Annotated Checklist of Vascular Flora Canyonlands National Park

Home , Achnatherum parishii, Antennaria marginata, Artemisia spinescens, Astragalus iselyi, Astragalus missouriensis, Astragalus paradoxus

National Park Service U.S. Department of the Interior

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Natural Resource Technical Report NPS/NCPN/NRTR—2009/221 ON THE COVER View of the Colorado River and White Rim from Island in the Sky, Canyonlands National Park, Utah. Photograph by Walter Fertig. Annotated Checklist of Vascular Flora Canyonlands National Park

Natural Resource Technical Report NPS/NCPN/NRTR—2009/221

Authors Walter Fertig Moenave Botanical Consulting 1117 W. Grand Canyon Dr. Kanab, UT 84741

Sarah Topp Northern Colorado Plateau Network National Park Service P.O. Box 848 Moab, UT 84532

Mary Moran Southeast Utah Group National Park Service P.O. Box 907 Moab, UT 84532

Editing and Design Alice Wondrak Biel Northern Colorado Plateau Network National Park Service P.O. Box 848 Moab, UT 84532

June 2009

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Please cite this publication as:

Fertig, W., S. Topp, and M. Moran. 2009. Annotated checklist of vascular ﬂ ora: Canyonlands National Park. Natural Resource Technical Report NPS/NCPN/NRTR—2009/221. National Park Service, Fort Collins, Colorado.

NPS 100066, June 2009 ii Annotated Checklist of Vascular Flora: Canyonlands National Park Contents

Figures ...... v Tables ...... v Abstract ...... vii Acknowledgements ...... ix

1 Introduction ...... 1 1.1 Background and justiﬁ cation ...... 1 1.2 Objectives ...... 1 1.3 Study area ...... 1 1.3.1 History and setting ...... 1 1.3.2 Geology ...... 2 1.3.3 Vegetation ...... 2 1.3.4 Previous botanical studies ...... 4

2 Methods ...... 7

3 Results ...... 9 3.1 Summary of the ﬂ ora of Canyonlands National Park ...... 9 3.2 Life form and biogeographic diversity ...... 10 3.3 Plants of special concern ...... 10 3.4 Non-native species and noxious weeds ...... 11 4 Discussion ...... 15

5 Literature Cited ...... 17

Appendices ...... 21 Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes...... 23 Appendix B. Potential (unconfi rmed) taxa of Canyonlands National Park ...... 82 Appendix C. Rejected (falsely reported or questionable) taxa of Canyonlands National Park. ...90

Appendix D. Conﬁ rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes...... 100

Contents iii

Figures

Figure 1.3.1. Location of Canyonlands National Park, Grand, Garﬁ eld, San Juan, and Wayne counties, Utah...... 3 Figure 1.3.4. Increase in the number of plant taxa conﬁ rmed as present in Canyonlands National Park, 1960–2007...... 6

Tables

Table 1.3.4. Chronology of plant-collecting efforts in Canyonlands National Park...... 5 Table 3.1. Statistical summary of the ﬂ ora of Canyonlands National Park...... 9 Table 3.3. Plant species of conservation concern from Canyonlands National Park...... 11 Table 3.4. Noxious weed species of Canyonlands National Park...... 14

Contents v

Abstract

In 2005–2006, the National Park Service’s Northern Colorado Plateau Network received funding to develop a revised checklist of the vascular plant fl ora of Canyonlands National Park. This entailed a thorough review of existing literature, re-examination of specimens from the Canyonlands National Park herbarium and other state and national museum collections, and additional fi eld work to corrob- orate unverifi ed species reports or locate new species. Based on this study, 594 vascular plant taxa have been confi rmed or reliably reported as occuring in the park (534 are known from validated herbarium specimens and 60 from unvouchered literature reports). Another 100 species formerly considered part of the park fl ora have now been shown to be falsely reported. The current fl ora of Canyonlands National Park represents 16.5% of the known fl ora of Utah, and 35.1% of the fl ora of the Colorado Plateau ecoregion within Utah. At least 45 of these species are either listed as sensitive by various agen- cies or considered species of concern by the state natural heritage program. Introduced plants make up 11.4% of the park fl ora (68 species), of which only seven species are considered noxious weeds by the State of Utah. Further surveys in Canyonlands National Park are likely to fi nd additional new species (at least 161 are known from similar habitats outside the park boundary), especially among late-fl owering, weedy, or wetland taxa.

Contents vii

Acknowledgements

We would like to thank the following individuals for their assistance with this project: Dr. Angela Evenden, botanist and former leader of the Northern Colorado Plateau Network (NCPN) program in Moab, Utah, was an active proponent of this project and helped secure funding for fi eld work and herbarium studies in 2006. Vicki Webster, curator of museum collections for the National Park Service Southeast Utah Group (SEUG) provided access to all mounted and unmounted herbarium specimens from Canyonlands National Park (CANY), as well as work space within the museum vaults. Melissa Powell, formerly of the NCPN, recorded specimen-label data from all mounted collections in the CANY herbarium. Charlie Schelz, biologist with SEUG, provided research permits for work in 2005–2006 and shared his unmounted plant collections. Libby Nance, former database special- ist with the NCPN, assisted with development and quality control of the park’s NPSpecies database. Libby Nance, Janet Coles, and Helen Thomas also provided data from recent vegetation mapping and wetland inventories on the park. Drs. Stanley Welsh and Duane Atwood provided access to herbarium collections at Brigham Young University. Dr. Michael Piep provided access to collections at the Utah State University (Intermountain) herbarium. Dr. Ron Hartman and Ernie Nelson provided access and specimen loans from the Rocky Mountain Herbarium at the University of Wyoming. Dr. John Ander- son, of the Bureau of Land Management’s Arizona offi ce, shared information from his 1983 rare plant fi eld work. Dr. Leila Shultz, of Utah State University, helped locate specimens and data from Loope’s 1977 dissertation. Thanks to the technical reviewers (Dr. Angela Evenden, Dr. Dusty Perkins, and Libby Nance) for their comments and suggestions. To any others whom we may have inadvertently forgotten, please accept our apologies (and thanks).

Research was conducted under permit #CANY-2006-SCI-0012 for study number CANY-00084. Funding was provided by the National Park Service through the University of Wyoming.

Contents ix

vertebrates and vascular plants for each park unit 1 Introduction in the network (Evenden et al. 2002). This eff ort has entailed reviewing existing species checklists, In 2006, the National Park Service (NPS) estab- corroborating the identifi cation of specimens in lished a cooperative agreement with the Univer- park museum and herbarium collections, locat- sity of Wyoming, through the Rocky Mountains ing additional specimen records from other mu- Cooperative Ecosystem Studies Unit, to produce seums, mining relevant literature and datasets, a series of technical reports summarizing the vas- and conducting fi eld surveys to locate new taxa cular plant fl ora of each of the 16 national park or confi rm unsubstantiated reports.1 All of these units of the Northern Colorado Plateau Network data have been synthesized in NPSpecies, the (NCPN). These reports are based on existing in- park service’s standardized database for docu- formation in the park service’s NPSpecies data- menting park status (present, probably present, base system and augmented by further data min- unconfi rmed, falsely reported), abundance, resi- ing, reviews of state and regional herbaria, and dency, nativity, and rarity. With the production of new fi eld surveys to resolve the status of many a technical series of reports on the fl ora of each reported and unconfi rmed species. The objec- park, the NPSpecies data will now be available to tive of these reports is to summarize the current a broader audience. state of knowledge of the fl ora of each park for the benefi t of NPS biologists and interpretive staff , outside researchers, and the general public. 1.2 Objectives The following report addresses the vascular fl ora The objectives of this project were to: of Canyonlands National Park, in southeastern Utah. Other NCPN park units are addressed in 1. Document the current status (present, separate reports. reported, potential, or falsely reported) of all vascular-plant species that have been cited as occurring in Canyonlands National Park, 1.1 Background and justifi cation based on a review of herbarium collections One of the primary missions of the NPS is “to and past literature. conserve unimpaired the natural and cultural resources and values of the national park sys- 2. Develop an annotated checklist of the tem for the enjoyment of this and future genera- fl ora of Canyonlands National Park, with tions” (NPS 2000). Due to their high degree of supplemental information on taxonomic protection and distribution across a wide variety synonyms, distribution, within-park abun- of habitat types, the nation’s national park units dance, fl owering period, growth form, nativ- make a signifi cant contribution to the protection ity, habitat, and sources for each record. of biological diversity (Noss and Cooperrider 1994). Unfortunately, the full contribution of the 1.3 Study area National Park System is not known, as many park units have traditionally lacked comprehensive 1.3.1 History and setting biological inventories or monitoring programs to Although hundreds of archeological sites indicate identify the composition and condition of their longtime human occupation of the Canyonlands biota. Knowing the status of park fl ora and fauna area (especially prior to 1300 A.D.), the arid me- is critical for eff ective park management and for sas and rugged canyons at the confl uence of the systematically identifying and fi lling gaps in the Green and Colorado rivers were some of the last nation’s protected area network (Margules and areas in the U.S. to be explored and settled by Pressey 2000). Europeans. During the Great Depression, U.S. Interior Secretary Harold Ickes proposed set- To rectify this shortcoming, Congress passed ting much of the canyon country of southeastern the National Parks Omnibus Management Act Utah aside as the Escalante National Monument, in 1998, to provide guidance and funding for to conserve watersheds and draw tourists. Action an inventory and monitoring program across on this proposal was stalled by World War II and the entire National Park System. Since 2000, the the uranium boom of the 1950s. Discussions re- NCPN Inventory and Monitoring Program has sumed in the 1960s, though centered on a smaller been developing and verifying species lists of

1Taxa refer to all named species, subspecies, and varieties.

Chapter 1: Introduction 1 area bordering the Colorado River corridor. Con- ing the Permian, the Canyonlands area fl uctuated gress and President Lyndon Johnson established between a shallow sea and coastal beaches, mud- Canyonlands National Park (NP) in 1964; its fl ats, and dunes. These deposits gave rise to the boundaries were expanded in 1971 (Evenden et Cedar Mesa Sandstone, Cutler Formation, and al. 2002). White Rim Limestone (Chronic 1988) that today are widely exposed in the Needles and Maze dis- Canyonlands National Park is located in north- tricts. In the Triassic, the region was a large delta, western San Juan, southwestern Grand, south- buried by river- and stream-borne mud, silt, and eastern Wayne, and northeastern Garfi eld coun- sand that became the Moenkopi and Chinle for- ties, Utah, and consists of four main subsections mations. These beds are exposed today above the (Figure 1.3.1) that total 136,610 hectares (337,570 White Rim in the ISKY area. Late in the Triassic acres). The Island in the Sky (ISKY) region is and into the Jurassic, the Canyonlands area was located at the north end of the park, in the V- enveloped by Sahara-like dune systems (Wingate shaped wedge above the confl uence of the Colo- and Navajo sandstones) interrupted by a period rado and Green rivers. It consists of a steep-sided of stream deposition (Kayenta Formation). The Wingate and White Rim sandstone plateau, con- Wingate, Kayenta, and Navajo formations form nected to mesas to the north by a slender isthmus the steep cliff s of the Island in the Sky. Quaterna- called The Neck. The Needles District occupies ry Aeolian deposits occur on the summit of ISKY, the southeastern quadrant of the park, on the the grassland parks of the Needles, and in the Salt east bank of the Colorado River, and is charac- Creek watershed (Baars 2003; Loope 1977). terized by extensive desert grasslands amid stone turrets, fi ns, toadstools, and arches carved from 1.3.3 Vegetation Cedar Mesa Sandstone. West of the Colorado River is The Maze District, an area of bewilder- Loope (1977) identifi ed six major vegetation ing and intricately carved canyons and gullies of types in Canyonlands NP. He found vegetation Cedar Mesa Sandstone. The Horseshoe Canyon patterns to be strongly correlated with soil depth- Unit is an isolated segment of the park located at to-bedrock and moisture availability. Areas with the head of Horseshoe Canyon (a tributary wash thin, lithic soils or exposed sandstone bedrock of the Green River); it protects the Great Gallery and cliff rims support dwarf forests of two-needle pictograph site. pinyon (Pinus edulis) and Utah juniper (Juniperus osteosperma), with an understory of dwarf moun- Canyonlands NP ranges in elevation from 1,190 tain mahogany (Cercocarpus intricatus), Utah meters (3,900 ft) at the bottom of Cataract Can- serviceberry (Amelanchier utahensis), cliff -rose yon to 2,188 meters (7,180 ft) in the Needles (Purshia mexicana var. stansburyana), and Big- (Evenden et al. 2002). Average maximum tem- elow’s sagebrush (Artemisia bigelovii). Pinyon-ju- peratures range from 64.1 to 68.3° F, and average niper woodlands are best developed on outcrops minimum temperatures from 37.8 to 42.2° F. An- of Kayenta, Navajo, Cedar Mesa, and White Rim nual precipitation averages 219–235 mm (8.6–9.3 sandstones or on rocky slopes of washes. in), with peaks in March and August (Evenden et al. 2002; Loope 1977). Mesa tops and alluvial benches with soils up to 30 cm deep to bedrock typically support blackbrush 1.3.2 Geology (Coloegyne ramosissima) communities. Black- brush is dependent on a subsurface hardpan or Uplifting of the Colorado Plateau in the Pleisto- bedrock layer to trap soil moisture that would cene, and downcutting of the Colorado River, otherwise percolate beyond the root layer. Com- has exposed over 3,000 feet of rock strata and 100 monly associated species in blackbrush stands million years of geologic history in Canyonlands include galleta (Hilaria jamesii), shadscale (Atri- NP. The oldest rocks in the park are thick beds of plex confertifolia), and Torrey’s ephedra (Ephedra salt and gypsum, of the Paradox Formation, that torreyana). Park lands with deeper soils (often outcrop along the lowermost reaches of the Col- associated with aeolian deposits over 50 cm thick orado River. The Paradox layer was derived from above bedrock) tend to support desert grasslands repeated cycles of evaporation of shallow marine dominated by needle-and-thread (Stipa coma- waters in a landlocked bay during the Pennsylva- ta), Indian ricegrass (S. hymenoides), blue grama nian (Baars 2003). The salt and gypsum beds were (Bouteloua gracilis), galleta, winterfat (Kraschen- later buried by sandy limestones of the Honaker innikovia lanata), and green ephedra (Ephedra Trail Member of the Hermosa Formation. Dur- viridis). Because of their diff use root systems,

2 Annotated Checklist of Vascular Flora: Canyonlands National Park Canyonlands National Park Utah

313

Island in the Sky HORSESHOE Visitor Center CANYON ISLAND UNIT IN THE SKY

G re e

i v

Hans Flat Visitor Center

THE MAZE Needles Visitor Center # 211

r Rive o d a r lo o C THE NEEDLES

Established September 12, 1964 Legend 1,189–2,188 m elevation Intermittent streams Dead Horse Point State Park (3,900–7,180 ft elevation) Rivers Glen Canyon NRA (NPS) 136,610 ha (337,570 ac) Paved roads Bureau of Land Management 02461 Unpaved roads Private Kilometers Unimproved/4WD roads State 02461 Park boundary (NPS) U.S. Forest Service Miles

MAP CREATED BY ANETH WIGHT, NCPN (DERIVED FROM EVENDEN ET AL. 2002) Figure 1.3.1. Location of Canyonlands National Park, Grand, Garﬁ eld, San Juan, and Wayne counties, Utah.

Chapter 1: Introduction 3 grass species are better adapted to exploit water Perhaps the rarest vegetation type documented that passes rapidly through deep, sandy soils. within Canyonlands NP consists of small patches Desert grasslands in much of the Needles region of Douglas-fi r (Pseudotsuga menziesii) and Rocky were grazed more intensively than remote and Mountain maple (Acer glabrum) on cool, shaded, aridic portions of the Maze and ISKY areas and, north-facing canyon slopes or alcoves (Welsh consequently, have less biological soil crust and 1970). lower vascular-plant species richness (Kleiner and Harper 1972). 1.3.4 Previous botanical studies Basin big sagebrush (Artemisia tridentata var. tri- The Canyonlands area was essentially unexplored dentata) communities are relatively infrequent botanically until Stanley Welsh, Glen Moore, and within Canyonlands NP, and occur mostly on their colleagues from Brigham Young Univer- alluvial benches and toeslopes with moderately sity (BYU) initiated baseline fl oristic studies in deep soils and a high water table (Loope 1977). 1964–1965. In two separate publications, Welsh Most stands in the park occur along benches and Moore (1965) documented 234 taxa from the bordering the Green and Colorado rivers, and in Needles District, and Moore and Welsh (1965) re- alluvial bottomlands in Salt Creek and Lavender ported 92 species from the Island in the Sky area. and Davis canyons. Frequently associated species Taking into account changes in species concepts, include fourwing saltbush (Atriplex canescens), misidentifi cations, and unvouchered reports, the greasewood (Sarcobatus vermiculatus), rubber fl ora of Canyonlands NP stood at 212 taxa in 1966 rabbitbrush (Chrysothamnus nauseosus), and (Table 1.3.4). In 1967, Edgar Kleiner and Kimball Gambel’s oak (Quercus gambelii). Harper, of the University of Utah, documented at least 13 new vascular-plant species as part of their Salt desert scrub vegetation predominates on research on grazing eff ects on desert grassland low benches with clayey soils. These stands typi- vegetation of Chesler and Virginia parks, in the cally consist of Garrett’s saltbush (Atriplex gar- Needles area (Kleiner and Harper 1972). Includ- rettii) and shadscale with budsage (Artemisia ing additional surveys by Welsh, Moore, and N. spinescens), Torrey’s ephedra, desert trumpet Duane Atwood (of BYU), the park fl ora reached (Eriogonum infl atum), and Hopi blanketfl ower 239 taxa at the end of the 1960s (Figure 1.3.4). (Gaillardia pinnatifi da) and are often associated with outcrops of the Chinle, Moenkopi, or Cutler Walter Loope, a graduate student from Utah State formations (Loope 1977). University, conducted a thorough vegetation study of Canyonlands NP from 1972 to 1974. Loope Riparian communities occur along perennial wa- (1977) discovered at least six new species for the tercourses and in hanging gardens within shady park, and reported four others while compiling a alcoves or cliff s. Coyote willow (Salix exigua var. checklist of 298 species. Over the remainder of stenophylla) and fi ve-stamen tamarisk (Tamarix the 1970s, Peg Johnson, David Johnson, and Di- chinensis) form a narrow strip of dense vegeta- ane Allen added 48 additional species while help- tion bordering much of the Colorado and Green ing to develop the park’s herbarium collections rivers. These stands are interspersed by wetlands (Table 1.3.4). of cattails (Typha domingensis or T. latifolia) or common reed (Phragmites australis). Moist ter- During the 1980s, fl oristic, rare-plant, and veg- races located slightly upslope of the active river etation monitoring studies added nearly 150 spe- channel support scattered Fremont cottonwood cies to the documented fl ora of Canyonlands NP (Populus fremontii), desert olive (Forestieria pube- (Figure 1.3.4). Anderson (1983) conducted a rare scens), and Goodding’s black willow (Salix good- plant study in the Needles area and discovered dingii). Hanging gardens occur infrequently in nine new species for the park fl ora. Stan Welsh, shady and permanently wet sandstone seeps, and Matthew Chatterley, and Larry Higgins, who support water birch (Betula occidentalis), golden conducted surveys along the Colorado River and sedge (Carex aurea), alcove columbine (Aquilegia in the ISKY area, contributed 42 new species to micrantha), cave primrose (Primula specuicola), the Canyonlands NP fl ora. Jayne Belnap and Tim alcove death-camas (Zigadenus vaginatus), East- Graham, of the NPS and U.S. Geological Sur- wood’s monkey-fl ower (Mimulus eastwoodiae), vey (USGS), added at least 60 new species in the and southern maidenhair fern (Adiantum capil- course of establishing a network of permanent lus-veneris). monitoring transects throughout the park. Other

4 Annotated Checklist of Vascular Flora: Canyonlands National Park NPS naturalists, rangers, and seasonal employees the Colorado Plateau. Beginning in 1996, NPS collected at least 35 new species for the park dur- Southeast Utah Group (SEUG) biologist Charlie ing the decade (Table 1.3.4). Schelz collected at least 24 new species and cited 46 others without vouchers in annual park check- From 1989 to 1991, Ken Heil and colleagues from lists (Schelz et al. 2006). Since 2003, 50 new spe- San Juan College, in New Mexico, surveyed na- cies have been confi rmed or reported for the park tional parks of the Colorado Plateau and discov- fl ora through vegetation mapping studies (NCPN ered 27 new species for Canyonlands NP. Fowler in ed.), weed inventories (Dewey and Anderson and others (2007) added two new wetland species 2005), and in the course of preparing this docu- during a fl oristic inventory of hanging gardens of ment (Table 1.3.4, Figure 1.3.4).

Table 1.3.4. Chronology of plant-collecting efforts in Canyonlands National Park. # of new Year Collector(s) Comments taxa 1964 S. L. Welsh, G. 202 Stan Welsh, Glen Moore, and colleagues from Brigham Young University (BYU) Moore, S. G. conducted a baseline fl oristic study of Canyonlands NP and documented 202 species Canter, and M. for the park. Among the more noteworthy collections was the type of Fisher Olsen milkvetch (Astragalus piscator) from Salt Canyon (Welsh 1982). 1965– S. L. Welsh, G. 10 Welsh and Moore (1965) published a checklist of the fl ora of the Needles area of 1966 Moore, and D. Canyonlands NP, citing 234 taxa. Moore and Welsh (1965) published a comparable Wiens list for the Island in the Sky (ISKY) region, with 92 taxa. Based on later corrections and excluding unvouchered taxa, the total park fl ora, through 1966, stood at 212 species. 1967 E. G. Kleiner 13 Edgar Kleiner and Kimball Harper, of the University of Utah, added at least 13 new and K. T. Harper species to the Canyonlands NP fl ora through their fi eld work describing grassland communities in Chesler and Virginia parks (Kleiner and Harper 1972). 1968– S. L. Welsh, G. 17 Welsh and colleagues from BYU added 14 species from 1968 to 1969, and 3 more in 1970 Moore, and N. 1970. D. Atwood 1972– W. Loope 6 (+ 4 Walter Loope, of Utah State University, sampled 157 vegetation plots throughout 1973 reported) the park and compiled a checklist of 298 species as part of his doctoral dissertation (Loope 1977). 1977– P. Johnson, D. 48 Peg Johnson, David Johnson, and Diane Allen made nearly 200 collections over a 1979 Johnson, and D. three-year period for the CANY herbarium and documented 48 new species for the Allen park. 1981– K. Kitchell and 3 Kathy Kitchell and Damian Fagan collected 14 specimens for the CANY herbarium, 1982 D. Fagan including fi rst collections of 3 taxa. 1983 J. Anderson, 55 (+ 2 U.S. Fish and Wildlife Service botanist John Anderson conducted a rare plant survey S. L. Welsh, M. reported) in the Needles Section of Canyonlands NP to assess potential impacts of a proposed Chatterly, L. nuclear waste facility and, in the process, discovered 9 new species. Ann DeBolt Higgins, and A. collected more than 30 specimens for the CANY herbarium, including 6 new taxa DeBolt for the park. Stan Welsh and colleagues made more than 60 collections along the Colorado River and in the ISKY area, and added 42 new species to the park fl ora. 1984– A. DeBolt, N. 23 Ann DeBolt, Nelson Boschen, and Judie Chrobak collected 45 specimens for the 1986 Boschen, J. CANY herbarium and added 23 new species to the park fl ora. Boschen (1985) Chrobak compiled a checklist for Canyonlands NP, with 429 species. 1987– J. Belnap, T. 65 Jayne Belnap and Tim Graham expanded on the monitoring studies initiated by 1988 Graham, K. Kleiner and Harper and added 60 new species to the park fl ora. Kaye Thorne and Thorne, L. colleagues added 3 new species, and Judie Chrobak-Cox collected 2 new taxa. Thomas, T. Inskip, and J. Chrobak-Cox

Chapter 1: Introduction 5 Table 1.3.4. Chronology of plant-collecting efforts in Canyonlands National Park, cont. # of new Year Collector(s) Comments taxa 1989– K. Heil, R. 24 (+ 3 Ken Heil, Rich Fleming, J. Mark Porter, and associates from San Juan College 1991 Fleming, and J. reported) collected more than 150 specimens for the CANY herbarium and documented 24 M. Porter new species for Canyonlands NP as part of a multi-year rare plant and fl oristic inventory of Colorado Plateau national parks. 1993 J. Fowler and C. 2 Jim Fowler and C. May discovered 2 new species during their comprehensive May inventory of hanging-garden communities in the Colorado Plateau (Fowler et al. 2007). 1996– C. Schelz and 21 (+46 Charlie Schelz, biologist with the Southeast Utah Group, added at least 20 new 2002 M. Miller reported) species for the park fl ora, and Mark Miller documented 1 additional species. Schelz and others (2006) compiled annual checklists of the park fl ora and cited at least 46 additional species that have not been vouchered within Canyonlands NP. 2003– K. Andersen, 23 (+ 5 Dewey and Andersen (2005) documented one new weed species and reported four 2004 S. Dewey, C. reported) others during their two-year inventory of exotic and noxious plants of the park. Schelz, S. Topp, Sarah Topp, Lisa Hahn, and Liz Ballenger added 18 new species and 1 additional L. Hahn, and L. species report while conducting vegetation mapping fi eld work (NCPN in ed.). Ballenger Charlie Schelz also documented 4 new species. 2005 G. Rink 4 Glen Rink made 19 collections for the CANY herbarium and added 4 new species to the park during a late-fall fl oat trip down the Colorado River. 2006– W. Fertig, M. 18 Walter Fertig added 9 new species in 2006, and Mary Moran, Sarah Topp, Becky 2007 Moran, S. Topp, Mann, and Joe Castello, of the NPS and the U.S. Geological Survey, collected 9 R. Mann, and J. additional species in 2007. Castello

600

500

400

300

200 Number of vascular taxa

100

0 1960 1970 1980 1990 2000 2007

Year

Figure 1.3.4. Increase in the number of plant taxa conﬁ rmed as present in Canyonlands National Park, 1960– 2007.

6 Annotated Checklist of Vascular Flora: Canyonlands National Park North America Editorial Committee 1993, 1997, 2 Methods 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c; Holmgren et al. 2005; and Weber and In July 2005, we reviewed all mounted specimens Wittmann 2001). The annotations to these lists in the Canyonlands National Park herbarium and label data from the CANY herbarium review (CANY), housed in the SEUG offi ce in Moab, were then incorporated into the NPSpecies data- Utah, to determine whether each was correctly base for the park. identifi ed to species, subspecies, or variety following the nomenclature of Welsh and others Work in 2006–2007 focused on confi rmation of (2003). As necessary, we made corrections and reported and potential species for the park fl ora annotations to each specimen and to the voucher through fi eld surveys and review of collections records in the park’s Automated National Cata- from state and regional herbaria. We made col- loging System (ANCS+) and NPSpecies data- lecting trips to Canyonlands NP in summer and base. fall 2006, and summer 2007, to target missing species, emphasizing undersampled wetland, weedy, Based on this herbarium work and a review of and late-fl owering taxa. Herbarium studies were park references (Anderson 1983; Moore and undertaken at Brigham Young University (BRY), Welsh 1965; Kleiner and Harper 1972; Loope Utah State University (USU), and the University 1977; Schelz et al. 2006; Springer et al. 2006; Welsh of Wyoming (RM) in March, June, and October and Moore 1965), we developed four lists: 2006 and February and March 2007. We also documented park specimens from the Virtual 1. A revised checklist of species that were pres- Herbarium website of the New York Botanical ent in the park (confi rmed by an authenti- Garden (http://sciweb.nybg.org/science2/hcol/ cated voucher; included in Appendices A intf/index.asp). Finally, we found additional new and D); species reports by examining unmounted col- 2. A list of species that were reported in the lit- lections by SEUG and NCPN researchers and erature but not corroborated with a voucher reviewing plot data from recent vegetation map- (equivalent to “probably present” in NPSpe- ping, weed, and wetland inventories (Dewey and cies terminology; included in Appendices A Andersen 2005; Fowler et al. 2007; NCPN in ed.; and D); Springer et al. 2006). 3. A list of potential species (“unconfi rmed” in NPSpecies) known from the vicinity of Canyonlands National Park using an unpublished list generated by Dr. N. Duane Atwood of Brigham Young University and Albee and others (1988) (included in Appen- dix B); and 4. A list of species that are now known to be misidentifi ed or highly questionable (Ap- pendix C). Each list was annotated with supplemental information on life form, geographic range (within the context of Utah), population size in the park (derived from Loope 1977; NCPN in ed.; and fi eld observations), fl owering period, distribution across general vegetation types, nativity, data source (collector and collection number from a sample specimen or literature citation), year of earliest collection (from herbarium records), and relevant synonymy and taxonomic problems (based on review of Barkworth et al. 2003, 2007; Barneby 1989; Cronquist 1994; Cronquist et al. 1972, 1977, 1984, 1997; Dorn 2001; Flora of

Chapter 2: Methods 7

a duplicate voucher deposited in the CANY her- 3 Results barium), Utah State University, and the University of Wyoming’s Rocky Mountain Herbarium. Field 3.1 Summary of the fl ora of surveys in 2004–2007 netted another 23 new species: 10 from NPS and USGS biologists Mary Mo- Canyonlands National Park ran, Sarah Topp, Becky Mann, and Joe Castello, 4 Of the 1,199 vascular plant vouchers in the CANY from Glenn Rink, and 9 from the fi eld collections herbarium that we examined in 2005 (excluding of Walter Fertig. Overall, the documented fl ora 363 specimens on loan), 65 were misidentifi ed of Canyonlands NP has increased by 83 species (5.4%), 62 were revised to a more current name (15.5%), bringing the total number of taxa to 534 (5.2%), 115 had variety or subspecies names (Table 3.1, Appendices A, D). added (9.6%), and 957 were found to be correctly determined (79.8%). Based solely on these Currently, 60 additional taxa have been reported collections, the confi rmed fl ora of Canyonlands for Canyonlands NP but have yet to be corrobo- National Park stood at 451 taxa (Fertig 2006). rated with a voucher specimen (Table 3.1, Ap- From 2005 to 2007, we located an additional 60 pendix A). Of these, 49 were reported by Schelz taxa from collections at Brigham Young Univer- and others (2006) (including three species from sity (mostly species reported in Welsh and Moore Heil, Fleming, and Porter, with vouchers at San [1965] and Moore and Welsh [1965], but without Juan College, that were not examined for this

Table 3.1. Statistical summary of the ﬂ ora of Canyonlands National Park. Number Number Category conﬁ rmed additionally Total in park reported for park Taxonomic diversity Total taxa 534 60 594 (including varieties and subspecies) Full species 510 55 565 (excluding varieties and subspecies) Families 70 2 72 Life form diversity Tree taxa 15 5 20 Shrub taxa 85 8 93 Perennial forb taxa 238 27 265 Annual forb taxa 99 9 108 Perennial graminoid taxa 72 10 82 Annual graminoid taxa 17 1 18 Fern taxa 8 0 8 Biogeographic diversity Introduced taxa 50 18 68 Native taxa Locally endemic taxa 15 1 16 Regionally endemic taxa 83 8 91 Disjunct taxa 0 0 0 Peripheral taxa 11 4 15 Sparse taxa 4 3 7 Widespread taxa 371 26 397 Total native taxa 484 42 526 The number of taxa and families is based on taxonomic concepts of Welsh and others (2003). See appendices for an explanation of the various categories.

Chapter 3: Results 9 study). The remainder were reported by Ander- forbs and grass-like plants comprise over 21% of son (1983), Dewey and Andersen (2005), Loope the fl ora. Like trees and shrubs, perennial gramin- (1977), and the NCPN (in ed.). Taken together, oids are ecologically important but relatively the number of confi rmed and reported vascular species-poor, representing about 14% of the total plant taxa in Canyonlands NP stands at 594 taxa fl ora. The diversity of ferns and fern-allies in Can- (Table 3.1, Appendix A). yonlands NP is especially low (1.3%) (Table 3.1).

Entering the 2006 fi eld season, we recognized Of the park’s 526 native taxa, 397 (75.5%) are 368 taxa as potentially occurring in Canyonlands widespread across Utah and western North Amer- NP, based on the proximity of the park to known ica. Only 16 species (3% of the park’s total native populations (Albee et al. 1988; Fertig 2006). At fl ora) have extremely limited global distributions least 45 of these species were found to occur in (local endemics found within an area of less than the park based on fi eld collections or relocated 1° of latitude × 2° of longitude; see appendices for herbarium specimens in 2006. Another 162 spe- complete defi nitions of biogeographic catego- cies were eliminated from the potential list based ries). Regional endemics (largely restricted to the on their low probability of occurring in the park. Colorado Plateau) comprise an additional 17.3% Most of these were species of montane conifer- of the native fl ora of the park (Table 3.1). Another ous forest or wetland habitats in the Abajo or La 22 taxa (4.2%) are sparsely distributed across the Sal ranges. The current number of potential spe- state or occur in the Canyonlands NP area at the cies that we are recognizing for Canyonlands Na- periphery of their continuous range. tional Park is 161 (Appendix B).

