AN INTERESTING POPULATION of Nucella Lapillus L.,1758 by John Llewellyn-Jones
Total Page:16
File Type:pdf, Size:1020Kb
Load more
Recommended publications
-
Nachtrag Zu "Ueber Abnormitäten Der Färbung, Der Windungsrichtung Und Der Gehäusebildung Bei Den Clausiliiden"
ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Archiv für Molluskenkunde Jahr/Year: 1928 Band/Volume: 60 Autor(en)/Author(s): Schlesch Hans Artikel/Article: Nachtrag zu "Ueber Abnormitäten der Färbung, der Windungsrichtung und der Gehäusebildung bei den Clausiliiden". 33-49 33 Nachtrag zu „lieber Abnormitäten der Färbung, der Windungsrichtung und der Gehäusebildung bei den Clausiliiden". Von Hans Schlesch, Kopenhagen. I. Ueber Albinismus der europäischen CI ausi lüden. Als ich meine Arbeit über die Abnormitäten der Clausiliiden veröffentlichte1), war ich mir dessen wohl bewußt, daß sie nie vollständig werden würde, denn die Angaben sind allzusehr in einer umfang reichen, oft nicht leicht zu beschaffenden Literatur versteckt. Ich kann bereits jetzt einen kleinen Nach trag liefern, den ich hauptsächlich den Zuschriften verschiedener Fachgenossen verdanke. Vor allem zu Dank verpflichtet bin ich jedoch den Herren Dr. Lajos Soös, Hofrat Dr. Rudolf Sturany und Dr. Wolfgang Adensamer, die mir bei meinen kürzlichen Besuchen in Budapest und Wien in lie benswürdiger Weise Zutritt zu den von ihnen betrauten Sammlungen verschafften, sodaß ich dieClausilii- dae dieser Museen durchsehen konnte. Zuerst sei noch einmal kurz auf die Einflüsse eingegangen, unter denen sich Albinos in einiger An zahl finden. Einmal nimmt anscheinend die Häufig keit solcher pigmentloser Exemplare unter bestimm ten ökologischen Bedingungen besonders zu, wie vor allem in kühlen, feuchten Gebirgstälern, wie sie sich am meisten in zerklüfteten Kalkgebirgen finden, ferner im Hochgebirge. Andererseits ist ja beobachtet, daß bei einer Anzahl von Gattungen(Fusulus, Laciniaria, Cochlodina, Delima) Albinismus nicht selten auftritt, während er bei andern kaum beobachtet wirdClau (- i) Arch. -
Bruno Sabelli * E Stefano Tommasini * Introduzione
Boll. Malacologico I Milano' 18 I (9-12) I 291-300 I settembre-dicembre 1982 Bruno Sabelli * e Stefano Tommasini * OSSERVAZIONI SULLA RADULA E SULLA PROTOCONCA DI BOLINUS BRANDARIS (L" 1758) E PHYLWNOTUS TRUNCULUS (L" 1758) Introduzione La sottofamiglia M u r i c i n a e comprende in Mediterraneo so- lo poche specie, per la precisione sei, di cui una (Bolinus cornutus (L., 1758)) segnalata solo dubitativamente per il nostro mare, due (Mu- rex tribulus L., 1758 e Aspella anceps (LAMARCK,1822)) sono ospiti re- centissimi essendo penetrati e forse acclimatati si in Mediterraneo so- lo dopo l'apertura del canale di Suez (BARASHe DANIN, 1972, 1977; GHI- SOTTI,1974) mentre le altre tre si possono definire come autoctone. Poichè le tre specie esotiche sono state reperite solo in numero esi- guo di esemplari e comunque mai viventi non si hanno informazioni sulla loro radula e sulla microscultura della loro protoconca e noi non siamo stati in grado di colmare questa lacuna. Per quanto con- cerne le altre tre specie Dermomurex scalaroides (BLAINVILLE,1826) è stato ampiamente investigato sotto il profilo tassonomico da FRAN- CHINI (1977) che ne ha anche illustrato la radula; mancherebbero co- munque dati sulla sua protoconca, ma gli oltre trenta esemplari esa- minati di questa specie non si sono prestati per questo studio essen- do tutti con l'apice abraso per cui la questione rimane insoluta. Solo le ultime due specie: Bolinus brandaris (L., 1758) e Phyllonotus trun- culus (L., 1758), sono dunque state studiate e se pur poco rappresen- tative dell'intera sottofamiglia presentano comunque alcuni interes- santi problemi che ci hanno indotto a redigere questa nota. -
Malaco Le Journal Électronique De La Malacologie Continentale Française
