Winter 2012/2013 Issue 12 Volume 2 2012 Summer Paleoinventory Results (Continued) Most of the Fossils Are Animals Organisms

Total Page:16

File Type:pdf, Size:1020Kb

Winter 2012/2013 Issue 12 Volume 2 2012 Summer Paleoinventory Results (Continued) Most of the Fossils Are Animals Organisms Great Basin National Park Park News National Park Service U.S. Department of the Interior The Midden The Resource Management Newsletter of Great Basin National Park Amazingly Successful Summer Paleontology Inventory During my first year after moving to GRBA, I had found a few good indications in the park’s backcountry. Also, park staff had recognized fossils in some of the remote drainages (see the Summer 2011 NPS Photo Midden article, “Significant index NPS Photo fossils found in park”). Some of the more than 1000 fossil In addition, geologic maps indicated specimens found this summer included these brachiopods, Orthambonites broad areas of rock outcrops within michaelis; Kanosh Shale, Ordovician the park where there was high An internal mold of a complete coiled Period. potential for fossils to be found. nautiloid cephalopod surprisingly similar to the living chambered nautilus. The dark By Gorden Bell, Supervisory These rocks are of the same age mass in the center is a bryozoan colony Environmental Protection Specialist and formations that are extremely that encrusted the shell after it fell to the and Paleontologist fossiliferous near Crystal Peak, Utah, seafloor; Lehman Formation, Ordovician Period. and in central Nevada. So, as the park If one were to judge the potential of is situated directly between those began to build an inventory of Great Basin National Park (GRBA) areas, it would seem likely that our paleontological resources heretofore to possess a significant amount of rocks might have the same kinds and unknown within the park. Over the paleontological resources based quantities of fossils. course of the summer we added 38 on available scientific literature, new paleontology localities to the it would not appear to be a good This past summer we were able to database, represented by 476 GPS bet. As part of my collateral test that possibility when we brought positions and more than 1000 fossil duties, I had been given the task in two Geoscientists-in-the Parks specimens. of first assessing the likelihood of interns (GIPs) through a program significant paleontological resources sponsored jointly by the NPS and So, what types of fossils are present being present within the park and the Geological Society of America in the park? Continued on Page 2 then searching for them if the results GeoCorps program. The two GIPs, appeared promising. Linda Sue Lassiter and Spencer Holmes, are enrolled in geology In This Issue My survey of the literature had degree programs at the University of turned up only one publication that Northern Arizona and California State Paleontology Inventory................1 identified fossil localities inside University - Chico, respectively. Rattlesnake Relocations.................4 the park boundaries. Another three Lehman Cave Restoration..............5 identified fewer than a half-dozen During an intense 12-week internship, 2012 Diptera BioBlitz.......................6 fossil localities in the southern these two intrepid students and I Forest Health.......................................8 Snake Range but outside of the park. covered approximately 600 hectares Johnson Lake Mine Recorded.....10 However, numerous publications (1500 acres) of the park, searching Fire Needed.....................................11 detailed exciting discoveries in the rock outcrops for fossils and using Remote Cameras.............................12 mountain ranges to the east, south, GPS units and cameras to document 2011 BioBlitz Update.....................14 and west of the park. what we found. Gradually, we Upcoming Events....................14 Winter 2012/2013 Issue 12 Volume 2 2012 Summer Paleoinventory Results (continued) Most of the fossils are animals organisms. We also spotted many Probably one of the most exciting and plants that lived in a marine types of molluscs, including some finds was a thick bed of corals at an environment in a relatively shallow clams, many species of marine elevation of almost 3350 m (11,000 sea that alternately covered and then snails, and plenty of straight ft), north of Granite Peak. About receded from this area many times nautiloids, which are cephalopods 470 million years ago during the over. Most are very old, spanning like squids and octopi but which Ordovician Period, living corals portions of the Cambrian and had a long conical external shell. may have covered the local sea Ordovician geologic periods from We did find one coiled nautiloid, floor like a patchwork blanket that about 510 to 470 million years ago. which does not appear to be much stretched for many kilometers. different than the chambered Geologists would call this type We found strange types of algae nautilus living in modern oceans of accumulation a “biostrome.” called receptaculitids that have today. While the bed we found would mineralized rod-shaped spicules not technically