©Naturhistorisches Museum Wien, download unter www.biologiezentrum.at Ann. Naturhist. Mus. Wien, B 112 265-313 Wien, März 2011

Typi Orchidacearum ab Augusto R. Endresio in Costa Rica lecti

F. Pupulin*, C. Ossenbach**, R. Jenny*** & E. Vitek****

Kurzfassung Auguste R. Endres sammelte von Ende 1866 bis 1874 in Costa Rica, für eine kurze Zeit war er auch in . In diesen sieben Jahren widmete er sich insbesonders der Aufsammlung von Orchideen. Nur eine geringe Zahl seiner neuen Funde wurden von ihm selbst publiziert. Elf Arten beschrieb er gemeinsam mit Heinrich Gustav Reichenbach, der weitere 22 Arten aufgrund von Endres' Material beschrieb. Andere Autoren, die mit diesem Material neue Taxa beschrieben, sind Rudolf Schlechter, Fritz Kränzlin und Carlyle A. Luer. Insgesamt wurden 109 Arten und 2 Varietäten auf der Basis von Endres' Sammlungen beschrieben. Hier wird eine kritische Evaluation seiner Orchideentypen in Reichenbach's Sammlungen, die heute im Naturhistorischen Museum Wien deponiert sind, vorgelegt.

A bstract Auguste R. Endres botanized in Costa Rica between the end of 1866 and the first months of 1874, spending a short time in Panama. In these seven years he devoted his main attention to the still unrevealed richness of Costa Rican . The results of his activity have still to be properly evaluated, but his contributions to the of Costa Rica are extraordinary in quantity and quality. Notwithstanding his immense labor, only a very small portion of the orchid he collected, studied and illustrated were published as new to the science. He co-authored eleven with Heinrich Gustav Reichenbach, who himself described another 22 species based on his collections. Other authors who described new taxa through the study of Endres’ plants were Rudolf Schlechter, Fritz Kränzlin and, more recently, Carlyle A. Luer. Including a few other minor contributions, the total number of orchids described as new to science on the basis of Endres’ collections amounts to 109 species and two varieties. In order to begin a complete evaluation of Endres’ botanical legacy, a critical revision of the information on the orchid types collected by him, and conserved in the Reichenbach at the Naturhistorisches Museum of Vienna, is presented. Types are associ- ated with Endres’ illustrations and other materials from his gatherings. Lectotypes are selected for chlorops, ionocentmm, pygmaea, castaneum, O. dielsiamim, O. globu- lifenim var. costaricense, O. rechingerianum, Restrepia prorepens, R. reichenbachiana, Sigmatostalix poikilostalix, endresianus, pfavii var. zonale, Warczewiczella picta. endresii is neotypified. Photographs of the lectotypes and the neotype are presented. Key words: Orchidaceae, typification, Auguste R. Endres, Costa Rica, Reichenbach Herbarium, Naturhis­ torisches Museum Wien

Franco Pupulin, Lankester Botanical Garden, University of Costa Rica, P.O. Box 302-7050 Cartago, Costa Rica; Harvard University Herbaria, Cambridge, MA, U.S.A.; Marie Selby Botanical Gardens, Sarasota, FL, U.S.A.; Andean Orchids Research Center, University Alfredo Perez Guerrero, Quito, - [email protected] - corresponding author Carlos Ossenbach, Andean Orchids Research Center, University Alfredo Perez Guerrero, Quito, Ecuador. *** . Rudolf Jenny, Schweizerische Orchideenstiftung am Herbarium Jany Renz, Botanisches Institut der ^ Universität Basel, Schönbeinstrasse 6, CH-4056 Basel, Schweiz - rjorchidgmx.ch Emst Vitek, Naturhistorisches Museum, Botanische Abteilung, Burgring 7, A-1010 Wien, Austria - [email protected] 266 ©Naturhistorisches Museum Wien,Annalen download des unter Naturhistorischen www.biologiezentrum.at Museums in Wien B j / •>

Introduction

As new information on Auguste R. Endres (1838-1874), his life and botanical activity has become available based on reliable sources (Ossenbach et al. 20 10 , Manning 2010), the time is ripe for a general review of Endres’ biography and the significance of his work for the history of Central American botany (Ossenbach et al., in preparation)

Results

The short career of A.R. Endres as a botanical explorer, author and illustrator, is con- centrated in just a few years that he spent in Costa Rica between the end of 1866 and the first months of 1874, with a brief parenthesis in Panama. In these seven years Endres botanized throughout the country, devoting his main attention to the still unrevealed richness of small orchid plants that are common in the canopies of Costa Rican (for a review of the main botanical itineraries of Endres, and the identification of his col- lecting localities, see O s s e n b a c h et al. 2010). The botanical results of his activity, largely neglected after his death in 1874, have still to be properly evaluated. However, from a preliminary assessment of the materials he prepared in Costa Rica, they must be considered extraordinary both in terms of quantity of vouchers, illustrations and descriptions, as well as in the quality of his meticulous work. Endres’ botanical legacy, almost exclusively preserved in the Herbarium and the Archives of the Natural History Museum of Vienna (Naturhistorisches Museum Wien), with a few duplicates kept at the Oakes Ames Orchid Herbarium, Harvard University, amounts to the astonishing figure of 3518 sheets, 1127 of which contain one or more drawings, plus a number of loose documents, scraps and notebooks that include other species descriptions. Of the Lepanthes alone, his preferred group among the orchids, Endres collected more than 250 specimens, corresponding to 63 species (or more than two thirds of all the taxa known from Costa Rica, according to L u e r 2003), and prepared 201 illustrations. The Herbarium of the Natural History Museum conserves Endres’ drawings, descriptions and exsiccata of more than half of the Costa Rican species of Masdevallia and Trichosalpinx, two thirds of the Oncidium species, three quarters of the known taxa of Platystele, almost half of the species of s.l., and so on. Notwithstanding his immense labor, only a very small portion of the orchid plants he collected, studied and illustrated during his Costa Rican travels (most of which still undescribed at this time) were published as new to Science. He co-authored 11 species with his main scientific correspondent, Heinrich Gustav Reichenbach (1823-1889), then professor of botany and director of the Botanic Garden of the Hamburg University and undisputed authority on orchid . During the time of their correspondence between Costa Rica and Germany, Reichenbach described another four species and one new variety based on collections by Endres, and another 18 new taxa after Endres’ death until 1886. After the death of Prof. Reichenbach and the relocation of his herbarium to the Court Imperial Museum in Vienna (now Naturhistorisches Museum Wien, herbarium W), Friedrich Richard Rudolf Schlechter (1872-1925) and Friedrich (Fritz) Wilhelm Lud­ wig Kränzlin (1847-1934) studied the material collected by Endres and described on [>iiih jl in et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Endresio Wien, download in Costa unter www.biologiezentrum.at Rica lecti 267

ihat basis 3 and 39 new orchid species respectively. Kränzlin, at least, removed from the herbarium of Reichenbach parts of those materials collected by Endres that served as the ivpes for his new species. The largest part of this collection was probably lost during the boinbing of the Berlin/Dahlem Herbarium in 1943, but Kränzlin had previously sold some of the specimens to Oakes Ames in 1924 to make the publication of his mono- M-aph on Masdevallia (K r ä n z l in 1925) possible. They are currently conserved at the Oakes Ames Orchid Herbarium, Harvard University Herbaria (AMES). The following important contribution to Endres’ orchidology was the work by Carlyle A. Luer (1922-), who between 1992 and 1999 published twenty species of Costa Rican (mostly belonging to the genus Lepanthes ) based on plants and draw­ ings of Endres kept at the herbarium in Vienna. If other, minor contributions resulting from the study of Endres’ materials are taken into account (i.e., N ic h o l s o n 1886, V eit c h 1890, P u p u l in 2001, P u p u l in & B o g a r in 2010), the total number of orchids described as new to Science on the basis of Endres’ collections barely amounts to 109 species (with two new genera) and two varieties. When compared with the substantial advances in the knowledge of Mesoamerican orchids produced by the exploratory work and the fine botanical activity of Endres, it is clear that his formal contributions to botany were much less than his commitment deserved. Endres himself was aware of this, and a few months before his death he bitterly wrote to his friend, the Captain John Melmoth Dow (1827— 1892): “Reichenbach has lately repeated hisproposal ofbuying my dried orch. collections and Ifear our first interview will be a stormy one. I begin to consider these cabinet-celebrities as vampires nourishing their inflated fame at the cost of the lifeblood o f those poor fools they condescendingly call “collectors”, and I am thoroughly disgusted, at moments, with the pursuits I have so passionately followed for seven years. / Rchb. lately proposed retarding the publication o f my plants to 1875, in order to adorn, as he says, the catalogue which will be published on occasion o f the — eth. anniversary o f the Hamburgh Botanical Gardens, o f which he is the Director. / My own opinion is that, if any further retardation takes place it will emanate from a resolu- tion of relinquishing his invaluable cooperation, and of working up the material myself and by myself. / Would that I had done this long ago I should then at least have enjoyed the public credit which is my just due in reward for the sacrifices I have made. If by sheer luck, not worn out bodily, my patience at least is exhausted" (E n d r e s 1874). The present paper is the first contribution to a complete evaluation of Endres’ botanical legacy, and it is aimed to critically revise the information on the orchid types that were collected by Endres, associating them with Endres’ illustrations and other materials from his gatherings, and lectotypifying his species when needed. Endres only rarely used collection numbers in the strict sense, but used numbers and annotations to correlate plants with figures and descriptions - these original notes are given in « ». Homotypic synonyms are given with =, all mentioned synonyms with the year of publication. Herbaria are given in [ ] with their acronyms according to Index Herbariorum: http://sweetgum.nybg.org/ih/. For databasing, the specimens in W have got a unique number („W “). The Orchid col­ lection of Reichenbach has an independent numbering system cited as „Rchb. Orch. “ Both numbers are cited to allow correlation with earlier publications. Collections with numbers „W 1889-...“ have been in the general collections of Reichenbach. All specimens can be found in Virtual Herbaria: http://herbarium.univie.ac.at/database/search.php. 268 ©Naturhistorisches Museum Wien,Annalen download des unter Naturhistorischen www.biologiezentrum.at Museums in Wien, B in

Orchid types based on collections by Auguste R. Endres

1. Barkeria lindleyana var. centerae Endres & RcHB.f., Gard. Chron. 1873: 1 597 (1873). = Epidendrum lindleyanum var. centerae H.J. V e i t c h (1890: “Introduced by us from Costa Rica, in 1873, through M. Endres”). Type: Costa Rica. “The original Barkeria Lindleyana was found in Costa Rica Now the lynx-eyed, indefatigable, and really scientific traveller, Mr. Endres, has discovered a very beautiful, prettier-coloured variety of it in Costa Rica I was most agreeably surprised to obtain fresh of this remarkable plant quite recently (beginning of November) from the Royal Exotic Nursery of Messrs. Veitch & Sons”, A.R. Endres s.n. Holotype: Costa Rica, without locality, «Barkeria centeraeana / lindleyana? / 2500’», A.R. Endres s.n. «Gen. Coll. Orch. N. 532» [W 0019041 / Rchb. Orch. 5540, isotype W 0019039 / Rchb. Orch. 13634]. Additional material associated with the type: Costa Rica, without locality, Endres’ coloured drawings of the and floral dissecions, «Barkeria centerana», A.R. Endres s.n. «Gen. Coll. Orch. N. 532, Barkeria N° 1, Sk. 169.a» [W 0018992 / Rchb. Orch. 36564]. In the original protologue of B. lindleyana var. centerae, Reichenbach expressly acknowledged that his description was “likewise the specimens and very complete and excellent descriptions by the friend Endres” (. Juxta specimina et descript. locupletissi- mam, optimam amici Endres ) (Reichenbach 1873).

2. Batemania burtii Endres & RcHB.f., Gard. Chron. 1872: 1099 (1872). = burtii (Endres & RcHB.f.) Pfitzer (1889). = Zygopetalum burtii (Endres & RcHB.f.) B enth . & Hook. f. ex H emsl. (1884). = Huntleya burtii (Endres & RcHB.f.) R olfe (1916), nom illeg. superfl. Type: Costa Rica. “It was discovered so early as June, 1867, in Costa Rica, by M. Endres, who proposed to name it in honor of one of his acquaintances” Holotype: Costa Rica, 1867, A.R. Endres s.n [W 0019751 / Rchb. Orch. 37976 (plant, Endres’ description «... Costa Rica / 1867 / Batemania Burtii», drawing of the flower and floral dissections)]. Additional Endres’ material: flowers and Endres’ description, «Batemania Burtii / ...», A.R. Endres 100 [W 0019752 / Rchb. Orch. 13599]; - Endres’ sketch of the plant and flower, with two descriptions, A.R. Endres 100 [W 0019759 / Rchb. Orch. 36019]; - Endres’ drawings of the flowers, floral dissections and descriptions, A.R. Endres 100 [W 0019758 / Rchb. Orch. 31952]; - Endres’ drawing of the plant, flower and floral dissections, A.R. Endres 100 [W 0019756 / Rchb. Orch. 36590]; - flowers, A.R. Endres s.n. [W 0019754 / Rchb. Orch. 37982]; - Endres’ drawing of a flower, A.R. Endres 109 [W 0019760 / Rchb. Orch. 34712]; - Endres’ drawing of the flower and descriptions, A.R. Endres s.n. [W 0019755 / Rchb. Orch. 18002]; - flowers, A.R. Endres s.n. [W 0019753 / Rchb. Orch. 13598]; - Endres’ sketch of the flower, A.R. Endres s.n. [W 0019757 / Rchb. Orch. 33319].

3. Brassia chlorops Endres & RcHB.f., Gard. Chron. 542. 1873. = chlorops (Endres & RcHB.f.) N.H.W illiams (1972) P u p u l i n et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Endresio Wien, download in Costa unter www.biologiezentrum.at Rica lecti 269

------« o s e w r s w r W 0019028 HERB. MUS. PA LAT. YfNDOB. 'firhroW •. 2 f3!) reserved Copyright

Ada chlorops (Sndr. & Rchb«f.) K.H. W illiam s, comb, nov.

d e t. Morris H. ü llia m s May 1971

Fig. 1: Lectotype of Brassia chlorops E n d r e s & RcHB.f. [W 0019028 / Rchb. Orch. 2439]. 270 ©Naturhistorisches Museum Wien,Annalen download des unter Naturhistorischen www.biologiezentrum.at Museums in Wien, B //i

Syntypes: “There are ... five dry specimens of it in my herbarium, gathered in Costa Rica by Mr. Endres, the skilful lynx-eyed investigator {Costa Rica, A.R. Endres 243, [W 0019028 / Rchb. Orch. 2439, W 0019029 / Rchb. Orch. 2435, W 0019030 / Rchb. Orch. 2434, W 0019031 / Rchb. Orch. 2437, W 0019032 / Rchb. Orch. 47630, W 0019034 / Rchb. Orch. 2438} and by Mr. Julian Carmiol {Costa Rica, J Carmiol s.n. [W 0019033 / Rchb. Orch. 18243]}. The fresh specimen come from Messrs. Veitch, who imported it likewise from Costa Rica {Costa Rica, cultivated by Veitch s.n., not located}” Lectotype (hic designatus): «near Mesospinidium / Septr», A.R. Endres 243 [W 0019028 / Rchb. Orch. 2439] (Fig. 1). Isolectotypes: [W 0019029 / Rchb. Orch. 2435, W 0019030 / Rchb. Orch. 2434, W 0019031 / Rchb. Orch. 2437, W 0019032 / Rchb. Orch. 47630 (including Endres’ drawing of the flower, floral dissections, and description), W 0019034 / Rchb Orch. 2438], Additional material associated with the type: Reichenbach’s manuscript description and drawing of material from Endres [W 0018985 / Rchb. Orch. 47631]; inflores- cences with flowers («Brassia (Mesospinidium) / Brachtia?»), A.R. Endres s.n. [W 0019035 / Rchb. Orch. 2436]. In the protologue, Reichenbach cited three different gatherings, for two of which mate­ rial is still in existence at W. His diagnostic drawings, however, were ostensibly pre­ pared from material sent by Endres. W 0019028 is the only sheet of Endres 243 that includes the vegetative parts of the plant, and for this reason it is selected as the lecto ­ type.

4. endresii S c h l t r . , Repert. S p . Nov. Regni Veg. 17: 14 (1921). = Chondroscaphe endresii (S chltr .) D ressler (2002). Type: Costa Rica. “Ohne nähere Standortsangabe”, A.R. Endres 166 [Holotype W 0018830/Rchb. Orch. 49751]. Additional material associated with the type: Endres’ drawings of the type and descriptions, A.R. Endres 166 [W 0018831 / Rchb. Orch. 10664]; - Endres’ draw­ ings of the flower and flower details, A.R. Endres 166 [W 0018833 / Rchb. Orch. 36691]; - Endres’ description, A.R. Endres 166 [W 0018832 / Rchb. Orch. 33497]. Among the materials referable to the type at W is a manuscript name by Reichenbach, who probably intended to publish the species with the name Chondrorhyncha umbon- ata, and the name «umbonata», in Reichenbach’s handwriting, is noted on an envelope mounted on the type sheet. A copy of the original drawings of the type by R. Schlechter, done under his supervision, is conserved at the Oakes Ames Orchid Herbarium, Harvard University [AMES 106743].

