ZOOTAXA

1799

Revision of the sap genus Pocadius Erichson, 1843 (Coleoptera: Nitidulidae: )

ANDREW R. CLINE

Magnolia Press Auckland, New Zealand Andrew R. Cline Revision of the genus Pocadius Erichson, 1843 (Coleoptera: Nitidulidae: Nitidulinae) (Zootaxa 1799) 120 pp.; 30 cm. 16 June 2008 ISBN 978-1-86977-217-8 (paperback) ISBN 978-1-86977-218-5 (Online edition)

FIRST PUBLISHED IN 2008 BY Magnolia Press P.O. Box 41-383 Auckland 1346 New Zealand e-mail: [email protected] http://www.mapress.com/zootaxa/

© 2008 Magnolia Press All rights reserved. No part of this publication may be reproduced, stored, transmitted or disseminated, in any form, or by any means, without prior written permission from the publisher, to whom all requests to reproduce copyright material should be directed in writing. This authorization does not extend to any other kind of copying, by any means, in any form, and for any purpose other than private research use.

ISSN 1175-5326 (Print edition) ISSN 1175-5334 (Online edition)

2 · Zootaxa 1799 © 2008 Magnolia Press CLINE Zootaxa 1799: 1–120 (2008) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA Copyright © 2008 · Magnolia Press ISSN 1175-5334 (online edition)

Revision of the sap beetle genus Pocadius Erichson, 1843 (Coleoptera: Nitidulidae: Nitidulinae)

ANDREW R. CLINE Plant Pest Diagnostics Center, California Department of Food & Agriculture, 3294 Meadowview Rd., Sacramento, CA 95832-1448, E-mail: [email protected]

Table of contents

Abstract ...... 4 Resumen ...... 4 Introduction ...... 5 Methods...... 7 Generic Redescription ...... 9 Key to Species ...... 11 Species (Re)descriptions ...... 16 Pocadius adustus Reitter, 1888...... 16 Pocadius africanus Kraatz, 1895 ...... 17 Pocadius antennuliferus Cline new species ...... 26 Pocadius ashei Cline new species ...... 31 Pocadius barclayi Cline new species ...... 33 Pocadius basalis Schaeffer, 1911...... 36 Pocadius bicolor Cline new species ...... 40 Pocadius brevis Reitter, 1873 ...... 41 Pocadius carltoni Cline new species ...... 43 Pocadius centralis Cline new species ...... 44 Pocadius cochabambus Cline new species ...... 46 Pocadius coxus Cline new species ...... 47 Pocadius crypsis Cline new species ...... 49 Pocadius decoratus Kirejtshuk, 1984 ...... 51 Pocadius dimidiatus Jelínek, 1977 ...... 52 Pocadius dominicus Cline new species ...... 54 Pocadius endroedyi Cline new species ...... 56 Pocadius ephite Leschen and Carlton, 1994 ...... 58 Pocadius falini Cline new species ...... 59 Pocadius fasciatus Cline new species ...... 61 Pocadius femoralis Cline new species ...... 63 Pocadius ferrugineus (Fabricius, 1775) ...... 64 Pocadius fulvipennis Erichson, 1843 ...... 66 Pocadius fumatus Jelínek, 1977 ...... 68 Pocadius fusiformis Cline new species ...... 70 Pocadius globularis Cline new species ...... 72 Pocadius helvolus Erichson, 1843 ...... 73 Pocadius insularis Cline new species ...... 75 Pocadius jelineki Leschen and Carlton, 1994 ...... 77

Accepted by D. Pollock: 7 Feb. 2008; published: 16 Jun. 2008 3 Pocadius kirejtshuki Cline new species ...... 79 Pocadius luisalfredoi Cline new species ...... 81 Pocadius majusculus Kirejtshuk, 1984 ...... 83 Pocadius maquipucunensis Leschen and Carlton, 1994 ...... 84 Pocadius martini Kirejtshuk, 1984 ...... 86 Pocadius monticola Lechanteur, 1959 ...... 88 Pocadius niger Parsons, 1936...... 89 Pocadius nigerrimus Cline new species ...... 91 Pocadius nobilis Reitter, 1873 ...... 93 Pocadius okinawaensis Cline new species ...... 94 Pocadius pecki Cline new species ...... 96 Pocadius peruensis Cline new species...... 98 Pocadius rubidus Erichson, 1843 ...... 99 Pocadius tepicensis Cline new species ...... 101 Pocadius testaceus Grouvelle, 1892 ...... 103 Pocadius torresi Jelínek, 1977 ...... 105 Pocadius wappesi Cline new species ...... 107 Pocadius yunnanensis Grouvelle, 1910 ...... 111 Acknowledgments ...... 116 Literature Cited ...... 118

Abstract

A total of 47 species are included in a revisionary treatment of Pocadius Erichson. Twenty-one species are redescribed, P. yunnanensis Grouvelle is resurrected, and 25 are described as new, including: P. antennuliferus (Brazil), P. ashei (Bolivia), P. barclayi (Sulawesi), P. bic o lo r (Brazil), P. carltoni (Nicaragua), P. centralis (Central America), P. cocha- bambus (Bolivia), P. coxus (Brazil), P. crypsis (Guyana), P. dominicus (Dominican Republic), P. endroedyi (southern Africa), P. falini (Paraguay), P. fasciatus (Borneo), P. femoralis (Vietnam), P. fusiformis (Sulawesi), P. globularis (Hon- duras), P. insularis (Trinidad), P. kirejtshuki (Australia), P. luisalfredoi (Mexico), P. nigerrimus (Paraguay), P. oki- nawaensis (Okinawa), P. pecki (Venezuela), P. peruensis (Peru), P. tepicensis (Mexico), P. w a pp e si (Bolivia). An adult and larval description for the genus also is provided and its phylogenetic position within Nitidulinae discussed. A key to all species is included, as well as a checklist of species and their distributions.

Key words: Nitidulidae, Pocadius, , monograph, Nitidulinae, new species

Resumen

Un total de 47 especies están incluidos en el tratamiento “revisionary” de Pocadius Erichson. Veintiuno de los especies están descrito de nuevo, P. yunnanensis Grouvelle fue resucitado, y 25 de los que están descritos como nuevos, incluy- endo: P. antennuliferus (Brazil), P. as he i (Bolivia), P. b ar c la y i (Sulawesi), P. b ic o lo r (Brazil), P. c a rlt oni (Nicaragua), P. centralis (Central America), P. cochabambus (Bolivia), P. c o xu s (Brazil), P. crypsis (Guyana), P. do m inic u s (Dominican Republic), P. endroedyi (southern Africa), P. falini (Paraguay), P. fa sc i atu s (Borneo), P. f e m or al is (Vietnam), P. fusifor- mis (Sulawesi), P. globularis (Honduras), P. i nsu lar is (Trinidad), P. k ir ej tshu ki (Australia), P. luisalfredoi (Mexico), P. nigerrimus (Paraguay), P. okinawaensis (Okinawa), P. pecki (Venezuela), P. peruensis (Peru), P. tepicensis (Mexico), P. wappesi (Bolivia). Una descripción de adulto y larval para el generó también esta proporcionada y su posición dentro del Nitidulinae esta discutido. Una clave a todas estas especies esta incluido, también una lista de comprobación de especies y sus distribuciones.

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