qualify as a reef forming an internal skeleton like because it only stood one to two that found in some sponges. One metes (three to six feet) above the type, Receptaculites, as shown in the seafloor, it nonetheless must have Summer 2011 issue of The Midden, functioned as reef-like habitat. The is shaped like a biscuit while another NPS Photo corals that grew in this biostrome, type, called Calathium, is conical Eofletcheria and Foerstephyllum, like an empty ice cream cone and are two of the three earliest forms often formed mounded colonies. The straight shell of a type of nautiloid of corals known in the fossil record. cephalopod, sometimes called an orthoceracone; Lehman Formation, Both are classified as members of We found plenty of trilobites, which Ordovician Period. a stem group known as tabulate are common in the surrounding corals. Corals would have to evolve areas, including a couple of tiny We were even lucky enough to pick blind forms called agnostids. up a single plate from an animal known as a chiton, a slug-shaped mollusc with a row of eight hard plates armoring its back. We found lots of stalked echinoderms, aka NPS Photo NPS Photo “sea lilies,” such as crinoids that stood well above the sea floor to This large trilobite tail is an undescribed catch food particles floating in the species of Cybelopsis (the quarter used water and cystoids that scooted Crowded and overgrown masses of the as scale is exactly an inch in diameter); coral, Eofletcheria, indicating a reefy Lehman Formation, Ordovician Period. around in the bottom muds to find type of habitat; Lehman Formation, food. Ordovician Period. We also found brachiopods, which much greater complexity and have bivalved shells. Some of these longevity before they could grow lived in muddy burrows and have into the massive shapes we know phosphatic shells, while others as true reefs. We were able to trace had calcium carbonate shells and one unbroken outcrop of this bed attached themselves to various NPS Photo of coral for a distance of about living animals or dead shells. We 370 m (1200 ft) where its edges located bryozoans, distantly related are interrupted by erosion or by to brachiopods but living in tiny faulting. We could pick it out again coral-like colonies that either grew A single plate from a cystid echinoderm, in nearby outcrops at about the probably Hadrocystis; Pogonip Group, independently or encrusted other Ordovician Period. Continued on Page 3 2 The Midden 2012 Summer Paleoinventory Results (continued) same horizon, suggesting it was indeed connected while growing. As it turns out, Eofletcheria has been reported as biostromal accumulations at about the same stratigraphic level in outcrops from Crystal Peak, in NPS Photo Utah all the way to the White Pine Range at the western edge of White Pine County in Nevada, a distance of more than 160 km (100 mi) today (map figure). The first vertebrate fossil from the park, a partial fin spine of an acanthodian fish Based on calculations of the amount tentatively identified as Nodocosta denisoni; Sevy Dolomite, Devonian Period. of geological extension that has As a final note, the occurrence of specimen is shown above and occurred during faulting in the Great Foerstephyllum at GBNP is only the is a fragmentary fin spine of an Basin, that distance would have been second record from Nevada – the acanthodian fish. It was tentatively approximately 100 km (60 mi) at the first being from the Ordovician rocks identified by Dr. David Elliot of time the coral was growing. Other of the Pioche area. Northern Arizona University as observations note that Eofletcheria Nodocosta denisoni. The specimen is found in float blocks at about the And last but not least, this summer was found on outcrops of the Sevy same stratigraphic level not far north we also found the first vertebrate Dolomite which is Early Devonian of Big Springs on the southern end fossil from the park! It is not much age and is about 405-400 million of the Snake Range. This suggests to look at and is only 13 mm (0.5 years old. Appropriately, the that the coral patch habitat might in) long, but some people place Devonian Period of geologic time is have originally grown in an east-west more significance on vertebrate also known as the “Age of Fishes.” trending tract at least 100 km (60 fossils than invertebrate fossils. Our mi) long and 16 km (10 mi) wide. So, in reflection, the amount and quality of fossil resources we found in such a short amount of time is amazing given so little indication in the scientific literature. This is due in large part to the efforts and dedication of our two GIPs, Linda Sue and Spencer. I want to extend a rousing round of thanks to them. It is also appaerent that GRBA contains a wealth of paleontological resources, but had simply been overlooked by researchers. After this past summer I am confident that we will make even more exciting discoveries next summer and that this paleontological inventory may very well spark a new era of geological and paleontological research at Great Basin National Distribution map of coral biostrome localities in the uppermost Lehman Formation Park.
