THE JEPSON GLOBE A Newsletter from the Friends of The Jepson Herbarium

VOLUME 21 NUMBERS 2 & 3, Fall 2011 Beginning in 2012, The Globe will be published twice a year.

Director’s Column: The Seed Plant Tree of Life: Herbaria Data Go Mainstream An Update By Brent D. Mishler By Anna Larsen, Outreach and Education Specialist for the Gymnosperm Tree of The University and Jepson Herbaria Life Project (UC/JEPS) has long been a leader in the nationwide effort to make data from Gymnosperms are an ecologically herbarium specimens available online, and evolutionarily diverse group of an effort that is rapidly gathering steam about a thousand species of plants that here and globally. The increased atten- bear seeds but, unlike flowering plants (angiosperms), they do not produce tion being paid to collections digitiza- flowers or fruits. Gymnosperms thrive tion across the United States and around in some of the world’s most physiologi- the world stems from several related cally stressful environments: alpine crises impacting natural biodiversity. and subalpine zones, swamps, deserts, Habitat loss through development, and boreal forests. To Californians, the pollution of the environment, and temperate conifers like pines and red- human-caused climate change have woods are familiar gymnosperms but all greatly impacted native plants and Wonderful News! these trees represent only a small frac- animals. To conserve biodiversity in The Jepson Manual, Second Edition tion of gymnosperm diversity. There a rapidly changing environment, it is is at the printer and will be published are many other gymnosperm groups. ever more essential to know the precise this winter! It reflects major improve- There are tropical conifers with fleshy makeup of taxonomic groups (includ- ments to plant taxonomy from phyloge- cones that are dispersed by birds and ing internal variation) and their past netic studies and includes treatments of bats. There are about 300 species of and current geographic ranges. Tools newly described or discovered taxa and insect-pollinated cycads, many of which for ready identification, understanding recently introduced plants. Nearly two- are threatened or endangered. Ginkgo variation within and among species, and thirds of the 7,600 species, subspecies, biloba is a familiar street tree, but is also mapping predicted distribution changes and varieties described are illustrated. the lone representative of an ancient lin- are needed more than ever. National ef- Geographic distributions, elevation eage now known in the wild only from forts such as the US Virtual Herbarium ranges, flowering times, nomenclature, plants growing near ancient Buddhist (http://www.usvirtualherbarium.org/), and the status of non-natives and native monasteries in China. The gnetophytes and international efforts such as GBIF taxa of special concern have been updat- are a particularly odd branch of the gymnosperm tree, which comprises (http://www.gbif.org/), depend on local ed throughout. The Second Edition also initiatives to generate and curate high- allows for identification of 240 alien (Continued on page 5.) quality data. taxa that are not fully naturalized but sometimes encountered. A new chapter ALSO IN THIS ISSUE Digitization efforts at UC/JEPS Field Work in Tierra del Fuego began with the SMASCH project that on geologic, climatic, and vegetation history of California is also featured. Graduate Student News had its roots back in the 1980s. Virtually Pre-orders (at a 20% discount) can be Archive News all the California specimen records of made via the UC Press web site. We vascular plants in UC/JEPS were digi- In Memory of Isabelle Tavares thank all of the Friends of the Jepson Visitors From Iraq tized in the 1990’s, supported by an NSF Herbarium for their loyal support dur- Workshop Year In Review (Continued on page 4.) ing this project! 1 Grad Student News Ben Carter, a graduate student in the comprised of species that are morpho- Mishler lab, has been awarded a pres- logically indistinguishable from other tigious Doctoral Dissertation Improve- species (i.e., “cryptic species”). Efforts ment Grant (DDIG) from the National to preserve and protect global diversity Science Foundation. His study, Niche rely on an understanding of how and differentiation among cryptic moss spe- why these cryptic species evolve and cies, is just part of his dissertation work coexist. that focuses on the molecular systemat- Additionally, most bryological ics, taxonomy, biogeography, and ecol- studies are focused on deep clades and ogy of the moss genus Scleropodium. there are only very limited examples Ben’s study will examine a pair of of studies like Ben’s that incorporate sympatric, cryptic moss species in the fine scale phylogenetic and ecological genus Scleropodium that are hypoth- data to understand recent and ongoing esized to occupy different niches along evolution and diversification. a strong ecological gradient (terrestrial- Ben’s interest in mosses is in part aquatic). To test whether habitat par- based on his belief that people’s envi- titioning is taking place, Ben’s work ronmental awareness and connection will include field collections in an area to conservation is in part facilitated by of known sympatry, sequencing of scientific research conducted in their molecular markers, and collection of own backyards. Because mosses are ecological data. common throughout local regional wild Ben Carter on Santa Catalina Island, Learning more about cryptic spe- lands, they can be a powerful tool for collecting Didymodon tophaceous and cies is important because a substantial scientists to engage students and the other mosses. Photo by Linda Farley fraction of the world’s biodiversity is public.

