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U.S. National Vegetation Classification: Advancing The
U.S. National Vegetation Classification: Advancing the Description and Management of the Nation’s Ecosystems Use of the NVC hierarchy to scale the GAP/LANDFIRE National Ecosystems Map Legend Don Long (U.S. Forest Service), Anne Davidson (GAP, BSU) Todd Earnhardt (GAP , NSCU) Alexa McKerrow (U.S. Geological Survey) . Background Methods Results A national inventory of the existing vegetation across the There are 551 natural vegetation classes represented in the 6 Classes 13 Subclasses 22 Formations U.S. has been central to the missions of both the GAP/LANDFIRE National Terrestrial Ecosystems Map for the a Landscape Fire and Resource Management Planning Tools conterminous U.S. The crosswalk allows for the Project (LANDFIRE) and the National Gap Analysis Program aggregation of the mapped classes into the hierarchical b (GAP). Over the past several years these two programs structure of the USNVC; specifically, the ecological systems a. Forest & Woodland a. Temperate & Boreal Forest & Woodland a. Temperate Flooded & Swamp Forest b. Desert & Semi-Desert b. Warm Desert & Semi-Desert Woodland, b. Warm Desert & Semi-Desert Scrub & have come together to collaborate on the next generation are crosswalked to the middle and upper levels of the Scrub & Grassland Grassland highly detailed existing vegetation maps for the U.S. This USNVC. Relationships between the two classification 51 Divisions 112 Macrogroups 240 Groups collaboration leverages the mapping and inventory to systems developed by NatureServe ecologists were used to meet needs for both fire and fuels management, as well link the mapped Ecological Systems to the Group level of as for wildlife habitat conservation planning. the USNVC. -
Phylogenetic Analyses of Juniperus Species in Turkey and Their Relations with Other Juniperus Based on Cpdna Supervisor: Prof
MOLECULAR PHYLOGENETIC ANALYSES OF JUNIPERUS L. SPECIES IN TURKEY AND THEIR RELATIONS WITH OTHER JUNIPERS BASED ON cpDNA A THESIS SUBMITTED TO THE GRADUATE SCHOOL OF NATURAL AND APPLIED SCIENCES OF MIDDLE EAST TECHNICAL UNIVERSITY BY AYSUN DEMET GÜVENDİREN IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF DOCTOR OF PHILOSOPHY IN BIOLOGY APRIL 2015 Approval of the thesis MOLECULAR PHYLOGENETIC ANALYSES OF JUNIPERUS L. SPECIES IN TURKEY AND THEIR RELATIONS WITH OTHER JUNIPERS BASED ON cpDNA submitted by AYSUN DEMET GÜVENDİREN in partial fulfillment of the requirements for the degree of Doctor of Philosophy in Department of Biological Sciences, Middle East Technical University by, Prof. Dr. Gülbin Dural Ünver Dean, Graduate School of Natural and Applied Sciences Prof. Dr. Orhan Adalı Head of the Department, Biological Sciences Prof. Dr. Zeki Kaya Supervisor, Dept. of Biological Sciences METU Examining Committee Members Prof. Dr. Musa Doğan Dept. Biological Sciences, METU Prof. Dr. Zeki Kaya Dept. Biological Sciences, METU Prof.Dr. Hayri Duman Biology Dept., Gazi University Prof. Dr. İrfan Kandemir Biology Dept., Ankara University Assoc. Prof. Dr. Sertaç Önde Dept. Biological Sciences, METU Date: iii I hereby declare that all information in this document has been obtained and presented in accordance with academic rules and ethical conduct. I also declare that, as required by these rules and conduct, I have fully cited and referenced all material and results that are not original to this work. Name, Last name : Aysun Demet GÜVENDİREN Signature : iv ABSTRACT MOLECULAR PHYLOGENETIC ANALYSES OF JUNIPERUS L. SPECIES IN TURKEY AND THEIR RELATIONS WITH OTHER JUNIPERS BASED ON cpDNA Güvendiren, Aysun Demet Ph.D., Department of Biological Sciences Supervisor: Prof. -
Morphology and Morphogenesis of the Seed Cones of the Cupressaceae - Part II Cupressoideae
