The Role of the IOBC in Modern Viticulture
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Entomofauna Ansfelden/Austria; Download Unter
©Entomofauna Ansfelden/Austria; download unter www.biologiezentrum.at Entomofauna ZEITSCHRIFT FÜR ENTOMOLOGIE Band 28, Heft 28: 377-388 ISSN 0250-4413 Ansfelden, 30. November 2007 Phytophagous Noctuidae (Lepidoptera) of the Western Black Sea Region and their ichneumonid parasitoids Z. OKYAR & M. YURTCAN Abstract Eleven agricultural and silviculturally important species of Noctuidae and their parasitoids were determined in 33 localities from the Western Black Sea region between 2001 and 2004. The ichneumonid biological control agents Enicospilus ramidulus, Barylypa amabilis and Itoplectis alternans were obtained by rearing the host larvae. K e y w o r d s : Lepidoptera, Noctuidae, Hymenoptera, Ichneumonidae, parasitoidism, Western Black Sea Region, Turkey Zusammenfassung 11 land- und forstwirtschaftlich bedeutende Noctuidae-Arten einschließlich ihrer Parasitoide aus 33 Standorten des Gebietes des westlichen Schwarzen Meeres wurden im Zeitraum 2001 bis 2004 studiert. Ichneumonidae der Arten Enicospilus ramidulus, Barylypa amabilis and Itoplectis alternans konnten durch Aufzucht der Wirtslarven festgestellt werden. 377 ©Entomofauna Ansfelden/Austria; download unter www.biologiezentrum.at Introduction The Noctuidae is the largest family of the Lepidoptera. Larvae of some species are par- ticularly harmful to agricultural and silvicultural regions worldwide. Consequently, for years intense efforts have been carried out to control them through chemical, biological, and cultural methods (LIBURD et al. 2000; HOBALLAH et al. 2004; TOPRAK & GÜRKAN 2005). In the field, noctuid control is often carried out by parasitoid wasps (CHO et al. 2006). Ichneumonids are one of the most prevalent parasitoid groups of noctuids but they also parasitize on other many Lepidoptera, Coleoptera, Hymenoptera, Diptera and Araneae (KASPARYAN 1981; FITTON et al. 1987, 1988; GAULD & BOLTON 1988; WAHL 1993; GEORGIEV & KOLAROV 1999). -
Jumping Mechanisms in Dictyopharid Planthoppers (Hemiptera
© 2014. Published by The Company of Biologists Ltd | The Journal of Experimental Biology (2014) 217, 402-413 doi:10.1242/jeb.093476 RESEARCH ARTICLE Jumping mechanisms in dictyopharid planthoppers (Hemiptera, Dicytyopharidae) Malcolm Burrows* ABSTRACT legs in the same plane underneath the body. A catapult-like The jumping performance of four species of hemipterans belonging to mechanism is used in which the trochanteral depressor muscles the family Dictyopharidae, from Europe, South Africa and Australia, contract slowly, energy is stored and is then released suddenly were analysed from high-speed images. The body shape in all was (Burrows, 2006a; Burrows, 2007b; Burrows, 2009). Despite these characterised by an elongated and tapering head that gave a important common features, each group has particular streamlined appearance. The body size ranged from 6 to 9 mm in specialisations of its own that define its jumping abilities. These length and from 6 to 23 mg in mass. The hind legs were 80–90% of include differences in body shape, in the length of the hind legs body length and 30–50% longer than the front legs, except in one and in the anatomy of the coxae. species in which the front legs were particularly large so that all legs Most leafhoppers have hind legs that are two to three times longer were of similar length. Jumping was propelled by rapid and than the other legs and are 90% of the body length (Burrows, simultaneous depression of the trochantera of both hind legs, powered 2007b). By contrast, froghoppers and planthoppers have hind legs by large muscles in the thorax, and was accompanied by extension of that are only 40–50% longer than the other legs and approximately the tibiae. -