News From the Archives The archives of the University and highlights how using the field books can When we realized this, it explained the Jepson Herbaria preserve and make often help archivists connect seemingly French binoculars in our Jepson ar- accessible information related to the unrelated events. chives—the Chevalier Opticien model Herbaria and the history of botany at In the Jepson Herbarium, there used by French soldiers during World UC Berkeley. The archives uniquely is odd specimen of a single plant, an War I, rumored to resemble those used complement the specimens held in Azolla (Mosquito Fern) collected by at the infamous Battle of the Little the University and Jepson Herbaria Willis Jepson in Barstow, California, on Bighorn. Jepson birded with these in and contain a wealth of materials. January 17, 1916 (Jep. No. 304, which Barstow. The archives are managed by Amy was not to be identified to species until Kasameyer, Archivist, and Richard G. 2006!). We wondered why on earth Beidleman, ecologist, author, and vol- Jepson would collect just one plant in unteer archivist. In this and future Globe the desert in the middle of winter. The issues, we’ll be bringing you stories that answer was found in his field books have been discovered in the collection. (Volumes 31 and 32). One of our most diverse collections That winter, Jepson was on leave are historical field books from over 60 at the Waterman Ranch, hostessed by botanists. The field books are a valuable former Governor Waterman’s daughter, resource to botanists revisiting histori- Abby (which might explain, in part, cal collections or verifying a collection why Volume 2 of his Flora of Califor- record. nia was dedicated to her); and actually There are countless examples of discovered that with no flowers to speak ways that field books have been used of (and hence only a single collection to provide more information about a of a fern during this trip), The Botany specimen. The following is one that Man essentially decided to watch birds!

2 The Importance of Planned Giving

Rimo Bacigalupi Helen-Mar Beard Mary Bowerman Production of the second edition of The Jepson Manual ment revenue is especially critical to the Jepson Herbarium, was made possible in part due to the foresight of Rimo Baci- where it is a major component of the annual budget and can galupi, Helen-Mar Beard, Mary L. Bowerman, Lawrence R. provide funding for projects that cannot be fully supported Heckard, Willis Linn Jepson, and Robert Ornduff, whom, with external funds. with estate gifts, established endowment funds that benefit A planned gift may enable you, too, to make a more the Jepson Herbarium. significant contribution than you thought possible while These visionaries knew the impact of planned gifts; they at the same time allowing you to achieve your financial, can ensure that the work that is important to the donor is con- philanthropic, and estate planning goals. tinued in perpetuity. With their gifts, Bacigalupi, Bowerman, If you would like to discuss these possibilities, please Heckard, Jepson, and Ornduff have all continued to support contact Staci Markos ([email protected] or 510-643- California floristics, a field they were deeply dedicated to. 7008) or the Office of Gift Planning ([email protected] With no support from the State of California, endow- or 510-642-6300).