The name is a synonym of Chondroscaphe bicolor (R olfe) D ressler (type: R. Pfau s.n., [K]). For other synonyms of the species, see Pupulin (2010).

5. Chondrorhyncha reichenbachiana Schltr ., Repert. Spec. Nov. Regni Veg. 17: 15 (1921). = Benzingia reichenbachiana (S chltr .) D ressler (2010). Type: Costa Rica. “Cataratas, blühend Marz-August”, A.R. Endres 557 [Holotype W 0018829 / Rchb. Orch. 49753]. I>it ij l i n et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Endresio Wien, download in Costa unter www.biologiezentrum.at Rica lecti 271

Additional material associated with the type: Endres’ drawings of the type and descriptions, A.R. Endres 557 [W 0018826 / Rchb. Orch. 28550]. Additional Endres’ material: Costa Rica, without locality, dissected flower, A.R. Endres 659 [W 0018823 /Rchb. Orch. 13614]; - without locality, A.R. Endres s.n. [W 0018827 ' Rchb. Orch. 13583], The original drawings by Endres of his «Zygopetalum N° 10» were used by Reichen­ bach to prepare a plate for his Xenia Orchidacea, under the intended name of «Chon- drorhyncha lamellata», finally never published. The same name is annotated in Reichen- bach’s handwriting on the sheet W 0018827 / Rchb. Orch. 13583. The drawings of the type, originally prepared by Schlechter, were published in 1931 by R. M a n s f e l d (1931).

6. Cryptarrhena quadricornu Kr a e n z l ., Pflanzern-. 4, 50 (Heft 80): 314 (1922). = Ciyptarrhena guatemalensis S c h l t r . (1911). Type: “Costa Rica, schwule, niedrig gelegene Gegenden an der atlantischen Seite”, A. R. Endres s.n. «N° 658 of gen. coll. Orch.» [Holotype W 0018849 / Rchb. Orch. 2427 (with Endres’ description and Kränzlin’s drawing of the flower].

7. Cryptophoranthus endresianus Kr a e n z l ., Repert. Spec. Nov. Regni Veg. 17: 437 (1921). = Zootrophion endresianum (K r a e n z l .) L u e r (1982). Type: Costa Rica. “Ohne genaueren Standort”, 1867, A.R. Endres s.n. «Orch. N° 7» [Holotype W 0018865 / Rchb. Orch. 15003 (annotated by Kränzlin, with Endres’ description and drawings and Reichenbach’s sketches), isotype: W 0018864 / Rchb. Orch. 15004]. Additional Endres’ material: Costa Rica. «San Ramon, camino a San Carlos», Endres’ descriptions and drawing of plant and flower, A.R. Endres s.n. «N° 3» [W 0018863 / Rchb. Orch. 36510]; - without locality, Endres’ coloured drawing of plant and flower, A.R. Endres s.n. «Orch. N° 3» [W 0018866]; - Endres’ coloured drawing of plant and flower, A.R. Endres s.n. «N° 3» [W 0018867 / Rchb. Orch. 36228]; - without locality, A.R. Endres s.n. [W 1889-29950]; without locality, A.R. Endres s.n. [W 1889-39082].

8. Dichaea cryptarrhena R cH B .f. ex Kr a e n z l ., Pflanzenr. 4 (50): 36 (1923). Type: “Subäquatoriale andine Provinz. Costa Rica, ohne genaueren Standort”, A.R. Endres 28 [Holotype W 0019083 / Rchb. Orch. 44067]. Additional material associated with the type: Costa Rica. «San Ramon», Endres’ drawings of the type and descriptions, A.R. Endres 28 [W 0019084 / Rchb. Orch. 38604], The publication of D. ciyptarrhena (March 1923) antedates that of the synonymous D. ovatipetala S c h l t r . (November 1923) by eight months.

9. Dichaea dammeriana Kr a e n z l ., Engler Pflanzenr. 4, 50 (83): 41 (1923). Syntypes: “Subäquatoriale andine Provinz. []. Sierra Nevada de S.ta Marta, H. Smith 2773 [AMES, US, NY]; - Costa Rica [Without specific locality, 1868], A. R. Endres 66” [AMES-26090, W 0019086 / Rchb. Orch. 18036, W 0019087/Rchb. Orch. 14751, W 0019090/Rchb. Orch. 14747, W 0019089 / Rchb. Orch. 10517, W 0019088 / Rchb. Orch. 14778]. Lectotype (P u p u l in 2007: 68): [AMES-26090, isolectotypes W 0019086 / Rchb. 272 ©Naturhistorisches Museum Wien,Annalen download des unter Naturhistorischen www.biologiezentrum.at Museums in Wien B / / •>

Orch. 18036 (with Endres’ and Reichenbach’s drawings), W 0019087 / Rchb Ord 14751, W 0019090 / Rchb. Orch. 14747, W 0019089 / Rchb. Orch. 10517- \v 0019088 / Rchb. Orch. 14778]. Additional material associated with the type: Costa Rica, «San Ramon», Endres' drawings and descriptions of the syntype, A.R. Endres 66 [W 0019085 / Rd^ Orch. 38600], Endres’ drawing of the syntype [W 0019092 / Rchb. Orch. 36738] The isolectotypes at W are sterile, while the specimen at AMES is fertile, and for this reason it was seleected as the lectotype.

10. Dichaea diandra R c h b .f., Bot. Zeit. 35: 41 (1877). Type: “Costa Rica. Endres”, A.R. Endres s.n. [Holotype to be found in the Reichen­ bach Herbarium, W, not located; no isotypes known to exist]. Endres’ type collection is likely referable to a species of sect. Dichaea, the only group that, according to the diagnosis, has abaxially verrucose (“ sepalis extus minute verm cosis”). However, while in the protologue Reichenbach first assigns the species to a group with muricate fruits (“ Echinocarpae ”), he later describes the ovary as glabrous (“ovano laeve”). Naked pollinia, without any stipe or viscidium (“ caudicula glandu- laque destitutis ”), are otherwise known in D. trichocarpa, but the lip of this species with its characteristic discoid hypochile is hard to define as “ humeratus ” The of D. diandra, with two lateral anthers (“zrc mediana oppositis ”) was likely a monster, and in the absence of a type specimen or any other material associated with the original proto­ logue, Reichenbach’s diandrous Dichaea cannot be assigned with certainty to any of the known taxa ( P u p u lin 2010).

11. Dichaea glabrescens K r a e n z l . , Pflanzenr. 4, 50: 41 (1923). Type: “Subäquatoriale andine Provinz, Costa Rica, [1867], A.R. Endres 118. [Holo­ type W 019150 / Rchb. Orch. 14776 (but without Edres number), isotype W 019146 /Rchb. Orch. 14776]. Additional material associated with the type: Costa Rica. Endres’ drawings of the type, A.R. Endres 118 [W 0019160 / Rchb. Orch. 38597]; - «Potrerillos», Endres’ description «Dichaea heteromorpha», A.R. Endres 118 [W 0019160 / Rchb. Orch. 36720]. Additional Endres’ material: Costa Rica. Endres’ drawings, A.R. Endres 12 [W 0021667 / Rchb. Orch. 38596]; -Costa Rica, A.R. Endres s.n. [W 0019145 / Rchb. Orch. 3628, W 0019147 / Rchb. Orch. 3634, W 0019148 / Rchb. Orch. 3636, W 0019149 / Rchb. Orch. 3630, W 0019151 / Rchb. Orch. 3629, W 0019152 / Rchb. Orch. 3635]. According to a manuscript annotation by Kränzlin on the type sheet, he changed the name from glabrescens to selaginella, but „it is the same species“ The name Dichaea selaginella was eventually not published by Kränzlin, because Schlechter had already used this name in 1920 for a species from Colombia (type: M. Madero s.n., B, destroyed), synonymous with Dichaea hystricina R c h b .f. (1865).

The name, D. glabrescens, is a synonym o f Dichaea tuerckheimii S c h l t r . (Orchis 10: 188. 1916), non K r a e n z l . 1923 = Dichaea muricatoides H a m e r & G a r a y .

12. Endresiella zahlbruckneriana S c h l t r . , Repert. Spec. Nov. Regni Veg. 17: 14 (1921). I>( i'ULiN et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Endresio Wien, download in Costa unter www.biologiezentrum.at Rica lecti 273

Trevoria zahlbruckneriana (S chltr .) Garay (1970). Type: “Costa Rica, an der Strasse von San Ramon nach San Carlos, Legua. Blühend im September”, A.R. Endres 512 [Holotype W 0019449 / Rchb. Orch. 43634]. The Oakes Ames Orchid Herbarium at the Harvard University hosts the copy of the Illustration of the type, originally prepared by Schlechter and traced under his supervi- sion [AMES 99111]. 13. Epidendrum endresii RcHB.f., Gard. Chron., n.s., 19: 432 (1883). = Oerstedella endresii (RcHB.f.) H ägsater (1981). Syntypes: Costa Rica. “First of all I had some branches from my poor friend Endres, who found it flowering in December at Costa Rica. The flowers were sim- ply described as white - lilac {Costa Rica, A.R. Endres 502 [W 0024422 / Rchb. Orch. 38657, W 0023883 / Rchb. Orch. 13657, W 0019970 / Rchb. Orch. 14548]}. Then my friend Mr. F.C. Lehmann sent me a glorious plant and some branches in spirit, which now lie before me, I had at the same time a veiy carefully prepared picture, which appears to have been taken from the very same individual I had in spirits. Mr Lehmann found it flowering in March, 1878, in Costa Rica {Costa Rica, F.C. Lehmann 177 [W 0019968 / Rchb. Orch. 50032], - drawing by F.C. Lehmann [W 0027548 / Rchb. Orch. 19679]}. Now, Mr. F. Sander sends me a fresh and twigs {Without locality, cultivate by F. Sander s.n. [W 0027532 / Rchb. Orch. 50032]}. Lectotype (Santiago & H ägsater 2007: 931): Costa Rica, F.C. Lehmann 177 [W 0019967 / Rchb. Orch. 50033]. Additional material associated with the type: drawings of Reichenbach «241» [W 0027549 / Rchb. Orch. 8657] - s.coll. (cf. Lehmann) 177 [W 0027545 / Rchb. Orch. 4485], Additional Endres’ material: Costa Rica, «Heights S.th of Cartago / Tablazo», Endres’ drawing of the flower and floral dissections, A.R. Endres 502 [W 0024422 / Rchb. Orch. 38657]; - plant, fertile, A.R. Endres 502 [W 0023883 / Rchb. Orch. 13657, W 0023885 / Rchb. Orch. 14727]; - plant and Endres’ description, A.R. Endres 502 [W 0019970 / Rchb. Orch. 14548]; - Endres’ drawing of the flower and floral dissections, A.R. Endres 502 [W 0024422 / Rchb. Orch. 38657]; - Endres’ drawing of the flower and floral dis­ sections, A.R. Endres «108» [W 0024490 / Rchb. Orch. 36565]; - «Tablazo of Cartago, Coralillo», A.R. Endres s.n. [W 1889-0031372]; - without locality, A.R. Endres s.n. [W 0019971 / Rchb. Orch. 13621]; - «Tablazo / Fila de Atarrazü», A.R. Endres s.n. [W 0019969 / Rchb. Orch. 13617]; - «Tablazo, Coralillos», A.R. Endres s.n. «sp. aff 83» [W 0019967 / Rchb. Orch. 50033]. 14. Epidendrum ionocentrum RcHB.f., Gard. Chron., n.s., 20: 8 (1883). = Epidendrum prismatocarpum var. ionocentrum (RcHB.f.) Teusch . (1969). - ionocentra (RcHB.f.) W.E.Higgins (1998). = Hormidium ionocentrum (RcHB.f.) B rieger (1977). - ionocentra (RcHB.f.) M ora -Re t . & J.Garcia (1991). = Pseudencyclia ionocentra (RcHB.f.) V.P.Castro & C hiron (2003). = Panarica ionocentra (RcHB.f.) W ithner & P.A.Harding (2004). s Encyclia prismatocarpa var. ionocentra (RcHB.f.) M.W olff & O.Gruss (2007). Syntypes: Costa Rica. “This curious and rare species was found by Messrs. Endres 274 ©Naturhistorisches Museum Wien,Annalen download des unter Naturhistorischen www.biologiezentrum.at Museums in Wien, B / /i

(R.A. Endres, various collections at W-Rchb. Orch. from Birris, Santa Maria de Dota, and Naranjo) and Pfau (R. Pfau s.n., not located). Lectotype (hic designatus): Costa Rica, Cartago, «Sitio de don / Eusebio Ortiz / Bir­ ris», plant with flowers, A.R. Endres s.n. (habit sp. N° 522) [W 0024579 / Rchb Orch. 5554] (Fig. 2). Isolectotypes: «Sitio Ortiz / Birris», inflorescence and flowers, A.R. Endres s.n. [W 0024551 / Rchb. Orch. 6499]; - «Sitio Ortiz / Birris», inflorescence and flower A.R. Endres s.n. [W 0024552 / Rchb. Orch. 5501]; «Sitio Ortiz / etc.», inflores- cence and flowers, A.R. Endres s.n. [W 0024415 / Rchb. Orch. 5508]; «Sitio Ortiz, Birris», inflorescence and flowers, A.R. Endres s.n. [W 0024419 / Rchb. Orch. 6500]; - «Sitio de don / Eusebio Ortiz / Birris», inflorescence and flowers, A.R. Endres s.n. [W 0024426 / Rchb. Orch. 5502]; - «Sitio Ortiz», inflorescence and flowers, A.R. Endres s.n. [W 0024429 / Rchb. Orch. 5510]. Additional material associated with the type: flower and Reichenbach’s sketch of the lip and the column, A.R. Endres s.n. (N° 522 of gen. coll.) [W 0020200 / Rchb Orch. 43669]. Additional Endres’ material: Costa Rica. San Jose: Dota, «around Sta. Maria ... Epiden- drum ionocentrum / Endr. Rchb.f.», flower and Endres’ description, A.R. Endres s.n. «N° 522, Epidendr. N° 55» [W 0020202 / Rchb. Orch. 14530]; - Alajuela: San Ramön, «Colorado near Naranjo. id. Birris», Endres’ description and sketch of the lip, A.R. Endres s.n. «N° 522 of gen. coll.» [W 0024580 / Rchb. Orch. 38661]; - Without locality, flowered inflorescence, A.R. Endres s.n. «N° 70 and 522» [W 0020201 / Rchb. Orch. 5543]. Hägsater (correction label, 2009) annotated Rchb. Orch. 5554 [W 0024579] and Rchb. Orch. 38661 [W 0024580] as isotypes, probably because Dressier (correction label, 2001) had previously annotated Rchb. Orch. 5543 [W 0020201] as the holotype. How­ ever, the specimen selected by Dressier has no indication of locality, and bears Endres reference to two different collections (numbers 70 and 522), while the assumed isotypes have indications of locality which only partially coincide. Neither the protologue nor any of the specimens annotated by Reichenbach have an indication of locality, but Endres’ collections of Epidendrum ionocentrum came at least from three different localities, i.e. Santa Maria de Dota (prov. San Jose), Naranjo [de Juan Vinas] (prov. Cartago), and Birris (prov. Cartago). Specimens from the last locality are more abundant, and include the only complete specimen (W 0024579), with the ter­ minal portion of the pseudobulb, , and flowered inflorescence). For this reason we propose to select this specimen as the species lectotype, regarding all the other collec­ tions from the same locality as isolectotypes, with the exception of W 0024580, which includes references to two different localities. The name, Epidendrum ionocentrum , was ostensibly suggested by Endres (annotation on W 0020202), who proposed himself as a co-author of the species. His name, how­ ever, was not included in the published description. 15. Epidendrum phyllocharis Rchb.f., Xenia Orchid. 3: 11-12, tab. 208, fig. 1-5 (1900) [“ 1878”]. Type: Costa Rica. “Diese in ihrer Art durch die prächtigen starren tief gefärbten Fig. 2: Lectotype of of Lectotype 2: Fig. ir i LiN PiirU et al.: Typi Orchidacearum Augusto R. Endresio in Costa Rica lecti Rica Costa in Endresio R. Augusto Orchidacearum Typi al.: et

Copyright reserved Epidendrum ionocentrum Epidendrum ©Naturhistorisches MuseumWien,downloadunterwww.biologiezentrum.at

R b h c . . W0259/Rh. rh 5554] Orch. Rchb. / 0024579 [W f. E:E ästr20 EP 42557+AMO ESP DET: 2009 Hägsater E. Isotype of: Epidendrum ionocentrum Rchb.f. ionocentrum Epidendrum Isotype of: AVR UIEST HERBARIA UNIVERSITY HARVARD Prosthechea ionocentra (Rchb.f.) W.E.Higgins (Rchb.f.) ionocentra Prosthechea '' _Herb. S Orchid. ö Nr. 'I'■' Gard. Chron. 20: 8. 1883. = 1883. 8. 20: Chron. Gard. HERB. «U S . PAI.AT. TOTOOB. TOTOOB. PAI.AT. . S «U HERB. eb Msi aa. Vindob. Palat. Musei Herb. O AMO M A IO R A B R E H NATlJRHISTGaiSCHES * MUSEUMWEN i/3(/iXt^S . le e \ E nd ras nd E \ e le z _ _ £<<■ J'n^ ______- 5 7 2 276 ©Naturhistorisches Museum Wien,Annalen download des unter Naturhistorischen www.biologiezentrum.at Museums in Wien, B / /1

Blätter ungemein stattliche, steife Pflanze führten wir im Hamburger Botanischen Garten aus Costa Rica von Herrn Endres ein”, 1877, cult. at Hamburg Botanic Gar­ den, A.R. Endres 2632 [Holotype W 0027534 / Rchb. Orch. 52931], Additional material associated with the type: drawing and engraved copy of tlr drawing intended for Xenia Orchidacea, [W 0027533 / Rchb. Orch. 375], Additional Endres’ material: Costa Rica, without locality, A.R. Endres s.n, «281» [ty 0027535 / Rchb. Orch. 43674],

16. Kefersteinia endresii Pupulin , Ann. Naturhist. Mus. Wien, B, 103: 543 (2001). Type: Costa Rica, without definite locality, A.R. Endres s.n. [Holotype W 27536 / Rchb. Orch. 18048]. This is the only known collection of a species apparently rare, with no close relatives within Kefersteinia. The callus of the lip, which extends from the base to about the half of the blade, formed by two inflate, pulvurulent keels that are abruptly reflexed at apex, where they become concave and form a 4-lobed tip, is unique and unmistakable.