Host Plants and Seasonal Presence of Dictyophara Europaea in the Vineyard Agro-Ecosystem
Bulletin of Insectology 61 (1): 199-200, 2008 ISSN 1721-8861 Host plants and seasonal presence of Dictyophara europaea in the vineyard agro-ecosystem Federico LESSIO, Alberto ALMA Di.Va.P.R.A., Entomologia e Zoologia applicate all’Ambiente “C. Vidano”, Facoltà di Agraria, Università di Torino, Italy Abstract Seasonal presence and host plants of Dictyophara europaea (L.), a candidate vector of phytoplasmas to grapevine, were studied in Piedmont during 2006 in different vine growing regions. Sampling consisted in net sweeping on different candidate host plants, and captures of adults with yellow sticky traps placed on grapevine. D. europaea nymphs and adults were collected on many weeds, showing how this planthopper should be considered a poly- phagous species, although Amaranthus retroflexus L. and Urtica dioica L. seem to be its preferred hosts, and may also bear phy- toplasmas. Larvae of Dryinidae were observed on almost 5% of collected individuals. The peak of adult presence was recorded in the middle of August, but few adults were captured on sticky traps placed on grapevine. Molecular analyses will be performed to detect the presence of phytoplasmas in captured individuals; however, given its scarce presence on grapevine, D. europaea does not seem capable to play a major role in the transmission of phytoplasmas to grapevine even if its vector ability were proved. Key words: Dictyophara europaea, vector, sweep net, Amaranthus retroflexus, grapevine. Introduction Holzinger et al. (2003). During 2007, collected nymphs and adults were put into a rearing cage made of plexi- The genus Dictyophara Germar is represented in Italy glas and insect-proof mesh, with a single plant of Ama- with four species: Dictyophara cyrnea Spinola (only in ranthus retroflexus L., to observe feeding behaviour and Sardinia), Dictyophara pannonica (Germar) (doubtful), ovoposition. -
Identification and Ecology of Alternative Insect Vectors Of
View metadata, citation and similar papers at core.ac.uk brought to you by CORE www.nature.com/scientificreportsprovided by AIR Universita degli studi di Milano OPEN Identifcation and ecology of alternative insect vectors of ‘Candidatus Phytoplasma solani’ to grapevine Fabio Quaglino1, Francesco Sanna2, Abdelhameed Moussa 1, Monica Faccincani3, Alessandro Passera1, Paola Casati1, Piero Attilio Bianco1 & Nicola Mori 2* Bois noir, a disease of the grapevine yellows complex, is associated with ‘Candidatus Phytoplasma solani’ and transmitted to grapevines in open felds by the cixiids Hyalesthes obsoletus and Reptalus panzeri. In vine-growing areas where the population density of these vectors is low within the vineyard, the occurrence of bois noir implies the existence of alternative vectors. The aim of this study was to identify alternative vectors through screening of the Auchenorrhyncha community, phytoplasma typing by stamp gene sequence analyses, and transmission trials. During feld activities, conducted in Northern Italy in a vineyard where the bois noir incidence was extremely high, nine potential alternative insect vectors were identifed according to high abundance in the vineyard agro-ecosystem, high infection rate, and harbouring phytoplasma strains characterized by stamp gene sequence variants found also in symptomatic grapevines. Transmission trials coupled with molecular analyses showed that at