanthocephala in the Liverpool School of Tropical Medicine. de Sciences (reference not precised). Annals of Tropical Medicine and Parasitologie, 19: 141-184. SKRJABIN(K.I.), 1913.- Zur Acanthocephalenfauna Russich Tur- SPENCERlONES (M.E.), 1990.- Heptamegacanthus niekerki n. g., n. kestans. A. Acanthocephalen der Sumpf- und Wasservogel. sp. (Acanthocephala: Oligacanthorhynchidae) from the south- Zoologische Jahrbiircher, Abteilung Systematik und Geograp- east African insectivore Chrysospalax trevelyani (Giinther, hie, 35 (4): 403-414. 1875). Systematic Parasitology, 15: 133-140. SKRJABIN(A.S.), 1966.- [Corynosoma mirabilis n. sp. parasite du SPREHN (C.E.W.), 1942.- Ein neuer Acanthocephale aus der cachalot]. In: [Faune helminthologique des animaux des mers Amsel. Deutsche Tieranliche Wochenschrift, Berlin und Mun- du Sud] (S.L. Delyamure edit.), Naukova Dumka, Kiev: 10-12. chen,23/24: 193. (in Russian) STfLES (C.W.) & HASSALL(A.), 1905.- The determination of generic SKRJABIN(A.S.), 1970.- [Une nouvelle espece dacanthocephale, types and a list of roundworms genera, with their original and type Bolbosoma tuberculata n. sp. (Polymorphidae Meyer, 1931), un species. Bulletin of the Bureaufor Industry, 79: 150 pp. parasite des baleines). Parazitologiya, 4 (4): 334-337. (in Rus- STOSSICH (M.), 1891.-Nuova serie di elminti veneti raccolti dal sian) Dr. P.A. Conte di Ninni. Bolletino della Societa Adriatica Sci. SKRJABIN (A.S.), 1972.- [Differences morphologiques chez les Natur. Trieste, 13: 109-116. acanthocephales Bolbosoma turbinella (Diesing, 1851)(fam. STRAND (E.), 1936.- Miscellanea nomenclatoria zoologica et pala- Polymorphidae) provenant soit de I'hemisphere Nord soit de eontologica. Folia Zoologica Hydrobiologia, Riga, 9: 167-176. l'hemisphere Sud). Paraziiologiya, 6 (1): 57-64.(in Russian) STRAND (E.), 1942.- Miscellanea nomenclatoria zoologica et pala- SKRJABIN(A.S.) & NIKOLSKY (O.R.), 1971.- [Corynosoma singu- eontologica. Folia Zoologica Hydrobiologia, Riga, 11: 387-388. laris n. sp. (Polymorphidae), un parasite des rnarnmiferes marins SUBRAHMANIAN(K.), 1927.- On a new genus of Acanthocephala de I'Antarctique]. Nauchnye Doklady Vysshei Shkoly, Biologi- from Rangoon. Annals and Magazine of Natural History, ser. 9, cheskie Nauki, 11: 7-9. (in Russian) 19: 175-179. SKRJABINA(E.S.), 1979.- [Neoechinorhynchus rutili (Acanthoce- SUBRAHMANIAN(K.), 1927.-On a small collection of Acanthoce- phala: Neoechinorhynchidae) des poissons de la zone littorale phala from Rangoon. Annals and Magazine of Natural History, marine glacee et de la zone interrnediaire de I'Amour]. Trudy ser. 9, 19 (9): 645-650. Gelminthologicheskoi Laboratorii, Akademii Nauk S.S.S.R. SUBRAHMANIAN(K.), 1937.- Pallisentis nagpurensis (Acanthocepha- (Gelminty Zhivotnykli Rastenii), 29: 131-135. (in Russian) la) (Bahlerao, 1931). Zoologischer Anzeiger, 119 (3/4): 111-112. SMALES (L.R.), 1986.- Polymorphidae (Acanthocephala) from SUMNER (F.B.), OSBORN (R.C.) & COLE (L.J.), 1913.- A biologi- Australian mammals with descriptions of two new species. Sys- cal survey of the waters of Woods Hole and vicinity. Part Il, tematic Parasitology, 8: 91-100. Section 3. Bulletin of the U.S. Bureau of Fisheries: 31. 