Larry Heckard Willis Linn Jepson Robert Ornduff Photo credits: Staci Markos (Bowerman), Jepson Herbarium archives (Bacigalupi, Beard, Heckard, Jepson, Ornduff)

We are halfway to our goal of purchasing a full class set of Leica EZ4 zoom microscopes and their carrying cases. We are looking forward to using these amazing new tools in our 2012 workshops. Many, many thanks to our latest donors: Anonymous Karen Markos Frank Ellis Wendy McClure Mary Ann Hannon Tom Schweich Alan Kaplan Yulan Tong Donald Lepley Desi & Karen Zamudio

3 (Director’s Column, cont. from page 1.) consortium whose goal is to digitize learning and developing techniques that grant, yet many specimens remained to bryophyte and lichen collections across I hope to apply next in California when be digitized at other institutions across the nation. This project will digitize we are ready. I will report my results in the state and a portal was needed to al- the label information for about 2.3 a future column. low searching across them all. million specimens across the United The Consortium for California States. Although unrelated, lichens Herbaria (CCH; http://ucjeps.berke- and bryophytes share many biological ley.edu/consortium/about.html) was traits (see my column on “Bryophyte formed in 2003, initially with major Biology” in The Jepson Globe 17(3), support from the California Digital 2007), which make them some of the Library of the system-wide UC Office most sensitive indicators of air quality of the President. The first major NSF and environmental change. The specific support for this regional network did goal of this project is to provide high not come until 2010, but when it came quality data to address how species it was a large, multi-institutional grant: distributions change with regards to nearly $2 million dollars to database major environmental events across time 300,000+ specimens that were not yet and space. New online presentation digitized and to georeference many techniques will allow interested mem- more specimens that had already had bers of the public to get involved and their basic data captured digitally. to learn about biodiversity alongside the professionals. The new data will be The plants targeted in this NSF grant Jonathan Baio and Catherine Wang us- include dominants in California habi- added to the following portals, which ing the light box and camera in room tats (i.e., woody plants and grasses) already provide access to hundreds of 1066. Photo by Andrew Doran and those most imperiled by threats to thousands of specimens: http://symbi- biodiversity, including climate change ota.org/nalichens/ and http://symbiota. (i.e., alpine plants and those listed org/bryophytes/. The Jepson Manual, Second because of conservation concern). The data generated by these digitiz- Edition, is now in press! Georeferencing is locating specimens ing projects, and the portals in which To celebrate the release of the Manual, on a map and putting their coordinates they are made available to scientists we are planning two events: in a database; such data are the most and the public alike, have already 1) A reception and book signing useful for the mapping studies men- been used for important studies, for at the CNPS conference Wednesday, tioned above. About half of the CCH example the famous 2008 Loarie et January 11th, from 7:00 PM – 9:00 PM. specimens are now georeferenced. al. study investigating threats to Cali- Details about the conference can be The next grant received to digitize fornia endemic vascular plants arising found on the CNPS web site. http:// UC/JEPS collections came in April, from different predictive models www.cnps.org/cnps/conservation/ 2011, and extended the effort to a new of climate change (available freely conference/2012/ habitat and new taxonomic groups of online at: http://www.plosone.org/ Advance registration for the re- organisms. This project will image, article/info%3Adoi/10.1371/journal. ception is requested. If you would like database, georeference, and curate over pone.0002502). Future studies of that to attend the reception and/or purchase 100,000 specimens of Pacific Coast type will be much better informed a book, please send your reply to Staci Markos ([email protected]) no seaweeds that are known to grow in by additional data, and new types of later than December 15, 2011. Your re- biodiversity and conservation studies California, Oregon, or Washington, ply will help ensure enough books are wherever in the world they occur. These will now be possible once geographic available for those who want a copy. coverage by digitized collection data is data will provide a major resource to 2) A release party at the Jepson those interested in the effects of climate robust enough. Herbarium, April 14th. While the full change on coastal environments, fisher- I am on sabbatical in Australia details have yet to be developed, we ies, invasive organisms, and water qual- (through December 2011) working are hoping for a day-long event that ity (The Jepson Globe, 21(1), 2011). on phylogenetic biodiversity analyses will include book signing, tours of the We have just received word of yet made possible by the advanced state herbaria, a presentation about the new another large grant, awarded through of herbarium digitization in that coun- Manual, and a silent auction featur- the new Advancing Digitization of try—Australia’s Virtual Herbarium ing duplicate books from the Jepson Biological Collections program at NSF. (http://www.chah.gov.au/avh/) has Library. Details will be sent early in We received funding as part of a large nearly complete coverage. I’m both