Recommended publications
  • Appendix a Table A.1. Information About Sampling and Management
    Appendix A Table A.1. Information about sampling and management types of the organic olive orchards. Orchard ID Type Year Month Sampling Locality DG1 Tilled 2011 6 1 Granada ON Tilled 2011 6 1 Deifontes DG2 Mowed 2011 6 1 Granada ODM Mowed 2011 6 1 Deifontes DG1 Tilled 2011 7 2 Granada ON Tilled 2011 7 2 Deifontes DG2 Mowed 2011 7 2 Granada ODM Mowed 2011 7 2 Deifontes DGA1 Tilled 2012 6 1 Granada DGB1 Tilled 2012 6 1 Granada ONA Tilled 2012 6 1 Deifontes ONB Mowed 2012 6 1 Deifontes ODM Mowed 2012 6 1 Deifontes DGA1 Tilled 2012 7 2 Granada DGB1 Tilled 2012 7 2 Granada ONA Tilled 2012 7 2 Deifontes ODM Mowed 2012 7 2 Deifontes ONB Mowed 2012 7 2 Deifontes DG1A Tilled 2013 6 1 Granada DG1B Tilled 2013 6 1 Granada DG3A Mowed 2013 6 1 Granada DG3B Mowed 2013 6 1 Granada ODM Mowed 2013 6 1 Deifontes ONB Mowed 2013 6 1 Deifontes ONA Mowed 2013 6 1 Deifontes DG1A Tilled 2013 7 2 Granada DG1B Tilled 2013 7 2 Granada DG3B Mowed 2013 7 2 Granada DG3A Mowed 2013 7 2 Granada ODM Mowed 2013 7 2 Deifontes ONA Mowed 2013 7 2 Deifontes ONB Mowed 2013 7 2 Deifontes Table A.2. Summary and formula* of the fitted generalized linear mixed-effects model (GLMM) with nested random effects fit by maximum likelihood with Laplace Approximation, including egg predation, type of management, and abundance of natural enemies (package “lme4”). Estimated Variable Variance SD coefficient SE z p Fixed effects: Intercept 4.723 0.3980 11.86 0.001 Management (tilled and mowed) 1.127 0.5299 2.12 0.033 Natural enemy 0.0002 0.00009 2.41 0.015 Random effects: Year : Site 1.100 1.049 * glmer (predated.eggs ~ management + natural.enemies + (1 | annual / ID), family = poisson) Table A.3.
    [Show full text]
  • Diptera Chamaemyiidae), an Overlooked Family of Biological Control Agents
    Bulletin of Insectology 68 (2): 173-180, 2015 ISSN 1721-8861 Seasonal habits of predation and prey range in aphidophagous silver flies (Diptera Chamaemyiidae), an overlooked family of biological control agents 1 2 3 1 4 2 Serdar SATAR , Alfio RASPI , Işıl ÖZDEMIR , Adnan TUSUN , Mehmet KARACAOĞLU , Giovanni BENELLI 1Department of Plant Protection, Faculty of Agriculture, University of Çukurova, Balcali, Adana, Turkey 2Department of Agriculture, Food and Environment, Insect Behaviour Group, University of Pisa, Italy 3Plant Health Central Research Institute, Yenimahalle, Ankara, Turkey 4Biological Control Research Station, Yüreğir, Adana, Turkey Abstract Aphids are among the most widespread and serious groups of pests in agro-ecosystems, and predaceous arthropods have been proposed as biological control agents against them, including parasitic Hymenoptera, lacewings, ladybugs, hoverflies and silver flies (Diptera Chamaemyiidae). Chamaemyiidae is a small family of predaceous flies, including aphidophagous and coccido- phagous species. Little is known about their ecology, and partial failures of Chamaemyiidae-based biological control programs against aphids may be due to poor synchronization of predator-prey seasonal habits. In this study, we investigated seasonality of predation and prey range of aphidophagous Chamaemyiidae. A field survey was conducted on crops and indigenous flora in seven agricultural sites in southern Turkey. Seventeen host plant families were surveyed for Chamaemyiidae presence and 371 silver flies were studied. All Chamaemyiidae larvae were observed to prey on adult and young instar aphids in the field. Silver fly total larval abundance reached a maximum in autumn and early winter. Six species were identified: Leucopis annulipes, L. formosana, L. glyphinivora, L. revisenda, L. rufithorax and L.