least eight species (Aphrodes makarovi, Dicranotropis hamata, Dictyophara europaea, Euscelis incisus, Euscelidius variegatus, Laodelphax striatella, Philaenus spumarius, and Psammotettix alienus/confnis) are alternative vectors of ‘Candidatus Phytoplasma solani’ to grapevines. These novel fndings highlight that bois noir epidemiology in vineyard agro-ecosystems is more complex than previously known, opening up new perspectives in the disease management. Bois noir (BN), a disease of the grapevine yellows (GY) complex, causes serious crop losses in wine-making grape varieties in the Euro-Mediterranean area and in other vine-growing countries. -
UNIVERSITÀ DEGLI STUDI DEL MOLISE Department
UNIVERSITÀ DEGLI STUDI DEL MOLISE Department of Agricultural, Environmental and Food Sciences Ph.D. course in: AGRICULTURE TECHNOLOGY AND BIOTECHNOLOGY (CURRICULUM: Sustainable plant production and protection) (CYCLE XXIX) Ph.D. thesis NEW INSIGHTS INTO THE BIOLOGY AND ECOLOGY OF THE INSECT VECTORS OF APPLE PROLIFERATION FOR THE DEVELOPMENT OF SUSTAINABLE CONTROL STRATEGIES Coordinator of the Ph.D. course: Prof. Giuseppe Maiorano Supervisor: Prof. Antonio De Cristofaro Co-Supervisor: Dr. Claudio Ioriatti Ph.D. student: Tiziana Oppedisano Matr: 151603 2015/2016 “Nella vita non c’è nulla da temere, c’è solo da capire.” (M. Curie) Index SUMMARY 5 RIASSUNTO 9 INTRODUCTION 13 Phytoplasmas 13 Taxonomy 13 Morphology 14 Symptomps 15 Transmission and spread 15 Detection 17 Phytoplasma transmission by insect vectors 17 Phytoplasma-vector relationship 18 Homoptera as vectors of phytoplasma 19 ‘Candidatus Phytoplasma mali’ 21 Symptomps 21 Distribution in the tree 22 Host plant 24 Molecular characterization and diagnosis 24 Geographical distribution 25 AP in Italy 25 Transmission of AP 27 Psyllid vectors of ‘Ca. P. mali’ 28 Cacopsylla picta Förster (1848) 29 Cacopsylla melanoneura Förster (1848) 32 Other known vectors 36 Disease control 36 Aims of the research 36 References 37 CHAPTER 1: Apple proliferation in Valsugana: three years of disease and psyllid vectors’ monitoring 49 CHAPTER 2: Evaluation of the current vectoring efficiency of Cacopsylla melanoneura and Cacopsylla picta in Trentino 73 CHAPTER 3: The insect vector Cacopsylla picta vertically -
Tracking Vectors of Bacteria and Phytoplasmas Threatening Europe’S Major Crops (VECTRACROP)
Euphresco Final Report Tracking vectors of bacteria and phytoplasmas threatening Europe’s major crops (VECTRACROP) Topic area Phloem and xylem feeding insect vectors, fruit and field crops, bacteria and phytoplasmas of phytosanitary concern - Topic Description 2015-D-168 Topic title Tracking vectors of bacteria and phytoplasmas threatening Europe’s major crops (VECTRACROP) 1. Administrative Details . Applicant / Coordinator – Partner 1 Organisation Institute for AgriculturaI and Fisheries Research - ILVO Name of contact Kris De Jonghe, Ph.D. Gender: M (incl. Title) Postal address Burg. Van Gansberghelaan 96, B- 9820 Merelbeke, Belgium E-mail [email protected]; [email protected] Phone ++32 9/ 272 24 48 Applicant – Partner 2 Organisation CRA-W Name of contact Thibaut Olivier, Ir Gender: M (incl. Title) Département Sciences du Vivant (CRAW), Unité Biologie des Postal address nuisibles et Biovigilance, Bâtiment Marchal, Rue de Liroux 4, B- 5030 Gembloux, Belgium E-mail [email protected] Phone ++32 81/ 62 03 39 Applicant – Partner 3 Organisation ANSES Name of contact Reynaud Philippe, Ph.D. Gender: M (incl. Title) Anses Laboratoire de la Santé des Végétaux [Plant Health Laboratory] Postal address 755 avenue du campus Agropolis CS 30016 FR-34988 Montferrier-sur-Lez Cedex E-mail [email protected] Phone + 33 (0)4 67 02 25 10 Applicant – Partner 4 Organisation INIAV Name of contact Célia Mateus- Researcher, Ph.D.; Esmeraldina Gender F (incl. Title) Sousa- Researcher, Ph.D. : Av. da República, Quinta do Marquês Postal address 2780-157 Oeiras – Portugal E-mail [email protected]; [email protected] Phone (+351) 214 403 500 Applicant – Partner 5 Organisation INRA-MOROCCO Name of contact Afechtal Mohamed, Ph.D.; Bouharroud Rachid, Gender: M (incl. -
