Contr. Tert. Geol. 7 fig'., 4 tab., 3 pis. Leiden, September 1990 Quatern. 27(2-3) 3981
A Late Oligocene (Chattian B) molluscan fauna from
the coastal cliff at Mogenstrup, North of Skive,
Jutland, Denmark
K.I. Schnetler
LANGA, DENMARK
and
C. Beyer
STAVANGER, NORWAY
Schnetler, K.I., & C. Beyer. A Late Oligocene (Chattian B) molluscan fauna from the coastal cliff at Mogenstrup, North of Skive,
Jutland, Denmark. — Contr. Tert. Quatern. Geol., 27(2-3): 39-82, 7 figs, 3 tabs, 3 pis. Leiden, September 1990.
rich Late molluscan fauna from coastal cliff North of Skive, was studied. A very Oligocene a at Mogenstrup, Jutland, Denmark,
A of the Late in NW is the and the section are des- summary Palaeogene sedimentary sequence Jutland given, locality exposed
cribed. Lithostratigraphical, magnetostratigraphical and biostratigraphical correlations are suggested. A list of the molluscan
species is given, comparisons with other Danish and German Late Oligocene localities are made, and palaeoecological interpreta-
the Andersondrillia is introduced within the Microdrillia. tions are suggested. In the systematic part subgenus subgen. nov. genus
Several taxa are discussed and the followingnew species and subspecies are described: Limopsis (Pectunculina) lamellata chattica subsp.
vonderhochti Collonia troelsi nov., Limopsis (Pectunculina) sp. nov., (Collonia) sp. nov., Lepetella helgae sp. nov., Lepetellajytteae sp. nov.,
Laiocochlis Tubiola subangulata sp. nov., Cerithiopsis (s. lat.) antonjansei sp. nov., (Laiocochlis) supraoligocaenica sp. nov., Triforis (Trituba)
Cirsotrema Searlesia ravni brueckneri danica Clavatula sorgenfreii sp. nov., (Opaliopsis) subglabrum sp. nov., sp. nov., Angistoma subsp. nov.,
Microdrillia Pleurotomella mogenstrupensis sp. nov., Microdrillia (Microdrillia) ingerae sp. nov., (Andersondrillia) brejningensis sp. nov.,
rasmusseni and ? nov. (Pleurotomella) sp. nov., Rimosodaphnella lappanni sp. nov. Actaeopyramis (s. lat.) pseudopunctata sp.
Key-words — Mollusca, Late Oligocene, Chattian B, new taxa, lithostratigraphy, magnetostratigraphy, biostratigraphy,
palaeoecology, Denmark.
K.I. Schnetler, Fuglebakken 14, Stevnstrup, DK-8870 Langa, Denmark; C. Beyer, CB-Magneto, P.O. Box 7015, Jorenholmen,
N-4004 Stavanger, Norway.
Contents
Material 45 Dansk sammendrag p. 40 p.
Introduction 41 List of molluscan 46 p. species p.
The 50 Late Oligocene sedimentary sequence Systematic part p.
other in NW Jutland p. 41 Comparison with Late Oligocene mol-
luscan faunas 71 Lithostratigraphical correlation p. 44 p.
correlation 44 73 Magnetostratigraphical p. Palaeoecological interpretations p.
The Mogenstrup section p. 44 Acknowledgements p. 75
References 75 Biostratigraphical correlation p. 45 p.
1 This study was supported by the Danish National Science Research Council through grant 91-3204 40
1 Location of the section. Part of the 12/6 III SV Kort- Fig. Mogenstrup 1:25,000 map-sheet Hvalpsund (C). og
Matrikelstyrelsen (A84-90). (Author: Schnetler).
Dansk sammendrag
En 0vre fra B Oligocaen molluskfauna Mogenstrup, cerer en Chattian alder. Faunalisten indeholder nord for Danmark. Fra den diskuteres Skive, Jylland, en kyst- 197 arter. I systematiske del en klint nord for Skive 0vre raekke afbildes raekke er en meget rig Oligo- arter, og pa tre plancher en
caen molluskfauna indsamlet. Denne fauna er karakteristiske arter. Inden for slaegten Microdrillia
raek- bemaerkelsesvaerdig ved at indeholde en lang opstilles den nye underslsegt Andersondrillia. ke der fundet i danske af- introduceres: Li- arter, aldrig tidligere er Folgende nye arter og underarter
fra denne del lamellata chattica lejringer periode, ligesom en arter, mopsis (Pectunculina) subsp. no v.,
der er andre her er vonderhochti Collonia yderst sjaeldne pa lokaliteter, Limopsis (Pectunculina) sp. nov.,
I det beskrives loka- troelsi hyppige. foreliggende arbejde (Collonia) sp. nov., Lepetella helgae sp. nov.,
0vre Oli- liteten, og der gives en oversigt over den Lepetella jytteae sp. nov., Tubiola subangulata sp.
i NV gocaene lagserie Jylland. Litho-, magneto- og nov., Cerithiopsis (s. lat.) antonjansei sp. nov., Laio-
korrelationer tolk- cochlis biostratigrafiske foreslas, og en (Laiocochlis) supraoligocaenica sp. nov., Triforis
af Pectiniderne indi- Cirsotrema ning aflejringsmiljoet gives. (Trituba) sorgenfreii sp. nov., (Opaliopsis) 41
Searlesia ravni does show affinities with subglabra sp. nov., sp. nov., Angistoma unpublished assemblages
brueckneri danica subsp. nov., Clavatula mogenstrupen- from the Vejle Fjord area, especially with regard
Microdrillia number of characteristic The sis sp. nov., (Microdrillia) ingerae sp. to a species. assem-
Microdrillia also shows affinities with the rich fauna nov., (Andersondrillia) brejningensis sp. blage very
Pleurotomella from F.R.G. nov., (Pleurotomella) rasmusseni sp. Glimmerode, (R. Janssen, 1978a, b;
nov., Rimosodaphnella lappanni sp. nov. og ? Actaeo- 1979a, b).
Faunasam- pyramis (s. lat.) pseudopunctata sp. nov.
THE LATE OLIGOCENE SEDIMENTARY SEQUENCE IN mensaetningen indicerer aflejring pa lavere vand NW JUTLAND (AUTHOR: BEYER) end de ovrige kendte danske 0vre Oligocaene
for raekke The Late in NW faunaer. Faunaen viser, isaer en Oligocene sedimentary sequence
sjaeldnere arters vedkommende, affinitet til fau- Jutland consists of the fine-grained Branden For-
faunaen fra mation naer fra Vejle Fjord omradet og til underlying a coarser grained sequence,
which several localities in the Glimmerode, Vesttyskland. Ud fra en palaso- outcrops at area.
foreslas sublittoralt These sediments okologisk tolkning et aflej- coarser (Fig. 2a) comprise an
of ringsmiljo. up to c. 120 m thick sequence shallow-marine
sands of deposits, overlain by Miocene quartz Schnetler Introduction (Authors: and Beyer) fluvial The Late be origin. Oligocene part may
The Mogenstrup section is located in NW divided into six facies types (indicated by letters F
the of to from to Jutland, Denmark, on east coast Sailing, ap- A, top bottom):
proximately 1 km South of the small village of
Mogenstrup (UTM coordinates: NH 074 814, F - Seven metres of alternating mud, silt and
A sand in storm-dominated tidal-flat Fig. 1). locality near Hakkildgârd, approx- deposited a
imately 1 km North of the Mogenstrup section, environment.
In E - in was already mentioned by Ravn (1906, 1907). Well-sorted, fine-grained sand, deposited
his extensive work on the molluscan faunas from near-shore, beach and back-barrier en-
the Oligocene and Miocene Formations that vironments. The thickness is up to 20 m.
D - and author considered the sediments to be of Middle Lagoonal protected bay deposits con-
this of Miocene age, but later (Ravn, 1924) age sisting black, homogeneous, organic mud
with horizons of calcite concretions. assignment was questioned. Subsequently, spherical
stratigraphie work was done in other localities in The thickness of this facies varies from a a few
60 the area (e.g. Sorgenfrei, 1940; Rasmussen, 1961; metres to about m.
and - and of Kristoffersen, 1972; Lieberkind, 1977), a C Shallow-bay shelf sediments consisting
Late Miocene for chamositic close to a Oligocene or Early age com- mud, probably deposited
parable sediments suggested. Since the publication river mouth. This facies is found only in the of Ravn's the at western of the where it papers, however, outcrops part area (Fig. 5)
had studied in reaches thickness of 8 Mogenstrup not been until 1987, a m.
which included in facies B - Shelfsediments of silt year they were a analysis consisting a glauconitic
with of siderite concretions in the of the Late Oligocene sedimentary sequence in layers up-
That included The thickness varies from less NW Jutland (Beyer, 1987). paper a permost metre.
the results of which 1 8 palaeomagnetic study, suggest than m to m.
Chattian for this A - Laminated silt, rich in diatoms, in a Late age sequence. deposited
of During field work along the coast North an anoxic environment, possibly an upwelling
Skive thin sand discovered The thickness is less than a glauconitic layer was zone. one metre.
in the coastal cliff at Mogenstrup. This sand was
rich in molluscs and The at least three very fossils, especially spines sequence comprises major of the echinoid ‘Cidaris’ The is hiatuses indicated in 2a. At least sp. present paper as Fig. two trans- the result of of the rnolluscan which occurred a study fauna, gressions during the time of deposition.
found 197 89 of which the first material was to comprise taxa, are During one, mainly fine-grained
for the recorded from the Danish Late Oligocene was deposited (facies A and B). After a regression first time. minor occurred. This a transgression transgres-
The molluscan is different sion the introduction of material into assemblage distinctly saw coarser from all known Danish Late but the to the establishment of barrier- Oligocene faunas, area, leading a 42
1
Fig. (see (author: Mogenstrup legend) for
at 2b sequence Fig. and Sedimentary location for Beyer).
3.
Fig.
to and 2a (author: Legend Figs Schnetler). 4
2b.
Fig.
the the (author: in Jutland. illustrating variable values NW column in highly maximum
are sequence with lithological which shown
Oligocene are
Composite Late Thicknesses area, Beyer).
2a.
Fig. 43
lagoon complex (C, D, E, and F), which was part- sediments are solely found in depressions around
salt and the thicknesses of the various ly destroyed during a subsequent regressive structures
This rests the Late facies short distances. period. sequence on Oligocene vary greatly over
Branden Formation in the eastern part of the area Palaeotopography was probably influenced by
while with main- (Fig. 2), a primary boundary to the synsedimentary halokinesis, deposition
Palaeocene/Eocene Fur Formation is further in the the salt seen ly occurring subsiding areas between
the where burrows extend from the Erosion of these to West, structures. on top structures may
sediment down into the Fur Formation. have removed of the while sub- Oligocene part sequence
of this between The deposition and preservation se- sidence the structures led to a high preser-
influenced halokinesis. The vation in these quence was partly by potential areas.
correlation of the Danish Late with the established of Hardenbol Fig. 4. Suggested Oligocene sequence stratigraphy Haq, &
should the in level Vail (1987). It be noted that short term changes global sea are not directly reflected in the
Danish facies associations: the shallow marine in of sedimentary sequence originated a period relatively high global
sea level, thus indicating that sedimentation was influenced by subsidence of the local basin rather than by changes
in global sea level (author: Beyer). 44
be correlated with chronozones LITHOSTRATIGRAPHICAL CORRELATION R4a may polarity
C7a and which of about 27 Ma (AUTHOR: BEYER) C8, implies an age
for the sediment that yielded the fauna described The sequence described is coeval with and very here. similar to the Late Oligocene Vejle Fjord Forma- tion (Larsen & Dinesen, 1959) but it is clearly not THE MOGENSTRUP SECTION (AUTHOR: BEYER) correlative with this the directly formation, type The sequence outcropping at Mogenstrup (Fig. 3) locality of which is situated 200 km SE of Mogens- be divided into 4 units: standard can trup. Thus, according to stratigraphical
these should be procedure two sequences given Unit 1 de- separate formation names. However, it is not Light-greenish, clayey silt, rich in glauconite, and sirable to add to the already existing confusion over containing fragments of shells. In the uppermost Danish Late Oligocene lithostratigraphical units. metre three horizons of siderite concretions occur. That is the Formation' is why name 'Vejle Fjord These contain flattened horizontal burrows, in- also applied to the sequence described here. dicating that compaction took place prior to Similarly, facies B will be referred to as Brejning cementation. The thickness of the unit is at least 2 Clay Member, because of the close lithological si- m. The base is not visible. The fauna described in milarity to the Brejning Clay Member of the Vejle the found the present paper was in uppermost 20 It reasonable Fjord Formation. seems to assume of this where cm unit, green sand, consisting en- that this facies was originally very widespread. of and tirely glauconite containing an extremely The Vejle Fjord Formation names will not be rich fauna, is associated with the concretions. for the other have used facies, since these clearly a
restricted very geographical occurrence. Unit 2
Facies D is the equivalent to Vejle Fjord Clay consists This unit has a sharp lower boundary and and facies E and F to the are equivalent Vejle Fjord of 30 of dark silt with mica and cm brown, clayey Sand. and contains C gypsum precipitations, organic
(2%), pyrite (c. 2%), siderite and rhodocrosite. MAGNETOSTRATIGRAPHICAL CORRELATION Vertical v-shaped cracks occur. (AUTHOR: BEYER)
Based molluscan on magnetostratigraphical data, Unit 3 faunas and facies 40 interpretations, Beyer (1987) About cm of light brown, silty clay with mica
for most of the and the lower of suggested a Late Chattian age se- jarosite precipitations, boundary
B and correlated which is Small horizontal burrows visi- quence (facies to F) polarity zones sharp. are
R4b, N4, R5 and N5 with polarity chronozone C7 ble. The sediment contains pyrite (8%), organic C
and and (Fig. 4). Polarity zones Nl, R2, N2, R3, N3, (4%), shell fragments.
Table 1. of the Mineralogy Mogenstrup sequence
Legend: Q-quartz, PL-plagioklas, KF-kalifelspar, SID-siderite, PY-pyrite, JA-jarosite, RH-
rhodocrosite, GI-gibbsite, FO-fosforite, VI-vivianite, GLA-glauconite, MS-monosulfides.
For the minerals calibration available and relative JA, RH, GI, FO no samples were only
vertical changes can thus be inferred from these results. The other values indicate percentage
of the mineral content determined by X-ray diffraction.
The lowermost two samples are from concretions.
Metres PL KF SID PY RH GI FO VI G LA MS C Q. clay JA org.
8.5 95 2 2 1 0 0 0 0 16 0 0 0 0
8.4 70 24 1 2 1 1 1 0 32 24 16 0 0
1 6.0 72 22 1 0 0 3 0 56 40 0 0 +
24 0 0 0 4.0 70 22 2 1 2 0 2 0 + 6
3.0 70 19 2 3 0 5 0 1 0 0 0 0 +
2.5 60 30 0 1 0 8 0 1 24 0 0 0 0 3
2.2 53 25 1 1 2 2 0 7 0 0 0 0 0 3
2.1 41 35 1 1 9 4 1 2 16 8 0 0 0
1.8 30 11 0 1 57 1 0 0 0 0 0 0 0
0.5 25 29 1 1 39 1 0 0 0 0 0 + 0 1 45
examined otolith Unit 4 comm., 1987) two samples (one
Six of silt with variable sand and from unit 1 and from the fossiliferous m a clay content. one very up-
the and of unit both could be Generally, grain size increases towards top permost part 1); assigned an
in the lenses of Late uppermost part longitudinal cross- early Oligocene age.
bedded The sediment is The sand occur. extensively biostratigraphy of the Chattian based on
be the bioturbated and rich in mica and contains two pectinids (R. Janssen, 1979b) can applied to
fauna since number of horizons of scattered, spherical calcite concretions. Mogenstrup a stratigraphi-
of vivianite and side- useful viz. Palliolum Also minor amounts pyrite, cally species were found,
rite The colour is brown but oxidation hausmanni P. limatum P. limatum occur. upon hausmanni, limatum,
it changes to black within a few minutes, which is ambignum, Pecten (Hilberia) bifidus bifidus and Pecten
the of monosulfides. Of these Palliolum probably due to presence (Hilberia) soellingensis. only
Based the limatum and P. on sedimentological analyses Mogen- ambignum hausmanni hausmanni are
section well P. limatum limatum and Pecten strup can be interpreted as follows (Fig. 4; preserved.
few Tab. 1): Unit 1, in the uppermost part of which (Hilberia) bifidus bifidus are represented by a
is the fauna described here was found, thought to fragments, while a single near-complete specimen
have been in marine shelf environ- of collected. The deposited a Pecten (Hilberia) soellingensis was
The environmentof unit 2 fauna whole indicates Chattian A/B ment. depositional was pectinid as a a
content of rhodocrosite Chattian since the Chattian A oxic, indicated by the high age, most likely B,
of cracks which show of and this mollus- and the occurrence v-shaped may species signs reworking,
indicate An is a supra-tidal marsh. oxygen-deficient can assemblage probably not a biocoenosis.
depositional environment is inferred for unit 3, The gastropod species Hinia schlotheimi in-
because of the mineral assemblages (e.g. rich in dicates, according to R. Janssen (1979b: 162), a
facies association and burrows with Chattian B The rather pyrite), the a or younger age. frequent
small diameter. This have been of this is in very may a occurrence species at Mogenstrup
shallow basin that into existence the with the determination came at start agreement age suggested
of that inundated the the a transgression low-lying by pectinids.
the salt Unit 4 Two from unit 1 and areas amongst structures. was samples, one proper one
from the mollusc-rich of unit 1 deposited in a brackish environment, probably a upper part were
This is based the facies for foraminifers Kaare Nor- lagoon. interpretation on analysed by Ulleberg,
the content of who concluded that both fall within association, monosulfides, organic way, samples
of the C, intensive bioturbation and the occurrence the Angulogerina gracilis Zone, as does Brejning
channel in the of the does 1987: small deposits uppermost part Clay (Ulleberg, 201) (K. Ulleberg,
unit. pers. comm.).
The of the geological history Mogenstrup sec- MATERIAL (AUTHOR: SCHNETLER) tion may thus be summarized as follows. Upon
in marine shelf environment The molluscs collected almost deposition a (unit 1) were exclusively
followed of and erosion from the of unit a period non-deposition uppermost part 1. Large samples
washed (unit 2). The succeeding transgression flooded the of glauconitic sand were taken, dried and
rise marine 0.5 mesh. The residues hand- area and gave to a shallow, protected on a mm were
basin As material intro- after and (unit 3). more sandy was picked drying fractioning.
duced into the barrier islands and in the area, developed During collecting field, larger specimens
Charonia Orthosurcula the subsiding basin became a lagoon. Gradually of e.g. flandrica, regularis,
the exceeded the of sub- and sedimentation rate rate Scaphella siemsseni Boreotrophon capito were
which led with but the bulk of the material sidence, to infill of the basin coarser regularly encountered,
material (unit 4). was collected by sediment processing. The
well but molluscs are generally preserved pyrite BIOSTRATIGRAPHICAL CORRELATION destruction of the disintegration causes rapid (AUTHOR: SCHNETLER) specimens, especially in the field.
is the non-molluscs The Late Oligocene age of the section imme- Very conspicuous among are
the molluscan fauna and number of of the echinoid diately apparent from the very large spines this determination is confirmed of ‘Cidaris’ of crinoid age by a study sp. Marginal plates asteroids, the otoliths. Dr P.A.M. Gaemers (Leiden, pers. brachials, Bryozoa and serpulids are also com- 46
Anthozoa LIST mon. Many are found, especially Cerato- OF MOLLUSCAN SPECIES (AUTHOR: SCHNETLER)
' cyathus granulatus (Goldfuss, 1826), Graphularia’ In Table 2 all molluscan species from the of the sp., and a few Flabellum sp. Many specimens listed. With few Mogenstrup outcrop are a excep- ‘Terebratula’ often in nest-like brachiopod sp. occur tions, nomenclature follows R. Tanssen (1978b; ' few of Terebratulina’ concentrations; a specimens 1979a, b). also collected. Teleost were otoliths are common; In column 1, species recorded from the Danish shark teeth, especially of the genus Pristiophorus, Late Oligocene for the first time are indicated with however, are less common. Foraminifers and an asterisk (*). but and ostracods are abundant, decapods In column 2, the frequency of the specimens is balanids seem to be absent. indicated on a scale from 1 to 7:
A small molluscan fauna was collected from unit
3. The molluscs are evenly distributed over the 1: Vi-l specimen sediment, but not common. The following very 2: 2-3 specimens species could be identified: 3: 3-10 specimens
4: 11-40 specimens
Nucula (Lamellinucula) comta 5: 41-125 specimens Yoldia (Yoldia) glaberrima 6: 126-350 specimens
Lentipecten (Lentipecten) corneus 7: more than 350 specimens Dentalium (Dentalium) geminatum
Drepanocheilus (Arrhoges) speciosus The number of scaphopods is estimated. Single
Polinices valve of bivalve = (Euspira) helicinus a 1/2 specimen.
of Ancilla (Ancillus) karsteni In column 3, the depository some rare species
is indicated. Fusiturris selysii. Abbreviations: see systematic part.
