THE JEPSON GLOBE A Newsletter from the Friends of The Jepson Herbarium

VOLUME 25 NUMBER 1, Spring 2015

Curator’s Column: Revisiting The Jepson Manual: Vascular Origins of Flora of California, Second By Bruce G. Baldwin Edition: Supplement II I was recently invited to author a By Thomas J. Rosatti, Scientific Editor, chapter for the Annual Review of Ecol- Jepson Flora Project ogy, Evolution, and Systematics (2014) When last we met here (The Jepson on “Origins of Diversity in the Globe 23(1):3-5. 2013), I discussed most California Floristic Province.” A vast of reasons why treatments in floras in literature on that topic had accumulated general need to be revised, giving spe- since Raven and Axelrod published cific examples from Supplement I. For their book-length masterpiece “Origins Richard Beidleman: From Supplement II (released in December, and Relationships of the California 2014 and posted online), we focused Flora” in the University of California the Field to the Archives our efforts on getting taxa new to the Publications in in 1978. Re- By Amy Kasameyer, Herbaria Archivist state incorporated, whether as taxa viewing that literature allowed for an Dr. Richard G. “Dick” Beidleman completely new to science, or as taxa updated perspective on evolution and lost his battle with cancer on August new only to California, as either natives assembly of our unusually diverse flora. 7, 2014. Most members of the Friends or naturalized aliens. Three of the general conclusions from knew Dick from the “50 Families in Taxa that are completely new to sci- the paper are discussed below. the Field” workshop he taught with his ence comprise two main kinds: those for (1) Recent studies of floristic diver- wife Linda. More than 200 California which no collections were known, and sity and endemism have reinforced the botanists took their first Jepson Work- those for which collections existed but importance of areas of high topographic shop (or even keyed their first plant) were previously identified as belonging and edaphic (substrate or soil) complex- with Dick and Linda. Many commented to an existing species. The latter is the ity as reservoirs of Californian plant on his sense of humor, but his depth of case for Lagophylla diabolensis B. G. biodiversity. Two studies are especially knowledge and vast experience teach- Baldwin, which was recently described noteworthy as pertinent to understand- ing natural history informed his relaxed from the southern Diablo Range by ing the basis for that pattern. In 2013, and welcoming spirit in the field. Bruce Baldwin, Curator of the Jepson Lancaster and Kay (in Evolution), after Fewer may be aware of Dick’s in- Herbarium (Madroño 60(3):249-254. estimating components of diversifica- valuable contribution to the University 2013). Plants that had been assigned to tion rates of California plants and their and Jepson Herbaria archives, which he L. dichotoma Benth. sensu Keck (1959, closest relatives, concluded that lower called the finest he had ever seen. Dick (Continued on page 4) extinction rates rather than accelerated was a dedicated volunteer at the ar- speciation may explain the high taxo- chives, beginning in the early 1990s. He ALSO IN THIS ISSUE nomic diversity within various Califor- prepared the first detailed inventory of Andrew Doran in Panama nia plant groups. They suggested in part the archives in 1993, which was used as California Naturalists Program that steep environmental gradients, as the foundation of a grant-funded project NSF grants for fungal collections found in montane areas, may have al- to catalog the archives beginning in Consortium at 2 million specimens lowed Californian plants to adjust their 2009. He cataloged and arranged the Chile collaboration ranges in response to climate change collections of notable botanists includ- New Mishler Lab members and thereby avoid extinction. In 2014, ing Willis Linn Jepson, John Gill and Members’ Night (Continued on page 6) (Continued on page 9) Botanists from around the globe convened in Panama to discuss the sustainability of the Global Plants project. Photo by Alfonso Jaén Fotógrafia, Panama. Documenting our most im- tizing type specimens for over 7 years main collection as isotypes (duplicate and are currently digitizing Central types) that are possible replacements portant specimens American type specimens (The Jepson in case the holotypes (principle types) Andrew Doran, Assistant Director Globe 19(1):3. 2008). Type specimens are destroyed or lost which has hap- for Collections, travelled to Panama (types) are the standard reference for pened a number of times at botanical in September 2014 to present the new the identification and naming of new institutions. database of UC/JEPS (CollectionSpace) species. Since the early 1990s, over To search our types online, go to and workflow to attendees of the 7th 300 new species have been discovered our home page and at the top menu Global Plants meeting and show inter- in California; this includes a number of bar, select DATABASES and UC/JEPS ested parties how we are handling the mosses and cacti just last year and all of SPECIMEN PORTAL and restrict your migration of data from type specimens these names have types, many of which search by checking on the box “include to the JStor Global Plants website, are housed in herbaria in California only type specimens”. which links the literature (where they and beyond. These types serve as criti- After our grant ends in June, we were first published) to the specimens. cal reference points for botanists who will still scan and database type speci- This project is part of our 3rd grant from work on the flora of California and the mens on request, but we also will look the Andrew W. Mellon Foundation for California Floristic Province. for other sources to sustain this effort. funding to document type specimens, Currently, we have almost 15,000 We are still exploring other funding some of the most valuable specimens digitized types in our new public por- options and would welcome input from in the Herbaria. tal (The Jepson Globe 24(2):2. 