17. Lepanthes atrata Endres ex Luer , Lindleyana 10: 135 (1995). Type: Costa Rica, without locality, ca. 1867, A.R. Endres 18 [Holotype W 1889- 0041458]. Additional material associated with the type: Costa Rica. Endres’ coloured drawing of the flower, A.R. Endres 18 [W 0019703 / Rchb. Orch. 7592]; - Endres’ drawings of the plant, flower and flower details, and description, A.R. Endres 18 [W 0019704 / Rchb. Orch. 38635]; - Endres’ pencil plate, A.R. Endres 18 [W 0019705 / Rchb. Orch. 36246], Additional Endres’ material: Costa Rica, without locality, A.R. Endres s.n. [W 1889- 40330]; - Uncertain locality, A.R. Endres 509 [W 1889-29962, W 1889-29970, W 1889-29895; W 1889-142252].

18. Lepanthes candida Endres ex Luer , Lindleyana 10: 138 (1995). Type: Costa Rica. Cartago: epiphytic in above San Cristobal Norte, alt. 1680 m, 26. 3. 1995, C. Luer, J. Luer, R. Dressier, K. Dressier & D. Mora de Retana 17476 [Holotype CR, isotype MO] Paratypes: Costa Rica. Alajuela: San Ramon, Portillo Prenzas [?], Endres’ drawings of the flower and floral details, with descriptions, A.R. Endres «N° 33 Gen. coll.» [W 0019679 / Rchb. Orch 27518 (mixed with L. confusa A mes & C. Schweinf ., according to Luer, correction label 1995) ]; - Endres’ description and floral analy­ sis, A.R. Endres 33 [W 0019677 / Rchb. Orch 27485]; - Endres’ drawings of the plant and flower, and description, A.R. Endres «N° 33 Gen. coll.» [W 0019684 / Rchb. Orch 36134]; - Endres’ drawings of the plant and flower, A.R. Endres «N° 33 Gen. coll.» [W 0019683 / Rchb. Orch 27542]; - Endres’ drawings of various flow­ ers, A.R. Endres «N° 33 Gen. coll. Orch.» [W 0019683 / Rchb. Orch 27542]. - Without collection data, Endres’ drawings of plant and flowers, A.R. Endres «657 Gen. coll. Orch.» [W 0019678 / Rchb. Orch 38590]. Additional Endres’ material: Costa Rica, without locality, Endres’ coloured drawing of the flower, A.R. Endres s.n. [W 0019676 / Rchb. Orch. 7593]; - Endres’ drawing of the flowered plant, A.R. Endres s.n. «sketch 228» [W 0019682 / Rchb. Orch. 38692]; - Endres’ pencil plate, A.R. Endres s.n. [W 0019680 / Rchb. Orch. 36152]; - Endres’ pencil I>iip u l in et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Endresio Wien, download in Costa unter www.biologiezentrum.at Rica lecti 277

drawing of a flower, A.R. Endres s.n. [W 0019681 / Rchb. Orch. 7631];-W ithout locality, pndres’ colored drawing of the flower of «Lepanthes müllneriana», A.R. Endres s.n. [W 0019675 / Rchb. Orch. 7618]; - Candelaria, crest of Cerro del Dragon (Legua de los npsamparados, Savannas), A.R. Endres s.n. [W 1889-29954]; - Without locality, A.R. Endres s.n. [W 1889-29912, W 1889-29916], 19 Lepanthes caroli-lueri B ogarin & Pupulin, Harvard Pap. Bot. 15 (1): 116-118, f. ' 4A-E (2010). Type: Costa Rica, Boundary between Alajuela and Heredia provinces, Grecia, Sara- piqui, Colonia Virgen del Socorro, road to Cariblanco, bridge on San Fernando river, 10°16’32“N 84°10’16“W, 750 m, shores of San Fernando river, 13 February 2004, D. Bogarin 741, H. Leön-Päez, F. Pupulin & E. Salas [Holotype JBL-spirit, isotypes CR, JBL-spirit]. Paratypes: Without collecting data, A.R. Endres s.n. [W 1889-29901]; - Endres’ drawings, A.R. Endres 62 [W 0019016 / Rchb-Orch 36220]; - Endres’ drawings of the plant and flower, with floral dissections, A.R. Endres 604 [W 0019017 / Rchb. Orch. 36205, W 0019018 / Rchb. Orch. 38503, W 0019015 / Rchb. Orch. 07635].

20. Lepanthes circularis Luer , Lindleyana 10(3): 140(1995). Type: Costa Rica, Cartago, “heights above El Tejar, alt. 4500 ft,” ca. 1867, A.R. Endres s.n. [Holotype W 1889-41448]. Additional material associated with the type: Illustration of type by C.A. Luer [W 0021489].

21. Lepanthes dichroma Luer , Lindleyana 11: 59 (1996). Type: Costa Rica, Cartago, “this side of San Cristobal and Santa Maria, 5 June”, ca. 1867, A.R. Endres s.n. [Holotype W 1889-142238],

22. Lepanthes dotae Endres ex Luer , Lindleyana 10 (3): 143 (1995). Type: Costa Rica, San Jose, “Dota, rainy season, Sept.-Nov.”, ca. 1867 [the date is not stated by Endres on the type material; it is probable that Luer assumed the year 1867 as a standardized reference], A.R. Endres s.n. «Lepanthes N° 56, sketch 253» [Holotype 0019635 / Rchb. Orch 38523]. Additional material associated with the type: Endres’ drawings of the flower and floral dissections, A.R. Endres s.n. «Lepanthes N° 56, sketch 253» [W 0019632 / Rchb. Orch. 38524]; - Endres’ coloured drawing of the flower, A.R. Endres s.n. [W 0019633 / Rchb. Orch. 7596]; - «Lepanthes, see sk. 253», A.R. Endres s.n. [W 0019634/Rchb. Orch. 7873]. 23. Lepanthes elegans Luer, Lindleyana 10 (3): 146 (1995). Type: Costa Rica. Cartago: “beyond the Laguna, Sitio Ortiz, Birris, Aug.”, ca. 1867, A.R. Endres s.n. (N° 586 gen. coll.) [Holotype W 1889-29930]. Paratype: Costa Rica, Cartago, “Palo Blanco, Sitio Ortiz, Birris, headwaters of Rio Turrialba, 5000-6000 ft., Aug.” [these are different localities: Palo Blanco lies to the South of Cartago, while Sitio Ortiz, Birris and “headwaters of Rio Turialba” are found to the East of Cartago, close to the village of Juan Vinas], ca. 1867, A.R. Endres s.n. [W 1889-142084]. Additional material associated with the type: Endres’ drawings of the flower and flo­ ral dissections, A.R. Endres s.n «N° 586 gen. coll.» [W 0019615 / Rchb. Orch. 278 ©Naturhistorisches Museum Wien,Annalen download des unter Naturhistorischen www.biologiezentrum.at Museums in Wien, B //■>

38672]; - Endres’ coloured drawing of the flower, A.R. Endres s.n «N° 586 gen coll. Orch., Lepanthes N° 42» [W 0019614 / Rchb. Orch. 7615].

24. Lepanthes endresii Luer , Lindleyana 7 (2): 106 (1992). Type: Costa Rica. Cartago: “heights above El Tejar south of Cartago, June” ca 1867, A.R. Endres 509 [Holotype W 1889-24850]. Additional material associated with the type: Costa Rica. Endres’ coloured drawing of the flower, A.R. Endres 509 [W 0019608 / Rchb. Orch. 7612]; - Endres’ sketch of the inflorescence, the flower and flower details, A.R. Endres 509 [W 0019609 / Rchb. Orch. 38537]; - Endres’ sketch of the plant, the flower and flower details A.R. Endres 509 [W 0019610 / Rchb. Orch. 38529], Additional Endres’ material: Costa Rica. «La Paz, etc.», A.R. Endres s.n. [W 1889- 30718].-Uncertain locality, A.R. Endres 509 [W 1889-29962, W 1889-29970, W 1889- 29895, W 1889-142252]. The name is probably a synonym of Lepanthes erinacea Rchb.f.

25. Lepanthes falcifera Luer , Lindleyana 10 (3): 148 (1995). Type: Costa Rica, Cartago, “beyond the Laguna, Sitio Ortiz, Birris”, ca. 1867, A.R. Endres s.n. «N° 663 Gen. coll. Orch.» [Holotype W 1889-46789, isotypes W 1889- 142079, W 1889-142080, W 1997-8536. Additional material associated with the type: Endres’ drawing of the flower, A.R. Endres s.n. «N° 663 Gen. coll. Orch., sketch 53» [W 0019394 / Rchb. Orch. 38685]; - Luer’s drawing of the plant and flower, based on A.R. Endres s.n. «N° 663 Gen. coll. Orch.» [W 0019408].

26. Lepanthes fascinata L uer , Lindleyana 10 (3): 148 (1995). Type: Costa Rica, Cartago, “San Cristobal, June 5”, ca. 1867, A.R. Endres s.n. [Holotype W 1889-24844], Additional material associated with the type: Costa Rica. “P. of Cartago”, A.R. Endres s.n. [W 1889-24866, specimen on the right]; Luer’s drawing of the plant and flower, based on A.R. Endres s.n. (P. of Cartago) [W 0019473].

27. Lepanthes forcipifera L uer , Lindleyana 10 (3): 151 (1995). Type: Costa Rica, without locality, ca. 1867, A.R. Endres s.n. [Holotype W 0019390/Rchb. Orch. 4317]. Additional material associated with the type: Luer’s drawing of the plant and flower from the type [nmw 0020585].

28. Lepanthes fugiens Luer , Lindleyana 10 (3): 151 (1995).

Type: Costa Rica, without locality, ca. 1867, A.R. Endres s.n. [Holotype W 0019389/Rchb. Orch. 4372], Additional material associated with the type: Luer’s drawing of the plant and flower from the type [0020586].

29. Lepanthes gracillima L uer , Lindleyana 10 (3): 154 (1995). Type: Costa Rica. Puntarenas: Monteverde, epiphytic in tree in a pasture, 1500 m, 21. 3. 1995, C. Luer, J. Luer, J. Atwood, W. Rhodehamel, G. Campbell & G. Bar- bosa 17447 [Holotype MO]. Pupulin et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Endresio Wien, download in Costa unter www.biologiezentrum.at Rica lecti 279

Paratype: Costa Rica. Cartago: “heighs back of Carpintaria”, ca. 1867, A.R. Endres s.n. (24, according to the protologue) [W 1889-29961]. Additional Endres’ material: Endres’ coloured drawing of the flower and floral details, «Lepanthes gracillima», A.R. Endres s.n. «Lepanthes N° 24» [W 0019386 / Rchb. Orch. 7600]; - Endres’ drawings of the plant and two flowers, A.R. Endres s.n. «59» [W 0019387 / Rchb. Orch. 36208]; - Endres’ drawings of the plant, flower, and floral dis­ sections, A.R. Endres s.n. «see fig. p. 59» [W 0019388 / Rchb. Orch. 38534].

30. Lepanthes guardiana Endres ex Luer , Lindleyana 10 (3): 156 (1995). Type: Costa Rica without locality, ca. 1867, A.R. Endres 670 «sketch 299» [Holo­ type W 0019373 / Rchb. Orch. 4319]. Paratype: Costa Rica, Without locality, ca. 1867, A.R. Endres s.n. [W 0019374 / Rchb. Orch. 4374]. Additional Endres’ material: Endres’ coloured drawing of the flower, «Lepanthes guardiana», A.R. Endres s.n. [W 0019376 / Rchb. Orch. 7619]; - Endres’ drawings of the flower, floral dissections and description, A.R. Endres s.n. «N° 670 gen. coll. Orch., sk. N° 299» [W 0019375 / Rchb. Orch. 36217],

L u e r (1995) considered the etymology of L. guardiana of unknown derivation, but the name honours General Tomäs Miguel Guardia Gutierrez (1831-1882), who was Presi­ dent of Costa Rica from 1870 to 1876, during Endres’ joumey in the country, and again from 1877 to 1881.

31. Lepanthes hamulifera Luer , Lindleyana 10 (3): 156 (1995). Type: Costa Rica, San Jose, “Taburcia, Candelaria”, ca. 1867, A.R. Endres s.n. [Holotype W 0019369 / Rchb. Orch. 4396], Additional material associated with the type: Luer’s drawing of the plant and flower from the type [W 0020587].

32. Lepanthes limbellata Endres ex Luer , Lindleyana 10 (3): 159 (1995). Type: Costa Rica, without locality, ca. 1867, A.R. Endres s.n. (12, according to Luer’s annotation) [Holotype W 1889-0041431]. Additional Endres’ material: Endres’ drawings of the plant and flower, «Lepanthes lim- bata», A.R. Endres s.n. [W-0019345 / Rchb. Orch. 38640]; - Endres’ drawings of the plant and flower, A.R. Endres s.n. «64» [W 0019344 / Rchb. Orch. 36300]; - Endres’ drawing of the flower, A.R. Endres s.n. «Lepanthes N° 12» [W 0019346 / Rchb. Orch. 7628],

33. Lepanthes minutissima Endres ex Luer , Lindleyana 10 (3): 159 (1995). Type: Costa Rica, without locality, ca. 1867, «Effusa», A.R. Endres s.n. (10, according to the protologue) [Holotype W 1889-0087090]. Additional Endres’ material: Endres’ drawings of the plant and flower, with descrip­ tions, «p. 10», A.R. Endres s.n. [W 0019339 / Rchb. Orch. 38642]; - Endres’ drawings of the plant and flower, and coloured drawing of the flower A.R. Endres s.n. «59» [W 0019338 / Rchb. Orch. 36193]; - Endres’ coloured drawing of the flower, «Lepanthes microscopica», A.R. Endres s.n. «Lepanthes N° 10» [W 0019337 / Rchb. Orch. 7609].

In the type drawing of L. minutissima by Luer (1995), the shape of the is inverted 280 ©Naturhistorisches Museum Wien,Annalen download des unter Naturhistorischen www.biologiezentrum.at Museums in Wien, B l / >

longitudinally. Based on living specimens, they are not obtriangular-flabellate, but trian­ gulär in outline.

34. Lepanthes orbella Luer , Lindleyana 11: 64 (1996). Type: Costa Rica, without locality, ca. 1867, A.R. Endres s.n. «75» [Holotype W 0019334 / Rchb. Orch. 36259 (Endres’ drawing of the plant, flower and floral dis­ sections)]. Additional material associated with the type: Costa Rica. Endres’ sketch of the flower, A.R. Endres s.n. «Lepanthes N° 46» [W-0019333 / Rchb. Orch. 7636]; Endres’ sketches of the plant, flower and floral dissections, A.R. Endres s.n. (46) [W 0019332 / Rchb. Orch. 38496]. 35. Lepanthesparvilabia Luer, Lindleyana 10 (3): 162 (1995). Type: Costa Rica, without locality, ca. 1867, A.R. Endres s.n. [Holotype W 1889- 0142082], Additional material associated with the type: Luer’s drawing of the plant and flower from the type [W 0019418], Additional Endres’ material: Costa Rica, without locality, A.R. Endres s.n. [W 1889- 0030669].

36. Lepanthespoasensis L uer , Lindleyana 10 (3): 164 (1995). Type: Costa Rica, Heredia, «Volcän Poäs, front crest», ca. 1867, A.R. Endres s.n. (11, according to the protologue) [Holotype W 0019331 / Rchb. Orch. 38641 - Endres’ drawings of the plant and flower, with description «see fig. p. 54»]. Additional material associated with the type: Costa Rica. A.R. Endres s.n. (see fig. p. 54) [W 0019330 / Rchb. Orch. 4318]; - Endres’ drawing of the plant, flower and floral dissections, A.R. Endres s.n. «p. 54» [W 0019328 / Rchb. Orch. 38206]. Additional Endres’ material: Costa Rica. «Volc. Poäs, on the crest of front ridge - June / Volc. Barba, Desengano, Sept.», A.R. Endres s.n. [W 1889-0046786]; - Endres’ draw­ ing of the flower, A.R. Endres s.n. «Lepanthes N° 11» [W 0019328 / Rchb. Orch. 7629].