Table 2. Molluscan species from Mogenstrup (Brejning Clay Member, Vejle Fjord Formation).
1 22 3
Bivalvia
LeionuculaLeionucula peregrina (Deshayes, 1852) 2
4 Nucula (Lamellinucula)(Lamellinucula) comta Goldfuss, 18371837 4
Nucula 3 Nucula (Lamellinucula)(Lamellinucula) sp.sp.
Nuculana (Saccella)(Saccella) westendorpi (Nyst, 1839) 3
PortlandiaPortlandia (Yoldiella) pygmaea (von Miinster,Munster, 1837) 41
Yoldia (Yoldia) glaberrima (von(von Miinster,Munster, 1837) 33
Acar aff. dentiens (Cossmann(Cossmann & Peyrot, 1912) 5 PI.Pl. 1
BathyarcaRathyarca bellula (Wiechmann, 1874)1874) 4
LimopsisLimopsis (Limopsis) aurita (Brocchi, 1814) 6
Limopsis (Pectunculina)(Peclunculina) lamellata chattica subsp. nov. 7 syst. partpart
LimopsisLimopsis (Pectunculina) retifera Semper, 1861 6
• vonderhochti 5 LimopsisLimopsis (Pectunculina) vonderhochti sp.sp. nov. syst. part
Glycymeris (Glycymeris) obovataobovata (Lamarck, 1819) 2
Modiolula 3 pygmaea (Philippi, 1843)
Musculus 3 sp.sp.
3 Arcoperna sp.
LentipectenLentipecten (Lentipecten) corneuscorneus (Sowerby, 1818) 4
* 44 PropeamussiumPropeamussium (Parvamussium) sp. syst. partpart
1 MNO PalliolumPalliolum (s. lat.) venosum (Speyer,(Speyer, 1864)
Palliolum (s. lat.) limatum limatum (Goldfuss, 1833)1833) 3
Palliolum (s. lat.) limatum ambignum (Anderson, 1958) 2
Palliolum (s. lat.) hausmanni hausmanni (Goldfuss,(Goldfuss, 1833) 3
PectenPecten (Hilberia) bifidus bifidus von Miinster,Münster, 1835 3
1 ENÁ PectenPecten (Hilberia) soellingensis von Koenen, 1868 1 ENA
Limea (Notolimea) nysti (Speyer, 1864) 2
Anomia (Anomia) ephippium (Linne,(Linné, 1758) 3
Pododesmus (Heteranomia) squamula Linne,Linné, 1758 2 47
1 22 3
33 Crassostrea cyathulacyalhula (Lamarck, 1806)
55 Cyclocardia (Cyclocardia) aff. kickxikickxi (Nyst & Westendorp, 1839)
1 MNOM NO Cyclocardia (Cyclocardia) depressa (von Koenen, 1884)
44 Astarte (Astarte)(Astarte) goldfussi praecursor Glibert,Glibcrt, 1957
1 MNO Astarte (Astarte)(Astarte) gracilis gracilis (von Miinster,Munster, 1837)1837) 1
41 Goodallia (Goodallia)(Goodallia) pygmaea (von Miinster,Munster, 1837) Parvicardium kochi (Semper, 1861)1861) 44
Gilbert && Poel, 1970 33 Laevicardium (Habecardium) excomatulum Glibert van dede Poel,
2 Angulus posterus (Beyrich, 1868) Abra (Abra) bosqueti (Semper, 1861) 44
Glossus subtransversus (d'Orbigny, 1852) 2
?.' 1 MNO Coralliophaga sp.
SpaniodontellaSpaniodonlella nilidanitida (Reuss, 1867) 2
1 Callista (Costacallista)(Coslacallista) beyrichi (Semper, 1861) 1 MNO
1 MNO Sphenia abscisa (Wiechmann, 1879) 1
Corbula (Varicorbula) gibba (Olivi, 1792) 22
1 MNO von Koenen, 1894 1 Spheniopsis depressa von
4 Hiatella (Hiatella) arctica (Linne,(Linné, 1767) 4
Teredinidae indet. 4 gen. et sp.
Jouannetia (Jouannetia) aff. neuvillei Cossmann, 1921 1 MNO
Koenen, 1868 1 AJB Lyonsia (Lyonsia) obovata vonvon Koenen, 1868 AJB
ThraciaThracia 1 ISLISL (Thracia) sp.sp.
3 Poromya (Poromya)(Poromya) hanleyana von Koenen, 1863
Cuspidaria (Cuspidaria) subcuspidata (d'Orbigny, 1852)1852) 3 3 Verticordia (Verticordia)(Verticordia) aff.aff. punctata Heering, 1950
PecchioliaPecchiolia 1 sp. AJB
3 Bivalvia indet.indct.
Scaphopoda
Dentalium (Dentalium) geminatum (Goldfuss,(Goldluss, 1841) 5
Dentalium (Dentalium) polypleurum (Seifert, 1959) 44
Rhabdus 2 sp.
Cadulus aff. subfusiformis (Sars, 1865) 1 MNO
Gastropoda
Scissurella (Anatoma) koeneniakoenenia R. Janssen, 1978 3
3 Lepetella compressiuscula (Karsten,(Karsten, 1849)1849)
2 LepetellaLepelella helgae sp. nov.nov. syst. part
* 4 nov. 4 Lepetella jytteae sp. nov. syst. part
Cocculina (Cocculina) dittmeri (Anderson, 1964) 3 PI. 11 ENÁ AstraeaAslraea (Bolma)(Bolma) infausta (Gicbel,(Giebel, 1852) 1 ENA
• 1 AstraeaAslraea (Lithopoma)pustulosa (von Miinster,Münster, 1844) 7 PI. 1
* Collonia troelsi 6 (Collonia) sp. nov. syst. part
Homalopoma (Boutillieria) simplex (Philippi, 1843) 3 Pl.l
1 Homalopoma (? Leptothyropsis) sp. syst. part
Tubiola 3 syst. part subangulata sp. nov. syst. part
* 2 Tubiola part sp. syst. part
? Skenea 3 sp. syst. part
2 Leucorhynchia rotellaeformisrotellaejormis (Grateloup, 1828) 2
Cirsope (Cirsope) multicingulata (Sandberger, 1859) 3
Cirsope (Pseudocirsope) subeffusasubejjusa (Sandberger, 1859) 2
1 MNO Cingula sp.
Rissoa (Persephona) karstenikarsteni R. Janssen, 1978 5
Rissoa (Persephona) punctatissima R. Janssen, 1978 4 Alvania (Arsenia) semperi Wiechmann, 1871 4
Alvania 2 sp. 48
1 22 3
Ovirissoa 1 sp. syst. part
Turritella (Haustator)(Haustalor) goettentrupensisgoeltentrupensis (Cossmann, 1899) 1 MNO
BittiumBiltium 1 MNO sp.
Cerithiopsis (Metaxia) aff. degrangeidegrangei (Cossmann & Peyrot, 1921)1921) 2
Cerithiopsis henckeliushhenckeliusii (Nyst,(Nyst, 1836) 3
Cerithiopsis jutensis Schnetler, 1985 4\
Cerithiopsis (s.(s. lat.) serrula R. Janssen, 1978 22
* Cerithiopsis (s. lat.) daphnelloides R. Janssen, 1978 3
Cerithiopsis (s. lat.) ariejansseni R. Janssen, 1978 22
* Cerithiopsis (s.(s. lat.)lat.) antonjansei sp. nov. 2 syst. part
* * Laiocochlis nov. 2 (Laiocochlis)(Laiocochlis) supraoligocaenicus sp. nov. syst. part
Cerithiopsida boelschei (von Koenen, 1891) 41
Cerithiella bitorquata (Philippi, 1843) 4
Seila (s. lat.) angustaangusla Tembrock, 1964 3
Seila (s. lat.) koeneni R. Janssen, 1978 1 MNO
* 3 Triforis (Trituba)(Trituha) sorgenfreii sp. nov. syst. part
* Biforina (Biforina) praeversa Griindel,Grundel, 1975 4
Norephora (Norephora) elatior (von Koenen, 1891) 4\
Opalia (Pliciscala)(Pliciscala) pusillapusilla (Philippi, 1843) 1
Turriscala (Turriscala) cf. rudis (Philippi, 1843) 2
Acirsa (Plesioacirsa) leunisii (Philippi, 1843) 2
Cirsotrema 2 (Opaliopsis) subglabrum sp. nov. syst. part
Cirsotrema (Opaliopsis) sp.lsp.l 1 syst. part
Cirsotrema 1 (Opaliopsis) sp.2sp.2 syst. part
Cirsotrema aff. koeneni A.W. 19671967 1 (? Opaliopsis) Janssen,Janssen, syst. part
Cirsotrema (s. lat.) crispatumcrispatum Harder, 191.31913 2 JHR
Amaea 1 ISL Amaea (Secalina) amoena (Philippi,(Philippi, 1843)
ArchileclonicaArchitectonica 2 sp.
Mathilda (Mathilda) sandbergensandbergeri (Koch, 1876) 2
AclisAclis (Aclis) vetusta Wiechmann, 1878 41
AclisAclis (Stilbe) proneglecta R. Janssen, 1978 2
AclisAclis (Graphis) hosiusi (Lienenklaus, 1891) 4
* Balcis (Balcis) ? lamberti (Cossmann, 1882) 11 MNO
Balcis alba naumanni (von Koenen, 1867) 41
Balcis (Polygyreulima) pseudonaumannipseudonaumanni R. Janssen, 1978 3
Eulima (Eulima)(Eulima) emersa Speyer, 1870 4
Niso (Niso)(Niso) minor Philippi, 1843184.3 3
Neojanacus planatus (Speyer, 1864) 3 PI. 22
Drepanocheilus (Arrhoges) speciosus (von(von Schlotheim, 1820) 7
Polinices (Euspira)(Euspira) helicinus (Brocchi, 1814) 7
** ? R. 1978 2 ApiocypraeaApiocypraea ? humbergihumbergi Janssen,Janssen, syst. part
Semicassis (Echinophoria)(Echinophoria) rondeletirondeleti (Basterot,(Basterot, 1825) 4
Charonia (Sassia) flandricaflandrica (de(de Koninck, 1837) 5
Ficus (Ficus) concinnus (Brongniart, 1822)1822) 3
Pterynotus (Pterochelus) tristichus (Beyrich, 1854) 2
BoreotrophonBoreolrophon (s. lat.) capitacapito (Philippi, 1843) 4
Lyrotyphis (Eotyphis) sejunctus (Semper,(Semper, 1861) 5
* Trophonopsis1'rophonopsis angustevaricataangustevancata (Gripp, 1915)1915) 5 syst. partpart
Coralliophila (Hirtomurex) kochi (Beyrich, 1854) 3 PI. 22
4 Scalaspira (Scalaspira) elegantulaeleganlula aequistnataaequistriata (Speyer, 1863)
Scalaspira (Scalaspira) ? waeli (Beyrich, 1856) 1 MNO
5 PI. 33 Scalaspira (Scalaspira) sp.
Searlesia ravni nov. 3 sp. syst. partpart
Exilioidea elatior (Beyrich, 1848) 2
Parvisipho (s. lat.)lat.) scrobiculatus (Boll, 1851) 1 ISL
Coptochetus (s. lat.) danicus Schnetler, 1985 2
Metula (Daphnobela) scabricula (Philippi, 1843) 41
Pisanella subgranulatasubgranulata (von Schlotheim, 1820) 3
* Buccinidae et sp. indet. 2 gen. et sp. syst. part 49
1 22 3
Hinia (Tritonella) schlotheimi (Beyrich, 1854) 4
StreptochetusSlreptochetus (Streptodictyon)(Streptodktyon) cheruscus cheruscus (Philippi, 1843) 6
Streptochetus (Streptolathyrus) soellingensis Tembrock, 1965 3
Latirus 1 (Pseudolatirus) sp. syst. part
Scaphella (Scaphella) siemssenisiemsseni (Boll, 1851) 4•1
Ancilla (Ancillus) karsteni (Beyrich, 1856) 55
Gibberula brevis (von Koenen, 1890) 4 PI. 3
Cancellaria (Merica) evulsa postera (Beyrich, 1856)1856) 3
BabylonellaBabylonella pusilla (Philippi, 1843) 5
Vexillum (Uromitra)(Uromitra) hastatumhaslatum (Karsten, 1849)1849) 41
ConomitraConomilra soellingensis (Speyer, 1864) 5
OrthosurculaOrlhosurcula regularisreguláris (de Koninck, 1837) 5
AcamptogenoliaAcamptogenotia morreni (de(de Koninck, 1837)1837) 2
Stenodrillia obeliscus (des Moulins, 1842)1842) 5
Cochlespira volgeri (Philippi, 1843) 41
Gemmula (Gemmula) laticlavia (Beyrich,(Beyrich, 1848) 4
Gemmula (Gemmula) trispiralis R. Janssen, 1979 5
Gemmula (Gemmula) pseudokoninckipseudokomncki R. Janssen, 1979 4
PleuroliriaPleuroliria konincki (Nyst, 1845) 3
Fusiturris duchastelii (Nyst, 1836) 6
Fusiturris selysii (de Koninck, 1837)1837) 4
Fusiturris enodis R. Janssen, 1979 2
GlibertturriculaGhberlturncula ariejansseni (Schnetler, 1987) 4
Clavatula 2 mogenstrupensis sp. nov. syst. part
* Cordieria plicata (Beyrich, 1848)1848) 1 ISL
RathytomaBathytoma (Bathytoma) leunisii (Philippi, 1843)1843) 5
* Asthenotoma holzapfeli (von Koenen, 1890) 3
Splendrillia koeneni (Speyer, 1867)1867) 4
* Microdrillia 3 (Microdrillia) ingerae sp. nov. syst.syst. partpart
* Microdrillia 4 (Andersondrillia) brejningensisbrejmngensis sp. nov. syst.syst. part
'Microdrillia'‘Microdrillia’ speyeri (Koch & Wiechmann, 1872) 3 PI. 3
Boreodrillia undatellaundalella (Speyer, 1867) 4
PleurotomellaPleuroiomella anderseni Schnetler, 1987 11 MNO
* Pleurotomella (Pleurotomella) margantatamargaritata R. Janssen, 1978 3 PI. 3
* PleurotomellaPleuroiomella rasmusseni 2 (Pleurotomella) sp. nov. syst. part
* 4 Rimosodaphnella lappanni sp. nov. syst. part
Conus (Leptoconus) semperi Speyer, 1862 4
Hastula (Haslula)(Hastula) beyrichi (Semper, 1861) 1 MNO
1 MNO Chrysallida sp.
Ebala 1 MNO sp. Evalea kochi (Gorges, 1952)1952) 3
• Evalea incrassata (von Koenen, 1867) 1 ISLISL
OdostomiaOdoslomia MNO sp. 11
Syrnola (Syrnola) subcylindracea (Philippi, 1843)1843) 5
* Syrnola (Puposyrnola) laevissima (Bosquet, 1859) 2
• Turbonilla (Strioturbonilla)(Slrioturbonilla) wiechmanni Speyer, 1870 1 AJB
Turbonilla (Pyrgolampros) jeffreysijejfreysi Koch & Wiechmann, 1872 3
* p? nov. 3 ActaeopyramisAclaeopyramis (s. lat.) pseudopunctata sp. syst. part
Actaeon (Actaeon) punctatosulcatuspunctatosulcatus (Philippi, 1843) 3
Crenilabium terebelloides (Philippi, 1843)1843) 2
Ringicula (Ringiculina) striata (Philippi, 1843) 4
Roxania utriculus (Brocchi, 1814) 4
Philine (Ossiania) kochi vonvon Koenen, 1882 1 MNO
Limacina hospes (Rolle,(Rollc, 1862)1862) 4
Limacina valvatina (Reuss, 1867)1867) 3
* Clio nielseni A.W. Janssen, 1990 4
inc. sed. 1 11 Gastropoda syst. part 50
Stratum typicum — Glauconitic sand, Brejning Clay
Member of the Vejle Fjord Formation, Late
Oligocène (Chattian B).
nominis — This Derivatio new subspecies of Limopsis
Chatt- lamellata is named after its occurrence in the
ian stage of the Oligocene.
Holotype — Pl. 1, Figs 2a-b, (coll. MGUH 20 023)
(leg. K.I. Schnetler).
Description — The shell is small, equivalve and
with somewhat rather convex a obliquely ovate to
subcircular outline. It is slightly inequilateral, and
the orthogyrate, dorsally projecting umbo has a
slightly prosocline position. The hinge line is
straight, the anterior and posterior dorsal margins
The of the are of almost the same length. transition Fig. 5. Other Danish localities mentioned in this paper anterior anterior dorsal margin into the convex (author: Schnetler). margin is gradual; the posterior dorsalmargin and
form obtuse the posterior margin together an SYSTEMATIC PART (AUTHOR: SCHNETLER) is less angle. The posterior margin more or convex. Danish localities mentioned in this section are The transitions of both the anterior and the poste- shown in Fig. 5. into the ventral also rior margin convex margin are indicated The depositories of the materialare by gradual. abbreviations: with The cardinal area is flat and triangular a
AHF - Coll. S.B. Andersen and H.C. rather small triangular resilifer below the umbo.
Flansen, Fredericia, DK. The dentition is divided into two series of teeth by
umbo. The anterior - NL. edentulous below AJB Coll. A.C. Janse, Brielle, an gap the
- four DGU Coll. Geological Survey of Den- series consists of three to teeth which are
mark, Copenhagen, DK. aligned almost dorso-ventrally. The anterior series consists ENA - Coll. E. Nielsen, Arhus, DK. is straight. The arched posterior series also
- teeth FFIK Coll. F. von der Hocht, Kerpen- of three to four which are placed obliquely.
ribs Balkhausen, F.R.G. The sculpture consists of flat concentric The - Coll. K.I. DK. ISL Schnetler, Langa, which are distributed somewhat irregularly.
- DK. much wider than Coll. Randers, between these ribs are the JHR J. Hillersborg, spaces
- DK. the LJL Coll. L.B. Jorgensen, Lime, ribs and finer ribs are visible in interspaces.
- Univer- weaker and the MGUFI Coll. Geological Museum, The radial ribs are sculptural com-
form less sity of Copenhagen, DK. ponents together a more or reticulate or-
MNO - Coll. M.S. Nielsen, Odense, DK. namentation.
crenulation which RGM - Coll. National Museum of Natural The shell margin has a fine
The crenulae History (formerly: Rijksmuseum corresponds to the radial ribs. are
in The line and the van Geologie en Mineralogie), fairly regular strength. palliai
distinct. The anterior Leiden, NL. adductor scars are not very
RLO - Coll. R. Lechner, Odense, DK. impression is small and rounded, the posterior one
SMF - Coll. Forschungsinstitut Sencken- is larger and rounded trapezoidal.
