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Geologica Hungarica Series Palaeontologica 60. Fasc. (2018.)

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Geologica Hungarica Series Palaeontologica 60. Fasc. (2018.) Geologica Hungarica series Palaeontologica, Fasciculus 60 47 Systematic descriptions In the following descriptions the systematics developed by TOZER (1981) is used, with a few exceptions. The genera Lardaroceras, Asseretoceras and Megaceratites, introduced afterwards, are systematically arranged according to the opinion of their author BALINI (1992a, b). Hyparpadites, due to its fastigate venter, is here removed from Arpaditidae to Paraceratitinae. Moreover, following the opinion of MANFRIN et al. (2005), Ticinites is removed from Danubitidae to Bulogitinae, Parasturia is removed from Sturiidae to Ptychitidae and, following MONNET & BUCHER (2005), Tropigymnites is removed from Japonitidae to Gymnitidae. The signs of open nomenclature were used according to BENGTSON (1988), except regarding the uncertainty of the generic attributions; here the opinions of RICHTER (1943) and MATTHEWS (1973) were followed, i.e. question marks “?” after the name, instead of “aff.” in front of the name, were used. The signs attached to the synonymy lists were applied from RICHTER (1943, p. 40–42) and MATTHEWS (1973). The majority of the material is deposited in the Department of Palaeontology and Geology of the Hungarian Natural History Museum (HNHM) under the inventory num- bers prefixed by M., INV, or PAL, and in the palaeontological collection of the Mining and Geological Survey of Hungary (MGSH) under the inventory numbers prefixed by T. A few figured specimens are kept in the private collections of K. TAMÁS (Kővágóörs) and L. VARGA (Úny), without inventory numbers. The author made comparative studies on ammonoid specimens deposited in several Figure 18. The axes of measure- museums abroad; the names of the collections and their acronyms (used in the present mono- ments (D = diameter, WH = whorl- graph) are the following: Geologische Bundesanstalt, Wien (GBAW), Natur historisches height, WW = whorl-width, U = diameter of umbilicus), and the ter- Museum, Wien (NHMW), Bayerische Staatssammlung, München (BSM), Paläonto - minology of nodes, used in the logisches Institut und Museum, Universität Zürich (PIMUZ), Natural History Museum, present monograph London (NHML). The dimensions of the measured ammonoid specimens (D = diameter, WH = whorl-height, WW = whorl-width, U = dia - meter of umbilicus) are given in millimetres (Figure 18). Order Ceratitida HYATT, 1884 Superfamily Noritoidea KARPINSKY, 1889 Family Noritidae KARPINSKY, 1889 Genus Norites MOJSISOVICS, 1878 Type species: Norites gondola (MOJSISOVICS, 1869) Norites gondola (MOJSISOVICS, 1869) Plate I: 1–5. * 1869 Ammonites Gondola MOJS. nov. sp. — MOJSISOVICS, Cephalopoden-Fauna des alpinen Muschelkalkes, p. 584, pl. XV, fig. 3a–b. 48 v 1900 [Ceratites] cordevolicus MOJS.— DIENER, Muschelkalk–Cephalopoden südl. Bakony, p. 25, pl. II, fig. 3. v 1901 Norites gondola MOJS. var. nov. — REIS, Fauna des Wettersteinkalkes I., p. 90, pl. IV, figs 24, 25. v 1903 Norites Dieneri ARTH. — ARTHABER, Neue Funden Muschelkalk des südl. Bakony, Revision, p. 26, pl. I, fig. 9. ? 1915 Norites gondola MOJS. — ARTHABER, Die Trias von Bithynien, p. 144, pl. XVII, fig. 3. v 1934 Norites gondola (MOJSISOVICS) — SPATH, Ammonoidea of the Trias, p. 281, fig. 98. v 1973 Norites dieneri ARTHABER, 1903 — RIEBER, Grenzbitumenzone, p. 71, pl. 17, figs 17, 18, 20. v 1989 Norites dieneri ARTHABER, 1903 ? — VÖRÖS & PÁLFY, Vászoly, p. 19. 1996 Norites dieneri ARTHABER, 1903 — FANTINI SESTINI, Calcare di Esino 2, p. 216, pl. 1, fig. 10. v 1998 Norites dieneri ARTHABER, 1903 — VÖRÖS, Balaton-felvidék, p. 26, 35, 38, 42. v 2003 Norites gondola (MOJSISOVICS, 1869) — VÖRÖS, Pelsonian, p. 79, pl. A–I, figs 5–7, text-fig. A–9. (cum syn.) 2008 Norites gondola (MOJSISOVICS 1869) — MONNET et al., Giudicarie, p. 69, text-figs 6, 7. Material 15 specimens of various state of preservation, from Felsőörs (1), Vörösberény (1), Szentantalfa (1), Vászoly (4), Mencshely (6), Sóly (1) and Tótvázsony (1). Measurements Description The specimens are rather small for the genus. The coiling is very involute; the whorl-section is high and narrow trapez - oidal. The umbilicus is narrow and deep. The umbilical wall is steep and forms a rather sharp shoulder with the flanks. The flanks are gently and evenly convex and meet the flat venter with a marked ventrolateral shoulder, carrying a definite, faint keel. The ornamentation is very weak. The details of the suture were not studied; a weathered specimen (Plate I: 5) shows a ceratitic suture with at least five lat- eral lobes at 27 mm diameter. Remarks N. gondola is the type species of the genus Norites. Its middle Anisian (Pelsonian) occurrences and the relationships to other Pelsonian forms of Norites