The Leafhoppers of Minnesota
Technical Bulletin 155 June 1942 The Leafhoppers of Minnesota Homoptera: Cicadellidae JOHN T. MEDLER Division of Entomology and Economic Zoology University of Minnesota Agricultural Experiment Station The Leafhoppers of Minnesota Homoptera: Cicadellidae JOHN T. MEDLER Division of Entomology and Economic Zoology University of Minnesota Agricultural Experiment Station Accepted for publication June 19, 1942 CONTENTS Page Introduction 3 Acknowledgments 3 Sources of material 4 Systematic treatment 4 Eurymelinae 6 Macropsinae 12 Agalliinae 22 Bythoscopinae 25 Penthimiinae 26 Gyponinae 26 Ledrinae 31 Amblycephalinae 31 Evacanthinae 37 Aphrodinae 38 Dorydiinae 40 Jassinae 43 Athysaninae 43 Balcluthinae 120 Cicadellinae 122 Literature cited 163 Plates 171 Index of plant names 190 Index of leafhopper names 190 2M-6-42 The Leafhoppers of Minnesota John T. Medler INTRODUCTION HIS bulletin attempts to present as accurate and complete a T guide to the leafhoppers of Minnesota as possible within the limits of the material available for study. It is realized that cer- tain groups could not be treated completely because of the lack of available material. Nevertheless, it is hoped that in its present form this treatise will serve as a convenient and useful manual for the systematic and economic worker concerned with the forms of the upper Mississippi Valley. In all cases a reference to the original description of the species and genus is given. Keys are included for the separation of species, genera, and supergeneric groups. In addition to the keys a brief diagnostic description of the important characters of each species is given. Extended descriptions or long lists of references have been omitted since citations to this literature are available from other sources if ac- tually needed (Van Duzee, 1917). -
Turkish Journal of Entomology)
Türkiye Entomoloji Dergisi (Turkish Journal of Entomology) Cilt (Vol.) 42 Sayı (No.) 3 Eylül (September) 2018 İnceleme ve Değerlendirmede Bilimsel Olarak Katkıda Bulunanlar (Scientific Advisory Board) ADAMSKI, Zbigniew, Poland İLKER, Ercan, Bursa ATAKAN, Ekrem, Adana KAÇAR, Gülay, Bolu AY, Recep, Isparta KARACA, İsmail, Isparta BAŞPINAR, Hüseyin, Aydın KAZAK, Cengiz, Adana BAYARTOGTOKH, Badamdorj, Mongolia KIVAN, Müjgan, Tekirdağ BOLU, Halil, Diyarbakır KOÇAK, Erhan, Isparta BORDERA, Santiago, Spain KOVANCI, Orkun Barış, Bursa CANHİLAL, Ramazan, Kayseri KUMRAL, Nabi Alper, Bursa CİVELEK, Hasan Sungur, Muğla KÜTÜK, Murat, Gaziantep ÇALMAŞUR, Önder, Erzurum MENNAN, Sevilhan, Samsun ÇETİN ERDOĞAN, Özlem, Edirne MIKUSKA, Alma, Croatia ÇETİN, Hüseyin, Antalya MILONAS, Panagiotis, Greece ÇETİNTAŞ, Ramazan, Kahramanmaraş MUMCUOĞLU, Kostas, İsrail ÇIKMAN, Emine, Hatay NAVARRO, Shlomo, Israel ÇİFTÇİ, Derya, Ankara ÖZDEMİR, Senem, Ankara ÇOBANOĞLU, Sultan, Ankara SAĞLAM, Özgür, Tekirdağ DAĞLI, Fatih, Antalya ŞENAL, Derya, Bilecik DAUTBASIC, Mirza, Bosnia-Herzegovina TEZCAN, Serdar, İzmir DURMUŞOĞLU, Enver, İzmir TİRYAKİ, Osman, Kayseri DONG, Shuang-Ling, China TOLUK, Ayşe, Kayseri EMEKÇİ, Mevlüt, Ankara TOMANOVIC, Zeljko, Serbia ERLER, Fedai, Antalya TOPRAK, Umut, Ankara FERİZLİ, Ahmet Güray, Ankara TREMATERRA, Pasquale, Italy GREGO, Nicola, Italy YOLDAŞ, Zeynep, İzmir GÜNCAN, Ali, Ordu YORULMAZ SALMAN, Sibel, Isparta GÜZ, Nurper, Ankara YURTCAN, Murat, Edirne HASSAN, Erol, Australia YURTSEVER, Selçuk, Edirne İçindekiler (Contents) Orijinal araştırmalar -