1 2 Bull. CCP 4 (2): 51-78. (10.2015) A. Jagel & V.M. Dörken Morphology and morphogenesis of the seed cones of the Cupressaceae - part II Cupressoideae Summary The cone morphology of the Cupressoideae genera Calocedrus, Thuja, Thujopsis, Chamaecyparis, Fokienia, Platycladus, Microbiota, Tetraclinis, Cupressus and Juniperus are presented in young stages, at pollination time as well as at maturity. Typical cone diagrams were drawn for each genus. In contrast to the taxodiaceous Cupressaceae, in Cupressoideae outgrowths of the seed-scale do not exist; the seed scale is completely reduced to the ovules, inserted in the axil of the cone scale. The cone scale represents the bract scale and is not a bract- /seed scale complex as is often postulated. Especially within the strongly derived groups of the Cupressoideae an increased number of ovules and the appearance of more than one row of ovules occurs. The ovules in a row develop centripetally. Each row represents one of ascending accessory shoots. Within a cone the ovules develop from proximal to distal. Within the Cupressoideae a distinct tendency can be observed shifting the fertile zone in distal parts of the cone by reducing sterile elements. In some of the most derived taxa the ovules are no longer (only) inserted axillary, but (additionally) terminal at the end of the cone axis or they alternate to the terminal cone scales (Microbiota, Tetraclinis, Juniperus). Such non-axillary ovules could be regarded as derived from axillary ones (Microbiota) or they develop directly from the apical meristem and represent elements of a terminal short-shoot (Tetraclinis, Juniperus). -
Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- ERICACEAE
Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- ERICACEAE ERICACEAE (Heath Family) A family of about 107 genera and 3400 species, primarily shrubs, small trees, and subshrubs, nearly cosmopolitan. The Ericaceae is very important in our area, with a great diversity of genera and species, many of them rather narrowly endemic. Our area is one of the north temperate centers of diversity for the Ericaceae. Along with Quercus and Pinus, various members of this family are dominant in much of our landscape. References: Kron et al. (2002); Wood (1961); Judd & Kron (1993); Kron & Chase (1993); Luteyn et al. (1996)=L; Dorr & Barrie (1993); Cullings & Hileman (1997). Main Key, for use with flowering or fruiting material 1 Plant an herb, subshrub, or sprawling shrub, not clonal by underground rhizomes (except Gaultheria procumbens and Epigaea repens), rarely more than 3 dm tall; plants mycotrophic or hemi-mycotrophic (except Epigaea, Gaultheria, and Arctostaphylos). 2 Plants without chlorophyll (fully mycotrophic); stems fleshy; leaves represented by bract-like scales, white or variously colored, but not green; pollen grains single; [subfamily Monotropoideae; section Monotropeae]. 3 Petals united; fruit nodding, a berry; flower and fruit several per stem . Monotropsis 3 Petals separate; fruit erect, a capsule; flower and fruit 1-several per stem. 4 Flowers few to many, racemose; stem pubescent, at least in the inflorescence; plant yellow, orange, or red when fresh, aging or drying dark brown ...............................................Hypopitys 4 Flower solitary; stem glabrous; plant white (rarely pink) when fresh, aging or drying black . Monotropa 2 Plants with chlorophyll (hemi-mycotrophic or autotrophic); stems woody; leaves present and well-developed, green; pollen grains in tetrads (single in Orthilia). -
Phylogeny and Phylogenetic Taxonomy of Dipsacales, with Special Reference to Sinadoxa and Tetradoxa (Adoxaceae)
PHYLOGENY AND PHYLOGENETIC TAXONOMY OF DIPSACALES, WITH SPECIAL REFERENCE TO SINADOXA AND TETRADOXA (ADOXACEAE) MICHAEL J. DONOGHUE,1 TORSTEN ERIKSSON,2 PATRICK A. REEVES,3 AND RICHARD G. OLMSTEAD 3 Abstract. To further clarify phylogenetic relationships within Dipsacales,we analyzed new and previously pub- lished rbcL sequences, alone and in combination with morphological data. We also examined relationships within Adoxaceae using rbcL and nuclear ribosomal internal transcribed spacer (ITS) sequences. We conclude from these analyses that Dipsacales comprise two major lineages:Adoxaceae and Caprifoliaceae (sensu Judd et al.,1994), which both contain elements of traditional Caprifoliaceae.Within Adoxaceae, the following relation- ships are strongly supported: (Viburnum (Sambucus (Sinadoxa (Tetradoxa, Adoxa)))). Combined analyses of C ap ri foliaceae yield the fo l l ow i n g : ( C ap ri folieae (Diervilleae (Linnaeeae (Morinaceae (Dipsacaceae (Triplostegia,Valerianaceae)))))). On the basis of these results we provide phylogenetic definitions for the names of several major clades. Within Adoxaceae, Adoxina refers to the clade including Sinadoxa, Tetradoxa, and Adoxa.This lineage is marked by herbaceous habit, reduction in the number of perianth parts,nectaries of mul- ticellular hairs on the perianth,and bifid stamens. The clade including Morinaceae,Valerianaceae, Triplostegia, and Dipsacaceae is here named Valerina. Probable synapomorphies include herbaceousness,presence of an epi- calyx (lost or modified in Valerianaceae), reduced endosperm,and distinctive chemistry, including production of monoterpenoids. The clade containing Valerina plus Linnaeeae we name Linnina. This lineage is distinguished by reduction to four (or fewer) stamens, by abortion of two of the three carpels,and possibly by supernumerary inflorescences bracts. Keywords: Adoxaceae, Caprifoliaceae, Dipsacales, ITS, morphological characters, phylogeny, phylogenetic taxonomy, phylogenetic nomenclature, rbcL, Sinadoxa, Tetradoxa. -
