BORNEO JULY 11–29, 2017

Western Tarsier photographed by Jimmy Stanley during our night-cruise on the Kinabatangan River.

LEADER: MACHIEL VALKENBURG LIST COMPILED BY: MACHIEL VALKENBURG

VICTOR EMANUEL NATURE TOURS, INC. 2525 WALLINGWOOD DRIVE, SUITE 1003 AUSTIN, TEXAS 78746 WWW.VENTBIRD.COM By Machiel Valkenburg

Our 2017 Borneo tour started in Kota Kinabalu, a lovely and picturesque city located on the South China Sea and on the western part of Sabah. On our first day we explored the Kota Belud area; due to the tip of a tail of a cyclone near Philippines, it had been raining the previous days, resulting in a very wet wetland with many interesting . More or less all possible egrets were found with Javan Pond-Heron, Rufous (Nankeen) Night-Heron, and Cinnamon Bittern being the best. The last was seen wonderfully, hunting in some rice fields. Some other species of birds related to water found during our outing were Buff-banded Rail, Watercock, Whiskered Tern, and some early returning Oriental Pratincoles. In the afternoon we headed for the beach where Green Imperial- Pigeon, Scaly-breasted Munia, and Blue-naped Parakeet were quickly found and gave good scope views. Around the hotel parking area, Collared Kingfisher and Sunda Woodpecker gave us a good show; these birds are truly magnificent.

Black-crowned Night-Herons were seen on a few occasions.

After leaving the city of Kota Kinabalu, we headed for Mount Kinabalu by way of the Rafflesia Center in the Crocker Range. The coming days meant mountain birding, but we were very unfortunate with a lot of rain, leftover from the previously mentioned cyclone. We did what we could and still got good looks at Bornean Bulbul, Mountain Barbet, and Bornean Leafbird in the Crocker Range—all of them desired endemics—as well as Golden-bellied Gerygone and Black- and-crimson Oriole. In the endemic-filled Kinabalu Park, the views were very poor, as this amazing was completely fogged in. During these days, we had a glimpse of the blue sky on only a few occasions, and when we did, we enjoyed some great birding. Some of the best birds we had splendid views of were Mountain Imperial-Pigeon, Bornean , Golden-naped Barbet,

Victor Emanuel Nature Tours 2 Borneo, 2017 Whitehead’s Broadbill, Gray-chinned , and Bornean . We walked over the main road and discovered a group of Sunda and Chestnut-hooded laughingthrushes foraging in the grassy patches along the road. The excitement level rose quickly when the endemic Bornean Green- was discovered; all of us had great views. All the way up at the gate of the mountain, a large group of Mountain Black-eyes gave us stiff necks from looking up; for that reason alone, a Bornean Stubtail must have decided to give us some excellent ground views of his features—he was so close we could almost touch him!

Wallace´s Hawk-Eagle was named after the legendary British naturalist Alfred Russel Wallace.

After some good days we left the mountains behind us and cruised for the lowlands. We all welcomed the warmth, and the birding proved to be hot as well. Sepilok was the first lowland destination; here the famous canopy walk was birded on four different occasions. This year many raptors were seen from the canopy with Besra being the best Bornean record, but Wallace’s Hawk- Eagle gave the best show as it ripped apart a big rat, with our group as first row spectators. An amazing experience! The first hornbills of the trip also passed through our bins with Oriental Pied- and Black Hornbill showing wonderfully. These two small hornbill species showed themselves abundantly. Several colorful species such as Blue-throated Bee-eater, Dollarbird, and White- crowned Shama were around. One evening we had the pleasure of taking a night walk in the reserve, and we were fortunate to find a flying Red Giant Flying Squirrel; some also saw the Western Tarsier. The resident Hooded Pitta was sleeping on a vine over the main road—very colorful and stunning! Naturally, owls also played a part this evening, and Oriental Bay-Owl, Sunda Scops-Owl, Reddish Scops-Owl, and Brown Boobook were either heard or seen.

Victor Emanuel Nature Tours 3 Borneo, 2017 We continued our travels towards the Kinabatangan River by way of the Gomantong caves. Around the limestone cave system we came across Black-and-red Broadbill and the similar stunning Black-and-yellow Broadbill, just before we enjoyed a large group of Maroon Langurs. This endemic species of langur was foraging quickly along the road and gave us a lot of time to observe its behavior and take many pictures. Our first primate of the trip was spotted here along the boardwalk: a female with young Bornean Orangutan was right above us and gave excellent photographic opportunities. Around five, the first Wrinkle-lipped Bats emerged from the cave, immediately initiating the evening feast of a pair of Bat Hawks.