37. Lepanthespulcherrima Endres ex B ogarin & Pupulin , Harvard Pap. Bot. 15 (1): 118-121, f. 5A -E , 6 ,7 (2 0 1 0 ). Type: Costa Rica, Alajuela, San Ramon, Santiago, flnca of Jesus Salas Jimenez, mountains toward the towers of Berlin, 10°02’21“N 84°12’02“W, 1300 m, epi- phytic in old trees along coffee plantations, R Pupulin 7159, E. Salas-Pupulin & Jesus Salas, 14. 9. 2008 [Holotype JBL-spirit]. Paratypes: Costa Rica, «South West of San Ramon, rainy season», A.R. Endres s.n. «see fig. p. 66 centre / vol. 1869-70» [W 1889-41417]; - Endres’ drawing of the plant and colored drawing of the flower, A.R. Endres s.n. «66 / N.° 273» [W 0019020 / Rchb. Orch. 36239]; - Endres’ drawings of the plant, flower and floral analysis, with description, A.R. Endres s.n. «N.° 273 gen. coll. Orch. / see fig. p. 66 centre / vol. 69-70 / descr. p. 9 Lepanthes» [W 0019019 / Rchb. Orch. 38644]; - Endres’ drawings of the plant and flower, 1869, A.R. Endres s.n. «N.° 273» [W 0020098 / Rchb. Orch. 44244]; - Endres’ colored drawing of the flower, A.R- Endres s.n. «N.° 273 gen. coll. Orch., / Lepanthes N.° 9» [W 0019021 / Rchb. Orch. 7589]; - Endres’ descriptions, A. R. Endres s.n. «9 / Lepanthes pulcherrima / see fig. p. 66 centre vol. 69-70» [W 0019348 / Rchb. Orch. 38643]; -Without collect- P u p u l in et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Endresio Wien, download in Costa unter www.biologiezentrum.at Rica lecti 281

ing data, a plant confiscated by MINAE, flowered in cultivation at Lankester Botanical Garden, Accession No. JBL-01304, 14. 7. 2006, D. Bogarin 2942 [JBL- spirit]. - Panama. Bocas del Toro: Culebra, collected by Beni and cultivated by Steve & Marjorie Samer, N. 862, Boquete, Panama, flowered in cultivation 19. 12. 2008, D. Bogarin 5992 [JBL-spirit].

38. Lepanthespurpurea Luer , Lindleyana 10 (3): 164 (1995). Type: Costa Rica, Alajuela, «La Paz San Ramon», ca. 1867, A.R. Endres s.n. [Holotype W 1889-0030740 (Endres’ drawings of the plant and flower, with description «see fig. p. 54»]. Paratype: Costa Rica, without locality, A.R. Endres s.n. [W 1889-0087076]. Additional material associated with the type: Luer’s drawing of the plant and flower from the type [W 0021491]. Additional Endres’ material: Costa Rica, without locality, A.R. Endres s.n. [W 1889- 0041418].

39. Lepanthes reflexa Luer & H ermans , Lindleyana 10 (3): 167 (1995). Type: Costa Rica, Cartago, Cerro de la Muerte, 2750 m, flowered in cultivation, 22. 3. 1994, J. Hermans & C. Hermans 2746 [Holotype MO, isotype K]. Paratype: Costa Rica, Cartago, «South of Cartago, beyond the lime kilns of Doha Ana Clara Mayorga, rainy season», ca. 1867, A.R. Endres s.n. [not located],

40. Lepanthes selliana Endres ex Luer , Lindleyana 10 (3): 167 (1995). Type: Costa Rica, without locality, A.R. Endres s.n. (638, sketch 279, according to the protologue) [Holotype W 0019321 / Rchb. Orch. 38499 - (Endres’ drawings of the plant and flower, «N° 638 gen. coll. Orch., Lepanthes N.° 49, sk. 279»)]. Additional material associated with the type: Endres’ coloured drawing of the flower, «N° 638 gen. coll. Orch., Lepanthes N.° 49 / Lepanthes Selliana Endr.» [W 0019319/Rchb. Orch. 75965], Additional Endres’ material: Endres’ drawing of the plant and flower flower, «89» (W 0019320 / Rchb. Orch. 36257].

41. Lepanthes similis Luer , Lindleyana 11: 64 (1996). Type: Costa Rica, without locality, ca. 1867, A.R. Endres s.n. [Holotype W 1889- 30678 (Endres’ drawing of the plant, flower and floral dissections)]. Additional Endres’ material: Costa Rica, without locality, A.R. Endres s.n. (Orch. N° 140) [W 0019333 / Rchb. Orch. 7636]; - without locality, A.R. Endres s.n. [W 1889- 0029965]; - without locality, A.R. Endres s.n. [W 1889-0030674]. 42. Lepanthes umbonifera Endres ex Luer, Lindleyana 10 (3): 167 (1995). Type: Costa Rica, without locality, A.R. Endres s.n. «figure 67» [Holotype W 0019300 / Rchb. Orch. 36215 (Endres’ drawings of the plant, flower and floral dis­ sections)]. Additional Endres’ material: Endres’ drawings of the plant, flower and floral dissec­ tions, «see fig. p. 66» [W 0019417 / Rchb. Orch. 38652]; - Endres’ drawing of the plant, flower and floral dissections, « 66» [W 0019415 / Rchb. Orch. 36238]; - Endres’ draw­ ings of the flower, « Lepanthes N° 3» [W 0019414 / Rchb. Orch. 7639]; - Endres’ draw­ ing of the flower and floral dissections, «67» [W0019416 / Rchb. Orch. 36250]. 282 ©Naturhistorisches Museum Wien,Annalen download des unter Naturhistorischen www.biologiezentrum.at Museums in Wien, B ij->

43. amoena Endres & Rchb.f., Gard. Chron. 1872: 666 (1872). Type: Costa Rica, “Discovered by Mr. Endres in Costa Rica. Now flow ering (March 16) in the Hamburgh Botanic Garden”, A.R. Endres 2527 [Holotype W 0010806 / Rchb.Orch 44339, isotype AMES 26060 (barcode 2554947)]. Additional material associated with the type: Reichenbach’s drawing o f the type based on A.R. Endres 2527 [W 0010807 / Rchb. Orch. 44351]. Additional Endres’ material: Costa Rica, without locality, A.R. Endres s.n. [W 0019586 / Rchb. Orch. 3000]; - A.R. Endres s.n. [W 0019587 / Rchb. Orch. 2996]; - A.R. Endres s.n. [W 0019588 / Rchb. Orch. 45080]; - A.R. Endres s.n. [W 0019589 / Rchb. Orch 2991]; - A.R. Endres s.n. [W 0019588 / Rchb. Orch. 45080]; - A.R. Endres s.n. [W 0019585 / Rchb. Orch. 2997]; - A.R. Endres s.n. [W 0019590 / Rchb. Orch. 2998]; - A.R. Endres s.n. [W 0019584 / Rchb. Orch. 45052].

44. Lockhartia grandibractea K r a e n z l . , Pflanzenr. 4, 50 (Heft 83): 15 (1923). Type: Costa Rica, “Subäquatoriale andine Provinz. Costa Rica”, A.R. Endres s.n. [holotype supposedly at W, not located]. Additional Endres’ material: Costa Rica. Endres’ drawings of the column, pollinarium and anther, A.R. Endres s.n. [W 0019581 / Rchb. Orch. 33709]; «Cartago», Endres’ description, A.R. Endres s.n. «N° 629 of gen. coll.» [W 0019580 / Rchb. Orch. 33713], Mario A. Blanco (correction label, 2006) noted that, notwithstanding the indication of Endres as the collector, as stated by Kränzlin in the protologue, no specimens of such collection have been found at W. However, among the specimens annotated by Kränzlin as type material, there are two Costa Rican collections made by Hübsch in 1882-1885 [W 0021971 / Rchb. Orch. 6159, W 0021972 / Rchb. Orch. 6139], which Blanco pro- posed for lectotypification purposes, waiting for formal designation.

A close relative of Lockhartia amoena, and frequently considered synonymous with it (i.e., Atwood & M ora de Retana 1999, H e l l e r 2001), L. grandibractea can be distin­ guished by the very reduced, almost rudimentary column wings, compared with the broadly dolabriform wings of L. amoena.

45. Lockhartia hercodonta R c h b .f. ex K r a e n z l . , Pflanzenr. 4, 50 (Heft 83): 8, t. 2A (1923). Type: Costa Rica, “Subäquatoriale andine Provinz. Costa Rica”, A.R. Endres s.n. «Orch. N° 167» [Holotype W 0010813 / Rchb. Orch. 18012 (with Reichenbach’s sketch of the type and the note «n. sp.», isotypes W 0010815 / Rchb. Orch. 2993, W 0010816 / Rchb. Orch. 2994, W 0010817 / Rchb. Orch. 2985, W 0010814 / Rchb. Orch. 2999], Additional material associated with the type: Costa Rica. Endres’ drawing of the rostellum and pollinarium, based on A.R. Endres s.n. «N° 167» [W 0019575 / Rchb. Orch. 33683]. Additional Endres’ material: Costa Rica. Endres’ drawings of the flower and floral dis­ sections, A.R. Endres s.n. [W 0019576 / Rchb. Orch. 33704; - Endres’ drawings of the plant, flower and lip, A.R. Endres s.n. [W 0019577 / Rchb. Orch. 33708; - Endres’ definitive drawing and engraved copy of the drawing intended for Xenia Orchidacea, A.R. Endres s.n. [W 0019579 / Rchb. Orch. 338; - flowers; plate intended for Xenia P u p u l in et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Endresio Wien, download in Costa unter www.biologiezentrum.at Rica lecti 283

Orchidacea, with Reichenbach’s note «hercodonta», A.R. Endres s.n. [W 0019578 / Rchb. Orch. 44372], 46. Lockhartia odontochila Kraenzl., Pflanzenr. 4, 50 (Heft 83): 17 (1923). Type: Costa Rica, “Subäquatoriale andine Provinz. Costa Rica”, A.R. Endres s.n. [Holotype W 1889-58236 (annotated by Kränzlin, 1917)]. 47. Lycaste dowiana E ndres ex R chb.f., Gard. Chron., n.s., 2: 194 (1874). Type: Costa Rica. “Lycaste dowiana was discovered in Costa Rica by the late lamented Warscewicz. The flowers. however, were not well preserved .... The plant was also found by the late collector Zahn. It is but now, however, that I am able to describe it with full satisfaction, from two good living plants, obtained from Mr. Endres, and beginning to flower in the Hamburgh Botanic Garden” Holotype: 1868, A.R. Endres s.n. «N° 89» [W 0019555 / Rchb. Orch. 44616 (anno­ tated by Reichenbach, flowers with Endres drawings of the flower and descrip­ tions)]. Isotypes: Flowers, A.R. Endres s.n. «N° 89 of gen. coll. Orch.» [W 0019552 / Rchb. Orch. 13596]; - Flowers, A.R. Endres s.n. «N° 89 of gen. coll. Orch.» [W 0019551 / Rchb. Orch. 13611]. Additional material associated with the type: Endres’ drawings and descriptions, A.R. Endres s.n. «N° 89 of gen. coll. Orch.» [W 0019554 / Rchb. Orch. 35945]. Other material: preparatory drawings intended for Xenia Orchidacea, based on Endres’ drawings [W 0019556 / Rchb. Orch. 23354].

48. Masdevallia anaristella K raenzl ., Repert. Spec. Nov. Regni Veg. 17 (19-30): 417^118 (1921). Type: “Costa Rica. Ohne genaueren Standort”, 1867, A. R. Endres s.n. [Holotype W 1889-0037485, isotypes W 1889-0037486, W 1889-0037480, AMES 41297].

The name is a synonym of Barbosella dolichorhiza Schltr .

49. Masdevallia anura Kraenzl ., Repert. Spec. Nov. Regni Veg. 17 (19-30): 433—434 (1921). Type: “Costa Rica. Ohne genaueren Standort”, A.R. Endres s.n. «N.° 21 gen. coll. Orch. / Masdevallia mucronata» [Holotype 0019518 / Rchb. Orch. 14510 (sterile)].

The name is probably a synonym of Masdevallia molossoides Kraenzl .

50. Masdevallia aperta K raenzl ., Repert. Spec. Nov. Regni Veg. 17 (19-30): 430-431 (1921). Type: “Costa Rica. Am Rio Pascita grande” («Rio Paquita, above the S. Marcos Bridge / Nov.ber - Dec.ber»), A.R. Endres s.n. (652 in the protologue) [Holotype W 1889-0030752, isotypes W 1889-0087087, AMES 41994 (with Kränzlin’s draw­ ing)]. Additional Endres’ material: Costa Rica, without locality. Endres’ drawings of the flow­ ered plant and floral details, A.R. Endres s.n. [W 0021291 / Rchb. Orch. 38513]; - «Cuesta from the saw mill of Birris up to the Sitio Ortiz / Aug.t-Dec.ber», «Birris, cuesta leading up to Sitio Ortiz / also near Pejivalle River», A.R. Endres s.n. «N.° 653 gen. coll. Orch. / M asdevallia N.° 22» [W 0021290 / Rchb. Orch. 38547]. 284 ©Naturhistorisches Museum Wien,Annalen download des unter Naturhistorischen www.biologiezentrum.at Museums in Wien, B / / >

The name is a synonym of Pleurothallis tripterantha Rchb.f. = Humboltia tripterantha (Rchb.f.) K u n t z e (1891) = Anthereon tripteranthus Rchb.f.) P r id g e o n & M.W.Chasi (2001) = Pabstiella tripterantha Rchb.f.) F.B a r r o s (2002) = Specklinia tripteranthu (Rchb.f.) L u e r (2004). 51. Masdevallia carpophora Kraenzl., Repert. Spec. Nov. Regni Veg. 17 (19-30)- 427 (1921). Type: “Costa Rica, Candelaria, el Rio Parrita”, «Taburcia, Candelaria, Dec.r / also along Rio Parrita / Dota», A.R. Endres s.n. [Holotype W 0020185 / Rchb. Orch 14507].

The name is a synonym of Pleurothallis tripterantha R chb .f. (1854) = see above. 52. Masdevallia funebris E ndres ex K ra en zl., Repert. Spec. Nov. Regni Veg. 17: 435 (1921). Type: “Costa Rica, ohne genaueren Standort” «Tablazo. i.s. heights / S.th of Cartago», A.R. Endres 501 [Holotype W 0019495 / Rchb. Orch. 39108 (three plants on the left), isotype W 1889-0142279 (sterile)]. Additional material associated with the type: Costa Rica. «Birris», A.R. Endres 50] [W 0019494 / Rchb. Orch. 14533]; - «Masdevallia funebris / (probably just a vari- ety of M. Reichenbachiana)», flowers, A.R. Endres s.n. «N° 645 of gen. coll. Orch.» [W 1889-0046840, W 1889-0046843); - Endres’ drawing of the flower, floral dis- section and description, A.R. Endres s.n. «N° 645 of gen. coll. Orch., Masdevallia N° 14, sk. 196» [W 1889-0046840],

The name is a synonym of Masdevallia reichenbachiana Endres ex R chb .f. (1875) 53. Masdevallia gorgo R chb.f. ex K ra en zl., Repert. Spec. Nov. Regni Veg. Beih. 34: 136-137(1925). Type: “Costa Rica: Cartago pr. San Ramon. Fl. Julio et Augusto”, A.R. Endres 134 «N.° 137 gen. coll. Orch» [Holotype W 0019178 / Rchb. Orch. 14523]. Additional material associated with the type: Costa Rica. A.R. Endres s.n. «N.° 137 gen. coll. Orch.» [W 0019424 / Rchb. Orch. 14520]; - Endres’ pencil plate of the plant, flower, and floral details, A.R. Endres s.n. «N.° 137 / 51» [W 0019184 / Rchb. Orch. 36046]; - Endres’ sketches of flower and floral analysis, with description, A.R. Endres s.n. «N.° 137 gen. coll. Orch. / see p. 51 vol. 1867-68» [W 0019183 / Rchb. Orch. 36175].

The name is likely a synonym ofDracula astuta (R chb .f.) Luer = Masdevallia astuta RcHB.f. (1886). 54. Masdevallia molossoides Kraenzl., Repert. Spec. Nov. Regni Veg. 17: 416-417 (1921). Type: “Costa Rica, ohne genauen Standort in 1200 m ü. D. M.”, «rarissima!, 3500”», 1867, A.R. Endres s.n. «Orch. N.° 21» [Holotype W 0019504 / Rchb. Orch. 15008 (with Endres’ drawing of the flowered plant and Reichenbach’s analysis of the flower), isotype W 0019503 / Rchb. Orch. 14539]. Additional material associated with the type: Endres’ drawings of the flower and floral dissections, A.R. Endres s.n. «N.° 21 gen. coll. Orch / Masdev. N.° 7» [W 0019501 / Rchb. Orch. 38512, W 0019500 / Rchb. Orch. 38511]. - Costa Rica. pu pul in et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Endresio Wien, download in Costa unter www.biologiezentrum.at Rica lecti 285

Jieiiilunbav.: Herb. Orchid. N r g (3 ( ) 5 5

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NATURHIS7 URISCHES MUSEUM WIEN W 0019476-

£ Copyright reserved

Fig. 3: Lectotype of Masdevallia pygmaea K r a e n z l . [W 0019476 / Rchb. Orch. 36055], 286 ©Naturhistorisches Museum Wien,Annalen download des unter Naturhistorischen www.biologiezentrum.at Museums in Wien B //i

«Queb.da Verde / March» A.R. Endres s.n. [W 1889-467500], identified by Kran- zlin as M. molossoides.