Eastern North American Plants in Cultivation
Eastern North American Plants in Cultivation Many indigenous North American plants are in cultivation, but many equally worthy ones are seldom grown. It often ap- pears that familiar native plants are taken for granted, while more exotic ones - those with the glamor of coming from some- where else - are more commonly cultivated. Perhaps this is what happens everywhere, but perhaps this attitude is a hand- me-down from the time when immigrants to the New World brought with them plants that tied them to the Old. At any rate, in the eastern United States some of the most commonly culti- vated plants are exotic species such as Forsythia species and hy- brids, various species of Ligustrum, Syringa vulgaris, Ilex cre- nata, Magnolia X soulangiana, Malus species and hybrids, Acer platanoides, Asiatic rhododendrons (both evergreen and decidu- ous) and their hybrids, Berberis thunbergii, Abelia X grandi- flora, Vinca minor, and Pachysandra procumbens, to mention only a few examples. This is not to imply, however, that there are few indigenous plants that have "made the grade," horticulturally speaking, for there are many obvious successes. Some plants, such as Cornus florida, have been adopted immediately and widely, but others, such as Phlox stolonifera ’Blue Ridge’ have had to re- ceive an award in Europe before drawing the attention they de- serve here, much as American singers used to have to acquire a foreign reputation before being accepted as worthwhile artists. Examples among the widely grown eastern American trees are Tsuga canadensis; Thuja occidentalis; Pinus strobus (and other species); Quercus rubra, Q. palustris, and Q. -
Maryum Bhatti1, Aaron Lee1, Hiba Rahman-Vyas1, Dianella G
NAC SUBFAMILY 1A GENE TREE Combining benchwork and bioinformatics to reconstruct the evolutionary history of CUP- SHAPED COTYLEDON in honeysuckles and relatives Maryum Bhatti1, Aaron Lee1, Hiba Rahman-Vyas1, Dianella G. Howarth2, Michael J. Donoghue3, Wendy L. Clement1 1Department of Biology, The College of New Jersey, Ewing, NJ; 2Department of Biological Sciences, St. John’s University, Jamaica, NY; 3Department of Ecology and Evolution, Yale University, New Haven, CT RECOVERING CUC FROM INTRODUCTION DIPSACALES Context Fusion among adjacent parts is a phenomena that occurs Sampling and Isolating CUC throughout flowering plants. CUP-SHAPED COTYLEDON, CUC, • 38 species were selected across the Dipsacales for a member of the NAC Subfamily 1a transcription factors, and has been shown to affect organ boundary formation.1,2 Honeysuckles, species that had no available genomic resources or Lonicera (Caprifoliaceae, Dipsacales), are known for fusing • Degenerate primers were designed to isolate exons petals into long tubes and also exhibit fusion among ovaries, bracts 1 and 2 of CUC1/2 and CUC3 based on reference and leaves. Variation in fusion across 160 species of Lonicera genomes (e.g., Arabidopsis, Petunia, Snapdragon) make them an excellent system to investigate the evolution of • CUC specific primers were also created including fusion. all introns and exons of CUC Goals • Successful reactions near 500 bp were isolated and • Recover CUC from the phylogenetic diversity of Dipsacales, cloned using PCR and cloning; sample target species in which no • Multiple clones per species were sequenced at the genomic data is available. Yale Sequencing on the Hill Facility • Reconstruct a gene tree for NAC Subfamily 1a and CUC using • Sequences were assembled and manually edited both direct sequences gained in this study and data extracted 3 from available genomic resources. -
Botanical Name Common Name