were discussed in detail by VÖRÖS (2003). N. dieneri ARTHABER, 1903 was taken as an independent species by some authors, including VÖRÖS (1998) mainly on the basis of its larger size and its occurrence in the higher Anisian beds (Avisianum Subzone), in contrast to N. gondola which was recorded prevalently from the Pelsonian. However the new collections at the Balaton Highland revealed N. gondola in some transitional horizons (Camunum, Pseudohungaricum and Felsoeoersensis subzones). The extremely large size of ARTHABER’s type specimen of N. dieneri is not considered enough to distinguish it as a separate species. The specimen figured as N. gondola by ARTHABER (1915, l. c.) seems to be poorly preserved and its ventrolateral should - er is rather rounded, without “keel”; therefore this item is included to the synonymy only with query. Several specimens of N. gondola listed by SPATH (1934) were examined in the collection NHML (London) and most of them (C. 5458, C. 13896, C. 21037–9, C. 23063, C. 37191 and C. 37206) are here regarded as proper representatives of that species. Distribution N. gondola was described from the Anisian Schreyeralm Limestone of the Northern Calcareous Alps. It was found also in Anisian localities of the Northern Calcareous Alps, the Southern Alps and the Dinarides (Han Bulog Limestone). At the Balaton Highland it ranges from the Pelsonian Balatonicus Subzone to the Illyrian Avisianum Subzone. Superfamily Ceratitoidea MOJSISOVICS, 1879 Family Ceratitidae MOJSISOVICS, 1879 Subfamily Beyrichitinae SPATH, 1934 Genus Beyrichites WAAGEN, 1895 Type species: Beyrichites reuttensis (BEYRICH, 1867) 49 Beyrichites cf. reuttensis (BEYRICH, 1867) Plate I: 6. * 1867 Ammonites Reuttensis. — BEYRICH, Über einige Cephalopoden, p. 113, pl. I, fig. 4a–c. v 1998 Beyrichites cf. reuttensis (BEYRICH, 1867) — VÖRÖS, Balaton-felvidék, p. 26, 59. v 2003 Beyrichites cf. reuttensis (BEYRICH, 1867) — VÖRÖS, Pelsonian, p. 95, pl. A–VII, fig.10 (cum syn.). Material Four fragmentary specimens from Szentantalfa. Measurements Description The specimens are medium-sized for the genus. Compressed shells with rather involute coiling. The whorl-section is high oval. The umbilical shoulder is rounded. The flank is gently convex and passes gradually into the highly arched, narrow venter. The ornamentation is weak; it consists of widely spaced, sinuous to falcoid ribs fading out near the venter. In the outer part of the flank, very weak secondary ribs are inserted between the primaries. Suture lines were not seen. Remarks The middle Anisian (Pelsonian) occurrences of B. reuttensis and the relationships to other species of Beyrichites were discussed in detail by VÖRÖS (2003). The figures published by VENZO & PELOSIO (1968, l. c.) are regarded as the most perfect and reliable illustrations of B. reuttensis. Distribution B. reuttensis was described from the Anisian of the Southern Alps. It is also known from the Northern Calcareous Alps (Wetterstein Limestone), the Dinarides and the Caucasus. At the Balaton Highland it ranges from the Illyrian Binodosus Subzone to the Pseudohungaricum Subzone. Genus Lardaroceras BALINI, 1992 Type species: Lardaroceras krystyni BALINI, 1992 Lardaroceras krystyni BALINI, 1992 Plate I: 9. * 1992 Lardaroceras krystyni sp. n. — BALINI, Lardaroceras, p. 12, text-figs 4A, 5A–E, pl. 1, figs 1–5. 1993 Lardaroceras krystyni BALINI, 1992 — GAETANI (ed.), Anisian/Ladinian boundary field workshop, p. 64, pl. 3, fig. 1. v 1998 Lardaroceras krystyni BALINI, 1992 — VÖRÖS, Balaton-felvidék, p. 20, 22, 59 (partim), (non pl. I, fig. 7). v 2010 Lardaroceras krystyni BALINI, 1992 — VÖRÖS, North Hungary, p. 6, pl. I, figs 4, 5. Material 50 specimens of various state of preservation, from Felsőörs (4), Vörösberény (10), Szentantalfa (6), Szent király - szabadja (2), Vászoly (23), Szentbékkálla (4), and Iszkaszentgyörgy (1). Measurements Description Small to large Lardaroceras with moderately involute, compressed conch. The whorl-section is high oval to subtrape- zoidal. The umbilical wall and the umbilical margin are poorly seen. The flanks are gently convex, almost flat and meet the arched venter at a blunt ventrolateral margin. The venter bears a definite rounded keel separated from the ventrolateral 50 shoulders. The ornamentation consists of weak, mostly sinuous radial ribs and nodes. The prominent umbilical nodes elev - ate from the umbilical margin. There are no lateral nodes. The ventrolateral nodes (around 20 on a half whorl) are of medium strength; on the body chamber they appear as somewhat adorally elongated clavi. Suture lines are not visible. Remarks This is the type species of the genus Lardaroceras and, according to the detailed description of BALINI (1992b, l. c.), it differs from the other species of that genus by the absence of lateral nodes. Previously this feature was not considered by the
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