VASCULAR PLANTS of the HEADWATERS SANCTUARY Patty Leslie Pasztor - April, 2013

VASCULAR PLANTS OF THE HEADWATERS SANCTUARY Patty Leslie Pasztor - April, 2013

* Non-native species Scientific names in parentheses indicate former names or synonyms

ACANTHACEAE ACANTHUS FAMILY

Dicliptera brachiata branched foldwing / dicliptera Justicia pilosella (Siphonoglossa pilosella) tube-tongue * Ruellia brittoniana mexican petunia Ruellia drummondiana Drummond's petunia Ruellia nudiflora wild petunia

ACERACEAE MAPLE FAMILY Acer negundo box elder

AMARANTHACEAE AMARANTH FAMILY * Alternanthera philoxeroides alligator-weed Amaranthus palmeri careless-weed / pigweed / redroot Celosia nitida slender celosia

ANACARDIACEAE SUMAC FAMILY Rhus aromatica (sw fence /281 entrance) aromatic or skunkbush sumac * Pistacia chinensis chinese pistache Toxicodendron radicans poison-ivy / poison oak

APIACEAE (UMBELLIFERAE) CARROT FAMILY * Torilis arvensis beggar's lice / sockbane Chaerophyllum tainturieri var. tainturieri chervil

AQUIFOLIACEAE HOLLY FAMILY Ilex decidua possum haw / deciduous holly

ARECACEAE PALM FAMILY Sabal mexicana sabal palm (native to deep S. TX, naturalized in the S.A. area) * Washingtonia filifera California fan palm

ASCLEPIADACEAE MILKWEED FAMILY Cynanchum barbigerum swallow-wort /climbing milkweed Matelia reticulata (St. Brigid's path slope) pearl milkweed vine

ASTERACEAE (COMPOSITAE) SUNFLOWER FAMILY Ageratina havanensis (bluff & sw fence area) bush boneset / white mistflower Ambrosia trifida giant ragweed Ambrosia psilostachya western ragweed Amphiachyris dracunculoides broomweed Baccharis neglecta false willow/ roosevelt weed Calyptocarpus vialis straggler daisy / horseherb Gaillardia pulchella firewheel / indian blanket Helianthus anuus annual sunflower * Lactuca seriola prickly lettuce Parthenium hysterophorus false ragweed Simsia calva bush sunflower (uplands) * Sonchus asper sow thistle Symphyotrichum drummondii var texanum (A. texanus) Drummond's aster / Texas aster Symphyotrichum subulatum (Aster subulatus) hierba del marrano / annual aster * Taraxacum officinale dandelion Thelesperma filifolium greenthread Thymophylla pentachaeta paralena / dogweed (uplands) Verbesina virginiana frostweed Viguiera dentata plateau goldeneye / goldeneye Wedelia texana (Zexmenia hispida) orange zexmenia / wedelia

BERBERIDACEAE BARBERRY FAMILY Berberis trifoliolata agarita * Nandina domestica nandina

BIGNONIACEA CATALPA OR TRUMPET-VINE FAMILY Campsis radicans trumpet-creeper / trumpet-vine * Macfadyena unguis-cati catclaw vine

BORAGINACEAE BORAGE FAMILY Ehretia anacua anacua Lithospermum matamorense rough gromwell

BRASSICACEAE (CRUCIFERAE) MUSTARD FAMILY * Capsella bursa-pastoris shepperd's-purse Lepidium virginicum virginia peppergrass * Rapistrum rugosum turnipweed / bastard cabbage

BROMELIACEAE PINEAPPLE FAMILY Tillandsia recurvata ball moss Tillandsia usneoides spanish moss

CACTACEAE CACTUS FAMILY Opuntia engelmannii var. lindheimeri Lindheimer's prickly pear Opuntia leptocaulis (sw, 281 fence area) tasajillo / pencil cactus

CAPRIFOLIACEA HONEYSUCKLE FAMILY Lonicera japonica japanese honeysuckle Sambucus nigra ssp. canadensis elderberry