2014) Friends. At UC/JEPS, we have been digi- but still many thousands exist in the Volunteers at the Jepson Herbarium Discovering a diversity of “buried treasures” is just one experience you might have during a volunteer Saturday. Volunteers are needed to mount, sort, and file specimens and to assist with related projects. No previous herbarium ex- perience is necessary! Volunteer Saturday begins at 10 a.m. and finishes by 4 p.m. (participants need not stay the full time). Upcoming dates include April 18th (Cal Day) and May 9th. We also welcome indi- vidual volunteers who can come in during our regular hours (M-F 8-5). We will try to match your unique interests, abilities, and schedule to one of the numerous curatorial projects that are available to work on. For more information about volunteer oppor- tunities, please call Ana Penny (510) 642-2465. Ana Penny, Volunteer Coordinator, David Margo- lies, David Gowen, and Elizabeth Brusati mount- ing specimens during a recent volunteer Saturday. 2 First Ever Graduates of the Jepson Herbarium’s California Naturalist Certification Talk about Their Motivation By Diana Rohini LaVigne science. UC academic credits were inspire interest in their conserva- What do a former environmental available to all students in the class. tion.” Melissa Hong, Alameda, lawyer, belly dancer, high school teach- Graduates of the program had User Experience Designer er, marathoner, choir singer, theater a wide variety of reasons to join the professional, journalist, surveyor, web- course; all had one common thread. “I want to help people appreciate master, wildlife biologist, Peace Corps They all aspire to motivate, impact, and the world in a whole new way.” alumna, filmmaker, microbiologist, and educate others in life sciences, citizen Adrian Cotter, Oakland, Co- an actor have in common? They were science, and their role within nature in curator San Francisco Natural all members of the California Natural- the . It was their History Series ist training program, sponsored by the personal stories that told the real story “I want to do a better job as a Jepson Herbarium. of this diverse and fascinating group of docent at the botanic gardens The Jepson Herbarium at UC people that came together to celebrate, where I volunteer and perhaps Berkeley held its first course for the study, and experience nature’s greatest branch out a bit beyond botany.” new UC California Naturalist training gifts. Barbara Steuart, Berkeley, Vol- that officially granted certification to unteer Docent, UC Berkeley Bo- 20 naturalists on November 6, 2014. “I want to help my students to develop their own environmental tanic Garden, East Bay Regional The UC California Naturalist Program Parks, Tilden Botanic Garden promotes environmental literacy and consciousness.” Amber Lancast- stewardship through discovery and ac- er, San Francisco, High School “I want to create engaging, tion. The Jepson Herbarium’s Califor- Science Teacher, June Jordan fun, and interactive digital mul- nia Naturalist classes met weekly at UC School for Equity timedia that uses the best of Berkeley and hosted several field trips “I want to share an appreciation storytelling, science, and outdoor both during class and on the weekend. of nature with children and visi- exploration to spark a new Aspiring Naturalists applied for tors to Devil’s Slide Trail Park.” generation of nature stewards.” admission into the program that en- Kathy Gesley, Palo Alto, Trail Catherine Lynn Butler, Rich- tailed a 40-hour course that combined Ambassador mond Heights, Chief Storyteller, classroom and field experience in sci- Greenexus LLC ence, problem solving, communication “I want to inspire youth to carry training, and community service to on the task of protecting the “I want to continue with citizen explore the unique ecology and natural earth’s living creatures.” Diana science projects, future bio- history of the Bay Area. In addition to Rohini LaVigne, San Francisco, blitzes, and, perhaps, bring CA the coursework, California Naturalists Chief Communications Officer, Naturalist activities to the school participated in 40 hours of volunteer Life Chiropractic College West where I currently volunteer.” service in one of four areas: program Patricia Denn, Oakland, happily support, interpretation/education, res- “I want to help others discover retired Clinical Microbiologist the value of natural spaces and toration/conservation, and/or citizen “I want to help others to real- ize and embrace our integral relation to the natural environ- ment, inspiring environmental consciousness and stewardship.” Shawna Casebier, San Francisco, California native, Aspiring Life & Soul Alignment Coach