    [Show full text]
  • Diptera Communities of Raptor (Aves) Nests in Nova Scotia, Canada
    The Canadian Entomologist (2020), page 1 of 13 doi:10.4039/tce.2020.26 ARTICLE Diptera communities of raptor (Aves) nests in Nova Scotia, Canada Valerie Levesque-Beaudin1* , Bradley J. Sinclair2, Stephen A. Marshall3, and Randolph F. Lauff4 1Centre for Biodiversity Genomics, University of Guelph, 50 Stone Road E, Guelph, Ontario, N1G 2W1, Canada, 2Canadian National Collection of Insects and Canadian Food Inspection Agency, Ottawa Plant Laboratory – Entomology, Central Experimental Farm, 960 Carling Avenue, Ottawa, Ontario, K1A 0C6, Canada, 3School of Environmental Sciences, University of Guelph, 50 Stone Road E, Guelph, Ontario, N1G 2W1, Canada and 4Department of Biology, St. Francis Xavier University, 4130 University Avenue, Antigonish, Nova Scotia, B2G 2W5, Canada *Corresponding author. Email: [email protected] (Received 3 December 2019; accepted 9 March 2020; first published online 27 April 2020) Abstract The identity, richness, and abundance of true flies (Diptera) from the nests of three cavity-nesting raptors (Aves) were investigated in northern Nova Scotia, Canada. After fledging, flies were extracted from the nest mate- rial using Berlese funnels within an emergence chamber. Thirty-one species/morphospecies from 14 families were collected, including eight new records for Nova Scotia and two new records for eastern North America. Introduction Bird nests are micro-ecosystems with diverse communities of invertebrates, from ectoparasites to commensal species. Most studies of the arthropods in bird nests have focussed on the presence and impact of ectoparasites (Møller et al. 1990; Loye and Zuk 1991; Krištofík et al. 2001, 2002, 2003, 2007; Fairn et al. 2014), including fleas (Siphonaptera) (Phipps and Bennett 1974); mites (Acari) (Wasylik 1971); and nest-associated Diptera in the families Muscidae (Lowenberg-Neto 2008), Calliphoridae (Bennett and Whitworth 1991; Whitworth and Bennett 1992), and Carnidae (Cannings 1986a, 1986b; Dawson and Bortolotti 1997).
    [Show full text]
  • The Genera Meoneura and Carnus (Diptera: Carnidae) in Israel
    ISRAEL JOURNAL OF ENTOMOLOGY, Vol. 47, pp. 173–214 (27 December 2017) The genera Meoneura Nitzsch and Carnus Rondani (Diptera: Carnidae) in Israel, with the description of ten new species, new records and identification keys JENS -HERMANN STUKE 1 & AMNON FREIDBERG 2 1Roter Weg 22, Leer, 26789 Germany. E-mail: [email protected] 2The Steinhardt Museum of Natural History, Israel National Center for Biodiversity Studies, Tel Aviv University, Tel Aviv, 69978 Israel. E-mail: [email protected] ABSTRACT The Carnidae of the genera Meoneura and Carnus from the collection of the Tel Aviv University have been investigated. Ten new species are described from Israel: Meoneura bilboi n. sp., M. brakeae n. sp., M. davidi n. sp., M. goldemari n. sp., M. grimmorum n. sp., M. lilliputensis n. sp., M. meszarosi n. sp., M. nilsholgerssoni n. sp. (also from Egypt), M. oskari n. sp., and M. wichtelorum n. sp. Ten species—M. acuticerca, M. furcata, M. glaberrima, M. hungarica, M. lacteipennis, M. maritima, M. moravica, M. neottiophila, M. prima, and M. triangularis—are recorded in the country for the first time. Three new synonyms are introduced: Meoneura palaestinensis Hennig, 1937 = Meoneura nitidiuscula Collin, 1949 (n. syn.), Meoneura moravica Gregor & Papp, 1981 = Meoneura pamphylica Ozerov, 2008 (n. syn.) and Meoneura prima (Becker, 1903) = Meo- neura baluchistani Duda, 1936 (n. syn.). A total of 22 species of Meoneura and one species of Carnus are now known from Israel. Four identification keys are presented for species of Meoneura species groups. KEYWORDS: Diptera, Carnidae, Meoneura, Carnus, Filth flies, Israel, Egypt, new species, new synonymy, identification keys.