the New Year. 4 (Tree of Life, cont. from page 1.) DNA from extinct gymnosperms means lion years ago as it was preserved in the angiosperm-like Gne- that DNA alone can’t tell the whole coal balls. Coal balls are rocks from the tum of tropical forests, evolutionary story; this is where plant Carboniferous Period (354-290 million the desert shrub Ephedra morphology and anatomy are critical. years ago) that contain fossil plant ma- (Mormon tea, ma huang), Using morphological features like the terials embedded in a calcium carbonate and Welwitschia, a plant arrangement of leaves on a stem or the matrix. Coal balls are useful because bearing only two gnarled leaves that pattern of leaf venation, comparisons they contain three dimensional plant sprawl on the sands of the Namib Desert. among living and fossil plants can be structures that were infused with miner- Despite the many advances in our made across the gymnosperm lineage. als as the rocks were forming, a process understanding of the evolutionary history Patterns in DNA sequences, morpho- called permineralization. Of all the tech- of the green plants, the relationships logical traits, and anatomical features niques available to paleobotanists, the among the major groups of gymnosperms provide the foundation for inferring so-called “coal ball peel” is one of the and a clear picture of how angiosperms hypotheses about the evolution and easiest and most informative about the are related to these other seed plants have diversification of seed plants. Variation anatomical structures of extinct plants remained elusive. A major challenge is in DNA sequences provides one lens that existed in a known time and place. that as many as eighteen major groups of for looking back through time to see The coal ball peel technique transfers a gymnosperms that have been identified how plants have diversified; variation in thin section of the fossilized plant mate- from the fossil record have no living physical traits provides a second. When rial from the prepared surface of the rock representatives; an estimated 70–80% the two lenses are used together, they to a clear cellulose acetate sheet, which of the seed-plant lineages that evolved provide a much more resolved picture students can examine using a microscope over the last 400 million years are of how—and ultimately why, where, and in the classroom. extinct. With so many missing pieces, the when—seed plants diversified. The Gymnosperm Tree of Life puzzle of seed plant evolution is difficult Over the lifetime of the project, project concluded this year. As a result to reconstruct. Analyses that focus on researchers associated with the Gym- of the project, four years of data col- comparisons of morphological features nosperm Tree of Life project have lection and analysis will be directed have suggested that angiosperms are dedicated time and resources to sharing toward the unanswered questions about most closely related to the gnetophytes information about gymnosperms and the the evolutionary history of seed plants. (the anthophyte hypothesis). In contrast, findings of the project with the public. The major findings were presented at comparison of DNA sequences has In 2010, these efforts culminated in a the International Botanical Congress indicated that gnetophytes are most series of teacher-training workshops in Melbourne, Australia. To learn more closely related to conifers (the gnetifer held around the country. The workshops about the project, please visit the project hypothesis) or even that they evolved were designed to help teachers integrate website: http://www.huh.harvard.edu/ from within the conifer lineage (the local resources, like the gymnosperms in research/mathews-lab/atolHtmlSite/ gnepine hypothesis). Botanists are still their local botanical gardens or school- index.html. actively debating this controversial yards, into their science curricula in issue, as much of the evidence has been order to teach key ecological and evo- obscured with the passage of time. lutionary concepts. Dean Kelch, a UC/ Over the last four years, botanists JEPS research associate and primary from more than a dozen institutions, investigator on the project, co-taught including the University and Jepson one such workshop at the San Francisco Herbaria, have been involved in a col- Botanical Garden with Matt Parks, a laborative effort to understand how the postdoctoral researcher at Oregon State different types of seed plants are related University, and Nathalie Nagalingum, a to each other. To this end, the Gym- postdoctoral researcher at UC Berkeley. nosperm Tree of Life project (part of “Gymnosperms represent more than a the National Science Foundation’s lens through which to examine the past Assembling the Tree of Life Program) evolution of life on earth,” said Kelch. has focused on assembling data sets “They also illustrate the challenges that that capture a comprehensive picture of organisms face as they adapt to changing gymnosperm diversity and include both ecological conditions today.” living and extinct plants. The experts as- In addition to touring the Garden’s sociated with the Gymnosperm Tree of living gymnosperm collections while Life project have collaborated to create Kelch introduced gymnosperm ecology, data sets that include DNA sequences teachers who attended the San Francisco A close-up of Podocarpus macrophyllus from nearly every living species of workshop had a first hand opportunity showing the fruit-like seed cone. gymnosperms. Of course, the lack of to look at plant diversity from 300 mil- Photo by Dean Kelch 5 Field Work in Southernmost Chile