Bornean Pygmy Elephants were seen foraging along the river.

Victor Emanuel Nature Tours 4 Borneo, 2017 In Kinabatangan (Sukau) we cruised the river, and this easy birding was thoroughly enjoyed by the entire group. In the narrow creeks you get very close to the birds, as on the overhanging branches attractive species such as Rufous-backed Dwarf-Kingfisher, Stork-billed Kingfisher, and Oriental Darter are common. We saw no less than five different individuals of Storm’s Stork, which is around 2% of the world’s population, currently estimated at 260–300 individuals left in the wild. Hornbills are a main attraction in this part of Borneo, and we saw four species, enjoying them to the fullest. We spotted a couple of endemic Bornean Pygmy Elephants feeding on the fresh grass along the river. In the evenings the took over, and on our night cruises we found a Common Palm Civet and three Malayan Colugos (Flying Lemurs), and a Slow Loris was a major highlight! We also came across a Buffy Fish-Owl that had just caught a large rat!

The endangered Storm’s Stork has a population of approximately 300 individuals left.

After the river area, we departed for the famous Danum Valley, which holds the oldest rainforest on our planet. The lavish Borneo Rainforest Lodge was our home for 4 full days (5 nights). From this beautiful lodge located in the middle of the rainforest we explored the surrounding using an extended trail system. During our days in the field we had many highlights, but some of the most memorable were an eye-to-eye meeting with a splendid male Blue-headed Pitta, which was very curious and sat on a vine for about 5 minutes studying our movements; a group of 8 Bornean Bristleheads, which were spotted from the amazing canopy walkway—we observed their movements and behavior for about half-an-hour; the calls of the Great Argus, a thrill for leader Machiel, were a simply wonderful sound in the early mornings; and extended magical views of a couple of Great Slaty Woodpeckers—full frame views were observed through the scope by all! Many species of abundant bulbuls and babblers, of which we saw most, some through the scope but most by bins, are another great feature of Danum. During our evenings we did not stop

Victor Emanuel Nature Tours 5 Borneo, 2017 exploring but continued on several night walks and drives. The aim was to spot Civets, more Flying Squirrels, Frogmouths, and deer, and yes, we succeeded! The best sighting of all was surely a Binturong, or Bearcat; this fig-eating was spotted high up in a fruiting tree near the lodge. All of these fantastic wildlife sightings were made from our amazing lodge, which I’d like to praise once again, as it is probably one of the best wildlife lodges in Asia! All the excellent meals in the lodge were accompanied by the sounds of calling Bornean Gibbons and honking Rhinoceros Hornbills in the background. An amazing time was had by all, and I cannot wait to return next year!

On our last day we drove back to Lahad Datu from where we flew back to Kota Kinabalu. Our final dinner was lovely, and it was time to say goodbye to this excellent group. I hope to see you all again soon and hope to have more wildlife adventures with you! Final praise goes to our knowledgeable and expert local staff who made our stay such a wonderful experience.

Our lovely group posing in the oldest rainforest on our planet, the Danum Valley.

Victor Emanuel Nature Tours 6 Borneo, 2017

BIRDS: This Checklist order, and common and scientific names are those given by James F. Clements in The Clements Checklist of Birds of the World, 6th Edition, Cornell University Press, with all current updates. Possible or debated species splits and alternative Common and Scientific names (particularly those of the Oriental Club) are given in ( ) or included in heavily bordered boxes. (I) = introduced species. (E) = Endemic.

Ducks, Geese & Waterfowl Wandering Whistling-Duck Dendrocygna arcuata

Pheasants, Grouse & Allies Red-breasted Partridge Arborophila hyperythra Chestnut-necklaces Partridge Arborophila charltonii Great Argus Argusianus argus Heard only Crimson-headed Partridge Haematortyx sanguiniceps Heard only