    [Show full text]
  • The Importance of Behavior and Venom System Morphology in Understanding Its Ecology and Evolution
    Toxins 2019, 11, 666; doi:10.3390/toxins11110666 S1 of S11 Supplementary Materials: The Diversity of Venom: The Importance of Behavior and Venom System Morphology in Understanding Its Ecology and Evolution Vanessa Schendel, Lachlan D. Rash, Ronald A. Jenner, and Eivind A. B. Undheim Table S1. Independently evolved venomous animal lineages and the primary ecological roles of their venoms. Taxa for which no direct support of their venomous nature could be found are shown in grey font. General Venom System Animal Group Venomous Lineage Primary Role References Morphology Predation, defense, Cnidarians All Nematocysts [1] intraspecific competition Coleoid Posterior and anterior glands, cephalopods, venom injected through salivary Predation [2,3] including octopus papilla. and squid Long duct/venom gland, venom Cone snails and injected through hollow radular Predation, [4] relatives (Conoidea) tooth on proboscis by a distal defense venom pump. Tritons, helmet Two-lobed salivary (venom) Molluscs shells, etc. glands that open through Predation [5] (Tonnoidea) common duct into buccal mass. Dwarf tritons, Single-lobed salivary (venom) including vampire glands that open through Predation [6] snails common duct into buccal mass. (Colubrariidae) Primary and accessory salivary Murex snails (venom) glands that open Predation [7] (Muricidae) through common duct into buccal mass. Proboscis with venom secreting cells, sometimes with stylet to Nemerteans Ribbon worms facilitate venom delivery Predation [8] (Enopla), or pseudocnidae with a potential role in venom delivery. Toxin-producing “lappets” secreting venom into large Blood worms muscular and glandular venom Predation [9] (Glyceridae) reservoir, which is presumably Annelids also involved in venom expulsion. Secretory cells dispersed along Predation, Leeches (Hirudinea) the buccal cavity in jawed [10–12] blood feeding leeches (Arhynchobdellida); Toxins 2019, 11, 666; doi:10.3390/toxins11110666 S2 of S11 presence of two paired salivary glands in jawless leeches (Glossiphoniidae).
    [Show full text]
  • Addenda to the Insect Fauna of Al-Baha Province, Kingdom of Saudi Arabia with Zoogeographical Notes Magdi S
    JOURNAL OF NATURAL HISTORY, 2016 VOL. 50, NOS. 19–20, 1209–1236 http://dx.doi.org/10.1080/00222933.2015.1103913 Addenda to the insect fauna of Al-Baha Province, Kingdom of Saudi Arabia with zoogeographical notes Magdi S. El-Hawagrya,c, Mostafa R. Sharafb, Hathal M. Al Dhaferb, Hassan H. Fadlb and Abdulrahman S. Aldawoodb aEntomology Department, Faculty of Science, Cairo University, Giza, Egypt; bPlant Protection Department, College of Food and Agriculture Sciences, King Saud University, Riyadh, Kingdom of Saudi Arabia; cSurvey and Classification of Agricultural and Medical Insects in Al-Baha Province, Al-Baha University, Al-Baha, Saudi Arabia ABSTRACT ARTICLE HISTORY The first list of insects (Arthropoda: Hexapoda) of Al-Baha Received 1 April 2015 Province, Kingdom of Saudi Arabia (KSA) was published in 2013 Accepted 30 September 2015 and contained a total of 582 species. In the present study, 142 Online 9 December 2015 species belonging to 51 families and representing seven orders KEYWORDS are added to the fauna of Al-Baha Province, bringing the total Palaearctic; Afrotropical; number of species now recorded from the province to 724. The Eremic; insect species; reported species are assigned to recognized regional zoogeogra- Arabian Peninsula; Tihama; phical regions. Seventeen of the species are recorded for the first Al-Sarah; Al-Sarawat time for KSA, namely: Platypleura arabica Myers [Cicadidae, Mountains Hemiptera]; Cletomorpha sp.; Gonocerus juniperi Herrich-Schäffer [Coreidae, Hemiptera]; Coranus lateritius (Stål); Rhynocoris bipus- tulatus (Fieber) [Reduviidae, Hemiptera]; Cantacader iranicus Lis; Dictyla poecilla Drake & Hill [Tingidae, Hemiptera]; Mantispa scab- ricollis McLachlan [Mantispidae, Neuroptera]; Cerocoma schreberi Fabricius [Meloidae, Coleoptera]; Platypus parallelus (Fabricius) [Curculionidae, Coleoptera]; Zodion cinereum (Fabricius) [Conopidae, Diptera]; Ulidia ?ruficeps Becker [Ulidiidae, Diptera]; Atherigona reversura Villeneuve [Muscidae, Diptera]; Aplomya metallica (Wiedemann); Cylindromyia sp.
