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Discovery of Wockia Koreana Sohn（Lepidoptera， Urodidae）

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Discovery of Wockia Koreana Sohn（Lepidoptera， Urodidae） The LepldopterologlcalLepidopterological SocSociety⊥ et 二y of Japan − 蝶 と蛾 Lepidoptera Science 66（2）：58 61， October 2015 日本 か らの Wockia koreana Sohn （メ ス ハ リオガ科 Urodidae）の発 見 お よび メ ス ハ リオガ科 と Wockia属 に つ い て 那須義次 D ・宮野昭彦2 ｝ 1｝ 648−0023 和歌山県橋本市中道153−29 2｝ 509−0104 岐阜県各務原市各務おが せ 町9−96−2 Discovery Wockia koreana of Sohn （Lepidoptera， Urodidae）fromJapan， Withnotes on thefamily Urodidae and the genus of Wockia Heinemann Y ｛）shitsugu NAsuo and Akihiko MlyANo2〕 D153 − − 2、 Nakado， Hashimoto， Wakayama ，648 0023 Japan 2） 9−96−2，Kakamiogase−cho ， Kakamigahara，509−OlO4 Japan Abstract Two Japanese species of W ∂ckia are treated． Wockia たo 厂ea ’za is newly recorded from Japan， with mustra【ions of adult and gen註alia ， A color photograph of adult 毘 magna is also provided． The phylogeny of the family Urodidae and themorpho 】Qgy and biology of the genus Wockia are summarized ． Key wor 〔ls genitalia， new record ， Wockia koreana， Wockia magna 、 ー メ ス ハ リ オ ガ 科Urodidaeは ヨ ロ ッ パ ，東 ア ジ ア ，東南 ア を受け，日本産 の ネマ ル ハ キ バ ガ 科Blastobasidaeに 似 た 未 ジ ア ，南北 ア メ リ カ に 分 布 し．6属 約70種が 記録 され て い 知 の 種 を検討 した と こ ろ ，韓 国 か ら記 載 さ れ たWockia る （那須，2011 ；Sohn，2014）．本科は 日本か らは従来記録 koreana Sohn，2008で あ る こ とが判 明 した，本報告 にお い て ， さ れ て い なか っ た （那須，2011）が，2014年 にSohn氏 によ wkoreana につ い て 成虫 と交尾器 の 図 を添 え て 記録す る と り，ア メ リ カ 国立 自然 史博物館に所 蔵 され て い る 日本産標 ともに，Wmagna に つ い て も成 虫 の 図 を示す，あ わせ て ， 一 本 （ 色 コ レ ク シ ョ ン ）お よ び 韓 国産標本 に基 づ きWoekia 日本 で はあ ま り知 られ て い ない メ ス ハ リオ ガ科 の 研究進展 magna Sohn，2014が 新種記載 され る と と もに，本科 の 日本 の ため，本科 の 系統 関係 お よびWockiaecの 特徴 と生態 に つ か ら の 初 記録 が な さ れ た （Sohn，2014），筆者 ら は 彼 の 発見 い て 紹介す る ． 2Figs − l 5．Adults of Wockia spp ． − （1；慨 magna ，2 4：Wkoreana ， Japan）．1．Holotype， male ， in the National Museum of Natural History USA by ， （photograph − J．C ． Sohn）．2． Male．3． Eye cap in male antenna base，4 ． Frenulum infemale．5． Second abdominal stemite ， 一 NNII-Electronic工 工 Electronlc LibraryL − brary Service The LepldopterologlcalLepidopterological 30cSociety⊥ ety of Japan 日本産Wockia属 59 ア メ リ カ 国立 自然 史博物館 の J．