Approved Approved & as a eligible to Not eligible to Approved as Frontage fulfill other fulfill other Type of plant a Street Tree Tree standards standards Heritage Tree Tree Heritage Species Botanical Name Common name Native Abelia x grandiflora Glossy Abelia Shrub, Deciduous No No No Yes White Forsytha; Korean Abeliophyllum distichum Shrub, Deciduous No No No Yes Abelialeaf Acanthropanax Fiveleaf Aralia Shrub, Deciduous No No No Yes sieboldianus Acer ginnala Amur Maple Shrub, Deciduous No No No Yes Aesculus parviflora Bottlebrush Buckeye Shrub, Deciduous No No No Yes Aesculus pavia Red Buckeye Shrub, Deciduous No No Yes Yes Alnus incana ssp. rugosa Speckled Alder Shrub, Deciduous Yes No No Yes Alnus serrulata Hazel Alder Shrub, Deciduous Yes No No Yes Amelanchier humilis Low Serviceberry Shrub, Deciduous Yes No No Yes Amelanchier stolonifera Running Serviceberry Shrub, Deciduous Yes No No Yes False Indigo Bush; Amorpha fruticosa Desert False Indigo; Shrub, Deciduous Yes No No No Not eligible Bastard Indigo Aronia arbutifolia Red Chokeberry Shrub, Deciduous Yes No No Yes Aronia melanocarpa Black Chokeberry Shrub, Deciduous Yes No No Yes Aronia prunifolia Purple Chokeberry Shrub, Deciduous Yes No No Yes Groundsel-Bush; Eastern Baccharis halimifolia Shrub, Deciduous No No Yes Yes Baccharis Summer Cypress; Bassia scoparia Shrub, Deciduous No No No Yes Burning-Bush Berberis canadensis American Barberry Shrub, Deciduous Yes No No Yes Common Barberry; Berberis vulgaris Shrub, Deciduous No No No No Not eligible European Barberry Betula pumila -
Old Fashioned Weigela (Caprifoliaceae) ------Weigela Florida Is Valued for Its Use As a Specimen Or Assets Mixed Border Plant
Weigela florida - Old Fashioned Weigela (Caprifoliaceae) ----------------------------------------------------------------------------------------------- Weigela florida is valued for its use as a specimen or Assets mixed border plant. Plant Old Fashioned Weigela -profusion of colorful flowers around windows so its abundant pink flowers can be -relatively low maintenance admired each spring. -adaptable and urban tolerant Liabilities FEATURES -becomes scraggly, unkempt, and larger than Form intended unless properly pruned and managed -deciduous, spreading, dense, rounded shrub reaching Habitat 6-9' tall x 9-12' wide, dense, arching, eventually to Zones 4 to 8 the ground Native to Japan -moderate to fast growth rate; low maintenance. Culture SELECTIONS -Weigela grows best in a sunny location and moist Alternates soil. Shaded plants are straggly. Weigela also -other spring-flowering shrubs (Viburnum, dislikes crowding. Rhododendron, Deutzia gracilis, etc.) -transplants well Cultivars - Variants - Related species -some annual dieback and winter injury make annual -many cultivars (at least 170), including variegated pruning necessary. and dwarfs: -becomes rangy unless pruned, no serious pests or -'Bristol Ruby' - red flowers produced sporadically diseases. throughout the growing season. Foliage -'Bristol Snowflake' - white flowers flushed with -leaves simple, opposite, elliptic, 2-4" long x 1" wide, pink. acuminate (pointed tip), rounded at base, serrate, -'Carnaval' - red, white and pink flowers are produced short petiole. on the same plant. -leaf color is medium to dark green in summer, -'Evita' - a slow growing, compact plant with bright turning brownish in autumn. red flowers produced in spring and then periodically. Flowers -'Java Red' - a hardy plant with purplish foliage and deep pink flowers. -'Minuet' - a hardy cultivar with purplish foliage and flowers that are red, purple and yellow. -
Abiotic Factors and Yushania Influences on Abies Forest Composition in Taiwan
Taiwania, 59(3): 247‒261, 2014 DOI: 10.6165/tai.2014.59.247 RESEARCH ARTICLE Abiotic Factors and Yushania Influences on Abies Forest Composition in Taiwan Cheng-Tao Lin(1), Tzu-Ying Chen(2), Chang-Fu Hsieh(3) and Chyi-Rong Chiou(1*) 1. School of Forestry and Resource Conservation, National Taiwan University, No. 1, Sect. 4, Roosevelt Rd., Taipei, 10617, Taiwan. 2. Department of Forestry and Natural Resources, National Ilan University, Sect. 1, Shen-Lung Rd., Ilan, 26047, Taiwan. 