Violaceae – Violet Family
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Towards an Understanding of the Evolution of Violaceae from an Anatomical and Morphological Perspective Saul Ernesto Hoyos University of Missouri-St
University of Missouri, St. Louis IRL @ UMSL Theses Graduate Works 8-7-2011 Towards an understanding of the evolution of Violaceae from an anatomical and morphological perspective Saul Ernesto Hoyos University of Missouri-St. Louis, [email protected] Follow this and additional works at: http://irl.umsl.edu/thesis Recommended Citation Hoyos, Saul Ernesto, "Towards an understanding of the evolution of Violaceae from an anatomical and morphological perspective" (2011). Theses. 50. http://irl.umsl.edu/thesis/50 This Thesis is brought to you for free and open access by the Graduate Works at IRL @ UMSL. It has been accepted for inclusion in Theses by an authorized administrator of IRL @ UMSL. For more information, please contact [email protected]. Saul E. Hoyos Gomez MSc. Ecology, Evolution and Systematics, University of Missouri-Saint Louis, 2011 Thesis Submitted to The Graduate School at the University of Missouri – St. Louis in partial fulfillment of the requirements for the degree Master of Science July 2011 Advisory Committee Peter Stevens, Ph.D. Chairperson Peter Jorgensen, Ph.D. Richard Keating, Ph.D. TOWARDS AN UNDERSTANDING OF THE BASAL EVOLUTION OF VIOLACEAE FROM AN ANATOMICAL AND MORPHOLOGICAL PERSPECTIVE Saul Hoyos Introduction The violet family, Violaceae, are predominantly tropical and contains 23 genera and upwards of 900 species (Feng 2005, Tukuoka 2008, Wahlert and Ballard 2010 in press). The family is monophyletic (Feng 2005, Tukuoka 2008, Wahlert & Ballard 2010 in press), even though phylogenetic relationships within Violaceae are still unclear (Feng 2005, Tukuoka 2008). The family embrace a great diversity of vegetative and floral morphologies. Members are herbs, lianas or trees, with flowers ranging from strongly spurred to unspurred. -
Evidence for Flower Mediated Assembly in Spring Ephemeral Understory Communities
Evidence for Flower Mediated Assembly in Spring Ephemeral Understory Communities by Stefan Weber A Thesis Presented to The University of Guelph In partial fulfilment of requirements for the degree of Master of Science. in Integrative Biology Guelph, Ontario, Canada © Stefan Weber, August, 2011 iv ABSTRACT EVIDENCE FOR POLLINATOR MEDIATED ASSEMBLY IN SPRING EPHEMERAL UNDERSTORY COMMUNITIES Stefan Weber Advisor: University of Guelph, 2011 Professor C. Caruso Plants with similar traits compete for resources. If related taxa share similar traits, phylogenetic relationships may predict competitive outcomes. Although plants compete for pollinators, flowers are rarely considered in community-assembly theory. I tested the hypothesis that plant communities are structured by competition for pollination. I inventoried communities at three spatial scales, measured seven flower traits, and tested the observed patterns against those generated by a null model to judge if community members were more or less similar in floral traits than expected by chance. I also measured the phylogenetic relatedness of community members to gauge trait-conservatism. Clustering of visually attractive traits suggests they promote facilitation of pollinators while over-dispersion of morphological traits suggests they partition pollinators in to avoid competition. Communities were phylogenetically even, but relatedness did not explain floral trait patterns. I suggest that flowers represent an ecological niche through which species can be sorted. iii AKNOWLEDGEMENTS This project was funded in part by a scholarship from the rare Charitable Organization. For assistance with the collection of historical inventories and specimens I would like to thank the Ontario Ministry of Natural Resources, The Royal Ontario Museum, The Royal Botanical Gardens, the Thames River Conservation Authority, the Michigan Department of Natural Resources, Oakland County Parks and Recreation, Michigan Metroparks, and herbaria at the University of Western Ontario, the University of Guelph, and Michigan State University. -
Outline of Angiosperm Phylogeny