Of the original pool of species documented or 3.3 Plants of special concern reported for Canyonlands NP, we found that 56 Canyonlands National Park apparently has no taxa were falsely reported based on misidentifi ed plants listed as threatened or endangered under voucher specimens in the CANY and BRY herbar- the U.S. Endangered Species Act, though one ia or literature reports based on collections made threatened species, Navajo sedge (Carex specuico- outside of the park. We consider an additional 44 la), has recently been reported from two wetland species cited (but not vouchered) by Schelz and sites in the Needles and Maze districts (Welsh et others (2006) and the NCPN vegetation mapping al. #2998 and Heil et al. #5916, both at CANY). team (NCPN in ed.) to be questionable based on Both collections were initially identifi ed as other the absence of suitable habitat or the distance of sedge species but were annotated as C. specuicola Canyonlands NP to the known or expected range by Sherel Goodrich, author of the Cyperaceae of each species (Appendix C). treatment of A Utah Flora, third edition. Goodrich now believes that all Utah material previously as- The confi rmed and reported fl ora of Canyonlands cribed to C. specuicola belongs to a more com- National Park represents 16.5% of the 3,594 na- mon species, Parry’s sedge (C. parryana) (Welsh tive and naturalized taxa documented for the state et al. 2003). Critical C. specuicola material from of Utah by Welsh and others (2003) and 47.7% of Canyonlands NP and southeast Utah should be the state’s 151 plant families (Table 3.1). Within evaluated by an independent sedge expert to con- Garfi eld, Grand, San Juan, and Wayne counties, fi rm their identity. the park fl ora captures 29.8% of the 1994 taxa reported by Welsh and others (2003). At the ecore- At least six park taxa were formerly candidates gional scale, Canyonlands NP contains 35.1% of for potential listing (Category 2 or 3C) under the the 1,694 species reported for the Utah portion of act (Table 3.3) (UDWR 1998). Schelz and Moran the Colorado Plateau (Shultz et al. 2006). (2007) cited 18 species known or suspected to occur in Canyonlands NP as NPS species of con- 3.2 Life form and biogeographic cern. Two of these species are now considered falsely reported for the park: Johnson’s cryptanth diversity (Cryptantha johnstonii) and Beck’s spring-parsley Although ecologically dominant, tree and shrub (Cymopterus beckii), Appendix C) due to misiden- species make up only 19% of the confi rmed and tifi cations. Five others on the SEUG list are not reported plant taxa of Canyonlands NP (Table presently known from Canyonlands NP, but are 3.1). Perennial forbs are the most species-rich on the potential species list (Appendix B): Utah life form in the park fl ora, with 265 known and bladder fern (Cystopteris utahensis), Canyonlands reported taxa (44.6% of the total fl ora). Annual lomatium (Lomatium latilobum), Entrada rush-

10 Annotated Checklist of Vascular Flora: Canyonlands National Park pink (Lygodesmia grandifl ora var. entrada), Trot- fat (Baccharis viminea var. atwoodii), and canyon ter’s spring-parsley (Oreoxis trotteri), and alcove spindly goldenbush (Haplopappus scopulorum rock-daisy (Perityle specuicola). Isely’s milkvetch var. canonis). (Astragalus iselyi), Cisco milkvetch (A. sabulosus), and Abajo penstemon (Penstemon lentus var. albi- 3.4 Non-native species and noxious fl orus) are listed as questionable for Canyonlands NP by Schelz and Moran (2007), but are not weeds known or expected to occur in the park due to an Introduced plant species (not historically native absence of likely habitat. to Utah or North America) make up 11.4% of the confi rmed and reported fl ora of Canyonlands NP The Utah Division of Wildlife Resources Con- (Table 3.1). This fi gure is lower than the statewide servation Data Center (UTCDC) tracks 51 spe- average of 13.5% non-native species (Fertig 2007; cies from Canyonlands NP (Table 3.3) as species Welsh et al. 2003). Of the 68 introduced plant spe- of concern based on global and statewide rarity cies confi rmed or reported for Canyonlands NP, (UDWR 1998). Although not currently cited on seven are listed as noxious by the State of Utah any agency list, three recently described narrow (SUDAF 2008) (Table 3.4). Dewey and Ander- endemics of Canyonlands NP and the Colorado sen (2005) mapped the distribution of selected River drainage may warrant attention as species noxious and non-native weeds within the park in of concern due to their apparently limited range 2003–2004, and found all but fi ve-stamen tama- (Welsh et al. 2003): San Rafael prickly-poppy (Ar- risk to occur in small and localized areas. gemone corymbosa var. parva), Atwood’s mule-

Table 3.3. Plant species of conservation concern from Canyonlands National Park. TNC Family Species Common name global Legal status UTCDC status rank Agavaceae Yucca toftiae Toft’s yucca G5T3 USFWS: former 3C Watch, S2S3

Asclepiadaceae Asclepias rusbyi Rusby’s milkweed G4? Add. Data, S1 Asclepiadaceae Sarcostemma climbing milkweed G5T5 Peripheral, S2? cynanchoides var. hartwegii Berberidaceae Berberis fendleri Fendler’s barberry G5 Add. Data, S2 Betulaceae Ostrya knowltonii western hop-hornbeam G3G4 Watch, S2 Cactaceae Echinocereus Glen Canyon hedgehog G5T3? Watch, S1S2 engelmannii var. cactus variegatus* Cactaceae Opuntia basilaris var. beavertail cactus G5T5 Peripheral, S2 basilaris* Cactaceae Opuntia phaeacantha large pricklypear G5T5 Peripheral, S2S3 var. major Chenopodiaceae Krascheninnikovia Ruin Park winterfat G5 T1?Q Rare, S1? lanata var. ruinina Compositae Aster spinosus Mexican devilweed G4G5 Peripheral, S1 (Asteraceae) Compositae Baccharis viminea var. Atwood’s mule-fat G5? TNR (Asteraceae) atwoodii Compositae Cirsium neomexicanum Utah thistle G5T?Q Add. Data, S3? (Asteraceae) var. utahense Compositae Cirsium rothrockii Rothrock’s thistle G3? Add. Data, S1 (Asteraceae)

Chapter 3: Results 11 Table 3.3. Plant species of conservation concern from Canyonlands National Park, cont.

TNC Family Species Common name global Legal status UTCDC status rank Compositae Cirsium rydbergii Rydberg’s thistle G3 USFWS: former 3C, Watch, S2S3 (Asteraceae) SEUG Species of Concern Compositae Grindelia fastigiata erect gumweed G3 Watch, S2 (Asteraceae) Compositae Haplopappus canyon spindly G4TNR (Asteraceae) scopulorum var. canonis goldenbush Compositae Pluchea sericea arrowweed G4G5 Peripheral, S3 (Asteraceae) Compositae Syntrichopappus Fremont’s G5 Peripheral, S2? (Asteraceae) fremontii syntrichopappus Compositae Thelesperma Hopi-tea G5 Peripheral, S1 (Asteraceae) megapotamicum Compositae Xylorhiza glabriuscula Moab woodyaster G4T1T2 Rare, S1S2 (Asteraceae) var. linearifolia Cruciferae Lepidium alyssoides var. Eastwood’s tall G5? T2T4 Add. Data, SR (Brassicaceae) eastwoodiae pepperwort Cucurbitaceae Cucurbita foetidissima buffalo gourd G5 Peripheral, S3? Cyperaceae Cyperus acuminatus taper-tip fl at-sedge G5 Peripheral, S1 Gramineae Andropogon bushy bluestem G5 Peripheral, S2 (Poaceae) glomeratus var. scabriglumis* Gramineae Bothriochloa silver bluestem G5TNR Peripheral, S1 (Poaceae) laguroides var. torreyana* Gramineae Spartina pectinata* prairie cordgrass G5 Add. Data, S? (Poaceae) Hydrophyllaceae Phacelia howelliana Howell’s phacelia G2 SEUG Species of Watch, S2 Concern Leguminosae Astragalus fl exuosus pliant milkvetch G5T5 Peripheral, S1 (Fabaceae) var. fl exuosus* Leguminosae Astragalus Monument milkvetch G4T2T3 Watch, S3? (Fabaceae) monumentalis Leguminosae Astragalus nidularius bird’s-nest milkvetch G3? Watch, S3? (Fabaceae) Leguminosae Astragalus piscator Fisher milkvetch G2G3 SEUG Species of Rare, S2 (Fabaceae) Concern Leguminosae Astragalus praelongus longstipe milkvetch G4T3? Watch, S1S2 (Fabaceae) var. lonchopus* Leguminosae Psoralidium slenderleaf scurfpea G5 T3?Q Tax. Prob., S3? (Fabaceae) lanceolatum var. stenophyllum Leguminosae Psorothamnus Thompson’s indigo- G3?T3? USFWS: former 3C Watch, S3? (Fabaceae) thompsoniae var. bush thompsoniae* Leguminosae Sophora nuttalliana* silky sophora G5 Peripheral, S1 (Fabaceae) Liliaceae Zigadenus vaginatus alcove death camas G2 SEUG Species of Rare, S2 Concern

12 Annotated Checklist of Vascular Flora: Canyonlands National Park Table 3.3. Plant species of conservation concern from Canyonlands National Park, cont.

TNC Family Species Common name global Legal status UTCDC status rank Malvaceae Sphaeralcea janeae Jane’s globemallow G1 UT BLM: Sensitive, Rare, S1 USFWS: former C2, SEUG Species of Concern Malvaceae Sphaeralcea moorei Moore’s globemallow G5T2T3Q Watch, S2S3 Nyctaginaceae Tripterocalyx carneus Wooton’s sand-verbena G4G5T2T4 Peripheral, S2? var. wootonii Orchidaceae Epipactis gigantea giant helleborine G3G4 SEUG Species of Concern Orchidaceae Habenaria zothecina alcove bog-orchid G2 UT BLM: Sensitive, Rare, S2 (Platanthera zothecina) USFWS: former C2, SEUG Species of Concern Papaveraceae Argemone corymbosa San Rafael prickly G3G4 TNR var. parva poppy Polemoniaceae Gilia imperialis (G. Cataract gilia G4T2 UT BLM: Rare, S2 latifolia var. imperialis) Sensitive,SEUG Species of Concern Polypodiaceae Asplenium resiliens ebony spleenwort G5 Peripheral, S1 Polypodiaceae Pellaea glabella var. western cliffbrake G5T4 Peripheral, S2 occidentalis Polypodiaceae Pellaea limitanea border cloak-fern G4G5 Peripheral, S1 Primulaceae Primula specuicola cave primrose G4Q USFWS: former 3C Watch, S3 Rhamnaceae Ceanothus greggii var. Franklin’s desert-lilac G5T1T2 Rare, S1S2 franklinii Rosaceae Potentilla rivalis brook cinquefoil G5 Infrequent, S1S2 Rubiaceae Galium multiﬂ orum Colorado bedstraw G4 Watch, S3? var. coloradoense Saxifragaceae Fendlera rupicola fendlerbush G5 Peripheral, S2S3 (Hydrangeaceae) Solanaceae Chamaesaracha false nightshade G5 Peripheral, S1 coronopus Solanaceae Solanum jamesii James’ potato G3G5 Peripheral, S1 Verbenaceae Phyla cuneifolia wedge-leaf fog-fruit G5 Peripheral, S3? Zygophyllaceae Kallstroemia californica yellow kallstroemia G5 Add. Data, S1 This list is derived from Schelz and Moran (2007) and the Utah Division of Wildlife Resources (1998) for Utah Conservation Data Center (UTCDC) status. Species are all conﬁ rmed as present (with vouchers) unless indicated by an * (reported).

Codes: TNC global rank assesses abundance and conservation priority on a scale of 1–5 (1 being extremely vulnerable and 5 being secure) for full species (G) and varieties or subspecies (T) across their entire range. A “?” indicates uncertainty in the rank, Q = taxonomic questions, U = unknown, and NR = not ranked. A comparable scoring system is used to measure state abundance (S) in Utah under the column UTCDC status. Under legal status, USFWS = U.S. Fish and Wildlife Service. C2 = Category 2 candidate (a former category used for taxa that might warrant being proposed for threatened or endangered status following additional research). 3C = Category 3 candidates (species dropped from consideration for listing). USFS = U.S. Forest Service. UT BLM = U.S. Bureau of Land Management, Utah ofﬁ ce. UTCDC status includes conservation categories adopted by the state natural heritage program to prioritize endemic and rare plant taxa (UDWR 1998). These categories include: Historical (presumed extinct or only known historically through their range), Rare (plants with rangewide viability concerns), Watch (regional endemics without rangewide viability concerns), Peripheral (rare or uncommon in Utah, but more common rangewide), Infrequent (plants occur infrequently over western U.S.), Taxonomic Problems (validity of species, subspecies, or variety has been questioned), Additional Data (species appear rare, but more information needed before being assigned to a different category).

Chapter 3: Results 13 Table 3.4. Noxious weed species of Canyonlands National Park. Family Species Common name Park status Compositae (Asteraceae) Carduus nutans musk thistle Reported Compositae (Asteraceae) Centaurea diffusa diffuse knapweed Present Compositae (Asteraceae) Centaurea repens Russian knapweed Present Compositae (Asteraceae) Cirsium arvense Canada thistle Reported Convolvulaceae Convolvulus arvensis ﬁ eld bindweed Present Cruciferae (Brassicaceae) Lepidium latifolium broad-leaf pepperplant Present Gramineae (Poaceae) Sorghum halepense Johnsongrass Present This list is derived from the State of Utah Department of Agriculture and Food (2008) and Dewey and Andersen (2005).

14 Annotated Checklist of Vascular Flora: Canyonlands National Park the documented fl ora of the park would increase 4 Discussion by 21.3%. Many of the “missing” potential or reported taxa consist of cryptic species that fl ower Since 1964, the known fl ora of Canyonlands Na- late in the growing season or occur in undersam- tional Park has nearly tripled, from 202 to 594 pled wetland or weedy habitats. Confi rmation of taxa, due to continuing fl oristic and vegetation reported species should remain a high priority, as research that has built on the baseline studies of some of these taxa may ultimately prove to be er- Kleiner and Harper (1972), Loope (1977), Moore roneous. Additional studies would also be useful and Welsh (1965), and Welsh and Moore (1965) to better refi ne the abundance and distribution of (Table 3.1, Appendices A and D). Based on avail- species of high management interest (especially able habitats and proximity, at least another 161 rare plants and invasive non-natives). plant taxa may occur within the park (Appendix B). Should all of these potential species be found,

Chapter 4: Discussion 15

Intermountain West, U.S.A. Bronx, N.Y.: 5 Literature Cited New York Botanical Garden. 270 pp. Albee, B. J., L. M. Shultz, and S. Goodrich. 1988. Cronquist, A., A. H. Holmgren, N. H. Holmgren, Atlas of the vascular plants of Utah. +Occa- J. L. Reveal, and P. K. Holmgren, eds. 1977. sional Publ. 7. Salt Lake City: Utah Museum Volume 6, The Monocotyledons. Inter- of Natural History. 670 pp. mountain fl ora: Vascular plants of the In- termountain West, U.S.A. Bronx, N.Y.: New Anderson, J. 1983. Rare plant survey of the York Botanical Garden. 584 pp. Needles Section, Canyonlands National Park. U.S. Fish and Wildlife Service. 24 pp. ——. 1984. Volume 4, Subclass Asteridae (except Asteraceae). Intermountain fl ora: Vascular Baars, D. L. 2003. Geology of Canyonlands Na- plants of the Intermountain West, U.S.A. tional Park. Pages 61–83 in Sprinkel, D. A., Bronx, N.Y.: New York Botanical Garden. T. C. Chidsey, Jr., and P. B. Anderson, eds., 573 pp. Geology of Utah’s parks and monuments. Utah Geological Association Publication 28 Cronquist, A., N. H. Holmgren, and P. K. Hol- (second edition). mgren, eds. 1997. Volume 3, Part A, Subclass Rosidae (except Fabales). Intermountain Barkworth, M. E., K. M. Capels, S. Long, and M. fl ora: Vascular plants of the Intermountain B. Piep, eds. 2003. Magnoliophyta: Com- West, U.S.A. Bronx, N.Y.: New York Botani- melinidae (in part): Poaceae, part 2. Flora of cal Garden. 446 pp. North America north of Mexico, Volume 25. New York: Oxford University Press. 783 pp. Dewey, S., and K. Andersen. 2005. An inventory of invasive non-native plants in Canyonlands Barkworth, M. E., K. M. Capels, S. Long, L. K. National Park (2004): Final report. Utah Anderton, and M. B. Piep, eds. 2007. Mag- State University, Logan Ut. Weed Science noliophyta: Commelinidae (in part): Poace- Research Project Report # SD0513. 40 pp. ae, part 1. Flora of North America north of Mexico, Volume 24. New York: Oxford Dorn, R. D. 2001. Vascular plants of Wyoming. University Press. 911 pp. Third edition. Cheyenne, Wy.: Mountain West Publishing. 412 pp. Barneby, R. C. 1989. Volume 3, Part B. Fabales. In Cronquist, A., A. H. Holmgren, N. H. Evenden, A., M. Miller, M. Beer, E. Nance, S. Holmgren, J. L. Reveal, and P. K. Holmgren, Daw, A. Wight, M. Estenson, and L. Cudlip. eds. Intermountain fl ora: Vascular plants of 2002. Northern Colorado Plateau Vital the Intermountain West, U.S.A. Bronx, N.Y.: Signs Network and Prototype Cluster plan New York Botanical Garden. 279 pp. for natural resources monitoring: Phase I report. October 1. Two volumes. National Boschen, N. 1985. Plant list, Canyonlands Na- Park Service, Northern Colorado Plateau tional Park, Utah. 16 pp. Network, Moab, Ut. 138 pp + app. Chronic, H. 1988. Pages of stone: Geology of Fertig, W. 2006. Herbarium and fi eld studies of western national parks and monuments: vascular plant fl ora of CANY for National Grand Canyon and the Plateau Country. Park Service Inventory and Monitoring Pro- Seattle, Wa.: The Mountaineers. 158 pp. gram. Investigator’s Annual Report, National Cronquist, A. 1994. Volume 5, Asterales. In Park Service. 2 pp. Cronquist, A., A. H. Holmgren, N. H. Hol- ——. 2007. Introduced and naturalized plants of mgren, J. L. Reveal, and P. K. Holmgren, eds. Utah. Sego Lily 30(5):7–11. Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: Flora of North America Editorial Committee. New York Botanical Garden. 496 pp. 1993. Pteridophytes and gymnosperms. Flora of North America north of Mexico, Cronquist, A., A. H. Holmgren, N. H. Holmgren, Volume 2. New York: Oxford University and J. L. Reveal, eds. 1972. Volume 1, Geo- Press. 475 pp. logical and botanical history of the region, its plant geography and a glossary. The ——. 1997. Magnoliophyta: Magnoliidae and vascular cryptogams and the gymnosperms. Hamamelidae. Flora of North America Intermountain fl ora: Vascular plants of the north of Mexico, Volume 3. New York: Ox-

Chapter 5: Literature Cited 17 ford University Press. 590 pp. Loope, W. J. 1977. Relationships of vegetation to environment in Canyonlands National Park. ——. 2000. Magnoliophyta: Alismatidae, Are- Doctoral dissertation, Utah State University, cidae, Commelinidae (in part), and Zin- Logan, Ut. giberidae. Flora of North America north of Mexico, Volume 22. New York: Oxford Margules, C. R., and R. L. Pressey. 2000. Sys- University Press. 352 pp. tematic conservation planning. Nature 405:243–253. ——. 2002a. Magnoliophyta: Commelinidae (in part): Cyperaceae. Flora of North America Moore, G., and S. L. Welsh. 1965. Plants of Can- north of Mexico, Volume 23. New York: yonlands National Park: II. Island in the Sky. Oxford University Press. 608 pp. Proceedings of Utah Academy of Sciences, Arts, and Letters 42(1). ——. 2002b. Magnoliophyta: Liliidae: Liliales and Orchidales. Flora of North America north National Park Service (NPS). 2000. Management of Mexico, Volume 26. New York: Oxford policies. Washington, D.C.: U.S. Department University Press. 723 pp. of Interior, National Park Service. Publica- tion # NPS D1416. 137 pp. ——. 2003. Magnoliophyta: Caryophyllidae, part 1. Flora of North America north of Mexico. Northern Colorado Plateau Network (NCPN). Volume 4. New York: Oxford University in ed. Canyonlands National Park vegetation Press. 559 pp. classifi cation and mapping project. Natural Resource Technical Report. National Park ——. 2005. Magnoliophyta: Caryophyllidae, part Service, Denver, Co. 2. Flora of North America north of Mexico. Volume 5. New York: Oxford University Noss, R. F., and A. Y. Cooperrider. 1994. Saving Press. 656 pp. nature’s legacy: Protecting and restoring biodiversity. Washington, D.C.: Island Press. ——. 2006a. Magnoliophyta: Asteridae, part 6: 416 pp. Asteraceae, part 1. Flora of North America north of Mexico. Volume 19. New York: Schelz, C., and M. Moran. 2007. Southeast Utah Oxford University Press. 579 pp. Group threatened, endangered and species of concern. Southeast Utah Group (Arches ——. 2006b. Magnoliophyta: Asteridae, part 7: NP, Canyonlands NP, Hovenweep NM, & Asteraceae, part 2. Flora of North America Natural Bridges NM), Moab, Ut. 3 pp. north of Mexico. Volume 20. New York: Oxford University Press. 666 pp. Schelz, C., M. Moran, and A. Lafever. 2006. Southeast Utah Group plants. National Park ——. 2006c. Magnoliophyta: Asteridae, part 8: Service Southeast Utah Group (Arches NP, Asteraceae, part 3. Flora of North America Canyonlands NP, Hovenweep NM, & Natu- north of Mexico. Volume 21. New York: ral Bridges NM), Moab, Ut. 16 pp. Oxford University Press. 616 pp. Shultz, L. M., R. D. Ramsey, and W. Lindquist. Fowler, J. F., N. L. Stanton, and R. L. Hartman. 2006. Revised atlas of Utah plants. College 2007. Distribution of hanging garden vegeta- of Natural Resources, Utah State University, tion associations on the Colorado Plateau, Logan, Ut. http://earth.gis.usu.edu/plants. USA. Journal of the Botanical Research Institue of Texas 1(1):585–607. Springer, A. E., L. E. Stevens, and R. Harms. 2006. Inventory and classifi cation of selected Holmgren, N. H., P. K. Holmgren, and A. Cron- National Park Service springs on the Colo- quist. 2005. Volume 2, Part B Subclass Dille- rado Plateau. Northern Arizona University, niidae. Intermountain fl ora: Vascular plants Flagstaff , Az. NPS Cooperative Agreement of the Intermountain West, U.S.A. Bronx, #CA 1200-99-009; Task # NAU-118. N.Y.: New York Botanical Garden. 488 pp. State of Utah Department of Agriculture and Kleiner, E. F., and K. T. Harper. 1972. Envi- Food (SUDAF). 2008. Utah noxious weed ronment and community organization in list. http://ag.utah.gov/divisions/plant/nox- grasslands of Canyonlands National Park. ious/documents/noxUtah.pdf. Ecology 53(2):299–309. Utah Division of Wildlife Resources (UDWR). 1998. Inventory of sensitive species and

18 Annotated Checklist of Vascular Flora: Canyonlands National Park ecosystems in Utah. Endemic and rare plants ——. 1982. Utah plant types—Historical per- of Utah: An overview of their distribution spective, 1840 to 1981—annotated list, and and status. Report prepared for the Utah bibliography. Great Basin Naturalist 42:1–44. Reclamation Mitigation and Conservation Welsh, S. L., N. D. Atwood, S. Goodrich, and L. Commission and U.S. Department of the C. Higgins. 2003. A Utah ﬂ ora. Third edition, Interior. 566 pp + app. revised. Provo, Ut: Brigham Young Univer- Weber, W. A., and R. C. Wittmann. 2001. Colo- sity. 912 pp. rado ﬂ ora: Western Slope. Third edition. Welsh, S. L., and G. Moore. 1965. Plants of Boulder: University Press of Colorado. 488 Canyonlands National Park: I. The Needles pp. region. Proceedings of Utah Academy of Welsh, S. L. 1970. Flora: “Canyonlands.” Natu- Sciences, Arts, and Letters. 42(1). ralist 21:26–37.

Chapter 5: Literature Cited 19

quist 1994; Cronquist et al. 1972, 1977, 1984, 1997; Dorn Appendices 2001; Flora of North America Editorial Committee 1993, 1997, 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c; The following appendices contain annotated checklists of Holmgren et al. 2005; Weber and Wittmann 2001). No ef- the vascular fl ora of Canyonlands National Park. fort has been made to identify additional synonyms from monographs and fl oras preceding the early 1970s. Appendix A lists all species that have been confi rmed for the fl ora with an authenticated voucher specimen (in- Common name (Appendices A, B, C, and D) cluding extant and historical records), or that have been reliably reported for the park in the literature (Anderson Common names are derived primarily from Welsh and 1983; Dewey and Andersen 2005; Kleiner and Harper others (2003) and the Intermountain Flora (Barneby 1989; 1972; Loope 1977; Moore and Welsh 1965; NCPN in ed.; Cronquist 1994; Cronquist et al. 1972, 1977, 1984, 1997; Schelz et al. 2006; Springer et al. 2006; Welsh and Moore Holmgren et al. 2005). 1965). Vouchers were examined from the CANY her- Life form (Appendices A, B, C, and D) barium (housed in the National Park Service’s Southeast Utah Group offi ce in Moab, Utah), and collections from Life form is based on the stature and growth habit of typi- Brigham Young University (BRY), Utah State University cal members of a species at reproductive maturity. Trees (USU), the University of Wyoming (RM), and the digi- are woody perennials with a single stem or trunk and are tal museum database of the New York Botanical Garden typically over 3.5 m tall. Shrubs are woody perennials (NY). Species entries in Appendix A are organized alpha- with one to many trunks arising from near the base of the betically by family and genus. plant and are usually less than 3.5 m tall. Perennial Forbs (PerF) are broad-leaved dicots or monocots with non- Appendix B includes species that may potentially occur woody stems (at least above ground level) that live for mul- within Canyonlands NP, based on the proximity of the park tiple years. Annual Forbs (AnnF) are broad-leaved dicots to other populations or the presence of suitable habitat. or monocots with non-woody stems that complete their life cycle (mature, fl ower, and die) in one year. Perennial Appendix C lists species that have been previously report- Graminoids (PerG) are linear-leaved, grass-like monocots ed for Canyonlands NP but are now considered errone- that live for multiple years. Annual Graminoids (AnnG) ous because they are based on misidentifi ed specimens or are linear-leaved, grass-like monocots that complete their questionable literature records. life cycle (mature, fl ower, and die) in one year. Ferns and fern-allies (Fern) are non-fl owering vascular plants that Appendix D contains the same confi rmed, historical, and reproduce by spores. reported taxa as in Appendix A, but organized by life form rather than by family in order to better meet the needs of Range (Appendices A, B, and C) NPS interpretive specialists. Range represents the distribution of a species within Utah The codes and references for each fi eld in the appropriate and the state’s contribution to the plant’s overall global appendices are summarized below. range. Disjunct (Disj) taxa have their Utah distribution separated from the main, contiguous portion of their range Family (Appendices A, B, C, and D) by a gap of more than 800 km (ca 500 miles). Introduced Family concepts and nomenclature follow A Utah Flora, (Intro) taxa are those that are not native to Utah or North third edition (Welsh et al. 2003). Alternative family names America but which have become naturalized (breeding and treatments from other regional fl oras (Dorn 2001; on their own without human assistance). Local Endemics Flora of North America Editorial Committee 1993, 2002b; (LocEn) are taxa whose entire global range is restricted Holmgren et al. 2005; Weber and Wittman 2001) are listed to an area of less than 16,500 km2 (ca 6,370 mi2, or 1° of in parentheses. latitude × 2° of longitude). Peripheral (Periph) taxa have a widespread global distribution but occur at the margin of Species name (Appendices A, B, C, and D) their contiguous range in Utah and occupy less than 5% Nomenclature for scientifi c names and taxonomic con- of the state’s area (usually only within a few miles of the cepts are derived from Welsh and others (2003). Scientifi c state border). Regional Endemics (RegEn) are taxa with names are listed in alphabetical rather than phylogenetic a global range of 16,500–250,000 km2 (an area about the order. size of the state of Wyoming). Sparse taxa occur widely across Utah or North America but their range within Utah Synonyms/Taxonomic notes (Appendices A, B, and C) is small and patchy, with populations restricted to special- Pertinent synonyms are included for species with alter- ized or uncommon habitats. Widespread (Wide) taxa have 2 native scientifi c names in other recent state and regional global ranges exceeding 250,000 km and occur over at fl oras (Barkworth et al. 2003, 2007; Barneby 1989; Cron- least 10% of the state.

Chapter 6: Appendices 21 Park status (Appendices A, C, and D) Year doc. (Appendix A) Park status identifi es whether a species is Present (Pres), Year documented indicates the fi rst year in which a spe- Historical (Hist), or Reported (Rep) for Canyonlands NP. cies was confi rmed or reported to occur in Canyonlands Species are considered Present if a confi rmed voucher NP. This year does not necessarily correspond with the or documented observation has been made within park date of the voucher specimen cited under Source. “NA” boundaries since 1970. Historical species have been con- indicates that the fi rst year is not known (used mostly for fi rmed with a voucher from the park but have not been “reported” taxa). relocated since before 1970. Reported species are listed for the park in the literature (Anderson 1983; Dewey and Comments (Appendices A, B, C, and D) Andersen 2005; Kleiner and Harper 1972; Loope 1977; This column includes additional comments on the status Moore and Welsh 1965; NCPN in ed.; Schelz et al. 2006; of a particular species in the park, the place of origin for Springer et al. 2006; Welsh and Moore 1965), but have not introduced species, legal status, and other data. been corroborated with a voucher specimen. Species that may potentially occur in Canyonlands NP are listed in Ap- Nativity (Appendix D) pendix B. Rejected taxa (listed in Appendix C) have either Nativity indicates whether a species is native to Utah or been falsely reported (FalsRep) in the literature based Introduced (Intro) based on Fertig (2007). on misidentifi ed herbarium specimens or are considered Questionable (Ques?) because park reports are from well Flower time (Appendix D) outside their known or expected range. Flower time refers to the months in which a species is most Pop. size (Appendix A) likely to be in fl ower. NA indicates a species that is a non- fl owering fern or gymnosperm. Population size refers to the abundance and frequency with which a species occurs within Canyonlands NP. Habitat (Appendix D) Abundance data are derived from Loope (1977), NCPN The distribution of species across generalized habitats in (in ed.), and personal observations. Abundant (Abun) taxa Canyonlands NP is indicated by an “x” in the columns have large populations, have a broad ecological amplitude BlckBr through Wet, and is derived from the Canyonlands (often being dominant in one-to-several vegetation types), NP vegetation classifi cation (NCPN in ed.), Loope (1977), or occur extensively across the park. Common (Com) taxa and Welsh and others (2003). BlckBr = Blackbrush, Salt have large local populations but may be restricted to a sin- = Salt desert scrub dominated by Atriplex spp., SageGrs gle vegetation type or do not occur across the entire park. = Desert grasslands and Basin big sagebrush vegetation Uncommon (Unc) taxa have low to medium-sized popu- types, PinJun = Pinyon-Juniper woodlands/rimrock, Wet lations, typically have a narrow ecological amplitude (or = Wetlands (including hanging gardens). Additional com- at least contribute relatively little to total vegetative cover), ments are included for species that are restricted to a par- or have a small range across the park. Rare taxa usually ticular soil/geologic type or are found mostly in disturbed have low population numbers and are restricted to a single sites or patches of Douglas-fi r alcove forest. location within the park. A “?” indicates that population size is unknown.

Source (Appendix A) Sources are the evidence for the assigned park status. In the case of Present and Historical taxa, sources are herbarium records (with collector, collection number, and repository) that were corroborated during this study. For reported taxa, the source is a published or unpublished reference citing the species within the park. Specimen repositories include the Canyonlands NP herbarium (CANY), Brigham Young University (BRY), Utah State University (USU), New York Botanical Garden (NY), and the University of Wyoming’s Rocky Mountain Herbarium (RM).

22 Annotated Checklist of Vascular Flora: Canyonlands National Park uence tropical America is ca 1 mi N of conﬂ of San Juan & Colorado rivers, San Juan Co., UT (Welsh 11935a BRY). USFWS: 3C is Helper, is Helper, Carbon Co., UT s.n. (Trelease MO) Comments c name with 1987 NA Native to 1983 1987 locality Type 1987 locality Type 1964 1964 1968 1983 1968 Year Year doc. s.n. (CANY) (2006) (CANY) (CANY) (CANY) & Canter 2910 (CANY) & Canter 2919 (CANY) (BRY) (BRY) (BRY) Source Pop. size Park status Range Shrub RegEn Pres Com Moore Welsh, Shrub Wide Pres Rare 8816 Welsh Life form prostrate pigweed AnnF Wide Pres Unc Chrobak-Cox tumble pigweed AnnF Intro Rep ? Schelz et al. sea purslane PerF Wide Pres Unc 22620 Welsh Toft’s yuccaToft’s Shrub LocEn Pres Unc Belnap 200 Harriman’s yuccaHarriman’s Shrub Wide Pres Unc Belnap 101 datil yucca Shrub Wide Pres Unc Moore Welsh, narrowleaved yucca box elder Tree Wide Pres Rare 7027 Welsh bigtooth maple Tree Wide Pres Rare Anderson s.n. Common name Rocky Mountain maple

var. var.

. var. var. & ssp toftiae baccata angustissima interius, Negundo aceroides Interius Synonyms/ notes Taxonomic Includes vars. neomexicanum, douglasii, tripartitum

var.

var. Amaranthus blitoides A. graecizans Amaranthus albus Sesuvium verrucosum Yucca toftiaeYucca angustissima Y. Yucca harrimaniae Yucca harrimaniae Yucca baccataYucca baccata Y. Yucca angustissima Yucca angustissima Y. Acer negundo A. negundo Acer grandidentatum Acer glabrum glabrum vascular plant taxa of Canyonlands National Park, organized by family and scientiﬁ rmed, historical, and reported Amaranthaceae Amaranthaceae Aizoaceae Agavaceae Agavaceae Agavaceae Agavaceae Aceraceae Aceraceae FamilyAceraceae Species name Appendix A. Conﬁ taxonomic and biogeographic notes.