55. Masdevallia pygmaea K raenzl ., Repert. Spec. Nov. Regni Veg. Beih. 34- 92 (1925). = Diodonopsis pygmaea (K raenzl .) Pridgeon & M.W. C hase (2001). Type: “Costa Rica”, A.R. Endres 289 [to be found at W, not located]. Lectotype (hic designatus): Costa Rica, without specific locality, A.R. Endres s.n. «N.° 289 / 64» [W 0019476 / Rchb. Orch. 36055 (Endres’ drawings of plant, flowers and floral dis­ sections)]. - Fig. 3. Additional material associated with the type: Costa Rica, without specific locality, tracings at actual size of the original flowered plants sent by Endres, prepared for Prof. Ames from Reichenbach’s herbarium, A.R. Endres s.n. [AMES 41307 / bar- code 00101298],

56. Masdevallia reichenbachiana Endres ex R chb .f., Gard. Chron., n.s., 4 (87): 257 (1875). Type: Costa Rica. “The beautiful plant now before me was discovered in Costa Rica by my late friend Endres, and was named by him”, A.R. Endres II 257 [Holotype W 0019495 / Rchb. Orch. 39108, plant on the upper right],

57. Masdevallia superflua Kraenzl ., Repert. Spec. Nov. Regni Veg. 17: 414 (1921). Type: “Costa Rica, ohne genauen Standort”, A.R. Endres s.n. [Holotype W 1889- 0076668], The name «superflua», with interrogation mark, but not in Kränzlin’s handwriting, is also annotated on a sheet that includes Endres’ accurate drawings of the flowered plant and floral analysis, and a botanical description [W 0019496 / Rchb. Orch. 38546]. The plant on which the drawings were made was collected «above the lime kilns near Tobosi, Cartago», and Endres annotated it as his sketch «N.° 256». According to Tropi- cos, the holotype of M. superflua (Costa Rica: Cartago: Tobosi, A. Endres 256) is kept at BR, but we have not found any specimens of Endres among those searchable through the online database of the Herbarium at the National Botanic Garden of Belgium. Another collection by Endres kept at W [W 1889-0046765] was determined by Kränzlin as the type of a new species with the intended name “Masdevallia otochila”, finally never published.

The name is a synonym ofMasdevallia striatella Rchb .f. (1886) (= Reichanta striatella R chb .f. ex Luer 2006). 58. Masdevallia triaristella Rchb.f., Gard. Chron., n.s., 6 (138): 226 (1876). Type: Costa Rica. “The discovery is due to the late M. Endres; the satisfaction of having flowered it for the first time in Europe is with Messrs. Veitch”, A.R. Endres s.n. «N° 260» [Holotype W 0021123 / Rchb. Orch. 44435 (plant, Endres’ drawing of the plant and flower, description)]. Additional material associated with the type: Endres’ drawing of the dissected flower, A.R. Endres s.n. «N° 260 gen. coll. Orch., Masdevallia N° 11» [W 0021328 / Rchb. Orch. 38557]; - Endres’ drawing of plant, flower, and floral dissection, A.R. Endres s.n. «N° 260 gen. coll. Orch., Masdevallia N° 11» [W 0021124 / Rchb. Orch. 38558]; - A.R. Endres s.n. «N°260» [W 1889-0142223]. Piü’ULiN et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Endresio Wien, download in Costaunter www.biologiezentrum.at Rica lecti 287

Additional Endres’ material: «potrero de Guayabo beyond Turrialba», flowers, A.R. Endres s.n. «N° 260 gen. coll. Orch., Masdevallia N° 11» [W 1889-0046717]; - Endres’ descriptions («San Ramon, Camino de San Carlos / Cataratas / La Paz»), A.R. Endres s n> «N° 260 gen. coll. Orch., Masdevallia N° 11» [W 0021329 / Rchb. Orch. 38557]. 59 Masdevallia zahlbruckneri K r a e n z l., Repert. Spec. Nov. Regni Veg. 17: 413 ’ (1921). Type: “Costa Rica, ohne genauen Standort”, 1870, A.R. Endres s.n. «N.° 463» [Holotype W 0019499 / Rchb. Orch. 15007 (with Endres’ drawings of the flowered plant and floral analysis, and Reichenbach’s sketch of the flower)]. Additional material associated with the type: Costa Rica. Endres’ drawings of plant, flowers and floral dissections, with description, A.R. Endres s.n. «N.° 463 gen. coll. Orch. / Masdevallia N.° 4» [W 0019498 / Rchb. Orch. 38508]; - Endres’ pencil plate of flowered plant and floral analysis, A.R. Endres s.n. «N.° 463 / 56» [W 0019497 / Rchb. Orch. 36060].

60. Maxillaria caespitifica R chb .f., Linnaea 41 (“ 1876”): 73 (1877). = Maxillariella caespitifica (R chb .f.) M .A.B lanco & C arnevali (2007). Type: “Costa Rica. Endres! (Viva in horto Hamburgensi)”, A.R. Endres s.n. [Holo­ type W 0018755 / Rchb. Orch. 44573 (with Reichenbach’s description and draw­ ings)].

61. Maxillaria endresii R chb .f., Gard. Chron., n.s., 25 (648): 680 (1886). Type: Costa Rica. “This flowered in 1870 in the Hamburgh Botanic Garden, having been sent alive with Sievekingia by the late Mr. Endres, to whose memory it is now dedicated. I made a sketch in colours, but never published it. Now it has reappeared, also from Costa Rica, in the establishment of Messrs. Hugh Low & Son, whence it has been kindly sent me by Mr. Edward Low” [Holotype W 0027464 (with Reichenbach’s drawings and description, May 1886, and additional flowers from plant cultivated at Hamburg Botanical Garden, E. Low s.n.), isotype W 0027463].

62. Maxillaria nasalis R chb .f., Refug. Bot. 2: t. 103 (1872), nom. illeg. superfl. Type: Costa Rica. “The original discoverer of this plant appears to be the late unfor- tunate L. Schlim, Director Linden’s half-brother, who found it near Lasita, in New Grenada; it is his 1752! Some time ago, Mr. Tucker, Mr. S. Low’s traveller, found it in Costa Rica, as did also Mr. Endres; finally it appeared in the Saundersian collec­ tion” Endres’ materials associated with the type: Costa Rica, without specific locality, A.R. Endres 266 [W 0018742 / Rchb. Orch. 44584, W 0018618 / Rchb. Orch. 44048, W 0010838 / Rchb. Orch. 45047].

The name is a sphalma for Maxillaria nasuta R chb .f. [type: Colombia, Lajitas, L.J. Schlim 1752 [W, K] = Nitidobulbon nasutum (R chb .f.) I. Ojeda & C arnevali .

63. Maxillaria reichenheimiana Endres & R chb .f., Gard. Chron. 1871: 1678 (1871). Type: “It was discovered by M. Endres, in 1867, at an altitude of 5500 feet, in Costa Rica”, A.R. Endres s.n. [Holotype mounted in 6 sheets: W 0018735 / Rchb. Orch. 44577 (specimen with Endres’ drawings of the plant, flower, and flower details, «N° 8»); - W 001873 / Rchb. Orch. 18355 («N° 8»); - W 0018737 / Rchb. Orch. 18361 («N° 8 gen. coll. Orch.»); -W 0018738 / Rchb. Orch. 18350 (« 8»); - W 288 ©Naturhistorisches Museum Wien,Annalen download des unter Natwhistorischen www.biologiezentrum.at Museums in Wien, B / / ■>

0018739 / Rchb. Orch. 33316 (Endres’ drawings of the plant, flower, and flower details, «Maxillaria Reichenheimiana»); - W 0019187 (Endres’ descriptions, «N° 8 gen. coll. Orch.»)]. Additional Endres’ material: Costa Rica, without locality, A.R. Endres s.n. «210» [W 0018734 / Rchb. Orch. 44626]; - A.R. Endres s.n. «N° 260» [W 1889-0142223],

64. Miltonia endresii G. N icholson , 111. Dict. Gard. 2: 368 (1886). Type: “Senor Endres was the first to introduce the species in a living state to this country”, A.R. Endres s.n. [Holotype probably not preserved] Neotype (hic designatus): Costa Rica, without specific locality, A.R. Endres s.n «Orch. N.° 200» [W 0019871 / Rchb. Orch. 18016], - Fig. 4. Additional material associated with the type: Costa Rica, without specific locality. Endres’ drawings of the column and lip, and pollinarium, A.R. Endres s.n. «N.° 200» [W 0019909 / Rchb. Orch. 36907], Additional Endres’ materials: Costa Rica, without specific locality, A.R. Endres s.n. ][W 0019868 / Rchb. Orch. 13792, W 0019870 / Rchb. Orch. 18015]; - Endres’ partially coloured drawing of the plant with flowers, «? sp. nov.», 5000 ft., 1869, A.R. Endres s.n. [W 0019869 / Rchb. Orch. 18].

The name is a synonym of warscewiczii (R chb .f.) Garay & D unst . (1976) = Odontoglossum warszewiczii R chb .f. (1852).

65. Oncidium angustisepalum K raenzl ., Pflanzenr. 4, 50 (Heft 80): 203, f. 18D (1922). Type: “Subäquatoriale andine Provinz. Costa Rica”, A.R. Endres s.n. [Holotype W 0019960/Rchb. Orch. 13181],

Atwood & M ora de R etana (1999) treated this taxon as a synonym of Oncidium klotzschianum R chb .f. It may well be that O. angustisepalum just represents an extreme form of that reputedly variable species.

66. Oncidium asparagoides K raenzl ., Pflanzenr. 4, 50 (Heft 80): 175 (1922). Type: “Subäquatoriale andine Provinz. Costa Rica, ohne genaueren Standort”, 1867, A.R. Endres 127 [Holotype W 0019895 / Rchb. Orch. 18386, isotypes W 0019923 / Rchb. Orch. 13204, W 0019922 / Rchb. Orch. 13225].

The name is a synomym ofOncidium biyolophotum R chb .f. (1871).

67. Oncidium calyptostalix Kraenzl ., Pflanzenr. 4, 50 (Heft 50): 260 (1922). Type: “Subäquatoriale andine Provinz. Costa Rica”, A.R. Endres s.n. [Holotype W 0019940/Rchb. Orch. 13201],

The name is a synonym of Oncidium obiyzatoides Kraenzl . (1922).

68 . Oncidium castaneum R chb .f. ex Kraenzl ., Pflanzenr. 4, 50 (Heft 80): 278 (1922). Syntypes: “Subäquatoriale andine Provinz. Costa Rica”, A.R. Endres s.n.; “Sub­ äquatoriale andine Provinz. Chiriqui”, R. Pfau s.n. Lectotype (hic designatus): Costa Rica, without specific locality, 1867, A.R. Endres s.n. «N.° 108 Oncd.m» [W 0019929 / Rchb. Orch. 18381]. - Fig. 5. Additional Endres’ materials: Costa Rica, without specific locality, A.R. Endres s.n. [W ©Naturhistorisches Museum Wien, download unter www.biologiezentrum.at 2 8 9 P u p u l in et al.: Typi Orchidacearum Augusto R. Endresio in Costa Rica lecti

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Fig. 4: Neotype of Miltonia endresii G. N i c h o l s o n [W 0019871 / Rchb. Orch. 18016], 290 ©Naturhistorisches Museum Wien,Annalen download unter des www.biologiezentrum.at Naturhistorischen Museums in Wien ß / / •>

0019925 / Rchb. Orch. 45515]; - A.R. Endres s.n. «2399» [W 0019926 / Rchb Orch 45513]; - A.R. Endres s.n. [W 0019927 / Rchb. Orch. 45514]; - A.R. Endres s n «2597» [W 0019928 / Rchb. Orch. 45512], The nam e Oncidium castaneum was first published as a nomen nudum by Lindi-n (1882) in his account on orchid plants newly introduced or first offered in commercc where under number 651 he makes reference to a letter by Reichenbach and to his cata- logue of Colombian plants for 1869. Among the materials of the Reichenbach orchid herbarium at W is kept a manuscript description with two drawings by Reichenbach himself. The upper one, coloured, has no provenance data. The other, which includes a manuscript description by Reichenbach, was prepared on the basis of a plant from Chiriqui, seemingly obtained through Linden. We have not found, however, any speci­ mens referable to these drawings. The name of the species is also annotated in Reichen­ bach’s handwriting on two specimens collected in Costa Rica by Endres around 1867 [W 0019929 / Rchb. Orch. 18381, W 0019925 / Rchb. Orch. 45515], The first of them, being a complete plant with inflorescence and flowers, is selected as the lectotype. Two other sheets include Endres’ material, one numbered «2597» and the other «2399»; they were both annotated as “Typus” by Kränzlin, but they do not have Reichenbach’s anno- tations.

The name is a synonym of Oncidium dichromaticum R chb .f. (1855).

69. Oncidium cheirophoroides K raenzl ., Pflanzenr. 4. 50 (Heft 80): 199, f. 17 (1922). Type: “Subäquatoriale andine Provinz. Costa Rica”, A.R. Endres 127 [Holotype W 0019924 / Rchb. Orch. 45055].

The name is a synomym of Oncidium luteum R olfe (1893). 70. Oncidium chelidonizon Kraenzl., Pflanzenr. 4, 50 (Heft 50): 217 (1922). Type: “Zentralamerikanische Provinz. Costa Rica, San Ramon längst des Rio Grande”, A.R. Endres s.n. [Holotype W 0019933 / Rchb. Orch. 13189]. Additional material associated with the type: Endres’ drawing of the flower and floral dissections, with description, A.R. Endres s.n. «N° 608 gen. coll.» [W 0019889 / Rchb. Orch. 33702]; - Endres’ drawing of flower, floral dissection, and description, «San Ramon, along Rio Grande» A.R. Endres s.n. «N° 608 gen. coll.» [W 0019934 / Rchb. Orch. 38509], Endres’ original label on the type sheet bears no locality data. The locality given in the protologue was apparently presumed from Endres’ sketches and descriptions, mentioned on other sheets.

The name is a synonym of ochmatochila (R chb .f.) N.H. W illiams & M.W. C hase (2001).

71. Oncidium dielsianum Kraenzl ., Pflanzenr. 4, 50 (Heft 80): 197, t. 198F (1922). Syntypes: “Subäquatoriale andine Provinz. Costa Rica”, A.R. Endres s.n.; “Sub­ äquatoriale andine Provinz. Costa Rica”, J. Carmiol s.n. Lectotype (hic designatus): sheet annotated by Kränzlin as “Typus”, Costa Rica, A.R. Endres s.n. «Orch. N° 306» [W 0020048 / Rchb. Orch. 18389 (Fig. 6); isolec- totypes W 0020049 / Rchb. Orch. 13211, W 0020047 / Rchb. Orch. 13226]. NATUHHi'KI C ft ft C NATUHHi'KI W W 001992H • • WEN MUSEUM HERB. MÜS. I'ALAT. VIN’DOB. Herb. Musei Herb. Palat. Vindob. Reichephach; Herb. Orchid. Nr. Orchid. Herb. Reichephach; Costa rica,Costa Rchb.f. ex Kraenzl. 0019929 [W / Rchb. Orch. 18381]. ©Naturhistorisches Museum Wien, download unter www.biologiezentrum.at Wien, download Museum ©Naturhistorisches Oncidium castaneum

Copyright reserved et al.: Typi Orchidacearum Augusto R. Endresio Rica Costa Augusto lecti in R. Endresio Typi Orchidacearum al.: et upulin P Fig. 5: Lectotype ofFig. 5: 292 ©Naturhistorisches Museum Wien,Annalen download unter des www.biologiezentrum.at Naturhistorischen Museums in Wien, B j p

Additional Endres’ material: Costa Rica, without locality, 1868, A.R. Endres s.n «M» 75» [W 0020046 / Rchb. Orch. 13202],

The name is a synomym of Oncidium cheirophorum R chb .f. (1852).