MalaCo Le journal électronique de la malacologie continentale française www.journal-malaco.fr MalaCo (ISSN 1778-3941) est un journal électronique gratuit, annuel ou bisannuel pour la promotion et la connaissance des mollusques continentaux de la faune de France. Equipe éditoriale Jean-Michel BICHAIN / Paris / [email protected] Xavier CUCHERAT / Audinghen / [email protected] Benoît FONTAINE / Paris / [email protected] Olivier GARGOMINY / Paris / [email protected] Vincent PRIE / Montpellier / [email protected] Les manuscrits sont à envoyer à : Journal MalaCo Muséum national d’Histoire naturelle Equipe de Malacologie Case Postale 051 55, rue Buffon 75005 Paris Ou par Email à [email protected] MalaCo est téléchargeable gratuitement sur le site : http://www.journal-malaco.fr MalaCo (ISSN 1778-3941) est une publication de l’association Caracol Association Caracol Route de Lodève 34700 Saint-Etienne-de-Gourgas JO Association n° 0034 DE 2003 Déclaration en date du 17 juillet 2003 sous le n° 2569 Journal électronique de la malacologie continentale française MalaCo Septembre 2006 ▪ numéro 3 Au total, 119 espèces et sous-espèces de mollusques, dont quatre strictement endémiques, sont recensées dans les différents habitats du Parc naturel du Mercantour (photos Olivier Gargominy, se reporter aux figures 5, 10 et 17 de l’article d’O. Gargominy & Th. Ripken). Sommaire Page 100 Éditorial Page 101 Actualités Page 102 Librairie Page 103 Brèves & News ▪ Endémisme et extinctions : systématique des Endodontidae (Mollusca, Pulmonata) de Rurutu (Iles Australes, Polynésie française) Gabrielle ZIMMERMANN ▪ The first annual meeting of Task-Force-Limax, Bünder Naturmuseum, Chur, Switzerland, 8-10 September, 2006: presentation, outcomes and abstracts Isabel HYMAN ▪ Collecting and transporting living slugs (Pulmonata: Limacidae) Isabel HYMAN ▪ A List of type specimens of land and freshwater molluscs from France present in the national molluscs collection of the Hebrew University of Jerusalem Henk K. -
Molluscs of the Dürrenstein Wilderness Area
Molluscs of the Dürrenstein Wilderness Area S a b i n e F ISCHER & M i c h a e l D UDA Abstract: Research in the Dürrenstein Wilderness Area (DWA) in the southwest of Lower Austria is mainly concerned with the inventory of flora, fauna and habitats, interdisciplinary monitoring and studies on ecological disturbances and process dynamics. During a four-year qualitative study of non-marine molluscs, 96 sites within the DWA and nearby nature reserves were sampled in cooperation with the “Alpine Land Snails Working Group” located at the Natural History Museum of Vienna. Altogether, 84 taxa were recorded (72 land snails, 12 water snails and mussels) including four endemics and seven species listed in the Austrian Red List of Molluscs. A reference collection (empty shells) of molluscs, which is stored at the DWA administration, was created. This project was the first systematic survey of mollusc fauna in the DWA. Further sampling might provide additional information in the future, particularly for Hydrobiidae in springs and caves, where detailed analyses (e.g. anatomical and genetic) are needed. Key words: Wilderness Dürrenstein, Primeval forest, Benign neglect, Non-intervention management, Mollusca, Snails, Alpine endemics. Introduction manifold species living in the wilderness area – many of them “refugees”, whose natural habitats have almost In concordance with the IUCN guidelines, research is disappeared in today’s over-cultivated landscape. mandatory for category I wilderness areas. However, it may not disturb the natural habitats and communities of the nature reserve. Research in the Dürrenstein The Dürrenstein Wilderness Area Wilderness Area (DWA) focuses on providing invento- (DWA) ries of flora and fauna, on interdisciplinary monitoring The Dürrenstein Wilderness Area (DWA) was as well as on ecological disturbances and process dynamics. -