Typical scenery in a seno (fiord), Isla Grande, Tierra del Fuego, Chile Photo by Jim Shevock Excerpted from letters sent by Jim Shevock dominated by the perennial juncaceous The scenery we encountered was Marsippospermum, and countless amazingly beautiful—glaciers were Earlier this year (Jan 5 to Feb 7, shallow marsh-like pools, the footing ever-present and streams cascaded 2011), I was fortunate to be one of the was somewhat unstable; it often down in all directions to the sea. bryologists participating in an NSF rippled beneath my feet as I bounced The vegetation was dominated by funded expedition to the Antártica across it. Upslope were granitic, open bryophytes in both numbers of species Chilena Province. The only way to landscapes but these, too, were slow and biomass. The tree-cover occurred access nearly all of this area is by boat; going because the peaks were steep primarily right along the fiords and there are no roads on the Chile side of and covered by glaciers. The highest bays. Occasionally, trees were seen Isla Grande de Tierra del Fuego. elevation I reached was 125 meters! in small pockets higher up slope but For the duration of the expedition, Despite these difficult collecting the ever-present snow and ice on the we lived on a seaworthy ship and were conditions, I was able to collect 30-40 higher slopes kept the forests restricted hosted by a very experienced crew. specimens per day. Life on the boat was cramped but was I focused on the offset by interesting colleagues, good species occurring food, fresh baked bread each day, and in streams and drinking water collected from melting rivulets, especially glaciers. The hold of the ship, normally those bryophytes containing king crabs, was retrofitted called rheophytes into a bryophyte drying facility. (seasonally We attempted to visit two submerged then collection sites per day but our ability exposed on to do so depended on the sometimes boulders). In total, unforgiving, cold, and wet weather. I collected 621 Each day, we disembarked in a new bryophyte numbers area, going ashore via zodiacs to in multiple sets, explore and collect. Once ashore, hiking and collectively in the forested areas was slow going the team obtained —the bryophytes can be up to waist- Making our way to shore on a zodiac. Kimmie Mighill (gradu- over 3000 numbers. high! While hiking in the ‘rushlands,’ ate student), Bill Buck (NYBG), crew member, & Jim Shevock Photo by Blanka Shaw 6 Map of Tierra del Fuego, courtesy of Wikipedia Commons mostly to the coastline. The dominant large fruits. This species is used in a type tree was Nothofagus betuloides of local beer called ‘calafante,’ a nice and another common tree was the darker colored beer with the juice of the small but handsome Drimys winteri. Berberis as part of the ingredients. One small shrub has spectacular magenta- Pilgerodendron was the only member Close-up of the ship with us getting of the Cupressaceae and in many areas colored tubular-bell-like flowers about drinking water from melting glacier it is only a ground cover. 3–4 inches long (Philesia magellanica, Photo by Jim Shevock In the Nothofagus and Drimys Philesiaceae). There was also a the most adventurous expedition I pockets of forest, the bryophytes spectacular shrub in the Proteaceae, have ever been on. I have never seen carpeted everything. Most of the Embothrium coccineum, with whorls anything that wild and pristine. No one understory shrubs appeared to be of red tubular flowers. Although it is a lives there and from the look of the members of Ericales, quite heath-like timber tree in Chile much farther north, land, it is also very rarely visited. Our and with many shrubs in full flower. it grows here as a small shrub (to 2 collections from the areas we visited The other important and fairly common meters) or prostrate over wind-swept are likely to be among the first made shrub was a species of Berberis with rocks. Herbaceous flowering plants were basically absent. by bryologists. This trip took us to glacier-filled Jim Shevock is a research associate at mountains, far away from shipping the California Academy of Sciences and lanes and civilization. It was by far the University of California Herbaruim