3. Institute of Ecology and Evolutionary Biology, National Taiwan University, No. 1, Sect. 4, Roosevelt Rd., Taipei, 10617, Taiwan. * Corresponding author. Tel.: +886-2-3366-4640; Fax: +886-2-2365-4520; Email: [email protected] (Manuscript received 20 March 2014; accepted 26 May 2014) ABSTRACT: Abies kawakamii forests are generally distributed above 3,000 m in Taiwanese high mountains. The community data used in our analysis were derived from the database of the National Vegetation Diversity Inventory and Mapping Project of Taiwan (NVDIMP), and environmental data were obtained from the WorldClim and NVDIMP databases. We used non-metric multidimensional scaling (NMDS) to identify vegetation composition of Abies communities and the structural equation models (SEMs) were used to examine the complex relationships between environmental factors and vegetation composition. The results of ordination showed the most important factors determining species composition of Abies forests involved habitat rockiness, heat load index, warmth index and summer and winter. SEM results approved the warmth index and winter precipitation were the main drivers determining the latent variable—climate, which significantly affect the overstory composition of Abies communities. -
Weigelaspecies and Cultivar Genome Size and Ploidy Estimations: Shrub Breeding©
Weigela species and cultivar genome size and ploidy estimations: shrub breeding© E. Pfarr and J.J. Rothleutnera Tree and Shrub Breeder, The Morton Arboretum, Lisle, Illinois 60532, USA. INTRODUCTION Weigela are among the most popular flowering shrubs for temperate landscapes as they tolerate a wide range of cultural conditions, propagate easily from cuttings, and flower heavily in late spring. The genus is composed of 10 species native to China, Japan, Manchuria, and the Korean peninsula. Since the genus was brought to western horticulture near 1860, over two hundred cultivars have been introduced (Dirr, 2009; Sheffield Botanical Gardens, 2015). Introductions continue today with breeding work emphasizing the development of compact plants, novel foliage colors, and recurrent blooming characteristics. One cultivar, ‘Courtalor’, Carnaval® weigela is widely promoted as a reblooming polyploid (Pantin, 2015; Wood). Because polyploidy may be associated with ornamental characteristics that breeders may be selecting for, such as reblooming, we set out to investigate the presence of polyploidy in natural populations and extent of polyploidy in available cultivars. This manuscript reports genome size and ploidy estimations for 10 species and 46 cultivars, from a total of 74 accessions. METHODS AND MATERIALS Plant material was sampled from plants growing at The Morton Arboretum, the Chicago Botanic Garden, and the Arnold Arboretum of Harvard University. Genome sizes were determined by using a flow cytometer (CyFlow® PloidyAnalyser; Partec. Münster, Germany) with materials and protocols from Cystain PI absolute P test kits (Partec. Münster, Germany). Tissue samples were collected from expanding leaves and co-chopped with an internal standard, a leaf sample of Pisum sativum ‘Ctirad’, with a known genome size of 8.76 pg (Greilhuber et al., 2007). -
IHCA Recommended Plant List
Residential Architectural Review Committee Recommended Plant List Plant Materials The following plant materials are intended to guide tree and shrub ADDITIONS to residential landscapes at Issaquah Highlands. Lot sizes, shade, wind and other factors place size and growth constraints on plants, especially trees, which are suitable for addition to existing landscapes. Other plant materials may be considered that have these characteristics and similar maintenance requirements. Additional species and varieties may be selected if authorized by the Issaquah Highlands Architectural Review Committee. This list is not exhaustive but does cover most of the “good doers” for Issaquah Highlands. Our microclimate is colder and harsher than those closer to Puget Sound. Plants not listed should be used with caution if their performance has not been observed at Issaquah Highlands. * Drought-tolerant plant ** Requires well-drained soil DECIDUOUS TREES: Small • Acer circinatum – Vine Maple • Acer griseum – Paperbark Maple • *Acer ginnala – Amur Maple • Oxydendrum arboreum – Sourwood • Acer palmation – Japanese Maple • *Prunus cerasifera var. – Purple Leaf Plum varieties • Amelanchier var. – Serviceberry varieties • Styrax japonicus – Japanese Snowbell • Cornus species, esp. kousa Medium • Acer rufinerve – Redvein Maple • Cornus florida (flowering dogwood) • *Acer pseudoplatanus – Sycamore Maple • Acer palmatum (Japanese maple, many) • • *Carpinus betulus – European Hornbeam Stewartia species (several) • *Parrotia persica – Persian Parrotia Columnar Narrow