(Approx) Mixed Micro Shells (22G Bags) Philippines € 10,00 £8,64 $11,69 Each 22G Bag Provides Hours of Fun; Some Interesting Foraminifera Also Included
Special Price £ US$ Family Genus, species Country Quality Size Remarks w/o Photo Date added Category characteristic (€) (approx) (approx) Mixed micro shells (22g bags) Philippines € 10,00 £8,64 $11,69 Each 22g bag provides hours of fun; some interesting Foraminifera also included. 17/06/21 Mixed micro shells Ischnochitonidae Callistochiton pulchrior Panama F+++ 89mm € 1,80 £1,55 $2,10 21/12/16 Polyplacophora Ischnochitonidae Chaetopleura lurida Panama F+++ 2022mm € 3,00 £2,59 $3,51 Hairy girdles, beautifully preserved. Web 24/12/16 Polyplacophora Ischnochitonidae Ischnochiton textilis South Africa F+++ 30mm+ € 4,00 £3,45 $4,68 30/04/21 Polyplacophora Ischnochitonidae Ischnochiton textilis South Africa F+++ 27.9mm € 2,80 £2,42 $3,27 30/04/21 Polyplacophora Ischnochitonidae Stenoplax limaciformis Panama F+++ 16mm+ € 6,50 £5,61 $7,60 Uncommon. 24/12/16 Polyplacophora Chitonidae Acanthopleura gemmata Philippines F+++ 25mm+ € 2,50 £2,16 $2,92 Hairy margins, beautifully preserved. 04/08/17 Polyplacophora Chitonidae Acanthopleura gemmata Australia F+++ 25mm+ € 2,60 £2,25 $3,04 02/06/18 Polyplacophora Chitonidae Acanthopleura granulata Panama F+++ 41mm+ € 4,00 £3,45 $4,68 West Indian 'fuzzy' chiton. Web 24/12/16 Polyplacophora Chitonidae Acanthopleura granulata Panama F+++ 32mm+ € 3,00 £2,59 $3,51 West Indian 'fuzzy' chiton. 24/12/16 Polyplacophora Chitonidae Chiton tuberculatus Panama F+++ 44mm+ € 5,00 £4,32 $5,85 Caribbean. 24/12/16 Polyplacophora Chitonidae Chiton tuberculatus Panama F++ 35mm € 2,50 £2,16 $2,92 Caribbean. 24/12/16 Polyplacophora Chitonidae Chiton tuberculatus Panama F+++ 29mm+ € 3,00 £2,59 $3,51 Caribbean. -
Fauna from Leaota Mountains - Romania
Muzeul Olteniei Craiova. Oltenia. Studii şi comunicări. Ştiinţele Naturii. Tom. 33, No. 1/2017 ISSN 1454-6914 DIVERSITY OF SOIL MITES (ACARI: MESOSTIGMATA) AND GASTROPODS (GASTROPODA) FAUNA FROM LEAOTA MOUNTAINS - ROMANIA MANU Minodora, LOTREAN Nicolae, CIOBOIU Olivia, POP Oliviu Grigore Abstract. In the period May-September 2016, the diversity of soil mites (Acari: Mesostigmata) and fauna of gastropods (Gastropoda) was investigated in Leaota Mountains. In total 44 transects were investigated, located in 13 types of habitats. Considering the mite fauna, 52 species were identified. The highest value of Shannon index was recorded in natural beech forest, in comparison with the planted spruce forest, where this parameter recorded the lowest value. From the conservative point of view, Pergamasus laetus Juvara-Bals, 1970 was identified as an endemic species for Romania. Another two rare species were signaled: Veigaia propinqua Willmann, 1936 and V. transisalae (Oudemans, 1902). Iphidosoma multiclavatum Willmann, 1953 was signaled for the first time in Romania. Taking into account the gastropod fauna (Gastropoda), 26 species were investigated. The most favorable habitats for these invertebrates were deciduous and spruce forest. One endemic species for Romania was identified: Mastus venerabilis (L. Pfeiffer, 1855). Keywords: abundance, diversity, gastropods, habitat, mites. Rezumat. Diversitatea faunei de acarieni edafici (Acari: Mesostigmata) și de gastropode (Gastropoda) din Munţii Leaota - România. În perioada mai - septembrie 2016, s-a realizat studiul diversităţii faunei de acarieni edafici (Acari: Mesostigmata) şi a gastropodelor (Gastropoda) din munţii Leaota. Au fost investigate 44 de transecte, localizate în 13 tipuri de habitate. Analizând fauna de acarieni, au fost identificate 52 de specii. Cea mai mare valoare a indicelui de diversitate Shannon s-a obţinut în pădurea naturală de fag, în comparaţie cu plantaţia de molid, unde acest parametru a scăzut semnificativ. -