Outline of angiosperm phylogeny: orders, families, and representative genera with emphasis on Oregon native plants Priscilla Spears December 2013 The following listing gives an introduction to the phylogenetic classification of the flowering plants that has emerged in recent decades, and which is based on nucleic acid sequences as well as morphological and developmental data. This listing emphasizes temperate families of the Northern Hemisphere and is meant as an overview with examples of Oregon native plants. It includes many exotic genera that are grown in Oregon as ornamentals plus other plants of interest worldwide. The genera that are Oregon natives are printed in a blue font. Genera that are exotics are shown in black, however genera in blue may also contain non-native species. Names separated by a slash are alternatives or else the nomenclature is in flux. When several genera have the same common name, the names are separated by commas. The order of the family names is from the linear listing of families in the APG III report. For further information, see the references on the last page. Basal Angiosperms (ANITA grade) Amborellales Amborellaceae, sole family, the earliest branch of flowering plants, a shrub native to New Caledonia – Amborella Nymphaeales Hydatellaceae – aquatics from Australasia, previously classified as a grass Cabombaceae (water shield – Brasenia, fanwort – Cabomba) Nymphaeaceae (water lilies – Nymphaea; pond lilies – Nuphar) Austrobaileyales Schisandraceae (wild sarsaparilla, star vine – Schisandra; Japanese -
Hydrastis Canadensis L.) in Pennsylvania: Explaining and Predicting Species Distribution in a Northern Edge of Range State
bioRxiv preprint doi: https://doi.org/10.1101/694802; this version posted July 8, 2019. The copyright holder for this preprint (which was not certified by peer review) is the author/funder. All rights reserved. No reuse allowed without permission. Title: Associated habitat and suitability modeling of goldenseal (Hydrastis canadensis L.) in Pennsylvania: explaining and predicting species distribution in a northern edge of range state. *1Grady H. Zuiderveen, 1Xin Chen, 1,2Eric P. Burkhart, 1,3Douglas A. Miller 1Department of Ecosystem Science and Management, Pennsylvania State University, University Park, PA 16802 2Shavers Creek Environmental Center, 3400 Discovery Rd, Petersburg, PA 16669 3Department of Geography, Pennsylvania State University, University Park, PA 16802 *telephone: (616) 822-8685; email: [email protected] bioRxiv preprint doi: https://doi.org/10.1101/694802; this version posted July 8, 2019. The copyright holder for this preprint (which was not certified by peer review) is the author/funder. All rights reserved. No reuse allowed without permission. Abstract Goldenseal (Hydrastis canadensis L.) is a well-known perennial herb indigenous to forested areas in eastern North America. Owing to conservation concerns including wild harvesting for medicinal markets, habitat loss and degradation, and an overall patchy and often inexplicable absence in many regions, there is a need to better understand habitat factors that help determine the presence and distribution of goldenseal populations. In this study, flora and edaphic factors associated with goldenseal populations throughout Pennsylvania—a state near the northern edge of its range—were documented and analyzed to identify habitat indicators and provide possible in situ stewardship and farming (especially forest-based farming) guidance. -
Evolutionary History of Floral Key Innovations in Angiosperms Elisabeth Reyes
Evolutionary history of floral key innovations in angiosperms Elisabeth Reyes To cite this version: Elisabeth Reyes. Evolutionary history of floral key innovations in angiosperms. Botanics. Université Paris Saclay (COmUE), 2016. English. NNT : 2016SACLS489. tel-01443353 HAL Id: tel-01443353 https://tel.archives-ouvertes.fr/tel-01443353 Submitted on 23 Jan 2017 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. NNT : 2016SACLS489 THESE DE DOCTORAT DE L’UNIVERSITE PARIS-SACLAY, préparée à l’Université Paris-Sud ÉCOLE DOCTORALE N° 567 Sciences du Végétal : du Gène à l’Ecosystème Spécialité de Doctorat : Biologie Par Mme Elisabeth Reyes Evolutionary history of floral key innovations in angiosperms Thèse présentée et soutenue à Orsay, le 13 décembre 2016 : Composition du Jury : M. Ronse de Craene, Louis Directeur de recherche aux Jardins Rapporteur Botaniques Royaux d’Édimbourg M. Forest, Félix Directeur de recherche aux Jardins Rapporteur Botaniques Royaux de Kew Mme. Damerval, Catherine Directrice de recherche au Moulon Président du jury M. Lowry, Porter Curateur en chef aux Jardins Examinateur Botaniques du Missouri M. Haevermans, Thomas Maître de conférences au MNHN Examinateur Mme. Nadot, Sophie Professeur à l’Université Paris-Sud Directeur de thèse M. -