Applications for the 2015 training will be available in May. For more informa- tion, please visit ucjeps.berkeley.edu/ workshops/2015/CalNat/. Participants of the 2014 California Naturalist Training sponsored by the Jepson Herbarium. Photo by Preston Merchant. 3 (Supplement, continued from page 1) A California Flora) from the Diablo years after the Ertter collection, dur- Mark Hershkovitz had presented data Range differ from plants elsewhere in ing the flowering period of the plants, in 2006 (Gayana Bot. 63:13-74) that the range of that species in a number with John L. Strother of the University showed that what had been called Cal­ of vegetative as well as reproductive Herbarium, again revealed no plants. andrinia ciliata (Ruiz & Pav.) DC. in characters, while phylogenetic analy- However, some were found in an area western North America (and TJM2) ses of DNA sequence data from both much closer to where Ertter 17810 is is phylogenetically distinct (although nuclei and chloroplasts have provided mapped in Berkeley Mapper, east- not very morphologically divergent) evidence that L. dichotoma sensu Keck northeast of the first area searched, on from ciliata in Central from the Diablo Range is more closely both sides of an evidently unnamed trail America and South America, where related to L. ramosissima Nutt. than approximately 100 m up-slope from a the type for the name was collected, to L. dichotoma sensu Keck from the point where it departs from Lake Anza and that whereas the North American San Joaquin Valley and Sierran foot- Trail, in numbers exceeding what would plants are closely related to Calandrinia hills. Examination of some of the type be suggested by “occasional,” although breweri S. Watson of California and specimens involved indicate that the in habit they could be described as lax; northern Baja California, plants be- species recognized now by Baldwin that they were not in (or fruit) longing to Cal ­andrinia ciliata from for the Great Central Valley and Sierra during a time they were in flower previ- Central America and South America Nevada foothills should go by the name ously could have been due to climatic are closely related to Calandrinia com- L. dichotoma Benth., and that for the variables. Essentially, we decided to pressa Schrad. ex DC., also of Central plants of the southern Diablo Range a treat these plants as naturalized because America and South America, and that new name (and formal description) was they occur in natural conditions and be- Calandrinia menziesii (Hook.) Torr. & needed; Baldwin selected L. diabolen- cause they not only have survived there A. Gray is available and should be used sis B. G. Baldwin to convey the idea that for at least 13 years but evidently have for the North American plants. the new species occurs in (and appar- increased their numbers there as well. A summary of all of the changes ently is restricted to) the Diablo Range. In addition to getting taxa new that comprise Supplement II follows. Regarding one of the two categories to the state incorporated, one of the Comparing it to the growing list of of taxa that are new only to California, revisions involved the moving of taxa revisions yet to be developed shows the naturalized aliens, there was a note from one to another (from Chei- rapid pace of basic botanical research under the genus in TJM2 (p. 160) that lanthes to Myriopteris), in response that underlies all floristics. Support rigidum Vent. is possibly to new understanding about their from the Friends, which has helped get naturalized in Tilden Park (SnFrB), relationships, with of course the at- us this far and is very much appreci- which is not far from where I sit as I tendant name changes. In some cases, ated, has never been more important to write this. A little research revealed names were changed for various other furthering our efforts. that Barbara Ertter, whose collection reasons. For example, Hydrophyllum no. 17810 (UC1789196, collected 29 capitatum Douglas ex Benth. var. al- October 2001) is the probable source pinum S. Watson is treated in Supple- of that note, considers the taxon to be ment II as a distinct species, which at “at least marginally naturalized” there. that taxonomic rank must be called Indeed, statements on the collection Hydrophyllum alpestre A. Nelson & P. label of Ertter 17810, including “under- B. Kenn. The change in epithet from story of mixed forest” and “Occasional “alpinum” to “alpestre” was required lax shrub,” strongly suggest that, but in this case because at the rank of spe- no such plants were seen by me in a cies, Hydrophyllum alpestre A. Nelson 2-hour search of the area (“along the & P. B. Kenn., published in 1908, has Sibley Trail between Lake Anza and priority over Hydrophyllum alpinum (S. the spot where the trail climbs up into Watson) Greene ex Brand, published in the Eucalyptus near the one-way road 1913, even though the latter was based below the Brazilian Room” according to on Hydrophyllum capitatum Douglas an e-mail of 23 July 2014 from David B. ex Benth. var. alpinum S. Watson, Wake, who first reported the occurrence published in 1871, because, according of the plant in Tilden to Ertter) on 23 to the International Code of Botanical July 2014; in fact, the search revealed no Nomenclature (Melbourne Code), Art. plants whatsoever with opposite simple 11.2, “A name has no priority outside Type specimen of Lagophylla diabolen- ! A search of the same area on the rank in which it is published”. In sis, a new species from the southern 29 October 2014, just about exactly 13 another case involving a name change, Diablo Range, California.