Chapter 6: Appendices 23 24

Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientiﬁ c name with taxonomic and Annotated Checklist ofVascular Flora: CanyonlandsNationalPark biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Amaranthaceae Amaranthus redroot pigweed AnnF Intro Pres Unc Moran s.n. 2007 Native to retrofl exus (CANY) Central America Anacardiaceae Rhus aromatica var. R. trilobata var. squawbush Shrub Wide Pres Com Heil, Porter, & 1964 simplicifolia simplicifolia Fleming 6002 (CANY) Anacardiaceae Rhus aromatica var. R. trilobata var. squawbush Shrub Wide Pres Unc Thorne & 1987 trilobata trilobata Thorne s.n. (BRY) Anacardiaceae Toxicodendron Rhus radicans, T. western poison ivy Shrub Wide Pres Unc Welsh Moore, 1964 rydbergii radicans & Canter 3012 (CANY) Apocynaceae Amsonia jonesii Jones’ amsonia PerF RegEn Rep ? Schelz et al. NA (2006) Apocynaceae Amsonia tomentosa A. tomentosa, A. tomentose PerF Wide Pres Unc Heil & 1979 Type locality var. stenophylla eastwoodiana amsonia Fleming 6089 of A. (CANY) eastwoodiana is Moab vicinity, Grand Co., UT (Rydberg & Garrett 8468 NY) Apocynaceae Apocynum Includes vars. common dogbane PerF Wide Pres Unc Belnap 274 1964 cannabinum cannabinum, (CANY) glaberrimum, & angustifolium Asclepiadaceae Asclepias asperula var. spider milkweed PerF Wide Pres Unc Debolt 382 1964 asperula (CANY) Asclepiadaceae Asclepias labriformis Circle Cliffs PerF RegEn Rep ? Loope (1977) NA Type locality is milkweed Capitol Wash, Wayne Co., UT (Jones 5650 POM) Asclepiadaceae Asclepias latifolia broadleaf PerF Wide Pres Unc Welsh, Moore 1964 milkweed & Olsen 3741 (CANY) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Asclepiadaceae Asclepias Included in A. bigseed milkweed PerF RegEn Pres Unc Heil, Porter, & 1964 macrosperma involucrata by Fleming 5893 some auhors (CANY) Asclepiadaceae Asclepias rusbyi A. engelmanniana Rusby’s milkweed PerF RegEn Pres Rare Welsh, Moore 1964 var. rusbyi & Canter 2930 (CANY) Asclepiadaceae Asclepias ruthiae A. uncialis ssp. Ruth’s milkweed PerF RegEn Pres Rare Cox s.n. 1998 Type locality ruthiae (CANY) is Calf Spring Canyon SE of Castledale, Emery Co., UT (Maguire 18310 UTC) Asclepiadaceae Asclepias speciosa showy milkweed PerF Wide Pres Unc Belnap 403 1987 (CANY) Asclepiadaceae Asclepias A. galioides whorled milkweed PerF Wide Pres Unc Kitchell 253 1982 subverticillata (CANY) Asclepiadaceae Asclepias tuberosa var. A. tuberosa ssp. orange milkweed PerF Wide Pres Unc Welsh 5392 1966 interior terminalis (CANY) Asclepiadaceae Sarcostemma Funastrum climbing milkweed PerF Periph Pres Rare Heil & 1990 cynanchoides var. cynanchoides ssp. Fleming 6072 hartwegii heterophyllum (CANY) Berberidaceae Berberis fendleri Fendler’s barberry Shrub RegEn Pres Unc Fowler & May 1965 1586 (CANY) Berberidaceae Mahonia fremontii Berberis fremontii Fremont’s mahonia Shrub Wide Pres Com Welsh, Moore 1964 & Canter 2960 (CANY) Berberidaceae Mahonia repens Berberis repens Oregon-grape Shrub Wide Pres Rare Topp 2004 ST082804001 Chapter 6:Appendices (CANY) Betulaceae Betula occidentalis B. fontinalis water birch Tree Wide Pres Rare Fowler & May 1983 1556 (RM) Betulaceae Ostrya knowltonii western hop- Tree RegEn Pres Unc Heil, Porter, & 1964 hornbeam Fleming 2961 (CANY) Bignoniaceae Catalpa bignonioides common catalpa Tree Intro Rep ? Schelz et al. NA Native to E (2006) North America 25 26

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Boraginaceae Cryptantha bakeri Oreocarya bakeri Baker’s cryptanth PerF RegEn Pres Rare Anderson s.n. 1983 (BRY) Boraginaceae Cryptantha cinerea Cryptantha San Juan cryptanth PerF RegEn Pres Rare Anderson s.n. 1983 Type locality var. pustulosa pustulosa (BRY) is Elk Mts, San Juan Co., UT (Rydberg & Garrett 9320 NY) Boraginaceae Cryptantha basin yellow PerF Wide Rep ? Loope (1977) NA confertifl ora cryptanth Boraginaceae Cryptantha thick-sepaled AnnF Wide Pres Unc Welsh, Moore 1964 crassisepala var. cryptanth & Canter elachantha 2834 (CANY) Boraginaceae Cryptantha fl ava Oreocarya fl ava plateau yellow PerF Wide Pres Unc Welsh, Moore 1964 cryptanth & Canter 2864 (CANY) Boraginaceae Cryptantha Oreocarya yellow-eye PerF Wide Pres Unc Belnap 210 1987 fl avoculata fl avoculata cryptanth (CANY) Boraginaceae Cryptantha Oreocarya plateau cryptanth PerF RegEn Pres Unc Welsh & 1964 Type locality of fulvocanescens fulvocanescens. Chatterley var. echinoides Includes vars. 21880 (CANY) is Paria Canyon, fulvocanescens & Garfi eld Co., UT echinoides (Jones 5297p POM) Boraginaceae Cryptantha gracilis slender cryptanth AnnF Wide Pres Rare Johnson 18 1979 (CANY) Boraginaceae Cryptantha kelseyana Kelsey’s cryptanth AnnF Wide Pres Rare Moore 233 1964 (BRY) Boraginaceae Cryptantha osterhoutii Oreocarya Osterhout’s PerF RegEn Pres Unc Heil, Fleming, 1964 Type locality osterhoutii cryptanth & Porter 5905 is Monument (CANY) Park [Colorado National Monument], near Grand Junction, Mesa Co., CO (Osterhout 6138 RM) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Boraginaceae Cryptantha pterocarya wing-nut cryptanth AnnF Wide Pres Unc Schelz s.n. 1997 var. pterocarya (CANY) Boraginaceae Cryptantha recurvata recurved cryptanth AnnF Wide Pres Unc Welsh & 1983 Chatterley 21864 (CANY) Boraginaceae Cryptantha tenuis slender cryptanth PerF RegEn Pres Unc Welsh 21806 1979 Type locality (CANY) is Courthouse Wash, Grand Co., UT (Eastwood s.n. CAS) Boraginaceae Heliotropium Euploca bindweed AnnF Wide Pres Rare Belnap 330 1987 convolvulaceum convolvulacea ssp. heliotrope (CANY) californica Boraginaceae Lappula occidentalis L. texana, L. cupseed stickseed AnnF Wide Pres Unc Belnap 245 1967 var. cupulata redowskii var. (CANY) cupulata, L. marginata Boraginaceae Lithospermum incisum showy stoneseed PerF Wide Pres Unc Welsh, Moore 1964 & Canter 2991 (CANY) Boraginaceae Tiquilia latior Coldenia matted tiquilia PerF RegEn Pres Unc Heil & 1983 (Ehretiaceae) hispidissima Fleming 6096 (CANY) Cactaceae Echinocereus E. engelmannii var. Glen Canyon PerF RegEn Rep ? Schelz et al. NA engelmannii var. decumbens. Vars. hedgehog cactus (2006) variegatus not recognized in Flora of North America (2003)

Chapter 6:Appendices Cactaceae Echinocereus Includes var. claretcup PerF Wide Pres Unc Debolt 168 1968 triglochidiatus var. inermis. Vars. not (CANY) melanacanthus recognized in Flora of North America (2003). Cactaceae Opuntia basilaris var. beavertail cactus PerF Periph Rep ? Schelz et al. NA basilaris (2006) 27 28

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Cactaceae Opuntia erinacea var. O. polyacantha var. Utah pricklypear PerF Wide Pres Com Welsh 5396 1966 utahensis hystricina. Var. not (CANY) recognized in Flora of North America (2003) Cactaceae Opuntia phaeacantha Vars. not large pricklypear PerF Wide Pres Unc Welsh 7031 1968 var. major recognized in Flora (CANY) of North America (2003) Cactaceae Opuntia polyacantha Flora of North plains prickly-pear PerF Wide Pres Unc Topp 2004 var. polyacantha America (2003) ST052204001 includes vars. (CANY) rufi spina & trichophora Cactaceae Sclerocactus whipplei S. parvifl orus, small-fl ower PerF Wide Pres Unc Welsh & 1968 Type locality of var. roseus S. whipplei var. fi shhook cactus Chatterley S. parvifl orus is intermedius. 21879 (CANY) Glen Canyon, Includes S. San Juan Co., contortus & S. UT (Clover & terrae-canyonae Jotter 2398 US) Capparaceae Cleome lutea yellow beeplant AnnF Wide Pres Unc Welsh, Moore 1964 (Cleomaceae) & Olsen 3711 (CANY) Capparaceae Cleome serrulata Rocky Mountain AnnF Wide Pres Unc Chrobak 407 1985 (Cleomaceae) beeplant (CANY) Caprifoliaceae Symphoricarpos long-fl ower Shrub Wide Pres Unc Heil & 1964 longifl orus snowberry Fleming 6088 (CANY) Caryophyllaceae Arenaria fendleri var. (Alsinaceae) eastwoodiae Eremogone Eastwood’s PerF Wide Pres Unc Welsh, Moore 1964 eastwoodiae var. sandwort & Canter eastwoodiae, A. 2902 (CANY) eastwoodiae Caryophyllaceae Silene antirrhina sleepy catchfl y AnnF Wide Pres Unc Heil, Fleming, 1964 & Porter 5975 (CANY) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Celastraceae Pachystima myrsinites Paxistima mountain-lover Shrub Wide Pres Rare Anderson s.n. 1983 myrsinites, (BRY) Pachistima myrsinites Chenopodiaceae Atriplex canescens var. A. canescens var. fourwing saltbush Shrub Wide Pres Com Welsh, Moore 1964 canescens occidentalis & Olsen 3688 (CANY) Chenopodiaceae Atriplex confertifolia shadscale Shrub Wide Pres Unc Welsh, Moore 1964 & Canter 2952 (CANY) Chenopodiaceae Atriplex dioica A. subspicata, thickleaf orache AnnF Wide Pres Rare Rink 4606 2005 Originally A. patula var. (CANY) reported as subspicata Endolepis dioica, a species not known from UT. Chenopodiaceae Atriplex gardneri var. A. cuneata Castle Valley Shrub RegEn Pres Unc Welsh 21805 1983 Type locality cuneata saltbush (CANY) is near Emery, Emery Co., UT (Jones 5443 RM) Chenopodiaceae Atriplex garrettii var. Garrett’s saltbush Shrub LocEn Pres Unc Belnap 517 1988 Type locality garrettii (CANY) is vicinity of Moab, Grand Co., UT (Rydberg & Garrett 8465 NY) Chenopodiaceae Atriplex heterosperma A. micrantha two-seed orache AnnF Intro Hist Rare Wiens s.n. 1965 (NY)

Chapter 6:Appendices Chenopodiaceae Atriplex rosea tumbling orach AnnF Intro Pres Unc Welsh, Moore 1964 Native to & Olsen 3742 Eurasia (CANY) Chenopodiaceae Bassia americana Kochia americana greenmolly PerF Wide Pres Unc Belnap 186 1987 (CANY) Chenopodiaceae Bassia hyssopifolia ﬁ ve-horn AnnF Intro Pres Unc Welsh, Moore 1964 Native to smotherweed & Olsen 3745 Eurasia (CANY) 29 30

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Chenopodiaceae Bassia scoparia Kochia scoparia, B. summer-cypress AnnF Intro Pres Unc Belnap 141 1987 Native to sieversiana (CANY) Eurasia Chenopodiaceae Chenopodium album C. album lambsquarter AnnF Intro Pres Unc Belnap 328 1978 Native to var. album (CANY) Eurasia Chenopodiaceae Chenopodium C. fremontii var. mountain AnnF Wide Pres Unc Welsh, Moore 1964 atrovirens atrovirens goosefoot & Olsen 3714 (CANY) Chenopodiaceae Chenopodium C. fremontii Fremont’s AnnF Wide Pres Unc Welsh, Moore 1964 fremontii var. goosefoot & Olsen 3743 fremontii (CANY) Chenopodiaceae Chenopodium narrow-leaf AnnF Wide Pres Unc Welsh, Moore 1964 leptophyllum goosefoot & Olsen 3764 (CANY) Chenopodiaceae Corispermum Includes vars. American bugseed AnnF Wide Pres Unc Welsh, Moore 1964 americanum americanum & & Olsen 3763 rydbergii in Flora (CANY) of North America (2003) Chenopodiaceae Cycloloma C. atriplicifolium winged pigweed AnnF Wide Pres Unc Belnap 433 1987 atriplicifolia (CANY) Chenopodiaceae Grayia spinosa Atriplex spinosa, A. spiny hopsage Shrub Wide Pres Unc Topp, S. 2007 grayi personal observation Chenopodiaceae Halogeton glomeratus halogeton AnnF Intro Pres Unc Welsh 22607 1978 Native to (CANY) Eurasia Chenopodiaceae Krascheninnikovia Ceratoides winterfat Shrub Wide Pres Unc Welsh, Moore 1964 lanata var. lanata lanata, Eurotia & Olsen 3730 lanata. Vars. not (CANY) recognized in Flora of North America (2003) Chenopodiaceae Krascheninnikovia lanata Ceratoides lanata Ruin Park Shrub LocEn Pres Unc Allen 106 1978 Type locality is var. ruinina var. ruinina. Vars. winterfat (CANY) Ruin Park, San not recognized Juan Co., UT in Flora of North (Welsh & Welsh America (2003). 22377 BRY). Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Chenopodiaceae Monolepis nuttalliana Nuttall’s poverty- AnnF Wide Pres Unc Schelz s.n. 2001 weed (CANY) Chenopodiaceae Salsola paulsenii barbwire Russian AnnF Intro Pres Unc Welsh 22629 1983 Native to thistle (CANY) Eurasia Chenopodiaceae Salsola tragus S. kali, S. iberica, S. Russian thistle AnnF Intro Pres Unc Allen 105 1967 Native to Asia pestifer, S. australis (CANY) Chenopodiaceae Sarcobatus greasewood Shrub Wide Pres Unc Debolt 237 1983 (Sarcobataceae) vermiculatus (CANY) Chenopodiaceae Suaeda torreyana var. S. moquinii, S. Torrey’s seepweed AnnF Wide Pres Unc Welsh, Moore 1964 torreyana nigra, S. fruticosa & Olsen 3685 (CANY) Chenopodiaceae Zuckia brandegeei var. Atriplex brandegei Brandegee’s Shrub RegEn Pres Rare Topp 1972 Type locality is brandegeei siltbush ST092604003 San Juan River, (CANY) San Juan Co., UT (Brandegee s.n. GH) Compositae Ambrosia Franseria bur ragweed AnnF Wide Pres Unc Welsh, Moore 1964 (Asteraceae) acanthicarpa acanthicarpa & Olsen 3681 (CANY) Compositae Ambrosia tomentosa Franseria skeleton-leaf PerF Wide Pres Rare Welsh 22645 1983 (Asteraceae) tomentosa bursage (CANY) Compositae Antennaria marginata Includes records of sandstone PerF Wide Pres Rare Anderson s.n. 1983 (Asteraceae) A. neglecta from pussytoes (BRY) UT Compositae Arctium minus burdock PerF Intro Pres Rare Moran s.n. 2004 Native to (Asteraceae) (CANY) Eurasia Compositae Artemisia bigelovii Bigelow’s Shrub Wide Pres Com Welsh 22606 1964 (Asteraceae) sagebrush (CANY)

Chapter 6:Appendices Compositae Artemisia campestris Oligosporus (Asteraceae) var. scouleriana sand wormwood PerF Wide Pres Unc Belnap 282 1964 campestris ssp. (CANY) pacifi cus, O. pacifi cus, A. campestris ssp. pacifi ca 31 32

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Artemisia dracunculus Oligosporus tarragon PerF Wide Pres Unc Welsh, Moore 1964 (Asteraceae) var. glauca dracunculus, Var. & Olsen 3676 not recognized in (CANY) Cronquist (1994) & Flora of North America (2006) Compositae Artemisia fi lifolia Oligosporus sand sagebrush Shrub Wide Pres Unc Welsh, Moore 1964 (Asteraceae) fi lifolius & Olsen 3776 (CANY) Compositae Artemisia frigida fringed sagebrush Shrub Wide Pres Unc Hahn 1987 (Asteraceae) LAH09140309 (CANY) Compositae Artemisia ludoviciana Louisiana PerF Wide Pres Unc Welsh, Moore 1964 (Asteraceae) var. albula wormwood & Olsen 3694 (CANY) Compositae Artemisia ludoviciana Louisiana PerF Wide Pres Rare Welsh, Moore 1964 CANY material (Asteraceae) var. ludoviciana wormwood & Olson 3694 intergrades (BRY) with var. albula. Compositae Artemisia ludoviciana Louisiana PerF Wide Pres Unc Welsh, Moore 1964 (Asteraceae) var. mexicana wormwood & Olsen 3713 (CANY) Compositae Artemisia nova var. A. arbuscula var. black sagebrush Shrub Wide Rep ? Schelz et al. NA (Asteraceae) nova nova, Seriphidium (2006) novum. Vars. not recognized in Flora of North America (2006) Compositae Artemisia spinescens Picrothamnus budsage Shrub Wide Pres Unc Chrobak 428 1973 (Asteraceae) desertorum (CANY) Compositae Artemisia tridentata Seriphidium basin big Shrub Wide Pres Com Welsh, Moore (Asteraceae) var. tridentata tridentatum ssp. sagebrush 1964 tridentatum & Canter 2958 (CANY) Compositae Aster ascendens A. chilensis ssp. Pacifi c aster PerF Wide Pres Unc Welsh, Moore 1964 (Asteraceae) adscendens, & Olsen 3751 Symphyotrichum (CANY) ascendens, Virgulaster ascendens Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Aster frondosus Symphyotrichum leafy aster AnnF Wide Pres Unc Welsh, Moore 1964 (Asteraceae) frondosum, & Olsen 3787 Brachyactis (CANY) frondosa Compositae Aster glaucodes Herrickia glauca blueleaf aster PerF Wide Pres Unc Topp 2003 (Asteraceae) var. glauca, Eurybia SET09260302 glauca, Eucephalus (CANY) glaucus Compositae Aster pansus A. ericoides white heath aster PerF Wide Pres Unc Welsh, Moore 1964 (Asteraceae) var. pansus, & Olsen 3770 Symphyotrichum (CANY) ericoides var. pansum, Virgulus ericoides Compositae Aster spathulatus A.occidentalis, western aster PerF Wide Pres Rare Schelz s.n. 1997 (Asteraceae) Symphyotrichum (CANY) spathulatum Compositae Aster spinosus Chloracantha Mexican devilweed PerF Wide Pres Unc Belnap 355 1983 (Asteraceae) spinosa var. spinosa (CANY) Compositae Baccharis salicina Includes B. emoryi Rio Grande Shrub Wide Pres Unc Welsh 22613 1983 (Asteraceae) seepwillow (CANY) Compositae Baccharis viminea var. Included in B. Atwood’s mule-fat Shrub LocEn Pres Unc Belnap 225 1987 Type locality is (Asteraceae) atwoodii salicifolia in Flora (BRY) 1 mi E of San of North America Juan River, (2006) San Juan Co., UT (Atwood et al. 22406 BRY). Duplicate at CANY is B. salicina. Chapter 6:Appendices Compositae Bidens cernua nodding bur- PerF Wide Pres Rare Rink 4607 2005 Originally (Asteraceae) marigold (CANY) reported as Bidens tripartita. Compositae Brickellia californica California Shrub Wide Pres Unc Welsh, Moore 1964 (Asteraceae) brickellbush & Olsen 3686 (CANY) 33 34

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Brickellia longifolia B. longifolia var. longleaf Shrub Wide Pres Unc Welsh 22628 1964 (Asteraceae) longifolia brickellbush (CANY) Compositae Brickellia microphylla rough brickellbush Shrub Wide Pres Unc Welsh, Moore 1964 (Asteraceae) var. scabra & Olsen 3692 (CANY) Compositae Brickellia oblongifolia Mohave Shrub Wide Pres Unc Belnap 208 1987 Type locality of (Asteraceae) var. linifolia brickellbush (CANY) var. linifolia is American Fork, Jordan Valley, Utah Co., UT (Watson 493 YU) Compositae Carduus nutans Includes ssp. musk thistle PerF Intro Rep Rare Dewey & 2004 Utah state (Asteraceae) leiophyllus & Andersen noxious weed. macrolepis (2005) Native to Eurasia Compositae Centaurea diffusa Acosta diffusa diffuse knapweed PerF Intro Pres Rare Andersen 2003 Utah state (Asteraceae) CANY V-1 noxious weed. (CANY) Native to Mediterranean Compositae Centaurea repens Acroptilon repens Russian knapweed PerF Intro Pres Unc Chrobak 449 1985 UT state (Asteraceae) (CANY) noxious weed. Native to Eurasia Compositae Chaenactis stevioides stevia dusty- AnnF Wide Pres Unc Allen 219 1964 (Asteraceae) maiden (CANY) Compositae Chaetopappa Leucelene rose-heath PerF Wide Pres Unc Debolt 222 1967 (Asteraceae) ericoides ericoides, Aster (CANY) arenosus Compositae Chrysopsis villosa var. C. villosa hispid golden-aster PerF Wide Pres Com Thorne, 1964 (Asteraceae) minor var. hispida, Thomas, & Heterotheca villosa Inskip 5160 var. hispida, H. (CANY) villosa var. minor Compositae Chrysopsis villosa var. H. villosa var. (Asteraceae) villosa hairy golden-aster PerF Sparse Rep ? Schelz et al. NA villosa (2006) Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Chrysothamnus Lorandersonia spear-leaf Shrub Wide Pres Unc Welsh, 1964 (Asteraceae) linifolius linifolia rabbitbrush Moore, & Olsen 3766 (CANY) Compositae Chrysothamnus Ericameria greenish Shrub Wide Pres Com Fertig 23013 2006 (Asteraceae) nauseosus var. nauseosa var. rabbitbrush (CANY) consimilis consimilis, E. nauseosa var. oreophila, C. nauseosus var. oreophilus Compositae Chrysothamnus C. nauseosus graystem Shrub Wide Pres Com Fertig 23015 2006 (Asteraceae) nauseosus var. var. hololeucus, rabbitbrush (CANY) gnaphalodes Ericameria nauseosa var. hololeuca Compositae Chrysothamnus Ericameria glabrate Shrub Wide Pres Unc Welsh & 1964 (Asteraceae) nauseosus var. nauseosa var. rabbitbrush Moore 3847 graveolens graveolens, E. (CANY) nauseosa var. glabrata Compositae Chrysothamnus Ericameria rush rabbitbrush Shrub RegEn Pres Com Welsh & 1964 (Asteraceae) nauseosus var. junceus nauseosa var. Moore 3848 juncea (CANY) Compositae Chrysothamnus Ericameria smoothseed (Asteraceae) nauseosus var. nauseosa var. rabbitbrush Shrub RegEn Pres Unc Hahn 2003 Type locality leiospermus leiosperma LAH09280302 is St. George, (CANY) Washington Co., UT (Palmer s.n. NY) Chapter 6:Appendices Compositae Chrysothamnus Included in var. slenderleaf Shrub Wide Pres Com Welsh, Moore 1964 (Asteraceae) viscidifl orus var. viscidifl orus in rabbitbrush & Olsen 3712 stenophyllus Cronquist (1994) (CANY) Compositae Chrysothamnus green rabbitbrush Shrub Wide Rep Unc NCPN (in ed.) 2004 (Asteraceae) viscidifl orus var. viscidifl orus 35 36

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Cirsium arvense Includes vars. Canada thistle PerF Intro Rep Rare Dewey & 2004 Utah state (Asteraceae) horridum & Andersen noxious weed. integrifolium, (2005) Native to Breea arvensis Eurasia Compositae Cirsium calcareum var. C. arizonicum var. wingless thistle PerF RegEn Pres Unc Moore 214 1964 (Asteraceae) bipinnatum bipinnatum. Var. (BRY) not recognized in Cronquist (1994) Compositae Cirsium calcareum var. Var. not recognized pretty thistle PerF Wide Pres Unc Belnap 256 1987 (Asteraceae) pulchellum in Cronquist (CANY) (1994), included in C. arizonicum var. bipinnatum in Flora of North America 2006) Compositae Cirsium Vars. not New Mexico thistle PerF Wide Pres Unc Welsh, Moore 1964 (Asteraceae) neomexicanum var. recognized in Flora & Canter neomexicanum of North America 2867 (BRY) (2006) Compositae Cirsium C. utahense. Not Utah thistle PerF RegEn Pres Unc Welsh, Moore 1964 Type locality (Asteraceae) neomexicanum var. recognized in Flora & Canter is Silver Reef, utahense of North America 2867 (CANY) Washington (2006). Co., UT (Jones 5163q GH) Compositae Cirsium rothrockii C. arizonicum var. Rothrock’s thistle PerF RegEn Pres Rare Chrobak 448 1985 (Asteraceae) rothrockii. Not (CANY) recognized for Utah in Flora of North America (2006) Compositae Cirsium rydbergii Rydberg’s thistle PerF RegEn Pres Unc Fowler & May (Asteraceae) 1539 (CANY)1993 Type locality is along San Juan River near Bluff, San Juan Co., UT (Rydberg & Garrett 10001 NY). USFWS: 3C Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Cirsium vulgare bull thistle PerF Intro Rep Rare Dewey & 2004 Native to (Asteraceae) Andersen Europe (2005) Compositae Conyza canadensis var. Vars. not Canadian AnnF Wide Pres Unc Welsh, Moore 1964 (Asteraceae) glabrata recognized in Flora horseweed & Olsen 3708 of North America (CANY) (2006) Compositae Crepis runcinata var. glaucous PerF Wide Pres Unc Fleming 863 1964 (Asteraceae) glauca hawksbeard (CANY) Compositae Dicoria brandegeei D. canescens ssp. Brandegee’s AnnF RegEn Pres Unc Rink 4605 2005 (Asteraceae) brandegei, D. sandplant (CANY) canescens ssp. wetherillii. Vars. not recognized in Flora of North America (2006) Compositae Encelia frutescens var. E. frutescens var. bush encelia Shrub Wide Pres Unc Fleming 805 1989 (Asteraceae) resinosa frutescens (CANY) Compositae Enceliopsis nudicaulis nakedstem PerF Wide Pres Unc Johnson 14 1979 Type locality (Asteraceae) var. nudicaulis (CANY) is “Southern Utah” (Bishop s.n. GH) Compositae Erigeron bellidiastrum E. bellidiastrum var. pretty daisy AnnF Wide Pres Unc Schelz s.n. 1983 (Asteraceae) bellidiastrum (CANY) Compositae Erigeron divergens spreading daisy PerF Wide Pres Com Allen 220 1978 (Asteraceae) var. divergens (CANY) Compositae Erigeron ﬂ agellaris trailing daisy PerF Wide Pres Rare Anderson s.n. 1983 (Asteraceae) (BRY) Compositae Chapter 6:Appendices Erigeron pumilus var. E. concinnus var. Navajo ﬂ eabane PerF Wide Pres Unc Welsh, Moore 1964 (Asteraceae) concinnus concinnus, includes & Canter var. subglaber 2920 (CANY) Compositae Erigeron utahensis var. E. sparsifolius slenderleaf daisy PerF RegEn Pres Unc Heil & 1987 Type locality is (Asteraceae) sparsifolius Fleming 6099 Willow Creek, (CANY) San Juan Co., UT (Eastwood 48 CAS) 37 38

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Erigeron utahensis var. E. utahensis Utah daisy PerF RegEn Pres Unc Belnap 117 1987 Type locality (Asteraceae) utahensis (CANY) is Kanab, Kane Co., UT (Thompson s.n. GH) Compositae Euthamia occidentalis Solidago western goldenrod PerF Wide Pres Unc Belnap 344 1987 (Asteraceae) occidentalis (CANY) Compositae Gaillardia pinnatifi da Hopi PerF Wide Pres Unc Allen 212 1973 (Asteraceae) blanketfl ower (CANY) Compositae Gaillardia spathulata basin PerF RegEn Pres Unc Chrobak 423 1972 Type locality is (Asteraceae) blanketfl ower (CANY) Rabbit Valley, Wayne Co., UT (Ward 401 GH) Compositae Gnaphalium G. chilense, cottonbatting AnnF Wide Pres Rare Schelz s.n. 2002 (Asteraceae) stramineum Pseudognaphalium cudweed (CANY) stramineum Compositae Grindelia fastigiata Included in G. erect gumweed PerF RegEn Pres Unc Heil & 1990 (Asteraceae) hirsutula in Flora Fleming 6082 of North America (CANY) (2006) Compositae Grindelia squarrosa Vars. not curly-cup PerF Wide Pres Unc Welsh 22618 1983 (Asteraceae) var. serrulata recognized in Flora gumweed (CANY) of North America (2006) Compositae Gutierrezia thread snakeweed Shrub Wide Pres Unc Welsh, Moore 1964 (Asteraceae) microcephala & Olsen 3693 (CANY) Compositae Gutierrezia sarothrae broom snakeweed Shrub Wide Pres Com Welsh & 1964 (Asteraceae) Moore 3836 (CANY) Compositae Haplopappus Stenotus (Asteraceae) armerioides var. armerioides thrift goldenweed PerF Wide Pres Unc Heil, Porter, & 1964 armerioides var. Fleming 5926 armerioides (CANY) Compositae Haplopappus rusbyi H. drummondii, Rusby’s Shrub Wide Rep ? Schelz et al. NA (Asteraceae) Isocoma rusbyi, I. goldenbush (2006) drummondii Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Haplopappus Vars. not canyon spindly Shrub LocEn Pres Rare Welsh 8813 1968 Type locality is (Asteraceae) scopulorum var. recognized in Flora goldenbush (BRY) Naturalist Cove, canonis of North America E of The Neck, (2006). Canyonlands NP, San Juan Co., UT (Welsh 8813 BRY) Compositae Haplopappus Hesperodoria spindly goldenbush Shrub RegEn Pres Unc Heil & 1990 Type locality is (Asteraceae) scopulorum var. scopulorum, Fleming 6188 Cedar Canyon, hirtellus Chrysothamnus (CANY) Iron Co., UT scopulorum (Garrett 6051 US) Compositae Helenium autumnale Vars. not fall sneezeweed PerF Wide Pres Unc Chrobak 871 1985 (Asteraceae) var. montanum recognized in Flora (CANY) of North America (2006). Compositae Helianthella purpledisk PerF RegEn Pres Rare Topp 2004 Type locality (Asteraceae) microcephala sunfl ower ST092304004 is Abajo Mtns, (CANY) San Juan Co., UT (Newberry s.n. GH) Compositae Helianthus annuus Ssp. not recognized common sunfl ower AnnF Wide Pres Unc Welsh 22481 1983 (Asteraceae) ssp. lenticularis in Flora of North (CANY) America (2006). Compositae Helianthus petiolaris Includes var. prairie sunfl ower AnnF Wide Pres Com Chrobak 444 1964 (Asteraceae) ssp. fallax petiolaris (CANY) Compositae Hymenopappus H. fi lifolius var. (Asteraceae) fi lifolius var. cinereus lugens, H. fi lifolius common PerF Wide Pres Com Debolt 224 1964 var. paucifl orus, hyalineherb (CANY) H. fi lifolius var. Chapter 6:Appendices

megacephalus, H. ﬁ lifolius var. tomentosus Compositae Hymenoxys acaulis var. H. argentea Canyonlands PerF Wide Pres Unc Welsh, Moore 1964 (Asteraceae) ivesiana var. ivesiana, hymenoxys & Canter Tetraneuris 2879 (CANY) ivesiana 39 40

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Hymenoxys Picardenia Colorado rubber- PerF Wide Pres Unc Schelz s.n. 1997 (Asteraceae) richardsonii var. richardsonii var. plant (CANY) fl oribunda fl oribunda Compositae Hymenoxys Picardenia Colorado rubber- PerF Wide Pres Unc Schelz s.n. 2003 (Asteraceae) richardsonii var. richardsonii var. plant (CANY) richardsonii richardsonii Compositae Iva axillaris I. axillaris var. poverty-weed PerF Wide Pres Unc Topp 1988 (Asteraceae) axillaris, Vars. not ST092404001 recognized in Flora (CANY) of North America (2006). Compositae Lactuca serriola prickly lettuce AnnF Intro Pres Unc Fertig 23011 2006 Native to (Asteraceae) (CANY) Europe Compositae Lactuca tatarica var. L. pulchella, L. blue lettuce PerF Wide Pres Unc Belnap 138 1987 (Asteraceae) pulchella oblongifolia, (CANY) Mulgedium pulchellum Compositae Lygodesmia L. grandifl ora showy rush-pink PerF Wide Pres Unc Johnson 61 1979 (Asteraceae) grandifl ora var. (CANY) grandifl ora Compositae Machaeranthera M. linearis, aristate aster PerF RegEn Pres Unc Welsh, Moore 1964 Type locality is (Asteraceae) canescens var. aristata M. rigida, M. & Olsen 3680 Willow Creek, canescens var. (CANY) San Juan Co., vacans, Dieteria UT (Eastwood canescens var. 45 CAS) aristata Compositae Machaeranthera Aster canescens hoary aster PerF Wide Rep ? Schelz et al. NA (Asteraceae) canescens var. (2006) canescens Compositae Machaeranthera Haplopappus gumweed aster PerF Wide Pres Unc Welsh, Moore 1964 (Asteraceae) grindelioides var. nuttallii, Xanthisma & Canter grindelioides grindelioides var. 2900 (CANY) grindelioides Compositae Machaeranthera tansyleaf aster AnnF Wide Pres Unc Belnap 363 1987 (Asteraceae) tanacetifolia (CANY) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Malacothrix glabrata Malacothrix fi liform desert- AnnF Wide Pres Rare Schelz s.n. 2001 (Asteraceae) californica var. dandelion (CANY) glabrata Compositae Malacothrix sowthistle desert- AnnF Wide Pres Unc Belnap 37 1964 (Asteraceae) sonchoides dandelion (CANY) Compositae Oxytenia acerosa Iva acerosa copperweed Shrub Wide Pres Unc Welsh & 1964 (Asteraceae) Moore 3850 (CANY) Compositae Petradoria pumila var. Solidago rock-goldenrod PerF Wide Pres Unc Welsh, Moore 1964 (Asteraceae) pumila petradoria & Olsen 3728 (CANY) Compositae Platyschkuhria Bahia nudicaulis desert bahia PerF RegEn Pres Unc Belnap 373 1986 Type locality is (Asteraceae) integrifolia var. var. desertorum (CANY) Cisco, Grand desertorum Co., UT (Jones s.n. US) Compositae Pluchea sericea Tessaria sericea arrowweed Shrub Wide Pres Unc Heil & 1990 (Asteraceae) Fleming 6074 (CANY) Compositae Prenanthella exigua Lygodesmia exigua prenanthella AnnF Wide Pres Rare Belnap 129 1987 (Asteraceae) (CANY) Compositae Ratibida columnifera prairie conefl ower PerF Sparse Pres Unc Fertig 23014 2006 (Asteraceae) (CANY) Compositae Senecio douglasii var. S. longilobus, Douglas’ groundsel PerF Wide Rep ? Schelz et al. NA (Asteraceae) longilobus S. fl accidus var. (2006) douglasii Compositae Senecio multilobatus Packera basin groundsel PerF Wide Pres Com Welsh, Moore 1964 (Asteraceae) multilobata & Canter 2853 (CANY)

Chapter 6:Appendices Compositae Senecio spartioides S. multicapitatus. (Asteraceae) var. multicapitatus Vars. not many-headed PerF Wide Pres Unc Boschen 22- 1984 recognized in Flora groundsel 84 (CANY)

of North America (2006) Compositae Senecio spartioides S. spartioides. Vars. broom groundsel PerF Wide Pres Unc Welsh & 1964 (Asteraceae) var. spartioides not recognized Moore 3678 in Flora of North (CANY) America (2006) 41 42

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Solidago canadensis S. lepida var. Canadian PerF Wide Pres Unc Topp 2004 (Asteraceae) var. salebrosa salebrosa, S. goldenrod ST090504001 altissima (CANY) Compositae Solidago velutina S. sparsifl ora, three-nerve PerF Wide Pres Com Welsh & 1964 (Asteraceae) S. velutina var. goldenrod Moore 3845 sparsifl ora (CANY) Compositae Sonchus asper spiny-leaf sow- AnnF Intro Pres Unc Heil, Porter, & 1990 Native to (Asteraceae) thistle Fleming 5908 Europe (CANY) Compositae Stephanomeria exigua white-plume wire- AnnF Wide Pres Com Welsh, Moore 1964 (Asteraceae) lettuce & Olsen 3709 (CANY) Compositae Stephanomeria brown-plume wire- PerF Wide Pres Unc Boschen 1984 (Asteraceae) paucifl ora lettuce 31017 (CANY) Compositae Stephanomeria S. minor var. minor slender wirelettuce PerF Wide Pres Unc Welsh, Moore 1964 (Asteraceae) tenuifolia var. & Olsen 3724 tenuifolia (CANY) Compositae Syntrichopappus Fremont’s AnnF Periph Pres Rare Belnap 379 1987 (Asteraceae) fremontii syntrichopappus (BRY) Compositae Taraxacum offi cinale common dandelion PerF Intro Pres Unc Belnap 112 1987 Native to (Asteraceae) (CANY) Eurasia Compositae Tetradymia canescens spineless Shrub Wide Rep ? Schelz et al. NA (Asteraceae) horsebrush (2006) Compositae Thelesperma Hopi-tea PerF Periph Pres Rare Schelz s.n. 2003 (Asteraceae) megapotamicum (CANY) Compositae Thelesperma Vars. not scapose PerF Wide Pres Unc Welsh & 1973 Type locality (Asteraceae) subnudum var. recognized in Flora greenthread Chatterley is “Southern subnudum of North America 21867 (CANY) Utah” (Parry (2006). 109 GH) Compositae Thymophylla Dyssodia scale glandweed PerF Wide Pres Unc Heil, Porter, & 1990 (Asteraceae) pentachaeta var. pentachaeta var. Fleming 6103 belenidium belenidium (CANY) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Townsendia annua annual townsendia AnnF Wide Pres Unc Debolt 220 1982 Type locality is (Asteraceae) (CANY) Cottonwood Wash, 1 ½ miles N of Bluff, San Juan Co., UT (Maguire 13509 GH) Compositae Townsendia incana hoary townsendia PerF Wide Pres Unc Heil, Porter, & 1978 (Asteraceae) Fleming 5936 (CANY) Compositae Tragopogon dubius T. dubius var. major yellow salsify PerF Intro Pres Unc Chrobak 460 1985 Native to (Asteraceae) (CANY) Europe Compositae Vanclevea stylosa Chrysothamnus resinbush Shrub RegEn Pres Unc Chrobak 446 1985 Type locality (Asteraceae) stylosa (CANY) is near Epsom Creek [between Bluff and Mexican Hat], San Juan Co., UT (Eastwood 36 CAS) Compositae Verbesina encelioides Ximenesia golden AnnF Wide Pres Unc Chrobak-Cox 1988 (Asteraceae) var. exauriculata encelioides crownbeard s.n. (CANY) Compositae Xanthium strumarium X. pensylvanicum, rough cocklebur AnnF Wide Pres Unc Welsh, Moore 1964 (Asteraceae) var. canadense X. saccharatum, & Olsen 3698 Vars. not (CANY) recognized in Flora of North America (2006). Compositae Xylorhiza glabriuscula Machaeranthera Moab woodyaster PerF LocEn Pres Unc Belnap 166 1987 Type locality is (Asteraceae) var. linearifolia linearifolia, (CANY) 6 miles NW of Chapter 6:Appendices Xylorhiza Moab, Grand linearifolia Co., UT (Watson 679 TEX) 43 44