Species List For: Valley View Glades NA 418 Species
Species List for: Valley View Glades NA 418 Species Jefferson County Date Participants Location NA List NA Nomination and subsequent visits Jefferson County Glade Complex NA List from Gass, Wallace, Priddy, Chmielniak, T. Smith, Ladd & Glore, Bogler, MPF Hikes 9/24/80, 10/2/80, 7/10/85, 8/8/86, 6/2/87, 1986, and 5/92 WGNSS Lists Webster Groves Nature Study Society Fieldtrip Jefferson County Glade Complex Participants WGNSS Vascular Plant List maintained by Steve Turner Species Name (Synonym) Common Name Family COFC COFW Acalypha virginica Virginia copperleaf Euphorbiaceae 2 3 Acer rubrum var. undetermined red maple Sapindaceae 5 0 Acer saccharinum silver maple Sapindaceae 2 -3 Acer saccharum var. undetermined sugar maple Sapindaceae 5 3 Achillea millefolium yarrow Asteraceae/Anthemideae 1 3 Aesculus glabra var. undetermined Ohio buckeye Sapindaceae 5 -1 Agalinis skinneriana (Gerardia) midwestern gerardia Orobanchaceae 7 5 Agalinis tenuifolia (Gerardia, A. tenuifolia var. common gerardia Orobanchaceae 4 -3 macrophylla) Ageratina altissima var. altissima (Eupatorium rugosum) white snakeroot Asteraceae/Eupatorieae 2 3 Agrimonia pubescens downy agrimony Rosaceae 4 5 Agrimonia rostellata woodland agrimony Rosaceae 4 3 Allium canadense var. mobilense wild garlic Liliaceae 7 5 Allium canadense var. undetermined wild garlic Liliaceae 2 3 Allium cernuum wild onion Liliaceae 8 5 Allium stellatum wild onion Liliaceae 6 5 * Allium vineale field garlic Liliaceae 0 3 Ambrosia artemisiifolia common ragweed Asteraceae/Heliantheae 0 3 Ambrosia bidentata lanceleaf ragweed Asteraceae/Heliantheae 0 4 Ambrosia trifida giant ragweed Asteraceae/Heliantheae 0 -1 Amelanchier arborea var. arborea downy serviceberry Rosaceae 6 3 Amorpha canescens lead plant Fabaceae/Faboideae 8 5 Amphicarpaea bracteata hog peanut Fabaceae/Faboideae 4 0 Andropogon gerardii var. -
Missouriensis Volume 28 / 29
Missouriensis Volume 28/29 (2008) In this issue: Improved Status of Auriculate False Foxglove (Agalinis auriculata) in Missouri in 2007 Tim E. Smith, Tom Nagel, and Bruce Schuette ......................... 1 Current Status of Yellow False Mallow (Malvastrum hispidum) in Missouri Tim E. Smith.................................................................................... 5 Heliotropium europaeum (Heliotropiaceae) New to Missouri Jay A. Raveill and George Yatskievych ..................................... 10 Melica mutica (Poaceae) New for the Flora of Missouri Alan E. Brant ................................................................................. 18 Schoenoplectus californicus (Cyperaceae) New to Missouri Timothy E. Vogt and Paul M. McKenzie ................................. 22 Flora of Galloway Creek Nature Park, Howell County, Missouri Bill Summers .................................................................................. 27 Journal of the Missouri Native Plant Society Missouriensis, Volume 28/29 2008 1 IMPROVED STATUS OF AURICULATE FALSE FOXGLOVE (AGALINIS AURICULATA) IN MISSOURI IN 2007 Tim E. Smith Missouri Department of Conservation P.O. Box 180, Jefferson City, MO 65102-0180 Tom Nagel Missouri Department of Conservation 701 James McCarthy Drive St. Joseph, MO 64507-2194 Bruce Schuette Missouri Department of Natural Resources Cuivre River State Park 678 State Rt. 147 Troy, MO 63379 Populations of annual plant species are known to have periodic “boom” and “bust” years as well as years when plant numbers more closely approach long-term averages. In tracking populations of plant species of conservation concern (Missouri Natural Heritage Program, 2007), there are sometimes also boom years in the number of reports of new populations. Because of reports of five new populations and a surge in numbers of plants at some previously-known sites, 2007 provided encouraging news for the conservation of the auriculate false foxglove [Agalinis auriculata (Michx.) Blake] in Missouri. -
BWSR Featured Plant: Downy Yellow Violet