C ． Sohn氏 に は Wmagna の チ ャ オ ビ メ ス ハ リ オガ （新 称）Wockia koreana Sohn，2008 − 成虫 図お よ び 文献 を提供 い た だ い た ．本論文 で 用 い た 豊山 （Figs 2 10） 市産 の 標 本 は す べ て 豊 田市 に よ る 新編 豊田市史編纂の た め Wockia korean‘l Sehn，2008 ：Sohn and Adamski ，2008 ；557， の 調査 によ り得 た もの で ，こ の 調査 で はあ ま市の 間野隆裕 − 二 い 氏 に お世話 に な っ た．こ の 両氏 に 厚 く御礼 申 し上 げる． 判別．成虫〔Figs 2 5）．性的 型 は 触角基部 に お て 顕著で ， ♂の 触角基 部は広 が り眼帽 を形 成 す る（Fig 3）．前翅 長 は イ ッ シ キ メ ス ハ リオ ガ （新称）Wockia magna Sohn ．2014 6 ．0−7．Omm ．前翅 は 披針形状，灰色 で 中央 に 茶 褐色 の 横帯 （Fig．1） ’ をもち，横帯上 の 鱗粉 は 盛 り上 が る ； そ の 帯 よ り先端部 分 Wo ‘kia magna Sohn，2014： Sohn，2〔〕14：260． に2 つ の 茶褐 色紋 を もつ ，本種 は 前種 Wmagna に 類 似す る 判 別．成 虫（Fig．1）の 前翅長 は 7．8−8．Onlm．前翅 は 披針 形 状 ， が ．♂ の 触角基 部 は 眼 帽 を もつ こ と，前翅 長 は よ り短 い こ 灰色 で 中央 に 暗褐灰色の 横帯を もち，横帯上 の 鱗粉 は 盛 り と，♂交尾器（Figs 6，7）の バ ル バ は大 き くくびれ， ク ク ル つ バ バ 上 が る ．本種 は 次種理 koreana に 類似す る が 1 識別点 は 次 ス は 三 角形状 で 針状 の 刺毛を も こ と （前種の ル 先端 種 の 判別文 を参照， は 尖り，ク ク ル ス は 発達 し な い ），バ ル バ の くび れ 部 の 腹 面は先端が尖 っ た強 く硬化 した突起をもつ こ と （前種 で は 分布．日本 （本州 ： 中部地 方），韓国． ・ こ の 突起 をもた な い ），♀交尾器（Figs 8−1｛｝）の ドゥ ク ツ ス 寄主植物．不明． ブ ル サ エ は全体が硬化する こ と，2個の li亅錐状 の シ グナを もつ こ と（Fig，10）（前種 の シ グ ナ は 歯状） で 区別で きる ． 生態．成虫は中部地 方 で は9月中旬 に 採集 され て い る ． 本種 の 原記載 に よ る と，前翅 の 長 さ は 13．3−15」 mnn と記述 され て い るが，こ れ は 前翅 開張 の 間違 い で あ る （Sohn，私信， 2015）． 〔 丶 ／ 、 ＼ 嚇 盛 〆 懸 鸛 駄 ノ 7 − Figs 6 10． Genitalia of Wockia koreana， Japan，6， Male． 7．Aedeagus in maEe ．8、 Female． 9．Ostiu皿 and du¢ tus bursae in 8 female． lo， Ductus bursae and signa in felnale． 一 NNII-Electronic工 工 Electronlc LibraryL − brary Service The LepldopterologlcalLepidopterological SocletySociety of Japan 60 那 須義次 ・宮 野 昭 彦 − 所 検標 本，成 虫，JAPAN ； Honshu ： Gifu Pref．， Hichisocho ， た め 識 別 叮 能 で あ る ， − Murogane，1♂，24， viii ．2002；Mitake cho ， Okugo ，1♂，21．vii ． − − 本属 で 寄主植物 が 判明 し て い る 3種 の 寄主 は い ずれ もヤ ナ 2005 ；Yamagata shi ， Miyama cho ，1♂，28． viii ．2007． Aichi ’ コ 毘 cheve hacca − − ギ 科 で あ る ． メ キ シ の Adamski，2009 は Pref．，Toyota shi ， Shimokawaguchi cho ，1♂，2， viii ．20121 ー − c αsearia nitida 〔L ）Jacq．を（Adamski et at ．，2009），北米 と ヨ Mitsukuricho ，1♀，30． vii ．20 ］2；Kise・cho ，1♂，21．vii，2011，1 ロ パ ーロ パ ッ の Wasperipunctella（Brし且and ．1851）は ヨ ッ ヤ マ ♀，26，viii．2011，1♀，3Lviii．2012，1♂，5．ix．