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Xylorhiza tortifolia Machaeranthera hurtleaf PerF RegEn Pres Unc Welsh, Moore 1964 Type locality (Asteraceae) var. imberbis tortifolia var. woodyaster & Canter is E bank of imberbis 2949 (CANY) Green River W of Moab (Cronquist 8994 NY) Compositae Xylorhiza venusta Machaeranthera Cisco woodyaster PerF RegEn Pres Rare Topp 2004 Type locality is (Asteraceae) venusta ST042504001 Cisco, Grand (CANY) Co., UT (Jones s.n. POM) Convolvulaceae Convolvulus arvensis fi eld bindweed PerF Intro Pres Unc Chrobak 463 1985 Utah state (CANY) noxious weed. Native to Eurasia Cornaceae Cornus sericea var. C. stolonifera, red-osier dogwood Shrub Wide Pres Unc Belnap 112 1983 sericea Swida sericea (CANY) Crossosomataceae Glossopetalon Forsellesia spiny greasebush Shrub RegEn Pres Unc Welsh, Moore 1964 (Celastraceae) spinescens var. meionandra & Canter meionandrum 2924 (CANY) Cruciferae Arabis holboellii var. Boechera holboellii Holboell’s rockcress PerF Wide Pres Unc Belnap 45 1987 (Brassicaceae) pinetorum var. pinetorum. (BRY) Name may be misapplied, according to Holmgren et al. (2005). Cruciferae Arabis perennans var. Boechera perennial rockcress PerF Wide Pres Unc Welsh, Moore 1964 (Brassicaceae) perennans perennans & Canter 2903 (CANY) Cruciferae Arabis pulchra var. Boechera pulchra desert rockcress PerF Wide Pres Unc Allen 133 1965 Type locality (Brassicaceae) pallens var. pallens (CANY) is Westwater, Grand Co., UT (Jones s.n. US) Cruciferae Arabis selbyi Boechera selbyi var. Selby’s rockcress PerF Wide Pres Unc Belnap 43 1979 (Brassicaceae) selbyi (CANY) Cruciferae Chorispora tenella (Brassicaceae) blue mustard AnnF Intro Pres Unc Chrobak-Cox 1989 Native to Asia s.n. (CANY) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Cruciferae Conringia orientalis hare’s-ear mustard AnnF Intro Rep ? Schelz et al. NA Native to (Brassicaceae) (2006) Europe Cruciferae Descurainia pinnata Included in western tansy- AnnF Wide Pres Unc Schelz s.n. 1996 (Brassicaceae) var. intermedia D. pinata var. mustard (CANY) osmiarum by some authors Cruciferae Descurainia pinnata D. pinnata ssp. western tansy- AnnF Wide Pres Unc Belnap 42 1967 (Brassicaceae) var. osmiarum halictorum mustard (CANY) Cruciferae Descurainia sophia fl ixweed AnnF Intro Pres Unc Belnap 509 1987 Native to (Brassicaceae) (CANY) Europe Cruciferae Dithyrea wislizenii Dimorphocarpa spectacle-pod AnnF Wide Pres Unc Debolt 169 1964 (Brassicaceae) wislizenii (CANY) Cruciferae Draba cuneifolia var. wedgeleaf draba AnnF Wide Pres Unc Welsh 8838 1969 (Brassicaceae) cuneifolia (BRY) Cruciferae Erysimum asperum E. asperum var. sand dune PerF Wide Pres Unc Welsh, Moore 1964 (Brassicaceae) var. purshii asperum, E. wallfl ower & Canter capitatum var. 2904 (CANY) purshii. Included in var. capitatum in Holmgren et al. (2005) Cruciferae Lepidium alyssoides L. montanum var. Eastwood’s tall PerF RegEn Pres Unc Welsh, Moore 1964 (Brassicaceae) var. eastwoodiae spathulatum, L. pepperwort & Olsen 3687 eastwoodiae (CANY) Cruciferae Lepidium densifl (Brassicaceae) orum Includes var. prairie pepperwort AnnF Wide Pres Unc Chrobak 870 1967 var. ramosum macrocarpum (CANY) Cruciferae Lepidium lasiocarpum St. George AnnF Wide Pres Unc Welsh 8840 1969 Type locality is (Brassicaceae) var. georginum pepperwort (BRY) valley of the Chapter 6:Appendices Virgin near St. George, Washington Co., UT (Parry 19 NY) Cruciferae Lepidium latifolium Cardaria latifolia broad-leaf PerF Intro Pres Unc Chrobak 844 1985 Utah state (Brassicaceae) pepperwort (CANY) noxious weed. Native to Eurasia 45 46

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Cruciferae Lepidium montanum Jones’ pepperwort PerF Wide Pres Unc Welsh, Moore 1964 Type locality (Brassicaceae) var. jonesii & Canter is St. George, 2847 (CANY) Washington Co., UT (Jones 1636 NY) Cruciferae Malcolmia africana African mustard AnnF Intro Pres Unc Allen 128 1978 Native to Africa (Brassicaceae) (CANY) Cruciferae Physaria acutifolia var. P. australis sharp-leaf twinpod PerF Wide Pres Unc Welsh, Moore 1964 (Brassicaceae) acutifolia & Canter 2898 (CANY) Cruciferae Physaria ludoviciana Lesquerella silver bladderpod PerF Wide Rep NA Schelz et al. NA (Brassicaceae) ludoviciana (2006) Cruciferae Physaria newberryi P. newberryi var. Newberry’s PerF RegEn Pres Unc Thorne & 1969 (Brassicaceae) racemosa twinpod Thorne s.n. (BRY) Cruciferae Physaria rectipes Lesquerella rectipes Colorado Plateau PerF RegEn Pres Unc Heil, Porter, & 1964 (Brassicaceae) bladderpod Fleming 5984 (CANY) Cruciferae Sisymbrium altissimum tumble mustard AnnF Intro Pres Unc Allen 127 1978 Native to (Brassicaceae) (CANY) Europe Cruciferae Stanleya pinnata S. pinnata var. prince’s-plume PerF Wide Pres Unc Welsh, Moore 1964 (Brassicaceae) pinnata & Canter 2878 (CANY) Cruciferae Streptanthella long-beak fi ddle- AnnF Wide Pres Com Belnap 506 1979 (Brassicaceae) longirostris mustard (CANY) Cruciferae Streptanthus cordatus S. cordatus var. heart-leaf PerF Wide Pres Unc Johnson 20 1979 (Brassicaceae) cordatus jewelfl ower (CANY) Cruciferae Thelypodiopsis candelabra PerF RegEn Pres Unc Belnap 501 1979 Type locality (Brassicaceae) divaricata thelypody (CANY) is 10 miles E of Hite, San Juan Co., UT (Cronquist 9033 GH) Cruciferae (Brassicaceae) Thelypodium Moab thelypody PerF RegEn Pres Unc Welsh, Moore 1964 integrifolium var. & Olsen 3771 gracilipes (CANY) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Cucurbitaceae Cucurbita foetidissima buffalo gourd PerF Wide Pres Rare Heil, Porter, & 1991 Fleming 7020 (CANY) Cupressaceae Juniperus Sabina Utah juniper Shrub Wide Pres Abun Welsh, Moore 1964 osteosperma osteosperma & Canter 2873 (CANY) Cyperaceae Carex aurea C. garberi golden sedge PerG Wide Pres Com Heil, Porter, & 1987 Fleming 5903 (CANY) Cyperaceae Carex nebrascensis Nebraska sedge PerG Wide Pres Unc Belnap 114 1987 (CANY) Cyperaceae Carex parryana C. parryana var. Parry’s sedge PerG Wide Pres Unc Welsh, Moore 1964 Includes parryana & Canter specimens 2998 (CANY) previously determined as C. specuicola Cyperaceae Carex pellita C. lanuginosa woolly sedge PerG Wide Pres Rare Welsh, Moore 1964 & Canter 2983 (CANY) Cyperaceae Carex praegracilis blackcreeper sedge PerG Wide Pres Rare Welsh, Moore 1964 & Canter 2985 (CANY) Cyperaceae Carex rossii C. defl exa var. Ross’ sedge PerG Wide Pres Rare Welsh, Moore 1964 boottii, C. brevipes, & Olsen 3727 C. pityophila (BRY) Cyperaceae Cyperus acuminatus taper-tip fl at-sedge AnnG Periph Pres Rare Welsh 22477 1983 (CANY) Cyperaceae Cyperus erythrorhizos redroot fl at-sedge AnnG Wide Pres Rare Rink 4598 2005 (CANY) Chapter 6:Appendices Cyperaceae Eleocharis palustris Includes E. common spikerush PerG Wide Pres Unc Belnap 283 1964 macrostachya, E. (CANY) erythropoda, & E. uniglumis Cyperaceae Eleocharis rostellata beaked spikerush PerG Wide Pres Unc Welsh 5384 1966 (CANY) 47 48

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Cyperaceae Scirpus acutus Schoenoplectus hardstem bulrush PerG Wide Pres Unc Welsh, 1964 acutus var. Moore & occidentalis, Canter 3773A Schoenoplectus (CANY) acutus var. acutus, Schoenoplectus lacustris ssp. acutus Cyperaceae Scirpus pungens var. Scirpus pungens three-square PerG Wide Pres Unc Heil, Porter, & 1964 longispicatus var. polyphyllus, bulrush Fleming 5891 Schoenoplectus (CANY) pungens var. longispicatus, Schoenoplectus pungens var. polyphyllus Cyperaceae Scirpus validus Schoenoplectus soft-stem bulrush PerG Wide Rep 1989 Schelz et al. NA tabernaemontani, (2006) Schoenoplectus lacustris ssp. validus Elaeagnaceae Elaeagnus angustifolia Russian-olive Tree Intro Pres Unc Welsh, Moore 1964 Native to & Canter Europe 2997 (CANY) Elaeagnaceae Shepherdia roundleaf Shrub RegEn Pres Unc Thorne, 1987 Type locality rotundifolia buffaloberry Thomas, & is upper valley Inskip 5158 of the Virgin, (CANY) Kane Co., UT (Siler s.n. ISC) Ephedraceae Ephedra torreyana Torrey’s ephedra Shrub Wide Pres Unc Welsh & 1968 Chatterley 21875 (CANY) Ephedraceae Ephedra viridis var. E. viridis green ephedra Shrub Wide Pres Com Belnap 387 1964 viridis (CANY) Ephedraceae Ephedra viridis var. E. cutleri Cutler’s ephedra Shrub RegEn Pres Unc Welsh, Moore 1964 viscida & Canter 2923 (BRY) Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Equisetaceae Equisetum arvense fi eld horsetail Fern Wide Pres Unc Welsh, 1964 Moore, & Canter 2993 (CANY) Equisetaceae Equisetum hyemale Hippochaete tall scouring-rush Fern Wide Pres Unc Topp 1987 var. affi ne hyemalis ssp. ST081604001 affi nis, E. (CANY) praealtum Equisetaceae Equisetum laevigatum Hippochaete smooth scouring- Fern Wide Pres Unc Welsh, Moore 1964 Some CANY laevigata, E. rush & Canter material (Schelz kansanum 2987 (CANY) s.n., 1997) may exhibit hybridization with E. hyemale. Euphorbiaceae Chamaesyce fendleri Euphorbia fendleri Fendler’s spurge PerF Wide Pres Unc Heil, Porter, & 1978 Fleming 5994 (CANY) Euphorbiaceae Chamaesyce Euphorbia ridge-seeded AnnF Wide Pres Unc Welsh 22503 1983 glyptosperma glyptosperma spurge (CANY) Euphorbiaceae Chamaesyce parryi Euphorbia parryi Parry’s spurge AnnF Wide Pres Unc Heil & 1964 Fleming 6075 (CANY) Euphorbiaceae Euphorbia brachycera E. robusta, E. shorthorn spurge PerF Wide Pres Unc Welsh, Moore 1964 brachycera var. & Canter robusta, Tithymalus 2962 (CANY) brachyceras Fagaceae Quercus gambelii var. Q. gambelii Gambel’s oak Tree Wide Pres Com Welsh, Moore 1964 gambelii & Canter

Chapter 6:Appendices 2957 (CANY) Fagaceae Quercus turbinella turbinella live-oak Shrub Wide Rep ? Schelz et al. NA (2006) 49 50

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Fagaceae Quercus welshii Q. havardii shinnery oak Shrub RegEn Pres Unc Welsh, Moore 1964 Type locality of var. tuckeri, Q. & Canter Q. havardii var. undulata 2907 (CANY) tuckeri is Low Pass E of Kens Lake, San Juan Co., UT (Welsh & Higgins 23630 BRY) Fumariaceae Corydalis aurea var. C. curvisiliqua ssp. golden corydalis PerF Wide Pres Unc Allen 205 1964 occidentalis occidentalis (CANY) Gentianaceae Gentiana affi nis var. Pneumonanthe Rocky Mountain PerF Wide Pres Unc Graham 1 1987 affi nis affi nis gentian (CANY) Gentianaceae Swertia radiata Frasera speciosa elkweed PerF Wide Rep Rare Moran s.n. 2007 (CANY) Gentianaceae Swertia utahensis Frasera paniculata Utah swertia PerF RegEn Pres Unc Chrobak 485 1985 Type locality is (CANY) Buckskin Mtns, Kaibab Plateau “on the S edge of Utah”, Kane Co., UT (Jones s.n. POM) Geraniaceae Erodium cicutarium stork’s-bill AnnF Intro Pres Unc Welsh, Moore 1964 Native to & Canter Europe 2972 (CANY) Gramineae Agropyron cristatum A. cristatum var. crested wheatgrass PerG Intro Pres Unc Boschen 23- 1978 Native to (Poaceae) cristatum, A. 84 (CANY) Eurasia. cristatum ssp. pectinatum, A. pectinatum, A. sibiricum, A. desertorum Gramineae Agrostis stolonifera A. alba, A. redtop PerG Intro Pres Rare Hahn 2003 Native to (Poaceae) gigantea LAH09140305 Eurasia and (CANY) Africa Gramineae Andropogon gerardii A. hallii Hall’s bluestem PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) var. chrysocomus & Olsen 3783 (BRY) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gramineae Andropogon gerardii A. gerardii big bluestem PerG Wide Pres Unc Welsh 22605 1983 (Poaceae) var. gerardii (CANY) Gramineae Andropogon glomer- bushy bluestem PerG Periph Rep ? Schelz et al. NA (Poaceae) atus var. scabriglumis (2006) Gramineae Aristida purpurea A. glauca, A. purple three-awn PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) fendleriana, & Olsen 3780 includes vars. (CANY) fendleriana & longiseta Gramineae Avena fatua var. sativa A. sativa oats AnnG Intro Pres Unc Welsh, 1970 Native to (Poaceae) Atwood & Eurasia Moore 10885 (BRY) Gramineae Bothriochloa cane bluestem PerG Wide Pres Unc Welsh 22620 1983 (Poaceae) barbinodis (CANY) Gramineae Bothriochloa silver bluestem PerG Periph Rep ? Schelz et al. NA (Poaceae) laguroides var. (2006) torreyana Gramineae Bouteloua barbata Chondrosum sixweeks grama AnnG Wide Pres Unc Johnson 74 1977 (Poaceae) barbatum. Includes (CANY) vars. barbata & rothrockii Gramineae Bouteloua Includes vars. sideoats grama PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) curtipendula curtipendula & & Olsen 3775 caespitosa (CANY) Gramineae Bouteloua gracilis Chondrosum blue grama PerG Wide Pres Unc Welsh & 1964 (Poaceae) gracilie Moore 3833 (CANY) Gramineae Bromus diandrus Anisantha diandra, ripgut brome AnnG Intro Pres Rare Moran s.n. 2007 Native to Chapter 6:Appendices (Poaceae) B. rigidus (CANY) Eurasia Gramineae Bromus inermis var. Bromopsis inermis (Poaceae) inermis smooth brome PerG Intro Pres Unc Hahn 2003 Native to LAH09140308 Eurasia (CANY) Gramineae Bromus japonicus B. arvensis. Includes Japanese chess AnnG Intro Pres Unc Heil & 1990 Native to (Poaceae) B. commutatus Fleming 6095 Eurasia (CANY) 51 52

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gramineae Bromus rubens red brome AnnG Intro Pres Unc Schelz 2154 2003 Native to (Poaceae) (CANY) Eurasia Gramineae Bromus tectorum Anisantha cheatgrass AnnG Intro Pres Abun Belnap 30 1967 Native to (Poaceae) tectorum (CANY) Eurasia Gramineae Calamagrostis Jones’ reedgrass PerG Wide Pres Unc Welsh, Moore 1964 Type locality (Poaceae) scopulorum & Olsen 3759 is Springdale, (CANY) Washington Co., UT (Jones 6075 POM) Gramineae Distichlis spicata D. spicata var. desert saltgrass PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) stricta, D. stricta & Canter 2989 (CANY) Gramineae Echinochloa crus-gallii Welsh et al. barnyard-grass AnnG Intro Pres Unc Boschen 7-84 1984 Native to (Poaceae) (2003) include E. (CANY) Eurasia. microstachya Gramineae Elymus canadensis E. canadensis var. Canada wildrye PerG Wide Pres Unc Heil & 1987 (Poaceae) canadensis Fleming 6094 (CANY) Gramineae Elymus elongatus E. elongatus tall wheatgrass PerG Intro Rep ? Schelz et al. NA Native to (Poaceae) var. ponticus, (2006) Eurasia Agropyron elongatum, Thinopyrum ponticum Gramineae Elymus elymoides E. elymoides var. squirreltail PerG Wide Pres Unc Heil, Porter, & 1964 (Poaceae) elymoides, Sitanion Fleming 6006 hystrix (CANY) Gramineae Elymus hispidus E. hispidus intermediate PerG Intro Rep ? Schelz et al. NA Native to (Poaceae) var. hispidus, wheatgrass (2006) Eurasia Agropyron intermedium, Elytrigia intermedia, Thinopyrum intermedium Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gramineae Elymus lanceolatus E. lanceolatus thickspike PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) var. lanceolatus, wheatgrass & Olsen 3718 Agropyron (CANY) dasystachyum var. dasystachyum, includes var. riparium, A. albicans var. albicans, A. albicans var. grifﬁ thsii Gramineae Elymus salinus E. salina, Leymus Salina wildrye PerG Wide Pres Unc Topp 2004 Type locality is (Poaceae) salinus, L. salina, L. ST092304003 top of Salina ambiguus (CANY) Pass, Sevier Co., UT (Jones 5447 POM) Gramineae Elymus smithii Agropyron smithii, western PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) Pascopyrum wheatgrass & Canter smithii, includes 3005 (CANY) vars. smithii & palmeri Gramineae Elymus trachycaulus E. trachycaulus slender wheatgrass PerG Wide Pres Unc Belnap 404 1987 (Poaceae) var. trachycaulus, (CANY) E. alaskanus ssp. latiglumis, Agropyron caninum, A. subsecundum, A. trachycaulum, A. latiglume

Chapter 6:Appendices Gramineae Elymus virginicus Virginia wildrye PerG Periph Pres Unc Welsh, Moore 1964 (Poaceae) & Olsen 3717 (CANY) Gramineae Enneapogon desvauxii spike pappusgrass PerG Sparse Pres Rare Miller s.n. 1997 (Poaceae) (CANY) Gramineae Eremopyrum triticeum Agropyron annual wheatgrass AnnG Intro Pres Unc Moran s.n. 2007 Native to Asia (Poaceae) triticeum (CANY) 53 54

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gramineae Erioneuron pilosum hairy tridens PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) & Canter 2833 (CANY) Gramineae Erioneuron Dasyochloa fl uffgrass PerG Wide Pres Unc Topp 2004 (Poaceae) pulchellum pulchella ST050804001 (CANY) Gramineae Festuca arundinacea Lolium tall fescue PerG Intro Pres Unc Topp 2004 Native to (Poaceae) arundinaceum, ST070504002 Europe Schedonorus (CANY) phoenix, S. arundinaceus Gramineae Festuca octofl ora Includes vars. six-weeks fescue AnnG Wide Pres Unc Welsh 21818 1967 (Poaceae) glauca, hirtella, & (CANY) octofl ora, Vulpia octofl ora Gramineae Hilaria jamesii Pleuraphis jamesii galleta PerG Wide Pres Com Welsh, Moore 1964 (Poaceae) & Canter 2839 (CANY) Gramineae Hordeum jubatum Critesion jubatum foxtail barley PerG Wide Rep ? Schelz et al. NA (Poaceae) (2006) Gramineae Hordeum murinum H. murinum rabbit barley AnnG Intro Pres Unc Belnap 280 1987 Native to (Poaceae) ssp. leporinum, (CANY) Europe H. leporinum, Critesion glaucum Gramineae Hordeum pusillum Critesion pusillum little barley AnnG Wide Pres Unc Kitchell s.n. 1964 (Poaceae) (CANY) Gramineae Leptochloa fascicularis Diplachne bearded AnnG Wide Pres Rare Welsh 22476 1983 (Poaceae) fascicularis, L. fusca sprangletop (CANY) var. fascicularis Gramineae Muhlenbergia andina foxtail muhly PerG Wide Pres Unc Welsh, 1964 (Poaceae) Moore, & Olsen 3762 (CANY) Gramineae Muhlenbergia scratchgrass PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) asperifolia & Olsen 3705 (CANY) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gramineae Muhlenbergia Includes reports of New Mexican PerG Wide Pres Rare Heil, Porter, & 1990 (Poaceae) pauciﬂ ora M. arsenei muhly Fleming 5999 (CANY) Gramineae Muhlenbergia porteri bush muhly PerG Wide Pres Unc Welsh 22494 1983 (Poaceae) (CANY) Gramineae Muhlenbergia sandhill muhly PerG Wide Pres Unc Welsh & 1964 (Poaceae) pungens Moore 3864 (CANY) Gramineae Muhlenbergia Includes M. Thurber’s muhly PerG Wide Pres Rare Topp 2003 Type locality of (Poaceae) thurberi curtifolia ST082804003 M. curtifolia is (CANY) between Kanab & Carmel, Kane Co., UT (Jones 6077j US) Gramineae Munroa squarrosa Monroa squarrosa false buffalograss AnnG Wide Pres Unc Topp 2003 (Poaceae) SET10120301 (CANY) Gramineae Oryzopsis micrantha Piptatherum littleseed ricegrass PerG Wide Pres Unc Heil, Porter, & 1990 (Poaceae) micranthum Fleming 5990 (CANY) Gramineae Panicum acuminatum P. occidentale, bundle panicgrass PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) P. tennesseense, & Olsen 3702 P. huachucae, (CANY) Dichanthelium acuminatum ssp. fasciculatum, D. lanuginosum var. fasciculatum Gramineae Panicum capillare P. capillare var. witchgrass AnnG Wide Pres Unc Welsh 22630 1983

Chapter 6:Appendices (Poaceae) occidentale (CANY) Gramineae Panicum obtusum vine mesquite PerG Wide Pres Unc Kitchell 67 1977 (Poaceae) (CANY) Gramineae Panicum virgatum switchgrass PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) & Olsen 3756 (CANY) Gramineae Phleum pratense timothy PerG Intro Rep NA Schelz et al. NA Native to (Poaceae) (2006) Eurasia 55 56

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gramineae Phragmites australis P. communis common reed PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) & Olsen 3754 (CANY) Gramineae Poa bigelovii Bigelow’s bluegrass AnnG Wide Rep NA Schelz et al. NA (Poaceae) (2006) Gramineae Poa bulbosa bulbous bluegrass PerG Intro Rep NA Schelz et al. NA Native to (Poaceae) (2006) Eurasia and Africa Gramineae Poa fendleriana P. longiligula, muttongrass PerG Wide Pres Com Welsh, Moore 1964 (Poaceae) P. fendleriana & Canter ssp. longiligula, 2911 (CANY) includes reports of P. cusickii & P. epilis Gramineae Poa secunda Includes P. Sandberg’s PerG Wide Pres Unc Topp 1999 (Poaceae) gracillima, P. bluegrass ST050904002 juncifolia, P. (CANY) sandbergii, P. canbyi, P. scabrella, P. nevadensis, P. ampla and all vars. of P. secunda Gramineae Polypogon rabbitfoot-grass AnnG Intro Pres Unc Belnap 182 1987 Native to (Poaceae) monspeliensis (CANY) Eurasia and Africa Gramineae Polypogon Agrostis water polypogon PerG Intro Pres Unc Welsh & 1964 Native to (Poaceae) semiverticillatus semiverticillata, P. Moore 3844 Eurasia and viridis (CANY) Africa Gramineae Puccinellia nuttalliana P. airoides Nuttall’s alkaligrass PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) & Canter 2994 (CANY) Gramineae Schizachyrium Andropogon little bluestem PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) scoparium scoparius, S. & Olsen 3761 scoparium var. (CANY) neomexicanum Gramineae Sorghum halepense Johnson-grass PerG Intro Pres Unc Belnap 367 1983 Utah state (Poaceae) (CANY) noxious weed. Native to Eurasia Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gramineae Spartina gracilis alkali cordgrass PerG Wide Pres Unc Heil & 1990 (Poaceae) Fleming 6181 (CANY) Gramineae Spartina pectinata prairie cordgrass PerG Sparse Rep ? Schelz et al. NA (Poaceae) (2006) Gramineae Sporobolus airoides S. airoides alkali sacaton PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) var. airoides & Olsen 3760 (CANY) Gramineae Sporobolus contractus spike dropseed PerG Wide Pres Unc Welsh 22615 1964 (Poaceae) (CANY) Gramineae Sporobolus sand dropseed PerG Wide Pres Unc Welsh 22641 1967 (Poaceae) cryptandrus (CANY) Gramineae Sporobolus ﬂ exuosus mesa dropseed PerG Wide Pres Unc Belnap 327 1964 (Poaceae) (CANY) Gramineae Sporobolus giganteus giant dropseed PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) & Olsen 3769 (CANY) Gramineae Stipa arida Achnatherum Mormon PerG Wide Pres Unc Welsh, Moore 1964 Type locality is (Poaceae) aridum needlegrass & Canter near Milford, 2892 (CANY) Beaver Co., UT (Jones 2106) Gramineae Stipa comata var. Hesperostipa needle-and-thread PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) comata comata var. comata & Canter 2890 (CANY) Gramineae Stipa coronata var. Achnatherum crested PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) parishii coronatum, needlegrass & Canter A. parishii ssp. 3011 (CANY) parishii, S. parishii

Chapter 6:Appendices Gramineae Stipa hymenoides Oryzopsis Indian ricegrass PerG Wide Pres Com Belnap 259 1964 (Poaceae) hymenoides, (CANY) Achnatherum hymenoides Gramineae Stipa neomexicana Hesperostipa New Mexico PerG Wide Pres Rare Topp (Poaceae) neomexicana feathergrass ST0510040012004

(CANY) 57 58

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gramineae Stipa speciosa Achnatherum desert needlegrass PerG Wide Pres Unc Belnap 150 1987 (Poaceae) speciosum, Jarava (CANY) speciosa Gramineae Stipa x bloomeri Stiporyzopsis Bloomer’s ricegrass PerG Wide Pres Unc Fertig 22632 2006 (Poaceae) x bloomeri, (CANY) Achnatherum x bloomeri. Hybrids between S. hymenoides and other Stipa species except S. comata Gramineae Tridens muticus T. muticus var. slim tridens PerG Wide Pres Unc Welsh, Moore 1964 (Poaceae) elongatus & Olsen 3784 (CANY) Hydrophyllaceae Nama densum var. leafy fi ddleleaf AnnF Wide Rep ? Schelz et al. NA parvifl orum (2006) Hydrophyllaceae Phacelia crenulata var. Var. not recognized corrugate phacelia AnnF Wide Pres Unc Debolt 234 1978 corrugata in Cronquist et al. (CANY) (1984) Hydrophyllaceae Phacelia howelliana Included in P. Howell’s phacelia AnnF RegEn Pres Unc Welsh 21815 1983 Type locality crenulata by some (CANY) is 0.4 miles N authors of Bluff, San Juan Co., UT (Atwood 2454 BRY). USFWS: 3C Hydrophyllaceae Phacelia ivesiana Ives’ phacelia AnnF Wide Pres Unc Belnap 390 1979 (CANY) Juncaceae Juncus arcticus J. arcticus var. Baltic rush PerG Wide Pres Unc Belnap 407 1987 balticus, J. balticus (CANY) var. montanus, J. arcticus ssp. ater, includes var. mexicanus Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Juncaceae Juncus ensifolius var. Included in J. Tracy’s rush PerG Wide Pres Unc Heil, Porter, & 1987 brunnescens ensifolius var. Fleming 5953 montanus by (CANY) some authors, J. saximontanus forma brunnescens Juncaceae Juncus ensifolius var. J. saximontanus. mountain rush PerG Wide Pres Unc Welsh, Moore 1964 montanus Includes J. tracyi, J. & Olsen 3755 xiphoides (CANY) Juncaceae Juncus longistylis longstyle rush PerG Wide Pres Unc Welsh, Moore 1964 & Canter 2956 (CANY) Juncaceae Juncus nodosus knotted rush PerG Wide Pres Unc Schelz s.n. 1997 (CANY) Juncaceae Juncus tenuis Includes vars. poverty rush PerG Wide Rep ? Schelz et al. NA dudleyi, congestus, (2006) & tenuis Juncaceae Juncus torreyi Torrey’s rush PerG Wide Pres Unc Belnap 345 1987 (CANY) Labiatae Dracocephalum Moldavica smallfl ower PerF Wide Pres Unc Welsh, Moore 1964 (Lamiaceae) parvifl orum parvifl ora dragonhead & Canter 2965 (CANY) Labiatae Hedeoma drummondii mock pennyroyal PerF Wide Pres Unc Heil, Porter, & 1964 (Lamiaceae) Fleming 5945 (CANY) Labiatae Lycopus americanus American PerF Wide Pres Rare Hahn 2003 (Lamiaceae) bugleweed LAH09140307 (CANY) Labiatae Marrubium vulgare common PerF Intro Pres Rare Castello s.n. 2007 Native to Chapter 6:Appendices (Lamiaceae) horehound (CANY) Eurasia Labiatae Poliomintha incana purple sage Shrub Wide Pres Unc Heil, Porter, & 1990 (Lamiaceae) Fleming 6003 (CANY) Leguminosae Albizia julibrissin (Fabaceae, silk-tree Tree Intro Pres Rare Fertig 22629 2006 Native to SC Mimosaceae) (CANY) Asia 59 60

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Leguminosae Astragalus amphioxys crescent milkvetch PerF Wide Pres Com Belnap 160 1979 (Fabaceae) var. amphioxys (CANY) Leguminosae Astragalus amphioxys Sheldon’s PerF RegEn Pres Unc Welsh 7073 1968 (Fabaceae) var. vespertinus milkvetch (BRY) Leguminosae Astragalus ceramicus painted milkvetch PerF Wide Pres Unc Welsh, Moore 1964 (Fabaceae) var. ceramicus & Canter 2846 (CANY) Leguminosae Astragalus A. chamaeleuce cicada milkvetch PerF RegEn Rep ? Anderson 1983 (Fabaceae) chamaeleuce var. chamaeleuce (1983) Leguminosae Astragalus coltonii var. Moab milkvetch PerF RegEn Rep ? Loope (1977) NA Type locality (Fabaceae) moabensis is near Monticello, San Juan Co., UT (Eastwood 9 POM) Leguminosae Astragalus desperatus rimrock milkvetch PerF RegEn Pres Unc Belnap 196 1983 Type locality (Fabaceae) var. desperatus (CANY) is near Cisco, Grand Co., UT (Jones s.n. POM) Leguminosae Astragalus episcopus A. episcopus var. Bishop’s milkvetch PerF RegEn Pres Rare Topp 2004 Type locality is (Fabaceae) episcopus ST050604003 ‘southern Utah’, (CANY) Kane Co?, UT (Bishop s.n. GH) Leguminosae Astragalus fl avus var. A. fl avus var. yellow milkvetch PerF Wide Rep ? Schelz et al. NA Type locality of (Fabaceae) fl avus candicans (2006) var. candicans is near Richfi eld, Sevier Co., UT (Ward 246 GH) Leguminosae Astragalus fl exuosus pliant milkvetch PerF Periph Rep ? Anderson 1983 (Fabaceae) var. fl exuosus (1983) Leguminosae Astragalus fucatus Hopi milkvetch PerF RegEn Pres Rare Heil, Porter, & 1990 (Fabaceae) Fleming 5880 (CANY)

Leguminosae Astragalus canyon PerF RegEn Pres Unc Heil, Porter, & 1990 (Fabaceae) kentrophyta var. kentrophyta Fleming 5904 coloradoensis (CANY) Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Leguminosae Astragalus straggling PerF RegEn Pres Unc Welsh, Moore 1964 Type locality (Fabaceae) lentiginosus var. milkvetch & Canter is Montezuma palans 2848 (CANY) Canyon, San Juan Co., UT (Eastwood s.n. POM) Leguminosae Astragalus great rushy PerF Wide Pres Unc Belnap 215 1985 Type locality is (Fabaceae) lonchocarpus milkvetch (CANY) ‘southern Utah’, (Parry 32 ISC) Leguminosae Astragalus Moenkopi PerF RegEn Pres Unc Fleming 801 1987 (Fabaceae) moencoppensis milkvetch (CANY) Leguminosae Astragalus mollissimus woolly milkvetch PerF Wide Pres Unc Welsh, Moore 1964 Type locality (Fabaceae) var. thompsoniae & Canter is Kanab, 2838 (CANY) Kane Co., UT (Thompson s.n. GH) Leguminosae Astragalus A. monumentalis Monument PerF LocEn Pres Unc Welsh, Moore 1964 Type locality is (Fabaceae) monumentalis var. monumentalis milkvetch & Canter White Canyon, 2888 (CANY) San Juan Co., UT (Harrison 11595 US). USFWS: 3C Leguminosae Astragalus musiniensis Ferron milkvetch PerF RegEn Pres Rare Welsh & 1970 Type locality (Fabaceae) Atwood 9919 is 2 miles S of (BRY) Ferron, Emery Co., UT (Jones 5454a POM) Leguminosae Astragalus nidularius birds-nest PerF LocEn Pres Rare Heil, Porter, 1990 Type locality is (Fabaceae) milkvetch & Fleming White Canyon, 5899a (CANY) 2 miles below Chapter 6:Appendices Kachina Bridge, San Juan Co., UT (Barneby 12778 CAS). USFWS: 3C Leguminosae Astragalus small-ﬂ owered AnnF RegEn Pres Unc Welsh 21820 1983 (Fabaceae) nuttallianus var. milkvetch (CANY) micranthiformis 61 62