4 Summary of changes made in Revision 2 of the Jepson eFlora, December 2014 PTERIDACEAE FAGACEAE Cheilanthes cooperae D. C. Eaton changed to Myriop- Quercus robur L. added, as waif teris cooperae (D. C. Eaton) Grusz & Windham, as native (Ruiz & Pav.) DC. replaced by Cheilanthes covillei Maxon changed to Myriopteris Calandrinia menziesii (Hook.) Torr. & A. Gray, covillei (Maxon) Á. Löve & D. Löve, as native added, as native Cheilanthes feei T. Moore changed to Myriopteris parryi A. Gray var. arizonicum J. T. gracilis Fée, as native Howell changed to Calyptridium arizonicum (J. T. Cheilanthes gracillima D. C. Eaton changed to Myriop- Howell) M. G. Simpson, M. Silveira, & Guilliams, teris gracillima (D. C. Eaton) J. Sm., as native as native Cheilanthes intertexta (Maxon) Maxon changed to MYRSINACEAE Myriopteris intertexta (Maxon) Grusz & Windham, L. changed to arvensis as native (L.) U. Manns & Anderb., as naturalized Cheilanthes newberryi (D. C. Eaton) Domin changed Anagallis minima (L.) E. H. L. Krause changed to to Myriopteris newberryi (D. C. Eaton) Grusz & (L.) U. Manns & Anderb., as Windham, as native native Cheilanthes parryi (D. C. Eaton) Domin changed to Anagallis monelli L. changed to Lysimachia monelli Myriopteris parryi (D. C. Eaton) Grusz & Windham, (L.) U. Manns & Anderb., as waif as native Glaux maritima L. changed to Lysimachia maritima Cheilanthes viscida Davenp. changed to Myriopteris (L.) Galasso, Banfi, & Soldano, as native viscida (Davenp.) Grusz & Windham, as native Trientalis europaea L. changed to Lysimachia euro- Cheilanthes wootonii Maxon changed to Myriopteris paea (L.) U. Manns & Anderb., as native wootonii (Maxon) Grusz & Windham, as native Trientalis latifolia Hook. changed to Lysimachia lati- folia (Hook.) Cholewa, as native L. added, as naturalized, replacing Viburnum edule (Michx.) Raf. eddyense Stubbs newly described, Viburnum rigidum Vent. added, as naturalized added, as native Polemonium occidentale Greene subsp. occidentale Dittrichia viscosa (L.) Greuter added, as naturalized no longer recognized, treated as synonym of Pol- Lagophylla diabolensis B. G. Baldwin added, as native emonium occidentale Greene Pallenis maritima (L.) Greuter added, as naturalized Polemonium pulcherrimum Hook. var. shastense Volutaria canariensis Wagenitz added, as naturalized (Eastw.) Stubbs recognized, added, as native BORAGINACEAE POLYGONACEAE Hydrophyllum capitatum Douglas ex Benth. var. alpi- Chorizanthe minutiflora R. Morgan, Styer, & Reveal num S. Watson changed to Hydrophyllum alpestre A. newly described, added, as native Nelson & P. B. Kenn. ROSACEAE Wigandia urens (Ruiz & Pav.) Kunth added, as natural- Rosa woodsii Lindl. var. glabrata (Parish) D. Cole ized added, as native, and as belonging to Rosa woodsii CACTACEAE Lindl. subsp. gratissima (Greene) W. H. Lewis & Cylindropuntia acanthocarpa (Engelm. & J. M. Big- Ertter elow) F. M. Knuth var. acanthocarpa becomes the Rosa woodsii Lindl. var. gratissima (Greene) D. Cole accepted name for Cylindropuntia acanthocarpa added, as native, and as belonging to Rosa woodsii (Engelm. & J. M. Bigelow) F. M. Knuth var. colo- Lindl. subsp. gratissima (Greene) W. H. Lewis & radensis (L. D. Benson) Pinkava, with the latter as a Ertter synonym CYPERACEAE Cylindropuntia chuckwallensis M. A. Baker & M. A. Calliscirpus brachythrix C. N. Gilmour, J. R. Starr, & Cloud-Hughes newly described, added, as native Naczi newly described, added, as native Cylindropuntia ×fosbergii (C. B. Wolf) Rebman, M. Eriophorum crinigerum (A. Gray) Beetle changed to A. changed to Cylindropuntia fosbergii (C. B. Wolf) Calliscirpus criniger (A. Gray) C.N. Gilmour, J.R. Rebman, M. A. Baker & Pinkava Starr, & Naczi, as native ERICACEAE POACEAE Pyrola aphylla Sm. added, as native Agrostis lacuna-vernalis P. M. Peterson & Soreng Pyrola crypta Jolles newly described, added, as native newly described, added, as native Pyrola dentata Sm. added, as native 5 Two new NSF grants for fungal value. Funding from a new grant will ~2.3 million microfungi specimens will collections at UC primarily be used to provide safe stor- be compiled and searchable through By Brent Mishler age (compactors and cases) in our main the MyCoPortal, which also holds the herbarium space for fungal collections results of the complementary Macro- The fungi are critical components that are now housed inadequately in a fungi Collections Consortium (MaCC) of all terrestrial ecosystems, where condemned building. project. The combination of MiCC and they play important roles as mutual- A second grant for mycology is MaCC will result in the digitization of ists, parasites, and commensals in similar to several others held by the nearly all existing specimen documen- both plants and animals. Most of the University and Jepson Herbaria dealing tation (~4 million records) for North world’s important plant pathogens, such with macroalgae, macrofungi, lichens, American fungi held by US institutions. as chestnut blight, Dutch elm disease, and bryophytes. The new Microfungi oak wilt, and white pine blister rust are Collections Consortium (MiCC) is a fungi. In addition, fungi are the primary nation wide collaboration that aims to decomposers responsible for recycling fill a critical gap in the national digiti- organic matter such as fallen leaves and zation effort through the digitization downed trees. In this latter role, they of over 1.2 million North American directly impact the global carbon cycle specimens of microfungi. The Consor- even on a geological time scale. tium is a collaborative effort among 38 The University Herbarium (UC) institutions in 31 states that includes has been expanding its mycological col- a variety of small to large university lections to meet the demands for better herbaria, natural history museums, understanding of fungi in California and botanical gardens. The MiCC and across North America. The cur- project will also incorporate existing Specimen image of Scorias spongiosa, rent fungal collection housed at UC has digitized microfungal specimen data, which feeds on honeydew excreted by roughly 215,000 specimens, and con- comprising over 1 million databased insects. This specimen was collected tains over 2,000 fungal type collections, records and ~53,100 specimen images from the campus of the University of and many other collections of historical of microfungi. An integrated dataset of Georgia on the twigs of a beech tree.