    [Show full text]
  • Notable Invertebrates Associated with Fens
    Notable invertebrates associated with fens Molluscs (Mollusca) Vertigo moulinsiana BAP Priority RDB3 Vertigo angustior BAP Priority RDB1 Oxyloma sarsi RDB2 Spiders and allies (Arachnida:Araeae/Pseudoscorpiones) Clubiona rosserae BAP Priority RDB1 Dolomedes plantarius BAP Priority RDB1 Baryphyma gowerense RDBK Carorita paludosa RDB2 Centromerus semiater RDB2 Clubiona juvensis RDB2 Enoplognatha tecta RDB1 Hypsosinga heri RDB1 Neon valentulus RDB2 Pardosa paludicola RDB3 Robertus insignis RDB1 Zora armillata RDB3 Agraecina striata Nb Crustulina sticta Nb Diplocephalus protuberans Nb Donacochara speciosa Na Entelecara omissa Na Erigone welchi Na Gongylidiellum murcidum Nb Hygrolycosa rubrofasciata Na Hypomma fulvum Na Maro sublestus Nb Marpissa radiata Na Maso gallicus Na Myrmarachne formicaria Nb Notioscopus sarcinatus Nb Porrhomma oblitum Nb Saloca diceros Nb Sitticus caricis Nb Synageles venator Na Theridiosoma gemmosum Nb Woodlice (Isopoda) Trichoniscoides albidus Nb Stoneflies (Plecoptera) Nemoura dubitans pNotable Dragonflies and damselflies (Odonata ) Aeshna isosceles RDB 1 Lestes dryas RDB2 Libellula fulva RDB 3 Ceriagrion tenellum N Grasshoppers, crickets, earwigs & cockroaches (Orthoptera/Dermaptera/Dictyoptera) Stethophyma grossum BAP Priority RDB2 Now extinct on Fenland but re-introduction to undrained Fenland habitats is envisaged as part of the Species Recovery Plan. Gryllotalpa gryllotalpa BAP Priority RDB1 (May be extinct on Fenland sites, but was once common enough on Fenland to earn the local vernacular name of ‘Fen-cricket’.)
    [Show full text]
  • Volume 2, Chapter 12-19: Terrestrial Insects: Holometabola-Diptera
    Glime, J. M. 2017. Terrestrial Insects: Holometabola – Diptera Nematocera 2. In: Glime, J. M. Bryophyte Ecology. Volume 2. 12-19-1 Interactions. Ebook sponsored by Michigan Technological University and the International Association of Bryologists. eBook last updated 19 July 2020 and available at <http://digitalcommons.mtu.edu/bryophyte-ecology2/>. CHAPTER 12-19 TERRESTRIAL INSECTS: HOLOMETABOLA – DIPTERA NEMATOCERA 2 TABLE OF CONTENTS Cecidomyiidae – Gall Midges ........................................................................................................................ 12-19-2 Mycetophilidae – Fungus Gnats ..................................................................................................................... 12-19-3 Sciaridae – Dark-winged Fungus Gnats ......................................................................................................... 12-19-4 Ceratopogonidae – Biting Midges .................................................................................................................. 12-19-6 Chironomidae – Midges ................................................................................................................................. 12-19-9 Belgica .................................................................................................................................................. 12-19-14 Culicidae – Mosquitoes ................................................................................................................................ 12-19-15 Simuliidae – Blackflies
    [Show full text]
  • Diptera) Diversity in a Patch of Costa Rican Cloud Forest: Why Inventory Is a Vital Science