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Leguminosae Astragalus piscator Fisher milkvetch PerF LocEn Pres Unc Welsh, Moore 1964 Type locality (Fabaceae) & Canter is Salt Canyon 2979 (CANY) above The Jump, Canyonlands NP, San Juan Co., UT (Welsh, Moore, & Canter 2979 BRY) Leguminosae Astragalus praelongus longstipe PerF RegEn Rep ? Schelz et al. NA (Fabaceae) var. lonchopus milkvetch (2006) Leguminosae Astragalus praelongus stinking milkvetch PerF Wide Pres Unc Welsh 21808 1983 (Fabaceae) var. praelongus (CANY) Leguminosae Astragalus preussii var. Included in var. Preuss’ milkvetch PerF RegEn Pres Unc Debolt 381 1983 Type locality (Fabaceae) latus preussii by some (CANY) is Green River, authors Emery Co., UT (Jones s.n. POM) Leguminosae Astragalus sabulonum gravel milkvetch AnnF Wide Pres Unc Heil & 1987 (Fabaceae) Fleming 6081 (CANY) Leguminosae Astragalus sandstone PerF RegEn Pres Rare Heil & 1990 Type locality is (Fabaceae) sesquifl orus milkvetch Fleming 6186 ‘southern Utah’ (CANY) (near Kanab), Kane Co., UT (Bishop s.n. GH) Leguminosae Dalea fl avescens var. Kanab prairie- PerF RegEn Pres Unc Welsh, Moore 1964 Type locality is (Fabaceae) fl avescens clover & Canter “Kanab” (but 3006 (CANY) probably from farther east. Thompson s.n. GH) Leguminosae Dalea oligophylla Petalostemon white prairie- PerF Wide Pres Unc Belnap 334 1987 (Fabaceae) candidum, P. clover (CANY) occidentale, D. candida var. oligophylla Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Leguminosae Glycyrrhiza lepidota G. lepidota var. Nuttall’s licorice PerF Wide Pres Unc Chrobak 835 1985 (Fabaceae) lepidota (CANY) Leguminosae Hedysarum boreale Includes var. northern PerF Wide Pres Unc Debolt 164 1983 (Fabaceae) var. boreale pabulare sweetvetch (CANY) Leguminosae Lathyrus brachycalyx Zion sweetpea PerF RegEn Pres Unc Belnap 459 1987 Type locality (Fabaceae) var. zionis (CANY) is 10 miles E of Zion NP, Kane Co., UT (Hitchcock 19013 isotype NY) Leguminosae Lupinus argenteus var. Moab lupine PerF RegEn Pres Unc Johnson 31 1979 Type locality is (Fabaceae) moabensis (CANY) Professor Valley Ranch, Grand Co., UT (Welsh & Taylor 14632 BRY) Leguminosae Lupinus brevicaulis shortstem lupine AnnF Wide Pres Unc Schelz s.n. 1997 (Fabaceae) (CANY) Leguminosae Lupinus pusillus var. dwarf lupine AnnF Wide Pres Unc Welsh, 1964 (Fabaceae) pusillus Moore, & Canter 2832 (CANY) Leguminosae Medicago sativa M. sativa var. sativa alfalfa PerF Intro Rep ? Schelz et al. NA Native to (Fabaceae) (2006) Europe Leguminosae Melilotus alba M. albus. Included white sweet-clover AnnF Intro Pres Unc Chrobak 516 1985 Native to (Fabaceae) in M. offi cinalis by (CANY) Europe some authors Leguminosae Melilotus offi cinalis yellow sweet- PerF Intro Pres Unc Chrobak 863 1985 Native to Chapter 6:Appendices (Fabaceae) clover (CANY) Europe Leguminosae Pediomelum Psoralea large-fl owered PerF RegEn Pres Unc Welsh, Moore (Fabaceae) megalanthum megalantha, breadroot & Canter 1964 Pediomelum megalanthum var. 2889 (CANY) megalanthum 63 64

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Leguminosae Peteria thompsoniae Thompson’s PerF RegEn Pres Unc Welsh 5382 1966 Type locality (Fabaceae) peteria (CANY) is Kanab, Kane Co., UT (Thompson s.n. GH) Leguminosae Psoralidium Psoralea lemon scurf-pea PerF Wide Pres Unc Welsh, Moore 1964 (Fabaceae) lanceolatum var. lanceolata. Vars. & Canter lanceolatum not recognized in 3010 (CANY) Barneby (1989) Leguminosae Psoralidium Vars. not slenderleaf PerF RegEn Pres Unc Welsh 5387 1966 Type locality (Fabaceae) lanceolatum var. recognized in scurfpea (CANY) is near Wilson stenophyllum Barneby (1989) Mesa, Grand Co., UT (Rydberg & Garrett 8367 NY) Leguminosae Psorothamnus Thompson’s Shrub LocEn Rep ? Schelz et al. NA Type locality (Fabaceae) thompsoniae var. indigo-bush (2006) in N AZ, near thompsoniae Kanab, or along the Colorado River in SE UT or AZ (Thompson s.n. US). USFWS: 3C Leguminosae Sophora nuttalliana silky sophora PerF Sparse Rep ? Schelz et al. NA (Fabaceae) (2006) Leguminosae Sophora stenophylla silvery sophora PerF Wide Pres Unc Welsh, Moore 1964 (Fabaceae) & Olsen 3683 (CANY) Leguminosae Vicia ludoviciana Louisiana vetch AnnF Wide Pres Unc Welsh, Moore 1964 (Fabaceae) & Canter 2982 (CANY) Liliaceae Allium macropetalum San Juan onion PerF Wide Pres Unc Allen 85 1978 (Alliaceae) (CANY) Liliaceae Allium nevadense Includes A. Nevada onion PerF Wide Pres Unc Johnson 47 1979 (Alliaceae) atrorubens var. (CANY) cristatum in Welsh et al. (2003). Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Liliaceae Androstephium Brodiaea paysonii pink funnel-lily PerF Wide Pres Unc Belnap 31 1987 Type locality (Alliaceae, brevifl orum (CANY) is near Kanab, Themidaceae) Kane Co., UT (Thompson s.n.) Liliaceae Asparagus offi cinalis asparagus PerF Intro Pres Unc Welsh 22621 1983 Native to (Asparagaceae) (CANY) Eurasia. Liliaceae Calochortus aureus golden mariposa PerF RegEn Pres Unc Graham 2 1987 Type locality (Calochortaceae) (CANY) is near Kanab, Kane Co., UT (Thompson s.n. US) Liliaceae Calochortus fl exuosus sinuous mariposa PerF Wide Pres Unc Chrobak 519 1985 Type locality (Calochortaceae) (CANY) is near Kanab, Kane Co., UT (Thompson s.n. GH) Liliaceae Calochortus nuttallii sego-lily PerF Wide Pres Unc Allen 87 1967 State fl ower of (Calochortaceae) (CANY) Utah Liliaceae Eremocrinum sand lily PerF RegEn Pres Unc Johnson 43 1979 Type locality (Anthericaceae) albomarginatum (CANY) is near Green River, Emery Co., UT (Jones s.n. POM) Liliaceae Smilacina stellata Maianthemum starry false PerF Wide Pres Unc Belnap 111 1987 (Convallariaceae stellatum Solomon’s-seal (CANY) or Ruscaceae) Liliaceae Zigadenus paniculatus Toxicoscordion foothills death PerF Wide Rep ? Loope et al NA (Melanthiaceae) paniculatum camas (1977) Liliaceae Zigadenus vaginatus Anticlea vaginatus. alcove death PerF LocEn Pres Unc Boschen S1- 1983 Type locality Chapter 6:Appendices (Melanthiaceae) Included in Z. camas 84 (CANY) is Natural elegans by some Bridges NM, authors San Juan Co., UT (Rydberg & Garrett 9407 NY). USFWS: 3C Linaceae Linum aristatum Mesynium broom-fl ax AnnF Wide Pres Unc Belnap 268 1964 aristatum (CANY) 65 66

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Linaceae Linum perenne ssp. L. lewisii blue fl ax PerF Wide Pres Unc Allen 151 1978 lewisii var. lewisii, (CANY) Adenolinum lewisii Linaceae Linum puberulum Mesynium downy fl ax AnnF Wide Rep ? Schelz et al. NA puberulum (2006) Linaceae Linum subteres Utah yellow-fl ax PerF RegEn Pres Unc Heil & 1983 Type locality is Fleming 6187 Willow Creek, (CANY) San Juan Co., UT (Eastwood s.n.) Loasaceae Mentzelia albicaulis M. obscura, white-stem AnnF Wide Pres Unc Debolt 379 1978 Acrolasia albicaulis, blazingstar (CANY) A. gracilis Loasaceae Mentzelia marginata M. marginata var. Colorado stickleaf PerF RegEn Pres Unc Welsh & 1964 Type locality is marginata. Includes Moore 3853 75 miles W of var. cronquistii, (CANY) Blanding and 10 M. cronquistii, miles E of Hite, Nuttallia San Juan Co., cronquistii UT (Cronquist 9030 NY) Loasaceae Mentzelia multifl ora M. multifl ora desert stickleaf PerF Wide Pres Unc Welsh, Moore 1964 var. multifl ora, & Olsen 3747 Nuttallia multifl ora (CANY) Malvaceae Malva neglecta common mallow AnnF Intro Pres Unc Welsh, Moore 1964 Native to & Canter Eurasia 2966 (CANY) Malvaceae Malvella leprosa alkali mallow PerF Wide Pres Unc Welsh 22487 1983 (CANY) Malvaceae Sphaeralcea coccinea Includes ssp. scarlet globe- PerF Wide Pres Unc Heil, Porter, & 1968 coccinea, elata, & mallow Fleming 5996 dissecta (CANY) Malvaceae Sphaeralcea S. grossulariifolia gooseberry-leaf PerF Wide Pres Unc Welsh, Moore 1964 grossulariifolia var. grossulariifolia globemallow & Canter 2852 (CANY) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Malvaceae Sphaeralcea janeae S. leptophylla var. Jane’s PerF LocEn Pres Unc Welsh & 1968 Type locality is janeae globemallow Chatterley along White 21878 (CANY) Rim Road, Canyonlands National Park, San Juan Co., UT (Welsh 7064 BRY). UT BLM: Sensitive, USFWS: former C2 candidate Malvaceae Sphaeralcea slenderleaf PerF Wide Pres Unc Allen 160 1967 leptophylla globemallow (CANY) Malvaceae Sphaeralcea moorei S. grossulariifolia Moore’s PerF LocEn Pres Unc Thorne, 1987 Type locality is var. moorei globemallow Thomas, & E side of Last Inskip 5171 Chance Bay, (CANY) Lake Powell, Kane Co., UT (Welsh & Atwood 11597 BRY) Malvaceae Sphaeralcea parvifolia small-leaf PerF Wide Pres Com Welsh 21809 1964 globemallow (CANY) Moraceae Morus alba white mulberry Tree Intro Rep ? Schelz et al. NA Native to China (2006) Nyctaginaceae Abronia fragrans A. elliptica, A. fragrant sand- PerF Wide Pres Com Welsh, Moore 1964 fragrans var. verbena & Canter elliptica, A. nana 2863 (CANY) var. harrisii Nyctaginaceae Allionia incarnata A. incarnata var. trailing four- PerF Wide Pres Unc Belnap 362 1981 Chapter 6:Appendices incarnata o’clock (CANY) Nyctaginaceae Mirabilis linearis var. Oxybaphus linearis, narrow-leaf four- PerF Wide Pres Unc Belnap 343 1978 linearis O. decumbens o’clock (CANY) Nyctaginaceae Mirabilis multiﬂ ora Includes vars. showy four-o’clock PerF Wide Pres Unc Welsh 5394 1964 multiﬂ ora, (CANY) pubescens, & glandulosa 67 68

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Nyctaginaceae Tripterocalyx carneus T. wootonii Wooton’s sand- AnnF RegEn Rep ? Schelz et al. NA var. wootonii verbena (2006) Nyctaginaceae Tripterocalyx Abronia micrantha, small-fl ower AnnF Wide Pres Unc Thorne, 1978 micranthus T. pedunculatus sandpuffs Thomas, & Inskip 5164 (CANY) Oleaceae Forestiera pubescens F. neomexicana desert olive Shrub Wide Pres Unc Heil, Porter, & 1964 Fleming 5954 (CANY) Oleaceae Fraxinus anomala singleleaf ash Shrub Wide Pres Com Welsh & 1964 Type locality Chatterley is Labyrinth 21889 (CANY) Canyon, San Juan Co., UT (Newberry s.n. NY) Onagraceae Calylophus C. hartwegii ssp. lavender-leaf PerF Wide Pres Com Welsh, Moore 1964 lavandulifolius lavandulifolius sundrops & Canter 2885 (CANY) Onagraceae Camissonia walkeri Walker’s AnnF Wide Pres Unc Welsh 22603 1983 Type locality is camissonia (CANY) Paradox Valley, Montrose Co., CO Onagraceae Epilobium ciliatum E. ciliatum northern willow- PerF Wide Pres Unc Schelz 7.14.98 1998 var. ciliatum, herb (CANY) E. watsonii, E. adenocaulon Onagraceae Gaura parvifl ora G. mollis small-fl owered AnnF Wide Pres Unc Belnap 135 1983 gaura (CANY) Onagraceae Oenothera albicaulis white-stem AnnF Wide Pres Unc Belnap 272 1987 evening-primrose (CANY) Onagraceae Oenothera caespitosa long-tube evening- PerF Wide Pres Unc Allen 172 1978 var. marginata primrose (CANY) Onagraceae Oenothera caespitosa Paria evening- PerF RegEn Pres Unc Johnson 11 1979 Type locality var. navajoensis primrose (CANY) is Paria River, Kane Co., UT (Cronquist 10202 MO) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Onagraceae Oenothera longissima Bridges evening- PerF RegEn Pres Unc Welsh, Moore 1964 Type locality is primrose & Canter Armstrong & 3757 (CANY) White canyons [Natural Bridges NM], San Juan Co., UT (Rydberg & Garrett 9410 NY) Onagraceae Oenothera pallida var. Includes var. pale evening- PerF Wide Pres Unc Belnap 167 1967 pallida runcinata primrose (CANY) Orchidaceae Epipactis gigantea giant helleborine PerF Wide Pres Unc Welsh 5389 1966 (CANY) Orchidaceae Habenaria zothecina Platanthera alcove bog-orchid PerF RegEn Pres Unc Fowler & May 1964 Type locality is zothecina, 1595 (CANY) ca 1 mile N of Limnorchis Moab, Grand zothecina Co., UT (Welsh & Higgins 23269 BRY). UT BLM: Sensitive, USFWS: former C2 candidate Orobanchaceae Orobanche fasciculata Aphyllon clustered PerF Wide Pres Unc Schelz 6.1.98 1983 fasciculatum broomrape (CANY) Orobanchaceae Orobanche Includes reports of Cooper’s PerF Wide Pres Unc Belnap 356 1978 ludoviciana var. O. multiﬂ ora, O. broomrape (CANY) cooperi cooperi Papaveraceae Argemone corymbosa San Rafael prickly- PerF RegEn Rep ? Schelz et al. NA Type locality is var. arenicola poppy (2006) 37.9 miles SW of Green River, Chapter 6:Appendices Emery Co., UT (Ownbey & Ownbey 2146 ) 69 70

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Papaveraceae Argemone corymbosa San Rafael prickly- PerF LocEn Pres Rare Thorne, 1987 Type locality var. parva poppy Thomas, & is bluffs along Inskip 5175 Colorado River (CANY) along Hwy 129, Grand Co., UT (Welsh 6857 BRY) Pinaceae Abies concolor white fi r Tree Wide Pres Rare Wilson s.n. 1970 (UTC) Pinaceae Pinus edulis two-needle pinyon Tree Wide Pres Abun Welsh, Moore 1964 & Olsen 3732 (CANY) Pinaceae Pinus ponderosa var. ponderosa pine Tree Wide Pres Unc Welsh, Moore 1964 scopulorum & Canter 3001 (CANY) Pinaceae Pseudotsuga menziesii Douglas-fi r Tree Wide Pres Unc Belnap 498 1987 var. glauca (CANY) Plantaginaceae Plantago major Includes var. common plantain PerF Intro Pres Unc Belnap 140 1987 Mostly native pachyphylla (CANY) to Europe. Type locality of the native phase “var. pachyphylla” is Salt Lake City, Salt Lake Co., UT (Jones 1030 POM) Plantaginaceae Plantago patagonica Includes vars. woolly plantain AnnF Wide Pres Unc Welsh, Moore 1964 patagonica, & Canter gnaphaloides, 2968 (CANY) spinulosa, & breviscapa Polemoniaceae Gilia hutchinsifolia Aliciella broadlobe gilia AnnF Wide Pres Rare Schelz s.n. 2004 hutchinsifolia (CANY) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Polemoniaceae Gilia imperialis G. latifolia var. Cataract gilia AnnF LocEn Pres Rare Higgins, 1983 Type locality imperialis, Aliciella Welsh, Welsh, is Cataract latifolia ssp. & Welsh Canyon N imperialis 14163 (CANY) of Imperial Canyon, San Juan Co., UT (Welsh 22507 BRY). UT BLM: Sensitive Polemoniaceae Gilia inconspicua Includes var. shy gilia AnnF Wide Pres Unc Heil, Porter, & 1967 CANY material inconspica & Fleming 5918 mostly var. sinuata, G. (CANY) sinuata (if ophthalmoides, G. recognized), sinuata, G. clokeyi, but also G. transmontana includes at least one collection of var. inconspicua. Polemoniaceae Gilia leptomeria var. Aliciella leptomeria common gilia AnnF Wide Pres Unc Belnap 247 1979 leptomeria (CANY) Polemoniaceae Gilia subnuda Aliciella subnuda carmine gilia PerF RegEn Pres Unc Belnap 391 1987 (CANY) Polemoniaceae Ipomopsis aggregata Gilia aggregata var. scarlet gilia PerF Wide Pres Unc Welsh 22608 1964 var. aggregata aggregata (CANY) Polemoniaceae Ipomopsis gunnisonii Gilia gunnisonii Gunnison’s gilia AnnF RegEn Pres Com Belnap 370 1987 Type locality (CANY) is “sand banks of the Green River”, UT (Cruetzfeldt s.n. NY) Chapter 6:Appendices Polemoniaceae Ipomopsis longifl ora Gilia longifl ora long-fl owered gilia PerF Wide Pres Unc Debolt 166 1964 (CANY) Polemoniaceae Ipomopsis polycladon Gilia polycladon spreading gilia AnnF Wide Pres Unc Welsh & 1983 Chatterley 21877 (CANY) Polemoniaceae Ipomopsis pumila Gilia pumila dwarf gilia AnnF Wide Pres Unc Johnson 42 1979 (CANY) 71 72

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Polemoniaceae Ipomopsis roseata Gilia roseata San Rafael gilia PerF RegEn Pres Unc Heil, Porter, & 1987 Fleming 5925 (CANY) Polemoniaceae Leptodactylon Leptodactylon prickly-phlox Shrub Wide Pres Unc Debolt 380 1984 pungens pungens var. (CANY) pungens, Linanthus pungens Polemoniaceae Leptodactylon Linanthus watsonii Watson’s prickly- PerF Wide Pres Unc Heil, Porter, & 1990 Type locality is watsonii phlox Fleming 5995 Cottonwood (CANY) Canyon, Salt Lake Co., UT (Watson s.n. GH) Polemoniaceae Phlox hoodii var. Includes ssp. carpet phlox PerF Wide Pres Unc Belnap 38 1979 canescens glabrata, hoodii, & (CANY) viscidula Polemoniaceae Phlox longifolia Includes vars. long-leaf phlox PerF Wide Pres Unc Schelz s.n. 1997 longifolia and (CANY) stansburyi Polygonaceae Eriogonum alatum E. alatum winged wild PerF Wide Pres Unc Belnap 249 1978 var. alatum, buckwheat (CANY) Pterogonum alatum Polygonaceae Eriogonum bicolor pretty wild PerF RegEn Pres Unc Heil, Porter, & 1968 Type locality buckwheat Fleming 5896 is Thompson’s b (CANY) Springs, Grand Co., UT (Jones s.n. POM) Polygonaceae Eriogonum cernuum Vars. not nodding wild AnnF Wide Pres Unc Allen 101 1967 var. cernuum recognized in Flora buckwheat (CANY) of North America (2005) Polygonaceae Eriogonum Includes var. crisp-leaf wild Shrub Wide Pres Unc Welsh & 1964 corymbosum var. velutinum and E. buckwheat Moore 3854 corymbosum lancifolium (CANY) Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Polygonaceae Eriogonum rimrock wild Shrub RegEn Pres Unc Fertig 22637 2006 Type locality corymbosum var. buckwheat (CANY) is Green River, orbiculatum Emery Co., UT (Jones s.n. POM) Polygonaceae Eriogonum defl exum E. defl exum var. skeletonweed wild AnnF Wide Rep ? Schelz et al. NA defl exum buckwheat (2006) Polygonaceae Eriogonum gordonii Gordon’s wild AnnF Wide Pres Unc Schelz s.n. 2001 buckwheat (CANY) Polygonaceae Eriogonum infl atum E. infl atum desert trumpet PerF Wide Pres Unc Welsh, Moore 1964 var. infl atum & Canter 2897 (CANY) Polygonaceae Eriogonum sand wild Shrub RegEn Pres Unc Welsh, Moore 1964 Type locality leptocladon var. buckwheat & Olsen 3731 is Green River, leptocladon (CANY) Emery Co., UT (Cruezfeldt s.n. NY) Polygonaceae Eriogonum E. microthecum var. slender wild PerF Wide Pres Com Welsh, Moore 1964 microthecum var. foliosum buckwheat & Olsen 3729 simpsonii (CANY) Polygonaceae Eriogonum E. ovalifolium var. cushion wild PerF Wide Pres Unc Topp 1999 ovalifolium ovalifolium buckwheat ST050904001 (CANY) Polygonaceae Eriogonum Palmer’s wild AnnF Wide Pres Unc Belnap 88 1987 palmerianum buckwheat (CANY) Polygonaceae Eriogonum shockleyi Includes vars. Shockley’s wild PerF Wide Pres Unc Heil, Porter, & longilobum & buckwheat 1990 Type locality of shockleyi Fleming 5968 var. longilobum (CANY) is near Price,

Chapter 6:Appendices Carbon Co., UT (Jones 5590j POM) Polygonaceae Eriogonum arid wild PerF Wide Rep ? Schelz et al. NA umbellatum var. buckwheat (2006) subaridum 73 74

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Polygonaceae Eriogonum wetherillii Wetherill’s wild AnnF RegEn Pres Unc Belnap 203 1987 Type locality is buckwheat (CANY) along the San Juan River, San Juan Co., UT (Eastwood 124 CAS) Polygonaceae Rumex hymenosepalus canaigre dock PerF Wide Pres Unc Debolt 223 1983 (CANY) Polygonaceae Rumex maritimus var. R. fueginus golden dock AnnF Wide Pres Rare Schelz s.n. 2002 fueginus (CANY) Polygonaceae Rumex venosus veiny dock PerF Wide Rep ? Schelz et al. NA (2006) Polypodiaceae Adiantum capillus- southern Fern Wide Pres Rare Heil, Porter, & 1964 (Adiantaceae) veneris maidenhair fern Fleming 5971 (CANY) Polypodiaceae Asplenium resiliens ebony spleenwort Fern Periph Pres Rare O’Kane & 1988 (Aspleniaceae) Carulli 3295 (BRY) Polypodiaceae Cheilanthes feei slender lip-fern Fern Wide Pres Unc Belnap 153 1987 (Adiantaceae) (CANY) Polypodiaceae Pellaea glabella var. P. occidentalis western cliffbrake Fern Periph Pres Unc Boschen 1984 (Adiantaceae) occidentalis PEBR8-14-84 (CANY) Polypodiaceae Pellaea limitanea Argyrochosma border cloak-fern Fern Periph Pres Rare Heil & 1990 (Adiantaceae) limitanea ssp. Fleming 6093 limitanea, (CANY) Notholanea limitanea Portulacaceae Portulaca oleracea common purslane AnnF Intro Pres Unc Moran s.n. 2007 Native to E (CANY) North America Portulacaceae Talinum brevifolium Phemeranthus sausageleaf PerF Wide Pres Unc Heil, Porter, & 1983 brevifolius talinum Fleming 5922 (CANY) Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Primulaceae Primula specuicola cave primrose PerF RegEn Pres Unc Debolt 227 1983 Type locality is (CANY) along San Juan River near Bluff, San Juan Co., UT (Rydberg 9882 NY). USFWS: 3C Ranunculaceae Aquilegia micrantha alcove columbine PerF RegEn Pres Unc Fowler & May 1978 Type locality (Helleboraceae) 1626 (CANY) is Bluff, San Juan Co., UT (Wetherill s.n. CAS) Ranunculaceae Clematis columbiana C. pseudoalpina, Columbian virgin’s- PerF Wide Pres Unc Belnap 413 1987 Atragene bower (CANY) columbiana Ranunculaceae Clematis ligusticifolia white virgin’s- PerF Wide Pres Unc Welsh, Moore 1964 bower & Olsen 3679 (CANY) Ranunculaceae Delphinium andersonii D. scaposum pale larkspur PerF Wide Pres Unc Welsh, Moore 1964 (Helleboraceae) var. scaposum & Canter 2896 (CANY) Ranunculaceae Delphinium D. nelsonii Nuttall’s larkspur PerF Wide Pres Unc Allen 118 1978 (Helleboraceae) nuttallianum (CANY) Ranunculaceae Ranunculus Halerpestes marsh buttercup PerF Wide Pres Unc Welsh, Moore 1964 cymbalaria cymbalaria ssp. & Canter saximontana, R. 2988 (CANY) cymbalaria var. cymbalaria. Vars. not recognized in Flora of North Chapter 6:Appendices America (1997) Ranunculaceae Ranunculus Ceratocephala bur buttercup AnnF Intro Rep ? Schelz et al. NA Native to testiculatus orthoceras, C. (2006) Eurasia testiculata 75 76

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Rhamnaceae Ceanothus greggii var. Franklin’s desert- Shrub LocEn Pres Unc Debolt 1983 1978 Type locality is franklinii lilac (CANY) Muley Point, San Juan Co., UT (Welsh, Welsh, & Welsh 22244 BRY) Rhamnaceae Rhamnus betulifolia Frangula birchleaf Shrub Wide Pres Unc Welsh 5385 1964 betulifolia buckthorn (CANY) Rosaceae Amelanchier utahensis Utah serviceberry Shrub Wide Pres Com Welsh, Moore 1964 Type locality & Canter is Toquerville, 2909 (CANY) Washington Co., UT (Jones 1716 GH) Rosaceae Cercocarpus intricatus C. ledifolius var. dwarf mountain Shrub Wide Pres Com Heil, Porter, & 1978 Type locality is intricatus mahogany Fleming 6004 American Fork (CANY) Canyon, Utah Co., UT (Watson 314 US) Rosaceae Cercocarpus ledifolius C. ledifolius var. curl-leaf mountain Shrub Wide Rep ? Schelz et al. NA Type locality var. intermontanus intercedens mahogany (2006) is ca 10 km E of Oak City, Millard Co., UT (Holmgren & Holmgren 11298 NY) Rosaceae Cercocarpus montanus alder-leaf Shrub Wide Pres Unc Welsh, Moore 1964 mountain & Canter mahogany 2917 (CANY) Rosaceae Coleogyne blackbrush Shrub Wide Pres Com Welsh, Moore 1964 ramosissima & Canter 2880 (CANY) Rosaceae Fallugia paradoxa Apache plume Shrub Wide Pres Unc Heil, Porter, & 1990 Fleming 5962 (CANY) Rosaceae Holodiscus dumosus H. discolor bush oceanspray Shrub Wide Pres Unc Belnap 485 1987 (CANY) Rosaceae Petrophyton Spiraea caespitosa rock spiraea PerF Wide Pres Unc Debolt 236 1983 caespitosum (CANY) Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Rosaceae Physocarpus alternans dwarf ninebark Shrub Wide Pres Unc Heil, Porter, & 1990 Fleming 5948 (CANY) Rosaceae Potentilla paradoxa P. supina ssp. bushy cinquefoil AnnF Wide Pres Unc Chrobak-Cox 1986 paradoxa s.n. (CANY) Rosaceae Potentilla rivalis brook cinquefoil AnnF Sparse Pres Rare Schelz 2002- 2002 0523.01 (CANY) Rosaceae Prunus persica peach Tree Intro Rep NA Schelz et al. NA Native to China (2006) Rosaceae Prunus virginiana var. Padus virginiana western Shrub Wide Pres Unc Belnap 36 1979 melanocarpa ssp. melanocarpa chokecherry (CANY) Rosaceae Purshia mexicana var. P. stansburiana, cliff-rose Shrub Wide Pres Unc Welsh, Moore 1964 stansburyana Cowania mexicana & Canter 2895 (CANY) Rosaceae Rosa woodsii R. woodsii var. Woods’ rose Shrub Wide Pres Unc Welsh, Moore 1964 ultramontana. & Canter Includes R. manca 2963 (CANY) Rubiaceae Galium multifl orum G. coloradoense Colorado bedstraw PerF RegEn Pres Unc Welsh, Moore 1964 var. coloradoense & Canter 2932 (CANY) Rubiaceae Galium multifl orum G. multifl orum shrubby bedstraw PerF Wide Pres Unc Fowler & May 1993 var. multifl orum 1571 (CANY) Salicaceae Populus alba white poplar Tree Intro Rep ? Schelz et al. NA Native to (2006) Eurasia Salicaceae Populus fremontii var. P. deltoides var. Fremont Tree Wide Pres Unc Welsh & 1964 fremontii fremontii cottonwood Moore 3855 (CANY)

Chapter 6:Appendices Salicaceae Populus x intercurrens P. acuminata, P. x acuminata. lance-leaf Tree Wide Pres Unc Welsh, Moore 1964 Type locality Hybrid cottonwood & Olsen 3749 is Jesse Ewing of P. fremontii x P. (CANY) Canyon, angustifolia Daggett Co., UT (Goodrich 24441 BRY) Salicaceae Salix bebbiana Bebb’s willow Shrub Wide Pres Unc Welsh, Moore 1964 & Olsen 3748

77 (CANY) 78

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Salicaceae Salix eriocephala var. S. ligulifolia strapleaf willow Shrub Wide Pres Rare Fertig 22635 2006 ligulifola (CANY) Salicaceae Salix eriocephala var. S. lutea, S. rigida yellow willow Shrub Wide Pres Unc Schelz s.n. 1997 watsonii var. watsonii (CANY) Salicaceae Salix exigua var. S. exigua var. coyote willow Shrub Wide Pres Unc Welsh, Moore 1964 stenophylla exigua & Canter 2940 (CANY) Salicaceae Salix gooddingii S. nigra var. Goodding’s black Tree Wide Pres Unc Belnap 275 1964 vallicola willow (CANY) Santalaceae Comandra umbellata C. pallida bastard toadfl ax PerF Wide Pres Unc Welsh, Moore 1964 var. pallida & Canter 2842 (CANY) Saxifragaceae Fendlera rupicola F. falcata fendlerbush Shrub Wide Pres Unc Belnap 159 1964 (Hydrangeaceae) (CANY) Saxifragaceae Philadelphus P. microphyllus littleleaf mock- Shrub Wide Pres Unc Welsh, Moore 1964 (Hydrangeaceae) microphyllus var. occidentalis, orange & Canter P. occidentalis var. 2935 (CANY) occidentalis Saxifragaceae Ribes aureum R. aureum var. golden currant Shrub Wide Pres Unc Fleming 866 1983 (Grossulariaceae) aureum (CANY) Scrophulariaceae Castilleja chromosa C. angustifolia desert paintbrush PerF Wide Pres Unc Belnap 102 1987 var. dubia, C. (CANY) applegatei ssp. martinii Scrophulariaceae Castilleja fl ava C. fl ava var. fl ava yellow paintbrush PerF Wide Rep ? Schelz et al. NA (2006) Scrophulariaceae Castilleja linariifolia C. linariaefolia Wyoming PerF Wide Pres Unc Fowler & May 1964 paintbrush 1567 (CANY) Scrophulariaceae Castilleja scabrida var. Eastwood’s PerF RegEn Pres Unc Belnap 198 1983 scabrida paintbrush (CANY) Scrophulariaceae Cordylanthus wrightii Wright’s bird’s- AnnF Wide Pres Unc Chrobak 584 1984 beak (CANY) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Scrophulariaceae Mimulus eastwoodiae Eastwood’s PerF RegEn Pres Unc Heil & 1983 Type locality is monkey-ﬂ ower Fleming 6191 San Juan River (CANY) near Bluff, San Juan Co., UT (Rydberg 9883 NY) Scrophulariaceae Pedicularis dwarf lousewort PerF Wide Pres Rare Anderson s.n. 1983 centranthera (BRY) Scrophulariaceae Penstemon dusty penstemon PerF RegEn Pres Unc Allen 203 1978 Type locality comarrhenus (CANY) is Aquarius Plateau, Garﬁ eld Co., UT (Ward 462 GH) Scrophulariaceae Penstemon bluestem PerF RegEn Pres Unc Fleming 807 1979 cyanocaulis penstemon (CANY) Scrophulariaceae Penstemon eatonii undulate PerF Wide Pres Unc Welsh, Moore 1964 Type locality var. undosus penstemon & Canter is St. George, 2912 (CANY) Washington Co., UT (Jones 5110h US) Scrophulariaceae Penstemon palmeri Palmer’s PerF Wide Pres Unc Chrobak-Cox 1987 var. palmeri penstemon s.n. (CANY) Scrophulariaceae Penstemon utahensis Utah penstemon PerF Wide Pres Unc Moore 292 1965 Type locality (BRY) is between Hatch’s Wash & Monticello, San Juan Co., UT (Eastwood s.n. CAS)

Chapter 6:Appendices Scrophulariaceae Verbascum thapsus woolly mullein PerF Intro Rep 1989 Schelz et al. NA Native to (2006) Eurasia Scrophulariaceae Veronica anagallis- water speedwell PerF Intro Pres Unc Schelz s.n. 2002 Native to aquatica (CANY) Europe Solanaceae Chamaesaracha false nightshade PerF Periph Pres Rare Mann s.n. 2007 coronopus (CANY) 79 80