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SOLONCHENKO (A.I.), 1968.- [Faune parasitaire du maquereau SZlDAT (L.) & GRAEFE (G.), 1967.- Estudios sobre la fauna de Scomber colias Gmelin habitant l'Ocean Atlantique pres de la parasites de peces antarticos. n. Los parasites de Parachae- cote Sud-Ouest de I'Afrique]. Biologiya Morya, 14: 90-95. (in nichthys charcoti. Servicio Hidrogrdfico Naval, Armada Argen- Russian) tina, Republica Argentina, Public. H, 911: 1-27. SOLOWJEW(P.T.), (1889) 1912.- Vers parasites des Oiseaux du TADROS (G.), 1966.- On three new Acanthocephala of the genera Turkestan Russe. Annales du Musee de Zoologie, St. Peters- Pallisentis Van Cleave, Saccosentis gen. novo and Acanthoce- bourg, 17: 86-115. phalus Koelreuther, from fishes. Journal of Helminthology, 40: SONSINO (P.), 1896.- Forme nuove 0 poco conosciute in parte 155-180. indeterminate di entozoi raccolti od osservati in Egitto. 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TADROS (G.), ISKANDER (A.R.) & WASSEF (N.A.), 1979.- On an TRAVASSOS (L.), 1923.- Gigantorhynchidos novos. AFolha Medi- intestinal cestode and acanthocephalan from the Nile and Red ca, Rio de Janeiro, 4 (2): 12. Sea fishes with histopathologic study of their habitat. Journal of TRAVASSOS(L.), 1923.-lnformac;:oes sobre a fauna helrninthologica the Egyptian Society of Parasitology, 9 (I): 143-157. de Mato Grosso (lI Nota). A Folha Medica, Rio de Janeiro, 4 (2): THAPAR (G.S.), 1927.- On Acanthogyrus n. gen. from the intesti- 12-55. ne of the Indian fish Labeo rohita, with a note on the classifi- TRAVASSOS(L.), (1920) 1923.- Contribucao para 0 conhecimento cation of the Acanthocephala. Journal of Helminthology, 5: dos acantocefalos da sub-familia Centrorhynchinae. A Folha 109-120. Medica, Rio de Janeiro, 4 (4): 29-4l. THAPAR (G.S.), 1930.- On Farzandia, a new genus of acanthoce- TRAVASSOS(L.), (1924) 1925.-Contribuicoes para 0 conhecirnen- phalid worms from the intestine of Ophiocephalus marulius. to da fauna helmintolojica brazileira. XVII. Revisao dos acan- Annals and Magazine of Natural History, ser. 10,6: 77-81. . thocephalos brazileiros. 1. Fam. Gigantorhynchidae Hamann, THATCHER (V.E.), 1979.- Uma nova especie de Gorytocephalus 1892. Suplemento. Memorias do lnstituto Oswaldo Cruz; 17 (2): Nickol e Thatcher, 1971 (Acanthocephala, Neoechinorhynchi- 365-387. dae) do acari bodo (Pisces: Loricariidae) da Arnazonia, Brasil. TRAVASSOS (L.), 1925.- Quelques acanthocephales nouveaux. Acta Amazonica, 9 (1): 199-202. Comptes Rendus Hebdomadaires des Seances de la Societe de THATCHER (V.E.), 1980.- Rhadinorhynchus plagioscionis n. sp. Biologie de Paris, 95: 935-937. (Acanthocephala: Rhadinorhynchidae) in «pescada» (Plagios- TRAVASSOS (L.), 1926.- Contribuicoes para 0 conhecimento da cion squamosissimusi da Arnazonia Brasileira. Acta Amazonica, fauna helmintolojica brazileira. XX. Revisao dos acanthocepha- 10 (4): 835-839. los brazileiros. Parte 11. Familia Echinorhynchidae Hamann, THATCHER(V.E.), 1981.- Neoechinorhynchus pterodoridis n. sp. 1892, sub-farn. Centrorhynchinae Travassos, 1919. Memorias (Acanthocephala: Neoechinorhynchidae) do bacu liso (Pterodo- do lnstituto Oswaldo Cruz; 19 (I): 31-125. ras granulosus) da Arnazonia Brasileira. Acta Amazonica, I J TRAVASSOS (L.), 1966.