ABSTRACT Title of Dissertation: PATTERNS IN
ABSTRACT Title of Dissertation: PATTERNS IN DIVERSITY AND DISTRIBUTION OF BENTHIC MOLLUSCS ALONG A DEPTH GRADIENT IN THE BAHAMAS Michael Joseph Dowgiallo, Doctor of Philosophy, 2004 Dissertation directed by: Professor Marjorie L. Reaka-Kudla Department of Biology, UMCP Species richness and abundance of benthic bivalve and gastropod molluscs was determined over a depth gradient of 5 - 244 m at Lee Stocking Island, Bahamas by deploying replicate benthic collectors at five sites at 5 m, 14 m, 46 m, 153 m, and 244 m for six months beginning in December 1993. A total of 773 individual molluscs comprising at least 72 taxa were retrieved from the collectors. Analysis of the molluscan fauna that colonized the collectors showed overwhelmingly higher abundance and diversity at the 5 m, 14 m, and 46 m sites as compared to the deeper sites at 153 m and 244 m. Irradiance, temperature, and habitat heterogeneity all declined with depth, coincident with declines in the abundance and diversity of the molluscs. Herbivorous modes of feeding predominated (52%) and carnivorous modes of feeding were common (44%) over the range of depths studied at Lee Stocking Island, but mode of feeding did not change significantly over depth. One bivalve and one gastropod species showed a significant decline in body size with increasing depth. Analysis of data for 960 species of gastropod molluscs from the Western Atlantic Gastropod Database of the Academy of Natural Sciences (ANS) that have ranges including the Bahamas showed a positive correlation between body size of species of gastropods and their geographic ranges. There was also a positive correlation between depth range and the size of the geographic range. -
MOLECULAR PHYLOGENY of the NERITIDAE (GASTROPODA: NERITIMORPHA) BASED on the MITOCHONDRIAL GENES CYTOCHROME OXIDASE I (COI) and 16S Rrna
ACTA BIOLÓGICA COLOMBIANA Artículo de investigación MOLECULAR PHYLOGENY OF THE NERITIDAE (GASTROPODA: NERITIMORPHA) BASED ON THE MITOCHONDRIAL GENES CYTOCHROME OXIDASE I (COI) AND 16S rRNA Filogenia molecular de la familia Neritidae (Gastropoda: Neritimorpha) con base en los genes mitocondriales citocromo oxidasa I (COI) y 16S rRNA JULIAN QUINTERO-GALVIS 1, Biólogo; LYDA RAQUEL CASTRO 1,2 , Ph. D. 1 Grupo de Investigación en Evolución, Sistemática y Ecología Molecular. INTROPIC. Universidad del Magdalena. Carrera 32# 22 - 08. Santa Marta, Colombia. [email protected]. 2 Programa Biología. Universidad del Magdalena. Laboratorio 2. Carrera 32 # 22 - 08. Sector San Pedro Alejandrino. Santa Marta, Colombia. Tel.: (57 5) 430 12 92, ext. 273. [email protected]. Corresponding author: [email protected]. Presentado el 15 de abril de 2013, aceptado el 18 de junio de 2013, correcciones el 26 de junio de 2013. ABSTRACT The family Neritidae has representatives in tropical and subtropical regions that occur in a variety of environments, and its known fossil record dates back to the late Cretaceous. However there have been few studies of molecular phylogeny in this family. We performed a phylogenetic reconstruction of the family Neritidae using the COI (722 bp) and the 16S rRNA (559 bp) regions of the mitochondrial genome. Neighbor-joining, maximum parsimony and Bayesian inference were performed. The best phylogenetic reconstruction was obtained using the COI region, and we consider it an appropriate marker for phylogenetic studies within the group. Consensus analysis (COI +16S rRNA) generally obtained the same tree topologies and confirmed that the genus Nerita is monophyletic. The consensus analysis using parsimony recovered a monophyletic group consisting of the genera Neritina , Septaria , Theodoxus , Puperita , and Clithon , while in the Bayesian analyses Theodoxus is separated from the other genera. -