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Solanaceae Datura wrightii D. meteloides angel’s trumpet AnnF Wide Pres Unc Welsh, Moore 1964 & Canter 2943 (CANY) Solanaceae Lycium andersonii Anderson’s Shrub Wide Pres Unc Welsh 22639 1983 wolfberry (CANY) Solanaceae Lycium pallidum pale wolfberry Shrub Wide Rep ? Schelz et al. NA (2006) Solanaceae Lycium torreyi Torrey’s wolfberry Shrub Wide Pres Unc Chrobak 592 1983 (CANY) Solanaceae Nicotiana N. obtusifolia, N. desert tobacco AnnF Wide Pres Unc Allen 197 1978 trigonophylla trigonophylla var. (CANY) palmeri Solanaceae Physalis hederifolia P. fendleri, P. Fendler’s ground- PerF Wide Pres Unc Welsh, Moore 1964 var. fendleri hederifolia var. cherry & Olsen 3746 cordifolia, P. (CANY) hederaefolia var. fendleri Solanaceae Physalis longifolia P. virginiana var. long-leaf ground- PerF Wide Rep NA Schelz et al. NA sonorae cherry (2006) Solanaceae Solanum jamesii James’ potato PerF Periph Pres Rare Moran s.n. 2007 (CANY) Solanaceae Solanum rostratum buffalobur AnnF Intro Rep ? Schelz et al. NA Native to C (2006) North America Solanaceae Solanum trifl orum cut-leaf nightshade AnnF Wide Rep 1991 Schelz et al. NA Native to C (2006) North America, may be adventive in UT. Tamaricaceae Tamarix chinensis T. pentandra, T. fi ve-stamen Shrub Intro Pres Com Welsh, Moore 1964 Native to ramosissima tamarisk & Canter Eurasia 2851 (CANY) Typhaceae Typha domingensis southern cattail PerG Wide Pres Unc Welsh, Moore 1964 & Olsen 3772 (CANY) Typhaceae Typha latifolia broad-leaved PerG Wide Pres Unc Fleming 862 1989 cattail (CANY) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Canyonlands National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Ulmaceae Celtis reticulata C. laevigata var. netleaf hackberry Tree Wide Pres Unc Welsh 21813 1964 (Celtidaceae) reticulata (CANY) Ulmaceae Ulmus pumila Siberian elm Tree Intro Rep Rare Loope (1977), 2004 Native to Asia Dewey & Andersen (2005) Umbelliferae Cymopterus acaulis C. fendleri, C. Fendler’s PerF Wide Pres Unc Welsh 21803 1983 (Apiaceae) var. fendleri glomeratus cymopterus (CANY) Umbelliferae Cymopterus bulbosus onion spring- PerF Wide Rep ? Schelz et al. NA (Apiaceae) parsley (2006) Umbelliferae Cymopterus newberryi sticky spring- PerF RegEn Pres Unc Johnson 44 1979 (Apiaceae) parsley (CANY) Umbelliferae Cymopterus purpureus C. purpureus Colorado Plateau PerF RegEn Pres Unc Schelz s.n. 1997 (Apiaceae) var. purpureus spring-parsley (CANY) Umbelliferae Cymopterus C. petraeus, Aletes rock spring-parsley PerF Wide Pres Unc Fleming 788 1983 (Apiaceae) terebinthinus var. petraea, Pteryxia (CANY) petraeus petraea Umbelliferae Lomatium parryi Parry’s lomatium PerF Wide Pres Unc Welsh, Moore 1964 Type locality (Apiaceae) & Canter is “Southern 2891 (CANY) Utah” (Parry 85) Verbenaceae Phyla cuneifolia Lippia cuneifolia wedge-leaf fog- PerF Sparse Pres Unc Belnap 353 1979 fruit (CANY) Verbenaceae Verbena bracteata prostrate vervain PerF Wide Pres Unc Chrobak-Cox 1985 s.n. (CANY) Viscaceae Arceuthobium pinyon dwarf- PerF Wide Pres Unc Fertig 22633 2006 Typically divaricatum mistletoe (CANY) parasitic on Pinus edulis and P. monophylla

Chapter 6:Appendices Viscaceae Phoradendron juniper mistletoe PerF Wide Pres Unc Welsh & 1964 Typically juniperinum Moore 3830 (CANY) parasitic on Juniperus osteosperma Zygophyllaceae Kallstroemia yellow kallstroemia AnnF Periph Pres Rare Belnap 365 1987 californica (CANY) Zygophyllaceae Tribulus terrestris puncture vine AnnF Intro Pres Unc Chrobak-Cox 1988 Native to s.n. (CANY) Eurasia 81 3C Range Comments Life form ower PerF Periph western stickseed AnnF Wide common pricklypear pricklypearNichol’s rufous-spine pricklypear PerF PerF PerF Wide Wide LocEn slenderspine pricklypear PerF Periph unequal cryptanthNevada cryptanthwing-nut cryptanth AnnF Periph AnnF AnnF Periph Periph northern arrowhead PerF Wide pallid milkweed milkweedCutler’s dwarf milkweed PerFJames’ cryptanth PerF Wide cushion cryptanth RegEn PerFsand dune cryptanth USFWS: Category PerF Periph AnnF Wide AnnF Wide Wide cardinal-ﬂ iodine bushmat saltbush orachBlue Valley stalked orachpitseed goosefoot Shrub AnnF Shrubdesert goosefoot Wide RegEn RegEn AnnF AnnF Wide RegEn AnnF Sparse

var. var. C. erinacea nicholii zschackei redowskii var. var. Flora of North Flora of North caput-medusae var. Included in var. by some authors in in O. polyacantha O. polyacantha var. (2003) (2003) Included in polyacantha America Included in polyacantha America L. redowskii C. jamesii, Oreocarya suffruticosa angustifolia C. berlandieri

var. var. var. var. var.

erinacea O. polyacantha var. saccaria A. saccaria var. var. ssp. var. ora rmed) taxa of Canyonlands National Park. spina Opuntia erinacea Opuntia nicholiiOpuntia polyacantha rufi Opuntia polyacantha trichophora O. polyacantha Lobelia cardinalis Cryptantha nevadensis Cryptantha pterocarya cycloptera Lappula occidentalis occidentalis Sagittaria cuneata Asclepias cryptoceras cryptoceras Asclepias cutleri Asclepias involucrata Cryptantha cinerea jamesii Cryptantha circumscissa Cryptantha fendleri Cryptantha inaequata C. holoptera. graminea Allenrolfea occidentalis Atriplex corrugata Atriplex gracilifl Atriplex saccaria Chenopodium album berlandieri Chenopodium desiccatum C. pratericola Cactaceae Cactaceae Cactaceae Cactaceae Campanulaceae Boraginaceae Boraginaceae Boraginaceae FamilyAlismataceae Species name notes Synonyms/Taxonomic Common name Asclepiadaceae Asclepiadaceae Asclepiadaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Appendix B. Potential (unconfi

82 Annotated Checklist of Vascular Flora: Canyonlands National Park Range Comments Life form ower AnnF Intro rewheel blanketﬂ rewheel whitestem rabbitbrush Shrub Wide pretty rabbitbrush Shrub Periph Bigelow’s rabbitbrushBigelow’s Shrub RegEn small-leaf pussyoes wormwoodCarruth’s PerF PerF Wide Wide Wyoming big sagebrush Shrub Wide sand rabbitbrush Shrub RegEn oakleaf goosefoot AnnF Wide western spiderwortcommon ragweed PerF AnnF Wide Wide smooth asteralkali aster asterWelsh’s PerF Periph PerF PerF Wide LocEn sticky seepwillow beggarsticksdevil’s- dwarf rabbitbrush Shrub PerF Wide Shrub Wide Wide lanceleaf rabbitbrushchicory Shrubnoddinghead Wide spreading daisymarshweedﬁ PerF PerF PerF RegEn Wide Intro AnnF Native to Eurasia Sparse spiny goldenweed PerF RegEn

var.

da ssp. speciosa bigelovii arenaria by some var. var. var. elatior occidentalis orus albicaulis, var. var. var. geyeri, B. salicina var. A. rosea geyeri Ericameria pulchella, C. baileyi C. nauseosus Ericameria nauseosa Seriphidium tridentatum wyomingense Ericameria nauseosa C. salinum Includes Symphyotrichum laeve, S. laeve var. Included in authors Ericameria nauseosa E. colomexicanus Machaearanthera pinnatiﬁ paradoxa

var. var. var. var. var.

var.

orus var. var. oruspaucifl Almutaster lanceolatus rmed) taxa of Canyonlands National Park, cont. Chrysothamnus pulchellus baileyi Chrysothamnus nauseosus speciosus Artemisia carruthii Artemisia tridentata wyomingensis Aster laevis A. laevis Chrysothamnus nauseosus bigelovii salinum occidentalisTradescantia occidentalis T. Chenopodium glaucum Ambrosia artemisiifoliaAntennaria microphylla A. artemisiifolia Aster paucifl Aster welshiiBaccharis glutinosa Symphyotrichum welshii Bidens frondosa Chrysothamnus depressus Chrysothamnus nauseosus arenarius Chrysothamnus viscidifl var. Cichorium intybus Enceliopsis nutans Erigeron divergens cinereus Flaveria campestris Gaillardia pulchella Haplopappus spinulosus paradoxus Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Commelinaceae Appendix B. Potential (unconfi FamilyChenopodiaceae Species name notes Synonyms/Taxonomic Common name Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 83 USFWS: former C2 Candidate. USFWS: former C2 Candidate. Range Comments Life form ower PerF RegEn ower PerF Periph ower AnnF RegEn eld dodder eld AnnF Wide sand sunfl tidytipsArizona rush-pinkEntrada rush-pink PerF RegEn PerF AnnF Wide LocEn UT BLM Sensitive, thorny wire-lettuce AnnF Wide piliferous turtleback AnnF RegEn thickleaf asteralcove rock-daisy PerF PerFgreenstem paperfl RegEn woolly paperfl LocEn UT BLM Sensitive, Holboell’s rockcressHolboell’s rockcressThompson’s PerF Wide PerF LocEn western tansy-mustardwestern tansy-mustard AnnF AnnF Wide Wide Rocky Mountain groundsel PerF Wide desert nest-strawdogweed townsendiaBeaman’s AnnF AnnF Periph LocEn PerF Wide rough mule’s-earsfi evolvulus PerF Wide PerF Wide

var. secunda, scabra var. by some ssp. retrofracta, includes vars. lipes var. ﬁ considered a synonym oodes var. H. deserticola & dianthopsis L. arizonica L. entrada, of var. authors Pleiacanthus spinosus Includes M. leucanthemifolia Packera streptanthifolia streptanthifola, B. retrofracta borealis Boechera holboellii

var. lipesincisa D. var. var.

fi var. var. secunda A. holboellii ora ora ora var. leucanthemifolia rmed) taxa of Canyonlands National Park, cont. Layia glandulosa grandifl Lygodesmia arizonica grandifl Lygodesmia entrada Lygodesmia spinosaLygodesmia Machaeranthera canescens var. Stephanomeria spinosa, Psathyrotes pilifera Psilostrophe sparsifl Helianthus anomalus Perityle specuicola Psilostrophe tagetina Senecio streptanthifolius streptanthifolius Arabis thompsoniiDescurainia pinnata Boechera thompsonii Descurainia pinnata nelsonii Stylocline micropoides Thymophylla acerosa beamanii Townsendia Wyethia scabra Dyssodia acerosa Scabrethia scabra Cuscuta pentagona Evolvulus nuttallianus Arabis holboellii Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconfi FamilyCompositae (Asteraceae) Species name notes Synonyms/Taxonomic Common name Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Convolvulaceae (Cuscutaceae) Convolvulaceae Cruciferae (Brassicaceae)

84 Annotated Checklist of Vascular Flora: Canyonlands National Park entrance road leading into ISKY, just outside of CANY boundary. 3C Range Comments Life form mbristylis PerG Sparse at-sedge AnnG Wide eld’s bluestem eld’s PerG Sparse ax-leaved plainsmustard ax-leaved PerF Wide yellow bluestem PerGSpringfi Intro Occurs along Payson’s tansymustardPayson’s ovalpurse AnnF Periph prairie pepperworthairy-pod pepperwort AnnF bladderpodFendler’s AnnF AnnF Wide fl Wide Wide PerF Periph smoothleaf prince’s-plume PerFRocky Mountain juniper Wide Tree Wide hot springs fi Canyonlands sedge PerG RegEnalkali bulrush USFWS: Category tiny spurgethyme-leaved spurge PerGUtah dove-weed Wide Goodhope oak AnnF Wide AnnF AnnF Wide RegEn Tree LocEn bottlebrush sedgebearded fl PerG Wide oakEastwood’s centauryBuckley’s Great Basin centauryjointed goatgrass Shrub AnnF PerF RegEn Periph Wide AnnG Intro Native to Eurasia

var. paludosus, var. songarica curatorum var. integrifolia var. paysonii var. var. D. incisa Hymenolobus procumbens Schoenoplectus maritimus, Bolboschoenus maritimus paludosus

var. var. bonina var. eldii utahensis var. var. orum eastwoodiae x orum rmed) taxa of Canyonlands National Park, cont. Bothriochloa springfi Descurainia pinnata lasiocarpum paysonii Hutchinsia procumbensLepidium densifl densifl Hornungia procumbens, Lepidium lasiocarpum Physaria fendleriSchoenocrambe linifoliaStanleya integrifolia Sisymbrium linifolium Lesquerella fendleri S. pinnata Juniperus scopulorumCarex curatorum Sabina scopulorum Fimbristylis spadicea C. scirpoidea Scirpus maritimus thermalis F. Scirpus maritimus Chamaesyce micromeriaChamaesyce serpyllifoliaCroton texensis Euphorbia micromeria Quercus gambelii Euphorbia serpyllifolia Quercus Carex hystericinaCyperus squarrosus C. hystricina C. aristatus Centaurium calycosumCentaurium exaltatum Aegilops cylindrica C. arizonicum Bothriochloa ischaemum B. ischaemum Cylindropyrum cylindricum Gramineae (Poaceae) Appendix B. Potential (unconfi FamilyCruciferae (Brassicaceae) Species name notes Synonyms/Taxonomic Common name Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cupressaceae Cyperaceae Cyperaceae Cyperaceae Euphorbiaceae Euphorbiaceae Euphorbiaceae Fagaceae Fagaceae Cyperaceae Cyperaceae Gentianaceae Gentianaceae Gramineae (Poaceae) Gramineae (Poaceae)

Chapter 6: Appendices 85 3C 3C Range Comments Life form eld mint eld PerF Wide eld sandbur eld AnnG Wide Hayden’s milkvetchHayden’s milkvetchEastwood’s milkvetchDiehl’s PerF PerF Wide milkvetchGeyer’s panther milkvetch RegEn PerFFort Wingate milkvetch AnnF RegEn PerF PerF Wide LocEn RegEn USFWS Category ﬁ camel-thorn PerF Intro Native to Asia seaside arrowgrass PerG Wide jointed rushtoad rush PerG Sparse AnnG Wide ditch polypogonweeping alkaligrasscultivated rye PerGprairie wedgegrass PerG namaHowell’s Intro Intro PerG AnnG Native to Eurasia Wide Intro AnnF Native to Eurasia RegEn USFWS: Category ﬁ Great Basin wildryetufted lovegrasswolftail PerGgreen muhlylittle ricegrass Wide AnnG Wide PerG PerG PerG Wide Wide Wide Zion milkvetchcreeping rush-peawestern redbud PerF PerF RegEn RegEn Shrub Wide

& P. monspeliensis P. canadensis obtusata bufonius, geyeri var. var. var. T. concinnum T. Agrostis stolonifera A. haydenianus Includes vars. occidentalis, halophilus, ambiguus Includes Hybrid between and C. orbiculata

var. var. zionis glabrata M. arvensis var. var. var. var. exuosus rmed) taxa of Canyonlands National Park, cont. haydenianus Astragalus eastwoodiae Astragalus ﬂ diehlii Astragalus geyeriAstragalus pardalinus Astragalus wingatanus Astragalus zionis A. geyeri Alhagi maurorum Astragalus bisulcatus Mentha arvensis Juncus articulatus Juncus bufonius maritima Triglochin Puccinellia distans Secale cereale Sphenopholis obtusata Nama retrorsum S. obtusata Cenchrus longispinus Elymus cinereusEragrostis pectinacea phleoidesLycurus Muhlenbergia racemosa Oryzopsis exigua Leymus cinereus Polypogon interruptus L. setosus Piptatherum exiguum Caesalpinia repensCercis occidentalis orbiculata repens Hoffmanseggia Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Labiatae (Lamiaceae) Juncaceae Juncaceae Juncaginaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Hydrophyllaceae Appendix B. Potential (unconﬁ FamilyGramineae (Poaceae) Species name notes Synonyms/Taxonomic Common name Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Leguminosae (Fabaceae, Caesalpiniaceae) Leguminosae (Fabaceae, Caesalpiniaceae)

86 Annotated Checklist of Vascular Flora: Canyonlands National Park Cat # 45531) from just outside park near Horseshoe Canyon. Sensitive, USFWS: former C2 Candidate. USFWS: former C2 Candidate. Range Comments PerF Wide Life form reweed PerF Wide at top wild buckwheat Shrub LocEn BLM Sensitive, ﬁ hairy-throat evening- primrose western evening-primrose PerF Wide thorny milkwortcushion milkwort Shrub Periph PerF Wide Cox s.n. (CANY silverleaf lupineGreat Basin lupine PerF Wide AnnF Wide Lambert’s locoweedLambert’s PerF Wide narrow-leaf dunebroom breadrootTuhy’s Shrub RegEn rush scurfpea PerF indigo-bushFremont’s LocEn Shrub UT BLM: Wide PerF RegEn American vetch mariposaGunnison’s PerF PerF Wide Wide ﬂ bent wild buckwheat AnnF RegEn blackbrush lupine AnnF Wide

ssp. var. pusillus var. var. gunnisonii exum var. lambertii hirsutissima included in var. ssp. circumvagum, C. danielsii P. acanthoclada P. L. caudatus, argenteus argophyllus, L. caudatus cutleri Vars. not recognized in Barneby Vars. (1989). Chamerion angustifolium canescens, C. angustifolium E. smithii Includes var. by some authors

var. var. var.

var.

rubens L. rubens, var. var. var. var. var. exumfl Stenogonum lifolia rmed) taxa of Canyonlands National Park, cont. Oenothera coronopifolia Oenothera elataPolygala acanthoclada O. elata acanthoclada Polygala subspinosa Lupinus caudatus argophyllus Lupinus pusillus intermontanus Lupinus pusillus Parryella fi Pediomelum aromaticum tuhyi Psoralidium junceumPsorothamnus fremontii Vicia americana Psoralea juncea americana Calochortus gunnisoniiEpilobium angustifolium canescens C. gunnisonii Eriogonum fl Eriogonum corymbosum smithii bigelovii Oxytropis lambertii Onagraceae Onagraceae Polygalaceae Polygalaceae Polygonaceae Appendix B. Potential (unconfi FamilyLeguminosae (Fabaceae) Species name notes Synonyms/Taxonomic Common name Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Liliaceae (Calochortaceae) Onagraceae Polygonaceae Leguminosae (Fabaceae)

Chapter 6: Appendices 87 Range Comments Life form ower PerF Wide ower AnnF Wide y wild buckwheat Shrub RegEn yard knotweedbulblet bladder fern AnnF Fern Intro Periph Native to Eurasia San Juan wild buckwheat Shrubsulfur wild buckwheat RegEn PerF Wide brittle bladder fernUtah bladder fern Fernblister buttercup Wide Fern mountain-lilacFendler’s antelope bitterbrush RegEn Shrub AnnFNew Mexico thimbleberry Wide Shrub Wide Shrub Periph Wide Rio Grande cottonwood Treered willow Periph Shrub Wide annual paintbrushreddish monkeyﬂ AnnF Wide glabrous monkeyﬂ veined penstemonshortest penstemon penstemonMoffatt PerF PerF RegEn PerF RegEn RegEn Hooker’s wild buckwheatHooker’s AnnF Wide bush penstemon PerF Wide

ssp. according by Welsh et by Welsh wislizenii C. tenuis minor var. & C. fragilis of WY & Great Plains ssp. C. utahensis, aviculare Includes reevesiana, to some authors Included in al. (2003) Hecatonia scelerata deltoides P. Misspelled in Welsh et al. (2003)Misspelled in Welsh butterﬂ Leiostemon ambiguus Sometimes confused with ssp. caudatus

var. var. var. var.

var. var.

var. var. dus rmed) taxa of Canyonlands National Park, cont. Cystopteris bulbifera Cystopteris fragilis Eriogonum leptocladon ramosissimum Eriogonum umbellatum umbellatum Polygonum aviculare aviculare arenastrum, P. P. Cystopteris utahensis Ranunculus sceleratus multiﬁ Ceanothus fendleri Purshia tridentata Rubus neomexicanus Populus fremontii wislizenii Salix laevigata Castilleja exilisPenstemon ambiguus laevissimus C. minor Mimulus glabratus Mimulus rubellus utahensis Penstemon breviculus Penstemon moffatii Eriogonum hookeri Eriogonum leptocladon papiliunculi Penstemon angustifolius venosus Polypodiaceae (Aspleniaceae) Polypodiaceae (Aspleniaceae) Polygonaceae Polygonaceae Polygonaceae Polypodiaceae (Aspleniaceae) Ranunculaceae Rhamnaceae Rosaceae Rosaceae Salicaceae Salicaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Appendix B. Potential (unconﬁ FamilyPolygonaceae Species name notes Synonyms/Taxonomic Common name Polygonaceae Scrophulariaceae

88 Annotated Checklist of Vascular Flora: Canyonlands National Park

rmed for the Sensitive, UT BLM Sensitive, USFWS: Former C2 candidate USFWS: former C2 Candidate. Atlas of the Vascular Flora of Utah Atlas of the Vascular Range Comments PerF Wide Life form d here. Brigham Young University, in 2001, and the University, Brigham Young Palmer’s penstemonPalmer’s PerF RegEn spring-parsleyTrotter’s PerFtobacco-root LocEnthicket-creeper UT BLM Sensitive, PerF Shrub Wide Wide blunt-leaf spike-mosscoyote tobaccopurple ground-cherry Fernbasin white-cup spring- parsley Wide PerFCanyonlands lomatium AnnF Disj Wide PerF LocEn USFS Region 4 eld, Grand, San Juan, and Wayne counties, Utah, but have not previously been reported or conﬁ been reported counties, Utah, but have not previously eld, Grand, San Juan, and Wayne mutica var.

var. rmed) taxa of Canyonlands National Park, cont. eglandulosus Selaginella mutica S. mutica Oreoxis trotteri edulis Valeriana Parthenocissus vitacea Cymopterus trotteri Penstemon palmeri Nicotiana attenuata Physalis lobataCymopterus purpurascens Lomatium latilobum Quincula lobata Aletes latilobus Selaginellaceae Umbelliferae (Apiaceae) Valerianaceae Vitaceae Appendix B. Potential (unconﬁ FamilyScrophulariaceae Species name notes Synonyms/Taxonomic Common name Solanaceae Solanaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) The vascular plant species listed here are known from the vicinity of Canyonlands NP, in Garﬁ the vicinity of Canyonlands NP, known from are The vascular plant species listed here N. Duane Atwood, of by Dr. a checklist of potential species for NCPN parks prepared derived from park. Species on this list are not include as potentially occurring in the park, but are Appendix C could be considered (Albee et al. 1988). Some species from

Chapter 6: Appendices 89 . C. C. O. A. C. tenuis

polyacantha. eastwoodiae. ed specimens of ed specimen of ed specimens of ed specimens of ed specimens of var. var. . . USFWS Category 3C var. var. Comments CANY is well outside the known range of this taxon. on an observation by Boschen in 1984, but CANY is outside the known range of this species. on misidentifi osterhoutii on misidentifi locality is Huntington Road, ca Type 15 mi W of US 50-6, Emery Co., UT USFWS: 3C (Higgins 1310 BRY). on misidentifi osterhoutii based on a specimen collected outside the park. on misidentifi polyacantha CANY is outside the known range of this taxon. CANY is outside the known range of this taxon. on misidentifi fendlerii is outside the known range of this taxon. Reports probably based on var. eastwoodiae. Park status Range Life form PerF Sparse Ques? Reported by Schelz et al. (2006), but Common name clasping-leaved dogbane swamp milkweed PerFdwarf cryptanth Wide Ques? PerF Reported in Schelz et al. (2006) based Wide FalsRep Reports from CANY Herbarium based Johnston’s cryptanthJohnston’s PerF LocEnParadox cryptanth FalsRep Report from CANY Herbarium based PerF RegEn FalsRep Reports from CANY Herbarium based salt heliotropebrittle prickly-pear PerF PerF Wide Wide FalsRep Report from Schelz et al. (2006) is FalsRep Reports from CANY Herbarium based littoral pricklypear PerF chollaWhipple’s RegEn Ques? Shrubball-head sandwort RegEn Reported by Schelz et al. (2006), but PerF Ques? Wide Reported by Schelz et al. (2006), but FalsRep Reports from CANY Herbarium based Fendler’s sandwortFendler’s PerF Periph Ques? Reported by NCPN (in ed.), but CANY

A. var. nana congesta, Synonyms/ notes Taxonomic hypericifolium Includes reports of fascicularis ssp. O. martiniana whipplei Eremogone congesta var. includes var. cephaloidea Eremogone fendleri

var. var.

var. congesta Apocynum sibiricum A. cannabinum Asclepias incarnata Cryptantha humilis Oreocarya humilis Cryptantha johnstonii Cryptantha paradoxa Oreocarya paradoxa Heliotropium curassavicum obovatum Opuntia fragilis Opuntia littoralis martiniana Opuntia whipplei Cylindropuntia Arenaria congesta var. Arenaria fendleri fendleri FamilyApocynaceae Species name Asclepiadaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Cactaceae Cactaceae Cactaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Canyonlands National Park. Appendix C. Rejected (falsely reported

90 Annotated Checklist of Vascular Flora: Canyonlands National Park S. A. B. C. A. rosea. Atriplex ed specimens of ed specimen of ed specimen of pulchellum. torreyana. ed specimens of ed specimens of var. var. ed specimen of var. var. . . A. pansus. Comments on misidentifi calcareum on misidentifi misidentifi dioica based on misidentifi torreyana on a misidentifi marginata. CANY is well outside the known range of this taxon. Reports might be based on CANY is well outside the known range of this taxon. on a misidentifi cernua CANY is well outside the known locality is range of this taxon. Type W of Leeds along road to Silver Reef Co., UT Campgrounds, Washington (Semple & Heard 7859 NY). but CANY is outside mapping crew, the known range of this taxon. Type locality is Canaan Ranch, Kane Co., UT (Jones 6066c POM). Park status Range Life form PerF SparsePerF Ques? Periph Reported by Schelz et al. (2006), but FalsRep Report from Rink 4607 (CANY) is based Common name Arizona thistle PerF Wide FalsRep Reports from CANY Herbarium based silver orach saltbushSuckley’s AnnF AnnF Wide Wide FalsRep FalsRep Report by Rink based on a Reports from CANY Herbarium based western seepweed AnnF Wide FalsRep Reports from CANY Herbarium are common pussytoes PerF Wide FalsRep Report from BYU herbarium is based falcate aster PerF Sparse Ques? Reported by Schelz et al. (2006), but Zion golden-aster PerF RegEn Ques? Reported by Schelz et al. (2006), but straw-stem beggarticks three-lobed beggarticks dune rabbitbrush Shrub RegEn Ques? Reported by NCPN (in ed.) vegetation

foliosa, & S. B. var. foliosa by some var. var. var. turbinata commutatus, Synonyms/ notes Taxonomic Includes vars. arizonicum nidulum some authors Included in calceoliformis falcatus, A. ericoides var. Symphyotrichum falcatum falcatum, Virgulus falcatus H. zionensis, villosa Included in tripartita Ericameria nauseosa var. authors

var. var. var. Endolepis dioica Atriplex suckleyi Atriplex argentea argentea Suaeda occidentalis Antennaria parvifolia Aster falcatus A. falcatus Chrysopsis zionensis C. villosa Bidens comosa Bidens tripartita Chrysothamnus nauseosus turbinatus Cirsium arizonicum Chenopodiaceae FamilyChenopodiaceae Species name Chenopodiaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix C. Rejected (falsely reported or questionable) taxa of Canyonlands National Park, cont.

Chapter 6: Appendices 91

D. C. M. H. G. Type locality Type . ed specimens of ed specimens of utahense. var. aristata. ed specimens of ed specimen of ed specimens of ed specimens of ed specimens of . ivesiana. . var. var. var. eld Co., UT (Jones 5695y US). H. armerioides. Comments on misidentifi brandegeei based on misidentifi Chaetopappa ericoides. on misidentifi marginata CANY is outside the known range of this taxon. CANY is well outside the known range of this species. Reports probably based on is Bromide Pass, Henry Mountains, Garfi on misidentifi neomexicanum CANY is well outside the known range of this taxon. based on misidentifi Stephanomeria exigua on misidentifi canescens CANY is outside the known range of this taxon. on misidentifi acaulis Park status Range Life form ower PerF Wide Ques? Reported by Schelz et al. (2006), but gray sandplant AnnF Periph FalsRep Reports from CANY Herbarium based silver daisysetose crustweed PerF AnnF Wide Periph FalsRep FalsRep Reports from CANY Herbarium Reports from CANY Herbarium based Rydberg’s gumweedRydberg’s PerF Periph Ques? Reported by Schelz et al. (2006), but stemless goldenweed PerF Wide Ques? Reported by Schelz et al. (2006), but Common name wavy-leaved thistle PerF Wide FalsRep Reports from CANY Herbarium based slender goldenweed AnnFlittle sunfl Periph Ques? Reported by Schelz et al. (2006), but rush skeleton-plant PerF Periph FalsRep Reports from CANY Herbarium are Bigelow’s asterBigelow’s PerF RegEn FalsRep Reports from CANY Herbarium based Arizona hymenoxys PerF Wide FalsRep Reports from CANY Herbarium based

var.

Included var. var. (2006). (2006) arizonica Flora of North G. squarrosa Flora of North Vars. not recognized Vars. in America in aphanactis. in America Stenotus acaulis acaulis Synonyms/ notes Taxonomic C. undulatum gracilis, Xanthisma gracile M. bigelovii commixta, Dieteria bigelovii commixta Tetraneuris acaulis Tetraneuris var.

ssp. ora acaulis undulatum arizonica canescens Erigeron argentatus Glyptopleura setulosa Grindelia aphanactis G. nuda Haplopappus acaulis var. Cirsium undulatum var. Dicoria canescens Haplopappus gracilis Machaeranthera Helianthella uniﬂ Lygodesmia juncea Lygodesmia Machaeranthera commixta Hymenoxys acaulis var. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix C. Rejected (falsely reported or questionable) taxa of Canyonlands National Park, cont.

92 Annotated Checklist of Vascular Flora: Canyonlands National Park A. P. C. L. C. . ed specimens of ed specimens of desertorum. C. parryana perennans. ed specimens of ed specimens of are now thought to be var. var. var. var. and according to Sherel Goodrich CANY is outside the known range of this taxon. CANY is well outside the known range of this taxon. Native to Europe. but CANY is outside mapping crew, the known range of this taxon. based on misidentifi perennans CANY is well outside the known range of this taxon. Native to Eurasia. CANY is outside the known range of this taxon. CANY is well out of the expected range of this species. herbarium is from outside the park. on misidentifi latifolium. on misidentifi Comments specuicola based on misidentifi integrifolia C. specuicola parryana et al. 2003). USFWS: (in Welsh Threatened Park status Range PerF Wide Ques? Reported by Schelz et al. (2006), but AnnF Wide FalsRep Reports from CANY Herbarium based Life form showy goldeneye PerF Wide Ques? Reported by Schelz et al. (2006), but pale madworthairy rockcress AnnF Introsmall-leaf rockcress PerF Ques? PerF WideEuropean wintercress Reported by Schelz et al. (2006), but Wide PerF Ques? FalsRep IntroCalifornia tansy- Reported by NCPN (in ed.) vegetation mustard Reports from CANY Herbaroum are Ques? pepperwortFremont’s PerF Reported by Schelz et al. (2006), but Periphclasping pepperwort Ques? AnnF poor-man’s Intro Reported in Schelz et al. 2006, but pepperwort FalsRep Native to Europe. Collection in CANY beautiful sedge PerG Wide FalsRep Reports from CANY Herbarium based Common name San Juan bahia PerF RegEn FalsRep Reports from CANY Herbarium are Navajo sedge PerG LocEn FalsRep Utah specimens previously considered

ora var. ora & multiﬂ microphylla Heliomeris multiﬂ var. Includes vars. glabrata pycnocarpa Boechera microphylla var. Synonyms/ notes Taxonomic Bahia nudicaulis oblongifolia

ora var. ora microphylla pubescens multiﬂ Alyssum alyssoides Arabis hirsuta Arabis microphylla var. Barbarea vulgaris Descurainia californica Lepidium fremontii Lepidium perfoliatum Lepidium virginicum var. Carex bella Viguiera multiﬂ var. Platyschkuhria integrifolia oblongifolia Carex specuicola Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cyperaceae Compositae (Asteraceae) FamilyCompositae (Asteraceae) Species name Cyperaceae Appendix C. Rejected (falsely reported or questionable) taxa of Canyonlands National Park, cont.

Chapter 6: Appendices 93 S. S. S. . C. B. Flora of ed specimens of ed specimens of A. purpurea (2003). CANY reports parishii. longispicatus. ed specimens of ed specimens of . . var. var. var. var. Comments (2006), but CANY is well outside the known range of this taxon. CANY is outside the known range of this taxon. on misidentifi pungens praegracilis. is well outside the known range of this taxon. Report probably based on on misidentifi japonicus CANY is outside the known range of this taxon. (2006), but CANY is well outside the known range of this taxon. CANY is well outside the known range of this taxon. based on misidentifi coronata based on misidentifi utahensis but this species is not considered to occur in Utah according to North America probably based on Park status Range Life form PerG RegEnPerG Ques? Wide Reported for CANY by Schelz et al. Ques? Reported by Schelz et al. (2006), but PerF Wide FalsRep Reports from CANY Herbarium are crenulate phacelia AnnF Wide Ques? Reported for CANY by Schelz et al. Common name Missouri milkvetch PerF RegEn Ques? Reported by Schelz et al. (2006), but rusty sedge PerG Sparse FalsRep Reports from CANY Herbarium based American bulrush PerG Wide Ques? Reported by Loope (1977), but CANY mountain brome PerG Wide FalsRep Reports from CANY Herbarium based tumble grass PerG Sparse Ques? Reported by Schelz et al. (2006), but depauperate needlegrass Letterman’s Letterman’s needlegrass Scribner needlegrass PerG Periph FalsRep Reports from CANY Herbarium are white-margined swertia Arizona three-awn PerG Periph Ques? Reported by Schelz et al. (2006), depauperatum, Synonyms/ notes Taxonomic Schoenoplectus americanus B. polyanthus, Ceratochloa carinata Achnatherum parishii ssp. A. coronatum lettermanii scribneri Frasera albomarginata

var. var. crenulata Phacelia crenulata var. Astragalus missouriensis amphibolus Carex subfusca Scirpus americanus Scirpus olneyi, Bromus carinatus B. marginatus, Schedonnardus paniculatus Stipa coronata depauperata Stipa lettermanii Achnatherum Stipa scribneri Achnatherum Swertia albomarginata Aristida arizonica Hydrophyllaceae Leguminosae (Fabaceae) FamilyCyperaceae Species name Cyperaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gentianaceae Gramineae (Poaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Canyonlands National Park, cont.

94 Annotated Checklist of Vascular Flora: Canyonlands National Park A. A. Z. ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of praelongus. . . . . var. var.

Comments based on misidentifi M. albicaulis based on misidentifi M. marginata CANY is outside the known range of this taxon. Report probably based on lanceolatum. P. based on misidentifi macropetalum based on misidentifi vaginatus. based on misidentifi M. marginata CANY is outside the known range of this taxon. based on misidentifi praelongus CANY is well outside the known range of this taxon. Native to Europe. CANY is outside the known range of this taxon. on specimen from CANY herbarium collected well outside the park. Type locality is W of Carmel, Kane Co., UT (Ferguson & Ottley 5613 NY). Park status Range Life form PerF Wide Ques? Reported by Schelz et al. (2006), but PerF Wide FalsRep Reports from CANY Herbarium are owered ower sweetpea PerF RegEn Ques? Reported by Schelz et al. (2006), but curvepod stickleaf AnnFwing-seed stickleaf Periph PerF FalsRep Reports from CANY Herbarium are Wide FalsRep Reports from CANY Herbarium are Wyoming stickleaf PerF RegEn FalsRep Reports from CANY Herbarium are slender-fl scurf-pea textile onionmountain death camas PerF Wide FalsRep Reports from CANY Herbarium are pepperwort Fern Wide Ques? Reported by Schelz et al. (2006), but Common name Patterson’s milkvetchPatterson’s PerFlicorice RegEn FalsRep Reports from the CANY Herbarium are whitefl PerFUtah trefoil Intro Ques? PerF Reported by Schelz et al. (2006), but RegEn FalsRep Report from Schelz et al. 2006 based ora vestita jonesii var. var. Nuttallia pterosperma M. pumila Psoralea tenuifl Synonyms/ notes Taxonomic L. leucanthus

var. orum leucanthus Mentzelia nitens M. nitens Mentzelia pterosperma Mentzelia pumila pumila Allium textile Zigadenus elegans Anticlea elegans Marsilea vestita M. vestita Glycyrrhiza glabra Lathyrus lanszwertii var. Lotus utahensis Psoralidium tenuiﬂ Astragalus pattersonii Loasaceae Loasaceae Loasaceae Liliaceae (Alliaceae) Liliaceae (Melanthiaceae) Marsileaceae Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) FamilyLeguminosae (Fabaceae) Species name Appendix C. Rejected (falsely reported or questionable) taxa of Canyonlands National Park, cont.