The Leafhoppers, Or Cicadellidae, of Illinois (Eurymelinae-Balcluthinae)
BULLETIN of the ILLINOIS NATURAL HISTORY SURVEY HARLOW B. MILLS, Chief The Leafhoppers, or Cicadellidae, of Illinois (Eurymelinae-Balcluthinae) D. M. DELONG PriDted by Authority of the STATE OF ILLINOIS DWIGHT H. GREEN, Govtrnor DEPARTMENT OF REGISTRATION AND EDUCATION FRANK G. THOMPSON, Dirtctor STATE t) F I 1. I, I N O I S DwiGiiT H. CiREES', Governor PEPARTMENT OF REGISTRATION ANi:) EDUCATION Frank G. Thompson, Director \^ ^- \' N A T U R A L HISTORY S U R E I ) I 1 S I O N Hari.o\\ B. Mii.i.s, (-liicf \ olumc 24 BULI^K TIN Article 2 The Leafhoppers, or Cicadellidae, of Illinois (Eurymelinae— Balcluthinae) ]). M. 1)1 f,c)Nc; Priulid hy Jul/iority of the Stall- of Illinois URBANA, ILLINOIS June 194S STATE OF ILLINOIS DwiGHT H. Green, Governor DEPARTMENT OF REGISTRATION AND EDUCATION Frank G. Thompson, Director BOARD OF NATURAL RESOURCES AND CONSERVATION Frank G. Thompson, Chairman A. E. Emerson, Ph.D., Rio/oxv George D. Stoddard, Ph.D., Litt.D., L.H.D., L. H Tiffany, Ph.D., Forestry LL.D., President of the Ihiivcrsily nj Illinois l' R. Howson, B.S.C.E., C.E., Walter H. Newhoi'isk, Ph.D., Geology Engineering Roger Adams, Ph.D., D.Sc, Chemistry NATURAL HISTORY SURVEY DIVISION Urbana, Illinois Scientific and Technical Staff H.^Ri.ow B. Mills, Ph.D., Chief Bessie B. Henderson, M.S., Assistant to the Chief Section of Economic Entomology Section of Forestry Entomologist George C. Decker, Ph.D., WiLLET N. Wandell, M.F., Forester and and Head Head M.S., Entomologist J. -
ZGRUPOWANIA PIEWIKÓW (HEMIPTERA: FULGOROMORPHA ET CICADOMORPHA) WYBRANYCH ZBIOROWISK ROŚLINNYCH BABIOGÓRSKIEGO PARKU NARODOWEGO Monografia
ZGRUPOWANIA PIEWIKÓW (HEMIPTERA: FULGOROMORPHA ET CICADOMORPHA) WYBRANYCH ZBIOROWISK ROŚLINNYCH BABIOGÓRSKIEGO PARKU NARODOWEGO Monografia LEAFHOPPER COMMUNITIES (HEMIPTERA: FULGOROMORPHA ET CICADOMORPHA) SELECTED PLANT COMMUNITIES OF THE BABIA GÓRA NATIONAL PARK The Monograph ROCZNIK MUZEUM GÓRNOŚLĄSKIEGO W BYTOMIU PRZYRODA NR 21 SEBASTIAN PILARCZYK, MARCIN WALCZAK, JOANNA TRELA, JACEK GORCZYCA ZGRUPOWANIA PIEWIKÓW (HEMIPTERA: FULGOROMORPHA ET CICADOMORPHA) WYBRANYCH ZBIOROWISK ROŚLINNYCH BABIOGÓRSKIEGO PARKU NARODOWEGO Monografia Bytom 2014 ANNALS OF THE UPPER SILESIAN MUSEUM IN BYTOM NATURAL HISTORY NO. 21 SEBASTIAN PILARCZYK, MARCIN WALCZAK, JOANNA TRELA, JACEK GORCZYCA LEAFHOPPER COMMUNITIES (HEMIPTERA: FULGOROMORPHA ET CICADOMORPHA) SELECTED PLANT COMMUNITIES OF THE BABIA GÓRA NATIONAL PARK The Monograph Bytom 2014 Published by the Upper Silesian Museum in Bytom Upper Silesian Museum in Bytom Plac Jana III Sobieskiego 2 41–902 Bytom, Poland tel./fax +48 32 281 34 01 Editorial Board of Natural History Series: Jacek Betleja, Piotr Cempulik, Roland Dobosz (Head Editor), Katarzyna Kobiela (Layout), Adam Larysz (Layout), Jacek Szwedo, Dagmara Żyła (Layout) International Advisory Board: Levente Ábrahám (Somogy County Museum, Kaposvar, Hungary) Horst Aspöck (University of Vienna, Austria) Dariusz Iwan (Museum and Institute of Zoology PAS, Warszawa, Poland) John Oswald (Texas A&M University, USA) Alexi Popov (National Museum of Natural History, Sofia, Bulgaria) Ryszard Szadziewski (University of Gdańsk, Gdynia, Poland) Marek Wanat (Museum -
Ichneumonid Wasps (Hymenoptera, Ichneumonidae) in the to Scale Caterpillar (Lepidoptera) [1]