Storks Storm’s Stork Ciconia stormi Lesser Adjutant Leptopilos javanicus

Anhingas Oriental Darter Anhinga melanogaster

Grebes Little Grebe Tachybaptus ruficollis

Herons, Egrets & Bitterns Yellow Bittern Ixobrychus sinensis Cinnamon Bittern Ixobrychus cinnamomeus Purple Heron Ardea cinerea Great Egret Ardea (Egretta) alba Intermediate Egret Mesophoyx intermedia Little Egret Egretta garzetta Pacific Reef-Heron Egretta sacra Cattle Egret (Eastern Cattle Egret) Bubulcus ibis (coromandus) Javan Pond-Heron Ardeola speciose Striated (Little) Heron Butorides striata Black-crowned Night Heron Nycticorax nycticorax Rufous (Nankeen) Night-Heron Nycticorax caledonicus

Hawks, Eagles & Kites Oriental Honey-buzzard Pernis ptilorhynchus Jerdon´s Baza Aviceda jerdoni Crested Serpent-Eagle Spilornis cheela Bat Hawk Macheiramphus alcinus Wallace’s Hawk-Eagle Nisaetus nanus

Victor Emanuel Nature Tours 7 Borneo, 2017 Rufous-bellied Eagle Lophotriorchis kienerii Black Eagle Ictinaetus malaiensus Crested Goshawk Accipiter trivirgatus Besra Accipiter virgatus Brahminy Kite Haliastur indus White-bellied Sea-Eagle Haliaeetus leucogaster Lesser Fish-Eagle Ichthyophaga humilis

Rails, Gallinules & Coots Buff-banded Rail Gallirallus philippensis White-breasted Waterhen Amaurornis phoenicurus Watercock Gallicrex cinerea Eurasian (Common Moorhen) Gallinula chloropus

Stilts & Avocets Black-winged Stilt Himantopus himantopus

Sandpipers & Allies Wood Sandpiper Tringa glareola Black-tailed Godwit Limosa limosa

Pratincoles & Coursers Oriental Pratincole Glareola maldivarum

Gulls, Terns & Skimmers Little Tern Sternula albifrons Whiskered Tern Chlidonias hybrida

Pigeons & Doves Rock (Feral) Pigeon Columba livia Spotted Dove Streptopelia chinensis Little Cuckoo-Dove Macropygia ruficeps Zebra Dove (I) Geopelia striata Little Green-Pigeon Treron olax Pink-necked (Green-) Pigeon Treron vernans Large Green-Pigeon Treron capellei Green Imperial Pigeon Ducula aenea Mountain Imperial Pigeon Ducula badia

Cuckoos Banded Bay Cuckoo Cacomantis sonneratii Plaintive Cuckoo Cacomantis merulinus Violet Cuckoo Chrysococcyx xanthorhynchos Square-tailed Drongo-Cuckoo Surniculus lugubris Black-bellied Malkoha Phaenicophaeus diardi Raffles’s Malkoha Phaenicophaeus chlorophaeus

Victor Emanuel Nature Tours 8 Borneo, 2017 Chestnut-breasted Malkoha Phaenicophaeus curvirostris Red-billed Malkoha Phaenicophaeus javanicus Bornean Ground-Cuckoo (E) Carpococcyx radiceus heard only Greater Coucal Centropus sinensis

Barn-Owls Oriental Bay-Owl Phodilus badius

Owls Reddish Scops-Owl Otus rufescens Mountain Scops-Owl Otus spilocephalus Heard only Sunda Scops-Owl Otus lempiji Buffy Fish-Owl Ketupa ketupa Brown Wood-Owl Strix leptogrammica Brown Boobook Ninox scutulata

Frogmouths Large Frogmouth Batrachostomus auritus Blyth´s Frogmouth Batrachostomus affinis

Swifts Silver-rumped Needletail Rhaphidura leucopygialis esculenta Bornean Swiftlet Collocalia dodgei Mossy-nest Swiftlet Aerodramus salangana

Treeswifts Gray-rumped Treeswift Hemiprocne longipennis Whiskered Treeswift Hemiprocne comate

Trogons Red-naped Trogon Harpactes kasumba Diard’s Trogon Harpactes diardii Heard only Scarlet-rumped Trogon Harpactes duvaucelii

Hornbills Helmeted Hornbill Buceros vigil Rhinoceros Hornbill Buceros rhinoceros Bushy-crested Hornbill Anorrhinus galeritus Black Hornbill Anthracoceros malayanus Oriental Pied Hornbill Anthracoceros albirostris Wreathed Hornbill Rhytoceros undulates Wrinkled Hornbill Rhabdotorrhunis corrugatus