Chapter 6: Appendices 95

G. E. P. hoodii P. ed specimen of . Varieties not . Varieties ed specimens of ed specimens of ed specimens of ed specimens of . Flora of North America ed specimens of . . Type locality is Springdale, . Type . rmation as CANY is well beyond canescens. based on misidentifi Habenaria zothecina recognized in 2003. confi the expected range of this species. based on a misidentifi Ipomopsis pumila based on misidentifi imperialis Comments based on misidentifi I. pumila Co., UT (Jones 5247 NY). Washington on misidentifi CANY is outside the known range of this taxon. CANY is outside the known range of this taxon. CANY is well outside the known range locality is “northern of this taxon. Type Arizona near Kanab” from Coconino Co., AZ or Kane UT (Thompson s.n. GH). based on misidentifi ciliatum var. var. Park status Range Life form Shrub RegEn Ques? Reported by Schelz et al. (2006), but slender bog orchid PerF Wide FalsRep Reports from CANY Herbarium singleleaf pinyon Treespreading eriastrum Wide AnnFspiny gilia Wide Ques? Report from Schelz et al. 2006 needs FalsRep Report by Moran from CANY is AnnF RegEn FalsRep Reports from CANY Herbarium are shrubby giliadesert phlox PerF RegEn FalsRep PerF Reports from CANY Herbarium are Wide FalsRep Reports from CANY Herbarium based Common name longleaf wild buckwheat pale evening-primrose PerF RegEn Ques? Reported by Schelz et al. (2006), but manylobe camissonia AnnF RegEn Ques?alpine willow-herb Reported by Schelz et al. (2006), but PerF Sparse FalsRep Reports from CANY Herbarium are

var. var. var.

Flora Vars. Vars. var. . Vars. not . Vars. (2005) ora Flora of North recognized in of North America (2002) latifolia, Aliciella latifolia Platanthera sparsiﬂ Gilia congesta frutescens not recognized Var. in Cronquist et al. (1984) Synonyms/ notes Taxonomic E. intermontanum, E. corymbosum humivagans. not recognized in America E. anagallidifolium

ora var. ora frutescens austromontana trichocalyx sparsiﬂ alpinum Pinus monophylla Eriastrum diffusum Gilia latifolia G. latifolia Ipomopsis congesta var. Phlox austromontana var. lonchophyllum Oenothera pallida var. Habenaria sparsiﬂ var. Epilobium alpinum var. Camissonia multijuga Eriogonum lonchophyllum Pinaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Onagraceae Orchidaceae Onagraceae FamilyOnagraceae Species name Polygonaceae Appendix C. Rejected (falsely reported or questionable) taxa of Canyonlands National Park, cont.

96 Annotated Checklist of Vascular Flora: Canyonlands National Park G. E. E. C. A. P. is Salix . occidentalis. ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of simpsonii. coloradoense. var. var. cernnum. E. watsonii var. var. ed specimens of var. var. . var. var. . orum Comments based on misidentifi multifl based on misidentifi microthecum based on misidentifi cernuum CANY is outside the known range of this taxon. based on misidentifi Physocarpus alternans based on misidentifi wrightii. based on misidentifi utahensis based on misidentifi paradoxa CANY is outside the known range of this taxon. on misidentifi not known from Utah, but is native to Nevada, Idaho, and Oregon. based on misidentifi Pellaea glabella gooddingii. CANY is outside the known range of this taxon. Park status Range Life form Shrub WideAnnF FalsRep RegEn Reports from CANY Herbarium are FalsRep Reports from CANY Herbarium are AnnF Wide FalsRep Reports from CANY Herbarium are ower bird’s- ower Common name cleavers AnnF Wide FalsRep Reports from CANY Herbarium are whiplash willow Shrub Wide Ques? Reported by Schelz et al. (2006), but slender wild buckwheat wild Watson’s buckwheat wax currant Shrub Wide FalsRep Reports from CANY Herbarium smallfl beak Saskatoon serviceberrySaskatoon Shrub Widebiennial cinquefoil FalsRep AnnF Reports from CANY Herbarium are Wide FalsRepCarolina poplar Reports from CANY Herbarium are peach-leaf willow Tree Intro Tree Ques? Wide Reported by Schelz et al. (2006), but FalsRep Reports from CANY Herbarium based Brewer’s cliff-brakeBrewer’s Fern Wide FalsRep Reports from CANY Herbarium are scarlet paintbrush PerF Wide Ques? Reported by Schelz et al. (2006), but

var. var. var. P. deltoides P. exum P. nigra P. Synonyms/ notes Taxonomic G. spurium x Hybird of S. lasiandra caudata watsonii R. cereum pedicellare

var.

var. var. ssp. canadensis x orus orum echinospermon Populus Eriogonum microthecum laxifl caudata Eriogonum watsonii E. defl inebrians Ribes cereum Amelanchier alnifolia Potentilla biennis Galium aparine Salix amygdaloides Pellaea breweri Salix lucida Castilleja miniata Cordylanthus parvifl Salicaceae FamilyPolygonaceae Species name Polygonaceae Saxifragaceae (Grossulariaceae) Rosaceae Rosaceae Rubiaceae Salicaceae Polypodiaceae (Adiantaceae) Salicaceae Scrophulariaceae Scrophulariaceae Appendix C. Rejected (falsely reported or questionable) taxa of Canyonlands National Park, cont.

Chapter 6: Appendices 97 C. P. M. C. Type Type D. wrightii. ed specimens of ed specimens of petraeus. var. var. ed specimens of ed specimens of . . Native to E North America. Comments locality is Fruita, Capitol Reef NP, locality is Fruita, Capitol Reef NP, USFS Co., UT (Beck s.n. BRY). Wayne Region 4 Sensitive, UT BLM USFWS former C2 candidate. on misidentifi eastwoodiae. CANY is outside the known range of this taxon. outside the known range of this taxon. Report probably based on Native to tropical America. no voucher seen in CANY herbarium. Species is not known to occur in Utah et al. 2003). (Welsh based on misidentifi reticulata on misidentifi (2006), but CANY is well outside the known range of this taxon. based on specimens collected outside the park. CANY is outside the known range of this taxon. based on misidentifi strictiformis terebinthinus Park status Range Life form PerF Wide Ques? Reported in Schelz et al. 2006, but PerF RegEnPerF FalsRep Reports from CANY Herbarium are Wide Ques? Reported by Schelz et al. (2006), but ower PerF Wide FalsRep Reports from CANY Herbarium based Common name yellow monkeyfl beardlip penstemon PerF Wide Ques? Reported by Schelz et al. (2006), but jimson weedVirginia ground- AnnFcherry Intro Ques?common hackberry Reported by Loope (1977) but CANY is Tree spring-parsleyBeck’s Intro PerF FalsRep LocEn Reports from CANY Herbarium are FalsRep Reports from CANY Herbarium based Fremont’s penstemonFremont’s PerF RegEn Ques? Reported for CANY by Schelz et al. Natural Bridges penstemon Rocky Mountain penstemon penstemonWatson’s PerF Wide FalsRep Reports from CANY Herbarium are

torreyi var. Synonyms/ notes Taxonomic Includes var. fremontii barbatus Mimulus guttatus Penstemon barbatus var. Penstemon fremontii fremontii P. Datura stramonium Physalis virginiana Celtis occidentalis Cymopterus beckii Penstemon strictiformis Penstemon strictus Penstemon watsonii FamilyScrophulariaceae Species name Scrophulariaceae Scrophulariaceae Solanaceae Solanaceae Ulmaceae (Celtidaceae) Umbelliferae (Apiaceae) Scrophulariaceae Scrophulariaceae Scrophulariaceae Appendix C. Rejected (falsely reported or questionable) taxa of Canyonlands National Park, cont.

98 Annotated Checklist of Vascular Flora: Canyonlands National Park f the species (Questionable or exilis. Comments CANY is well outside the known range of this taxon. Native to W North America. CANY is well outside the known range parasitic on of this taxon. Typically Pinus fl Park status et al. 2006) or from herbarium specimens, but are now believed to be er- herbarium specimens, but are et al. 2006) or from Range Life form PerF Sparse Ques? Reported by Schelz et al. (2006), but matted fog-fruit PerFlimber pine dwarf- Intromistletoe Ques? Reported by Schelz et al. (2006), but Common name cyanocarpum A. campylopodium var. Synonyms/ notes Taxonomic ed or made from outside the park (False Reports or FalsRep), because is well known expected range o ed or made from ora Arceuthobium cyanocarpum Phyla nodifl Viscaceae FamilyVerbenaceae Species name Appendix C. Rejected (falsely reported or questionable) taxa of Canyonlands National Park, cont. The vascular plant species listed here have been reported for Canyonlands NP from published and unpublished literature (Schelz published and unpublished literature for Canyonlands NP from have been reported The vascular plant species listed here misidentifi because collections were roneous Ques?).

Chapter 6: Appendices 99 r r r r forest forest forest forest Disturbed areas Sandstone bedrock Shady rock crevices X X X X X Wet Comments XX X X Jun Pin- X X X SgeGrs XX Des Salt Br Blk- Flower time Park status Native Pres Apr–May r Native Pres NA Douglas-fi r Native Pres NA Douglas-fi bigtooth maple Native Pres Apr–May Douglas-fi box elder Native Pres Apr–Jun Siberian elm Intro Rep Mar–Apr lance-leaf cottonwood Native black Goodding’s willow Presnetleaf hackberry Apr–Jun Native Pres Apr–May X silk-treewhite mulberrywhite fi Introtwo-needle pinyon Intro ponderosa pine Pres Native RepDouglas-fi Jun–Aug Pres Apr–Jun Nativepeach NA Preswhite poplarFremont cottonwood NA Native Pres Intro Intro Mar–Jun Rep Rep Mar–Jun Apr–May Disturbed areas Cultivated Douglas-fi Disturbed areas Cultivated water birchwestern hop-hornbeam Native Pres Native Jul Pres Apr–May X Hanging garden common catalpaRussian-olive oakGambel’s Intro Rep Intro Native Jun–Jul Pres Pres May–Jul Apr–Jun Cultivated X

var. var. var. intercurrens x glauca Acer grandidentatum Acer negundo Ulmus pumila Salix gooddingii Populus Celtis reticulata Morus alba Abies concolor Populus alba Populus fremontii Albizia julibrissin Pinus edulis Pinus ponderosa scopulorum Pseudotsuga menziesii var. Prunus persica fremontii Ostrya knowltonii Betula occidentalis Elaeagnus angustifolia Quercus gambelii gambelii Catalpa bignonioides taxa of Canyonlands National Park, organized by life form with ecological notes. rmed, historical, and reported FamilyTrees Aceraceae Species name Common name Nativity Aceraceae Ulmaceae Salicaceae Salicaceae Ulmaceae (Celtidaceae) Moraceae Pinaceae Salicaceae Salicaceae Leguminosae (Fabaceae, Mimosaceae) Pinaceae Pinaceae Pinaceae Rosaceae Betulaceae Betulaceae Elaeagnaceae Fagaceae Bignoniaceae Appendix D. Conﬁ

100 Annotated Checklist of Vascular Flora: Canyonlands National Park r r forest forest X X X Wet Comments X X X X XX XX XX X X X X X Jun Pin- X X X X SgeGrs X X X XX XX XX XX XX Des Salt XX X XX XX X XX X XX X Br Blk- Flower time Park status ower snowberry ower Native Pres May–Jul Rocky Mountain maple Native Pres Apr–Jun Douglas-fi narrowleaved yucca Native Pres May–Jul squawbush Native Pres May–Jul western poison ivy barberryFendler’s mahoniaFremont’s Native Native Pres Native Pres May–Jun Pres May–Jun Apr–Jul X X X datil yucca yuccaHarriman’s yuccaToft’s Native Native Pressquawbush Pres Apr–May Native Apr–Jun Pres May–Jul NativeOregon-grape Preslong-fl May–Jul mountain-lover Nativefourwing saltbush Rep Native Apr–Jun shadscale Native Pres saltbushCastle Valley Pres May–Aug Native saltbushGarrett’s May–Sep Presspiny hopsage Native May–Sep Nativewinterfat Pres PresRuin Park winterfat May–Sep Native May–Sep greasewood Native Pres Native Pres Apr–Jul Pres May–Jul Native May–Jul Douglas-fi Pres May–Aug

var. var. var.

var. var. var. var. orus lanata ruinina Acer glabrum glabrum Yucca angustissima Yucca Rhus aromatica simplicifolia Berberis fendleri Mahonia fremontii Toxicodendron rydbergii Toxicodendron Yucca baccata Yucca harrimaniae Yucca harrimaniae toftiae Yucca Rhus aromatica trilobata Mahonia repens Symphoricarpos longiﬂ Pachystima myrsinites Atriplex canescens canescens Atriplex confertifolia Atriplex gardneri cuneata Atriplex garrettii garrettii Grayia spinosa Krascheninnikovia lanata var. Krascheninnikovia lanata var. Sarcobatus vermiculatus rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyShrubs Aceraceae Species name Common name Nativity Agavaceae Anacardiaceae Berberidaceae Berberidaceae Anacardiaceae Agavaceae Agavaceae Agavaceae Anacardiaceae Berberidaceae Caprifoliaceae Celastraceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae (Sarcobataceae)

Chapter 6: Appendices 101 Rock crevices Rocky areas Slickrock X X X X X Wet Comments XX X X X X X X X XX X Jun Pin- X X SgeGrs X X XX XX XX XX XX XX XX XX Des Salt X Br Blk- Flower time Park status Bigelow’s sagebrushBigelow’s Native Pres Aug–Oct Brandegee’s siltbushBrandegee’s Native Pres Jun–Aug Atwood’s mule-fatAtwood’s California brickellbush Native Native Pres Pres May–Jun rough brickellbush Aug–Sep Native Pres Aug–Oct greenish rabbitbrush Native Pres Aug–Oct fringed sagebrushblack sagebrush Nativebudsage Pres Native Jul–Sep basin big sagebrush RepRio Grande seepwillow Native Sep–Oct Native Pres Native Pres Pres Jul–Sep May–Nov Apr–Jun longleaf brickellbush Native Pres Aug–Sep rabbitbrushspear-leaf Native Presgraystem rabbitbrush Jul–Sep Nativeglabrate rabbitbrush Pres Native Aug–Oct Pres Aug–Oct sand sagebrush Native Pres Aug–Oct Mohave brickellbush Native Pres Jun–Aug

var. var.

var. nova var. var. var. lifolia consimilis gnaphalodes graveolens Artemisia bigelovii Zuckia brandegeei brandegeei Baccharis viminea atwoodii Brickellia californica Brickellia microphylla scabra Chrysothamnus nauseosus var. Artemisia frigida Artemisia nova Artemisia spinescens Artemisia tridentata tridentata Baccharis salicina Brickellia longifolia Chrysothamnus linifolius Chrysothamnus nauseosus var. Chrysothamnus nauseosus var. Artemisia ﬁ Brickellia oblongifolia linifolia rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) FamilyShrubs, cont. Chenopodiaceae Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Conﬁ

102 Annotated Checklist of Vascular Flora: Canyonlands National Park r forest Slickrock Hanging gardens X X Wet Comments X X X X X X X XX X Jun Pin- XX X X SgeGrs X XX XX XX XX XX XX XX XX XX Des Salt X XX X Br Blk- Flower time Park status Native Pres Aug–Oct Native Pres Aug–Sep Douglas-ﬁ green rabbitbrush Native Rep Jul–Sep rush rabbitbrush Native Pres Aug–Oct red-osier dogwoodspiny greasebush Native Pres Native May–Jul Pres Mar–Jun X smoothseed rabbitbrush thread snakeweedbroom snakeweed Native goldenbush PresRusby’s Native Jul–Oct Prescanyon spindly Nativegoldenbush Rep Jul–Oct spindly goldenbush Aug–Sep copperweed Native Pres Aug–Sep Native Pres Jun–Oct Utah juniperroundleaf buffaloberry Native Pres Native Mar–May Pres NA bush encelia Native Pres Apr–Jun slenderleaf rabbitbrush Native Pres Jul–Sep resinbush Native Pres Jul–Sep spineless horsebrush Native Rep Jun–Sep arrowweed Native Pres May–Jun

sericea var.

var. var. var. orus orus orus junceus meionandrum leiospermus canonis hirtellus Chrysothamnus nauseosus var. Cornus sericea Glossopetalon spinescens var. Chrysothamnus nauseosus var. Chrysothamnus viscidifl viscidifl Encelia frutescens Gutierrezia microcephala Gutierrezia sarothrae Haplopappus rusbyi Haplopappus scopulorum var. Haplopappus scopulorum var. Oxytenia acerosa Juniperus osteosperma Shepherdia rotundifolia resinosa Chrysothamnus viscidifl stenophyllus Vanclevea stylosa Vanclevea Tetradymia canescens Tetradymia Pluchea sericea rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyShrubs, cont. Compositae (Asteraceae) Species name Common name Nativity Cornaceae Crossosomataceae (Celastraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Cupressaceae Elaeagnaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 103 r forest Slickrock Slickrock Wet Comments X X X X X XX X X X X X X XX X Jun Pin- X X X SgeGrs XX XX XX XX XX XX XX XX XX XX XX Des Salt X X X XX XX X XX Br Blk- Flower time Park status Native Rep May–Aug Native Pres Jul–Oct Native Pres Apr–Jun Native Pres May–Jun Native Rep May–Jun Thompson’s indigo- Thompson’s bush desert olivesingleleaf ash Native Native Pres Pres Apr–May Apr–May X Torrey’s ephedraTorrey’s Native Pres NA Apache plume Native Pres Apr–Aug green ephedra ephedraCutler’s turbinella live-oakshinnery oak Native Nativepurple sage Native Pres Pres Rep NA NA Native Apr–Jun Pres Native Pres Apr–Jun prickly-phlox May–Jun crisp-leaf wild buckwheat rimrock wild buckwheat Native Native Pres Pres X Aug–Oct desert-lilacFranklin’s May–Jul Nativebirchleaf buckthornUtah serviceberry Pres Nativedwarf mountain Apr–May mahogany Pres Native May–Jun Pres Apr–Jun blackbrushbush oceanspray Native Native Pres Pres Apr–Jun Jun–Aug X Rock crevices Douglas-ﬁ sand wild buckwheat Native Pres Jun–Oct mountain alder-leaf mahogany curl-leaf mountain mahogany

var. var. viridis viscida

var. var. var. corymbosum orbiculatum leptocladon Fraxinus anomala thompsoniae Forestiera pubescens Ephedra torreyana Fallugia paradoxa Quercus welshii Amelanchier utahensis Ephedra viridis Ephedra viridis Quercus turbinella Poliomintha incana Psorothamnus thompsoniae Leptodactylon pungens Eriogonum corymbosum var. Eriogonum corymbosum var. Ceanothus greggii franklinii Rhamnus betulifolia Cercocarpus intricatus Coleogyne ramosissima Holodiscus dumosus Eriogonum leptocladon var. Cercocarpus ledifolius Cercocarpus montanus intermontanus rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Oleaceae Oleaceae FamilyShrubs, cont. Ephedraceae Species name Common name Nativity Rosaceae Fagaceae Rosaceae Ephedraceae Ephedraceae Fagaceae Labiatae (Lamiaceae) Leguminosae (Fabaceae) Polemoniaceae Polygonaceae Polygonaceae Rhamnaceae Rhamnaceae Rosaceae Rosaceae Rosaceae Polygonaceae Rosaceae Rosaceae Appendix D. Conﬁ

104 Annotated Checklist of Vascular Flora: Canyonlands National Park X X X Wet Comments X X X X XX X X X X X X Jun Pin- X SgeGrs XX X XX XX XX XX XX Des Salt X X XX X XX X XX X Br Blk- Flower time Park status ve-stamen tamarisk ve-stamen Intro Pres Apr–Sep X western chokecherry Native Pres May–Jul dwarf ninebark Native Pres May–Aug X X Rock crevices common dogbanespider milkweed Native Pres Native Jun–Aug Pres May–Jul X yellow willowcoyote willow NativeFendlerbush Pres Native Apr–Jun Pres Native Apr–Aug Pres Apr–Jun strapleaf willow Native Pres Apr–Jun cliff-rose roseWoods’ willowBebb’s Native Native Native Pres Pres Pres May–Jun May–Jul May–Junlittleleaf mock-orange Nativegolden currant Pres wolfberryAnderson’s May–Sep pale wolfberry Native Native wolfberryTorrey’s Pres Presﬁ Feb–May Mar–Jul Native Native X sea purslane X Rep PresJones’ amsoniatomentose amsonia Mar–May Mar–May Native Native Native Pres Pres X Rep milkweedCircle Cliffs Apr–May May–Sep Apr–Jun broadleaf milkweed Native Rep Native Pres May–Aug Jun–Aug X

var. var. var. var. var. var.

var. melanocarpa Physocarpus alternans Prunus virginiana Apocynum cannabinum Asclepias asperula asperula Salix eriocephala watsonii Salix exigua stenophylla Fendlera rupicola ligulifola Salix bebbiana Salix eriocephala chinensis Tamarix Sesuvium verrucosum Amsonia jonesii Purshia mexicana stansburyana Rosa woodsii Philadelphus microphyllus Ribes aureum andersonii Lycium pallidum Lycium torreyi Lycium Amsonia tomentosa stenophylla Asclepias labriformis Asclepias latifolia rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyShrubs, cont. Rosaceae Rosaceae Species name Common name Nativity Apocynaceae Asclepiadaceae Salicaceae Salicaceae Saxifragaceae (Hydrangeaceae) Salicaceae Salicaceae Tamaricaceae Forbs Perennial Aizoaceae Apocynaceae Rosaceae Rosaceae Saxifragaceae (Hydrangeaceae) Saxifragaceae (Grossulari-aceae) Solanaceae Solanaceae Solanaceae Apocynaceae Asclepiadaceae Asclepiadaceae

Chapter 6: Appendices 105 Disturbed areas Wet Comments X X X X X X X X X X X X X X X X Jun Pin- XX X X X X X X SgeGrs XX XX X X XX XX XX XX XX XX Des Salt X XX XX X XX X XX XX X Br Blk- Flower time Park status Native Pres Apr–Jun Native Rep Mar–Jul bigseed milkweed Native Pres Apr–Jul Rusby’s milkweedRusby’s milkweedRuth’s showy milkweed Nativewhorled milkweed Pres Native Native Pres Native Jun–Jul climbing milkweed Pres Pres Apr–Jun May–Aug Native May–Sep cryptanthBaker’s PresSan Juan cryptanth Apr–Sep Nativebasin yellow cryptanth Native Native Presplateau yellow Prescryptanth Rep May–Jun yellow-eye cryptanth May–Aug plateau cryptanth Apr–Jun Native cryptanth PresOsterhout’s Native X slender cryptanth Native Apr–Jun Presshowy stoneseed Presmatted tiquilia May–Jun Native May–Jun Native PresGlen Canyon hedgehog cactus Pres Native May–Jun claretcup Apr–Jul Pres May–Aug beavertail cactusUtah pricklypear Native Native Pres Rep Native Apr–Jun Feb–Jun Pres May–Jun orange milkweed Native Pres May–Aug

ora var. var.

var.

var. var. var. ava avoculata variegatus Asclepias macrosperma Asclepias subverticillata Asclepias rusbyi Asclepias ruthiae Asclepias speciosa Sarcostemma cynanchoides hartwegii Cryptantha bakeri Cryptantha cinerea pustulosa Cryptantha confertifl Cryptantha fl Cryptantha fl Cryptantha fulvocanescens Cryptantha osterhoutii Cryptantha tenuis Lithospermum incisum latior Tiquilia Echinocereus engelmannii var. Echinocereus triglochidiatus melanacanthus Opuntia basilaris basilaris Opuntia erinacea utahensis Asclepias tuberosa interior rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. FamilyPerennial Forbs, cont. Asclepiadaceae Species name Common name Nativity Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae (Ehretiaceae) Cactaceae Cactaceae Cactaceae Cactaceae Asclepiadaceae Appendix D. Confi

106 Annotated Checklist of Vascular Flora: Canyonlands National Park r forest Disturbed areas X X X X X Wet Comments X X X XX XX X XX XX XX XX Jun Pin- X X X X SgeGrs XX XX XX XX Des Salt XX X XX X XX Br Blk- Flower time Park status Native Pres Apr–May shhook ower fi ower c aster c Native Pres Jul–Oct cactus large pricklypearplains prickly-pear Native Pres Native Pres Apr–Jul May–Jun sandstone pussytoes Nativeburdock Pres May–Jul Intro Pres Jul–Sep Douglas-fi skeleton-leaf bursage Native Pres Jun–Oct western asterMexican devilweed Native Native Pres Pres Jul–Sep Mar–Oct small-fl sandwortEastwood’s Nativegreenmolly Pres May–Sep Native Pres Aug–Oct tarragonLouisiana wormwood NativeLouisiana wormwood Pres Native NativeLouisiana wormwood Jul–Oct Pres Pres NativePacifi Jul–Oct Jul–Oct Presblueleaf aster Jul–Oct white heath aster Native Native Pres Pres Jul–Sep Aug–Oct sand wormwood Native Pres Jul–Sep

var. var. var. var. var. var. var.

var. var. major Opuntia polyacantha polyacantha Antennaria marginata Arctium minus Opuntia phaeacantha Ambrosia tomentosa Aster spathulatus Aster spinosus Sclerocactus whipplei roseus Arenaria fendleri eastwoodiae Bassia americana Artemisia dracunculus glauca Artemisia ludoviciana albula Artemisia ludoviciana ludoviciana Artemisia ludoviciana mexicana Aster ascendens Aster glaucodes Aster pansus Artemisia campestris scouleriana rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Cactaceae Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Conﬁ FamilyPerennial Forbs, cont. Cactaceae Species name Common nameCompositae Nativity (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Cactaceae Caryophyllaceae (Alsinaceae) Chenopodiaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 107 Disturbed areas Hanging gardens Disturbed areas X X X X X X Wet Comments X X X XX X X X Jun Pin- XX X X SgeGrs XX XX XX XX Des Salt XX X XX X Br Blk- Flower time Park status musk thistle Intro Rep Jun–Sep nodding bur-marigold Native Pres Jul–Sep wingless thistlepretty thistle Native Pres Native Jun–Sep Pres Jun–Sep nakedstem Native Pres May–Jun Canada thistle Intro RepNew Mexico thistle Jul–Sep Utah thistle Native Pres thistleRothrock’s May–Jun thistleRydberg’s Native Native Pres Pres Native May–Jun glaucous hawksbeard Jun–Aug Pres Native May–Aug Pres May–Aug hispid golden-asterhairy golden-aster Native Pres Native May–Sep Rep May–Sep diffuse knapweeddiffuse Russian knapweed Intro Intro Pres Jul–Sep Pres Jun–Sep Disturbed areas bull thistle Intro Rep Jul–Aug rose-heath Native Pres May–Sep

var.

var. var. var. var. var. neomexicanum utahense Carduus nutans Bidens cernua bipinnatum pulchellum Cirsium calcareum Cirsium calcareum Enceliopsis nudicaulis nudicaulis Cirsium rydbergii Chrysopsis villosa Chrysopsis villosa Cirsium arvense Cirsium neomexicanum var. Cirsium neomexicanum var. Cirsium rothrockii glauca minor villosa Centaurea repens Centaurea diffusa Cirsium vulgare Crepis runcinata Chaetopappa ericoides rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Conﬁ

108 Annotated Checklist of Vascular Flora: Canyonlands National Park X X X Wet Comments XX X X X X X X X X X Jun Pin- X X X X SgeGrs XX XX X XX XX XX XX XX XX XX Des Salt XX X Br Blk- Flower time Park status ower Native Pres Jul–Sep ower Native Pres May–Oct ower Native Pres Apr–Aug eabane Native Pres May–Aug Navajo fl trailing daisy Native Pres May–Aug spreading daisy Native Pres May–Sep curly-cup gumweed Native Pres Jul–Sep thrift goldenweed Native Pres May–Jun Hopi blanketfl purpledisk sunfl slenderleaf daisyUtah daisy Nativewestern goldenrod Pres Native Jun–Sep Native Presbasin blanketfl Pres Jul–Oct Apr–Jun fall sneezeweed Nativecommon hyalineherb Pres Native Jul–Sep PresColorado rubber-plant May–Oct NativeColorado rubber-plant Pres Native May–Aug Pres May–Aug canyonlands hymenoxys Native Pres Apr–Jun erect gumweed Native Pres Aug–Sep

var.

var. var. var. var. var. da var. lifolius agellaris oribunda armerioides cinereus fl richardsonii concinnus divergens serrulata Erigeron divergens Grindelia squarrosa Haplopappus armerioides var. Gaillardia pinnatifi Helianthella microcephala Erigeron fl Erigeron pumilus Erigeron utahensis sparsifolius Erigeron utahensis utahensis Euthamia occidentalis Gaillardia spathulata Helenium autumnale montanum Hymenopappus fi var. Hymenoxys acaulis Hymenoxys richardsonii var. Hymenoxys richardsonii var. ivesiana Grindelia fastigiata rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 109 Disturbed areas X X Wet Comments X X X X X X X X XX X Jun Pin- X X SgeGrs X XX XX XXX XX XX XX XX XX XX XX Des Salt XX X XX X XX X Br Blk- Flower time Park status Native Pres Jun–Oct Native Pres May–Oct ower Native Pres Jun–Aug showy rush-pink Native Pres May–Jun blue lettuce Native Pres Jun–Sep poverty-weed Native Pres May–Sep three-nerve goldenrod Native Pres Aug–Oct aristate asterhoary astergumweed aster Native Pres Nativerock-goldenrod Native Jul–Oct Repdesert bahia Pres May–Sep Native May–Jul prairie coneﬂ Pres Native Jul–Sep Presbasin groundsel May–Jun many-headed groundsel Nativebroom groundsel PresCanadian goldenrod Apr–Aug Native Native Pres Presbrown-plume wire- Jun–Oct lettuce Aug–Oct slender wirelettuce Native Pres Jun–Sep Douglas’ groundsel Native Rep May–Oct

ora

var.

var. var. ora var. var.

var. aristata canescens var. ora var. var. grandifl desertorum tenuifolia Lygodesmia grandifl Lygodesmia var. pulchella Lactuca tatarica Iva axillaris Solidago velutina multicapitatus spartioides Machaeranthera canescens Machaeranthera canescens Machaeranthera grindelioides grindelioides Petradoria pumila pumila Platyschkuhria integrifolia var. Ratibida columnifera Senecio multilobatus Senecio spartioides Senecio spartioides Solidago canadensis salebrosa Stephanomeria paucifl Stephanomeria tenuifolia var. Senecio douglasii longilobus rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Confi

110 Annotated Checklist of Vascular Flora: Canyonlands National Park Disturbed areas X X Wet Comments X X X X X X X X XX X Jun Pin- XX X X SgeGrs X XX XX XX XX XX XX XX Des Salt X X XX XX X XX XX X Br Blk- Flower time Park status Native Pres Jul–Oct ower Native Pres Apr–Aug eld bindweed Intro Pres Jun–Aug Disturbed areas Hopi-tea Native Pres Jun–Aug common dandelion Introscapose greenthread Pres Native Apr–Dec Pres May–Aug hurtleaf woodyaster Native Presﬁ Apr–Jun rockcressHolboell’s Native Pres May–Jul Jones’ pepperwort Native Pres Apr–Sep scale glandweedhoary townsendia Nativeyellow salsify Pres Native Pres Aug–Oct Apr–Jul IntroCisco woodyaster Pres May–Jul Native Pres Apr–Jun desert rockcress rockcressSelby’s Nativesand dune wallﬂ Pres Native tall Eastwood’s Mar–Jun Prespepperwort broad-leaf pepperwort May–Aug Intro Pres Jun–Oct perennial rockcress Native Pres Mar–Jul Moab woodyaster Native Pres Apr–May

var. var.

var. var. var. pallens var. var. var. cinale subnudum belenidium Thelesperma megapotamicum Taraxacum ofﬁ Taraxacum Thelesperma subnudum var. Xylorhiza tortifolia Convolvulus arvensis Arabis holboellii pinetorum imberbis Lepidium montanum jonesii Thymophylla pentachaeta var. incana Townsendia dubius Tragopogon Xylorhiza venusta Arabis perennans Arabis pulchra Arabis selbyi Erysimum asperum purshii Lepidium alyssoides eastwoodiae Lepidium latifolium perennans Xylorhiza glabriuscula linearifolia rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) Appendix D. Conﬁ FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species nameCompositae (Asteraceae) Common name Nativity Compositae (Asteraceae) Convolvulaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Compositae (Asteraceae)

Chapter 6: Appendices 111 Disturbed areas X X X Wet Comments X X X X XX X XX X X X X Jun Pin- X XX X X X X SgeGrs XX XX XX XX XX XX XX Des Salt X X XX X XX Br Blk- Flower time Park status Native Pres Apr–Aug Native Pres Jun–Aug Native Pres Jun–Sep ower Native Pres Apr–Jul ower sharp-leaf twinpod Native Pres May–Jul buffalo gourdbuffalo Native Pres May–Aug common horehound Intro Pres May–Oct silver bladderpod twinpodNewberry’s Native NativeColorado Plateau Repbladderpod Presprince’s-plume May–Jun Mar–Jun heart-leaf jewelﬂ Nativecandelabra thelypody Pres NativeMoab thelypody Pres Apr–Sep Apr–Jun Native spurgeFendler’s Presshorthorn spurge Jun–Sep golden corydalis Native Native Pres Native Pres May–Sep Pres May–Jun smallﬂ Apr–Sep dragonhead mock pennyroyalAmerican bugleweed Native Native Pres Pres May–Aug Jul–Aug elkweedUtah swertia Native Native Pres Pres Jun–Jul Jun–Aug X Rocky Mountain gentian

var.

var. var. var. nis orum nis acutifolia Physaria acutifolia Cucurbita foetidissima Marrubium vulgare Physaria ludoviciana Physaria newberryi Physaria rectipes Stanleya pinnata Streptanthus cordatus Thelypodiopsis divaricata Thelypodium integrifolium gracilipes Chamaesyce fendleri Euphorbia brachycera Corydalis aurea occidentalis Gentiana affi Dracocephalum parvifl Hedeoma drummondii americanus Lycopus Swertia utahensis Swertia radiata affi rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. FamilyPerennial Forbs, cont. Cruciferae (Brassicaceae) Species name Common name Nativity Cucurbitaceae Labiatae (Lamiaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Euphorbiaceae Euphorbiaceae Fumariaceae Gentianaceae Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Gentianaceae Gentianaceae Appendix D. Confi

112 Annotated Checklist of Vascular Flora: Canyonlands National Park Rock crevices X Wet Comments X X X X X X X X X X X X Jun Pin- X X X X X SgeGrs XX X XX XX XX XX XX XX XX Des Salt XX XX XX X XX X Br Blk- Flower time Park status crescent milkvetch Native Pres Mar–Jun straggling milkvetch Native Pres Apr–Jun great rushy milkvetch NativeMoenkopi milkvetch Pres Native May–Jul Pres May–Jul Sheldon’s milkvetchSheldon’s painted milkvetch Native Prescicada milkvetch Native Mar–Jun Moab milkvetch Pres Native Apr–Jun rimrock milkvetch Rep Native Apr–Jun Native Rep Pres Apr–Jun May–Jul pliant milkvetchHopi milkvetch Nativecanyon kentrophyta Rep Native Native May–Aug Pres Pres May–Aug Jun–Sep woolly milkvetchMonument milkvetch Native NativeFerron milkvetch Pres Pres Feb–Jun May–Jun Native Pres Apr–Jun yellow milkvetch Native Rep Apr–Jul Bishop’s milkvetchBishop’s Native Pres May–Jun

var. var. var. var.