2019 June Plant of the Month BWSR Featured Plant Name: Downy yellow violet (Viola pubescens) Plant family: Violet (Violaceae) Downy yellow Downy yellow violet, AKA hairy yellow violets are an important early violet or smooth yellow violet, is food source for a short (4- to 12-inch-tall), native, pollinators. Fine hairs along the herbaceous perennial that blooms in rounded teeth woodlands, gardens edge of the leaf are a distinguishing and shady areas Plant Stats feature. Brown lines starting in April. It STATEWIDE on the flower petals provides an early lead pollinators to WETLAND nectar and pollen. splash of color and INDICATOR Photo Credits: important early STATUS: FACU Heather Holm season nectar and PRIMARY USES: pollen. Like other Ground cover, shade/pollinator Viola species, this gardens, edibles, plant produces both woodland showy, open cross- restorations pollinating flowers at the top of the plant, and fully closed, self-pollinating flowers that may be found aboveground or underground. The showy flowers bloom before trees leaf out. The closed flowers bloom once the tree canopy leafs out. Planting Recommendations Range Downy yellow violets and can be used as Downy yellow violet is found may not be as an alternative to turf throughout Minnesota. Records aggressive as some grass, along paths and exist in all but a handful of other violets in a woodland borders, counties. It is mostly found east garden, but will spread and can be mixed with of the Missouri River, with a few over time in ideal other short woodland records west of the Missouri. conditions — part plants such as sedges, Its range stretches into New shade and medium to anemones and wild England and north into central dry soils. -
Flora of New Jersey Project Approved Nomenclature
FLORA OF NEW JERSEY PROJECT APPROVED NOMENCLATURE FNJP Authority Name: William Olson Date: March 14, 2010 Violaceae – Violet Family New Jersey Violet Family (synonyms as indented) Hybanthus Jacq. Hybanthus concolor (T.F. Forst.) Spreng. Eastern Green-Violet Syn. – Cubelium concolor (T.F. Forst.) Raf. Syn. – Viola concolor T.F. Forst. Viola L. Viola affinis Le Conte - Sand Violet, Le Conte’s Violet Syn. – Viola chalcosperma Brainerd Syn. – Viola rosacea Brainerd Syn. – Viola sororia ssp. affinis (Le Conte) R.J. Little Syn. – Viola sororia var. affinis (Le Conte) McKinney Viola arvensis Murr. - European Field Pansy EXOTIC Syn. – Viola tricolor var. arvensis (Murr.) Boiss. Viola bicolor Pursh - Field Pansy EXOTIC Syn. – Viola kitaibeliana var. rafinesquei Fern. Syn. – Viola kitaibeliana auct. non J.A. Schultes Syn. – Viola rafinesquei Greene Viola blanda Willd. - Sweet White Violet var. blanda var. palustriformis Gray RARE Syn. – Viola incognita Brainerd Syn. – Viola incognita var. forbesii Brainerd THE FLORA OF NEW JERSEY PROJECT IS A VOLUNTEER EFFORT AIMED AT THE PRODUCTION OF A MANUAL TO THE VASCULAR FLORA OF NEW JERSEY. Viola brittoniana Pollard - Northern Coastal Violet RARE var. brittoniana RARE Syn. – Viola pedatifida ssp. brittoniana (Pollard) McKinney var. pectinata (Bickn.) Alexander RARE Syn. – Viola pectinata Bickn. Viola canadensis L. - Canadian White Violet, Canada Violet var. canadensis RARE Syn. – Viola canadensis var. corymbosa Nutt. ex Torr. & Gray Viola cucullata Ait. - Marsh Blue Violet, Marsh Violet Syn. – Viola cucullata var. microtitis Brainerd Syn. – Viola obliqua Hill Viola hirsutula Brainerd - Southern Woodland Violet RARE Viola labradorica Schrank - Alpine Violet, American Dog Violet Syn. – Viola adunca var. minor (Hook.) Fern. Syn. – Viola conspersa Reinchenb. -
Dispersal Effects on Species Distribution and Diversity Across Multiple Scales in the Southern Appalachian Mixed Mesophytic Flora
DISPERSAL EFFECTS ON SPECIES DISTRIBUTION AND DIVERSITY ACROSS MULTIPLE SCALES IN THE SOUTHERN APPALACHIAN MIXED MESOPHYTIC FLORA Samantha M. Tessel A dissertation submitted to the faculty at the University of North Carolina at Chapel Hill in partial fulfillment of the requirements for the degree of Doctor of Philosophy in Ecology in the Curriculum for the Environment and Ecology. Chapel Hill 2017 Approved by: Peter S. White Robert K. Peet Alan S. Weakley Allen H. Hurlbert Dean L. Urban ©2017 Samantha M. Tessel ALL RIGHTS RESERVED ii ABSTRACT Samantha M. Tessel: Dispersal effects on species distribution and diversity across multiple scales in the southern Appalachian mixed mesophytic flora (Under the direction of Peter S. White) Seed and spore dispersal play important roles in the spatial distribution of plant species and communities. Though dispersal processes are often thought to be more important at larger