2 CARYOPHYLLACEAE PINK FAMILY * Stellaria media chickweed

CHENOPODIACEAE GOOSEFOOT FAMILY * Chenopodium album lamb's quarters

COMMELINACEAE DAYFLOWER FAMILY Commelina erecta widow's tears / dayflower Tinatia anomala (Commelinantia anomala) false dayflower Tradescantia sp. spiderwort

CONVOLVULACEAE MORNING GLORY FAMILY Convolvulus equitans Texas bindweed Dichondra carolinensis Carolina ponyfoot Ipomoea cordatotriloba var. cordatotriloba morning glory/ purple bindweed Merremia dissecta alamo vine

CORNACEAE DOGWOOD FAMILY Cornus drummondii rough-leaf dogwood

CUCURBITACEAE GOURD FAMILY Cucurbita foetidissima buffalo gourd Melothria pendula meloncito

CUPPRESACEAE CYPRESS FAMILY Juniperus sp. juniper Taxodium distichum bald cypress (planted)

CYPERACEAE SEDGE FAMILY Carex perdentata meadow sedge Carex sp. sedge sp. Cyperus ochraceus umbrella sedge / pond flat sedge * Cyperus rotundus purple nutsedge Eleocharis sp. spikerush / spikesedge

DRYOPTERIDACEAE WOOD FERN FAMILY * Cyrtomium falcatum Holly Fern / Japanese holly fern

EUPHORBIACEAE SPURGE FAMILY Acalypha phleoides (A. lindheimeri) three-seeded mercury Bernardia myricifolia oreja de raton / mouse ears Chamaesyce prostrata (Euphorbia prostrata) prostrate spurge Euphorbia dentata toothed spurge * Triadica sebifera (Sapium sebiferum) Chinese tallow tree Tragia racemosa nose-burn

3 EBENACEAE EBONY FAMILY Diospyros texana Texas persimmon

FABACEAE (LEGUMINOSAE) LEGUME FAMILY Acacia farnesiana (A. smalii) huisache Acacia roemeriana (bluff, sw fence) cat claw acacia / roemer's acacia Eysenhardtia texana kidneywood * Medicago polymorpha bur-clover * Melilotus albus white sweet clover * Melilotus minima least bur-clover Parkinsonia aculeata retama Prosopis glandulosa mesquite Rynchosia minima snoutbean Styphnolobium affine (Sophora affinis) eve’s necklace Sophora secundiflora TX mountain laurel / mescalbean Vicia ludoviciana vetch / deer vetch

FAGACEAE BEECH OR OAK FAMILY Quercus buckleyi (Q. texana) Texas red oak / Spanish oak Quercus fusiformis plateau live oak Quercus shumardii (possibly naturalized) Shumard Oak

FLACOURTIACEAE FLACOURTIA FAMILY *Xylosma congestum xylosma

FUMARIACEA FUMITORY FAMILY Corydalis curvisiliqua scrambled eggs

GERANIACEAE GERANIUM FAMILY * Erodium cicutarium stork's bill / filaree Erodium texanum Texas stork's bill Geranium carolinianum wild geranium

HYDROPHYLLACEAE WATERLEAF FAMILY Nemophila phacelioides baby blue eyes Phacelia congesta blue curls

IRIDACEAE IRIS FAMILY Sisyrinchium langloisii Blue-eyed grass

JUGLANDACEAE WALNUT FAMILY Carya illinoinensis pecan Juglans major, J. nigra walnut

LAMIACEAE MINT FAMILY * Lamium amplexicaule henbit

4 LILIACEAE LILY FAMILY Allium canadense wild onion Allium drummondii wild garlic Cooperia drummondii and C. pedunculata rainlily Nothoscordum bivalve crow poison

MALVACEAE MALLOW FAMILY Abutilon fruticosum indian mallow Callirhoe leiocarpa winecup / tall or annual winecup Malvastrum coromandelianum threelobe false mallow Malvaviscus arboreus var. drummondii turk’s cap Sida abutifolia (S. procumbens) spreading sida

MELIACEAE MAHOGANY FAMILY *Melia azedarach chinaberry

MENISPERMACEAE MOONSEED FAMILY Cocculus carolinus carolina snailseed

MORACEAE MULBERRY FAMILY *Broussonetia papyrifera paper mulberry *Morus alba white mulberry Morus rubra red mulberry