var. var. var. avus exuosus palans coloradoensis thompsoniae exuosus avus Astragalus amphioxys Astragalus lentiginosus var. amphioxys Astragalus lonchocarpus Astragalus moencoppensis Astragalus amphioxys vespertinus Astragalus ceramicus ceramicus Astragalus chamaeleuce Astragalus coltonii moabensis Astragalus desperatus desperatus Astragalus fl Astragalus fl fl Astragalus fucatus Astragalus kentrophyta var. Astragalus mollissimus var. Astragalus monumentalis Astragalus musiniensis fl Astragalus episcopus rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Leguminosae (Fabaceae) Species name Common name Nativity Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae)

Chapter 6: Appendices 113 X Wet Comments X X X X X XX XX X X X X Jun Pin- X X X X X SgeGrs XX XXX XX XX XX XX XX XX Des Salt XX X Br Blk- Flower time Park status Native Pres Apr–Jun owered birds-nest milkvetch Native Pres Apr–Jun northern sweetvetch Native Pres May–Aug Fisher milkvetch Native PresPreuss’ milkvetch Apr–Jun sandstone milkvetch Native NativeKanab prairie-clover Pres Pres Nativewhite prairie-clover Apr–Jun May–Aug Pres licoriceNuttall’s Native May–Jul Pres Native May–Sep Zion sweetpea PresMoab lupine May–Jul Nativealfalfa Presyellow sweet-clover Native Apr–Jun Preslarge-ﬂ Introbreadroot May–Aug peteriaThompson’s Pres Intro May–Octlemon scurf-pea Native Rep Pres May–Sep Native Apr–Jun Pres Apr–Aug Disturbed areas Disturbed areas longstipe milkvetchstinking milkvetch Native Rep Native Apr–Jul Pres Apr–Jul

var. var.

var.

var. var. var. orus var. cinalis avescens lanceolatum avescens Astragalus nidularius Hedysarum boreale boreale Melilotus offi Pediomelum megalanthum Astragalus piscator Astragalus praelongus Astragalus praelongus Astragalus preussii latus Astragalus sesquifl Dalea fl fl Dalea oligophylla Glycyrrhiza lepidota Lathyrus brachycalyx zionis Lupinus argenteus moabensis Medicago sativa Peteria thompsoniae Psoralidium lanceolatum var. lonchopus praelongus rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. FamilyPerennial Forbs, cont. Leguminosae (Fabaceae) Species name Common name Nativity Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix D. Confi

114 Annotated Checklist of Vascular Flora: Canyonlands National Park X X X Wet Comments X X X X X X X XX X X X X Jun Pin- X X X X SgeGrs X XX XX XX XX XX XX XX Des Salt XX X XX X XX X XX X Br Blk- Flower time Park status Native Pres Apr–Jun ax Native Pres May–Aug ax Native Pres Apr–Aug silvery sophora Native Pres Jul–Aug silky sophora Native Rep May–Jun slenderleaf scurfpea Native Pres Apr–Aug starry false Solomon’s- seal golden mariposa Native Pres May–Jun blue fl San Juan onionNevada onion Nativepink funnel-lily Pres Native May–Jun asparagus Pres Native Pres Apr–Jul Apr–May sinuous mariposa Introsego-lily Native Pressand lily Pres Jun–Jul Apr–Jun Nativefoothills death camas Pres Native Nativealcove death camas Jun–Jul Pres Rep Native Apr–Jun Mar–Jun Utah yellow-fl PresColorado stickleaf Jul–Aug desert stickleaf Native Pres Native Apr–Oct Pres Apr–Sep lewisii ora ssp. cinalis exuosus orum stenophyllum Sophora stenophylla Sophora nuttalliana Psoralidium lanceolatum var. Smilacina stellata Calochortus aureus Linum perenne Allium macropetalum Allium nevadense Androstephium brevifl Asparagus offi Calochortus fl Calochortus nuttallii Eremocrinum albomarginatum Zigadenus paniculatus Zigadenus vaginatus Linum subteres Mentzelia marginata Mentzelia multifl rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix D. Confi FamilyPerennial Forbs, cont. Leguminosae (Fabaceae) Species name Common name Nativity Liliaceae (Convallariaceae or Ruscaceae) Liliaceae (Calochortaceae) Linaceae Liliaceae (Alliaceae) Liliaceae (Alliaceae) Liliaceae (Alliaceae, Themidaceae) Liliaceae (Asparagaceae) Liliaceae (Calochortaceae) Liliaceae (Calochortaceae) Liliaceae (Anthericaceae) Liliaceae (Melanthiaceae) Liliaceae (Melanthiaceae) Linaceae Loasaceae Loasaceae

Chapter 6: Appendices 115 Hanging gardens X X Wet Comments X X X X X XX X X X X X X Jun Pin- X X X X SgeGrs X X XX XX XX XX Des Salt X XX XX XX XX X XX X XX X XX X XX X Br Blk- Flower time Park status Native Pres May–Sep Native Pres May–Aug Native Pres Mar–Oct Native Pres Apr–Jun Native Pres Jul–Sep Native Rep May–Jul alkali mallowscarlet globe-mallow Native Native Presslenderleaf Pres Apr–Oct globemallow May–Novnarrow-leaf four- o’clock X pale evening-primrose Native Pres May–Jul gooseberry-leaf globemallow globemallowJane’s Native globemallowMoore’s Pressmall-leaf globemallow Native Native May–Jul fragrant sand-verbena Pres Nativetrailing four-o’clock Pres May–Jul Pres Mar–Oct Nativeshowy four-o’clock Apr–Sep Pres sundropslavender-leaf Apr–Sep X Nativenorthern willow-herb Nativelong-tube evening- Native Pres Presprimrose Pres May–Aug Paria evening-primrose Apr–Jul Native Jun–Aug XBridges evening- Presprimrose X Apr–Jun Xgiant helleborine Native X broomrapeCooper’s Pres NativeSan Rafael prickly- Apr–Aug poppy X Pres May–Jul alcove bog-orchidclustered broomrape Native Native Pres Pres Jul–Aug Jun–Jul Xgardens Hanging

var.

var. var.

var. var. ora cooperi Malvella leprosa Sphaeralcea coccinea Sphaeralcea leptophylla Mirabilis linearis linearis Oenothera pallida pallida Sphaeralcea parvifolia Calylophus lavandulifolius Oenothera caespitosa Sphaeralcea grossulariifolia Sphaeralcea janeae Sphaeralcea moorei Abronia fragrans Allionia incarnata Mirabilis multiﬂ Epilobium ciliatum marginata Oenothera caespitosa navajoensis Oenothera longissima Epipactis gigantea Orobanche ludoviciana var. Argemone corymbosa arenicola Orobanche fasciculata Habenaria zothecina rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. FamilyPerennial Forbs, cont. Malvaceae Malvaceae Species nameMalvaceae Common name Nativity Nyctaginaceae Onagraceae Malvaceae Onagraceae Onagraceae Malvaceae Malvaceae Malvaceae Nyctaginaceae Nyctaginaceae Nyctaginaceae Onagraceae Onagraceae Onagraceae Orchidaceae Orobanchaceae Papaveraceae Orobanchaceae Orchidaceae Appendix D. Conﬁ

116 Annotated Checklist of Vascular Flora: Canyonlands National Park Slickrock Hanging gardens X Wet Comments X X X X X X X X X X X X X X X Jun Pin- X X X X X X SgeGrs X XX XX XX XX XX XX Des Salt XX XX X XX X XX X XX XX X Br Blk- Flower time Park status Native Pres May–Jul Native Pres May–Aug Native Pres May–Sep owered gilia owered Native Pres May–Aug common plantaincarmine gilia Intro Pres Native May–Aug Pres May–Jun c X Disturbed areas San Rafael prickly- poppy veiny dock Native Rep Apr–Jun scarlet gilialong-ﬂ San Rafael gilia prickly-phloxWatson’s Native Native Pres Native Preslong-leaf phlox May–Aug Pres May–Aug winged wild buckwheat Native May–Jun pretty wild buckwheat Pres Native Nativedesert trumpet Pres Pres Jun–Oct slender wild buckwheat Native Apr–Jul Apr–Jul Nativecushion wild buckwheat Pres Native Pres wild Shockley’s Jun–Oct buckwheat Pres Mar–Oct arid wild buckwheat Apr–Aug Nativecanaigre dock Repsausageleaf talinum Jun–Oct cave primrose Native Nativealcove columbine Pres Pres Apr–Jun Native Native Jun–Oct Pres Pres May–Jun Feb–Jul X Slickrock Xgardens Hanging carpet phlox Native Pres Apr–Jun Columbian virgin’s- bower white virgin’s-bower Native Pres Jul–Aug X

var. var. var.

ora atum var. simpsonii subaridum atum Gilia subnuda Plantago major Argemone corymbosa Rumex venosus parva Primula specuicola Aquilegia micrantha Ipomopsis aggregata aggregata Ipomopsis longifl Ipomopsis roseata Leptodactylon watsonii Phlox longifolia Eriogonum alatum Eriogonum bicolor Eriogonum infl infl Eriogonum microthecum var. Eriogonum ovalifolium Eriogonum shockleyi Eriogonum umbellatum var. Rumex hymenosepalus brevifolium Talinum Phlox hoodii canescens Clematis columbiana Clematis ligusticifolia rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Polemoniaceae Plantaginaceae Appendix D. Confi FamilyPerennial Forbs, cont. Papaveraceae Species name Common name Nativity Polygonaceae Primulaceae Ranunculaceae (Helleboraceae) Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Portulacaceae Polemoniaceae Ranunculaceae Ranunculaceae

Chapter 6: Appendices 117 Rock crevices Disturbed areas Hanging gardens X X Wet Comments X X X X XX X X XX X X X X X X Jun Pin- X X X X X SgeGrs X XX XX XX XX XX XX XX XX XX Des Salt X XX X Br Blk- Flower time Park status Native Pres May–Sep ax Native Pres Apr–Aug ower yellow paintbrushWyoming paintbrush paintbrushEastwood’s Native Native Native Rep Pres Pres Jun–Aug Jul–Aug May–Jul X pale larkspur Native Pres Apr–Jun undulate penstemon Native Pres May–Jul Nuttall’s larkspurNuttall’s marsh buttercup Nativerock spiraea Pres Native Apr–Jul shrubby bedstraw Pres Native May–Sepbastard toadﬂ Native Presdesert paintbrush Pres Aug–Sep May–Aug Native Pres monkey- Eastwood’s Apr–Jun ﬂ X bluestem penstemon Native penstemonPalmer’s Pres May–Jun NativeUtah penstemonwoolly mullein Pres Native May–Aug X Pres Intro Apr–Jun Rep Jun–Aug X Colorado bedstraw Native Pres May–Aug water speedwellfalse nightshade Intro Native Pres Pres Jun–Aug Apr–Jul dwarf lousewortdusty penstemon Native Native Pres Pres Apr–Jun Jun–Aug X

var. var. var. var. var. var. orum orum ava orum scaposum Castilleja linariifolia scabrida Delphinium andersonii var. undosus Petrophyton caespitosum multifl Penstemon eatonii Delphinium nuttallianum Ranunculus cymbalaria Galium multifl Comandra umbellata pallida Castilleja chromosa Castilleja fl Castilleja scabrida Mimulus eastwoodiae Penstemon cyanocaulis Penstemon palmeri palmeri Penstemon utahensis thapsus Verbascum coloradoense Galium multifl anagallis- Veronica aquatica Chamaesaracha coronopus Penstemon comarrhenus Pedicularis centranthera rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Scrophulariaceae FamilyPerennial Forbs, cont. Ranunculaceae (Helleboraceae) Species name Common name Nativity Rosaceae Scrophulariaceae Ranunculaceae (Helleboraceae) Ranunculaceae Rubiaceae Santalaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Rubiaceae Scrophulariaceae Solanaceae Scrophulariaceae Scrophulariaceae Appendix D. Confi

118 Annotated Checklist of Vascular Flora: Canyonlands National Park Wet Comments X XX X X X X X X X X X X X X X X Jun Pin- X X X SgeGrs XX XX XX XX XX XX XX XX XX XX XX Des Salt Br Blk- Flower time Park status Native Pres Apr–Jun Fendler’s ground-cherryFendler’s Native Pres May–Oct Fendler’s cymopterusFendler’s Native Pres Apr–May long-leaf ground-cherry NativeJames’ potato Rep Apr–Aug onion spring-parsley Native Nativesticky spring-parsley Pres Rep NativeColorado Plateau Jul–Sepspring-parsley Apr–May Presrock spring-parsley Apr–May lomatiumParry’s Native Preswedge-leaf fog-fruit Nativeprostrate vervain X Apr–May Nativepinyon dwarf-mistletoe Pres Native Pres X Native Apr–Jun juniper mistletoe Pres May–Jul Pres Aug–Sep Native May–Sep tumble pigweedprostrate pigweed Presredroot pigweed Jul–Sep Introthick-sepaled cryptanth Native Native Pres Rep Introslender cryptanth Pres cryptanthKelsey’s Jun–Sep Jun–Oct Pres Apr–Jun wing-nut cryptanth Native Jun–Sep Native X Presrecurved cryptanth Native Pres Apr–Jul Pres Native Jun–Jul Apr–Jul Pres Apr–Jun Disturbed areas Disturbed areas Disturbed areas

var. var. exus elachantha purpureus petraeus pterocarya fendleri Physalis hederifolia fendleri Cymopterus acaulis bracteata Verbena Amaranthus blitoides Amaranthus retroﬂ Cryptantha crassisepala var. Physalis longifolia Solanum jamesii Cymopterus bulbosus Cymopterus newberryi Cymopterus purpureus var. Cymopterus terebinthinus var. Lomatium parryi Phyla cuneifolia Arceuthobium divaricatum Phoradendron juniperinum Amaranthus albus Cryptantha gracilis Cryptantha kelseyana Cryptantha pterocarya var. Cryptantha recurvata rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyPerennial Forbs, cont. Solanaceae Species name Common name Nativity Umbelliferae (Apiaceae) Verbenaceae Amaranthaceae Amaranthaceae Boraginaceae Solanaceae Solanaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Verbenaceae Viscaceae Viscaceae Annual Forbs Amaranthaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae

Chapter 6: Appendices 119 Disturbed areas Disturbed areas Disturbed areas Disturbed areas X X X Wet Comments XX X XX X X XX X X XX Jun Pin- XX X X X SgeGrs XX XX XX XX XX XX XX XX XX Des Salt Br Blk- Flower time Park status Native Pres May–Oct y Native Pres May–Aug X X X ve-horn smotherweed ve-horn Intro Pres Aug–Oct bindweed heliotrope Nativecupseed stickseed Pres Native May–Sep Pres Apr–Jun yellow beeplantRocky Mountain beeplant Native Pres Apr–Sep goosefootFremont’s Nativenarrow-leaf goosefoot Native PresAmerican bugseed Pres Aug–Oct winged pigweedhalogeton Jul–Sep Native poverty-weedNuttall’s Pres Native Nativebarbwire Russian thistle Intro Aug–Oct PresRussian thistle Pres seepweedTorrey’s Pres Intro Aug–Oct Apr–Jul bur ragweed Jun–Sep Native Pres Intro Presleafy aster Jul–Aug X Pres Jun–Sep Native Jun–Sep Pres X Native Jul–Oct X Pres Aug–Oct X Disturbed areas Disturbed areas lambsquartermountain goosefoot Native Intro Pres Pres Jul–Sep Jun–Sep summer-cypress Intro Pres Aug–Oct sleepy catchﬂ thickleaf orachetwo-seed orachetumbling orache Nativeﬁ Intro Pres Intro Hist Jun–Sep Pres Jun–Sep Jul–Sep X X X X Disturbed areas

var. var. var. fremontii Heliotropium convolvulaceum Lappula occidentalis cupulata Halogeton glomeratus Monolepis nuttalliana Salsola tragus Suaeda torreyana Cleome lutea Cleome serrulata Chenopodium fremontii var. Chenopodium leptophyllum Corispermum americanum Cycloloma atriplicifolia Salsola paulsenii torreyana Ambrosia acanthicarpa Aster frondosus Chenopodium album album Chenopodium atrovirens Bassia scoparia Atriplex heterosperma Atriplex rosea Bassia hyssopifolia Silene antirrhina Atriplex dioica rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. FamilyAnnual Forbs, cont. Boraginaceae Species nameBoraginaceae Common name Nativity Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Capparaceae (Cleomaceae) Capparaceae (Cleomaceae) Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Compositae (Asteraceae) Compositae (Asteraceae) Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Caryophyllaceae Chenopodiaceae Appendix D. Conﬁ

120 Annotated Checklist of Vascular Flora: Canyonlands National Park Disturbed areas X X X Wet Comments X X X X X X X X X X X Jun Pin- X X X SgeGrs XX XX XX XX XX XX XX XX Des Salt XX X XX XX X Br Blk- Flower time Park status Native Pres May–Sep Native PresNative Apr–Jul Pres Apr–Jun Native Pres Mar–May ower Native Pres Jun–Sep ower Native Pres Jun–Sep liform desert- liform common sunfl golden crownbeard Native Pres May–Oct stevia dusty-maiden Native Pres Apr–Jul white-plume wirelettuce Brandegee’s sandplantBrandegee’s Nativepretty daisy Prescottonbatting cudweed Aug–Oct Native Pres Nativeprairie sunfl Jun–Oct Pres May–Aug prickly lettuce Introfi dandelion Pressowthistle desert- dandelion Jul–Sep prenanthellaspiny-leaf sow-thistle Intro Native Pres Pres Fremont’s syntrichopappus Jul–Oct May–Jun annual townsendia Native Pres Apr–Jun Canadian horseweed Native Pres Jul–Sep tansyleaf aster Native Pres May–Sep

var. ssp. ssp. var. lenticularis exauriculata Chaenactis stevioides encelioides Verbesina Helianthus annuus Stephanomeria exigua Conyza canadensis Dicoria brandegeei Erigeron bellidiastrum Gnaphalium stramineum Helianthus petiolaris fallax Lactuca serriola Malacothrix glabrata Malacothrix sonchoides Prenanthella exigua Sonchus asper Syntrichopappus fremontii annua Townsendia glabrata Machaeranthera tanacetifolia rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyAnnual Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 121 Disturbed areas Disturbed areas Disturbed areas Disturbed areas X Wet Comments X X X X X X X X X X X XX X X X Jun Pin- X X X X X SgeGrs X XX XX XX XX XX XX XX XX XX XX XX Des Salt XX X Br Blk- Flower time Park status Native Pres Mar–Jun ddle- ddleleaf Native Rep May–Aug ixweed Intro Pres Mar–Aug wedgeleaf draba Native Pres Feb–Jul blue mustard Intro Pres Apr–Jun rough cocklebur Native Pres Apr–Oct hare’s-ear mustardhare’s-ear Intro Rep Apr–Aug western tansy-mustard Nativefl Pres Mar–Aug prairie pepperwortSt. George pepperwort Native Native PresAfrican mustard Pres May–Jul tumble mustard Mar–Jun Intro Intro Presridge-seeded spurge Apr–Jul spurgeParry’s Pres Nativestork’s-bill May–Aug Pres Jun–Aug Nativecorrugate phacelia Pres Intro phaceliaHowell’s Native May–Sep Pres Pres Native May–Jun Feb–Oct Pres Apr–Jun Disturbed areas leafy fi western tansy-mustard Native Pres Mar–Aug spectacle-pod Native Pres Apr–Sep long-beak fi mustard

var. var. var.

var. orum var. orum cuneifolia Chorispora tenella Xanthium strumarium canadense Conringia orientalis georginum Sisymbrium altissimum Descurainia pinnata osmiarum Descurainia sophia Draba cuneifolia Lepidium densifl ramosum Lepidium lasiocarpum Malcolmia africana Chamaesyce glyptosperma Chamaesyce parryi Erodium cicutarium Nama densum Phacelia crenulata corrugata Phacelia howelliana parvifl Descurainia pinnata intermedia Dithyrea wislizenii Streptanthella longirostris rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Cruciferae (Brassicaceae) FamilyAnnual Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Euphorbiaceae Euphorbiaceae Geraniaceae Hydrophyllaceae Hydrophyllaceae Hydrophyllaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix D. Confi

122 Annotated Checklist of Vascular Flora: Canyonlands National Park Disturbed areas X Wet Comments X X X X X X X X X X X X X Jun Pin- X SgeGrs X XX XXX XX XX XX XX XX XX XX XX XX XX XX XX XX Des Salt X XX X Br Blk- Flower time Park status Native Pres Apr–Jun Native Pres Mar–Jun ax Native Rep May–Jun X X ax Native Pres Apr–Jun ower sandpuffs ower Native Pres Apr–Sep owered gaura owered Native Pres May–Aug X X Ives’ phacelia Native Pres Apr–Jun camissoniaWalker’s Native Pres Apr–Jun Cataract giliashy gilia Native Pres Native Jun–Sep Pres Apr–Jul X small-fl Wooton’s sand-verbenaWooton’s Native Rep Apr–Sep small-fl milkvetch gravel milkvetchshortstem lupine Nativedwarf lupine Pres Nativewhite sweet-clover Feb–Jul Pres May–Jul Native Intro Pres Presdowny fl Apr–Jul May–Oct white-stem blazingstar Nativecommon mallow Pres Intro Apr–Aug Pressmall-fl white-stem evening- Apr–Octprimrose woolly plantain broadlobe gilia Native Nativecommon gilia Pres Pres Mar–Aug giliaGunnison’s Xspreading gilia Apr–Jul Native X Native Pres Disturbed areas Pres Native May–Jun Pres May–Jun Apr–Jun Louisiana vetchbroom-fl Native Pres Apr–May

var.

var. var. ora micranthiformis Phacelia ivesiana Camissonia walkeri Gilia imperialis Gilia inconspicua Tripterocalyx micranthus Tripterocalyx wootonii Mentzelia albicaulis carneus Tripterocalyx Oenothera albicaulis Astragalus nuttallianus var. Astragalus sabulonum Lupinus brevicaulis Lupinus pusillus pusillus Melilotus alba Linum puberulum Malva neglecta Gaura parviﬂ Plantago patagonica Gilia hutchinsifolia Gilia leptomeria leptomeria Ipomopsis gunnisonii Ipomopsis polycladon Vicia ludoviciana Linum aristatum rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyAnnual Forbs, cont. Hydrophyllaceae Species name Common name Nativity Onagraceae Polemoniaceae Polemoniaceae Nyctaginaceae Loasaceae Nyctaginaceae Onagraceae Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Linaceae Malvaceae Onagraceae Plantaginaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Leguminosae (Fabaceae) Linaceae

Chapter 6: Appendices 123 Disturbed areas Rock crevices X Wet Comments X X X X X X X X X X Jun Pin- X X X SgeGrs X XX XX XX XX XX XX Des Salt XX X XX XX X XX Br Blk- Flower time Park status Native PresNative Apr–Oct RepNative Apr–Oct PresNative May–Oct PresNative Mar–Oct Pres Apr–Oct dwarf gilia Native Pres Apr–Jun nodding wild buckwheat skeletonweed wild buckwheat wild Gordon’s buckwheat wild Palmer’s buckwheat wild Wetherill’s buckwheat common purslanebur buttercup Native Pres Intro trumpetangel’s Jun–Sepdesert tobacco Rep Mar–Jun Nativeyellow kallstroemia Pres Nativepuncture vine Pres Apr–Oct Native X Mar–Oct golden sedge PresNebraska sedge Intro Jun–Oct sedgeParry’s woolly sedge Pres Native Nativeblackcreeper sedge Disturbed areas May–Sep PresRoss’ sedge Pres Native Nativecommon spikerush Apr–Aug Native May–Jul Pres Pres Pres Native May–Jul May–Aug Native Pres May–Sep Pres May–Aug May–Aug Disturbed areas X X X X X X golden dock Native Pres May–Sep bushy cinquefoilbrook cinquefoil bird’s-beakWright’s Native Native Native Pres Pres Pres Jun–Jul Jul–Sep Jun–Aug X X buffaloburcut-leaf nightshade Native Rep Intro May–Sep Rep Jun–Sep Disturbed areas

var. var. exum orum Ipomopsis pumila Ranunculus testiculatus Carex aurea Carex nebrascensis Carex parryana Carex pellita Carex praegracilis Carex rossii Eriogonum cernuum cernuum Eriogonum defl Eriogonum gordonii Eriogonum palmerianum Eriogonum wetherillii Rumex maritimus Portulaca oleracea Datura wrightii Nicotiana trigonophylla Kallstroemia californica terrestris Tribulus Eleocharis palustris fueginus Potentilla rivalis Cordylanthus wrightii Potentilla paradoxa Solanum trifl Solanum rostratum rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. FamilyAnnual Forbs, cont. Polemoniaceae Species name Common name Nativity Ranunculaceae Graminoids Perennial Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Portulacaceae Solanaceae Solanaceae Zygophyllaceae Zygophyllaceae Cyperaceae Rosaceae Scrophulariaceae Rosaceae Solanaceae Solanaceae Appendix D. Confi

124 Annotated Checklist of Vascular Flora: Canyonlands National Park Disturbed areas Disturbed areas X X X X X X X Wet Comments X XX X XX X X Jun Pin- X X SgeGrs XX XX XX XX XX Des Salt XX X XX Br Blk- Flower time Park status soft-stem bulrushcrested wheatgrass Native Intro Rep Pres Jun–Aug Jun–Aug X beaked spikerushhardstem bulrushthree-square bulrush Native Native Native Pres Pres Pres Jun–Aug Jun–Aug May–Aug X X tall wheatgrass Intro Rep Jun–Aug Canada wildrye Native Pres Jul–Aug purple three-awn Native Pres Mar–Sep Jones’ reedgrassdesert saltgrass Native Pres Native Jul–Aug Pres Jun–Sep big bluestembushy bluestem Native Native Prescane bluestem Rep Jul–Sep silver bluestem Apr–Oct Nativesideoats grama Pres Nativeblue grama Apr–Nov Rep Nativesmooth brome Aug–Sep Pres Native Jul–Sep Intro Pres Pres Jul–Sep Jun–Aug Disturbed areas redtop bluestemHall’s Native Intro Pres Pres Jul–Sep Jun–Sep

var. var.

var. var. scabriglumis torreyana Agropyron cristatum Scirpus validus Eleocharis rostellata Scirpus acutus Scirpus pungens longispicatus Elymus elongatus Elymus canadensis Aristida purpurea Calamagrostis scopulorum Distichlis spicata Bouteloua curtipendula Andropogon gerardii gerardii Andropogon glomeratus var. Bothriochloa barbinodis Bothriochloa laguroides var. Bouteloua gracilis Bromus inermis inermis Agrostis stolonifera Andropogon gerardii chrysocomus rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Gramineae (Poaceae) Cyperaceae Appendix D. Conﬁ FamilyPerennial Graminoids, cont. Cyperaceae Cyperaceae Species nameCyperaceae Common name Nativity Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

Chapter 6: Appendices 125 Disturbed areas Slickrock Disturbed areas X X Wet Comments XX X X X X XX X X X X X XX X Jun Pin- X X X X SgeGrs X XX XX XX XX XX Des Salt XX X XX X XX X XX X Br Blk- Flower time Park status Intro Rep Jun–Aug squirreltail Native Pres May–Aug scratchgrassNew Mexican muhly Native Native Pres Pres Aug–Sep Jul–Aug intermediate wheatgrass Salina wildryewestern wheatgrass Native Nativeslender wheatgrass Pres PresVirginia wildrye Native Jun–Jul Jun–Aug Presspike pappusgrass Native Jun–Aug Native Presfuffgrass Pres Jul–Sep Aug–Sep tall fescue Nativefoxtail barley Pres Introfoxtail muhly Apr–Aug Pres Native May–Jul Rep Native Jun–Aug Presbush muhly Jul–Sep Native Pres Jul–Sep thickspike wheatgrass Native Pres Jun–Aug hairy tridens Native Pres May–Jul galleta Native Pres May–Aug ora Elymus elymoides Muhlenbergia asperifolia Muhlenbergia pauciﬂ Elymus hispidus Elymus salinus Elymus smithii Elymus trachycaulus Elymus virginicus Enneapogon desvauxii Erioneuron pulchellum Festuca arundinacea Hordeum jubatum Muhlenbergia andina Muhlenbergia porteri Elymus lanceolatus Erioneuron pilosum Hilaria jamesii rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. FamilyPerennial Graminoids, cont. Gramineae (Poaceae) Species name Common name Nativity Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix D. Conﬁ

126 Annotated Checklist of Vascular Flora: Canyonlands National Park Hanging gardens X X X X X X X Wet Comments XX X XX X X X XX XX Jun Pin- XX X X X X X X X SgeGrs X XX Des Salt XXX XX Br Blk- Flower time Park status Thurber’s muhlyThurber’s Native Pres Jul–Sep Sandhill muhly Native Pres Jul–Sep little bluestem Native Pres Jul–Sep bulbous bluegrass Intro Rep Apr–Jul alkaligrassNuttall’s Native Pres May–Aug prairie cordgrass Native Rep Jun–Sep littleseed ricegrassbundle panicgrass Native Pres Native Jun–Aug Pres Jul–Sep common reed Nativemuttongrass Pres bluegrassSandberg’s Jul–Sep Nativewater polypogon Native Pres Pres Intro Apr–Sep May–Aug PresJohnson-grass Jul–Aug alkali cordgrass Intro Native Pres Pres Jul–Sep Jun–Sep switchgrasstimothy Native Pres Intro Jun–Sep Rep Jun–Sep vine mesquite Native Pres Aug–Sep Muhlenbergia thurberi Muhlenbergia pungens Schizachyrium scoparium Poa bulbosa Puccinellia nuttalliana Spartina pectinata Oryzopsis micrantha Panicum acuminatum Phragmites australis Poa fendleriana Poa secunda Polypogon semiverticillatus Sorghum halepense Spartina gracilis Panicum virgatum Phleum pratense Panicum obtusum rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Gramineae (Poaceae) Appendix D. Conﬁ FamilyPerennial Graminoids, cont. Gramineae (Poaceae) Species name Common name Nativity Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

Chapter 6: Appendices 127 X X Wet Comments XX X X X X X X X X X X X Jun Pin- X X X SgeGrs XX XX XX XX XX XX XX XX Des Salt XX X XX X Br Blk- Flower time Park status Native Pres Jun–Jul alkali sacaton Native Pres Jun–Aug mountain rushlongstyle rushknotted rush Nativepoverty rush Pres Native Jun–Aug Native Pres Native Pres Jun–Aug Rep Jul–Sep Jun–Aug X X X Tracy’s rushTracy’s Native Pres Jun–Aug spike dropseedsand dropseed Nativemesa dropseed Pres Nativegiant dropseed Jun–Sep Pres NativeMormon needlegrass Jun–Aug Pres Native Nativeneedle-and-thread Jun–Sep Pres Pres Native Jun–Sep May–Jul PresIndian ricegrass May–Jul New Mexico feathergrass Nativedesert needlegrass Pres ricegrassBloomer’s Native May–Jul Presslim tridens Native Apr–Jun PresBaltic rush Jun–Aug Native Pres Native May–Sep Pres May–Jul X crested needlegrass Native Pres Jun–Jul

var.

var. var. comata var. var. exuosus bloomeri x Sporobolus airoides airoides Juncus nodosus Juncus tenuis Juncus ensifolius montanus Juncus longistylis brunnescens Juncus ensifolius Sporobolus contractus Sporobolus cryptandrus Sporobolus ﬂ Sporobolus giganteus Stipa arida Stipa comata Stipa coronata Stipa hymenoides Stipa neomexicana Stipa speciosa Stipa muticus Tridens Juncus arcticus parishii rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. FamilyPerennial Graminoids, cont. Gramineae (Poaceae) Species name Common name Nativity Juncaceae Juncaceae Juncaceae Juncaceae Juncaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Juncaceae Appendix D. Conﬁ

128 Annotated Checklist of Vascular Flora: Canyonlands National Park Disturbed areas X X X X Wet Comments XX X X X X X Jun Pin- X SgeGrs X XX XX XX XX XX Des Salt XX XX X XX X XX X Br Blk- Flower time Park status at-sedge Native Pres Jul–Oct X at-sedge Native Pres Jun–Sep X redroot fl oatssixweeks grama Native Pres Intro Aug–Sep Pres Jun–Aug Disturbed areas Torrey’s rushTorrey’s southern cattailbroad-leaved cattail Native Native Native fl taper-tip Pres Pres Pres Jun–Aug Jun–Aug Jul–Augcheatgrass X X X Intro Pres May–Jul ripgut brome Intro Pres Mar–Jun Japanese chess Intro Pres Jun–Aug annual wheatgrass Intro Pres May–Jul false buffalograss Native Pres Jun–Sep red bromebarnyard-grass Intro Pres Introsix-weeks fescue Mar–Jun Presrabbit barley Native Aug–Sep little barley Presbearded sprangletop Intro Apr–Jun Native Pres Native Preswitchgrass May–Jul Pres Jul–Sep May–Aug Native Pres Jun–Sep sativa ora var. Cyperus erythrorhizos fatua Avena Bouteloua barbata Juncus torreyi domingensis Typha latifolia Typha Cyperus acuminatus Bromus tectorum Bromus diandrus Bromus japonicus Eremopyrum triticeum Munroa squarrosa Bromus rubens Echinochloa crus-gallii Festuca octofl Hordeum murinum Hordeum pusillum Leptochloa fascicularis Panicum capillare rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. Cyperaceae Gramineae (Poaceae) Gramineae (Poaceae) Appendix D. Confi FamilyPerennial Graminoids, cont. Juncaceae Typhaceae Species nameTyphaceae Annual Graminoids Cyperaceae Common name Nativity Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

Chapter 6: Appendices 129 crevices Rock crevices Hanging gardens X X X X Wet Comments X X Jun Pin- X SgeGrs XX Des Salt Br Blk- Flower time Park status Native Pres NA eld horsetail eld Native Pres NA X X X Bigelow’s bluegrassBigelow’s Native Rep Mar–May ﬁ tall scouring-rushsmooth scouring-rush Nativesouthern maidenhair Nativefern Pres Presebony spleenwort NA NAslender lip-fern Native Pres Native NA Pres NA X Shady rock rabbitfoot-grass Intro Pres Jun–Aug western cliffbrakeborder cloak-fern Native Pres Native NA Pres NA Sandstone cliffs

var.

var. ne Poa bigelovii Equisetum hyemale afﬁ Adiantum capillus-veneris Equisetum arvense Equisetum laevigatum Asplenium resiliens Cheilanthes feei Polypogon monspeliensis Pellaea glabella Pellaea limitanea occidentalis rmed, historical, and reported taxa of Canyonlands National Park, organized by life form with ecological notes, cont. FamilyAnnual Graminoids, cont. Gramineae (Poaceae) Species name Common name Nativity Equisetaceae Polypodiaceae (Adiantaceae) Ferns and Allies Equisetaceae Equisetaceae Polypodiaceae (Aspleniaceae) Polypodiaceae (Adiantaceae) Gramineae (Poaceae) Polypodiaceae (Adiantaceae) Polypodiaceae (Adiantaceae) Appendix D. Conﬁ

130 Annotated Checklist of Vascular Flora: Canyonlands National Park The Department of the Interior protects and manages the nation’s natural resources and cultural heritage; provides scientiﬁ c and other information about those resources; and honors its special responsibilities to American Indians, Alaska Natives, and afﬁ liated Island Communities.

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