72. Oncidium globuliferum var. costaricense R chb .f., Gard. Chron. 1871: 1678 (1871). Syntypes: “It is a new discovery that it occurs also in Costa Rica, where it has been found both by Endres [A.R. Endres s.n.] and by the late M. Zahn, who sent it to Messrs. Veitch, with whom it has just flowered [Zahn s.n., ex hört. Veitch]” Lectotype (hic designatus): Costa Rica. «All around San Ramon, January-February, Oncidium scandens», A.R. Endres s.n. [W 0020057 / Rchb. Orch. 13212] - Fig. 7. Additional Endres’ material: Costa Rica, without locality data, «Oncidium scandens», A.R. Endres s.n. [W 0020058 / Rchb. Orch. 13213]; - Endres’ drawings of the flower and floral dissections, «Oncid. scandens», A.R. Endres s.n. [W 0020060 / Rchb. Orch. 32393]; - Endres’ description, «Oncidium scandens», A.R. Endres s.n. [W 0020059 / Rchb. Orch. 33529], Even though the drawings of the flower and the floral dissections made by Endres, as well as the detailed description he prepared, have no labels indicating the collecting locality, they were ostensibly prepared from the same collection sine numero from the vicinity of San Ramon de Alajuela. The only sheet with locality data is selected as the lectotype, but the description and illustrations probably refer to the same gathering. Among the sheets of Reichenbach’s herbarium, there is a series of other specimens col­ lected by Endres, under the number 256 of his general collection of orchids. One of these is annotated in Reichenbach’s handwriting as «globuliferum», and it is likely he did not refer them to his new variety: Costa Rica, without known provenence, A.R. Endres s.n. «256» [W 0020056 / Rchb. Orch. 18387, W 0020052 / Rchb. Orch. 44070]; - A.R. Endres s.n. «Orch. N° 256» [W 0020055 / Rchb. Orch. 13218]; - A.R. Endres s.n. «256» [W 0020056 / Rchb. Orch. 18387]; - A.R. Endres s.n. «N° 256 of gen. coll. Orch.» [W 0020054 / Rchb. Orch. 13168]; - A.R. Endres s.n. «N°. 256 of gen. coll.» [W 0020053 / Rchb. Orch. 13190].

The name is a synonym of globuliferum (Kunth ) N.H.W illiams & M.W. C hase (2001).

73. Oncidium henrici-gustavi Kraenzl ., Pflanzenr. 4, 50 (Heft 80): 187 (1922), non Schltr . (1923). Type: “Subäquatoriale andine Provinz. Costa Rica”, A.R. Endres s.n. «Orch. N.° 292» [Holotype W 0020062 / Rchb. Orch. 18391].

The name is a synomym of Oncidium bracteatum Warsz . ex R chb .f. (1852). 74. Oncidium macrorhynchum K raenzl., Pflanzenr. 4, 50 (Heft 80): 200, t. 198J (1922). Type: “Subäquatoriale andine Provinz. Costa Rica, San Juan de Candelaria”, A.R. Endres s.n. [Holotype W 0020086 7 Rchb. Orch. 13191]. Additional material associated with the type: Costa Rica, without specific locality, annotated by Kränzlin as “Typus”, A.R. Endres s.n. «Orch. N° 176» [W 0020087 / Rchb. Orch. 13233].

The name is a synomym ofOncidium cheirophorum R chb .f. (1852). ©Naturhistorisches Museum Wien, download unter www.biologiezentrum.at P u p u l i n et al.: Typi Orchidacearum Augusto R. Endresio in Costa Rica lecti 293

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Fig. 6: Lectotype of Oncidium dielsianum K r a e n z l . [W 0020048 / Rchb. Orch. 18389], 294 ©Naturhistorisches Museum Wien,Annalen download unter des www.biologiezentrum.at Naturhistorischen Museums in Wien, B // ->

75. Oncidium microphyton Kraenzl ., Pflanzenr. 4, 50 (Heft 80): 204 (1922). Type: “Subäquatoriale andine Provinz. Costa Rica”, A.R. Endres s.n. [Holotype W 0020030 / Rchb. Orch. 13206 (sterile)].

The name is a synomym of Oncidium bryolophotum R chb .f. (1871).

76. Oncidium obryzatoides Kraenzl ., Pflanzenr. 4,. 50 (Heft 80): 240 (1922). Type: “Subäquatoriale andine Provinz. Costa Rica”, A.R. Endres s.n. [Holotype W 0019980 / Rchb. Orch. 13182, isotype W 0019982 / Rchb. Orch. 13230 (A.R. Endres s.n. «Orch. N.° 285»)]. Additional Endres’ material: Costa Rica, without locality data, «325», A.R. Endres s n [W 0019981 / Rchb. Orch. 13227].

77. Oncidium rechingerianum Kraenzl ., Pflanzenr. 4, 50 (Heft 80): 202, f. 18C, a-c (1922). Type: “Subäquatoriale andine Provinz. Costa Rica”, A.R. Endres s.n. Lectotype (hic designatus): Costa Rica, without specific locality, «See the sp. fig. at p. 108 / Fl. yellow, blotched with chocolate brown», A.R. Endres s.n. [W 0020077 / Rchb. Orch. 45066] - Fig. 8. Additional material associated with the type: Costa Rica, without specific locality, annotated by Kränzlin as “Typus”, «See the sp. fig. at p. 108 / Fl. white, spotted with bright carmine», A.R. Endres s.n. [W 0020078 / Rchb. Orch. 45067] / nmw): - A.R. Endres s.n. [W 0020076 / Rchb. Orch. 45084]. Three sheets at W are annotated by Kränzlin as “Typus” Two of them belong to the typ- ical form of O. dichromaticum, with yellow sepals blotched with chestnut brown and a yellow lip, while the other is the form lacking flavonoids, with white sepals blotched with purple, and white lip. Apparently, Kränzlin did not select any of them as the holo­ type, and a lectotype has to be designated due to the differences between the specimens and Endres’ annotations. The specimen selected as the lectotype belongs to the more common, yellow form, and has a label in Endres’ handwriting.

The name is a synonym of O. dichromaticum R chb .f. (1855). 78. Oncidium sclerophyllum Kraenzl., Pflanzenr. 4, 50 (Heft 80): 204 (1922). Type: “Subäquatoriale andine Provinz. Costa Rica: San Ramon”, April, «see analy­ sis p. 108. upper outer vol. 1869-1870», A.R. Endres s.n. [Holotype W 0020074 / Rchb. Orch. 45069, Isotype W 0020074 / Rchb. Orch. 45069 («see sk. p. 108 upper outer angle»)].

Atwood & M ora de R etana (1999) treated this taxon as a synonym of Oncidium klotzschianum RCHB.f. (1852).

79. Oncidium stenobulbon Kraenzl ., Pflanzenr. 4, 50 (Heft 80): 281 (1922). Type: “Costa Rica: am Rio Jesus” («San Ramon towards Rio Jesus / May-June»), A.R. Endres s.n. [Holotype W 0020073 / Rchb. Orch. 45062]. Additional material associated with the type: Endres’ description and drawing of the flower and floral dissection, A.R. Endres s.n. [W 0020085 / Rchb. Orch. 33508]; - Endres’ drawing of the flower and floral dissection, improved copy [W 0020084 / Rchb. Orch. 33729]. P u p u l in et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Wien, Endresio download in unter Costa www.biologiezentrum.at Rica lecti 2 9 5

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Fig. 7: Lectotype of Oncidium globuliferum var. costaricense R c h b . f. [W 20057 / Rchb. Orch. 13212], 296 ©Naturhistorisches Museum Wien,Annalen download unter des www.biologiezentrum.at Naturhistorischen Museums in Wien, B ] p

Additional Endres’ material: Without locality, plant with flowers, AR. Endres s n [ty 0020083 / Rchb. Orch. 13183],

80. Oncidium tetraskelidion Kraenzl ., Pflanzenr. 4, 50 (Heft 80): 259, f. 22C (1922) Type: “Subäquatoriale andine Provinz. Costa Rica”, A.R. Endres s.n. «Orch N 0 285» [Holotype W 0020026 / Rchb. Orch. 18383], The name is a synonym of Oncidium obryzatoides Kraenzl. (1922).

81. Ornithidium strumatum Endres & R chb .f., Gard. Chron., n.s., 2 (51): 772 (1874). Type: Costa Rica. “It has just flowered (November) in Hamburgh Botanic Garden. One of the very numerous discoveries of Senor Endres in Costa Rica”, 1867, A.R. Endres s.n. (N° 281 gen. coll. Orch.) [Holotype W 0018730 / Rchb. Orch. 14353 and W 0018729 / Rchb. Orch. 18044 (with Endres drawings), isotype AMES 45387 (barcode 5165995)]. Additional Endres’ material: Costa Rica, Endres’ drawing of the flower and floral dis­ section, A.R. Endres 90 [W 0018728 / Rchb. Orch. 36586].

The name is the basionym of Camaridium strumatum (Endres & R ch b .f.) M.A. B lanco (2007).

82. Pleurothallis endotrachys R chb .f., Linnaea 41: 95 (1876) Annot.: non F. Lehm. & Kraenzl . 1899, nom. illeg. (= Pleurothallis lehmanniana Schltr .). Type: “Costa Rica. Endres”, A.R. Endres s.n. «N° 92» [Holotype W 0020331 I Rchb. Orch. 44221 (sterile, with Reichenbach’s manuscript description)], Isotypes: San Ramon, along Barranca River, «Pleurothallis mellifera»,, A.R. Endres s.n. «N.° 92 gen. coll. Orch.» [W 0021581 / Rchb. Orch. 12806 (with Endres’ description)]; - A.R. Endres s.n. «92» [W 0020150 / Rchb. Orch. 12806]; - A.R. Endres s.n. «92» [AMES 118500]. Additional material associated with the type: Endres’ sketches of the plant and flower, A.R. Endres s.n. «N.° 92» [W 0020152 / Rchb. Orch. 44198; - Endres’ sketches of the plant and flower, A.R. Endres s.n. «92» [W 0020151 / Rchb. Orch. 33221]; - A.R. Endres s.n. «92» [W 1889-33651], Additional Endres’ material: A.R. Endres s.n. «See fig. p. 81 Below vol. 1869-70» [W 1889-46833],

83. Pleurothallis endresii L uer , Monogr. Syst. Bot. Missouri Bot. Gard. 76: 172 (1999). Type: Costa Rica. Cartago: «Pizirres, along road to San Cristobal», ca. 1867, A.R. Endres s.n. «N° 555 gen coll. Orch.» [Holotype W 0020145 7 Rchb. Orch. 12798 (Endres’ drawing of the plant, flower and floral dissections)]. Additional material associated with the type: Luer’s drawing of the plant, flower and floral dissection from the type [W 0020144]. Additional Endres’ material: Costa Rica. Santa Maria, June 4, A.R. Endres s.n. «Orch. N° 140» [W 1889-0024860].

84. Pleurothallis fulgens R chb .L, Gard. Chron., n.s., 4 (95): 516 (1875). = Specklinia fulgens (RcHB.fi) Pridgeon & M.W. C hase , Lindleyana 16: 257 (2001). ©Naturhistorisches Museum Wien, download unter www.biologiezentrum.at P u p u l in et al.: Typi Orchidacearum Augusto R. Endresio in Costa Rica lecti 2 9 7

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= Sarcinula fulgens (R chb .f.) Luer , Monogr. Syst. Bot. M issouri Bot. Gard. 105-213 (2006). Type: Costa Rica. “The plant was discovered in Costa Rica by Endres, and had flowered several times at the Hamburgh Botanic Garden”, A.R. Endres 2512 [H olo­ type W 0020353 / Rchb. Orch. 25509, isotype AMES 118517 (barcode 5224566)] Additional material associated with the type: Reichenbach’s manuscript description [W 0020245 / Rchb. Orch. 25510], Additional Endres’ material: Costa Rica. «Boca de Dota», A.R. Endres s.n. «N° 246 gen. coll. Orch.; 51» [W 0020241 / Rchb. Orch. 38482]; - Endres’ drawings of the flower, floral dissection and description, A.R. Endres s.n. «N° 51 gen. coll. Orch.» [W 0020240 / Rchb. Orch. 38492]; - A.R. Endres s.n. «51» [W 0020244 / Rchb. Orch. 44226]: - Endres’ drawings of the plant, flower, and floral dissections, A.R. Endres s.n. «51» [W 0020242 / Rchb. Orch. 33418]; - «San Ramon», A.R. Endres s.n. «N° 252 gen. coll. Orch.» [W 0020243 / Rchb. Orch. 12839]; - «San Ramon», A.R. Endres s.n. «51» [W 1889-0041435]; - A.R. Endres s.n. «52» [W 1889-0033634); - «Calera», A.R. Endres s.n. «51» [W 1889-0041397].

85. Pleurothallis lateritia Endres ex R chb .f., Gard. Chron. 731 (1872). Type: Costa Rica. “It was sent from Costa Rica to the Hamburgh Botanic Garden by M. Endres”, A.R. Endres 52 [Holotype W 0019852 / Rchb. Orch. 24750 (with Reichenbach’s drawings and description) and W 0019851 / Rchb. Orch. 44241 (with Endres’ drawings), isotype AMES 69077 (barcode 5225584)]. Additional Endres’ material: Costa Rica. A.R. Endres s.n [W 0020313 / Rchb. Orch. 13154];-A .R . Endres s.n. «218» [W 0020314 / Rchb. Orch. 44170];-A .R . Endres s.n. [W 0020316/Rchb. Orch. 13156];-A .R . Endres s.n. «51» [W 1889-0033628]; - A.R. Endres s.n. «208» [W 1889-0037461].

The name is a synonym of Pleurothallis lanceola (Sw.) Spreng . (1826).

86. Pleurothallis luctuosa R chb .f., Linnaea 41: 48 (1877), emend. in Xenia Orch. 3: 15-16, pl. 210(1878). Type: Costa Rica. “Viv. observavi in horto Hamb.”, (Alajuela, San Ramon, 3500“, 1867), A.R. Endres s.n. «N.° 2537» [Holotype W 0020310 / Rchb. Orch. 53791 (with Endres’ and Reichenbach’s drawings of the plant and flower), isotype AMES 118528], Additional Endres’ material: Costa Rica. San Ramon, May-June, Endres’ sketches of the plant and flowers, floral dissections and description, A.R. Endres s.n «N.° 50 gen. coll.» [W 0020306 / Rchb. Orch. 33417]; - Endres’ pencil plate of the plant, flowers and floral dissections, A.R. Endres s.n «N.° 50 gen. coll.» [W 0020308 / Rchb. Orch. 36294]; May-June, A.R. Endres s.n «N.° 50 gen. coll. Orch.» [W 0020305 / Rchb. Orch. 2805]; - A.R. Endres s.n «N.° 50» [W 0020307 / Rchb. Orch. 44222]; - Without specific local­ ity, A.R. Endres s.n [W 0020309 / Rchb. Orch. 2808; - A.R. Endres s.n. «218» [W 0020314 / Rchb. Orch. 44170]; - A.R. Endres s.n. (W-0020316 / Rchb. Orch. 13156]; - A.R. Endres s.n. «51» [W 1889-33628]; - A.R. Endres s.n. «208» [W 1889-37461].

87. Pleurothallis moschata R chb .f., Xenia Orchid. 3 (2): 42—43, t. 217 (1881). = Trichosalpinx moschata (R chb .f.) Luer (1983). Type: Costa Rica. “Diese recht zierliche Pleurothallis blüht seit einigen Jahren im Pin>ULiN et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Wien, Endresio download in unter Costa www.biologiezentrum.at Rica lecti 299

hamburger Botanischen Garten, Wir haben sie von Endres aus Costa Rica erhalten” [Holotype W 0020892 / Rchb. Orch. 53825 (with Reichenbach’s drawings, A.R. Endres s.n. «N.° 2653»]. Additional Endres’ material: Costa Rica, without specific locality, «Pleurothallis / see p. 65 lower outer vol. 1867-68», A.R. Endres s.n. [W 1889-46826]; Endres’ sketches of the flowered plant, flower and floral analysis, with description, «See p. 65 lower outer vol. 1867-68, descr. p. 62», A.R. Endres s.n. [W 0020891 / Rchb. Orch. 38471]; - Endres’ pencil plate of the flowered plant, flower and floral analysis, A.R. Endres s.n. [W 0020890 / Rchb. Orch. 36255]; - Endres’ pencil plate of the flowered plant, flower an d floral analysis, A.R. Endres s.n. «N.° 65 gen. coll.» [W 0020889 / Rchb. Orch. 36269]; - A.R. Endres s.n. «Orch. N.° 239» [W 0020888 / Rchb. Orch. 12786].

88. Pleurothallispolyliria Endres & R chb .f., Gard. Chron. 731 (1872). Humboldtia polyliria (Endres & R chb .f.) Kuntze , Revis. Gen. Pl. 2: 668 (1891). Syntypes: “Abeautiful specimen at hand is kindly sent by M. Wilson Saunder, Esq., who remarks that the plant is totally cespitose. Quite the same plant... is at hand in dried specimens, gathered at Costa Rica by Mr. Endres” [Holotype not found].

The name is a synonym ofPleurothallis gelida Lindl . (1841).

89. Polycycnis gratiosa Endres & R chb .f., Gard. Chron. 1451 (1871). Type: Costa Rica. “It is a very elegant thing, discovered by Mr. Endres in Costa Rica. Last autumn, it flowered in the celebrated collection of Messrs. Veitch” [Holotype W 0029866 / Rchb. Orch. 19116 (with Reichenbach’s drawings and description, cultivated by Veitch, A.R. Endres s.n. «136»)]. Additional Endres’ material: Costa Rica, without locality, A.R. Endres s.n «N.° 2560» [W 0020865 / Rchb. Orch. 33819]; - A.R. Endres s.n «135» [W 0020864 / Rchb. Orch. 19117].