Frankfurt F.R.G. Measurements — Most have of berg, am Main, specimens a height
- maximum is 3 WLH Coll. W. Lappann, Heiligenhaus- about 2 mm. The height c. mm.
Isenbiigel, F.R.G. Paratypes — Mogenstrup (coastal cliff): 1/2
specimen (Pl. 1, Figs 3a-b) (leg. K.I. Schnetler; Limopsis (Pectunculina) lamellata chattica coll. MGUH 20 024); 3/1 and 300/2 specimens subsp. nov. (leg./coll. ISL); 1/1 and 48/2 specimens (leg./coll. PI. 1, Figs 2a-b, 3a-b JHR); 7/2 specimens (leg./coll. LJL); 11/2
2/2 Locus typicus — Coastal cliff at Mogenstrup, North specimens (leg./coll. ENA); specimens (leg. coll. 1/1 and 32/2 of Skive, Jutland, Denmark (Fig. 1). K.I. Schnetler, MNO); speci- 51
292/2 Derivatio nominis — The is named after Mr mens (leg./coll. ACJ); specimens (leg./coll. species
der MNO); 5/2 specimens (leg. K.I. Schnetler; coll. Fritz von Hocht, geologist, Kerpen-Balkhau-
K.I. F.R.G. RGM 229 789); 5/2 specimens (leg. sen,
— MGUH 20 Schnetler; coll. SMF 308 405). Holotype Pl. 1, Figs 4a-b, (coll. 025)
K.I. Hadsten (road excavation): 1/2 specimen (leg. Schnetler).
— with The shell is equivalve a (leg./coll. ISL). Description
Brejning (beach exposure): 40/2 specimens rounded-ovate, slightly oblique outline, inequila-
umbo has (leg./coll. MNO); 3/2 specimens (leg./coll. ISL). teral. The slightly opisthogyrate a pro-
socline it Bogeskov (coastal cliff): 2/2 specimens position; projects only slightly beyond
(leg./coll. MNO). the dorsal margin. The shell is strongly convex,
maximum at about 2/3 of the — and L. lamellata the Discussion The new subspecies with convexity
close with shell The is less obvious in juve- Lehmann, 1885 s. str. are very regard height. convexity
and How- nile to size, outline, convexity sculpture. specimens. The of the from the numerous available The line is straight. length ever, judging hinge
of anterior dorsal is about two thirds of that specimens of the new subspecies and specimens margin
dorsal and these L. lamellata s.str. from the Miocene of Dingden of the posterior margin, margins
almost in line. Both the anterior and the (type locality!), Twistringen (F.R.G.), and Win- are poste-
almost in their dorsal terswijk-Miste (The Netherlands) (coll. ISL) a rior margins are straight
in the The transitions into the ventral are number of features seem to be different parts. margin
Danish material. gradual, posteriorly with a somewhat stronger cur-
than In L. lamellata s.str. the cardinal area is higher vature anteriorly.
and with and umbonal The cardinal area is flat and subtriangular, a more projecting stronger
The number of teeth with fine striation the region and a larger resilifer. provided a parallel to hinge
The rather resilifer is below the in L. lamellata s.str. in the anterior and posterior line. large placed
umbo. Its outline is that of series is 4-6 and these teeth are somewhat a right-angled triangle
the the with the shorter cathetus placed The stronger. Furthermore, sculpture on anteriorly. resilifer is with striation similar that Miocene form is generally stronger, especially the provided a to
of the cardinal The dentition into concentric ribs, and finally its maximum size is area. is divided about 5 these dif- series of teeth an edentulous below the mm. Although not very obvious, two by gap
Because of the dif- umbo. The anterior series consists of 4-6 teeth ferences seem to be constant.
which almost The ference in I the Danish material are age prefer to treat aligned dorso-ventrally.
of L. lamellata. The arched series consists of 4-8 as a stratigraphical subspecies posterior, slightly
distributed in the the 2-3 of nominal subspecies is widely more obliquely placed teeth, posterior
line. The Middle Miocene of the North Sea Basin (Hem- which are almost parallel to the hinge
number of teeth in all the moorian and Reinbekian). is variable, but cases
of teeth is than the anterior posterior row longer
vonderhochti Adult with 4-5 anterior and 5-6 Limopsis (Pectunculina) sp. nov. one. specimens
PI. 1, Figs 4-7 posterior teeth are most frequent.
The sculpture is dominated by numerous con-
— in & lines. The ? 1884 Limopsis Goldfussi Nyst von Koenen, Speyer centric riblets which follow the growth
von Koenen, 31, 9-10 pi. figs (non Nyst). number of these riblets is about one hundred on
— 141 ? 1937 Limopsis Goldfussi Nyst Eriksen, p. (non Nyst with of 10 concentric specimens a height mm. The ?)• sculpture elements frequently alternate in 1941 — 160 Limopsis Goldfussi Nyst Gorges, p. (partim, The radial is much weaker and non Nyst). strength. sculpture
1952 — 17 the Limopsis Goldfussi Nyst Gorges, p. (non Nyst). almost exclusively visible in between concen-
1861 — R. 1979b Limopsis (Pectunculina) retifera Semper, tric the serrated ribs, giving interspaces a ap- 16 Janssen, p. 30, pi. 1, fig. (partim, nonSemper). pearance.
Locus typicus — Coastal cliff at Mogenstrup, North The inner shell margin has a coarse crenulation
of the This of Skive, Jutland, Denmark (Fig. 1). on the anterior part ventral margin.
— consists of knobs which Stratum typicum Glauconitic sand, Brejning Clay crenulation 4-6 coarse are
in dorsal direction ribs Member of the Vejle Fjord Formation (Late elongated as radiating on
material Oligocène, Chattian B). the inner shell surface. In the available 52
these ribs are somewhat variable in strength. The Kirstinebjerg Skov (beach exposure): 1/2 speci-
of the ventral is remaining part margin usually men (leg./coll. ISL).
but in few shells in- A and smooth, a additional knobs are Krefeld-Linn (water well) (Chattian B):
dicated, especially near the transition into the pos- 1/2 specimen, PI. 1, Fig. 6a-b, (leg. WHL, coll.
teriormargin. The crenulations do not correspond SMF 309 204), 7/2 specimens (leg.coll. FHK).
radial ribs the outside of the shell. Krefeld-Linn with the on ('Maizena' water well) (Chattian
line adductor The pallial and the scars are not A): 1/2 specimen, (leg./coll. FFIK).
distinct. A is situ- Krefeld 1/2 very well-developed myophore (borehole GLA-1): specimen (Chat-
ated below the anterior of the The tian 1/2 part hinge. ante- A), (leg./coll. FFIK); specimen (Chattian
much smaller rior adductor scar is subovate and B), (leg./coll. FHK).
than the 1/2 rounded-trapezoidal posterior one. Matzerath(Schacht S.J. 8): specimen (310.5-
Range of variation — The Danish material demon- 312.5 m) (leg./coll. FHK); 7/2 specimens (330-332
strates a rather wide range of variationwith regard m); 1/1 + 1/2 specimen (328-330 m) (both leg./
ratio varies to the convexity. The width/height coll. FHK).
from 0.33 to 0.46, this figure being higher in larger Dalheim (borehole 50/19): 1/2 specimen (Chat-
specimens. tian A) (leg./coll. FHK); 1/2 specimen (Chattian
The Danish material was compared with speci- B) (leg./coll. FHK).
from the German localities Ahnetal Meerbusch-Ilverich 1/1 and 1/2 mens (coll. (water well):
SMF), Krefeld-Linn (coll. WLH) and Meerbusch- specimens (Chattian A) (leg./coll. FHK); 4/2
Osterath (coll. FHK). Those from Ahnetal and specimens (Chattian B) (leg./coll. FHK).
lesser of 1/1 and Krefeld-Linn demonstrate a degree con- Meerbusch-Osterath (water well): 38/2
vexity (length/height ratio 0.27-0.35). Also the specimens (Chattian A + B) (leg./coll. FHK).
crenulation of the anterior ventral margin is Willich (water well): 2/2 specimens (Chattian A)
knobs coarser, with the situated more posteriorly. (leg./coll. FHK).
the is accentuated. In addition, sculpture more Neukirchen-Vluyn (Schacht 5): 1/2 specimen
From of and Meerbusch-Osterath a sample 38/2 (Chattian A) (leg./coll. FHK).
1/1 available. This Wittlaer-Bockum 1/2 specimens was population (water well): specimen
with the Danish material. The agrees more closely (Chattian A) (leg./coll. FHK).
length/height ratio varies from 0.28 to 0.39, and Moers-Kapellen (Kieswerk): 2/2 specimens
the crenulationof the anterior ventral margin and (Chattian B) (leg./coll. FHK).
the rather similar the Moers-Rumeln and external sculpture are to (Schacht Rumeln): 1/1 3/2
Danish specimens. specimens (Chattian A) (leg./coll. FHK); 3/2
5/2 The differences in the length/height ratios may specimens (coll. SMF); specimens (leg./coll.
be the result of local environmental influences and AJB).
therefore I confidently refer the German popula- Ahnetal: 3/1 and 9/2 specimens (coll. SMF 308
tions the coll. SMF 308 to new species. 407) (leg. J. Gorges; 408).
Paratypes — Mogenstrup, coastal cliff: 1/2 Niederkaufungen: 1/2 specimen (coll. SMF 308
specimen (Pl. 1, Fig. 5a-b) (leg. K.I. Schnetler; 409).
coll MGUH 20 026); 69/2 specimens and 21 frag- Discussion — This large Limopsis species has
often been referred L. ments (leg./coll. ISL); 20/2 specimens (leg./coll. previously to goldfussi (Nyst,
restricted the A JHR); 5/2 specimens (leg./coll. LJL); 1/2 1845), a species to Rupelian.
L. found specimen (leg. K.I. Schnetler; coll. MNO); 2/2 specimen of goldfussi was by Harder
specimens (leg. K.I. Schnetler; coll.RGM 229 (1913: 52, pi. 3, fig. 18a-c) in the Middle
fine Arhus. This 790); 2/2 specimens (leg. K.I. Schnetler; coll. SMF Oligocene sand at specimen can-
308 406); 1/2 specimen, leg. K.I. Schnetler, coll. not be referred to L. retifera Semper, 1861, as was
WLH; 3/2 specimens and 2 fragments (leg./coll. assumed by R. Janssen (1979: 30).
2/2 The L. AJB), 86/2 specimens (leg./coll. MNO), new species differs from goldfussi by the
specimens (coll. FHK, leg. K.I. Schnetler). lower number of teeth in the posterior series, a dif-
Brejning (beach exposure): 1/1 and 5/2 speci- ferent outline, a crenulated margin, a different
shell. mens (leg./coll. MNO); 2/2 specimens (leg./coll. sculpture and a more convex R. Janssen
ISL); 1/2 specimen (leg./coll. RLO); 1/2 specimen (1979: 30, pi. 1, fig. 16) included specimens of the
in L. from (leg./ coll. AHF). new species retifera, which it differs by 53
has several features: L. retifera a regularly Lepetella helgae sp. nov.
fine PI. 9a-b. crenulated margin, consisting of numerous 1, Fig.
which almost and denticles, are equal in strength
Locus — Coastal cliff North correspond with the radial sculpture on the outside typicus at Mogenstrup,
the the series of the shell. In new species posterior of Skive, Jutland, Denmark (Fig. 1).
— Glauconitic of hinge teeth is always the longer, while in L. Stratum typicum sand, Brejning Clay
Member of the Late retifera the anterior series is. In the new species the Vejle Fjord Formation, Oli-
concentric sculpture is dominated by the ribs, gocène (Chattian B).
the cancellate. Derivatio nominis — This is named after while in L. retifera sculpture is more new species
the is L. Lund Schnetler. Finally, new species generally larger than my daughter, Helga
the auricles less distinct and the shell is — Pl. MGUH 20 retifera, are Holotype 1, Fig. 9a-b, (coll. 028)
than that of L. The M.S. more equilateral retifera. new (leg. Nielsen).
— The shell is small and species differs from the Miocene L. anomala auct. Description cap-shaped.
1830 The is elevated and situated (non von Eichwald, ?) by having more con- apex highly precisely
with above the The vex valves, a coarser sculpture more widely posterior apertural margin. proto-
The knobs conch resembles small button. spaced concentric elements. coarse on a
is almost circular. In lateral view the inner ventral margin are placed more poste- The aperture
the anterior outline of the shell is whereas riorly in L. anomala and the entire ventral margin convex,
in anomala the There is is crenulate. The resilifer L. has an posterior outline is almost straight. no
well acute-angled triangular outline, while the outline external sculpture. The growth lines are
is concentric and somewhat variable in of the resilifer in the new species right-angled marked, triangular. strength.
On the inside of the shell the horseshoe-shaped
muscular is distinct. Propeamussium (Parvamussium) sp. impression fairly
Paratypes — Mogenstrup (coastal cliff): 1 specimen
Remarks — A rather limited material, consisting of (leg./coll. MNO).
11 defective valves and fragments, is available. Mr Discussion — The new species is related to the very
that the variable L. A.C. Janse (pers. comm., 1988) noted on species compressiuscula (Karsten, 1849).
the shell is fragments in his collection from Mogenstrup the In L. compressiuscula, however, distinctly
number of radial ribs the inner side of the shell and the is situated above on compressed laterally apex
the of the L. differs, in comparison to P. pygmaeum (von centre elliptical aperture. compressius-
Miinster, 1835) from the German and Danish cula is also relatively less elevated.
Late Oligocene. The Mogenstrup specimens all
have radial ribs the inside of both the left 11 on Lepetella jytteae sp. nov
lOa-b and right valve, while specimens of P. pygmaeum PI. 1, Fig.
ribs the valve and 12-14 show 15-17 on right on
the left also R. 1978a: Locus — cliff North one (see Janssen, 48). typicus Coastal at Mogenstrup,
valve Denmark The sculpture on the outside of the right of Skive, Jutland, (Fig. 1).
is rather similar to that of P. pygmaeum. The left Stratum typicum — Glauconitic sand, Brejning Clay
valve of Member of the Late Oli- has a cancellate sculpture, consisting Vejle Fjord Formation,
concentric ribs and line radial about 35 numerous gocène (Chattian B).
lines. Derivatio nominis — This species is named after Mrs
be- who found and donated the Apparently, there are consistent differences Jytte Hillersborg,
tween the Mogenstrup material and P. pygmaeum. holotype.
More and better preserved material is needed for Holotype— Pl. 1, Fig. lOa-b (coll. MGUH 20 029)
of the form. a proper interpretation Mogenstrup (leg. J FIR).
referred to — The shell is rather and Mr A.C. Janse (pers. comm., 1989) Description large cap-
be related: P. The is with its two taxa that might duodecemlamella- shaped. aperture suboval, greatest
width in the anterior The shell tum (Bronn, 1831), from the Early Pliocene of Italy part. height equals
about half the diameter of the and P. duodecemlamellatum bronni (Mayer, 1861) greatest aperture.
of The is situated about five sixths of the shell from the Middle Oligocene Hungary. apex at 54
The anterior outline is visible. The is subcircular with length. (lateral view) con- aperture a sharp
while the is almost labrum. The columella is with vex, posterior part straight. concave, a some-
The protoconch is button-shaped and smooth. what thickened inner lip demarcating the umbili-
On shell A is visible the the juvenile numerous growth-lines are cus. vertical impression on columel-
that it had almost circular which indicates the of visible, indicating an la, most probably position
About three millimetres from the the aperture. proto- operculum.
concentric The whorls have An conch a rather strong sculpture appears, a glossy appearance. ex- consisting of strong ribs, separated by interspaces tremely fine spiral furrow is present near the adapi-
almost cal Prosocline rather dis- of equal strength. On the largest available suture. growth-lines are
tinct. On the former specimen about 20 such ribs are present. The con- largest specimen two apertures centric ribs are crossed by weaker radial ribs, the are visible as rather prominent prosocline lines.
number of which increases towards the apertural Paratypes — Mogenstrup (coastal cliff): 3 specimens
25 margin, because secondary radialribs appear in be- (leg./coll. ISL); specimens (coll. ISL/leg.
the The 200 tween primary ones. radial ribs are rather MNO); specimens (leg./coll. MNO).
the side of the Discussion — The differs from Collonia prominent on adapical concentric new species ribs, but are also visible in the interspaces. (C.) minima (Philippi, 1843) by several features,
On the inside of the shell the horseshoe-shaped although juvenile specimens at first look rather muscular impression is fairly distinct. similar to this species. The protoconch of C. minima
— the nucleus is rather small Paratypes Mogenstrup (coastal cliff): 25 comprises one whorl, specimens (leg./coll. MNO); 2 specimens (coll. and the transition to the teleoconch is not sharp. In
C. minima the whorls in ISL, leg. MNO). increase more gently
Remarks — This rather differs diameter. its base furrow is large Lepetella species On a sharp present from its close the rib. This congeners by strong sculpture. to demarcating spiral spiral, as
well as those within the umbilicus, are coarser in C.
Collonia troelsi minima. its has (Collonia) sp. nov. Adapically, aperture adapically a
PI. 2, Fig. 2a-c sharper edge, and it is generally smaller than the
new species.
Locus typicus — Coastal cliff at Mogenstrup, North of Denmark Skive, Jutland, (Fig. 1). Homalopoma (? Leptothyropsis) sp.
Stratum typicum — Glauconitic sand, Brejning Clay PI. 1, Figs 12a-c.
Memberof the Vejle Fjord Formation, LateOligo- cène (Chattian B). Description — A rather limited material, consisting
Derivatio nominis — The is named after of a defective shell and seven ofthe species my single fragments
Schnetler. son, Troels Lund same specimen, was collected (leg. E. Nielsen,
Holotype — PI. 2, Fig. 2a-c (coll. MGUH 20 030) Arhus). Unfortunately, the protoconch is missing,
K.I. and the of is rather for (leg. Schnetler). state preservation poor,
— The shell is rather thin-walled which it has been make Description small, reasons not possible to a and wider than high, juvenile specimens very much final interpretation of the material.
of The shell is rather about so. The protoconch consists of three quarters a small, trochiform, as
wide. The defective consists of whorl, the nucleus being relatively large and slight- high as specimen on-
The with the teleoconch is 1 which and ly depressed. boundary ly 3/4 whorls, are moderately convex
and indicated almost orthocline radial increase sharp by an separated by deep sutures. They very furrow. rapidly in diameter.
In the largest specimen the teleoconch consists of The body whorl equals about five sixth of the
in total shell two convex whorls, rapidly increasing diameter, (estimated) height; the aperture is about and The half shell. The separated by deep sutures. last whorl as high as the complete base is only
about nine the of the with rather wide equals tenths, height aperture slightly convex, a umbilicus, sur-
six tenths of the total shell The rounded distinct rib. The is about height. peri- by a spiral aperture phery is regularly convex. The base of the shell is oval, the labrum is broken. The columella is con-
is the provided with a rather narrow umbilicus, which cave, and inner lip has a rather thick callus.
from base The whorls separated the by two close-set spiral ribs. spiral sculpture on the middle con-
In about is the umbilicus four weaker spiral ribs are sists of a thin, sharp spiral carina, which project- 55
visible. The is almost ing and situated immediately above the abapical spiral is aperture circular;
the labrurn is the columellar suture. On the base of the shell three further, not thickened, but
four of the inner is similar spirals are present. The spirals are part lip slightly consolidated.