    Zootaxa 4402 (1): 053–090 ISSN 1175-5326 (print edition) http://www.mapress.com/j/zt/ Article ZOOTAXA Copyright © 2018 Magnolia Press ISSN 1175-5334 (online edition) https://doi.org/10.11646/zootaxa.4402.1.3 http://zoobank.org/urn:lsid:zoobank.org:pub:C2FAF702-664B-4E21-B4AE-404F85210A12 Remarkable fly (Diptera) diversity in a patch of Costa Rican cloud forest: Why inventory is a vital science ART BORKENT1, BRIAN V. BROWN2, PETER H. ADLER3, DALTON DE SOUZA AMORIM4, KEVIN BARBER5, DANIEL BICKEL6, STEPHANIE BOUCHER7, SCOTT E. BROOKS8, JOHN BURGER9, Z.L. BURINGTON10, RENATO S. CAPELLARI11, DANIEL N.R. COSTA12, JEFFREY M. CUMMING8, GREG CURLER13, CARL W. DICK14, J.H. EPLER15, ERIC FISHER16, STEPHEN D. GAIMARI17, JON GELHAUS18, DAVID A. GRIMALDI19, JOHN HASH20, MARTIN HAUSER17, HEIKKI HIPPA21, SERGIO IBÁÑEZ- BERNAL22, MATHIAS JASCHHOF23, ELENA P. KAMENEVA24, PETER H. KERR17, VALERY KORNEYEV24, CHESLAVO A. KORYTKOWSKI†, GIAR-ANN KUNG2, GUNNAR MIKALSEN KVIFTE25, OWEN LONSDALE26, STEPHEN A. MARSHALL27, WAYNE N. MATHIS28, VERNER MICHELSEN29, STEFAN NAGLIS30, ALLEN L. NORRBOM31, STEVEN PAIERO27, THOMAS PAPE32, ALESSANDRE PEREIRA- COLAVITE33, MARC POLLET34, SABRINA ROCHEFORT7, ALESSANDRA RUNG17, JUSTIN B. RUNYON35, JADE SAVAGE36, VERA C. SILVA37, BRADLEY J. SINCLAIR38, JEFFREY H. SKEVINGTON8, JOHN O. STIREMAN III10, JOHN SWANN39, PEKKA VILKAMAA40, TERRY WHEELER††, TERRY WHITWORTH41, MARIA WONG2, D. MONTY WOOD8, NORMAN WOODLEY42, TIFFANY YAU27, THOMAS J. ZAVORTINK43 & MANUEL A. ZUMBADO44 †—deceased. Formerly with the Universidad de Panama ††—deceased. Formerly at McGill University, Canada 1. Research Associate, Royal British Columbia Museum and the American Museum of Natural History, 691-8th Ave. SE, Salmon Arm, BC, V1E 2C2, Canada. Email: [email protected] 2.
    [Show full text]
  • Diptera Chamaemyiidae) Described by Camillo Rondani: a Revision of Eight Silver Fly Species
    Bulletin of Insectology 69 (2): 199-219, 2016 ISSN 1721-8861 On the identity of Leucopis (Diptera Chamaemyiidae) described by Camillo Rondani: a revision of eight silver fly species Alfio RASPI, Giovanni BENELLI Department of Agriculture, Food and Environment, University of Pisa, Italy Abstract The dipterologist Camillo Rondani described a wide number of new Diptera species, including 12 species of silver flies (Diptera Chamaemyiidae). The first Rondani description of Chamaemyiidae was carried out in 1847 in his research “Osservazioni sopra parecchie specie di esapodi afidicidi e sui loro nemici”. Among them, Ochtiphila obscuripes Rondani 1875 was considered by Czerny (1936) synonymous of Chamaemyia geniculata (Zetterstedt 1838). Later on, Raspi shed light on the identity of Leucopis palumbii Rondani 1872 (Raspi, 1983b; 1988); Leucopis bursaria Rondani 1848 (Raspi, 2003) and Ochtiphila frontella Rondani 1875 (Raspi, 2006). In this research, the identity of the other Leucopis species described by Rondani is estabilished. The valid species are Leucopis (Leucopis) aphidivora Rondani 1847, of which Leucopis fiorii Raspi 1986 is a new synonym; Leucopis (Leucopis) aphidiperda Rondani 1848; Leucopis (Leucopis) misaphida Rondani 1848; Leucopis (Leucopis) minuscula Rondani 1875; Leucopis (Leucopis) talaria Rondani 1875; Leucopomyia palliditarsis (Rondani 1875), of which Leucopis alticeps Czerny 1936 is a new synonym. Leucopis (Leucopis) armillata Rondani 1875 is a new synonym of Leucopis palumbii Rondani 1872, and Leucopis ballestrerii Rondani 1875 is a new synonym of Leucopomyia silesiaca (Egger 1862). Overall, concerning the twelve species of Chamaemyiidae described by Camillo Rondani, nine of them are still considered valid species. This study adds basic knowledge to silver fly systematics and has helpful implications concerning Chamaemyiidae-based biological control programs.