NYCTAGINACEAE FOUR-O’CLOCK FAMILY Boerhavia diffusa scarlet spiderling * Mirabilis jalapa four-o’clock

OLEACEAE ASH FAMILY Forestiera pubescens elbowbush / spring herald Fraxinus berlandieriana berlandier ash / fresno Fraxinus pennsylvanica green ash Ligustrum sinense chinese privet * Ligiustrum lucidum ligustrum * Ligustrum japonicum ligustrum

ONAGRACEAE EVENING PRIMROSE FAMILY Oenothera speciosa evening primrose Ludwigia octovalvis water primrose

OXALIDACEAE WOOD-SORREL FAMILY Oxalis dillenii wood sorrel / sour clover (yellow) Oxalis drummondii wood sorrel / sour clover (purple)

PASSIFLORACEAE PASSION-FLOWER FAMILY Passiflora lutea yellow passionflower Passiflora foetida corona de cristo / passionflower

PHYTOLACCACEAE POKEWEED FAMILY Phytolacca americana pokeweed Rivina humilis pigeonberry

PLATANACEAE SYCAMORE (PLANE-TREE) FAMILY Platanus occidentalis sycamore

POACEAE (GRAMINAE) GRASS FAMILY * Bothriochloa ischaemum var. songarica King Ranch (KR) bluestem Bothriochloa laguroides (B. saccharoides) silver bluestem * Bromus arvensis (B. japonicus) Japanese brome * Bromus catharticus (B. unioloides) rescue grass Chasmanthium latifolium inland sea oats / river oats * Cynodon dactylon bermudagrass * Echinochloa colona jungle rice * Eleusine indica goosegrass Elymus virginicus Virginia wildrye * Lolium perenne perennial ryegrass Nasella leucotricha Texas wintergrass /Speargrass * Paspalum dilatatum dallisgrass Paspalum langei rustyseed paspalum Setaria leucopila bristlegrass Setaria scheelei southwestern bristlegrass * Sorghum halapense johnson grass Tridens albescens white tridens

POLYGONACEAE BUCKWHEAT FAMILY Polygonum pensylvanicum (Persicaria bicornis) smartweed * Rumex crispus curly dock

PTERIDACEAE MAIDENHAIR FERN FAMLY Adiantum capillus-veneris maidenhair-fern

RANUNCULAEACE BUTTERCUP FAMILY Anemone berlandieri (A. heterophylla) anemone / wind flower Clematis drummondii old man's beard / barba de viejo * Clematis terniflora sweet-autumn clematis Ranunculus muricatus buttercup

RHAMNACEAE BUCKTHORN FAMILY Condalia hookeri brasil / bluewood condalia Colubrina texensis hogplum Ziziphus obtusifolia lotebush

6 ROSACEAE ROSE FAMILY * Eriobotrya japonica loquat Geum canadense white avens * Photinia serrulata chinese photinia Prunus caroliniana cherry laurel (native to eastern Tx) Rubus trivialis dewberry

RUBIACEAE MADDER FAMILY Galium aparine Bedstraw/ cleavers

RUTACEAE CITRUS FAMILY Ptelea trifoliata wafer ash / hop-tree

SALICACEAE WILLOW FAMILY Populus deltoides cottonwood, eastern Salix nigra black willow

SAPINDACEAE SOAPBERRY FAMILY * Koelreuteria paniculata rain tree / golden rain-tree Ungnadia speciosa mexican buckeye Sapindus saponaria var. drummondii soapberry

SAPOTACEAE SAPODILLA FAMILY Sideroxylon lanuginosum (Bumelia lanuginosa) gum bumelia

SCROPHULARIACEA SNAPDRAGON FAMILY Maurandya antirrhiniflora snapdragon vine

SIMAROUBACEAE QUASSIA FAMILY * Ailanthus altissima tree of heaven

SMILACACEAE GREENBRIAR FAMILY Smilax bona-nox greenbriar

SOLANACEAE NIGHTSHADE FAMILY Capsicum annuum var. gabriusculum chile pequín / chile petín physalis cinarescens ground cherry Solanum eleagnifolium silver-leaf nightshade Solanum triquetrum Texas nightshade