Estudio De Referencia Del Johnny Cay Regional Park En La Isla De San Andrés - Reserva De La Biosfera Seaflower
Estudio de referencia del Johnny Cay Regional Park en la Isla de San Andrés - Reserva de la Biosfera Seaflower Item Type Book/Monograph/Conference Proceedings Authors Vides Casado, Martha; Alonso Carvajal, David Publisher INVEMAR Download date 08/10/2021 21:39:16 Link to Item http://hdl.handle.net/1834/15822 ESTUDIO DE REFERENCIA del Johnny Cay Regional Park en la isla de San Andrés - Reserva de la Biosfera Seaflower RESERVA DE BIOSFERA SEA FLOWER ESTUDIO DE REFERENCIA del Johnny Cay Regional Park en la isla de San Andrés - Reserva de la Biosfera Seaflower RESERVA DE BIOSFERA SEA FLOWER Financiado por el Proyecto Biodiversidad Fondo de Compensación Ambiental 2018 Director EQUIPO TÉCNICO: Francisco A. Arias Isaza Gloria Murcia, Andrés Acosta Chaparro, Juan González Co- rredor, Laura Sánchez Valencia, Raúl Navas Camacho, Dia- Subdirector na Isabel Gómez, Oswaldo Coca Domínguez, Constanza Coordinador de Investigaciones (SCI) Ricaurte Villota, Daniela Yepes Gaurisas, Adibe Cárdenas Jesús Antonio Garay Tinoco Oliva, Bibian Martínez, Catalina Arteaga Flórez, Fernando Dorado Roncacio, Karen Ayala Galván, Maria Alejandra Subdirectora Administrativa (SRA) Mutis Martinezguerra, Andrea Dueñas, Martha Vides, San- Sandra Rincón Cabal tiago Alonso Millán Cortés, Felipe Valencia Ospina, Julián Pizarro Pertuz y David Alejandro Alonso C. Coordinadora de Investigación e Información para la Gestión Marina y Costera (GEZ) Cartografía: Santiago Millán y Felipe Valencia Paula Cristina Sierra Correa Apoyo en campo: Equipo Logístico CORALINA Coordinador Programa Biodiversidad y Ecosistemas Marinos (BEM) Imagen portada: Vista aérea de Johnny Cay, 2017 David Alonso Carvajal (Martha Vides) Coordinadora Imágenes contraportada: Juan Carlos Márquez Programa de Geociencias Marinas (GEO) Constanza Ricaurte Villota Montaje: John Khatib/Daiver García (ediprint.com.co) Coordinadora Se imprimen 320 ejemplares, Programa Calidad Ambiental Marina (CAM) Bogotá D.C., Colombia • Noviembre de 2018 Luisa Fernanda Espinosa Impresión: Ediprint S.A.S. -
Guide to the Systematic Distribution of Mollusca in the British Museum
PRESENTED ^l)c trustee*. THE BRITISH MUSEUM. California Swcademu 01 \scienceb RECEIVED BY GIFT FROM -fitoZa£du^4S*&22& fo<?as7u> #yjy GUIDE TO THK SYSTEMATIC DISTRIBUTION OK MOLLUSCA IN III K BRITISH MUSEUM PART I HY JOHN EDWARD GRAY, PHD., F.R.S., P.L.S., P.Z.S. Ac. LONDON: PRINTED BY ORDER OF THE TRUSTEES 1857. PRINTED BY TAYLOR AND FRANCIS, RED LION COURT, FLEET STREET. PREFACE The object of the present Work is to explain the manner in which the Collection of Mollusca and their shells is arranged in the British Museum, and especially to give a short account of the chief characters, derived from the animals, by which they are dis- tributed, and which it is impossible to exhibit in the Collection. The figures referred to after the names of the species, under the genera, are those given in " The Figures of Molluscous Animals, for the Use of Students, by Maria Emma Gray, 3 vols. 8vo, 1850 to 1854 ;" or when the species has been figured since the appear- ance of that work, in the original authority quoted. The concluding Part is in hand, and it is hoped will shortly appear. JOHN EDWARD GRAY. Dec. 10, 1856. ERRATA AND CORRIGENDA. Page 43. Verenad.e.—This family is to be erased, as the animal is like Tricho- tropis. I was misled by the incorrectness of the description and figure. Page 63. Tylodinad^e.— This family is to be removed to PleurobrancMata at page 203 ; a specimen of the animal and shell having since come into my possession. -
Late Eocene and Early Oligocene) of the Hampshire Basin