The The dominant element is rather regularly spaced with concave interspaces. sculptural a
of these short distance from distance between the lowermost spirals strong spiral rib, lying at a the
On the and the whorl into two and the umbilical spiral is distinctly larger. upper suture dividing
be- flat and lower which largest fragment two secondary spirals develop parts: a adapical ramp, a part
is almost and becomes tween the two adapical spirals. adapically flat, slightly con-
below. The of the whorl The radial sculpture consists of about 30 pro- vex two parts together
in- form The into the base socline folds, separated by somewhat narrower an obtuse angle. transition
lines Towards the these folds is rounded. The are terspaces. upper suture regularly growth barely but almost orthocline. are slightly accentuated, gradually becoming visible, weaker and less well-defined downwards, and they Paratypes — Mogenstrup (coastal cliff): 2
The 1 M. disappear between the two adapical spirals. specimens (leg./coll. MNO); specimen (leg.
small the radial sculpture forms projections on S. Nielsen; coll. ISL).
Distinct lines Discussion — The differs from the rather spirals. prosocline growth are pres- species ent. From the suture run un- similar Tubiola 1 W. 1984: 128, upper they slightly sp. (A. Janssen, pi.
the whorls. dulating downwards, especially well visible on 6, fig. 3a-b) by having more angular
1964 has radial folds. Also between the spirals they are Daronia dingdensis Anderson, a concave
and carina which distinct. adapical ramp a demarcating
Remarks — The similar to the Below this carina the shell seems to be rather projects beyond ramp.
downwards Homalopoma (Lepthothyropsis) philipiana Dall (see whorl is slightly concave, changing to
Its Wenz, 1938: 341, fig. 799) in general outline, but strongly convex. apex projects only slightly additional material is needed. over the last whorl.
Tubiola Tubiola subangulata sp. nov sp.
PI. PI. 1, Fig. 14 2, Fig. 1
— — North The shell is small and consists Locus typicus Coastal cliff at Mogenstrup, Description very
whorls which of Skive, Jutland, Denmark (Fig. 1). of 2 3/4 moderately convex are
Stratum typicum — Glauconitic sand, Brejning Clay separated by deep, almost canaliculate sutures.
Late The wide. The Member of the Vejle Fjord Formation, shell is almost as high as pro-
than half Oligocène (Chattian B). toconch comprises slightly more a whorl,
— = and is from the teleoconch Derivatio nominis L. subangulata ( near- clearly separated by a
difference in colour. angular). The species is named after the obtusely radial orthocline rib, and by a angular whorls. The nucleus is relatively large.
— MGUH 20 The whorl of Holotype Pl. 1, Fig. 14; (coll. 032) body equals about seven eighths is (leg. M.S. Nielsen). the total shell height. The periphery regularly
— with transition into the base. Description The shell is very small and relatively convex a gradual
The rather but wide umbilicus is thick-walled, about 1 /% times as wide as high. It deep, not very
which distinct rib which is consists of about three convex whorls rapid- separated by a spiral ly increase in diameter. The protoconch consists of granulated by the growth lines. One additional half whorl The rather voluminous and in- rib is visible in the umbilicus. a only. spiral
surface. The is almost and neither flated nucleus has a very finely granulated aperture circular,
The boundary with the teleoconch is only in- labrum nor inner lip are thickened. The shell's
weak al- dicated the of rib on the surface is smooth and with rather by appearance a spiral glossy,
of the first whorl. orthocline lines. adapical part post-embryonic most growth
of the Material — 2 The body whorl equals about seven eighths Mogenstrup (coastal cliff): specimens total shell height. (leg./coll. MNO), 1 specimen (PI. 2, Fig. 1) (leg.
The base of the shell is slightly convex and MNO, coll. MGUH 20 033).
Discussion — The shell differs from Tubiola 2 demarcated from the deep and rather wide um- sp.
rib. In the umbilicus further from the Hemmoorian of bilicus by a spiral one Winterswijk-Miste 56
(A. W. Janssen, 1984: 128, pi. 6, fig. 8) by having Description — The shell is rather small and slender
lower and less columella. a apex a concave turriculate; about three times as high as wide. The
less than third the last whorl equals slightly one of
? Skenea sp. total shell height. PI. 2, Figs 3a-b The protoconch consists of four and a half
moderately convex whorls which are separated by Description — Only five juvenile specimens are rather distinct sutures. The nucleus is small. From available. The shell is very small and planispiral. the second protoconch whorl onwards the whorls The consists the protoconch of half a whorl, nu- with of are provided a radial sculpture, consisting cleus is voluminous. The transition into the teleo- numerous fine riblets. Initially these riblets are al- conch is and demarcated radial rib. sharp by a most orthocline, but they become slightly opistho-
The teleoconch consists of 3/4 convex whorl cline the last whorls. A on protoconch spiral It has and al- only. a very convex periphery, an sculpture consisting of five or six spiral threads is most flat part, which is from adapical separated the last and half present on two a protoconch the lower of the whorl The transi- part by a spiral. the whorls. These spirals are situated on abapical tion into the base is The umbili- convex gradual. of the whorl part and, although they increase in rib. The cus is wide and demarcated by a spiral strength in abapical direction, they remain weaker is subcircular and the labrum is broken. aperture than the spiral ribs. The two sculptural elements Through the umbilicus the base of the nucleus is result in reticulate of the together a ornament visible. of the abapical part whorl. The transition to the The sculpture consists of fine, thread-like very teleoconch is sharp and opisthocline. A the in spiral riblets. spiral near adapical suture The teleoconch consists of five and a halfwhorls
is rather distinct, while those on the particular in the largest specimen. These whorls are convex weaker. On the base periphery are the spiral and separated by deep sutures. The last whorl the wide umbilicus is distinct. demarcating very equals slightly less than one third of the total shell The lines the growth are opisthocline near the with the about height, aperture siphonal canal adapical suture. The is the one fourth. aperture rounded-oval,
Remarks — The material shows a stronger spiral labrum is broken and the columella is concave. ? Skenea sculpture than does sp. (A.W. Janssen, The two prominent spiral ribs (see below) cause an 1984: 129, pi. 6, fig. 9a-b). angular periphery. The base is almost flat.
The spiral sculpture starts immediately behind Ovirissoa sp. the boundary with the protoconch with three
of which the is thread-like spirals, adapical one — The found is im- Description only specimen very
and situated below the suture. This and shows immediately mature no teleoconch sculpture. Yet, does increase in the follow- spiral not strength on the apical angle would indicate that the specimen
whorls, but it becomes more distant is related ing gradually more closely to O. gottscheana (von from the adapical suture. The two abapical spirals Koenen, 1882) than to O. westfalica (A.W. the which is which has wider are prominent, especially upper one, Janssen, 1967), a distinctly apical of the whorl. The lowermost placed on the centre angle. lies distance above the spiral at a short abapical
suture. In-between the spirals the whorl is con- Cerithiopsis (s. lat.) antonjansei sp. nov.
so between the two lower PI. 2, Fig. 9. cave, especially spirals. A rather weak spiral rib, demarcating the base of
Locus — Coastal North the is visible the last typicus cliff at Mogenstrup, whorl, only on whorl. Close of Denmark this weaker rib is Skive, Jutland, (Fig. 1). to demarcating spiral a spiral
— the base. Stratum typicum Glauconitic sand, Brejning Clay present on
Member of the Vejle Fjord Formation, Late Oli- The radial sculpture consists of slightly opistho-
cline ribs which much the gocène (Chattian B). are narrower than
Derivatio — This after between them. The number of these ribs is nominis species is named my spaces friend Anton C. The Netherlands. first whorl Janse, Brielle, 15 on the post-embryonal and 14 on the
Holotype — PI. 2, Fig. 9, (coll. MGUH 20 040) body whorl. At the points of intersection of the
M.S. radial ribs and the (leg. Nielsen). two stronger spirals acute knobs 57
which are most on the one to the teleoconch is not distinct, but indicated occur, prominent upper very of these The rib the of and two spirals. adapical spiral con- by disappearance the spiral sculpture a
the radial without knobs darker colour. tinues across ribs, being change from a light to a
the formed. In-between the radial sculpture rather Even largest specimen has only one and a
lines half whorls. weak opisthocline growth are visible. teleoconch They are moderately con-
— 1 and The flat base Paratypes Mogenstrup (coastal cliff): specimen vex separated by deep sutures.
(leg./coll. MNO); 1 specimen (leg./coll. JHR). is demarcated by a spiral rib. The labrum is
the inner is well defined. Kirstinebjerg skov (beach exposure); 1 speci- broken, and lip not very
The canal is rather short and men (leg./coll. ISL). siphonal wide, slight-
Remarks — The turned new species differs from the ly backwards.
other of the The the teleoconch consists numerous species genus Cerithiopsis spiral sculpture on
Sea the of smooth the from the North Basin by glossy appea- two spiral ribs, accompanying upper
the rather and the lower suture the shell's sur- rance of the whorls which is caused by respectively,
The face in between these is for narrow spiral and radial sculpture. proto- spirals smooth, except
resembles that Ceri- about ten fine furrows. The radial conch of the new species of spiral sculpture
but differs about almost orthocline ribs thiopsis (s. lat.) serrula R. Janssen, 1978, consists of 12 strong
number of threads six in the whorl which between the ribs in the spiral (five to per run two spiral
but in Growth lines new species, only two C. serrula). Further- without crossing them. are visible,
teleoconch is much in the base In the available more, the sculpture coarser on they are prosocyrt.
related has C. serrula. There seems to be no closely juvenile specimens the base no further
species in the North Sea Basin. sculpture.
Paratypes — Mogenstrup (coastal cliff): 1
Laiocochlis (Laiocochlis) supraoligocaenica specimen, (leg./coll. MNO).
Discussion — The outline and of the sp. nov. sculpture pro-
PI. 8 toconch with the and illustration 2, Fig. agree description
in Gründel (1980: 251, fig. 35) of the protoconch
— Laiocochlis Dunker 1874. Locus typicus Coastal cliff at Mogenstrup, North of the genus & Metzger,
this is four of Skive, Jutland, Denmark (Fig. 1). In NW Europe genus represented by
Stratum typicum — Glauconitic sand, Brejning Clay species in the Lutetian of the Paris Basin (Coss-
Oli- 1889: the Pliocene Member of the Vejle Fjord Formation, Late mann, 41, pi. 2, figs 20-23), by
L. woodi van 1954 = gocène (Chattian B). species Regteren Altena, (
— This is named after its Cerithium and Derivatio nominis species granosum Wood, 1848, non Borson)
in the Late because this the Pliocene and Recent L. occurrence Oligocene, by species sinistrata chronostratigraphic unit is often referred to as (Nyst, 1835) (see Grundel, 1980: 251, erroneously
sinistria None of 'Upper' Oligocene. cited as L. Nyst). these, however,
— PL MGUH 20 be related the for Holotype 2, Fig. 8, (coll. 039) seems to to new species. Except
(leg. M.S. Nielsen). the Eocene L. passyi (Deshayes), which has smooth
Description — Only two juvenile specimens were teleoconch whorls, all these species have a strong
and sinis- and weaker lack- collected. The shell is turriculate, small spiral ornament a or completely
The last whorl about half radial tral. present comprises ing ornament. the total shell-height. The new species is the first representative of the
Laiocochlis be from the Late The relatively large protoconch consists of about genus to reported
of the In 2 x/i convex whorls which are separated by deep Oligocene North Sea Basin. the Oligo-
The nucleus is and inflated. of the Basin sutures. large The cene (Rupelian) Aquitaine (SW
the larval shell consists of of the Laiocochlis lesbarritzensis sculpture of a group France) species four inequidistant spiral bands, lying alongside the Vergneau, 1963 occurs. A lot consisting of 23
from the Gaas adapical suture, and about 15 very delicate spiral specimens locality (Lagouarde) was
the of the whorl. The loan Dr P. Lozouet grooves on remaining part kindly sent on to me by
bands national d'Histoire naturelle, upper spiral are separated by extremely (Museum Paris).
of The French differs from the narrow grooves. A radial sculpture, consisting species present one by about almost smaller smaller 13 rather strong, orthocline ribs, is its much protoconch and relatively
after whorl. The transition nucleus. A radial is absent. On the teleo- present % protoconch sculpture 58
three which riblets united conch primary spirals are present are are by a further spiral rib, below
than the radial L. somewhat stronger sculpture. Obviously which the radial riblets are weaker, for-
lesbarritzensis is related L. not closely to supraoligo- ming an angle of about 30° with the abapical
caenica. suture. The number of riblets is about 24 on the
penultimate protoconch whorl and it decreases to
Triforis (Trituba) sorgenfreii sp. nov. about 20 on the last whorl. The transition to the
PI. 2, Figs 5-7 teleoconch is sharp and opisthocline. The largest
available specimen comprises 6'A teleoconch
1978b Triforis (Trituba) aff. raulini Cossmann & Peyrot 1921 whorls which low and distinct are separated by
— R. Janssen, 171 p. (partim). These almost The last sutures. sutures are straight. ?1978b Triforis (Trituba) aff. raulini Cossmann & Peyrot 1921 whorl equals less than % of the total shell height. — R. 51 Janssen, p. 171, pi. 12, fig. The is aperture rounded rectangular. The siphonal
Locus typicus — Coastal cliff at Mogenstrup, North canal is broken in all available specimens, but
of Denmark be Skive, Jutland, (Fig. 1). seems to directed obliquely backwards. The
Stratum — Glauconitic base is flat and with typicum sand, Brejning Clay provided numerous prosocyrt
Member of the Vejle Fjord Formation, Late growth lines.
Oligocène (Chattian B). The spiral sculpture consists of two rather strong
Derivatio nominis — This is in the species named honour ribs, adapical one being situated at about three
of the late Professor Dr Th. the the Sorgenfrei fourths of the height of whorl, abapical one
who contributed about fourth of this The (Copenhagen), substantially to at one height. spirals are
the knowledge of Tertiary molluscs. much narrower thantheir interspaces. Between the
Holotype— PI. 2, Fig. 5, (coll. MGUH20 038) (leg. two spirals the shell wall is concave. A third weak
M.S. Nielsen). spiral is situated very close to the lower suture, but
Description — At Mogenstrup a rather limited ma- is usually covered by the next whorl. On the body
of five terial consisting fragmentary shells and one whorl this spiral demarcates the base of the shell.
juvenile individual with its protoconch preserved The radial sculpture consists of about 14-18
collected. The is ribs. At their was juvenile specimen designated opisthocline points of intersection the
holotype here, since it demonstrates the typical spiral and radial sculpture elements form solid
teleoconch the teleo- knobs. These knobs almost in sculpture on two preserved are equal strength on
conch whorls. Additional material from Kirstine- the two spirals. The abapical knobs almost touch
bjerg Skov, consisting of three fragmentary shells the suture. The spiral demarcating the base is
and three also used for the follow- fragments, was smooth. Opisthocline growth lines are especially
well visible the of whorl ing description. on concave part the and on
The dextral shell is rather small, turriculate to the base, which is otherwise smooth.
and about four times — defective cylindrical slender, as high as Paratypes Mogenstrup (coastal cliff): 1
of wide. The shell is very fragile because the high specimen; PI. 2, Fig. 6, (coll. MGUH 20 037) (leg.
numberof for which whorls, reason only fragmen- K.I. Schnetler); 1 defective specimen (leg./coll.
shells be tary or juvenile could collected. A recon- ISL); 1 defective specimen (leg./coll. MNO); 2
of the struction fragments suggest a shell height of defective specimens (leg./coll. JHR).
to 20 for adult Skov defective up mm specimens. Kirstinebjerg (beach exposure): 1
The protoconch comprises about six low and specimen, PI. 2, Fig. 7 (coll. MGUH 20 038 (leg. moderately convex whorls. These whorls are K.I. Schnetler); 2 defective specimens, 3
separated by deep sutures. The nucleus is small. fragments (leg./coll. ISL).
The last whorl about third Freden: 1 protoconch equals one specimen (coll. SMF 309 205), men- of the total The last five larval tioned in 171 protoconch height. R. Janssen, 1978b, p. (erroneously whorls with mentioned are provided a radial sculpture, con- two specimens were there).
sisting of numerous thread-like riblets which run Volpriehausen: ? 2 specimens (one of which il- from one suture to the other. On the second pro- lustrated in R. Janssen, 1978b, pi. 12, fig. 51) (leg. toconch whorl riblets A. coll. these are almost orthocline, von Koenen, Geologisch-Palaontolo-
become but on the last protoconch whorl they opis- gisches Institut, Gottingen).
At the time rib above Krefeld-Linn 2 defective thocyrt. same a spiral appears (water well): specimens the the The of the abapical suture. adapical parts (leg./coll. WLH). 59
mentioned — The of the Discussion protoconch sculpture new 171; two specimens were erroneously),
shows similar in loaned Dr This species a development as Triforis kindly to me by Janssen. specimen
aff. bitubulatus 1856 from the with the Danish material and (Trituba) Baudon, agrees consequently
Hemmoorian of Winterswijk-Miste in The Ne- it is referred to the new species. therlands (see A.W. Janssen, 1984: 160, pi. 7, fig.
of 6), but it differs by having a higher number Cirsotrema (Opaliopsis) subglabrum sp. nov.
in the the PI. 10 radial riblets. Furthermore, new species 2, Fig. protoconch whorls have their greatest convexity
it is 1987 Cirsotrema (Opaliopsis) — Schnetler, in Schnetler & about halfway between the sutures, whereas sp. 12 Beyer, p. 211, pi. 1, fig. situated more abapically in the species from
Winterswijk-Miste. The teleoconch whorls of the Locus typicus — Norre Vissing, clay-pit of Galten
have smooth in the Denmark. new species a peripheral spiral, Brickworks, Jutland, species from Miste this spiral is provided with Stratum typicum — Brejning Clay Member of the
rather Late knobs. In other features the two species are Vejle Fjord Formation, Oligocene (Chattian similar. B).
nominis — = almost I have compared the Danish material with topo- Derivatio (L.) subglabrum typical specimens of Triforis (Trituba) raulini Coss- smooth.
from — in Schnetler mann & Peyrot, 1921 Peyrehorade-Peyrere Holotype Schnetler, & Beyer, 1987,
loan Mr A.W. MGUH 17 (France), kindly sent on to me by pl. 1, fig. 12, (coll. 612).
and with such — Since first of this Janssen (coll. ROM), two topotypes Description my description in coll. ISL (leg. R. Janssen). The protoconch in species additional material has been collected, con-
whorls Its of three defective shells and the French species consists of three only. sisting a protoconch
whorls from and with nucleus is relatively large, the are highly Mogenstrup, a juvenile specimen
the second and and its from Skov. convex, especially so the third, protoconch preserved Kristinebjerg
The material matches the from provided with a collabral sculpture. On the initial new specimen protoconch whorl 12 rather weak orthocline ribs Norre Vissing perfectly. The largest specimen
consists of 3 teleoconch which number increases to 16 the from A are present, on Mogenstrup
the transition from the following whorls. The radial sculpture becomes whorls, including pro-
the last half toconch to the teleoconch. The number of radial stronger downwards, on protoconch
ribs is about 20 the last whorl. A fine whorl the ribs become opisthocline. on very spiral
The teleoconch of T. raulini differs in several ornament, consisting of eight spiral bands,
The Danish has delicate is respects. species relatively higher separated by spiral grooves, faintly
The discernible. For detailed the whorls and the two spirals are further apart. a more description
of knobs almost in in reader is referred to the above-mentioned two rows are equal strength paper. the Danish species, while in the French form the Paratypes — Mogenstrup (coastal cliff): 1 defective
knobs and also coll. abapical are stronger more specimen (Pl. 1, Fig. 13) (leg. J. Hillersborg; pointed. Finally, the peripheral spiral of the MGUH 20 041); 2 defective specimens (leg./coll.