Kingfishers Blue-eared Kingfisher Alcedo meninting

Victor Emanuel Nature Tours 9 Borneo, 2017 Rufous-backed Dwarf-Kingfisher Ceyx rufidorsa Banded Kingfisher Lacedo pulchella Stork-billed Kingfisher Pelargopsis capensis Collared Kingfisher Todiramphus chloris Rufous-collared Kingfisher Actenoides concretus Heard only

Bee-eaters Blue-throated Bee-eater Merops viridis

Rollers Dollarbird Eurystomus orientalis

Asian Barbets Brown Barbet (E) Calorhamphus fuliginosus Blue-eared Barbet Psilopogon duvaucelii Red-throated Barbet Psilopogon mystacophanos Golden-naped Barbet (E) Psilopogon pulcherrima Mountain Barbet (E) Psilopogon monticola Gold-faced Barbet Psilopogon chrysopsis

Woodpeckers Rufous Piculet Sasia abnormis Sunda Woodpecker Dendrocopus moluccensis White-bellied Woodpecker Drycopus javensis Banded Woodpecker Picus miniaceus Checker-throated Woodpecker Picus mentalis Rufous Woodpecker Micropterus brachyurus Buff-rumped Woodpecker Meiglyptes tristis Buff-necked Woodpecker Meiglyptes tukki Maroon Woodpecker Blythipicus rubiginosus Orange-backed Woodpecker Reinwardtipicus validus Gray-and-buff Woodpecker Hemicircus concretus Great Slaty Woodpecker Mulleripicus pulverulentus

Falcons & Caracaras White-fronted Falconet (E) Microhierax latifrons

Old World Parrots Long-tailed Parakeet Psittacula longicauda Blue-naped Parrot (I) Tanygnathus lucionensis Blue-crowned Hanging-Parrot Loriculus galgulus

African & Green Broadbills Whitehead’s Broadbill Calyptomena whiteheadi

Asian & Grauer’s Broadbills

Victor Emanuel Nature Tours 10 Borneo, 2017 Black-and-red Broadbill Cymbirhynchus macrorhynchus Banded Broadbill Eurylaimus javanicus Black-and-yellow Broadbill Eurylaimus ochromalus

Pittas Black-crowned Pitta (E) Erythropitta ussheri Bornean Banded-Pitta Hydrornis schwaneri Blue-headed Pitta Hydrornis baudii Hooded Pitta Pitta sordida

Thornbills & Allies Golden-bellied Gerygone (Flyeater) Gerygone sulphurea

Vangas, Helmetshrikes & Allies Black-winged Flycatcher-shrike Hemipus hirundinaceus Rufous-winged Philentoma Philentoma pyrhoptera Maroon-breasted Philentoma Philentoma velata

Woodswallows White-breasted Woodswallow Artamus leucorynchus

Bristlehead Bornean Bristlehead (E) Pityriasis gymnocephala

Ioras Common Iora Aegithina tiphia Green Iora Aegithina viridissima

Cuckooshrikes Fiery Minivet Pericrocotus igneus Gray-chinned Minivet Pericrocotus solaris Scarlet Minivet Pericrocotus speciousus Sunda Coracina larvata Pied Triller Lalaga nigra Lesser Cuckooshrike Lalage fimbriata

Whistlers & Allies Bornean Whistler (E) Pachycephala hypoxantha

Vireos, Shrike-Babblers & Erpornis Pied (White-browed) Shrike-Babbler Pteruthius flaviscapis

Old World Orioles Dark-throated Oriole Oriolus xanthonotus Heard only Black-and-crimson Oriole Oriolus cruentus

Victor Emanuel Nature Tours 11 Borneo, 2017 Drongos Ashy Drongo Dicrurus leucophaeus Hair-crested Drongo Dicrurus hottentottus Greater Racket-tailed Drongo Dicrurus paradiseus

Fantails Spotted Fantail Rhipidura perlata Malaysian Pied-Fantail Rhipidura javanica White-throated Fantail Rhipidura albicollis

Monarch Flycatchers Black-naped Monarch Hypothymis azurea Blyth´s Paradise-Flycatcher Terpsiphone affinis

Crows, Jays & Crested Platylophus galericulatus Black Magpie Platylophus leucopterus Bornean Green-Magpie (E) jefferyi (E) cinerascens Slender-billed Crow enca