- Serrasentis sagittifer (Linton, 1889) (3): 445-448. (Acanthocephala). Memorias do Instituto Oswaldo Cruz, 64 (I): THATCHER (V.E.) & NICKOL (B.B.), 1972.- Some acanthocephalan 1-10. from Panama and Colombia. Proceedings of the Helmiruhologi- TRAVASSOS(L.), ARTIGAS (P.) & PEREIRA (C.), 1928.- Fauna hel- cal Society of Washington, 39: 245-248. mintologica dos peixes de agua doce do Brasil. Arquivos do Ins- TOUMA OFF (C.), 1947.- Sur un acanthocephale nouveau du genre tituto Biologico, Sao Paulo, 1: 5-68. Oncicola (Oncicola malayana n. sp.) rencontre chez une panthe- TRESHCHEV (V.V.), 1966.- [Une nouvelle espece d' Acanthocepha- re noire (Felis melas Desm.) provenant de Malaisie. Revue le de la Mer de Chukotsk]. Problemy Parazitologii, Trudy Ukr. d'Elevage et de Medecine Veterinaire des Pays Tropicaux, Respubl. Nauch. Obshch. Parazit., 5: 112-115. (in Russian) Paris, I: 231-236. TRIPATHI (Y.R.), (1956) 1959.- Studies on parasites of Indian fis- TRAVASSOS (L.), 1913.- Contribuicoes para 0 conhecimento da hes. V. Acanthocephala. Records of the Indian Museum, 54 fauna helmintolojica brazileira. I. Gigantorhynchus aurae n. sp. (1/2): 61-99. Memorias do Instituto Oswaldo Cruz, 5: 253-255. TROCY (P.M.), (1969) 1970.- Contribution a l'etude des helrnin- TRAVASSOS(L.), 1915.- Revisao dos acantocephalos brazileiros. 1. thes d' Afrique, principalement du Tchad. Bulletin du Museum Fam. Gigantorhynchidae Hamann, 1892 ( ota previa). Brazil National d'Histoire Naturelle de Paris, 41 (6): 1487-1511. Medico, Rio de Janeiro, 29 (12): 89. TRO CY (P.M.), 1978.- Nouvelles observations sur les parasites TRAVASSOS(L.), 1915.- Revisao dos acantocephalos brazileiros. 1. des poissons du bassin tchadien. Bulletin de l'Institut Fonda- Fam. Gigantorhynchidae Hamann, 1892 (Nota previa). Brazil mental d'Afrique Noire, 40 (3): 528-552. Medico, Rio de Janeiro, 29 (14): 105. TRONCY (P.M.) & V ASSILIADES (G.), 1973.- Acanthocephales TRAVASSOS (L.), 1915.- Revisao dos acantocephalos brazileiros. 1. parasites des poissons d' Afrique. Bulletin de L'Lnstitut Fonda- Fam Gigantorhynchidae Hamann, 1892 (2" Nota previa). Brazil mental d'Afrique Noire, 35 (3): 522-539. Medico, Rio de Janeiro, 29 (18): 137. TRONCY(P.M.) & V ASSILIADES (G.), 1974.- Acanthocephales TRAVASSOS(L.), 1915.- Revisao dos acantocefalos brasileiros. H. parasites des poissons d' Afrique. Deuxierne note. Bulletin de Familia Echinorhynchidae Hamann, 1892. Brazil Medico, Rio l'Institut Fondamental d'Afrique Noire, 36 (4): 902-910. de Janeiro, 34 (48): 377. TSIMBALYUK (E.M.), 1968.- [Acanthocephales des animaux des TRAVASSOS(L.), 1917.- Gigantorhynchidae brasileiras. Annais do I iles de la Mer de Behring]. In: [Vers parasites des animaux Congreso Medicina Paulista (l916), Sec. Biologia, Il: 181-19l. domestiques et sauvages. Articles d'Helminthologie presentes a TRAVASSOS (L.), 1917.- AIguns helrninthos da collecao do Institu- l'occasion du 40eme anniversaire des activites d'enseignement to Bacteriologico de S. Paulo. Brazil Medico, Rio de Janeiro, 31 et scientifiques du Prof A.A. Sobolev], Dalnevostoschnii Gosu- (12): 99-100. darstevennii Universitet, Vladivostok: 343-347.