Danish species is smooth, while the corresponding JHR); 1 juvenile specimen (protoconch only)
Skov spiral in the French species has distinct knobs, (leg./coll. MNO). Kirstinebjerg (beach ex-
undulated the — 1 causing a more suture. Thus, new posure) specimen (leg./coll. ISL).
demonstrates closer the species a relationship to
than the aff. koeneni species from Winterswijk-Miste to one Cirsotrema (? Opaliopsis)
from Peyrehorade. A.W.Janssen, 1967
From the German Late Oligocene R. Janssen PI. 2, Fig. 11
illustrated (1978b: 172, pi. 12, fig. 51) a specimen
from — At Volpriehausen, indicated as Trifora (Trituba) Description Mogenstrup a single specimen aff. raulini Cossmann 1912. This of two whorls & Peyrot, was found, consisting protoconch
and 4 3/4 teleoconch whorls. specimen might be conspecific with T. sorgenfreii, but the available for The shell is slender about three specimen was not yet study. turriculate,
rather times wide. The last whorl about I have compared a defective specimen as high as equals
(about two teleoconch whorls) from Freden (coll. one third of the total shell height. The preserved
whorls and SMF 309 205), mentioned in R. Janssen (1978b: protoconch are slightly convex sepa- 60
increase in of close-set threads be rated by deep sutures. They gradually very delicate, spiral can
with about 30 diameter. The whorls are provided noted. On the lower part of the whorls about ten
the fine The opisthocline radial riblets. The transition to extremely spirals are discernible. spiral
teleoconch appears to be sharp. rib demarcating the basal disc is visible in the
The teleoconch consists of 4 3/4 convex, some- suture.
The radial consists of what angular whorls which are separated by deep sculpture 18-20 promi-
smooth basal the almost orthocline ribs which the sutures. The shell has a disc, nent, run across
the labrum is broken whorls from the other. ribs aperture is rounded oval, one suture to These are
and the inner is The callus is thin. lip concave. almost as wide as their interspaces. In-between the
The is visible from the second spiral sculpture ribs growth lines are visible. On the adapical part
consists of three of the whorls the delicate teleoconch whorl onwards. It they cross spiral threads,
of which the is the in fine spiral ribs, adapical one resulting a granulation (magnification
strongest. This rib is situated at about two thirds x 50).
of whorl The three Remarks — The shell has overall height. spirals are regularly an Turbonilla-like
A smooth demarcates the spaced. further, spiral appearance because of the slender outline, the
basal disc, it is partly covered by the subsequent radial sculpture and the weak spiral sculpture. volution, but visible on all teleoconch whorls. No Opalia (Pliciscala) pusilla (Philippi, 1843) (see R.
from the Lower secondary spirals are inserted. The radial sculp- Janssen, 1978b, pi. 12, fig. 57)
16 Rhine has rather radial ture consists of about ribs which are stronger area, F.R.G., a similar
than the almost wide their but base spirals. They are as as sculpture, no spiral on the at the transi-
interspaces. On the first teleoconch whorl the tion from the base to the canal. Cirsotrema (Opaliop-
radial ribs are opisthocline, but they gradually sis) turbonillaeforme A.W. Janssen, 1967 is quite
become almost orthocline the whorls. similar in but differs its lower on younger many respects, by
At the of intersection between the two number of radial ribs instead of 18-20 ribs points (14 per
sculptural elements rounded knobs form, most whorl).
prominently on the adapical spiral rib.
Remarks — The Late de- Oligocene specimen Cirsotrema 2. (Opaliopsis) sp. scribed here differs from the Miocene C. koeneni by PI. 2, Fig. 13 having three primary spirals instead of two.
— A found which Description single protoconch was
differs from Cirsotrema (Opaliopsis) subglabra sp. Cirsotrema 1 (Opaliopsis) sp. nov. The specimen comprises five moderately con- PI. 2, Fig. 12 vex whorls, separated by deep sutures. The
nucleus is relatively large and inflated. The last
— A collected three whorls have fine radial Description single specimen was at a very sculpture, con-
The and of riblets 60 Mogenstrup. protoconch the first teleo- sisting numerous flexuous (about on conch whorls are missing and the aperture is very the penultimate whorl). In between this radial
fine poorly preserved. The shell is slender turriculate, sculpture numerous extremely spiral grooves
wide. The whorls The base of the shell is The about four times as high as ex- are suggested. convex.
in The consists is rounded the inner somewhat pand regularly diameter. specimen aperture oval, lip of 4 whorls which 3/4 are separated by deep, thickened. slightly undulating sutures. The height of the body Remarks — I refer this larval shell to Cirsotrema whorl measures about one third of the total shell of its to the (Opaliopsis) on account similarity pro-
The flat basal disc demarcated toconchs of C. height. is by a (O.) subglabrum sp. nov., as des- smooth spiral rib which is thickened in adapical cribed above, and of C. (O.) turbonillaeforme A.W.
1967 the Reinbekian of In direction in between the radial ribs. The aperture Janssen, from Dingden. is the lab the it be excluded that rounded-oval, rum and abapical part fact, cannot the present of the broken off. A further lies does the Cir- aperture are spiral specimen belong to same species as at the transition from the base to the canal. sotrema (Opaliopsis) sp. 1, the protoconch of which
At about of their the three quarters height is unknown. This assumption seems to be sup- whorls show faint Above this the weak teleoconch of a angulation. angula- ported by very sculpture
in the of the number the last-named tion, adapical part whorls, a species. 61
Apiocypraea (Apiocypraea) The protoconch is paucispiral and consists of
cf. R. 1978 humbergi Janssen, two whorls, separated by deep sutures. The
PI. 3, Fig. 4a-b nucleus is large and vesicular. On the last % whorl
five The boun- orthocline radial ribs are present.
Description — A rather limited material, consisting dary with the teleoconch is sharp and marked by
of radial rib. The one protoconch, one juvenile shell with half a a slightly flexuous protoconch sur-
teleoconch whorl and isolated face delicate preserved one shows an extremely microsculpture,
lab is available. of about 20 furrows whorl. rum, consisting spiral per
The protoconch is conical and comprises about This microsculpture is visible on well-preserved
five whorls which moderately convex are separated specimens only.
by rather shallow sutures. The nucleus is relatively The largest specimen available has 3 3/4
and the whorls of large inflated, following are provid- teleoconch whorls which are angular because a
ed with a cancellated sculpture, produced by ring of projecting spines placed just above mid-
fine and thread- whorl. The of the whorl numerous opisthocline prosocline adapical part is almost
like riblets. The transition the teleoconch whereas the is to can- flat, lower part slightly convex. The
not be studied on the juvenile specimen. base of the body whorl is regularly constricted.
The the half teleoconch The columella is sculpture on preserved slightly flexuous, the aperture is
whorl consists of ribbon-like ribs which rounded-oval and constricted below spiral are to the long,
than their and canal. The inner narrower interspaces. Secondary spirals narrow straight siphonal lip
develop rapidly. The radial sculpture consists of is simple, smooth and well defined; the labrum is
riblets which the somewhat thickened. prosocyrt are weaker than spiral
ribs. The combination of the two sculptural The spiral sculpture starts with three weak
elements results in reticulate The the of which is the a ornament. spirals, upper one strongest.
radial in The sculpture disappears soon apertural spirals are separated by much narrower in-
direction. On the whorl five terspaces. body spirals are pres-
The isolated labrum lacks its The adapical part. ent below the strong adapical one.
outline of the labrum, its size, and also the number The radial sculpture consists of laminate ribs
which almost orthocline. On the first and the shape of the teeth, agree very well with the are teleo-
illustration of whorl description and Apiocypraea (A.) conch there are 15 such ribs, increasing to
in R. 18 20 the whorl. At the of inter- humbergi Janssen (1978b: 197, pi. 14, fig. to on body points
86). The protoconch closely resembles the descrip- section with the spirals the ribs become crispate.
On tion and illustration of the species A. (A.) subphy sis the strong upper spiral spines are formed that
ionalis from the Hem- somewhat in direction. On the septemtr (Schilder, 1929) project adapical
moorian The Netherlands base similar is but it becomes of Winterswijk-Miste, a sculpture present,
(A.W. Janssen, 1984: 193, pi. 82, fig. 2). I there- gradually weaker downwards. In-between the
fore refer the Danish with to A. radial material, a query, sculpture fine, slightly flexuous growth lines
visible. humbergi. are
Remarks — I compared the fairly rich material
from with of the Trophonopsis (Pagodula) angustevaricata Mogenstrup specimens genus
from number of localities. (Gripp, 1912) Trophonopsis a From the
PI. 2, Fig. 14a-b Danish Late Oligocene I studied material from
Skanderborg (1 specimen, coll. ISL) and Brejning
1912 Murex in angustevaricata nov.sp., Gripp, Koch, Gripp & (2 specimens, coll. ISL, and 2 specimens, coll. Franke, 22, fig. 4. p. These lots with MNO). seem to agree the material 1963 Trophonopsis (Pagodula) semperi angustevaricata (Gripp, from Mogenstrup with regard to the protoconch — 6. 1912) Tembrock, p. 314, pi. 5, fig. on the two from The
1968 — Trophonopsis semperi (Koenen, 1872) Rasmussen, p. (worn specimens Brejning).
number of radial ribs is lower the 125 (partim, non von Koenen). on Brejning
but specimens (10 to 12 ribs, 18 on those from
— is slender and its In these Description The shell fusiform; Mogenstrup). addition, ribs are
about width height equals 2 lA times the opisthocline abapically. Thus, the Brejning mate-
(estimated, because the siphonal canal is broken in rial probably does not belong to Trophonopsis
all but be specimens). angustevaricata, seems to closer to T. semperi 62
Derivatio nominis — This is named in (von Koenen, 1872) (compare Tembrock, 1963, new species
who the pi. 5, fig. 5). honour of the late Dr J.P.J. Ravn, was
From the Danish Late Miocene one specimen of first to describe this species.
— 14a-b Trophonopsis semperi from Gram was studied (coll. Holotype Ravn, 1907, pi. 5, fig. (coll.
ISL). Its protoconch differs from the Mogenstrup MGUH 505).
smaller - The shell is fusiform and rather form by having a nucleus, a slightly angu- Description large,
of thick-walled. The times the lar last whorl and a microsculpture consisting height equals two
three of the prosocline radial threads and spirals that cross the diameter, the last whorl about quarters
and the and canal radials at right angles (Rasmussen, 1968: 123). On total shell-height, aperture the
from the half of the total shell the specimens Mogenstrup microsculp- slightly more than height. have but ture consists of spiral furrows only. On the teleo- The adult shell may up to eight whorls, conch of the Gram specimen the angulation of the no complete specimen was found. On one defec-
the and tive shell the is but it is rather whorl is situated on the middle of whorl, protoconch present, there The number of It consists of two are only two spirals present. poorly preserved. relatively large radial ribs is about 12 on the shell from Gram and whorls, of which the first one is flattened and
than in while second is the spines are projecting more adapically dome-shaped, the one more con-
These whorls the Mogenstrup material. From Made (Danish vex. are separated by deep sutures,
Gram Clay, Late Miocene) Rasmussen (1968: the nucleus is large and apparently vesicular. The
second whorl with 124) mentioned a large specimen having only eight protoconch is provided a spiral
nine radial ribs. of about 11 weak or ornament, consisting spiral
transition the teleoconch is distinct Rasmussen (1968: 125) united T. semperi semperi bands. The to five close-set orthocline (von Koenen, 1872) and T. semperi angustevaricata and marked by very
since vari- lines. behind this (Gripp, 1912) the number of spirals was growth Immediately boundary
the radial ribs and the number of able in his material from Gram. In my opinion T. occur, spiral teleoconch semperi and T. angustevaricata should be considered bands increases markedly on the first
because of the difference in whorl. two separate species
The teleoconch consists of about seven protoconch and the considerably higher number of up to
ribs the whorls which and radial on specimens from Mogenstrup. are moderately convex
The of radial Tembrock (1963: 314) treated T. angustevaricata separated by deep sutures. presence
that both makes the line undulate as a subspecies of T. semperi and stated oc- ribs (see below) suture on
first cur in the 'Upper Miocene' of Liineburg, F.R.G. the teleoconch whorls.
This of action does The is oval and constricted into the course not comply with the aperture
and well-defined canal which is present-day subspecies concept seems to con- siphonal relatively
short and somewhat twisted backwards. The firm that T. semperi and T. angustevaricata are
labrum is and both separate species. strongly regularly thickened,
The specimens from Brejning seem to belong to interiorly and exteriorly. Exteriorly the apertural
inner is T. semperi and thus T. semperi and T. angustevaricata margin is abruptly constricted. The lip
in the North Sea Basin from the Late well and the columella is are present defined, smooth, slightly
thickened the transition the canal. A Oligocene to the Late Miocene. at to solid,
rounded parietal tooth is present. The transition to
Searlesia ravni the base with the canal is and sp. nov gradual slightly
PI. 2, Figs 16, 17a-b concave.
The spiral sculpture of the teleoconch starts with
1907 aff. Koninckii — Ravn, bands which Fusus Nyst p. 119, pi. 5, fig. 22-23 flat spiral are separated by nar-
14a-b. The row furrows. number of these spirals is cons-
1987 Angistoma brueckneri (Beyrich, 1856) — Schnetler, in but the whorl of tant, on body a generation Schnetler & Beyer, p. 213 (partim, non Beyrich, non The 3-4 brueckneri danica secondary spirals develops. adapical pi. 2, fig. 3 = Angistoma subsp. nov.).
primary spiral bands are weaker than the lower
— the canal have similar Locus typicus Cilleborg (former clay-pit), ones. The base and a spiral
Jutland, Denmark. ornament.
— The radial consists of 16 radial ribs Stratum typicum Glauconitic clay, LateOligocene sculpture on
(Chattian A). the first teleoconch whorl, decreasing to 13-15 on 63
wide their show number of dif- the last whorl. These ribs are about as as and Fakkegrav specimens a
On the whorls the radial ferences with well similarities A. interspaces. younger as as to typical sculpture gradually becomes weaker, and on the brueckneri, for which reasons I treat this material as
of penultimate whorl it fades completely. Weak, al- a geographical subspecies A. brueckneri.
lines visible. Below the with other the — most orthocline growth are Affinities some species of 'Fusus' group
curved backwards. The koninckii 1845 adapical suture they are slightly Rupelian ‘Fusus’ Nyst, (sensu
Paratypes — Mogenstrup (coastal cliff): 1 specimen Beyrich) has more angular and more convex
of (PI. 2, Fig. 16a-b, (coll. MGUH 20 047, leg. S.B. whorls, and radial ribs on the adapical part all
whorls. Its and tooth Andersen); 1 specimen (PI. 2, Fig. 15) (coll. size, aperture parietal are
MGUH 20 K.I. 1 similar Searlesia ravni which seems to be 046, leg. Schnetler); specimen to sp. nov.,
‘Fusus’ koninckii (leg./coll. ISL); 1 specimen (leg./coll. MNO). closely related. In my opinion
also be considered the Brejning (beach exposure); 1 specimen should to belong to genus
(leg./coll. MNO). Searlesia, rather than to Euthria, as was suggested
Discussion — I have recently discussed the Danish by R. Janssen (1979a: 290). Species of Euthria
and have subsutural without dis- material of the genus Angistoma pointed out a concave depression
brueckneri tooth and some differences between the German A. tinct spiral sculpture, no parietal a
canal. (Beyrich, 1856) and the Danish material (Schnet- longer, narrow siphonal
1987: More from the Ger- ler, in Schnetler & Beyer, 213). mate- Angistoma feldhausi (Beyrich, 1856)
it Middle rial is available now, and seems justified to man Oligocene (Neustadt-Magdeburg)
the Danish material from has 10-13 3-4 denticles the columella separate Cilleborg, Brej- spirals, on
and is smaller than Searlesia ravni ning and Mogenstrup from A. brueckneri. The sp. nov. specimens from Arhus, Norre Vissing and Fakke- Searlesia mitgaui (von Koenen, 1867) from the
should be treated as a German Late has radial ribs grav Badehotel, however, Oligocène (Söllingen)
geographical subspecies of A. brueckneri. In my on all whorls and two generations of secondary
referred that do reach the of the opinion the new species is better to spirals not strength
weak denticles Searlesia, in view of its protoconch, spiral sculpture primary ones. On the inner lip two
that the wall and the and aperture. are present, is, on parietal at
Searlesia ravni differs from A. transition to the canal. sp. nov. typical
number of brueckneri by a considerably higher For the material from Arhus, Norre Vissing and
rather Badehotel I here introduce the spiral bands which are equal in strength, by Fakkegrav name the absence of knobs the columella and danica and on the Angistoma brueckneri subsp. nov.,
of solid tooth. The whorls the illustrated Schnetler presence a parietal are designate specimen by (in
and the shell attains Schnetler coll. MGUH more regularly convex a & Beyer, 1987, pl. 2, fig. 3;
dif- 17 The larger size. Finally the protoconchs are very 618) holotype. following paratypes are
ferent: has available: Arhus - 1 A. brueckneri a relatively small, (former railway excavation) multispiral protoconch, whereas in Searlesia ravni specimen, coll. D.G.U. (leg. P. Harder); Norre the larval shell is large and consists of only two Vissing (clay-pit of Galten Brickworks) - 1 whorls. specimen (leg./coll. ISL), 1 specimen (coll. MNO,
The specimens from Arhus (Harder, 1913, p. leg. K.I. Schnetler) 2 specimens (leg./coll. MNO);
78, pi. 6, Figs 9-10), from Norre Vissing Fakkegrav Badehotel (coastal cliff) - 1 specimen
(Schnetler, in Schnetler & Beyer, 1987, p. 213, pi. (leg./coll. MNO).
2, Fig. 3) and from Fakkegrav Badehotel (coll.
with the German A. brueckneri Buccinidae et indet. MNO) agree species gen. sp. in size and general outline, and in the absence of Text-Fig. 6
Danish a parietal tooth. Unfortunately, no specimen has its protoconch preserved. They have Description — Only two defective juvenile slightly angular whorls with about 16 spiral bands specimens are available (leg. M.S. Nielsen). In the
better of these the and the first of almost equal strength and two denticles on the one protoconch
have columella, whereas the German specimens teleoconch whorl are preserved. The second
whorls 6 9 of the second non-angular with about to spiral bands, specimen, demonstrating a part somewhat variable in strength, and four denticles teleoconch whorl, is very defective because of a
the axis of on the columella. Thus, the Arhus, Norre Vissing split parallel to the shell. 64
1 from collection. All these The protoconch comprises % whorls, my private specimens
The nucleus is and therefore their separated by a deep suture. large, are full-grown apical parts are
globular and slightly vesicular. On the last half worn. The Mogenstrup material seems to agree
whorl al- L. of protoconch fine, irregularly distributed, best with rarus with regard to the convexity
orthocline crossed most growth-lines are present, the whorls and the number of primary spirals. On
delicate which are the the number of is 12, by some ten very spiral furrows, Brejning specimen spirals
of from Gram merely suggested. These furrows are most promi- while the specimens L. ventrosus
of transition nent on the upper part the whorl. The have only 7 primary spirals. The very juvenile
is indicated about 15 of the final in- to the teleoconch gradual, by state material, however, prevents a close-set growth-lines. terpretation.
Dr R. Janssen (SMF) kindly examined the illus-
trated specimen. In his opinion it cannot be re-
ferred to Liomesus rarus (pers. comm., 1990), which
differs much 2.5 by a larger protoconch (diameter
mm), which is flat and dome-like. Furthermore
the spiral sculpture of that species is much coarser.
Latirus (Pseudolatirus) sp.
Pl. 3, Fig. 2
Description — One juvenile specimen, consisting of
the col- protoconch and one teleoconch whorl, was
lected at Mogenstrup (leg./coll. JHR). In addi-
four known from tion, juvenile specimens are
Kirstinebjerg Skov (leg./coll. ISL).
The shell is fusiform, about three times as high
wide. The of the whorl is about six as height body
tenths of the total shell height.