Swallows Pacific Swallow Hirundo tahitica

Nuthatches Sittidae Velvet-fronted Nuthatch Sitta frontalis

Bulbuls Puff-backed Bulbul Pycnonotus eutilotus Black-headed Bulbul Pycnonotus atriceps Straw-headed Bulbul Pycnonotus zeylanicus Leader only Bornean (Black-crested) Bulbul (E) Pycnonotus montis Yellow-vented Bulbul Pycnonotus goiavier Olive-winged Bulbul Pycnonotus plumosus Cream-vented Bulbul Pycnonotus simplex Red-eyed Bulbul Pycnonotus brunneus Spectacled Bulbul Pycnonotus erythropthalmus Hairy-backed Bulbul Tricholestes criniger Finsch´s Bulbul Alophoixus finschii Ochraceous Bulbul Alophoixus ochraceus Gray-cheeked Bulbul Alophoixus bres Yellow-bellied Bulbul Alophoixus phaeocephalus Buff-vented Bulbul Lole olivacea Streaked Bulbul Ixos malaccensis

Bush-Warblers & Allies

Victor Emanuel Nature Tours 12 Borneo, 2017 Bornean Stubtail Urusphena whiteheadi cucullatus Heard only Sunda Bush-Warbler vulcania

Leafwarblers Mountain Warbler Phylloscopus trivirgatus Yellow-breasted Warbler Seicercus montis

Grassbirds & Allies Striated Grassbird Megalurus palustris

Cistocolas & Allies Ashy Tailorbird Orthotomus ruficeps Rufous-tailed Tailorbird Orthotomus sericeus Yellow-bellied Prinia Prinia flaviventris

White-eyes, Yuhinas & Allies Chestnus-crested Yuhina (E) Yuhina everetti Mountain Black-eye (E) Chlorocharis emiliae Black-capped White-eye Zostrerops atricapilla

Tree-, Scimitar-Babblers & Allies Bold-striped Tit-Babbler Mixornis bornensis Fluffy-backed Tit-Babbler Macronus ptilosus Chestnut-winged Babbler Cyanoderma erythropterum Chestnut-backed Scimitar Babbler Pomatorhinus montanus Chestnut-rumped Babbler Stachyris maculate Gray-throated Babbler Stachyris poliocephala

Ground Babblers & Allies Moustached Babbler Malacopteron magnirostre Sooty-capped Babbler Malacopteron affine Scaly-crowned Babbler Malacopteron cinereum Rufous-crowned Babbler Malacopteron magnum Black-capped Babbler Pellorneum capistratum White-chested Babbler Pellorneum rostratum Ferruginous Babbler Pellorneum bicolor Striped Wren-Babbler Kenopia striata Bornean Wren-Babbler (E) Ptilocichla leucogrammica Horsfield´s Babbler Turdinus sepiarius Black-throated Wren-Babbler (E) Turdinus atrigularis Mountain Wren-Babbler (E) Turdinus crassus

Laughingthrushes & Allies Brown Fulvetta Alcippe brunneicauda Sunda Laughingthrush Garrulux palliatus

Victor Emanuel Nature Tours 13 Borneo, 2017 Bare-headed Laughingthrush (E) Garrulux (Melanocichlia ) calcus Chestnut-hooded Laughingthrush (E) Ianthocincla treachery

Fairy-bluebirds Asian Fairy-bluebird Irena puella

Old World Flycatchers Dark-sided Flycatcher Muscicapa sibirica Asian Brown Flycatcher Muscicapa latirostris -Robin Copsychus saularis White-rumped Shama Copsychus malabaricus (White-crowned Shama) (E) (C. malabaricus stricklandii) White-tailed Flycatcher Cyornis concretus Malaysian Blue-Flycatcher Cyornis turcosus Bornean Blue-Flycatcher (E) Cyornis superbus Indigo Flycatcher Eumyias indigo Verditer Flycatcher Eumyias thalassinus Eye-browed Jungle-Flycatcher Vauriella gularis White-browed Shortwing Brachypteryx montana Bornean Whistling-Thrush Myophonus borneensis White-crowned Forktail Enicurus leschenaultia (Bornean Forktail) (E. leschenaultia borneensis) Chestnut-naped Forktail Enicurus ruficapillis Heard only Snowy-browed Flycatcher Ficedula hyperythra Little Pied Flycatcher Ficedula westermanni Rufous-chested Flycatcher Ficedula dumetoria