(Curator’s Column, continued from page 1) with endemic diversity in the CA-FP species diversity for which divergence Anacker and Strauss (in Proceedings descended from North American an- times have been estimated appear to of the Royal Society B: Biological Sci- cestors. Although deep ancestry for have undergone diversification in the ences) concluded from examining the much of our flora can be traced back to CA-FP since mid-Miocene (~15 million geographic and ecological distribu- Eurasian or southern sources, the most years ago), when the transition toward tions of Californian plant species in immediate ancestry of various groups the summer-dry climate that character- light of their phylogenetic relationships long thought to have their closest rela- izes much of the CA-FP was underway. that habitat and soil heterogeneity on tives in the Old World, or that were of Phylogenetic studies also have been a regional scale were most strongly uncertain relationship until recently, is helpful in corroborating more ancient associated with diversification. Ongo- closer to home, within more extensive divergence of iconic paleoendemic ing work in the University and Jepson North America clades. That pattern is conifers, such as bristlecone fir Abies( Herbaria on the phylodiversity and evident, for example, for major clades bracteata) and Brewer spruce (Picea phyloendemism project discussed in within a diversity of tribes of Astera- breweriana), as well as various xeric- the last Jepson Globe is aimed at gain- ceae and Fabaceae and for other clades adapted (or preadapted) evergreen or ing more understanding of the spatial spanning multiple genera of Apiaceae, drought-deciduous woody eudicot distribution of California plants in a Campanulaceae, and Lamiaceae. Those lineages, such as those represented phylogenetic context. findings are in line with results of work by California buckeye (Aesculus cali- (2) The evident importance of in multiple labs indicating that western fornica), chaparral-pea (Pickeringia), regional-scale diversification in the North American plants with disjunct tree-anemone (Carpenteria), and laurel California Floristic Province (CA-FP) is relatives in both Eurasia and eastern sumac (Malosma). mirrored at a higher level by evidence North America are usually most closely If you are interested in reading that North America, and especially related to their eastern North America more, the review chapter can be freely western North America, has been a counterparts and appear to stem from downloaded at the “Read it online” link cradle for more extensive flowering North American ancestors. at the following site: ucjeps.berkeley. plant radiations than earlier believed (3) As for the age of our flora, edu/cgi-bin/searchpub.pl?Baldwin. and that most Californian plant groups most Californian clades with endemic 6 The Consortium of California wealth and availability of plant speci- a truly collaborative effort to provide Herbaria now serving 2 million men data allows researchers to use this California plant biodiversity data to specimen records information for large-scale compara- the world. The CCH database has been tive studies that would not be possible accessed by users in 148 countries and By David Baxter otherwise. has been cited in over 90 scholarly This February, the CCH reached These 2 million records are diverse publications in local, national, and in- a landmark: with the addition of its in many respects, covering wide ranges ternational journals (Google Scholar 33rd participant, the Sweeney Granite in time, geography, and plant diversity. page: ucjeps.berkeley.edu/consortium/ Mountains Desert Research Center The CCH is able to serve a diversity of citations/). (GMDRC), the CCH is now serving specimen records thanks to its diverse To learn more about the CCH and over 2 million plant specimen records, roster of participant herbaria. Large participating institutions, as well as ac- all through a single interface. The herbaria serve specimens collected cess the main search interface, please throughout the state, while smaller visit: ucjeps.berkeley.edu/consortium/. herbaria have stronger local focus and fill in botanical gaps not covered by larger herbaria. Furthermore, herbaria of managed lands such as National Park Service and Bureau of Land Manage- ment have specimens from areas that are otherwise very difficult to collect from. The CCH also has some out-of- state participants such as the Harvard University Herbaria, which contain thousands of California type specimens and other historically important col- lections. The CCH began in 2003 as a proj- ect supported by the California Digital Density map of specimens served by the Library and hosted at UC/JEPS to CCH. Only records with coordinates database botanical specimens held in Locations of California herbaria par- are mapped (over 70% of total). Map University of California herbaria. In the ticipating in the CCH. Map by David courtesy of GBIF, www.gbif.org following 12 years, it has grown into Baxter.