The protoconch is conical, it comprises 4-4
whorls which moderately convex are separated by
The nucleus is small. The whorls deep sutures. are
smooth and glossy, except for the last half whorl, 6. Buccinidae frontal b: Fig. sp., Mogenstrup; a: view, rather on which opisthocyrt growth lines are apical view. Coll. MGUH 20 276 (A.W. Janssen distinct and three rather weak ribs del., 1990; bar length is 1 mm). spiral appear.
The adapical one of these is the weakest. A further
The teleoconch whorls moderate- fine is all whorls in preserved are spiral present on protoconch
and almost canali- in front of the ly convex separated by a deep, the abapical suture. Immediately
with the teleoconch 2 culate suture. The aperture is ovoid, abapically boundary or 3 slightly
into rather wide and si- radial ribs The transition passing a short, oblique opisthocline are present. phonal canal. to the teleoconch is gradual.
The consists of about teleoconch whorl is in spiral sculpture ten prim- Only one convex present
which the available The of the round- ary spiral bands, are separated by narrower specimens. height
furrows. The adapical spiral is somewhat more ed oval aperture is about halfthe total shell height.
canal is The prominent. A radial sculpture is absent, except for The siphonal long and rather narrow.
inner is smooth weak orthocline growth-lines. lip and not very strongly protrud-
— A weak fold is col- Discussion The features of protoconch and ing. oblique present on the
teleoconch rather well with the umella. agree description and illustration of Liomesus The of the teleoconch consists of rarus (Beyrich) (see spiral sculpture
Beyrich, 1856: 250, pi. 17, figs 6a-b). I compared three rather narrow spiral bands which are the
materialwith of this continuations of the the the present a specimen species spirals on protoconch.
from with two of L. Below the weak band Brejning and specimens ven- adapical suture a spiral ap-
from both of whorl. The radial trosus (Beyrich, 1856) Gram, samples pears on a quarter a orna- 65
nine ribs the first ment comprises strong on whorl separated by a rather distinct spiral furrow; four to
which weaken the transi- five distinctly downwards, on very weak, merely indicated spiral bands are
tion the canal. The base and the above the lower to siphonal canal present two ones. The transition
are with a similar to the teleoconch is difficult but provided spiral sculpture. to detect, sug-
Discussion — The limited Danish material consists gested by close-set growth lines.
of for which On the two found the teleoconch juvenile specimens only, reason a spe- specimens con-
cific identification is the sists of 2 whorls which impossible. Yet, proto- up to Vi are slightly convex,
conch characteristics differ from those of the Mio- almost flat below the having an part adapical
North Sea Basin Latirus The whorls cene species (Pseudolatirus) suture. are separated by deep sutures.
rothi (Beyrich, 1856) by a number of features. The The base is strongly constricted below the peri-
of the Danish has smaller The is rounded-oval protoconch specimens a phery. aperture and rather
and constricted apical angle only two or three radial ribs on its abruptly at the transition to a rather
terminal while in L. rothi the ultimate wide and canal. The labrum part complete short, open siphonal
protoconch whorl is provided with radial ribs. Fur- is broken, the inner lip and the left margin of the
the columellarfold to be weaker in canal The callus is thermore, seems are S-shaped. thin, lying deeper
the Danish specimens. Sorgenfrei (1958: 232, pi. and thus showing traces of resorption of calcareous
described and illustrated the 50, fig. 157) proto- matter.
conch of L. rothi from the Arnum and The with six Formation, spiral sculpture starts to seven
he also found the radial of the ornament protoconch primary spiral bands which are separated by
be restricted the last third half whorl. The to to one or rather deep furrows. adapical spiral band is
This feature also noted in of 20 from the was a sample juve- separated second one by a somewhat
nile of L. rothi from well furrow and into sutural band specimens a water at Ny deeper develops a on
Lifstrupvej (South ofVarde, Denmark). Thus, it the second teleoconch whorl. The spiral bands
well be that extensive number might possible a more mate- two and seven are weaker than the others.
rial from Denmarkallows ataxonomic subdivision. The number of increases spirals by one on each of
the teleoconch whorls because of further spirals ap-
Clavatula from below the mogenstrupensis sp. nov. pearing abapical suture. No secon-
5 PI. 3, Fig. dary spirals are inserted. The base and the canal
with similar are provided a spiral sculpture.
Locus — Coastal cliff North The On the first typicus at Mogenstrup, radial ribs are opisthocyrt.
of Denmark teleoconch whorl Skive, Jutland, (Fig. 1). they run from one suture to the
Stratum — but the second whorl become typicum Glauconitic sand, Brejning Clay other, on they
Member of the Late Oli- Vejle Fjord Formation, weaker, especially so below the adapical suture
where the band gocène (Chattian B). sutural is developed. The radial
Derivatio nominis — This species is named after the sculpture is rather similar to that of the well-
type locality. known species Fusiturris duchastelii (Nyst, 1836).
PI. 5 MGUH 20 The number of 18 the last whorl. Holotype— 3, Fig. (coll. 051) (leg. ribs is about on
K.I. The radial ribs follow the of the Schnetler). course growth
— The shell is rather slender visible between the Description small, lines which are radial ribs,
about fusiform, two and a half times as high as with the deepest point of the sinus situated be- wide. The of the last whorl tween band number three and height equals slightly spiral four, as than half the total the of the height, height aper- counted from the adapical suture.
is about fifths of the total — from the ture two height. Paratypes One specimen type locality The protoconch comprises about 5 % slightly to (leg./coll. MNO).
whorls which Discussion — The moderately convex are separated by new species shows some resem- deep sutures. The nucleus is small and slightly he- blance with C. chattica R. Janssen, 1979, but dif-
The whorls fers whorls terostrophic. protoconch are smooth, by having more convex protoconch
for the last which is with and radial ribs the last except one provided stronger opisthocyrt on pro- about 18 radial ribs. The opisthocyrt last proto- toconch whorl. On the teleoconch the radial sculp- conch whorl is and has weak is more convex a spiral ture stronger. Finally, the number of primary
bands is six in ornament, consisting of two weak spiral spirals to seven the new species, whereas above the abapical suture. These bands are C. chattica has only five primary spirals. 66
Stratum — Member of the Microdrillia (Andersondrillia) subgen. nov. typicum Brejning Clay
Vejle Fjord Formation, Late Oligocene (Chattian
Diagnosis — A subgenus of Microdrilliashowing the B).
features the On the Derivatio nominis — This is named after its same protoconch as genus. te- species
leoconch the sickle-shaped riblets on the subsutural type locality.
— 20 flat of the whorl are absent. On the PI. MGUH 054) or concave part Holotype 3, Fig. 8a-b, (coll.
lower of the whorls the combinationof M. S. part spiral (leg. Nielsen).
results in less — The shell is rather small and slender and radial ornament more or promi- Description
the carina. about 2.8 times than wide. The nent knobs, especially so on fusiform, higher of Type species — Microdrillia (Andersondrillia) grippi An- height of the last whorl equals about two thirds
of the derson, 1964 (Miocene, North Sea Basin). the total shell height, the height aperture is
total shell The — The the about half the Discussion new subgenus resembles ge- height. largest
Harris 1891 and consists of about whorls of which nus Asthenotoma & Burrows, espe- specimen seven
cially the species A. obtusangula (Brocchi, 1814) in 3% belong to the protoconch.
The is conical with general outline and sculpture of the teleoconch, as protoconch broadly convex
A.W. The whorls which sutures. The pointed out by Janssen (1984: 299). pro- are separated by deep
with Microdrillia nucleus is inflated and toconch, however, agrees entirely small, slightly apparently
of Microdrillia smooth. whorls s.str. From typical representatives Succeeding protoconch are provid-
M. M. ed with radial which consists of s.str., such as bicingulata (Sandberger, 1860), a sculpture fine,
serratula (Bcllardi, 1878) and M. teretiaeformis A.W. opisthocyrt ribs, numbering about twenty on the
the differs first whorl. The ribs backwards below the ada- Janssen, 1972, new subgenus markedly run
by the complete absence of the sickle-shaped riblets pical suture and form an opisthocyrt curve across
in the subsutural excavated of the the whorl. Near the the ribs bend part whorl. These abapical suture
their The radial species generally are also more slender and forwards and disappear. sculpture gra-
increases in the last spiral ornament is more prominent. dually strength; on protoconch
The is in the North whorl the ribs are more and new subgenus represented widely spaced promi- terminal half whorl about Sea Basin by two species, viz. Microdrillia (Anderson- nent. On the protoconch
1964 and M. such ribs below the drillia) grippi Anderson, (A.) brejningen- seven are present. Immediately
sis from the and illus- suture lies a fine rib; on the middle sp. Nov. Judging description adapical spiral
the of the whorl few furrows indicated. tration in Peyrot (1932: 71, pi. 7, figs 6-7) spe- part a spiral are
described from Late of the whorls cies M. venusta (Peyrot, 1932), On the abapical part protoconch an is of Oligocene sediments at Peyrere (SW France), prob- extremely fine sculpture present, consisting
W. thinlines the radial ribs, visible ably also belongs to the new subgenus. A. Jans- crossing exclusively
this be in in front sen (1984: 299) considered taxon to closely well-preserved specimens. Immediately
with M. of the the teleoconch several flexuous related to, or even conspecific grippi. transition to
‘Microdrillia’ speyeri (Koch & Wiechmann, 1872) ribs are present.
The 3 3/4 teleoconch whorls has a teleoconch rather similar to Microdrillia s.str., largest specimen has
which due central but the paucispiral protoconch excludes a close are carinate to a prominent,
divides whorl into distinct- relationship. According to A.W. Janssen (pers. spiral rib. This rib the a
and almost vertical that should better be referred concave an comm., 1990) species ly adapical part
The base is convex and to another, possibly new genus. abapical part. slightly
Derivatio — The is named rather constricted canal of nominis new subgenus abruptly to a siphonal
medium The is afterProfessor H. -J. Anderson(Marburg, F.R.G.) length. aperture elongated-oval
contributions with transition into the canal. The for his substantial to the knowledge of a gradual
lab is broken all available The Tertiary molluscs. rum on specimens. inner lip is smooth, the callus is thin and excavated,
Microdrillia (Andersondrillia) brejningensis thus showing traces of resorption of calcareous
sp. nov. matter.
PI. 3, Figs 9a-b, 90 The spiral sculpture starts with three primary
is the spirals, of which the adapical one strongest.
Locus — Beach at South This accentuated rib, or better, carina, is typicus exposure Brejning, spiral
of Vejle Fjord, Jutland, Denmark. situated at mid-whorl, and the two other spiral ribs 67
distributed the of Microdrillia are regularly on abapical part (Microdrillia) ingerae sp. nov.
the whorl. Immediately above the abapical suture PI. 3, Figs 7a-b
a weak spiral rib is seen from the second teleo-
whorl onwards. The Locus — Coastal cliff North conch spirals are distinctly typicus at Mogenstrup,
than their A of Denmark narrower interspaces. large specimen Skive, Jutland, (Fig. 1).
Stratum — Glauconitic from Mogenstrup shows an aberrant spiral sculp- typicum sand, Brejning Clay
above the ture in that the third spiral from is com- Member of Vejle Fjord Formation, Late
absent. In all other features the shell pletely agrees Oligocène (Chattian B).
with the material. In the Derivatio nominis — This is named after completely remaining species my
whorl four Schnetler. concave subsutural part of the spiral wife Inger Lund
— PI. MGUH 20 bands are present on the first teleoconch whorl, Holotype 3, Fig. 6a-b, (coll. 053)
but K.I. separated by narrow furrows, they soon disap- (leg. Schnetler).
On the whorl weak — The shell is rather pear. body only one very spiral Description small, fusiform,
is indicated immediately above the adapical spiral about 2.4 times higher than wide, rather solid, the
rib. No inserted. The base is of the less than secondary spirals are height aperture equals slightly half
with end the total shell provided a similar sculpture to the of the height.
canal, a further seven spirals are present. The protoconch is broadly conical and consists
The radial consists of 15-18 ribs which of five whorls which sculpture moderately convex are
than their These ribs The nucleus is small are narrower interspaces. are separated by deep sutures.
almost orthocline and only visible below the and inflated. The initial one and a half protoconch
whorls carina. At the points of intersection of spiral and are smooth, the following ones are provid-
with radial sculpture knobs are formed, most promi- ed a weak spiral rib above the abapical suture
the carina. The lines and with radial ribs which nently on growth are numerous narrow
especially well visible above the carina, where gradually increase in strength. Near the adapical
radial sculpture is absent. The rather deep sinus is suture these ribs run strongly backwards, but they
situated above the carina. On the become last whorl immediately opisthocyrt on the protoconch
base the lines form flat and here about fine in- growth a very prosocyrt eight spiral grooves are
dicated. An of curve. oblique microsculpture, consisting
Paratypes — Brejning (beach exposure): 3 very fine divergent lines crossing the radial ribs at
less is specimens (leg./coll. MNO); 1 specimen (leg. more or right angles present in between the
M.S. ribs shell In Nielsen; coll.ISL). (preserved on one only). very juvenile
1 PI. it be that the base of the Mogenstrup (coastal cliff): specimen, 3, specimens can seen pro-
K.I. toconch is Fig. 9, (coll. MGUH 20 055) (leg. Schnetler); demarcated by a strong, smooth spiral
weaker rib below. 1 specimen (leg./coll. LJL); 5 specimens (leg./coll. rib followed by a spiral On the
base of the the lines form ISL); 4 specimens (leg./coll. MNO); 1 specimen protoconch growth a pro-
socline The transition the teleoconch is (leg./coll. JHR). curve. to
Discussion — This resembles Microdrillia distinct but not species very sharp.
but number of grippi Anderson, 1964, differs by a The teleoconch of the largest specimen consists
features. The has slender of three whorls which new species a more pro- are separated by deep
toconch with the radial sculpture starting im- sutures. The body whorl equals about two thirds
after the nucleus. The teleoconch has of the total shell The whorls divided in- mediately a height. are
distinct of the whorl without and almost concave adapical part to a slightly concave adapical part an
The below the carina spiral sculpture. spirals are ilat abapical part by a carina which lies at mid- there sutural band. The narrower and is no spiral whorl. The base is constricted to a rather short
radial sculpture consists of almost orthoclinic ribs, canal. The callus is thin and the columella is
while M. grippi has prosocline ribs. The sinus in smooth, showing traces of resorption of calcareous
the lines of the is situated oval and growth new species matter. The aperture is narrows gradual-
below the middle of the of the the canal. concave part whorl, ly into siphonal
while in M. grippi the sinus lies at mid-whorl. The The spiral sculpture starts with a strong carina
shell has because of the a glossier appearance on the middleof the whorl. Immediately below the
radial and sharper sculpture, the weak sculpture adapical suture a relatively broad sutural band is
the absence of and the weak secondary spirals. situated above abapical suture a 68
spiral rib is visible. From the second teleoconch Pleurotomella (Pleurotomella) rasmusseni whorl onwards the carina consists of close-set two sp. nov. spirals. Both the sutural band and the spiral rib PI. 3, Figs lla-b below the carina gradually increase in strength. A secondary spiral rib develops in the centre of the Locus typicus — Coastal cliff at Mogenstrup, North
between the carina and the of Denmark space adapical spiral. Skive, Jutland, (Fig. 1).
This becomes than — secondary spiral soon stronger Stratum typicum Glauconitic sand, Brejning Clay the abapical spiral rib. On the base and the canal Member of the Vejle Fjord Formation, Late Oli- a further eleven spirals are present. A generation gocène (Chattian B).
base and Derivatio nominis — The is in honour of secondary spirals develops on the on species named the neck of the canal. of State Geologist Dr Leif Banke Rasmussen,
The radial sculpture is rather complex. From Geological Survey of Denmark, Copenhagen, who the form accentuated carina, on which they has contributed substantially to the knowledge of knobs, thickened opisthocline ribs (following the the molluscs and the stratigraphy of the Danish
of the direction growth lines) are present, quickly Neogene. diminishing in strength downwards and disap- Holotype — PI. 3, Fig. lOa-b, (coll. MGUH 20
the second below the K.I. pearing already near spiral 056) (leg. Schnetler). carina. Their number increases from about 18 on Description — The shell is rather small and thin- the first teleoconch whorl 24 26 the to to on body walled, subfusiform, about two and a half times whorl. The than wide. The the last whorl growth-lines are distinct, especially so higher height of
of three fifths of the total shell The on the concave subsutural part the whorls, equals height. where and has of which they form numerous sharp regularly largest specimen about seven whorls, spaced, sickle-shaped riblets (curved backwards). 4% belong to the protoconch. This larval shell is
Below the carina they are curved in an anterior broadly conical with convex whorls separated by direction and, gradually becoming weaker, they deep sutures. The nucleus is small, slightly continue downwards in flat with a prosocyrt curve. heterostrophic and provided an extremely
Paratypes — Mogenstrup (coastal cliff): 4 juvenile fine microgranulation arranged in eight spirals. specimens (leg./coll. ISL); 1 juvenile specimen The last three whorls of the protoconch have a
(leg./coll. JL). 2 specimens (leg./coll. MNO); 2 cancellate ornament on the abapical two third of juvenile specimens (leg./coll. JHR). the whorl, while the abapical third of the whorl has
Kirstinebjerg Skov (beach exposure): 2 collabral riblets only. On the last protoconch
whorl carina the middleof the whorl specimens (leg./coll. ISL). a develops on
Discussion — The is and in front of the transition the new species obviously closely immediately to related Microdrilliaserratula but teleoconch rib both sides of the to (Bellardi, 1878) a spiral appears on differs features. M. has carina. On the last whorl by a number of ingerae a % numerous
of radial ribs the lines visible. The transi- higher number on protoconch. opisthocline growth are
The sutural band the tion the teleoconch is but more strongly developed, to sharp not very promi- carina is less sharp and the two spirals below the nent. The diagonal sculpture fades away and a carina rather weak. In M. serratula the carina fine the time are granulation appears at same is sharp and consists of one spiral only. More- (magnification x 50).
this has number of The has 2 3/4 teleoconch over, species a higher spiral largest specimen
base ribs on the and its general outline is more whorls which are very convex and separated by slender. deep, undulating sutures. Below the adapical
In Microdrillia the the bicingulata (Sandberger, 1860) suture whorls are slightly concave. The last two spirals below the carina are prominent and whorl comprises about three fifths of the total shell more widely spaced. Its radial sculpture is much height. The base is convex and abruptly con- weaker. M. teretiaeformis A.W. Janssen, 1972 has stricted into a rather short siphonal canal. The
carina of is rounded-oval. The labrum is broken a smooth consisting only one spiral. aperture
all and ‘Microdrillia’ speyeri (Koch & Wiechmann, 1872), on specimens the columella is smooth with
is also inner callus is show- which present at Mogenstrup, has a a well-defined lip. The thin, paucispiral protoconch. ing traces of resorption of calcareous matter. 69
The with three The of the whorls is excavated spiral sculpture starts primary concave part more
which of the carina and the and the is spirals are continuations on new species, finally granulation
3 whorl of the from the two spiral ribs on the last A proto- weaker than in P. mioweberi. Pleurotomella sp.
conch. The rib remains all the Late adapical spiral weak on German Oligocene (R. Janssen, 1979a:
whereas the middle the conti- differs the absence of whorls, one, being 329, pi. 18, fig. 77) by secon-
nuation of the carina, remains the strongest. The dary spirals and by its more angular whorls.
abapical spiral increases in strength but remains
weaker than the middle above one. Immediately Rimosodaphnella lappanni sp. nov.
the further weak rib PI. 13a-b abapical suture a spiral ap- 3, Fig.
pears on a quarter of a whorl. In the upper concave
of the whorl Locus — Badehotel part four narrow spirals are present typicus Fakkegrav (coastal cliff),
from the first teleoconch whorl onwards. Their North of Vejle Fjord, Jutland, Denmark
number increases to seven on the body whorl. In- Stratum typicum — Brejning Clay Member of the
between the of Late primary spirals a generation secon- Vejle Fjord Formation, Oligocene (Chattian
dary spirals develops from the second teleoconch B).
whorl The base and the canal Derivatio nominis — This is named after onwards. are provid- new species
ed with similar Mr. Werner a spiral ornament. my friend, Lappann, Heiligenhaus-
The radial sculpture consists of nine to ten slight- Isenbügel, F.R.G.
ly opisthocline ribs separated by about equally Holotype— PI. 3, Fig. 12a-b; (coll. MGUH 20 057)
wide interspaces. These ribs are almost invisible on (leg. M.S. Nielsen).
the of the whorls but — The shell is rather small and concave part prominent on Description
of intersection of 2.5 The the convex part. At the points fusiform, about times higher than wide.
and radial knobs form. last whorl three the spiral ornament elongated comprises fifths, aperture
The radial ribs gradually disappear on the base. slightly less than half the total shell height.