Cainozoic Research, 4(1-2), pp. 27-39, February 2006 The Neritidae of the Solent Group (Late Eocene and Early Oligocene) of the Hampshire Basin M.F. Symonds The Cottage in the Park, Ashtead Park, Ashtead, Surrey KT21 1LE, United Kingdom Received 1 June 2003; revised version accepted 7 March 2005 Gastropods of the family Neritidae in the Solent Group of the Hampshire Basin, southern England are reviewed and two previously unde- scribed taxa are described. New genus: Pseudodostia. New species: Clithon (Pictoneritina) cranmorensis and Clithon (Vittoclithon) headonensis. Neotypes designated for Neritina planulata Edwards, 1866 and Neritina tristis Forbes, 1856. Amended diagnosis: subgenus Vittoclithon. New combinations: Pseudodostia aperta (J. de C. Sowerby, 1823), Clithon (Pictoneritina) concavus (J. de C. Sowerby, 1823), Clithon (Pictoneritina) planulatus (Edwards, 1866) and Clithon (Pictoneritina) bristowi Wenz, 1929. KEY WORDS: Mollusca, Gastropoda, Neritidae, Palaeogene, Hampshire Basin. Introduction Systematic Palaeontology Family Neritidae Rafinesque, 1815 Although the number of species of Neritidae in the Solent Genus Pseudodostia gen. nov. Group is rather limited, specimens are common at certain horizons and they have received the attention of numerous Type species — Nerita aperta J. de C. Sowerby, 1825. Eo- authors in the past. In particular Curry (1960, 265-270) cene, Headon Hill Formation. dealt in detail with the taxonomy of Theodoxus concavus (J. de C. Sowerby, 1823), Theodoxus planulatus (Edwards, Derivatio nominis — The name reflects the close resem- 1866) and Theodoxus bristowi Wenz, 1929. The purpose of blance between the shell of the type species of this genus this paper is to update Curry’s work and to cover additional and that of Nerita crepidularia Lamarck, 1822, the type species. -
Mechanical Basis of Morphogenesis and Convergent Evolution of Spiny Seashells
Mechanical basis of morphogenesis and convergent evolution of spiny seashells Régis Chirata, Derek E. Moultonb,1, and Alain Gorielyb aCentre National de la Recherche Scientifique 5276, Université Lyon, 69622 Villeurbanne Cedex, France; and bOxford Centre for Collaborative Applied Mathematics, Mathematical Institute, University of Oxford, Oxford OX1 3LB, United Kingdom Edited by George Oster, University of California, Berkeley, CA, and approved February 22, 2013 (received for review November 26, 2012) Convergent evolution is a phenomenon whereby similar traits that is, we must ask whether the morphogenetic processes that evolved independently in not closely related species, and is often actually shape these structures make them likely to reappear. In interpreted in functional terms. Spines in mollusk seashells are this study, we focus on the mechanistic approach and the natural classically interpreted as having repeatedly evolved as a defense in mechanical process that shapes spines in mollusk shells and response to shell-crushing predators. Here we consider the mor- underlies their repeated emergence. phogenetic process that shapes these structures and underlies their Spines, like other ornamentations found in seashells, are in- repeated emergence. We develop a mathematical model for spine crementally secreted and represent the spatiotemporal record of morphogenesis based on the mechanical interaction between the the shape modifications of the shell edge during growth. Spines secreting mantle edge and the calcified shell edge to which the first emerge as a bulge of the shell edge that curves both longi- mantle adheres during shell growth. It is demonstrated that a large tudinally and transversely as growth proceeds, resulting in a fold diversity of spine structures can be accounted for through small that eventually closes in on itself as the lateral edges converge variations in control parameters of this natural mechanical process.