90. Ponera mesocopis Endres & R chb .f., Xenia Orchid. 2: 222, t. 200 (1874). = Scaphyglottis mesocopis (Endres & R chb .f.) B enth . & H ook. f. ex Hemsl. (1884). Type: Costa Rica. “Diese Art, welche stark an manche Arten der GattungHexades- mia erinnert empfing ich von Hem Endres aus Costa Rica”, A.R. Endres s.n. «N.° 2617» [Holotype W 0020735 / Rchb. Orch. 52835 (with Reichenbach’s drawings)]. Additional Endres’ material: Costa Rica, without locality, «Ponera heterocopis Endr. Rbf.», Endres’s sketch of the flowered plant, A.R. Endres s.n «714» [W 0020731 / Rchb. Orch. 52815]; - «Hexadesmia heterocopis», Reichenbach’s sketch of the flower, A.R. Endres s.n «N.° 714» [W 0020733 / Rchb. Orch. 51951]; - A.R. Endres s.n «Orch. N.° 237» [W 0020738 / Rchb. Orch. 13798]; - A.R. Endres s.n «237» [W 0020739 / Rchb. Orch. 13842]; - with Reichenbach’s sketch, «Ponera cirrhata», A.R. Endres s.n «Orch. N.° 337» [W 0020737 / Rchb. Orch. 43741]; - Endres’ sketch of the flower and floral dissection, with description, «Ponera cirrhata Endr. Rchb.f.», A.R. Endres s.n «N.° 337» [W 0020736 / Rchb. Orch. 37039]; - A.R. Endres s.n [W 0020734 / Rchb. Orch. 51952]; - A.R. Endres s.n «N.° 2655» [W 0020732 / Rchb. Orch. 51950].

91. Restrepia dayana R chb .f., Gard. Chron., n.s., 4 (87): 257 (1875). Type: “The plant is a discovery of the late M. Endres in Costa Rica, where it grows at an altitude of 3000 feet, but is very scarce” [Holotype W 0020817 / Rchb. Orch. 300 ©Naturhistorisches Museum Wien,Annalen download unter des www.biologiezentrum.at Naturhistorischen Museums in Wien B //1

44199 (sterile, with Reichenbach’s description and Endres’ drawings o f the plam flower, and lip, 1868, A.R. Endres «39», isotype W 0020822 / Rchb. Orch. 13150] Additional material associated with the type: Costa Rica. «San Carlos Valley, pejc var. picta, a pretty var.? of 263», Endres’ drawings and descriptions, A.R. Endres s.n. «Restrepia N°. 3, N°. 39 gen. coll. Orch.» [W 0020818 / Rchb. Orch. 38581 and W 0020819 / Rchb. Orch. 38580]; - «Restrepia concinna», Endres’ drawing o f the connate with column, A.R. Endres s.n. «N° 39 gen. coll. Orch.» [W 0020821 / Rchb. Orch. 36076]; - «Restrepia concinna», Endres’ descriptions, A.R. Endres s.n. «N° 39» [W 0020820 / Rchb. Orch. 33424]; - Endres’ definitive pencil plate «62» A.R. Endres s.n. «N° 39 gen. coll.» [W 0020814 / Rchb. Orch. 36063], Additional Endres’ material: Costa Rica. «Angostura, May», A.R. Endres s.n. [W 1889- 0039086, W 1889-0039075]; - «San Cristobal, Sta. Maria, Nov.ber-Dec.ber», A.R. Endres s.n., W-Rchb. Orch. 1889-0030738; - «Sta. Maria, San Cristobal, December», A.R. Endres s.n., W-Rchb. Orch. 1889-0030744. - «Pr. of Cartago», A.R. Endres s.n. [W 1889-0024852]; - Without locality, «Restrepia xanthophtalma var. picta, N° 39 gen. coll. Id. var. concinna», Endres’ drawings of the plant with flower and lip, and descrip­ tion, A.R. Endres s.n. «Restrepia N° 1» [W 0020816 / Rchb. Orch. 36074]; - «Restrepia xanthophtalma var. picta [and] Restrepia N° 1 var. obscura», Endres’ descriptions, A.R. Endres s.n. «Restrepia N° 1» [W 0020815 / Rchb. Orch. 32237]; - A.R. Endres s.n. [W 1889-0037467]; - Endres’ drawing of the plant, A.R. Endres s.n. [W 0020823 / Rchb. Orch. 36560]; - Endres’ definitive pencil plate «104» A.R. Endres s.n. «N° 263» [W 0020813 / Rchb. Orch. 36064]; - «see fig. p.104», A.R. Endres s.n. «N°. 263» [W 1889- 0041455, W 1889-0041456]; - «see fig. p.104», A.R. Endres s.n. «N° 263 of gen. coll.» [W 1889-0041455, W 1999-0000196],

The name is a synonym of Restrepia muscifera (Lindl .) R chb . f. ex Lindl . (1859)

92. Restrepiaprorepens R chb .f., Gard. Chron., n.s., 7 (183): 810 (1877). = Barbosella prorepens (R chb .f.) Schltr . (1918) Type: “It is one of the Costa Rican discoveries of the late M. Endres. It has flowered several times in the Hamburgh Botanic Garden”, A.R. Endres 2618. Lectotype (annotated by Luer on correction label 1992, hic designatus): Costa Rica, without specific locality, A.R. Endres 2618 [W 0018995 / Rchb. Orch. 29375 (with Reichenbach’s manuscript descriptions and drawings)] - Fig. 9. Additional Endres’ material: Costa Rica, without locality data, 1867, with Endres’ pen­ cil plate of plant, flowers and floral dissections, «Malaxidieae N° 71», A.R. Endres s.n. [W 0019003 / Rchb. Orch. 44201]; - Endres’ pencil plate of plant, flowers and floral dissections, «52», A.R. Endres s.n. «N° 71» [W 0018993 / Rchb. Orch. 36078]; - Endres’ preparatory sketches of the flowered plant and floral dissections, A.R. Endres s.n. «N° 71» [W 0019000 / Rchb. Orch. 33217]; - Endres’ pencil plate of plant, flowers and floral dissections, «53», A.R. Endres s.n. «N° 271» [W 0018993 / Rchb. Orch. 36078]; - Endres’ floral dissections, «Restrepia teretifolia», A.R. Endres s.n. «Restrepia N° 8, N° 71 gen. coll. Orch.» [W 0018999 / Rchb. Orch. 38575]; - Endres’ pencil plate of plant, flowers and floral dissections, « Restrepia», A.R. Endres s.n. «N° 271 gen. coll. Orch.» [W 0019002 / Rchb. Orch. 36077]; - Endres’ preparatory sketches of plant, flowers and floral dissections, «Restrepia violodora», A.R. Endres s.n. «Restrepia N° 9, N° 271 gen. coll. Orch.» [W 0019003 / Rchb. Orch. 44201]; - Endres’ preparatory P i i p u l i n et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Wien, Endresio download inunter Costa www.biologiezentrum.at Rica lecti 3 0 1 reserved Copyright

Fig. 9: Lectotype of Restrepia prorepens R c h b . f. [W 0018995 / Rchb. Orch. 29375]. 302 ©Naturhistorisches Museum Wien,Annalen download unter des www.biologiezentrum.at Naturhistorischen Museums in Wien B y yi

sketches of plant, flowers and floral dissections, «Restrepia violodora», A.R. Endres s «Restrepia N° 9, N° 271 gen. coll. Orch.» [W 0018997 / Rchb. Orch. 38573]. The material from Endres at W include at least two different gatherings, one of which is identified by him as «Restrepia teretifolia» or «Restrepia N° 8», corresponding to his general collection No. 71, and the other annotated as «Restrepia violodora» or «Restrepia N° 9», corrsponding to his general collection No. 271.

93. Restrepia reichenbachiana E n d r e s & R c h b .f., Gard. Chron., n.s., 4 (90)- 355 (1875). = Restrepiopsis reichenbachiana (E n d r e s & R c h b .f.) L u e r (1 9 8 1 ). = Pleurothallopsis reichenbachiana (E n d r e s & R c h b .f.) P r id g e o n & M .W .C hasl; (2001). Type: “It was discovered in 1868 by Endres in Costa Rica, and has flowered several times in Hamburgh Botanic Garden”, A.R. Endres s.n. Lectotype (hic designatus): Costa Rica, without locality, A.R. Endres s.n. «Orch. N° 46», [W 0020803 / Rchb. Orch. 44207 (with Endres’ final pencil plate, paitly coloured, of plant, flowers and floral dissections) (Fig. 10), isolectotype W 1889- 0037462]. Additional material associated with the type: Costa Rica. Endres’ pencil plate of p lant, flowers and floral dissections, «Restrepia Reichenbachiana E n d r .» , A.R. Endres s.n. «N° 46 gen. coll. Orch.» [W 0020802 / Rchb. Orch. 36073]; - «R. Reichenbachiana», Endres’ sketches and descriptions, A.R. Endres s.n. «Restrepia N° 7, N° 46 of gen. coll. Orch.» [W 0020801 / Rchb. Orch. 38576]. Additional Endres’ material: A.R. Endres s.n. [W 1889-0039081]; - Endres’ sketch of the flowered plant, A.R. Endres s.n. [W 0021312 / Rchb. Orch. 33415]. Among the materials referrable to this taxon at W are collections by Endres which are identified by his general collection number 46, corresponding to «Restrepia N° 7», and others with no number. As Reichenbach does not include this information in the proto­ logue, the sheet 0020803 of the general collection at Vienna is selected as the lectotype: it includes a fertile specimen, as well as detailed drawings by Endres, showing the char­ acteristic features of the species.

94. Scaphosepalum endresianum K r a e n z l ., Repert. Spec. Nov. Regni Veg. 17 (28): 435-436(1921). Type: “Costa Rica. Ohne genaueren Standort”, (1868, 4500’), A. R. Endres s.n. (Orch. N.° 55) [Holotype W 0020770 / Rchb. Orch. 15005 (with Endres’ drawing of the plant, flower and floral dissections, and Reichenbach’s analysis of the flower), isotype W 1889-30746]. Additional material associated with the type: Costa Rica. «San Ramon. Camino a San Carlos, Legua. Rainy season», with Endres’ description, A.R. Endres s.n. «N.° 55 gen. coll. Orch.» [W 0020776 / Rchb. Orch. 18284]; - «San Ramon. Scarce / Rainy season», Endres’ drawings of the plant, flowers and floral dissections, A.R. Endres s.n. «N.° 55 gen. coll. Orch. / Masdevallia N.° 19» [W 0020774 / Rchb. Orch. 38549],

The name is a synonym of Scaphosepalum anchoriferum (R c h b .f.) R o l f e (1890) = M as­ devallia anchorifera R chb.f. (1884). P u p u l in et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Wien, Endresio download in unter Costa www.biologiezentrum.at Rica lecti 303

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Fig. 10: Lectotype of Restrepia reichenbachiana E n d r e s & Rchb.f. [W 0020803 / Rchb. Orch. 44207]. 304 ©Naturhistorisches Museum Wien,Annalen download unter des www.biologiezentrum.at Naturhistorischen Museums in Wien, B //■>

95. Scaphosepalum naviculare Kraenzl., Repert. Spec. Nov. Regni Veg. 17 n%\- 435-436(1921). 1 Type: “Costa Rica. Ohne genaueren Standort”, A.R. Endres s.n. [Holotype W 0020777 / Rchb. Orch. 14515], isotype «N.° 506 gen. coll. Orch. / see p. 57 y0| 1869-70» W 0020765 / Rchb. Orch. 14508], Additional material associated with the type: Endres’ pencil plate of the plant, flow­ ers and floral dissections, A.R. Endres s.n. «p. 57» [W 0020772 / Rchb. Orch 36058]; - Endres’ sketches of the plant, flowers and floral dissections, A.R. Endres s.n. «N.° 506 gen. coll. Orch. / see p. 57 vol. 1869-70» [W 0020775 / Rchb. Orch 38552]; - Endres’ descriptions, A.R. Endres s.n. «N.° 506 gen. coll. Orch. / see fig p. 57 vol. 1869-70» [W 0021727 / Rchb. Orch. 38552], The name is a synonym of Scaphoseplaum anchoriferum.

96. Sievekingiaßimbriata R chb .f., 69: 449^1-50 (1886). Type: “Costa Rica. Endres”, «San Ramon. Legua / Dec. January», A.R. Endres s.n. [Holotype W 0020709 / Rchb. Orch. 44723 (with Reichenbach’s sketches and description)]. Additional material associated with the type: Endres’ sketches of the flower and flo­ ral analyses, «San Ramon. Legua / Dec. January», A.R. Endres s.n. [W 0020708 / Rchb. Orch. 37032].

97. Sievekingia suavis R chb .f., Beitr. Syst. Pflanzenk. 3 (1871). Type: Costa Rica. “Sie blühte zuerst 1868 im hiesigen botanischen Garten und wurde am 29. Juni 1869 bei einer unserer Ausstellungen angemeldet, hatte aber lei­ der bereits dem Verblühungsprocesse ihren Tribut gezollt. Sie wurde aus Costa Rica von Herrn Endres eingesendet, der dieser Tage berichtete, er habe eine zweite Art entdeckt”, A.R. Endres s.n. [Holotype W 0027462 / Rchb. Orch. 44725 (with Rei­ chenbach’s drawings and description)]. 98. Sigmatostalix macrobulbon Kraenzl., Pflanzenr. 4, 50 (Heft 80): 307, f. 27B (1922). Type: “Zentralamerikanische Provinz. Costarica”, 3^4000”, 1867, A.R. Endres s.n. «20» [Holotype W 0020696 / Rchb. Orch. 18047 (with Endres’ drawings of the plant and flowers), isotypes W 0020705 / Rchb. Orch. 2057; W 0020704 / Rchb. Orch. 2059; W 0020703 / Rchb. Orch. 2045; W 0020697 / Rchb. Orch. 2052; W 0020703 / Rchb. Orch. 2045; W 0020699 / Rchb. Orch. 2043], Additional material associated with the type: Costa Rica, without locality, Endres’ floral analyses, A.R. Endres s.n. «N.° 20 ge, coll. » [W 0020700 / Rchb. Orch. 37054]; - Endres’ sketches of the plant and flowers, A.R. Endres s.n. «20» [W 0020702 / Rchb. Orch. 37057]; - Reichenbach’s sketches of the , A.R. Endres s.n. «Endr. 20» [W 0020701 / Rchb. Orch. 39351]. Additional Endres’ material: Costa Rica, without locality, A.R. Endres s.n. [W 0020693 / Rchb-Orch 2050; W 0020694 / Rchb-Orch 2051; W 0020695 / Rchb-Orch 2055; W 0020698 / Rchb-Orch 2061]; - Costa Rica, without specific locality, Reichenbach’s description and drawings, «Sigmatostalix endresii», A.R. Endres s.n. «E 20» [W 0020706 / Rchb. Orch. 18046]. The sheet selected as the lectotype has a fertile specimen and a detailed illustration by Endres. Pupulin et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Wien, Endresio download inunter Costa www.biologiezentrum.at Rica lecti 305 Copyright reserved

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Fig. 11: Lectotype of Sigmatostalix poikilostalix K r a e n z l . [W 0020692 / Rchb. Orch. 18043], 306 ©Naturhistorisches Museum Wien,Annalen download unter des www.biologiezentrum.at Naturhistorischen Museums in Wien B / /1

Some authors treat this name as the basionym of Oncidium macrobulbon (Kraenzl \ M.W. C hase & W illiams (2008).

99. Sigmatostalixpoikilostalix K raenzl ., Pflanzenr. 4, 50 (Heft 80): 310, f. 26D (1922) Lectotypus (hic designatus): 1868, A.R. Endres 97 [W 0020692 / Rchb-Orch 18043 (with Endres drawings of the plant, flower, and floral analysis)] (Fig. m isolectotypes [W 0020689 / Rchb. Orch. 2097, W 0020687 / Rchb. Orch. 2044 \y 0020686 / Rchb. Orch. 2040, W 0020684 / Rchb. Orch. 2048]. Syntypes: Costa Rica. “Zentralamerikanische Provinz. Costarica”, A.R. Endres 38 [W 0020691 / Rchb. Orch. 2041], Additional material associated with the type: Costa Rica, without locality, Endres’ sketch of the flowered plant, flower, and floral details, A.R. Endres 97 [W 0020690 / Rchb. Orch. 37055]; - Endres’ sketch of the lip, A.R. Endres 97 [W 0020685 / Rchb. Orch. 37056].

Some authors treat this name as the basionym of Oncidium poikilostalix (K raenzl ) M.W. C hase & N.H. W illiams (2008). 100. endresii Kraenzl., Pflanzenr. 4, 50 (Heft 80): 317 (1922). = Mesospinidium endresii (Kraenzl .) Garay (1973). Type: “Heimat nicht genau bekannt, vermutlich Costa Rica”, A.R. Endres s.n. [Holotype W 0027461 / Rchb. Orch. 32817].