Herbarium Botanists Visit A highlight of the Chile for Collaborative Stud- visit was a field trip that made a transect from ies With Chilean Scientists the Andean alpine down Brent Mishler, Bruce Baldwin, and through the coast ranges, David Ackerly traveled to Chile Nov. and finally to the coastal 11-20, 2014, to begin a research col- prairie. The physiog- laboration on comparative studies of nomy of the vegetation the Mediterranean-climate regions of was surprisingly famil- California and Chile with colleagues at iar to a Californian, un- the University of Chile: Rosita Scher- til one looked close! A son, Paulette Naulin, and Pato Pliscoff. number of amphitropical They also gave a two-day workshop on disjuncts were seen, but phylogenetic approaches to biodiversity many more convergently Brent Mishler and Bruce Baldwin looking at Quillaja studies to a group of enthusiastic stu- similar plants were seen. (the sole genus in Quillajaceae) in mattoral, a shrub- dents from across Chile. land community type characteristic of central Chile. Photo by Rosita Scherson.

7 Welcome New Members of the Mishler Lab Andrew Thornhill National University. His project inves- a taxonomic group such as the plant tigated the pollen morphology of the phylogeny of the Daintree Rainforest. Andrew Thornhill is an Austra- Myrtaceae family using a combination After eighteen months in Cairns, lian who joined the Mishler Lab as a of microscopy and morphological and Andrew and his wife Naomi drove over postdoc at the start of February. He phylogenetic techniques. 3,000 miles to get back to Canberra and grew up in Melbourne in the southern After finishing his PhD, Andrew then Melbourne and Sydney to then fly Australian state of Victoria, where he began work as a postdoc for Joe Miller here to California to join the University attended Monash University for his at the Australian National Herbarium and Jepson Herbaria. Andrew’s work undergraduate degree and for a Masters based in CSIRO, Canberra. He worked is supported by an NSF grant awarded degree project on carnivorous pitcher on building large species level phy- to Brent Mishler, Bruce Baldwin, and plants. He moved to Canberra, ACT, logenies of Australian Acacia and David Ackerly (Jepson Globe 24(1):1- to undertake his PhD at the Australian Eucalyptus as well as 6.) to study phylogenetic diversity and many other projects on phylogenetic endemism in the Cali- smaller groups such as fornia flora. Andrew will construct a Bryophytes, Palms, and phylogeny of the Californian vascular Gymnosperms. After two plant flora and combine it with spatial years Andrew began his data available from the Consortium of second postdoc for Dar- California Herbaria. He has already ren Crayn at the Austra- begun sampling plants from UC/JEPS lian Tropical Herbarium to complete the phylogeny and is look- based out of Cairns in ing forward to getting out into botanic Far North Queensland. gardens in Berkeley and further afield He continued working on to find additional specimens for the making large phylogenies project. If you would like to get in con- with a focus more on spe- tact with Andrew, his email address is Andrew Thornhill at Thala Beach Lodge, Oak Beach, cies within a geographi- [email protected]. (Far North) Queensland. Photo by Naomi Brydon. cal region rather than