(in Russian) TRAVASSOS(L.), 1917.- Helminthos da collecao do Museu Paulis- Tso (W.K.), CHEN(C.F.), CHIEN (H.H.), Wu (F.H.) & DENG ta. Brazil Medico, Rio de Janeiro, 31 (15): 121 - I22. (T.B.), 1974.- [Acanthosentis shashiensis n. sp. et traitement TRAVASSOS (L.), 1917.- Contribuicoes para 0 conhecimento da des poissons infestes par ce parasite]. Acta Zoologica Sinica, 20 fauna helmintolojica brazileira. VI. Revisao dos acantocefalos (4): 409-413.(in Chinese) brazileiros. Parte I. Fam. Gigantorhynchidae Hamann, 1892. TUBA GUI (M.A.), 1933.- Notes on Acanthocephala in the Philip- Memorias do Instituto Oswaldo Cruz; 9 (I): 5-62. pines. Philippine Journal of Science, 50 (2): 115-128. TRA VASSOS(L.), 1918.- Pesquizas sobre as Gigantorhynchidae. TUBAGUI (M.A.), 1935.- Additional notes on Philippine Acantho- Revista de la Societa Brasileira de Sciencias, 2: 79-80. cephala. Philippine Journal of Science, 56 (1): 13-17. TRAVASSOS (L.), (1919) 1920.- Urn novo tipo de acantocefalo. TUBANGUI (M.A.) & MASILUNGA (V.), 1937.- Diplosentis am- Revista de la Societa Brasileira de Sciencias, 3: 209-215. phacanthi gen. et sp. nov., an acanthocephalan parasitic in a TRAVASSOS (L.), 1919.- Inforrnacoes sobre 0 materal helrnintolo- marine fish. Philippine Journal of Science, 62 (2): 183-189. gico colJectionado na ilha da Trinidade em 1916. Arquivos do TUBA GUI (M.A.) & MASILUNGA (V.), 1938.- Nephridiorhyn- Museo Nacional, 22: 159-167. ch us palawanensis sp. nov., an acanthocephalan parasite of TRAVASSOS (L.), 1921.- Contribucao para 0 conhecimento dos Manis javanica Desmarest. Philippine Journal of Science, 66 acantocefalos da sub-familia Centrorhynchinae. A Folha Medi- (1): 1-5. ca, Rio de Janeiro, 2 (6): 42-43. TuBANGUI (M.A.) & MASlLUNGAN(V.), 1946.- On two Acanthoce- 202 Y.J. GOLVAN

phala from the Philippines. Journal of Parasitology, 32: 145-155. Academy of Natural Sciences of Philadelphia, 76: 279-334. V ANBENEDEN (PJ.), 1871.- Les poissons des cotes de la Belgique, V AN CLEAVE(HJ.), 1924.- Additional notes on the Acanthocepha- leurs parasites et leurs commensaux. Memoires de l'Academie la from America described by J.E. Kaiser (1893). Centralblatt Royale de Belgique, 38 (4): 1-100. fur Bakteriologie, Parasitenkunde, Infektionskrankheiten und VAN CLEAVE (H.J.), 1913.- The genus Neorhynchus in North Hygiene, 94: 279-334. America. Zoologischer Anzeiger, 43: 177-190. VAN CLEAVE (HJ.), 1925.- Acanthocephala from Japan. Parasito- VAN CLEAVE (HJ.), 1914.- Eorhynchus: a proposed new name for logy, 17 (2): 149-156. Neorhynchus Hamann preoccupied. Journal of Parasitology, 1: V AN CLEAVE (HJ.), 1928.- Acanthocephala from China. I. New 50-51. species and new genera from Chinese fishes. Parasitology, 20 VAN CLEAVE (H.J.), 1915.- Acanthocephala in North American (I): 1-9. amphibia. Journal of Parasitology, I: 175-178. V AN CLEAVE (H.J .), 1928.- Two new genera and species of Acan- VAN CLEAVE (HJ.), 1916.- Filicollis botulus n. sp. with notes on thocephala from fishes of India. Records of the Indian Museum, the characteristics of the genus. Transactions of the American 30(2): 147-149. Microscopical Society, 35 (2): 131-134. V AN CLEAVE (H.J.), 1929.