Central JSM Anatomy & Physiology Bringing Excellence in Open Access Research Article *Corresponding author Bui Tuan Viet, Institute of Ecology an Biological Resources, Vietnam Acedemy of Science and Ichneumonid Wasps Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam, Email: (Hymenoptera, Ichneumonidae) Submitted: 11 November 2016 Accepted: 21 February 2017 Published: 23 February 2017 Parasitizee a Pupae of the Rice Copyright © 2017 Viet Insect Pests (Lepidoptera) in OPEN ACCESS Keywords the Hanoi Area • Hymenoptera • Ichneumonidae Bui Tuan Viet* • Lepidoptera Vietnam Academy of Science and Technology, Vietnam Abstract During the years 1980-1989,The surveys of pupa of the rice insect pests (Lepidoptera) in the rice field crops from the Hanoi area identified showed that 12 species of the rice insect pests, which were separated into three different groups: I- Group (Stem bore) including Scirpophaga incertulas, Chilo suppressalis, Sesamia inferens; II-Group (Leaf-folder) including Parnara guttata, Parnara mathias, Cnaphalocrocis medinalis, Brachmia sp, Naranga aenescens; III-Group (Bite ears) including Mythimna separata, Mythimna loryei, Mythimna venalba, Spodoptera litura . From these organisms, which 15 of parasitoid species were found, those species belonging to 5 families in of the order Hymenoptera (Ichneumonidae, Chalcididae, Eulophidae, Elasmidae, Pteromalidae). Nine of these, in which there were 9 of were ichneumonid wasp species: Xanthopimpla flavolineata, Goryphus basilaris, Xanthopimpla punctata, Itoplectis naranyae, Coccygomimus nipponicus, Coccygomimus aethiops, Phaeogenes sp., Atanyjoppa akonis, Triptognatus sp. We discuss the general biology, habitat preferences, and host association of the knowledge of three of these parasitoids, (Xanthopimpla flavolineata, Phaeogenes sp., and Goryphus basilaris). Including general biology, habitat preferences and host association were indicated and discussed. -
ARTHROPODA Subphylum Hexapoda Protura, Springtails, Diplura, and Insects
NINE Phylum ARTHROPODA SUBPHYLUM HEXAPODA Protura, springtails, Diplura, and insects ROD P. MACFARLANE, PETER A. MADDISON, IAN G. ANDREW, JOCELYN A. BERRY, PETER M. JOHNS, ROBERT J. B. HOARE, MARIE-CLAUDE LARIVIÈRE, PENELOPE GREENSLADE, ROSA C. HENDERSON, COURTenaY N. SMITHERS, RicarDO L. PALMA, JOHN B. WARD, ROBERT L. C. PILGRIM, DaVID R. TOWNS, IAN McLELLAN, DAVID A. J. TEULON, TERRY R. HITCHINGS, VICTOR F. EASTOP, NICHOLAS A. MARTIN, MURRAY J. FLETCHER, MARLON A. W. STUFKENS, PAMELA J. DALE, Daniel BURCKHARDT, THOMAS R. BUCKLEY, STEVEN A. TREWICK defining feature of the Hexapoda, as the name suggests, is six legs. Also, the body comprises a head, thorax, and abdomen. The number A of abdominal segments varies, however; there are only six in the Collembola (springtails), 9–12 in the Protura, and 10 in the Diplura, whereas in all other hexapods there are strictly 11. Insects are now regarded as comprising only those hexapods with 11 abdominal segments. Whereas crustaceans are the dominant group of arthropods in the sea, hexapods prevail on land, in numbers and biomass. Altogether, the Hexapoda constitutes the most diverse group of animals – the estimated number of described species worldwide is just over 900,000, with the beetles (order Coleoptera) comprising more than a third of these. Today, the Hexapoda is considered to contain four classes – the Insecta, and the Protura, Collembola, and Diplura. The latter three classes were formerly allied with the insect orders Archaeognatha (jumping bristletails) and Thysanura (silverfish) as the insect subclass Apterygota (‘wingless’). The Apterygota is now regarded as an artificial assemblage (Bitsch & Bitsch 2000).