2013． ・ ー ナ ラ シ Pt）pulus trtmt ‘la L ．， ヨ ロ ッ パ ク ロ ヤ マ ナ ラ シ R 標本 は すべ て 宮野昭彦 に よ っ て 灯火採集 で 採集 され た もの nigra L ．とSalix eleagn θ s Scop，を（Kyrki，1988；Heppner，1997； で ．大 阪府立大学昆虫学研究室 に保管 さ れ る ． Parenti，2000），そ して チ ャ オ ビ メ ス ハ リオ ガ は韓国 で は コ ’ ウ ラ イ シ ダ レ ヤ ナ ギSalix pseudolasiog．yne H ． L6v．を（Sohn 分布．囗本 （本州 中部地方），韓国．日本新記録． and Adamski ，2008）摂食 して い る ．本属 の 種 で は本科の 他 寄 主 植 物． ヤ ナ ギ 科 ： コ ウ ラ イ シ ダ レ ヤ ナ ギSalix の 属 と同様 に ，幼虫は葉を巻 くこ となく外部か ら摂食し， pseudolasiogyne H ． L6v．（韓 国 の 記録） 繭 は 粗 い 目 の 籠状 で 羽化時 に は 蛹 は 繭か ら突出す る （Kyrki， 1988；Heppner，1997；Adamski et at ．，2009）， 生態．本種成虫 は夜 間灯火 に 誘 引 され る ．中部地方 で は 7 月下 旬か ら9月上旬 に 採集 され て い る ．韓国で は コ ウ ラ イ 引用文献 シ ダ レ ヤ ナ ギ の 葉を巻 くこ とな く外部か ら摂食す る （Sobn and Adamski，2008）． Adamski， D ，， K ． Boege， J． E Landry and J． C ． Sohn，2009． Two new species of Wockia Heinemanl1（Lepidoptera： Urodidae）from メ ス ハ リオ ガ科 と Wockiare − coastal dry forests ill western Mexico ． Proc． Entomol． Soc． Wash ，111 − メ ス ハ リオ ガ 科の 種はか つ て ス ガ科Yponomeutidaeとし て ； 166 182． Cho ． A ． Zwick ． ． Regier Mitter Yao 扱 わ れ て い たが，ス ガ 上科Yponomeutoideaの 有望 な固有新 ，S ， ．J C ，C， ，M ．R Cummings，J． ， Z ．Du． H ． ZhaQ A ， Y， Kawahara S， Weller D ． R ， Davis J， 形質 で あ る ♂成虫 の 腹部第8節 の 側板葉を欠 き，ス ガ 上科 ， ， ， ， Baixeras，」． W ． Brown and C ． Parr，2011，Can deliberately に 見 られな い い くつ か の 特徴 （♀の 翅刺 は 3本（Fig．4），腹 incQmplcte gene sample augmentation improve a phylogeny 部 第2節 腹 板 は ハ マ キ ガ 型 （Fig，5），幼虫 の 鉤 爪 は 縦 帯状， 一 estimate for the advanced moths and buしヒerflies （Hexapoda ： 幼虫 腹部 の Ll と L2刺 毛 は同 の 刺 毛基板 上 に あ る ，蛹 は − LepidoP 匸era ）？Syst． Bioi・60：782 796． 背方に刺列 をもつ ） を有す る ため に ，ス ガ上科とは独 立 し Dugdale， J． S．， N P Kristensen， G ． S． Robinson and M ． J． Scoble， た科 とされ た（Kyrki，1984，1988）．後 に ，本科 1科 で メ ス ハ − 1998．The smaller Microlepidop重era grade superfamilies ． In リオ ガ 上科Urodoideaと扱 われ た（Nielsen，1989）．本上科 は KristensenN ．P ． 1： 一 ， （ed ）， Lep玉doptera， moths alld butterflies 般 的 に は被蛹 類Obtectmeraを 二 Apeditrysiaの 除く新 門類 Evolution systematics and biogeography． Handbook of 一 ， ， と し て は わ れ て い る が は で る − 員 扱 ，類縁関係 不明 瞭 あ Zoology！Handbuch der Zoologte 4 （35）： 217 232． Walter de et at et al （Dugdale ．，1998；Kristensen ．，2007），ちなみ に ス ガ Gruyter， Berlin＆ New York ． 上 科 は 二 門 類Ditrysiaで あ る が 新二 門類 で は な い ．