- A new genus and new species of VAN CLEAVE(HJ.), 1916.- A revision of the genus Arhythmorhyn- Acanthocephala from the Antarctic. Annals and Magazine of chus, with descriptions of two new species from North America. Natural History, ser. 10,4: 229-231. Journal of Parasitology, 2: 167-174. VAN CLEAVE (H.J.), 1931.- Acanthocephala from Japan. n. Two VAN CLEAVE (HJ.), 1916.- Acanthocephala of the genera Cen- new species of the genus Acanthocephalus. Annotationes Zoolo- trorhynchus and Mediorhynchus (new genus) from North Ame- gicae Japonenses, 13 (2): 33-37. rican birds. Transactions of the American Microscopical VAN CLEAVE (H.J.), 1931.- Acanthocephala in North American Society, 35 (4): 221-232. amphibia. 11. A new species of the genus Acanthocephalus. Tran- VAN CLEAVE (HJ .), 1918.- Centrorhynchus pinguis n. sp., from sactions of the American Microscopical Society, 50 (I): 46-47. China. Journal of Parasitology, 4: 164-169. V AN CLEAVE (HJ.), 1931.- Heterosentis, a new genus of Acantho- VAN CLEAVE (H.J.), 1918.- Acanthocephala of the sub-family cephala. Zoologischer Anzeiger, 93 (5/6): 144-146. Rhadinorhynchinae from American fishes. Journal of Parasito- VAN CLEAVE (HJ.), 1931.- New Acanthocephala from fishes of logy,5: 17-24. Mississippi and taxonomic reconsideration of forms with unusal VAN CLEAVE (HJ.), 1918.- The Acanthocephala of North Ameri- numbers of cement glands. Transactions of the American can birds. Transactions of the American Microscopical Society, Microscopical Society, 50 (4): 348-362. 37 (1): 19-48. V AN CLEAVE (H.J.), 1934.- Observations on the status of certain VAN CLEAVE (HJ.), 1919.- Preliminary survey of the Acanthoce- genera of Acanthocephala, chiefley from birds (Abstract). Jour- phala from fishes of the Illinois River. Transactions of the Illi- nal of Parasitology, 20 (6): 324. nois State Academy of Science, 12: 151-156. VAN CLEAVE (H.J.), 1936.- Tenuisentis, a new genus of Acantho- V AN CLEAVE (H.J.), 1919.- Acanthocephala from the Illinois cephala and its taxonomic position. Parasitology, 28: 446-451. River, with descriptions of new species and a synopsis of the VAN CLEAVE (HJ.), 1936.- Acanthocephala from amphibians and family Neoechinorhynchidae. Bulletin of the Illinois Natural reptiles of China. Journal of Parasitology, 22: 523-530. History Survey, 13: 225-257. VAN CLEAVE (HJ.), 1937.- Acanthocephala of the genus Coryno- VAN CLEAVE (HJ.), 1919.- Acanthocephala from fishes of Dou- soma from the-Dyer Island, South Africa. viuerhets-Samhailes glas Lake, Michigan. Occasional Papers of the Museum of Zoo- Handlingar, Femte Foljden, ser. B, 5 (2): 1-6. logy, University of Michigan, 72: 1-12. VAN CLEAVE (HJ.), 1937.- Acanthocephala from China. n. Two VAN CLEAVE (H.J.), 1920.- Two new genera of Acanthocephala new species of the genus Acanthocephalus from amphibia. from Venezuelan fishes. Anatomical Records, Abstracts, 17: 334. Parasitology, 29: 395-398. V AN CLEAVE (HJ .), 1920.- Acanthocephala of the Canadian Arc- VAN CLEAVE (H.J.), 1937.