    [Show full text]
  • Flies) Benjamin Kongyeli Badii
    Chapter Phylogeny and Functional Morphology of Diptera (Flies) Benjamin Kongyeli Badii Abstract The order Diptera includes all true flies. Members of this order are the most ecologically diverse and probably have a greater economic impact on humans than any other group of insects. The application of explicit methods of phylogenetic and morphological analysis has revealed weaknesses in the traditional classification of dipteran insects, but little progress has been made to achieve a robust, stable clas- sification that reflects evolutionary relationships and morphological adaptations for a more precise understanding of their developmental biology and behavioral ecol- ogy. The current status of Diptera phylogenetics is reviewed in this chapter. Also, key aspects of the morphology of the different life stages of the flies, particularly characters useful for taxonomic purposes and for an understanding of the group’s biology have been described with an emphasis on newer contributions and progress in understanding this important group of insects. Keywords: Tephritoidea, Diptera flies, Nematocera, Brachycera metamorphosis, larva 1. Introduction Phylogeny refers to the evolutionary history of a taxonomic group of organisms. Phylogeny is essential in understanding the biodiversity, genetics, evolution, and ecology among groups of organisms [1, 2]. Functional morphology involves the study of the relationships between the structure of an organism and the function of the various parts of an organism. The old adage “form follows function” is a guiding principle of functional morphology. It helps in understanding the ways in which body structures can be used to produce a wide variety of different behaviors, including moving, feeding, fighting, and reproducing. It thus, integrates concepts from physiology, evolution, anatomy and development, and synthesizes the diverse ways that biological and physical factors interact in the lives of organisms [3].
    [Show full text]
  • Vertical Distribution of Insect Populations in the Free Air Space of Beech Woodland Ent
    CORRECTION I RETTELSE. To be placed with paper by I Bedes indlagt ved artiklen af B. OVERGAARD NIELSEN Vertical distribution of insect populations in the free air space of beech woodland Ent. Meddr 54,2: 169-178. 1987. Please note I Bemcerk: All figure texts are printed in the correct sequence, but due to an unfortunate oversight, the diagrams (figures) for Figs l and 2 have been interchanged. Samtlige figurtekster er trykt i korrekt rcekkefelge, men pa grund af en beklagelig uagtsomhed er diagrammerne tit Figs 1 og 2 blevet byttet om. With apologies I Med beklagelse The Editor I Redakteren Vertical distribution of insect populations in the free air space of beech woodland B. OVERGAARD NIELSEN Nielsen, B. Overgaard: Vertical distribution of insect populations in the free air space ofbeech woodland. Ent. Meddr 54: 169-178. Copenhagen, Denmark, 1987. ISSN 0013-8851. The vertical distribution of nocturnal aerial insect populations - especially Diptera Nematocera - in a high-holed beech stand was recorded by means of light traps placed at four levels from the forest floor to the top canopy. The insects were mainly trapped near the forest floor and to some extent in the beech canopy; the number recorded in the trunk space and above the top canopy was distinctly lower. In Tipulinae and Mycetophilidae, the vertical distribution declined with increasing height irrespective of season and weather; 80-85% of all specimens were recorded at ground level. In Limoniinae, Cecidomyiidae, Psychodidae and Sciaridae the vertical distribution pattern changed repeatedly during the season, being unimodal with a peak near the forest floor or bimodal with another one in the beech canopy.
    [Show full text]