Philesia magellanica Photo by Jim Shevock Mountain scenery with Magellanic grassland. Photo by Jim Shevock 7 Kurdish Delegation Visits UC/JEPS By Barbara Ertter On April 21, 2011, the University and Jepson Herbaria were visited by a botanical delegation from the semi-autonomous Kurdistan Regional Government (KRG) in northeastern Iraq. The delegation is working to establish botanical gardens in Erbil and Suleimaniah (both in northern Iraq), and they will also oversee a new regional herbarium that will be contributing to a new Flora of Iraq project. According to KRG prime minister Barham Salih, “The KRG places a great deal of emphasis on improving the standards of higher education, as well as giving students the chance to go and continue Collecting team trudging up mountain in Iraqi Kurdistan their studies abroad, a right denied Senior Science Consultant for The and Iran. One of the participants in that to them by the dictatorial regime of American University of Iraq–Sulaimani, expedition was Dr. Ihsan Al-Shehbaz of Saddam Hussein”. and former KRG Deputy Minister of the Missouri Botanical Garden, an Iraqi- Members of delegation were Dr. Agriculture & Irrigation. His equally American who was a former colleague of Sarbagh Salih, Dr. Bahram K. Maulood, accomplished wife, Dr. Hamdi, is a Dr. Maulood’s at Salahaddin University and Dr. Bushra Ahmed Hamdi. Dr. zoologist and Chair of the Department in Iraq. This connection led to both Salih, in addition to being a human of Laboratory Analysis at Hawler Al-Shehbaz and Ertter being invited rights activist and wife of KRG prime Medical University. Accompanying to participate in the inaugural floristic minister, is a horticulturalist who has the delegation were Dr. Barbara Ertter survey of a mountain range in the KRG worked at the Agricultural Research (UC/JEPS), Dr. Christopher Davidson, in June 2010. Davidson and Christoph Center in Beltsville, Maryland. Dr. and his wife Sharon Christoph. took advantage of the opportunity as Maulood wears an even greater number The delegation represented the well, furthering their ambitious goal of of hats, including research limnologist, latest spin-off of the American- photographing all plant families in the Iranian Botanical world (floraoftheworld.org). Program (AIBP), The visit by the Kurdish delegation established by Dr. in April 2011 provided an opportunity for Fosiee Tahbaz the American participants to reciprocate (coordinator, the outstanding hospitality they were AIBP) and Barbara given in Iraq, with the added potential Ertter after their of broadening future collaborations. joint trip to Iran in A guided tour of the University and 1999. Among the Jepson Herbaria was only one of many more successful during a 3-week whirlwind tour that accomplishments included botanical gardens, herbaria, of the AIBP has museums, and experiment stations in been an expedition Texas, Arizona, California, Idaho, and to Iran in 2004, Missouri. Time in the herbaria included which included a sumptuous reception hosted by Dr. participants from Tahbaz, which allowed the delegation various institutions to interact with interested faculty, staff, in the United States and students. Bahram Maulood, Barbara Ertter, Brent Mishler, Sarbagh Salih, Fosiee Tahbaz, and Bushra Ahmed Hamdi in the herbarium.