Caleb Caswell-Levy at UC Santa Cruz, Caleb became inter- became interested in the implications of ested oak communities, the ecology of poikilohydry and dessication tolerance, Caleb Caswell-Levy is a new invasive species, and natural history of two fundamental aspects of the bryo- graduate student in the Mishler Lab the bryophytes, lichens, and vascular phyte lifestyle that affect their ecology interested in the ecology and evolution plants of California. and biogeography, which he hopes to of California bryophytes. Originally After graduating from UC Santa research further here at Berkeley. from Los Angeles, as an undergraduate Cruz in 2009, Caleb pursued a vari- ety of hobbies includ- ing playing and teaching bluegrass music, hiking, surfing, and skiing, be- fore returning to botany. He worked in Southern California on a tidal salt marsh restoration project, and spent several years as a field botanist with the National Park Service in Point Reyes National Seashore and Pinnacles Syntrichia ruralis, a common desicca- National Park. He then tion-tolerant moss in California. Photo Caleb Caswell-Levy on a backpacking trip in Denali by John Game. NP on the Toklat River. Photo by Madison Allen. 8 (Beidleman, continued from page 1) down bits of information. While re- The Jepson Herbarium Sara Plummer Lemmon, and Kate searching John Gill Lemmon’s time Projects & Resources and T. S. Brandegee. Dick transcribed in Sierra Valley, he contacted the Sier- hundreds if not thousands of letters, raville postmistress who turned out to 2,200,000+ Worldwide Plant Specimens diary entries, and writings from those be the town historian. She sent him a Director: Brent D. Mishler collections. On June 3, 2013, we dedi- large packet of information about the Deep Moss: Reconstructing the early evolu- cated the renovated archives stacks to Lemmons, including a photograph of tion of mosses from comparative genomics him in honor of his years of volunteer the cabin where Lemmon lived with his Moorea Biocode Project (a complete inven- service and his 90th birthday. brother and a photograph she thinks is tory of an island ecosystem) His research in the archives re- Lemmon with his students at the town Systematics and ecology of Syntrichia sulted in more than 20 published works school. Curator: Bruce G. Baldwin including articles such as “Willis Linn I can’t imagine doing this job Systematics and Evolution of Calif. tarweeds Jepson and the 1899 Unalaska Caper,” without Dick and the volumes of back- and relatives (tribe , Compositae), “Rowboat botanizing with Willis Jep- ground material about our archives that Chaenactis (Chaenactidieae, Compositae), and Collinsia (Plantaginaceae). son on the Colorado River, 1912,” and he compiled and readily shared. I felt Curator of Ecology: David Ackerly “John Charles Fremont and his floral so fortunate to not have had to start Ecology and evolution of California flora; forays into Oregon,” and his well- from scratch when I began working Climate change impacts and conservation received book, California’s Frontier with the archives. From my very first strategies Naturalists (UC Press, 2006). I highly day, he was here to show me the ropes Curator of Monocots: Chelsea D. Specht recommend California’s Frontier and fill me in on all the idiosyncrasies Evolution and biogeography of Calif. Naturalists to anyone interested in the of the botanists represented in our monocots (including Allium, Nolina) history of botany in California. As a re- collections. Whenever I’m asked a Systematics and evolution of Heliconiaceae, viewer for Bay Nature magazine wrote: reference question for our archives, Costaceae, and Zingiberaceae “The book treats California’s natural I turn first to the material Dick cre- Floral developmental evolution in the tropi- world with such wonder, curiosity, and ated while he was here, as usually he cal gingers (Zingiberales) respect, it will leave you reluctant to had already answered a similar ques- Curator of W. N. Am. Botany: Barbara return to the real world.” tion, transcribed a pertinent letter, or Ertter, Flora of Mount Diablo and flora of Dick enjoyed answering reference written an article on the topic. With the East Bay, North American Potentilleae questions about the archives, no mat- his notebooks, transcriptions, and Trustees: ter what the topic, and never failed research files, he has left a rich and Vice Chancellor Emeritus Beth Burnside; to be generous with his knowledge enduring legacy to the archives. We UC Botanical Garden Director Paul Licht; and time. Our visitors always enjoyed miss him and fondly remember his Cathy Park; Professors John Taylor and chatting with him about their research, love of research, history, and nature, Brent D. Mishler (ex officio) and many made sure to schedule their and especially how he loved to share Asst. Director for Collections: Andrew return visits at times when he would his enthusiasm with others. Doran, Cultivated plants, UK flora be at the Herbaria so they could see To learn more about Dick’s amaz- Asst. Director for Development & Outreach: him again. ing life and career before he joined the Staci Markos, Jepson eFlora, CCH & He was incredibly detailed in his Herbaria, please read his obituary in Globe editor own research. For example, as he came the Pacific Grove Cedar Street times; a Biodiversity Informatics Manager: David across information in the Jepson or link is available in the “News” section Baxter Lemmon collections regarding specific of ucjeps.berkeley.edu. Collections Staff & Plant Identification: dates, he transferred that information Kim Kersh, Clare Loughran, Ana Penny, to a detailed chronology notebook, al- and Margriet Wetherwax lowing us to see exactly what Jepson Jepson eFlora & Online Interchange was doing on a specific day. He also for California Floristics maintained lists of names he came Scientific Editor: Tom Rosatti across in the archives, and often went Constancea: UC Publications in Botany (online) Archivist and Librarian: Amy Kasameyer, to other archives and historical societ- Botanical Library and Archives ies to learn more about people who Public Programs: Jeanne Marie Acceturo corresponded with Jepson and the Membership, workshop enrollment, and Lemmons. Globe design: Edith Summers He had a knack for connecting The cabin where John Lemmon lived in Staff Research Associate: Bridget Wessa with people who could help him track Sierra Valley. 9 Members’ Night On September 12, 2014, the Jepson Herbarium hosted a Members’ Night. Over 100 guests attended a lively reception and book sale followed by a presentation by Dr. Peter Raven titled, “Plant Conservation: De- veloping a Strategy.” The full audio version is online thanks to a generous Friend who made the recording! You Peter H. Raven with Brent D. Mishler, can find a link to the audio file on the Andrew Doran, Assistant Director for Director, University and Jepson Her- Friends membership page. Collections, leading a tour. Photo by baria. Photo by Ana Penny Ana Penny

Memorials and Special Gifts The Jepson Herbarium is pleased to offer thanks to those who chose to honor or remember others with gifts to the herbarium.