ULMACEAE ELM FAMILY Celtis laevigata var. laevigata sugarberry Celtis laevigata var reticulata netleaf hackberry Celtis ehrenbergiana (C. pallida) spiny hackberry / granjeno Ulmus americana american elm Ulmus crassifolia cedar elm

7 URTICACEAE NETTLE FAMILY Parietaria pensylvanica pellitory / cucumber plant

VERBENACEAE VERBENA FAMILY Aloysia gratissima whitebrush / beebrush Phyla nodifera frogfruit Glandularia bipinnatifida (Verbena bipinnatifida) prairie verbena Glandularia cannescens gray verbena / vervain Glandularia pumila low vervain / verbena * Lantana camara lantana / calico-bush * Lantana montevidensis purple trailing lantana Lantana urticoides (L. horrida) Texas native lantana * Verbena brasiliensis brazilian verbena or vervain Verbena cannescens gray vervain / verbena Verbena halei Texas vervain / verbena * Vitex agnus-castus chaste-tree / monk's chaste

VIOLACEAE VIOLET FAMILY Viola missouriensis (V. sororia var. missourensis) missouri violet

VISCACEAE MISTLETOE FAMILY Phoradendron tomentosum mistletoe

VITACEAE GRAPE FAMILY Ampelopsis arborea peppervine Ampelopsis cordata heart-leaf ampelopsis Cissus trifoliata (C. incisa) cow-itch vine / ivy treebine Parthenocissus quinquefolia virginia creeper Vitis cinerea var. helleri wild grape Vitis mustangensis mustang grape

ZYGOPHYLLACEA CALTROP FAMILY Guajacum agustifolium (Guaiacum angustifolium) Guayacan

Plant list compiled from informal field surveys from 2010-2011 by Patty Leslie Pasztor and for the Biodiversity Study 2012-2013 by Patty Leslie Pasztor, with baseline vegetation sampling by Jennifer Ince of Trinity University and Hannah Peterson of University of the Incarnate Word. Previous informal surveys conducted by Bill Carr, March, 2008, Bill Carr and Lacey Halstead, 2007, and the Ladybird Johnson Wildflower Center, 2006.

Scientific names follow Jones and Wipff (2003), Diggs, Lipscomb, and O'Kennon (1999), and updates from Flora North America Editorial Committee (1993+). Common names follow Correll & Johnston (1970), Diggs et al. (1999) and various other sources.

References

Correll, D. S., and Marshall C. Johnston. 1970. Manual of the Vascular Plants of Texas. Texas Research Foundation, Renner. 1881 pp.

Cox, P. W. and P. Leslie. 1991. Texas Trees - A Friendly Guide. Corona Publishing Co., San Antonio. 374 pp.

Diggs, G. M., Jr., B. L. Lipscomb and R. J. O'Kennon. 1999. Shinners and Mahler's Illustrated Flora of North Central Texas. Botanical Research Institute of Texas, Ft. Worth. 1626 pp.

Enquist, Marshall. 1987. Wildflowers of the Texas Hill Country. Lone Star Botanical Press, Austin. 275 pp.

Flora of North America Editorial Committee, eds. 1993+. Flora of North America North of Mexico. 16+vols. New York and Oxford.

Gould, F. W. 1975. The grasses of Texas. Texas A & M University Press, College Station. 653 pp.

Johnson, M.C. 1990. The Vascular Plants of Texas: A List Updating The Manual Of The Vascular Plants of Texas. 2nd edition. Privately published, Austin. 165 pp.

Jones, S. D. & J. K. Wipff. 2003. A 2003 updated checklist of the vascular plants of Texas. Botanical Research Center, Bryan, TX, 712 pp.

Jones, S. D., J. K. Wipff and P. M. Montgomery. 1997. Vascular plants of Texas: a comprehensive checklist including synonymy, bibliography, and index. University of Texas Press, Austin. 404 pp.