8 2011 Workshop Year In Review

Clockwise, from top left: Removing Cholla spines in transit on San Clemente Island, Little Duck Lake at sunrise, a pack mule carries gear to camp, intensive Asteraceae keying, examining hydric soils for Wetland Delineation, Mimulus pulchellus, and Linda Beidleman in the field. Photos by Jeanne Marie Acceturo

9 In Memory of Dr. Isabelle Tavares The Jepson Herbarium Projects & Resources Laboulbeniales, an order of minute fungi that parasitize The Jepson Flora Project insects. This research con- The Jepson Manual & Jepson Desert Manual tinued after her dissertation Online Interchange for California Floristics A Flora of California (electronic) and eventually resulted in her Educational Services & Resources magnum opus: The Laboul- Botanical Workshops & Courses beniales, published in 1985. Plant Identification She began working in the 2,200,000+ Worldwide Plant Specimens University Herbarium while Botanical Library & Archives still in graduate school, and Publications & Research Projects continued uninterruptedly Constancea: U.C. Publications in Botany until long past her retirement Director: Brent D. Mishler in 1993. She curated the fungi Deep Green Plant Phylogenetics: Novel (including lichens) and the Analytical Methods for Scaling Data from Genomics to Morphology bryophytes, and participated Dr. Isabelle Tavares died on May Moorea Biocode Project (a complete inventory in all day-to-day operations of the of an island ecosystem) 21, 2011. Dr. Tavares had been associ- Herbarium. Curating lichens led her Systematics and ecology of Syntrichia ated with the University Herbarium for to her second major research interest: Curator: Bruce G. Baldwin 59 years. Her academic career began the taxonomy of Usnea, a widespread Systematics and Evolution of Calif. tarweeds at City College of Los Angeles, was and notoriously taxonomically diffi- and relatives (tribe Madieae, Compositae) interrupted by service in the Women’s cult genus of lichens. Dr. Tavares was Unravelling the dynamics of mating-system Army Corps during World War II, extensively involved in the California evolution in tribe Collinsieae and continued at Berkeley after the Botanical Society, especially in editing Curator of Ecology: David Ackerly War. She received B.A., M.A., and its journal Madroño. She was a found- Niche conservatism, functional trait evo- lution, and the diversification of the Cali- Ph.D. degrees from the University of ing member of the California Lichen California. For her doctoral disserta- fornia vernal pool flora Society, and an active promoter of Ecological Flora of California tion, under the guidance of Profes- California lichenology. Curator of Monocots: Chelsea D. Specht sor Lee Bonar, she investigated the Evolution and biogeography of Calif. alliums Systematics and evolution of Heliconia What took so long? Floral developmental evolution in the tropical gingers (Zingiberales) Some of you may have wondered (over 7600 terminal taxa), a large group Curator of W. N. Am. Botany: Barbara Ertter why it took so long to produce the of authors (over 300 from around the Flora of Mount Diablo & Flora of the East Bay second edition of The Jepson Manual. world), a concise format (treatments North American Potentilleae There are many reasons: a huge flora were edited to adhere to conventions), Trustees: Vice Chancellor Emeritus Roderic a small staff (equivalent Park, Chairman; Vice Chancellor Emeritus to about four positions), a Beth Burnside; UC Botanical Garden Direc- dispersed editorial board tor Paul Licht; Professors John Taylor and (members from various Brent D. Mishler (ex officio) regions of California and Administrative Curator: Andrew Doran beyond), and an extensive Jepson Flora Project Staff: proof process. Pictured Project Research Specialist: Scott Simono here is Tom Rosatti, one Project Manager: Staci Markos Manager of Collections Data: Richard Moe of the scientific editors, Scientific Editor: Tom Rosatti with the first and second Research Associate: Bridget Wessa sets of proofs (there were Archivist: Amy Kasameyer five sets in total). Each Collections Management Staff: Kelly tab represents at least one Agnew, Kim Kersh, Ana Penny, change to the manuscript. and Margriet Wetherwax We made hundreds of Public Programs: Jeanne Marie Acceturo changes to the proofs, a Admin. Assistant and Globe design: process that took about Edith Summers six months! Development & Globe Editor: Staci Markos

