Chapter 2. Human Evolution
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The Place of the Neanderthals in Hominin Phylogeny ⇑ Suzanna White, John A.J
Journal of Anthropological Archaeology 35 (2014) 32–50 Contents lists available at ScienceDirect Journal of Anthropological Archaeology journal homepage: www.elsevier.com/locate/jaa The place of the Neanderthals in hominin phylogeny ⇑ Suzanna White, John A.J. Gowlett, Matt Grove School of Archaeology, Classics and Egyptology, University of Liverpool, William Hartley Building, Brownlow Street, Liverpool L69 3GS, UK article info abstract Article history: Debate over the taxonomic status of the Neanderthals has been incessant since the initial discovery of the Received 8 May 2013 type specimens, with some arguing they should be included within our species (i.e. Homo sapiens nean- Revision received 21 March 2014 derthalensis) and others believing them to be different enough to constitute their own species (Homo Available online 13 May 2014 neanderthalensis). This synthesis addresses the process of speciation as well as incorporating information on the differences between species and subspecies, and the criteria used for discriminating between the Keywords: two. It also analyses the evidence for Neanderthal–AMH hybrids, and their relevance to the species Neanderthals debate, before discussing morphological and genetic evidence relevant to the Neanderthal taxonomic Taxonomy debate. The main conclusion is that Neanderthals fulfil all major requirements for species status. The Species status extent of interbreeding between the two populations is still highly debated, and is irrelevant to the issue at hand, as the Biological Species Concept allows -
Hominin Occupation of the Chinese Loess Plateau Since About 2.1
LETTER https://doi.org/10.1038/s41586-018-0299-4 Hominin occupation of the Chinese Loess Plateau since about 2.1 million years ago Zhaoyu Zhu1,2*, Robin Dennell3*, Weiwen Huang2,4, Yi Wu5, Shifan Qiu6, Shixia Yang4,7, Zhiguo Rao8, Yamei Hou2,4, Jiubing Xie9, Jiangwei Han10 & Tingping Ouyang1,11 Considerable attention has been paid to dating the earliest recording of polarity reversals in loess20, we date each unit to within a appearance of hominins outside Africa. The earliest skeletal and precessional cycle rather than to the 1-thousand-year precision of the artefactual evidence for the genus Homo in Asia currently comes Chiloparts timescale. from Dmanisi, Georgia, and is dated to approximately 1.77–1.85 The main palaeoanthropological limitation of the Loess Plateau is million years ago (Ma)1. Two incisors that may belong to Homo that it largely lacks evidence of the Early Palaeolithic, or associated erectus come from Yuanmou, south China, and are dated to 1.7 Ma2; hominin remains. The main exception is Lantian county in which a the next-oldest evidence is an H. erectus cranium from Lantian mandible and cranium of H. erectus have been found at Chenjiawo (Gongwangling)—which has recently been dated to 1.63 Ma3— and Gongwangling, respectively. Both were dated by palaeomagne- and the earliest hominin fossils from the Sangiran dome in Java, tism and correlation with the Luochuan loess–palaeosol sequence21. which are dated to about 1.5–1.6 Ma4. Artefacts from Majuangou At Chenjiawo the mandible occurs in palaeosol S6 (684–710 thousand III5 and Shangshazui6 in the Nihewan basin, north China, have years ago), and at Gongwangling the layer containing the cranium is also been dated to 1.6–1.7 Ma. -
Homo Habilis
COMMENT SUSTAINABILITY Citizens and POLICY End the bureaucracy THEATRE Shakespeare’s ENVIRONMENT James Lovelock businesses must track that is holding back science world was steeped in on surprisingly optimistic governments’ progress p.33 in India p.36 practical discovery p.39 form p.41 The foot of the apeman that palaeo ‘handy man’, anthropologists had been Homo habilis. recovering in southern Africa since the 1920s. This, the thinking went, was replaced by the taller, larger-brained Homo erectus from Asia, which spread to Europe and evolved into Nean derthals, which evolved into Homo sapiens. But what lay between the australopiths and H. erectus, the first known human? BETTING ON AFRICA Until the 1960s, H. erectus had been found only in Asia. But when primitive stone-chop LIBRARY PICTURE EVANS MUSEUM/MARY HISTORY NATURAL ping tools were uncovered at Olduvai Gorge in Tanzania, Leakey became convinced that this is where he would find the earliest stone- tool makers, who he assumed would belong to our genus. Maybe, like the australopiths, our human ancestors also originated in Africa. In 1931, Leakey began intensive prospect ing and excavation at Olduvai Gorge, 33 years before he announced the new human species. Now tourists travel to Olduvai on paved roads in air-conditioned buses; in the 1930s in the rainy season, the journey from Nairobi could take weeks. The ravines at Olduvai offered unparalleled access to ancient strata, but field work was no picnic in the park. Water was often scarce. Leakey and his team had to learn to share Olduvai with all of the wild animals that lived there, lions included. -