Thrushes & Allies Orange-headed Thrush Geokickla citrina Fruit-hunter (E) Chlamydochaera jefferyi Heard only

Starlings Asian Glossy Starling Aplonis panayensis Common Hill Myna Gracula religiosa Javan Myna Acridotheres javanicus

Leafbirds Greater Green Leafbird Chloropsis sonnerati Lesser Green Leafbird Chloropsis cyanopogon Bornean Leafbird (E) Chloropsis kinabaluensis

Flowerpeckers Yellow-breasted Flowerpecker Prionochilis maculatus Yellow-rumped flowerpecker (E) Prionochilis xanthopygius Yellow-vented Flowerpecker Dicaeum chrysorrheum Orange-bellied flowerpecker Dicaeum trigonostigma

Victor Emanuel Nature Tours 14 Borneo, 2017 Black-sided Flowerpecker Dicaeum monticolum Scarlet-backed flowerpecker Dicaeum cruentatum

Sunbirds & Spiderhunters Ruby-cheeked Sunbird Chalcoparia (Anthreptes) singalensis Plain Sunbird Anthreptes simplex Plain-(Brown)throated Sunbird Anthreptes malacensis Red-throated Sunbird Anthreptes rhodolaemus Copper-throated Sunbird Leptocoma calcostetha Olive-backed Sunbird Cinnyris (Nectarina) jugularis Temminck’s Sunbird Aethopyga temminckii Crimson Sunbird Aethopyga siparaja Long-billed Spiderhunter Arachnothera robusta Little Spiderhunter Arachnothera longirostra Purple-naped Spiderhunter Arachnothera hypogrammicum Yellow-eared Spiderhunter Arachnothera chrysogenys Spectacled Spiderhunter Arachnothera flavigaster Streaky-breasted Spiderhunter Arachnothera affinis

Old World Sparrow Eurasian Tree Sparrow (I) Passer montanus

Waxbills & Allies Dusky Munia (E) Lonchura fuscans Scaly-breasted Munia (I) Lonchura punctulata Chestnut Munia Lonchura atricapilla

MAMMALS

Tree Shrews glis Mountain Treeshrew Tupaia montana

Colugos (Flying Lemurs) Malayan Colugo Cynocephalus variegatus

Lorises, Pottos & Galagos Slow Loris Nycticebus coucang

Old World Fruit Bats Wrinkle-lipped free-tailed Bat Chaerephon plicatus Hairless Bat Cheiromeles torquatus

Victor Emanuel Nature Tours 15 Borneo, 2017 Tarsiers Western Tarsier (E) Tarsius bancanus

Old World Monkeys Maroon Langur (E) Presbytis rubicunda Silvered Langur Presbytis cristata Proposcis Monkey (E) Nasalis larvatus Long-tailed Macaque Macaca fasciciularis Southern Pig-tailed Macaque Macaca nemistrina

Primates Bornean Gibbon (E) Hylobates muelleri

Primates Bornean Orang-Utan (E) Pongo pygmaeus

Squirrels Pale Giant Squirrel Ratufa affinus Prevost’s Squirrel Callosciurus prevostii Plantain Squirrel Callosciurus notatus Bornean Black-banded Squirrel (E) Callosciurus orestes Jentink’s Squirrel (E) Sundasciurus jentinki Borean Mntn. Ground Squirrel (E) Dremomys everetti Whitehead´s Pigmy Squirrel (E) Exilisciurus whiteheadi Plain Pigmy Squirrel (E) Exilisciurus exilis Thomas’s Flying Squirrel (E) Aeromys thomasi Red Giant Flying Squirrel Petaurista petaurista

Civets, Genets & kin Malay Civet Viverra tangalunga Common Palm Civet Paradoxurus hermaphrodites Binturong Arctictis binturong

Elephants Bornean Pygmy Elephant (E) Elaphas borneensis

Pigs Bearded Pig Sus barbatus

Deer Lesser Mouse Deer Tragulus javanicus Barking Deer Muntiacus muntjac Sambar Deer Cervus unicolor

REPTILES Saltwater (Estuarine) Crocodile Crocodylus porosus

Victor Emanuel Nature Tours 16 Borneo, 2017 Crested Green Lizard Bronchocela cristatella Yoshi’s Bent-toed Gecko Crytodactylus yoshii Asian House Gecko Hemidactylus frenatus Water Monitor Lizard Varanus salvator

Victor Emanuel Nature Tours 17 Borneo, 2017