The lines in the The is and growth are very prominent con- protoconch high-conical multispiral,
of with the 3 -4 whorls cave part the whorls where together consisting of Zi moderately convex
form which The spiral ribs they a regular reticulate pattern. are separated by deep sutures. whorls
width is Downwards they decrease in strength. On the base have their greatest abapically. The nucleus
The while three they form a very shallow prosocyrt curve. en- small and apparently smooth, the last tire teleoconch has fine between the whorls with On a granulation are provided a diagonal sculpture.
elements. The sinus is rather shallow the last half whorl of the carina is sculptural protoconch a
and has its below the middle somewhat below the middle of the deepest point slightly present, lying of the whorl. last concave part of the whorl. On the % whorl before the transition to
— 1 the between Paratypes Mogenstrup (coastal cliff): specimen teleoconch a weak spiral rib develops
(leg./coll. ISL). the carina and the abapical suture. Immediately in
2 further weak Jensgard (coastal cliff): specimens (leg./coll. front of the transition a spiral occurs
1 above the carina. These three continue ISL); specimen (leg./coll. MNO). spirals as
Discussion — This is related to the the teleoconch. The transi- new species closely primary spirals on
P. mioweberi described and il- tion the teleoconch is demarcated (Nordsieck, 1972), as to sharp, by a lustrated by A.W. Janssen (1984: 325, pi. 74, fig. sigmoidal rib.
I have the The teleoconch consists of 3 3/4 whorls which 10; pi. 82, fig 5). compared new species with materialof the latter from Gram have and lower species (Late a concave adapical a convex part.
The whorls The Miocene, coll. ISL) and Miste (Middle Miocene; are separated by deep sutures. coll. A number of differences be but rather ISL). seem to con- base of the shell is convex, abruptly con-
The of the consists of stricted The stant. protoconch new species to a relatively long siphonal canal.
in callus is thin and the inner is 4 Yi whorls, whereas 5 such whorls are present lip not very sharply
P. mioweberi. The protoconch of P. rasmusseni is demarcated (which could indicate that the avail-
conical. The in the able not The more broadly sculpture concave specimens are yet fully-grown). aper-
whorl is is rather narrowed adapical part of the much more prominent ture oval, suddenly at the in the and the sinus of the lines the canal. The new species growth transition to siphonal labrum is
mioweberi. is situated more abapically than in P. broken. 70
with three The spiral sculpture starts spirals on continues as the second spiral. Furthermore, the
German has somewhat thick-set the convex part of the whorl which are continua- species a more tions of the carina and the conical two spirals on the pro- outline, a more broadly protoconch, a
of whorl rather toconch. On a quarter a a further, wider concave zone, a less sharp sculpture and is weak above the spiral rib appears immediately generally larger. abapical suture. The second spiral from above is
of second situated on the middle the whorl. The ? Actaeopyramis (s. lat.) and third are than the others. On spirals stronger pseudopunctata sp. nov.
teleoconch whorl the second secondary spirals are PI. 3, Figs 14-15 inserted between the and the primary ones, on
whorls further Locus — Water well younger two spiral generations are typicus at Krefeld-Linn, present. Ultimately, the spirals become nearly Nordrhein-Westfalen, F.R.G. equal in strength. The base and the canal are fur- Stratum typicum — Grafenberg Sands, Late Oligo-
A nished with a similar sculpture. All spirals are cène (Chattian or B).
threadlike and than their Derivatio nominis — The is named after narrower interspaces. species its
On the columella the visible below the distinctive = spirals are very sculpture (pseudopunctata falsely callus. On the concave upper part of the whorls punctate). no spirals are present. Holotype — PI. 3, Figs. 15 (coll. MGUH 20 059)
The radial is restricted the F. sculpture to convex (leg. von der Flocht).
of the whorls. It teleoconch — At six part starts on the first Description Mogenstrup juvenile speci- whorl with about ribs which collected. An adult from 14 opisthocline are mens were specimen about wide their The number of Krefeld-Linn is as as interspaces. designated holotype. ribs is constant all the whorls. On the base of The shell is small and rather on very thick-walled, the body whorl the radial sculpture gradually ovoid-conical. In juvenile specimens the height
downwards. At the of intersec- about times the width. In the disappears points equals 1.3 presum- tion of spiral and radial ornament small, elongate ably adult holotype the shell height is about 1.9 knobs the first teleo- times the width. The of the form, most prominently on height aperture equals conch whorl. On the of the about third of the total shell The adult concave adapical part one height. whorls the lines accentuated and form about four of growth are specimen comprises whorls, one
riblets with backward sickle-shaped a curvature. which belongs to the heterostrophic protoconch,
for- of which the terminal 3/4 whorl is On the convex part of the whorls they curve only visible, wards to opisthocline, in the same direction as the projecting only slightly above the teleoconch. The radial sculpture. An extremely fine granulation is transition to the teleoconch is sharp and slightly
between the elements all present sculptural over prosocline.
The three teleoconch whorls the teleoconch, especially well visible (magnifica- are slightly to tion x 50) on the concave part of the whorls. moderately convex and separated by deep, some-
Paratypes — Mogenstrup (coastal cliff): 15 speci- what canaliculate sutures. The body whorl equals mens (leg./coll. ISL); 13 specimens (leg./coll. about 6/10 of the total shell height. The base is
1 ISL/coll. 1 the umbilicus is rather The MNO); specimen (leg. WLH); convex, narrow. aper-
SMF is specimen (leg. ISL/coll. 308 411); 1 ture oval, with a regularly rounded lower
ISL/coll. RGH 229 The labrum is with specimen (leg. 791). margin. sharp-edged a
Skov transition into the columellar The Kirstinebjerg (beach exposure): 1 speci- gradual part.
columella is and bears men (leg./coll. MNO). concave a rather weak, ob-
Discussion — The differs new species from Rimoso- lique fold opposite the umbilicus.
nomina daphnella sp. [described and illustrated sub The spiral sculpture starts immediately after
Philbertia sinuosula Sorgenfrei, 1958 by Anderson the boundary with the protoconch and consists of
(1964: 312, pi. 42, figs 268, 268a) from the Mio- nine to ten ribbon-like spirals which are separated
of the of furrows. A weak subsutural is cene Dingden, F.R.G.] by development by narrower ramp the primary spirals. In the species described by demarcated by the adapical spiral band; on this
the carina the last whorl fine furrows visible. The Anderson, on protoconch ramp two very spiral are
the of develops into uppermost primary spiral the base of the shell has a similar spiral sculpture.
whereas in the this carina The radial teleoconch, new species sculpture consists of about 30 opis- 71
which weaker than the is wide much like the thocline, flat riblets are and open, very shaped con-
the side the The of the spiral bands. The combination of two cave apical of shell. base
elements results in nucleus is sculptural a pattern of almost visible in the umbilicus.
The consists of visible punctate appearance. The depressions are ellip- spiral sculpture barely
their diameter in the direc- furrows. On the side of the whorl fur- tical, having greatest upper one tion of the The lines fine row can be on the and on spirals. growth are very seen, periphery three, and be observed few the base two. The lines sometimes visi- can on a specimens only. growth are
orthocline and have ble the They are almost appear to a as sigmoidal lines, especially so on
flat sinus under the very adapical suture. periphery.
Paratypes — Mogenstrup (coastal cliff): 1 specimen
COMPARISON WITH OTHER LATE OLIGOCENE FAUNAS (PI. 3, Fig. 14) (coll. MGUH 20 058, leg. M.S. (AUTHOR: SCHNETLER) Nielsen); 4 specimens (leg./coll. MNO); 1 specimen (leg. M.S. Nielsen, coll. ISL). Danish Late Oligocene molluscan faunas were
Discussion — I refer the the Ravn Harder new species to genus published by (1907, 1909), (1913),
Eriksen and Schnetler Actaeopyramis (s. lat.) with some hesitation, since Madsen (1918), (1937) & the sculpture is somewhat untypical. In Actaeopyra- Beyer (1987). These faunas are characterised by a mis smooth and and in of like the spirals are projecting, high frequency species Drepanocheilus (Arrho- the the radial lamellae rise Polinices helicinus and Fusiturris interspaces give to a ges) speciosus,
the lines punctation. Moreover, growth are pro- socline. The outline of the general new species cor-
well to the so do the responds genus Actaeopyramis ,
the and the but it shape of protoconch aperture,
resembles the referred closely species to as
Pyramidellidae indet. and described from Norre
in: Schnetler 1987: Vissing (Schnetler, & Beyer,
this form the 220, pi. 2, figs 16-17). From new
differs its weaker subsutural less species by ramp,
whorls and less convex a regular punctate sculp- ture. In all other features the two species appear closely related. They should probably be united in
Introduction of such is a new genus. a new genus deferred to another occasion.
Gastropoda inc. sed.
Text-Fig. 7
Description — Only a single (? juvenile) specimen is available (leg./coll. M.S. Nielsen). The shell is
small and 2.8 times wider than very discoidal, high. The protoconch consists of 1 % whorls which are separated by a distinct suture. The nucleus is small. The transition to the teleoconch is sharp, demarcated flexuous rib. by a distinct, slightly
The teleoconch consists of somewhat more than
1 % whorls, slowly expanding in diameter and cir-
distinct cular in transverse section, separated by a
The of the whorls lies suture. upper junction on the periphery of the preceding whorl, the lower
which in one touches below the periphery, results Fig. 7. Gastropoda inc. sed. , Mogenstrup; a: apical view, concave and lower sides of the shell. The upper b: frontal umbilical view, c: view, d: lateral view. transition into the base is The is gradual. aperture Coll. MGUH 20 275 (A.W. Janssen del., 1990; subcircular, the labrum is broken. The umbilicus bar length is 1 mm. 72
duchastelii. Other very common species are Nucula stinebjerg Skov, Jensgârd and Skanderborg
aff. kickxi and comta, Limopsis aurita, Cyclocardia (glacial flow)], or even absent from other Danish
Gemmula spp. In comparison to the Arhus fauna Late Oligocene localities (e.g. Arhus, Cilleborg
the absent Norre these (Harder, 1913) following species are at and Vissing). Thus, rare species point
Mogenstrup: out certain affinities with the faunas from the Vejle
rather Fjord area, than with the faunas from
Nucula Norre harderi Schnetler, in Schnetler & Arhus, Cilleborg and Vissing.
Beyer, 1987 When compared to the German Late Oligocene
Emarginula punctulata Philippi, 1843 faunas the Mogenstrup association shows affinities
Unitas with granulata (Nyst, 1845) those from the Lower Rhine area, Glimme-
Conomitra ravni (Harder, 1913) rode, Freden and the Sternberger Gestein. Asso-
Microdrillia bicingulata (Sandberger, 1860). ciations described from Sollingen, Niederkau-
Ahnetal and fungen, Doberg, however, appear to
In comparison to the recently described fauna be different.
from Norre the The fauna from Vissing (Schnetler & Beyer, 1987) the Lower Rhine area has 110
characteristic absent: in with following species are species common Mogenstrup, among
which are the following characteristic species:
Nucula harderi Schnetler, in Schnetler &
Beyer, 1987 Limopsis aurita
Solariella spp. Limopsis vonderhochti
Unitas granulata Lepetella compressiuscula
Pleurotomoides naumanni (Speyer, 1867). Cirsope multicingulata
Cirsotrema crispatum
The faunas from Norre Vissing and Mogens- Conomitra soellingensis
have 97 in but trup species common, frequencies ‘Microdrillia’ speyeri
of common species differ. A fairly large number of
found at is absent at Norre E. species Mogenstrup Moreover, Emarginula spp. (except punctulata
absent from both faunas. Ob- Vissing, e.g. : Philippi, 1843) are
vious differences the absence of Astraea are spp.
Limopsis lamellata chattica and the lower diversity of the Cerithiopsidae in the
vonderhochti Lower Limopsis Rhine area.
Acar aff. dentiens The fauna from Glimmerode has 105 species in
Pecten with the bifidus bifidus common Mogenstrup association, among
Pecten soellingensis which are:
Astraea pustulosa
Cocculina dittmeri Acar aff. dentiens
Astraea Cirsope subeffusa spp.
Triforis sorgenfreii Scissurella koeneniana
Neojanacus planatus Gibberula brevis
Trophonopsis angustevaricata Cerithiopsidae
Pterynotus tristicus Muricidae.
Coralliophila kochi
the Gibberula brevis At Glimmerode following species which are Microdrillia spp. common at Mogenstrup, are missing:
Pleurotomella margaritata
Pleurotomella rasmusseni Limopsis aurita
Rimosodaphnella lappanni Limopsis vonderhochti.
pteropods.
Additionally, the Mogenstrup fauna did not
of these are in the Many species extremely rare yield any Amphineura or Emarginula spp.
of localities in the The fauna 100 Brejning Clay Vejle Fjord area from Freden has species in com-
Kir- with [Brejning, Fakkegrav Badehotel, Bogeskov, mon Mogenstrup, among which are: 73
Limopsis aurita Vexillum hastatum
Astraea pustulosa Gemmula trispiralis
Turbonilla Cirsope subeffusa. spp.
The following species from Mogenstrup are Emarginula spp. are present in the Söllingen
missing at Freden: fauna.
The fauna has 68 in Doberg species common
vonderhochti with Limopsis Mogenstrup: e.g. :
almost all Cerithiopsidae
‘Microdrillia’ speyeri. Acar aff. dentiens
Astraea spp.
An obvious difference is the presence of
fauna. Emarginula spp. in the Freden The following species, common at Mogenstrup,
fauna from the 'Kasseler Meeres- absent from the fauna: The typical are Doberg
sand' (Niederkaufungen and Ahnetal) has 80
in with aurita species common Mogenstrup, among Limopsis
which are: Limopsis vonderhochti
Rissoa spp.
Exilioidea elatior many Cerithiopsidae
Metula scabricula Muricidae, except Lyrotyphis cuniculosus
Streptochetus soellingensis (Nyst, 1836)
Conomitra soellingensis. Vexillum hastatum
Conomitra soellingensis
Emarginula spp. are absent from both faunas. In Gemmula trispiralis
Gemmula comparison to the Mogenstrup fauna, a large pseudokonincki.
number of species are absent, e.g. :
The Doberg fauna comprises a large number of
Acar aff. dentiens bivalve species, especially Pectinidae. Emarginula
aurita also Limopsis spp. are present.
Palliolum hausmanni hausmanni The 'Sternberger Gestein' fauna has 100 species
Astraea in with As pustulosa common Mogenstrup. this fauna origi-
nates from boulders of different Cirsope spp. ages (Chattian A,
Rissoa B and further made. spp. C) analyses were not
almost all Cerithiopsidae.
PALAEOECOLOGICAL INTERPRETATIONS
from has 75 in (AUTHOR: SCHNETLER) The fauna Sollingen species com-
with which In unit 1 molluscs mon Mogenstrup, among are: are generally not very common
and regularly distributed in the sediment. In the
Acar aff. dentiens uppermost part of unit 1 molluscs and other fossils
Scissurella koeneniana The molluscan is are abundant. assemblage most
Astraea the fossils pustulosa probably not a biocoenosis, as are con-
Pterynotus tristrichus. centrated in a thin horizon which has a sharp
lower boundary; Fusiturris duchastelii, Gemmula
the that do Polinices helicinus and cheruscus Among many Mogenstrup species spp., Streptochetus are
the be found almost not occur at Sollingen following may men- exclusively as juvenile specimens.
tioned: of molluscs both and However, many juvenile
the adult specimens are found, e.g. gastropods
Limopsis aurita Astrea pustulosa (dominant), Lyrotyphis sejunctus,
Limopsis vonderhochti Hiniaschlotheimi, Charonia flandrica and the bivalves
Rissoa spp. Limopsis spp.
Triphoridae The abundant occurrence of Astraea pustulosa is
Ancilla karsteni very conspicuous. According to R. Janssen 74
the Astraea littoral con- The Charonia the (1978a: 161) genus suggests gastropod species flandrica, on
of would indicate and ditions. This is also suggested by the occurrence other hand, deeper water the
Gibberula brevis and of Cocculina dittmeri indicate Alvania semperi, Cirsope spp., occurrence might even
The rather The Cocculina and related Vexillum hastatum. common species Fusiturris deep water. genus duchastelii and helicinus be rather forms live wood Polinices seem to on waterlogged (Marshall, 1985)
below 150 with ‘Teredo’ facies independent since they are common in all at depths m together sp.,
and since ‘Teredo’ is rather facies. sp. common at
Mogenstrup this might indicate deeper water.
Table 3. of bivalve and Frequencies gastropod species. Very remarkable is the high diversity of Murici-
dae and Cerithiopsidae, since representatives of number of these other Danish Late groups are very rare at specimens % Norre Arhus and Oligocene localities, e.g. Vissing, Bivalvia We Skanderborg. may assume that, generally 1. Limopsis lamellata chattica 155155 24.8 speaking, the gastropod assemblage indicates lit- 2. Limopsis retifera 135 21.6
toral to sublittoral conditions. 3.3. Limopsis aurita 62 9.99.9 higher 4.4. Cyclocardia aff.aff. kickxi 48 7.7 The bivalve assemblage is dominated by Limop-
5. Limopsis vonderhochtivonderhochli 4111 6.5 sis of this make of spp.: species genus up 63.9% 6.6. AstarteAstarle goldfussi praecursor 17 2.7 the bivalve total number of specimens. Limopsis is 7. Teredinidae indet. 14 2.2 gen. etet sp. epibenthic and lives in a sublittoral environment. 8.8. Portlandia pygmaea 11 1.8
L. is often found at 9.9. Acar aff. dentiens 11 1.8 retifera very Mogenstrup ar-
ticulated low 10. BathyarcaBalhyarca bellula 10 1.6 specimens, indicating current
velocities and hardly The species Gastropoda any transport. Nucula Nuculana and Yoldia 1. Collonia troelsi 229 12.7 spp., westendorpi glaber-
2.2. Astraea pustulosapustulosa 201 11.2 rima are rather rare at Mogenstrup, but very com-
3.3. Fusiturris duchasteliiduchastelii 200 11.1 in the sediments of the Danish Late mon clayey
4. Polinices helicinus 131131 7.3 the localities Norre Oligocene, e.g. Àrhus, Vissing 5.5. Streptochetus cheruscus 127 7.1 and These endoben- Skanderborg. specimens are 6. Lyrotyphis sejunctus 68 3.83.8 thic. The endobenthic Astarte 7.1. Drepanocheilus speciosus 63 3.53.5 equally species
and aff. kickxi 8. Stenodrillia obeliscus 53 2.92.9 goldfussi praecursor Cyclocardia are
9. Gemmula trispiralistrispiralis 46 2.62.6 rather is the common. Very interesting relatively 10.10. Orthosurcula regularisreguláris 40 2.22.2 common occurrence of Acar aff. dentiens, which is
Table 4. UTM coordinates of Danish localities mentioned in this paper
No. Kind of Coordinates Locality outcrop Mapsheet
1. Mogenstrup coastal cliff 1216 III SV NH 074 814
2. Cilleborg former clay-pit 1216 II S0 NH 550 784
3. Hadsten former road-excavation 1315 III NV NH 650 442
4. Arhus former railway-cut 1314 I VN0 NH 745 235
5. Norre Vissing working clay-pit 1214 I N0 NH 525 212
6. Skanderborg former motorway-excavation 1214 I S0 NH 573 120
7. Jensgârd coastal cliff 1313 I NV NG 664 851
8. Fakkegrav Badehotel coastal cliff 1213 I S0 NG 490 711
9. beach 1213 I SV NG 439 696 Brejning exposure
10. Bogeskov coastal cliff 1213 I S0 NH 480 646
11. Skov beach 1213 II N0 NG 505 616 Kirstinebjerg exposure
also common in the near-shore sediments at Glim- since remains of these teresting, groups are very
bivalve merode. In all, the assemblage indicates rare at other Danish Late Oligocene localities (for higher sublittoral conditions. instance Norre Vissing, Arhus and Skanderborg).