spatial scales, the distribution patterns of species and plant communities even at small scales can be determined, at least in part, by dispersal. I studied the influence of dispersal in southern Appalachian mixed mesophytic forests by categorizing species by dispersal morphology and by using spatial pattern and habitat connectivity as predictors of species distribution and community composition. All vascular plant species were recorded at three nested sample scales (10000, 1000, and 100 m2), on plots with varying levels of habitat connectivity across the Great Smoky Mountains National Park. Models predicting species distributions generally had higher predictive power when incorporating spatial pattern and connectivity, particularly at small scales. Despite wide variation in performance, models of locally dispersing species (species without adaptations to dispersal by wind or vertebrates) were most frequently improved by the addition of spatial predictors. -
Sand Violet & Endangered Species Viola Adunca Sm
Natural Heritage Sand Violet & Endangered Species Viola adunca Sm. Program www.mass.gov/nhesp State Status: Special Concern Federal Status: None Massachusetts Division of Fisheries & Wildlife GENERAL DESCRIPTION: Sand Violet (Viola adunca; family Violaceae) is a low-growing perennial herb with dense rosettes of egg-shaped leaves and showy purple-violet flowers borne on leafy stems. It grows in disturbed habitats, usually in full sun, on moist to very dry soils. It competes poorly with other plants and relies on periodic disturbance to remove potential competitors. Violets produce long-lived soil seed banks (ca.100 years in species that have been assessed). AIDS TO IDENTIFICATION: Sand Violet produces showy purple-violet flowers about 2 cm (~0.75 in.) long at the tops of leafy stems in the early spring. The lateral petals are bearded (i.e., they have tufts of white hairs on their inner faces). The spur (a long, horn-like extension from the base of the flower) is short and blunt. The leaves and stems emerge from the crown of a buried stem and form a low-growing rosette. The leaves are Left: Close-up of flower. Middle inset: View of style showing egg-shaped to nearly round in outline, mostly 1 to 2.5 characteristic low, cylindrical bumps at the head. Right: Habit of cm (0.4–1 in.) wide and are barely rounded to very flowering and fruiting plant. Illustration by Elizabeth Farnsworth. slightly heart-shaped at the base. Leaves and flower stalks are covered in short hairs. The stipules (bracts SIMILAR SPECIES: Sand Violet is one of only three near the leaf bases) have comb-like appendages along New England “stemmed violets” (a group of species their upper edges. -
SPRING WILDFLOWERS of OHIO Field Guide DIVISION of WILDLIFE 2 INTRODUCTION This Booklet Is Produced by the ODNR Division of Wildlife As a Free Publication
SPRING WILDFLOWERS OF OHIO field guide DIVISION OF WILDLIFE 2 INTRODUCTION This booklet is produced by the ODNR Division of Wildlife as a free publication. This booklet is not for resale. Any By Jim McCormac unauthorized reproduction is prohibited. All images within this booklet are copyrighted by the Division of Wild- life and it’s contributing artists and photographers. For additional information, please call 1-800-WILDLIFE. The Ohio Department of Natural Resources (ODNR) has a long history of promoting wildflower conservation and appreciation. ODNR’s landholdings include 21 state forests, 136 state nature preserves, 74 state parks, and 117 wildlife HOW TO USE THIS GUIDE areas. Collectively, these sites total nearly 600,000 acres Bloom Calendar Scientific Name (Scientific Name Pronunciation) Scientific Name and harbor some of the richest wildflower communities in MID MAR - MID APR Definition BLOOM: FEB MAR APR MAY JUN Ohio. In August of 1990, ODNR Division of Natural Areas and Sanguinaria canadensis (San-gwin-ar-ee-ah • can-ah-den-sis) Sanguinaria = blood, or bleeding • canadensis = of Canada Preserves (DNAP), published a wonderful publication entitled Common Name Bloodroot Ohio Wildflowers, with the tagline “Let Them Live in Your Eye Family Name POPPY FAMILY (Papaveraceae). 2 native Ohio species. DESCRIPTION: .CTIGUJQY[ƃQYGTYKVJPWOGTQWUYJKVGRGVCNU Not Die in Your Hand.” This booklet was authored by the GRJGOGTCNRGVCNUQHVGPHCNNKPIYKVJKPCFC[5KPINGNGCHGPYTCRU UVGOCVƃQYGTKPIVKOGGXGPVWCNN[GZRCPFUKPVQCNCTIGTQWPFGFNGCH YKVJNQDGFOCTIKPUCPFFGGRDCUCNUKPWU