- Status of the generic name Profilicollis tic Expedition. Report of the Canadian Arctic Expedition of A. Meyer (Abstract). Journal of Parasitology, 23 (6): 563. J9J3/J918. 9. Annelids, Parasitic worms, Protozooans, etc. V AN CLEAVE (H.J.), 1939.- A new species of the acanthocephalan (Part E.), Southern Party, 1913-1916: 1-11. genus Polymorphus and notes on the status of the name Profili- V AN CLEAVE (HJ.), 1920.- Two new genera and species of acan- collis.Journal of Parasitology, 25: 129-131. thocephalous worms from Venezuelan fishes. Proceedings of VAN CLEAVE (HJ.), 1940.- Some comparisons of Acanthocephala the United States National Museum, 58: 455-466. of marine fishes of the Atlantic and Pacific coasts (Abstract). VAN CLEAVE (HJ.), 1921.- Preliminary survey of the Acanthoce- Journal of Parasitology, 26 (6, Suppl.): 40-41. phala from fishes of the Illinois River. Journal of Parasitology, VAN CLEAVE (HJ.), 1940.- The Acanthocephala collected by the 12: 151-156. Allan Hancock Pacific Expedition, 1934. Allan Hancock Foun- VAN CLEAVE (HJ.), 1921.- Acanthocephala collected by the Swe- dation Publications, University of Southern California Press, dish Expedition to the Juan-Fernandez Islands (1916-1917). The Los Angeles, ser. 1, 2 (15): 501-527. Natural History of Juan-Fernande; and Easter Island (c. V AN CLEAVE (H.J.), 1942.- Reconsideration of Plagiorhynchus Skottsberg edit.), 3: 75-80. formosus and observations on Acanthocephala with atypical VAN CLEAVE(HJ.), 1921.- Acanthocephala from the eel. Transac- lemnisci. Transactions of the American Microscopical Society, tions of the American Microscopical Society, 40 (1): 1-13. 61 (2): 206-210. VAN CLEAVE (HJ.), 1923.- Notes on Acanthocephala from Japan. VAN CLEAVE(HJ.), 1945.- A new species of the acanthocephalan American Society of Zoology, Program (Abstract No. 21), Ana- genus llliosentis (Rhadinorhynchidae). Journal of Parasitology, tomical Research, 24 (6): 371. 31: 57-60. VAN CLEAVE (HJ.), 1923.- Telosentis, a new genus of Acanthoce- V AN CLEAVE (H.J.), 1945.- A new genus of the acanthocephalan phala from Southern Europe. Journal of Parasitology, 9: 174-175. genus Polymorphus from the American coot. Journal of Parasi- V AN CLEAVE (HJ .), 1923.- Acanthocephala from the fishes of tology, 31: 128-130. Oneida Lake, New York. Roosevelt Wild-Life Forest Experi- V AN CLEAVE (H.J.), 1945.- The status of the acanthocephalan mental Station, Roosevelt Wild-Life Bulletin, 2 (1): 73-84. genus Arhythmorhynchus, with particular reference to the vali- VAN CLEAVE (HJ.), 1924.- A critical study of the Acanthocephala dity of A. brevis. Transactions of the American Microscopical described and identified by Joseph Leidy. Proceedings of the Society, 64 (2): 133-137. Nomenclature of the Acanthocephala 203