The abundance of and other remains of The ‘Terebratula’ is Ser- spines brachiopod sp. common. the sea-urchin ‘Cidaris’ of are also but balanids absent. regular sp., marginal pulids common, are
of asteroids and of Anthozoa is in- Teeth of some shark were found: plates very species Pristiopho- 75
acutissima and Mr K. is rus sp., Synodontaspis (Agassiz, 1843) Ulleberg (Galleberg, Norway) kindly
indet. der thanked for information on the Foraminifera Odontaspididae gen. et sp. (F. von
zonation. Hocht, pers. comm., 1987). These species might
but fauna We Dr indicate deeper water, as the shark is furthermore like to thank C. Heilmann-
limited and the of Clausen of very at Mogenstrup state preser- (Geological Institute, University
vation is rather the teeth for critical remarks poor are possibly Arhus, Denmark) stimulating
reworked. and helpful advice.
otoliths collected Mr S.L. and Mrs M. A number of teleost were (coll. Jakobsen Bjerreskov
This material examined Dr of ISL, AJB). was by (Geological Museum, University Copenhagen,
with the P.A.M. Gaemers (Leiden), who identified 17 spe- Denmark) were very helpful regard to
cies, of which Gadichtys attenuatus (Koken, 1891) is MGUH collections. Mr O.B. Berthelsen (Geo-
the Dr is thanked for most common. According to Gaemers logical Institute, Copenhagen)
the otolith fauna indi- of the work. (pers. comm., 1987, 1989) doing part photographic
The cates rather deep-water (100-200 m). The two MrJ.W.M. Jagt (Venlo, Netherlands), co-
and editor of this corrected the deep water genera Diaphus (lantern-fishes) periodical, kindly
Coelorinchus (grenadier-fishes) are represented by a English text.
assistance of Mr S.B. Andersen few specimens and all other species, except for Col- The substantial
liolus indicate rather water. The state of Institute, of Arhus, Den- spp., deep (Geological University
indicates this in and the preservation that material is not re- mark) critically reading correcting
is thus indication of material worked and a good deposition manuscript, providing and, especially,
for the is in a sublittoral environment. doing photographic work, gratefully
acknowledged.
ACKNOWLEDGEMENTS REFERENCES
The authors the are very grateful to following per- Anderson, H.J., 1964. Die miocane Reinbek-Stufe in Nord- for molluscan material in sons (information on) und Westdeutschland und ihre Molluskenfauna. —
Bo Andersen their collections: Mr Sten (Frederi- Fortschr. Geol. Rheinld. u. Westf., 14: 31-368.
1987 0. Miocaen i Nordvest cia, Denmark); Mr H.C. Hansen (Fredericia, Beyer, C., . Oligocasn-N. Jylland. Faciesanalyse and magnetostratigrafi. Arhus Denmark); Mrs J. Hillersborg (Randers, Den- of 165 (thesis University Arhus), pp. (unpublished). Mr A.C. (Brielle, The Netherlands); mark); Janse Beyrich, E., 1856. Die Conchylien des norddeutschen Ter- Mr F. der Hocht von (Kerpen-Balkhausen, tiargebirges, 4/5. Berlin (Hertz): 177-296, pis 16-25. shark F.R.G.), who also supplied information on Cossmann, M. 1889. Catalogue illustre des coquilles de
l'Eocene des environs de Paris. — Ann. Soc. r. Malacol. material; Mr A.W. Janssen (National Museum of Belg., 24: 3-381, 12 pis. Natural History, Leiden, The Netherlands), who Eriksen, K., 1937. En forekebig meddelelse om Tertiaeret ved also kindly examined some problematic molluscs, Brejning pa sydsiden af Veile Fiord. — Bull. geol. Soc.
6 and an earlier prepared text-Figs 7, improved Denmark, 9(2): 137-150.
and studied the Mo- Die Rumeln version of the manuscript Gorges, J., 1941. Oberoligocanfauna von am
for Dr R. Niederrhein. — Decheniana, 100(A): 115-186, 3 pis. genstrup pteropods a separate paper; Gorges, J., 1952. Die Lamellibranchiaten und Gastropoden Janssen (Senckenberg Museum, Frankfurt am
des Oberoligocanen Meeressandes von Kassel. — Main, who also examined some F.R.G.), kindly Abhandl. hess. L.-Amt 4: 134 3 Bodenforsch., pp, pis.
molluscs; Mrs L.B. (Lime, Uberfamilie problematic Jorgensen Griindel, J., 1980. Bemerkungen zur Cerithiop-
Mr W. H.A. 1854 sowie Denmark); Lappann (Heiligenhaus-Isen- sacea Adams, (Gastropoda) zur Fassung
ihrer — Zool. biigel, F.R.G.); Mrs R. Lechner (Odense, Den- einiger Gattungen. Anz., lena, 204(3-4): 209-264, 41 figs. mark); Dr P. Lozouet (Musee national d'Histoire Haq, B.U., J. Hardenbol & P R. Vail, 1987. Chronology of naturelle, Paris, France); Mrs E. Nielsen (Arhus,
levels since the Triassic. — fluctuating sea Science, 235: and Mr M.S. Nielsen Den- Denmark) (Odense, 1156-1167.
mark), who kindly placed his important collection Harder, P., 1913. De oligocaene Lag i Jaernbanegennem-
skaeringen ved Aarhus Station. — Danm. Geol. Unders., from Mogenstrup at my disposal and in this way (2)22: 140 5 tabs, 4 figs, 9 pis. contributed this pp., substantially to paper. Win- Janssen, A.W., 1984. Mollusken uit het Mioceen van Dr P.A.M. Gaemers (National Museum of terswijk-Miste. Een inventarisatie, met beschrijvingen en
Natural Leiden, The sup- History, Netherlands) afbeeldingen van alle aangetroffen soorten. Amsterdam
information otolith material. 451 82 plied on (K.N.N.V., N.G.V., R.G.M.), pp., pis. 76
0verste Janssen, A.W., 1990. Pteropod species (Mollusca, Gastro- Lieberkind,K., 1977. Ostracodstratigrafi i Eocaen og
1 from the Late of Mo- i Danmark - ud fra poda, Euthecosomata) Oligocene Oligocaen belyst Viborg boringen og
— enkelte Arhus of genstrup, Jylland, Denmark. Contr. Tert. Quatern. daglokaliteter. (thesis University Arhus),
10 in this 112 23 Geol., 27(2-3): 83-92, figs, 1 pi. (printed issue). pp, pis. (unpublished)
Janssen, R., 1978a. Die Scaphopoden und Gastropoden des Madsen, V., 1918. Om Tertieeret ved Mariager Fjord. —
Kasseler Meeressande Glimmerode Danm. Unders. 41 von (Niederhessen). geol. (4)1(8): pp.
— Geol. Jb., (A)4: 3-195, 3 figs, 3 tabs, 7 pis. Marshall, B.A., 1986. Recent and Tertiary Cocculinidae and
Pseudococculinidae from Janssen, R., 1978b. Die Mollusken des Oberoligozans (Chat- (Mollusca: Gastropoda) New
tium) in Nordseebecken, 1. Scaphopoda, Archaeogastro- Zealand and New South Wales. — New Zealand Journal
poda, Mesogastropoda. — Arch. Moll., 109(1/3): of Zoology, 12(1985): 505-546, 24 tabs, 14 figs.
137-227, 1 fig., pis 9-14. Peyrot, A., 1932. Conchologie neogenique de l'Aquitaine, 6.
Janssen, R., 1979a. Die Mollusken des Oberoligozans (Chat- — Actes Soc. linn. Bordeaux, 84: 5-128, 10 pis.
tium) im Nordseebecken, 2. Neogastropoda, Euthyneura, Rasmussen, L.B., 1961. De miocasne formationer i Dan-
— Arch. 5 mark. — Danm. 45 Cephalopoda. Moll., 109(4/6): 277-376, pis. geol. Unders., 4(5), pp.
Janssen, R. 1979b. Revision der Bivalvia des Oberoligozans Rasmussen, L.B., 1968. Molluscan faunas and biostratigra-
Kasseler — Geol. Abhandl. of the marine Miocene formations in Den- (Chattium, Meeressand). phy younger
Hessen, 78: 1-181, 1 fig., 4 pis. mark, 2. Palaeontology. — Danm. geol. Unders., 2(92):
K. A. 1912. Die staatlichen Tief- 265 27 Koch, E., Gripp & Franke, pp., pis.
bohrungen XIV, XV, XVI, XVII in den Vierlanden bei Ravn, J.P.J., 1906. Nogle bemaerkninger om de Oligocsene
Hamb. wiss. Anst. 4. Beih.: 33 Miocsene i — Bull. Geol. Soc. Den- Hamburg. —Jb. 29, pp., og aflejringer Jylland.
8 figs, 1 pi. mark, 12: 1-6.
KristofTersen, F. N., 1972. Miocaen i Ravn, 1907. Molluskfaunaen i Oligocaen og Novling, J.P.J., Jyllands Tertiaer-aflej-
1. — Danm. geol. Unders., 3 (40): 63-70. ringer. — Kgl- Danske Vid. Selskab Skr., 7 (nat. mat.)
180 8 Larsen, G., & Dinesen, A., 1959. Vejle Fjord Formationen 3(2): pp., pis.
ved Foraminiferfaunaen - 1909. Om findesteder for Tertiasrforstenin- Brejning. Sedimenterne, (Olig Ravn, J.P.J., nye
— Danm. 114 i Danmark. - Bull. Soc. 3: 331-336. Mioc.). geol. Unders., (2)82, pp. ger geol. Denmark,
PLATE 1
Fig. 1. Acar aff. dentiens (Cossmann & Peyrot, 1912); a: left valve, interior; b: left valve, exterior; x 10. Coll. MGUH 20
022.
left valve, interior; b: left valve, exterior; x Fig. 2. Limopsis (Pectunculina) lamellata chattica subsp. nov., holotype; a: 9.5.
Coll. MGUH 20 023.
b: Fig. 3. Limopsis (Pectunculina) lamellata chattica subsp. nov., paratype; a: right valve, interior; right valve, exterior; x
9.5. Coll. MGUH 20 024.
4. vonderhochti b: 3. Coll. Fig. Limopsis (Pectunculina) sp. nov., holotype; a: right valve, interior; right valve, exterior; x
MGUH 20 025.
5. vonderhochti b: left 3. Coll. Fig. Limopsis (Pectunculina) sp. nov., paratype; a: left valve, interior; valve, exterior; x
MGUH 20 026.
6. vonderhochti left b: left 3. Coll. SMF Fig. Limopsis (Pectunculina) sp. nov., paratype; a: valve, interior; valve, exterior; x
309 204.
vonderhochti b: 3. Coll. Fig. 7. Limopsis (Pectunculina) sp. nov., paratype; a: right valve, interior; right valve, exterior; x
SMF 308 407.
MGUH Fig. 8. Cocculina dittmeri (Anderson, 1964); apical view; x 5. Coll. 20 027.
9. a: lateral b: x 20. Coll. MGUH 20 028. Fig. Lepetella helgae sp. nov., holotype; view; apical view;
10. lateral b: 2. Coll. MGUH 20 029. Fig. Lepetella jytteae sp. nov., holotype; a: view; apical view; x
Fig. 11. Astraea (Lithopoma) pustulosa (von Münster, 1844); apertural view; x 10. Coll. MGUH 20.030.
Fig. 12. Homalopoma (Boutillieria) simplex (Philippi, 1843); apertural view; x 10. Coll. MGUH 20 031.
13. b: umbilical 5. Coll. ENÅ. Fig. Homalopoma (? Leptothyropsis) sp.; a: apertural view; view; c: apical view; x
14. Tubiola 30. Coll. MGUH 20 032. Fig. subangulata sp. nov., holotype; apertural view; x
Fig. 6 from Krefeld-Linn, F.R.G. (water well). Fig. 7 from Ahnetal, F.R.G. All other specimens from
Mogenstrup (coastal cliff), Denmark. 77\1
PLATE 1 78
P. 1924. Tertiaer. In Danmarks Koenen. — Abhandl. Ravn, J. J. : Oversigt over Tafel-Erklarungenvon A. von geol.
geologi (ed. V. Madsen) — Danm. geol. Unders., 5(4), Spez.-Karte Preussen, 4(4): i-xii, 31 pis.
208 2 1963. Muriciden dem Mittel- und pp., pis. Tembrock, M.-L., aus
Schneder, K.I., & C. Beyer, 1987. A Late Oligocene (Chat- Oberoligocan und den Vierlandschichten des Nord-
tian B) mollusc fauna from the clay-pit of Galten Brick- seebeckens. — Palaont. Abhandl., 1: 299-351, 3 tabs, 10
works at Norre Vissing, Jylland, Denmark. — Meded. pis.
Werkgr. Tert. Kwart. Geol., 24(3): 193-224, 3 figs, 1 Ulleberg, K., 1987. Foraminiferal zonation of the Danish
tab., 2 pis. Oligocene sediments. — Bull. geol. Soc. Denmark, 36:
Sorgenfrei, T., 1940. Marint Nedre-Miocaen i Klintingeho- 191-202.
ved Als. — Danm. 143 8 1938-1944. Teil und paa geol. Unders., 2(65), pp., Wenz, W., Gastropoda. Allgemeiner
pis. Prosobranchia. — Handbuch der Palaozoologie, 6: xii +
1958. Molluscan from the 1639 4211 Sorgenfrei, T., assemblages pp., figs.
marine Middle Miocene of South Jutland and their en-
1-2. — Danm. Unders. 79: 503 vironments, geol. (2) pp.,
76 pis.
Speyer, O., & A. von Koenen, 1884. Die Bivalven der Manuscript received 23 October 1989, revised version ac-
Casseler Tertiar-Bildungen, mit einem Vorwort und cepted 3 August 1990.
PLATE 2
1. Tubiola 30. Coll. MGUH 20 033. Fig. sp.; apertural view; x
2. Collonia troelsi b: umbilical Coll. Fig. (Collonia) sp. nov., holotype; a: apertural view; view; c: apical view; x 20.
MGUH 20 034.
3.? b: umbilical 30. Coll. MGUH 035. Fig. Skenea sp.; a: apical view; view; x 20
Coll. Fig. 4. Neojanacusplanatus (Speyer, 1864); apertural view; x 3. LJL.
5. 30. Coll. MGUH 20 036. Fig. Triforis (Trituba) sorgenfreii sp. nov., holotype; apertural view; x
6. 11. Coll. MGUH 20 037. Fig. Triforis (Trituba) sorgenfreii sp. nov., paratype; apertural view; x
7. 4. Coll. MGUH 20 038. Fig. Triforis (Trituba) sorgenfreii sp. nov., paratype; apertural view; x
8. Laiocochlis 20. Coll. MGUH 20 039. Fig. (Laiocochlis) supraoligocaenica sp. nov., holotype; apertural view; x
9. x 10. Coll. MGUH 20 040. Fig. Cerithiopsis (s. lat.) antonjansei sp. nov., holotype; apertural view;
10. Cirsotrema 10. Coll. MGUH 20 041. Fig. (Opaliopsis) subglabrum sp. nov., paratype; apertural view; x
Cirsotrema aff. koeneni Fig. 11. (? Opaliopsis) A.W. Janssen, 1967; apertural view, x 20. Coll. JHR.
12. Cirsotrema 6. Coll. MGUH 20 042. Fig. (Opaliopsis) sp. 1; apertural view; x
13. Cirsotrema 30. Coll. MGUH 20 043. Fig. (Opaliopsis) sp. 2; apertural view; x
14. Coll. Fig. Trophonopsis (Pagodula) angustevaricata (Gripp, 1912); a: apertural view; b: dorsal view; x 5. MGUH 20 044.
Fig. 15. Coralliophila (Hirtomurex) kochi (Beyrich, 1854); a: apertural view; b: dorsal view; x 5. Coll. MGUH 20 045.
16. Searlesia ravni dorsal 1. Coll. MGUH 20 046. Fig. sp. nov., paratype; view; x
16. Searlesia ravni lateral 1. Coll. MGUH 20 047. Fig. sp. nov., paratype; a apertural view; b: view; x
Fig. 7 from Kirstinebjerg Skov (beach exposure), Denmark. All other specimens from Mogenstrup (coastal cliff),
Denmark. 79\2
PLATE 2 80
PLATE 3
1. 9. Coll. MGUH 20 048. Fig. Scalaspira (Scalaspira) sp.; apertural view; x
2. Latirus view; x 11. Coll. Fig. (Pseudolatirus) sp.; apertural JHR.
Fig. 3. Gibberula brevis (von Koenen, 1890); apertural view; x 8. Coll. MGUH 20 049.
x Coll. MGUH Fig. 4. Apiocypraea (Apiocypraea) cf. humbergi R. Janssen, 1978; a: apertural view; b: dorsal view; 10. 20
050.
5. Clavatula 11. Coll. MGUH 20 051. Fig. mogenstrupensis sp. nov., holotype; apertural view; x
Fig. 6. Babylonella pusilla (Philippi, 1843); apertural view; x 8. Coll. MGUH 20 052.
7. Microdrillia b: dorsal view; x 8.5. Coll. MGUH 20 053. Fig. ingerae sp. nov., holotype; a: apertural view;
Fig. 8. Microdrillia’ speyeri (Koch & Wiechmann, 1872); ‘apertural view; x 8. Coll. LJL.
9. Microdrillia b: dorsal 8. Coll. MGUH Fig. (Andersondrillia) brejningensis sp. nov., holotype; a: apertural view; view; x
20 054.
10. Microdrillia view; x 8. Coll. MGUH 20 055. Fig. (Andersondrillia) brejningensis sp. nov., paratype; apertural
11. Pleurotomella rasmusseni 8 dorsal 8. Coll. Fig. (Pleurotomella) sp. nov., holotype; a: apertural view, x ½; b: view, x
MGUH 20 056.
Coll. Fig.R.Janssen,12. Pleurotomella (Pleurotomella) margaritata 1978; apertural view; x 8. JHR.
13. b: dorsal 5. Coll. MGUH 20 057. Fig. Rimosodaphnella lappanni sp. nov., holotype; a: apertural view; view; x
14.? 34. Coll. MGUH 20 058. Fig. Actaeopyramis (s. lat.) pseudopunctata sp. nov., paratype; apertural view; x
15.? view, 23. Coll. MGUH 20 059. Fig. Actaeopyramis (s. lat.) pseudopunctata sp. nov., holotype; apertural x
Fig. 9 from Brejning (beach exposure), Denmark; Fig. 13 from Fakkegrav Badehotel (coastal cliff), Denmark;
Fig. 15 from Krefeld-Linn (water well), F.R.G. All other specimens from Mogenstrup (coastal cliff), Denmark. 81\3
PLATE 3