In memory of Dr. L.V. Bardunov In memory of Paul Silva Yelena Kosovich-Anderson In memory of James Hickman Nancy R. Morin Carole S. Hickman In memory of Wallace Beck Marcia H. Wolfe In memory of Robert Soost Kathryn Beck Jean Soost In honor of Job Kuijt In memory of Mary Bowerman Alan I. Kaplan In memory of May Stekel Alfred & Barbara Sattler Peter Stekel In honor of Paul Licht In memory of Brenda Butner Ramona Davis In honor of John L. Strother Jean & David Struthers Dale E. & Marie Johnson In memory of Robert Lloyd In memory of Annetta Carter Theodora Lee Gregg In memory of Chancellor Tien and Lincoln Constance Mo-Mei Chen Claudia R. Lindsay In honor of Staci Markos Betsy Ringrose & Edward Adasiak In honor of Yulan Tong In honor of Walter Cecatto Karen Markos Philip Reed Thorston Henrich In honor of Richard Moe In memory of Bob and Pat West In memory of Lincoln Constance Diana Hickson J. Fraser & Helen Muirhead Dale E. & Marie Johnson In memory of Rod Park In memory of May Stekel In memory of Lewis A. Coveler Paul Licht Peter Stekel Alan I. Kaplan In memory of Marian and Roger Reeve Paul & Diane Reeve In memory of Thomas C. Fuller Special Gifts: Kenneth Fuller In memory of Jean Ann (Seely) Rosatti Over 55 books from In memory of Emma Gunterman and Edward James Rosatti, Jr. Tim Lukaszewksi and Paul Preston Hazel Gordon Thomas Rosatti 30 books from the In memory of Larry Heckard In memory of John Sawyer Milo Baker Chapter Susan Cochrane Levitsky Jane Cole of the California Native Plant Society Cherie Wetzel

10 You can shape the future: Planned giving for a lasting impact Donors who establish endowment funds with gifts from their estate are visionaries who know the value of planned gifts. Endowment funds ensure that the work that is important to them is continued in perpetuity. The endowment funds of the University and Jepson Herbaria have provided support for existing programs and created new ones. Without them, the herbaria would not be what they are today⎯world-renowned research collections at a public university. Making a bequest can be as simple as including a few sentences in your will. “I give $ (or % of the residue of my estate) to the University of California, Berkeley Foundation, a Cali- fornia non-profit public benefit corporation, to be added to The Herbaria Futures Endowment Fund (R14891000) to support the University and Jepson Herbaria at the Berkeley campus of the University of California.” Bequests of any amount are welcomed and appreciated. Gifts over $50,000 may be used to establish a new endowment to support work that is of particular interest to the donor.

Below, we have highlighted two endowment funds that support graduate students. In 2015, representatives from the Her- baria will be talking with our Friends about programs they would like to support with an estate gift. For more information, please contact Staci Markos ([email protected]) or the Office of Planned Giving [email protected]( ; 510-642-6300).

The Robert Ornduff The Lawrence R. Heck- Fellowship of the Uni- ard Fund of the Jepson versity and Jepson Herbarium Herbaria A biosystematist and taxono- Robert Ornduff (1932- mist, Larry Heckard (1923- 2001) was a long-time 1991) was Curator of the Botany and Integrative Jepson Herbarium for over Biology Professor, ex- 20 years. This endowment Lasthenia californica. pert on plant evolution, awards grants and encourag- and supporter of the Herbaria as well as the UC Botanical es research in the systematics Garden. This fellowship supports UC Berkeley students who of vascular plants of Califor- travel to professional meetings to give an oral presentation on Castilleja wightii. nia and their close relatives in his or her study of plant systematics, evolution, or ecology. Photos by Neal Kramer. North America.

Support the Jepson Herbarium Name(s) ______Amount $______Visa __ Mastercard __ Address ______Card # ______City, State Zip ______Signature ______Telephone/ Email ______Exp. Date ______Basic Membership ($35 individual, $50 family) Basic members receive The Jepson Globe and discounts on Weekend Workshops. Sustaining Membership ($200) Receive basic membership benefits plus acknowledgement in the Jepson eFlora. Lifetime Membership ($5,000 total, or pledge a minimum of $250/year) Demonstrate your dedication and commitment to the Jepson Herbarium with a lifetime membership. Gain recog- nition for your support in The Jepson Globe and the Jepson eFlora. Share your ideas with the Director and Curator at special, invitation-only events. My or my spouse’s employer will match this gift. (Please enclose company form.) This gift is ___ in honor of / ___ in memory of ______Please send me information about including the Herbarium in my will. Please make your check payable to the UC Regents, charge your gift, or give online at: give.berkeley.edu/fund/?f=FU0840000 Jepson Herbarium, 1001 Valley Life Sciences Building #2465, University of California, Berkeley, CA 94720-2465

11 The Jepson Globe, Vol. 25 No. 1 Nonprofit Organization University of California, Berkeley U.S. Postage PAID Friends of the Jepson Herbarium 1001 Valley Life Sciences Building # 2465 University of California, Berkeley, CA 94720-2465 Berkeley

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There is still space available in these workshops! Plant Families of the World in Bay Area Public Gardens May 1-3 Fire Ecology in the Central Sierra Nevada September 24-27 Restoration Ecology October 17-18 Evolution and Diversity of Mushrooms December 11-13 For details go to: ucjeps.berkeley.edu/workshops/2015/index.html Photo by Jeanne Marie Acceturo

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