According to Kränzlin (1922), in the Reichenbach herbarium there were seven dried inflorescences with the label “Sigmatostalix endresii”, without author. Kränzlin considered that the species was not related to Sigmatostalix, and described it under the genus Solenidium. The name «Sigmatostalix endresii» was in fact suggested by Reichenbach, who prepared a manuscript description and illustrated a flower, ostensibly from Endres s.n. «20» (on the upper right side of the leaflet, Reichenbach annotated «E 20»), which corresponds to the type of the species eventually described by Kränzlin himself as Sig­ matostalix macrobulbon. This drawing is actually mounted alone on sheet W 0020706 / Rchb. Orch. 18046, but until at least late 2000 it was mounted on a composite sheet including specimens of Mesospindium collected by Endres (now on W 0027461), as well as a drawing of the flower by Reichenbach. It is probably studying these specimens and the drawing from a flower, which are completely unrelated to the name «Sig­ matostalix endresii» sensu Reichenbach, that Kränzlin commented about his difficulty to understand the generic placement suggested by the Hamburg professor (Kränzlin 1922).

The name is a synonym ofMesospinidium warscewiczii R chb . f.

101. Stanhopeapulla R chb .f., Gard. Chron., n.s., 7: 810 (1877). Type: “We obtained it from our friend, the late M. Endres, who discovered it in Costa Rica” Holotype: Costa Rica, without specific locality, with Reichenbach’s description, A.R. Endres s.n. [W 0027462]. Additional Endres’ material: Costa Rica. «Calvario / Camino a Limon, Juy August», Endres’ drawings and description, A.R. Endres s.n. «N° 612 gen. coll. Orch» [W 0020589 / Rchb. Orch. 32019] - Without locality, A.R. Endres s.n. [W 0021219 / Rchb. Orch. 8883]; - A.R. Endres s.n. «277» [W 0021220 / Rchb. Orch. 44262]; - A.R. Endres s.n. [W 1889-0041472]; - A.R. Endres s.n. «see fig. p. 91» [W 1889-0046718]. ©Naturhistorisches Museum Wien, download unter www.biologiezentrum.at P u p u l in et al.: Typi Orchidacearum Augusto R. Endresio in Costa Rica lecti 3 0 7

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Fig. 12: Lectotype of Telipogon endresianus K r a e n z l . [W 0020980 / Rchb. Orch. 2008]. 308 ©Naturhistorisches Museum Wien,Annalen download unter des www.biologiezentrum.at Naturhistorischen Museums in Wien B I/ ■>

According to Jenny (2010), S. pulla was described from a dried inflorescence sent from Costa Rica by Endres. The species was later illustrated from Reichenbach’s own sketches, on the basis of material cultivated at Hamburg Botanical Garden, also col­ lected by Endres (R eichenbach 1878). Besides Endres’ original gathering, on the typc sheet at W are also mounted five other inflorescences, probably corresponding to the plants flowered at Hamburg in June 1877 and used by Reichenbach for his illustrations intended for Xenia Orchidacea, plus a single flower which belongs to Stanhopea con- nata (Jenny 2010).

102. endresii R chb .f., Gard. Chron. 1870 (42): 1373 (1870). Type: “It was discovered in Costa Rica by M. Endres, and flowered in the Ham- burgh Botanic Garden in December, 1869, and July, 1870” Holotype: Costa Rica. Cultivated at Hamburg Botanic Garden, 1869-1870, A.R. Endres s.n. [W 0021221 / Rchb. Orch. 18468 (sterile, with Reichenbach’s drawings and description)]. Additional Endres’ material: Costa Rica. «Sarsero [Zarcero], Sapote [Zapote], May- June» and «old road to San Carlos, on the height beyond Tapisco river, May», with Endres’ drawings of the flower, floral dissections and description, A.R. Endres s.n. [W 1889-014213]; - Without locality, A.R. Endres s.n. «Orch. N° 277» [W 0021219 / Rchb. Orch. 8883]; A.R. Endres s.n. «277» [W 0021220 / Rchb. Orch. 44262]; A.R. Endres s.n. [W 1889-0041472]; - A.R. Endres s.n. «see fig. p. 91» W-Rchb. Orch. 1889- 0046718; The name is probably a synonym ofStelis argentata Lindl. (1842).

103. Telipogon buenavistae Kraenzl ., Ann. Naturhist. Mus. Wien 33: 33-34 (1920). Type: “Costa Rica. Buenavista, near the spot called „ojo de aqua“ in 3000 m”, [Cerro de Buena Vista], A.R. Endres s.n. [Holotype W 0020978 / Rchb. Orch. 45087].

The name is a synonym of Telipogon costaricensis Schltr . (1911).

104. Telipogon christobalensis Kraenzl ., Ann. Naturhist. Mus. Wien 33: 30 (1920). Type: “Costa Rica. Bei San Cristobal”, Tablazo, road to S. Cristobal / June-July, A.R. Endres 512 [Holotype W 0020975 / Rchb. Orch. 2003], isotypes [W 0020974 / Rchb. Orch. 2009, W 0020976 / Rchb. Orch. 2010, «S. Crist. + S. Maria» W 0020977 / Rchb. Orch. 2005].

105. Telipogon endresianus K raenzl ., Ann. Naturhist. Mus. Wien 33: 31 (1920) Lectotype (hic designatus): Costa Rica. [Cartago: Birris, Sitio de Don Eusebio Ortiz, August], A.R. Endres s.n. [W 0020980 / Rchb. Orch. 2008, Fig. 12, isolecto- type AMES 26049]. Syntype: “Costa Rica. Ohne genaueren Standort”, 1882, Hübsch s.n. [W 22702]. Additional material associated with the type: Endres’ pencil plate of the lectotype, including plant habit with flowers, pollinarium and anther, A.R. Endres s.n. «p. 41» [W 0020979 / Rchb. Orch. 9960],

The sheet selected as the lectotype includes two fertile specimens, and it is accompanied (on another sheet) by a detailed drawing by Endres.

The name is a synonym of Telipogon biolleyi Schltr . (1911). P u p u l in et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Wien, Endresio download inunter Costa www.biologiezentrum.at Rica lecti

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Fig. 13: Lectotype of Trichocentrum pfavii var. zonale R c h b . f. [W-0020947 / Rchb. Orch. 37150]. 310 ©Naturhistorisches Museum Wien,Annalen download unter des www.biologiezentrum.at Naturhistorischen Museums in Wien B / / ->

106. Telipogon minutiflorus Kraenzl ., Ann. Naturhist. Mus. Wien 33: 14 (1920). Type: “Costa Rica. Ohne genaueren Standort”, A.R. Endres s.n. [Holotype W 0021327 / Rchb. Orch. 2011, isotypes W 0021326 / Rchb. Orch. 2006, W 0021325 / Rchb. Orch. 2007]. 107. Trichocentrum pfavii var. zonale Rchb. f., Gard. Chron., n.s., 19: 44 (1883). Type: Costa Rica. “A fresh inflorescence just sent by Mr. Harry Veitch”, A.R Endres s.n. Lectotype (hic designatus): Costa Rica. San Jose: Dota, «Boca de Dota / Oct.- December», «Trichocentrum saundersii», A.R. Endres s.n. [W 0020947 / Rchb Orch. 37150 (Endres’ drawings and description)], Fig. 13. Additional material associated with the type: Costa Rica. Endres’ drawings of the ovary, with sections, and descriptions, «Boca de Dota, along the creeks / on the trunks of young trees / 1- 2000”, beyond Ramon Zuniga house», «Trichocentrum saundersianum», A.R. Endres s.n., [W 0020946 / Rchb. Orch. 37148]; - Endres’ coloured drawing of flower, A.R. Endres s.n. [W 0020952 / Rchb. Orch. 37153]; - Without locality, «Trichocentrum», A.R. Endres s.n. «N° 16 gen. coll. » [W 0020948 / Rchb. Orch. 14171]; - with Endres’ analysis of the flower, «Trichocen­ trum saundersii», A.R. Endres s.n. [W 0020949 / Rchb. Orch. 18376]; - Endres’- drawing of the plant, A.R. Endres s.n. [W 0020945 / Rchb. Orch. 37152]; - Endres’ drawing of the pollinarium, «Trichocentrum saundersii», A.R. Endres s.n. [W 0020951 / Rchb. Orch. 37151]. In the protologue, Reichenbach refers to an “unpublished lithographic plate belonging to one of my unpublished pamphlets”, which corresponds to an engraving intended for his series of Xenia Orchidacea, still conserved in his herbarium with the title “Trichocen­ trum zonale” This lithograph was ostensibly prepared from the drawings sent by Endres under the intended name of “Trichocentrum saundersii” and “Trichocentrum saunder­ sianum”, from material originally collected at «Boca de Dota, along the creeks / on the trunks of young trees / 1-2000”, beyond Ramon Zuniga house», which have to be con­ sidered part of the type material studied by Reichenbach. In absence of any actual spec­ imens, the illustrations sent by Endres and used by Reichenbach to prepare the litho­ graph for his T. zonale are selected as the lectotype.

The name is a synonym of Trichocentrum pfavii R chb .f. (1881).

108. Trichocentrum caloceras Endres & R chb .f., Gard. Chron. 1257 (1871). Type: “Mr. Endres, the excellent Costa Rica traveller, has found since 1867, in Costa Rica, another Trichocentrum” Holotype, to be found at W, not located. Neo­ type, designated by Pupulin (1995): Costa Rica. Puntarenas: Coto Brus, F. Pupulin & D. Castelfranco 1 [USJ]. 109. dasyandra Rchb.f., Xenia Orchid. 3: 64, t. 230 (1883). = dasyandra (R chb .f.) D ressler & N.H. W illiams (1970). Type: Costa Rica. “Endres”, 1868, «Trichopiliastrum», A.R. Endres s.n. «Orch. N.° 93». [Holotype W 0019850 / Rchb. Orch. 18032 (with Endres’ and Reichenbach’s drawings), isotype: A.R. Endres s.n. «93» W 0020939 / Rchb. Orch. 2982]. Additional material associated with the type: Costa Rica. Endres’ sketches and pencil plate of flowered plant, flower and floral analysis, with description, «Aspa- P u p u l in et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Wien, Endresio download inunter Costa www.biologiezentrum.at Rica lecti

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Fig. 14: Lectotype of Warczewiczella picta R c h b . f. [W 0019049 / Rchb. Orch. 13582], 312 ©Naturhistorisches Museum Wien,Annalen download unter des www.biologiezentrum.at Naturhistorischen Museums in Wien B / / ->

sia», A.R. Endres s.n. «N. 93 gen. coll. Orch.» [W 0018815 / Rchb. Orch. 35949]. - Endres’ sketches of floral parts, «Trichopiliastrum», A.R. Endres s.n. «93» 0018816 / Rchb. Orch. 35950 (engravings from Endres’ drawings, intended for Xenia Orchidacaea, not published)]. Additional Endres’ material: Costa Rica, without locality, «», A.R. Endres s n [W 0018814/Rchb. Orch. 38570]. 110. Warczewiczella picta R chb.f., Gard. Chron., n.s., 20: 8 (1883). = Stenotyla picta (RcHB.f.) D ressler (2005). Lectotype (hic designatus): “This very fine plant was discovered by late Endres“ Costa Rica. A.R. Endres s.n. «N° 575 gen. coll. Orch.» [W 0019049 / Rchb. Orch 13582], Fig. 14. Syntype: „..., and lately found again by M. Pfau“ Costa Rica, R. Pfau s.n. [not located]” Additional material associated with the type: Costa Rica. Endres’ drawings and descriptions, «Cerro de los Guatusos, headwater of La Paz River ...», A.R. Endres s.n. «N° 575 gen. coll. Orch.» [W 0019022 / Rchb. Orch. 14629]; - Endres’ draw­ ings, A.R. Endres s.n. «N° 575 gen. coll. Orch., Zygopetalum N° 5» [W 0019023 / Rchb. Orch. 10623]. 111. Zygopetalum lacteum R chb.f., Gard. Chron. 1290 (1872). = Kefersteinia lactea (R chb .f.) Schltr . (1923). Syntypes: “It was originally discovered by Mr. Wallis during his adventurous expe­ dition to Chiriqui [Panama, Wallis s.n., not located]. Later, it was collected by M. Endres [Costa Rica, A. R. Endres 334, W 0019731 / Rchb. Orch. 49620]. It flow'- ered some years ago with Director Linden [Panama or Costa Rica, J. Linden s.n., W 0019731 / Rchb. Orch. 49620], and is now once more in flower with Messrs. Veitch [without locality, cult. Veitch s.n., W 0019731 / Rchb. Orch. 49620]” Lectotype (Pupulin 2001: 548): Panama or Costa Rica. Chiriqui, cultivated by Lin­ den s.n. [W 0019731 / Rchb. Orch. 49620]. Reichenbach planned to publish the species after correspondence with Endres, who sent him several sketches and tentative descriptions based on Costa Rican collections made around 1868 [W 0019832 / Rchb. Orch. 44590, W 0019831 / Rchb. Orch. 14705, W 0019830 / Rchb. Orch. 33240] with the proposed name „Zygopetalum lacteum Endr. & Rchb.f.“ Even though the manuscript protologue by Reichenbach expressly indicates the co-autorship of the new species with its discoverer, the name of Endres was eventu- ally deleted before publication.

Excluded species

Aspasia papilionacea R chb.f., Gard. Chron., n.s., 6: 100 (1876). Type: Costa Rica. “Its discoverer, the late Senor Endres, appears not to have guessed what it was. He sent it with other things without ever mentioning the plant that beats all his Orchidaceous discoveries. He may have obtained it shortly before leaving Costa Rica, and the plant took a long rest before it developed two splendid flowers at once. It may be found with Messrs. Veitch, who got a lot of unnamed P u p u l in et al.: Typi Orchidacearum©Naturhistorisches Augusto Museum R. Wien, Endresio download in unter Costa www.biologiezentrum.at Rica lecti 313

things from Senor Endres”, cult Veitch s.n. [Holotype not located; drawing of the type, copy of Reichenbach’s sketches, AMES 24849 - barcode 4747525].

The name is a later synonym of L indl . {Edwards’s Bot. Reg. 22: t. 1907 (1836)}, described on the basis of a plant collected in by M. Descourtilz. This small sized, unmistakable epiphyte is only known from Brazil, and the data about the original collection of A. papilionacea given by Reichenbach in the protologue were surely based on a mislabeled plant obtained through the nurseries of Veitch.

R eferen ces Atwood J.T. & Mora de Retana D.E., 1999: Family #39 Orchidaceae: Tribe Maxillarieae: sub- tribes Maxillariinae and . - Fieldiana, Bot., n.s. 40: i-vi, 1-182.

E n d r e s A.R., 1874: Letter to Capt. J. Dow, January 11, 1874. - John Melmoth Dow Papers. Division of Rare and Manuscript Collections, Comell University Library. H eller F., 2001: Orchidaceae. - In: Stevens W.D., U llo a U. C., P o o l A. & M o n tiel O.M.: Flora de Nicaragua. - Monogr. Syst. Bot. Missouri Bot. Gard. 85.

J e n n y R., 2010: The Stanhopea book. - Quito: Imprenta Mariseal. Kränzlin F., 1922: Orchidaceae-Monandrae, Tribus Oncidiinae - Odontoglosseae Pars II. - In: A. Engler (ed.): Das Pflanzenreich, Vol. 4, 50. Kränzlin F., 1925: Monographie der Gattungen Masdevallia Ruiz et Pavon. - Repert. Sp. Nov. Regni Veg. Beih. 34. Linden J., 1882: Plantes introduites ou mises pour la premiere fois dans le commerce. -111. Hort. 29: 98-100. Luer C.A., 2003: Lepanthes. - In: Hammel B.E., Grayum M.H., H errera C. & Zam ora N. (eds.): Manual de plantas de Costa Rica, vol. 3. - Monogr. Syst Bot. Missouri Bot. Gard. 93: 216-255 Manning S., 2010: Discovering New World orchids. -Nantwich, Cheshire: Manning Publ. M ansfeld R., 1931: Blütenanalysen neuer Orchideen von R. Schlechter. II. Mittelamerikani­ scher Orchideen. - Repert. Sp. Nov. Regni Veg. Beih. 49 (2). Ossenbach, C., Pupulin F. & JennyR., 2010: Orchid itineraries of Augustus R. Endres in Cen­ tral America: a biographic and geographic sketch. - Lankesteriana 10(1): 19—47. Pupulin F., 1995: A revision of the genus Trichocentrum (Orchidaceae: Oncidiinae). - Lind­ leyana 10 (3): 183-210. Pupulin F., 2001: Contributions to a reassessment of Costa Rican (Orchidaceae). The genus Kefersteinia Rchb.f. - Ann. Naturhist. Mus. Wien, B, 103: 525-555. Pupulin F., 2007: Contributions toward a reassessment of Costa Rican Zygopetalinae (Orchi­ daceae) 3. A systematic revision of Dichaea in Costa Rica. - Harvard Pap. Bot. 12 (1): 15-153.

Pupulin F., 20 1 0 : Flora Costaricensis. Family # 3 9 Orchidaceae: Tribe : Subtribe Zygopetalinae. - Fieldiana, Bot. n.s. 49: 1 -6 0 . Pupulin F. & B ogarin D., 2010: Illustration and studies in Neotropical Orchidaceae. The Lep­ anthes jimenezii group (Pleurothallidinae). - Harvard Pap. Bot. 15 (1): 111-121. Reichenbach H.G., 1873: New Garden Plants. - Gard. Chron. 1873: 1597. Reichenbach H.G., 1878: Stanhopeapulla. - Xenia Orchidacea 3: pl. 205.