Archaeologist Volume 44 No
OHIO ARCHAEOLOGIST VOLUME 44 NO. 1 WINTER 1994 Published by THE ARCHAEOLOGICAL SOCIETY OF OHIO The Archaeological Society of Ohio MEMBERSHIP AND DUES Annual dues to the Archaeological Society of Ohio are payable on the first of January as follows: Regular membership $17.50; husband and wife (one copy of publication) $18.50; Life membership $300.00. EXPIRES A.S.O. OFFICERS Subscription to the Ohio Archaeologist, published quarterly, is included in 1994 President Larry L. Morris, 901 Evening Star Avenue SE, East the membership dues. The Archaeological Society of Ohio is an incor Canton, OH 44730, (216) 488-1640 porated non-profit organization. 1994 Vice President Stephen J. Parker, 1859 Frank Drive, BACK ISSUES Lancaster, OH 43130, (614) 653-6642 1994 Exec. Sect. Donald A. Casto, 138 Ann Court, Lancaster, OH Publications and back issues of the Ohio Archaeologist: 43130, (614)653-9477 Ohio Flint Types, by Robert N. Converse $10.00 add $1.50 P-H 1994 Recording Sect. Nancy E. Morris, 901 Evening Star Avenue Ohio Stone Tools, by Robert N. Converse $ 8.00 add $1.50 P-H Ohio Slate Types, by Robert N. Converse $15.00 add $1.50 P-H SE, East Canton, OH 44730, (216) 488-1640 The Glacial Kame Indians, by Robert N. Converse.$20.00 add $1.50 P-H 1994 Treasurer Don F. Potter, 1391 Hootman Drive, Reynoldsburg, 1980's& 1990's $ 6.00 add $1.50 P-H OH 43068, (614) 861-0673 1970's $ 8.00 add $1.50 P-H 1998 Editor Robert N. Converse, 199 Converse Dr., Plain City, OH 1960's $10.00 add $1.50 P-H 43064, (614)873-5471 Back issues of the Ohio Archaeologist printed prior to 1964 are gen 1994 Immediate Past Pres. -
Hands-On Human Evolution: a Laboratory Based Approach
Hands-on Human Evolution: A Laboratory Based Approach Developed by Margarita Hernandez Center for Precollegiate Education and Training Author: Margarita Hernandez Curriculum Team: Julie Bokor, Sven Engling A huge thank you to….. Contents: 4. Author’s note 5. Introduction 6. Tips about the curriculum 8. Lesson Summaries 9. Lesson Sequencing Guide 10. Vocabulary 11. Next Generation Sunshine State Standards- Science 12. Background information 13. Lessons 122. Resources 123. Content Assessment 129. Content Area Expert Evaluation 131. Teacher Feedback Form 134. Student Feedback Form Lesson 1: Hominid Evolution Lab 19. Lesson 1 . Student Lab Pages . Student Lab Key . Human Evolution Phylogeny . Lab Station Numbers . Skeletal Pictures Lesson 2: Chromosomal Comparison Lab 48. Lesson 2 . Student Activity Pages . Student Lab Key Lesson 3: Naledi Jigsaw 77. Lesson 3 Author’s note Introduction Page The validity and importance of the theory of biological evolution runs strong throughout the topic of biology. Evolution serves as a foundation to many biological concepts by tying together the different tenants of biology, like ecology, anatomy, genetics, zoology, and taxonomy. It is for this reason that evolution plays a prominent role in the state and national standards and deserves thorough coverage in a classroom. A prime example of evolution can be seen in our own ancestral history, and this unit provides students with an excellent opportunity to consider the multiple lines of evidence that support hominid evolution. By allowing students the chance to uncover the supporting evidence for evolution themselves, they discover the ways the theory of evolution is supported by multiple sources. It is our hope that the opportunity to handle our ancestors’ bone casts and examine real molecular data, in an inquiry based environment, will pique the interest of students, ultimately leading them to conclude that the evidence they have gathered thoroughly supports the theory of evolution. -
Human Evolution Timeline 1