最近 の Heppner， J， B ．，1997． Wockia aspe ゆ un ごtella in North America ー っ い ： ： Holarct． Lepid ．4： − 分子 デ タ を使 た 系統解析 に お て も T 本科 の 類縁性 に 〔LepidopteruUrodidaeGalacticinae）． 73 関 し て は 定 ま っ た 説 は な い （Sohn，2014）． Mutanen et al ， 74． （2010）は新 二 門類 の ホ ソ マ イ コ ガ 上科Schredkensteiniidaeと Heppner， J． B ．，2008． The genus Wockia and a new species from southern Brazil Lepidoptera ： Urodidae ： Galacticinae ． Lepid． 近縁 としたが，Cho et al ．（2011）は ，新二 門類 の ニ ジ ュ ウ シ （ ） Nov ．1： li6−ll8， ト リバ ガ科Alucitidae，ハ マ キ モ ド キ ガ 科Choreutidae，トリ Heppner， J． B ．，2010． Notes on Vietnam rnoths ，10． Notes on Wockia バ ガ 科Pterophoridaeある い は ハ マ キ ガ 科Tortricidaeに近 い と 一 in Vietnam（Lepidoptera；Urodidae；Galacticinae）． Lepid． No 肌 し て い る． 方，Regier 8 ’α 奴 2013）は 新二 門類 の 姉妹群 と 3； 165−168． し て い る， Kristensen N ， P ， M ． J． Scoble and O ， KarshQlt，2007， Lepidoptera ー Wockiavaは束 ア ジ ア ，東南 ア ジア ，南北 ア メ リ カ お よび ヨ phylogeny and systematics ： the state of inventorying mDth and ロ ッ パ （イギ リス を除 く） に 分布し，10種 が記載され て い butterny diversity，1n Zhang， Z ， Q， and W ． A ． Shear （eds ）， る （Sohn，2014）．本属 の 前翅 は披 針 形状 灰 色 ある い は褐 Linnaeus tercentenary ； progress in invertebrate taxQnomy ． Zootaxa 1668 − 灰 色 で ，中央 部 に黒 褐色 の 隆起 鱗片 か らな る 横帯 を もち ；699 747， Kyrki，1，，1984， The Yponomeutoidea ： Areassessment of the （Kyrki，1986），外 部表徴 は お互 い に よ く似 て い る （Kyrki ， superfamily and its suprageneric groups （Lepidoptera）， 1986；Heppner，1997，2008，2010；Landry，1998；SQhn and Entomol． Scand、15：71 −84， Adamski，2008；Sohn and Park，2009；Parenti，2000；Adamski et Kyrki， J，，1986． Wockia わα 〃kpapane〃α sp ， n 、 from Borneo al ．，2009 ；Sohn，2013，2014）．♂交尾器 の バ ル バ は コ ス タ か （Lepidoptera， Yponometidae auct ．）， Ann ．Entomot ． Fenn ．52： らの 有刺毛突起 をもち，有刺毛 の ク ク ル ス が 発達 す る こ と 42−43． があ る ．♀交尾器 の 腹 部第8節以降 の 産卵器 は 非常 に 長 く Kyrki ， J．，1988 ， The systematic posi丘on of Wockia Heincmann ， 伸長す る ．本属の 種は外部表徴 で はBlastobasis属 （ネ マ ル 1870 ，and related genera 〔Lepidoptera： Dilrysia ： ハ キ バ ガ科）の 種 に類似す る が ，口 吻 は 鱗粉で 被わ れ ない − Yponomeutidae aucし）、1Vo∫α 吻 祕 11：45 69． 一 NII-ElectronicN 工 工 Electronlc LibraryLlbrary Service TheTheLepidopteiologicalSociety Lepidopterological Society of Japan H 7ts XUlockiaos 61 Landry, J, F., 1998.
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