V A CLEAVE (HJ.), 1945.- The acanthocephalan genus Corynoso- Florida. Journal of Parasitology, 33: 487-490. ma. I. The species found in water birds in North America. Jour- VAN CLEAVE(H.I.) & MUELLER (J.F.), 1932.- Parasites of the nal of Parasitology, 31: 332-340. Oneida Lake fishes. Part I. Descriptions of new genera and spe- V AN CLEAVE(HJ .), 1946.- Remarques sur le genre Moniliformis cies. Roosevelt Wild Life Annals, 3 (I): 71 pp. (Acanthocephales) et particulierernent sur les especes parasites VA CLEAVE (H.I.) & PRATT (E.M.), 1940.- A new species of the des rats. Annales de Parasitologie Humaine et Comparee, 21: genus Centrorhynchus (Acanthocephala) from the barred . 142-147. Journal of Parasitology, 26: 297-299. VA CLEAVE (HJ.), 1947.- The Eoacanthocephala of North Ame- V AN CLEAVE (HJ.) & RAUSCH (R.L.), 1950.- A new species of the rica, including the description of Eocollis arcanus, new genus acanthocephalan genus Arhythmorhynchus from sandpipers of and new species, superficially resembling the genus Pom- Alaska. Journal of Parasitology, 36: 278-283. phorhynchus. Journal of Parasitology, 33: 285-296. VAN CLEAVE (HJ.) & RAUSCH (R.L.), 1951.- The Acanthocephala V A CLEAVE (H .J.), 1947.- The acanthocephalan genus of eider ducks. Proceedings of the Helminthological Society of Mediorhynchus, its history and a review of the species occurring Washington, 18: 81-84. in the United States. Journal of Parasitology, 33: 297-315. V AN CLEAVE (H.J.) & STARRETT (W.c.), 1940.- The Acanthoce- VAN CLEAVE (H.I.), 1947.- Travassosia tumida n. sp., first record phala of wild ducks in Central Illinois, with descriptions of two of the occurrence of this acanthocephalan genus in North Ame- new species. Transactions of the American Microscopical rica. American Midland Naturalist, 38 (2): 427-433. Society, 59 (3): 348-353. VAN CLEAVE (H.I.), 1947.- On the occurrence of the acanthoce- VAN CLEAVE (HJ.) & TIMMONS (H.F.), 1952.- An additional new phalan genus Telosentis in North America. Journal of Parasito- species of the acanthocephalan genus Neoechinorhynchus. Jour- logy, 33: 126-133. nal of Parasitology, 38: 53-56. VAN CLEAVE (H.I.), 1947.- Analysis of distinction between the VAN CLEAVE (HJ.) & TOWNSEND(L.H.), 1936.- On the assignement acanthocephalan genera Filicollis and Polymorphus, with des- of Echinorhynchus dirus to the genus Acaruhocephalus. Procee- cription of a new species of Polymorphus. Transactions of the dings of the Helminthological Society of Washington, 3: 63. American Microscopical Society, 66 (3): 302-313. V AN CLEAVE (HJ.) & WILLlAMS (R.B.), 1951.- Acanthocephala VA CLEAVE (HJ.), 1949.- Pseudoporrorchis teliger, a new spe- from passerine birds in Alaska. Journal of Parasitology, 37: cies of Acanthocephala from Java. Parasitology, 39: 214-217. 151-159. V AN CLEAVE(HJ .), 1949.- The acanthocephalan genus Neoechi- VERMA rs.c.: & DATTA (M.N.), 1928.- Two acanthocephalous norhynchus in the catostomid fishes of orth America, with des- worms from Indian fishes. Proceedings of the 15th Indian Scien- criptions of two new species. Journal of Parasitology, 35: 500-512. ce Congress, Calcutta: 194. V A CLEAVE(HJ .), 1952.- Acanthocephalan nomenclature intro- VERMA(S.C.) & DATTA (M.N.), 1929.- Acanthocephala from duced by Lauro Travassos. Proceedings of the Helminthological orthern India. 1. A new genus Acanthosentis from a Calcutta Society of Washington, 19: 1-8. fish. Annals of Tropical Medicine and Parasitology, Liverpool, VA CLEAVE(HJ.), 1953.- A preliminary analysis of the acantho- 23 (4): 483-494. cephalan genus Corynosoma in mammals of orth America. 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