Name: Human Origins Web Inquiry 1. Got to the website: http://humanorigins.si.edu/evidence 2. Read the paragraph under “Evidence of Evolution.” 3. In the maroon box on the left, click on “Timeline Interactive” 4. Start by clicking on the red bands at the bottom of the timeline, beginning with Homo sapiens and the rest of the Hominids. Read the information boxes that pop up when you click. 5. When you come across a species you find particularly interesting, record that information (including dates) in the box below. Record facts of at least 5 interesting hominid species, including Homo sapiens. 6. Then begin to explore the rest of the timeline. As you go, record at least 5 other interesting species. 7. Use the magnifier tool at the bottom of the timeline. Click on “color key” to identify the different color dots. Each dot represents a specific piece of evidence scientists have discovered in their study of evolution. 8. Click on various colored dots to learn about tools, events, geology, climate information, behavior, and skeletal adaptations discovered by scientists that add to the evidence for human evolution. Record 7 pieces of evidence you find interesting or particularly important in the box on the back. Make sure to put down different types of evidence (different colors). 9. Take 15-20 minutes to explore the whole timeline and record interesting facts. 10. Build your own timeline on the next page, filling in the interesting species and evidence you recorded in their proper place in the timeline. Hominids Species Dates Interesting fact(s) Homo sapiens Name: Evidence Type of Date Interesting fact(s) Evidence Name: Human Evolution Timeline 1. -
The Adaptive Significance of Human Language
University of Tennessee, Knoxville TRACE: Tennessee Research and Creative Exchange Senior Thesis Projects, 1993-2002 College Scholars 2000 The Adaptive Significance of Human Language Nathan Oesch Follow this and additional works at: https://trace.tennessee.edu/utk_interstp2 Recommended Citation Oesch, Nathan, "The Adaptive Significance of Human Language" (2000). Senior Thesis Projects, 1993-2002. https://trace.tennessee.edu/utk_interstp2/52 This Project is brought to you for free and open access by the College Scholars at TRACE: Tennessee Research and Creative Exchange. It has been accepted for inclusion in Senior Thesis Projects, 1993-2002 by an authorized administrator of TRACE: Tennessee Research and Creative Exchange. For more information, please contact [email protected]. The Adaptive Significance of Human Language Nathan Oesch Department of Psychology University of Tennessee, Knoxville noesch @ utk. edu Abstract Many experts have argued that human language is fundamentally incompatible with the principles of traditional Darwinian evolutionary theory. According to conventional Darwinian explanations, specific traits evolved among species according to gradual and incremental genetic changes, each of which that were in some way so favorable to the survival and reproduction of ancestral generations that they were ultimately preserved within successive generations of those species. Human language, it has been said, is simply to complex to be explained as a result of Darwinian explanations, since each successive step in the evolution of language would confer no obvious survival benefits to its recipients. According to this idea, language is such an "all-or none system," that it could not possibly have existed in any immediately beneficial intermediate forms and thus could not have evolved according to conventional Darwinian modes of explanation. -
Homo Erectus Infancy and Childhood the Turning Point in the Evolution of Behavioral Development in Hominids
10 Homo erectus Infancy and Childhood The Turning Point in the Evolution of Behavioral Development in Hominids Sue Taylor Parker In man, attachment is mediated by several different sorts of behaviour of which the most obvious are crying and calling, babbling and smiling, clinging, non-nutritional sucking, and locomotion as used in approach, following and seeking. —John Bowlby, Attachment The evolution of hominid behavioral ontogeny can be recon - structed using two lines of evidence: first, comparative neontological data on the behavior and development of living hominoid species (humans and the great apes), and second, comparative paleontolog- ical and archaeological evidence associated with fossil hominids. (Although behavior rarely fossilizes, it can leave significant traces.) 