10 Friends of the Jepson Herbarium Become a Friends Donor $725 Dissecting microscope Name(s) ______$65 Carrying case for microscope Address ______Other $ ______

City, State Zip ______Join the Friends (annual membership) Telephone / email ______$35 Individual ___ $50 Family ___ New Member ___ Renewal My or my spouse’s employer will match this gift. (Please enclose company form.) This gift is in honor of in memory of ______

Please make your check payable to UC Regents, charge your gift (Visa or Mastercard), or give online (http://givetocal.berkeley.edu/browse/?u=71).

___ Visa ___ MasterCard Total Amount $______MAIL TO: The Jepson Herbarium Account # ______1001 VLSB #2465 University of California Exp. Date ______Signature ______Berkeley, CA 94720-2465

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Memorials and Special Gifts The Jepson Herbarium is pleased to offer thanks to those who chose to honor or remember others with gifts to the herbarium.

In memory of Margaret Ashworth In honor of Thomas McCabe Wendy McClure In memory of Sue Hossfeld Thorston Henrich Wilma & William Follette In memory of Milo Baker In honor of Gary Monroe G Douglas Barbe & Ann Evans In memory of Milton B. Irvine Cynthia & John Perrine Ruth Irvine In memory of Mary Bowerman In memory of Robert Ornduff Alfred & Barbara Sattler In memory of Ramona Davis William S. Davis In memory of Esperance John S Mooring In memory of Emily Reid JK Anderson Sheila McMahon In memory of Robert Lloyd In honor of M. Helen Guerber Theodora Lee Gregg In memory of Maurice T. Resnicow Jay Guerber Alan I. Kaplan In honor of Staci Markos In memory of Larry Heckard Karen Markos In honor of Paul Silva Susan Cochrane Levitsky Susan Brawley L. Maynard Moe In honor of Steve Matson and Eileen Burger Cherie Wetzel David Matson & Susan Gerson In memory of Willis E. Simms Ellen Simms In memory of James C. Hickman In memory of Elizabeth McClintock Carole S Hickman Carlyn Halde In memory of G. Ledyard Stebbins Charles E. Blair

11 Friends of the Jepson Herbarium Nonprofit Organization The Jepson Globe, Vol. 21 Nos. 2 & 3 1001 Valley Life Sciences Building #2465 U.S. Postage PAID University of California, Berkeley University of California Berkeley, CA 94720-2465

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Jepson Herbarium Public Programs We’re still hard at work planning our 2012 workshop season, and it’s shaping up to be a great one! The following workshops are confirmed, with more on the way:

Wetland Restoration

Death Valley’s Endemic Flora Astragalus funereus Flora of San Luis Obispo County Seaweeds of Central California Bees and Pollination Ecology of Spring Wildflowers Late-Season Flora of the White Mountains

Introduction to Bryophytes

The full 2012 schedule will be available in late fall. Stay tuned for updates! For more information, contact Jeanne Marie Acceturo 510-643-7008, [email protected] or visit our Web site: http://ucjeps.berkeley.edu/workshops/

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