1 In this chapter I focus on paleontological and neontological evi - dence relevant to modeling the evolution of the following hominid adaptations: (1) bipedal locomotion and stance; (2) tool use and tool making; (3) subsistence patterns; (4) growth and development and other life history patterns; (5) childbirth; (6) childhood and child care; and (7) cognition and cognitive development. In each case I present a cladistic model for the origins of the characters in question. 2 Specifically, I review pertinent data on the following widely recog - nized hominid genera and species: Australopithecus species (A. afarensis , A. africanus , and A. robustus [Paranthropus robustus]) , early Homo species (Australopithecus gahri , Homo habilis , and Homo rudolfensis) , and Middle Pleistocene Homo species (Homo erectus , Homo ergaster , and others), which I am calling erectines . Copyrighted Material www.sarpress.org 279 S UE TAYLOR PARKER Table 10.1 Estimated Body Weights and Geological Ages of Fossil Hominids _______________________________________________________________________ Species Geologic Age Male Weight Female Weight (MYA) (kg) (kg) _______________________________________________________________________ A. -
New Fossils of Australopithecus Anamensis from Kanapoi, West Turkana, Kenya (2003E2008)
Journal of Human Evolution 65 (2013) 501e524 Contents lists available at SciVerse ScienceDirect Journal of Human Evolution journal homepage: www.elsevier.com/locate/jhevol New fossils of Australopithecus anamensis from Kanapoi, West Turkana, Kenya (2003e2008) C.V. Ward a,*, F.K. Manthi b, J.M. Plavcan c a Department of Pathology and Anatomical Sciences, M263 Medical Sciences Building, University of Missouri, Columbia, MO 65212, USA b Department of Earth Sciences, National Museums of Kenya, P.O. Box 40658, Nairobi, Kenya c Department of Anthropology, 330 Old Main, University of Arkansas, Fayetteville, AR 72701, USA article info abstract Article history: Renewed fieldwork from 2003 through 2008 at the Australopithecus anamensis type-site of Kanapoi, Received 29 January 2013 Kenya, yielded nine new fossils attributable to this species. These fossils all date to between 4.195 and Accepted 7 May 2013 4.108 million years ago. Most were recovered from the lower fluvial sequence at the site, with one from Available online 30 August 2013 the lacustrine sequence deltaic sands that overlie the lower fluvial deposits but are still below the Kanapoi Tuff. The new specimens include a partial edentulous mandible, partial maxillary dentition, two Keywords: partial mandibular dentitions, and five isolated teeth. The new Kanapoi hominin fossils increase the Australopithecus anamensis sample known from the earliest Australopithecus, and provide new insights into morphology within this Kanapoi New fossils taxon. They support the distinctiveness of the early A. anamensis fossils relative to earlier hominins and Dentition to the later Australopithecus afarensis. The new fossils do not appreciably extend the range of observed variation in A. -
Human Evolution: a Paleoanthropological Perspective - F.H
PHYSICAL (BIOLOGICAL) ANTHROPOLOGY - Human Evolution: A Paleoanthropological Perspective - F.H. Smith HUMAN EVOLUTION: A PALEOANTHROPOLOGICAL PERSPECTIVE F.H. Smith Department of Anthropology, Loyola University Chicago, USA Keywords: Human evolution, Miocene apes, Sahelanthropus, australopithecines, Australopithecus afarensis, cladogenesis, robust australopithecines, early Homo, Homo erectus, Homo heidelbergensis, Australopithecus africanus/Australopithecus garhi, mitochondrial DNA, homology, Neandertals, modern human origins, African Transitional Group. Contents 1. Introduction 2. Reconstructing Biological History: The Relationship of Humans and Apes 3. The Human Fossil Record: Basal Hominins 4. The Earliest Definite Hominins: The Australopithecines 5. Early Australopithecines as Primitive Humans 6. The Australopithecine Radiation 7. Origin and Evolution of the Genus Homo 8. Explaining Early Hominin Evolution: Controversy and the Documentation- Explanation Controversy 9. Early Homo erectus in East Africa and the Initial Radiation of Homo 10. After Homo erectus: The Middle Range of the Evolution of the Genus Homo 11. Neandertals and Late Archaics from Africa and Asia: The Hominin World before Modernity 12. The Origin of Modern Humans 13. Closing Perspective Glossary Bibliography Biographical Sketch Summary UNESCO – EOLSS The basic course of human biological history is well represented by the existing fossil record, although there is considerable debate on the details of that history. This review details both what is firmly understood (first echelon issues) and what is contentious concerning humanSAMPLE evolution. Most of the coCHAPTERSntention actually concerns the details (second echelon issues) of human evolution rather than the fundamental issues. For example, both anatomical and molecular evidence on living (extant) hominoids (apes and humans) suggests the close relationship of African great apes and humans (hominins). That relationship is demonstrated by the existing hominoid fossil record, including that of early hominins. -
Morphological Affinities of the Sahelanthropus Tchadensis (Late Miocene Hominid from Chad) Cranium
Morphological affinities of the Sahelanthropus tchadensis (Late Miocene hominid from Chad) cranium Franck Guy*, Daniel E. Lieberman†, David Pilbeam†‡, Marcia Ponce de Leo´ n§, Andossa Likius¶, Hassane T. Mackaye¶, Patrick Vignaud*, Christoph Zollikofer§, and Michel Brunet*‡ *Laboratoire de Ge´obiologie, Biochronologie et Pale´ontologie Humaine, Centre National de la Recherche Scientifique Unite´Mixte de Recherche 6046, Faculte´des Sciences, Universite´de Poitiers, 40 Avenue du Recteur Pineau, 86022 Poitiers Cedex, France; §Anthropologisches Institut, Universita¨t Zu¨rich-Irchel, Winterthurerstrasse 190, 8057 Zu¨rich, Switzerland; †Peabody Museum, Harvard University, 11 Divinity Avenue, Cambridge, MA 02138; and ¶Department de Pale´ontologie, Universite´deNЈDjamena, BP 1117, NЈDjamena, Republic of Chad Contributed by David Pilbeam, November 5, 2005 The recent reconstruction of the Sahelanthropus tchadensis cra- cross-sectional ontogenetic samples of Pan troglodytes (n ϭ 40), nium (TM 266-01-60-1) provides an opportunity to examine in Gorilla gorilla (n ϭ 41), and Homo sapiens (n ϭ 24) (see Table detail differences in cranial shape between this earliest-known 3, which is published as supporting information on the PNAS hominid, African apes, and other hominid taxa. Here we compare web site). In addition, we digitized as many of the same land- the reconstruction of TM 266-01-60-1 with crania of African apes, marks as possible on a sample of available relatively complete humans, and several Pliocene hominids. The results not only fossil hominid crania: the stereolithograhic replica of AL 444-2 confirm that TM 266-01-60-1 is a hominid but also reveal a unique (Australopithecus afarensis) (9); CT scans of Sts 5 and Sts 71 mosaic of characters. -
Paranthropus Boisei: Fifty Years of Evidence and Analysis Bernard A
Marshall University Marshall Digital Scholar Biological Sciences Faculty Research Biological Sciences Fall 11-28-2007 Paranthropus boisei: Fifty Years of Evidence and Analysis Bernard A. Wood George Washington University Paul J. Constantino Biological Sciences, [email protected] Follow this and additional works at: http://mds.marshall.edu/bio_sciences_faculty Part of the Biological and Physical Anthropology Commons Recommended Citation Wood B and Constantino P. Paranthropus boisei: Fifty years of evidence and analysis. Yearbook of Physical Anthropology 50:106-132. This Article is brought to you for free and open access by the Biological Sciences at Marshall Digital Scholar. It has been accepted for inclusion in Biological Sciences Faculty Research by an authorized administrator of Marshall Digital Scholar. For more information, please contact [email protected], [email protected]. YEARBOOK OF PHYSICAL ANTHROPOLOGY 50:106–132 (2007) Paranthropus boisei: Fifty Years of Evidence and Analysis Bernard Wood* and Paul Constantino Center for the Advanced Study of Hominid Paleobiology, George Washington University, Washington, DC 20052 KEY WORDS Paranthropus; boisei; aethiopicus; human evolution; Africa ABSTRACT Paranthropus boisei is a hominin taxon ers can trace the evolution of metric and nonmetric var- with a distinctive cranial and dental morphology. Its iables across hundreds of thousands of years. This pa- hypodigm has been recovered from sites with good per is a detailed1 review of half a century’s worth of fos- stratigraphic and chronological control, and for some sil evidence and analysis of P. boi se i and traces how morphological regions, such as the mandible and the both its evolutionary history and our understanding of mandibular dentition, the samples are not only rela- its evolutionary history have evolved during the past tively well dated, but they are, by paleontological 50 years.