Montellano Ballesteros, M., Y J. Arroyo-Cabrales. 2002. Avances En Los

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Montellano Ballesteros, M., Y J. Arroyo-Cabrales. 2002. Avances En Los Á¿'ttilces en las estudios f t n le rxt t a sl oz t>ológic o s e n II é **icct iCt TCCICNCi[,NTIFICA Auancesen losestudios paleomastozoológicos en México Nlalisoli'lontellano Ballesteros Joaquín Arrovo Cabrales Cr¡ctrdinctdores SERIEARQUEOLOCIA INSTITUTONACIONAL DE ANTRoPoT-ocinE HtsroRtA Indice Fresentación Panoramade los marníferosrresczoiccs en México lv4ar i s a ! lvlo nt e I ! an o Bailesferos 11 Ei registrc paleogenicode mamífercs*;le México y su signiticaeiéngeologica - pale*rrtológica Is r¡tae i Fe r r,¡ s e uí a -\,/ i| | af ra n c a, Ed t arcj c,, J : ¡r,é n ez Ltid a I g o, '!trg* A. Añiz't\lcndieiay Víctori4anuei Bre'¡J Cueyas. 25 lii legrstr* rr¡asiu"r!aunisticon:iocérric* ue Lñéxico y sil* rmplie:aci*nesqeolcgico - paieentológicas Ft-i :t a, i,: -ii m e n e:z l'I i 11 ai gc. lsnaei Ft. :t t; sq tt ía-V rI I af ran c a y ,,'i:..:!:,ti,4anuei *r¿v;.: C{,,g,,rár. 47 inmigrantessi¡clamericanos en i** fr*nas de¡Terc¡ario tardío del ce¡rtrode Méxicc üscar CarranzüCa*sfa¡eda ¡ Wadt {. lvitiier ..... ..... 69 irnportanceof Mexice s iate iertiervmer;rmalien faunas vtade F Milier y Oscar Carrar,zaCe.sla¡roc¡-a Bs L-amastofal¡na del cuaternar¡atardía de fuléxieo Jaaqu:nArrolic. Cabrales, Csr:a, _i. !1t:lact:'.; Erieen Johnson.. .. i03 Evoluticnaryhistont of the fossitnlarlne mammals of Mexico LarryBarnes. ... ... ..125 Fossilxenarthra of Mexico:A ;'eview H. GregoryMacDanald ..227 Presentación Hanpasado más de 30 añosdesde que lostrabajos sintéticos acerca del estadodel conocimientode la paleomastofaunamexicana por Álvarez (1965)y Silva-Bárcenas(1969) fueron publicados. Dichos trabajos fueron importantespara evaluar el desarrollode losestudios sobre los fósiles de mamÍferosen México,así como paraestablecer prioridades a las inves- tigacionessubsecuentes. Desde entonces han aparecidoalgunos estu- diosque resumenel estadodel conocimientopara algunas paleomasto- faunasya searegionales (p.e. Carranza- Miller y Castañeda,1984) o tem- porales(p.e., Montellano, 1991). Sin embargo, hasta ahora no se habían tratadode sintetizarlos conocimientosque se tienenpara todo el perio- do en que los mamíferoshan estadopresentes en México "Paleomammalogy Estevolumen se originaa partirdel simposio titulado in México", que se realizó en el marco del Vll Congreso Teriológico Internacionalefectuado en Acapulco,Guerrero, México en el mes de sep- tiembrede 1997.La mayorparte de los participantesen dicho simposio aportaronlos manuscritoscorrespondientes a sus presentaciones,por lo quese ha podidocubrir la temporalidad desde el Mesozoicoal Cenozoico tardíoo Cuaternario,asícomo a las mastofaunasde ambientesterrestres y acuáticos.Infortunadamente la revisiónhistórica del desarrollodel campoen México,de O. J. Polaco,no fue entregadadentro del periodo que se lesdio a los autorespor lo que no apareceaquí. Montellanoaporta una síntesisde lo poco que se conoceacerca de mamíferosmesozoicos en el país.Falta mucho por haceren estetema, desde realizarlas prospeccionespara localizaráreas con fósilesde esta edad.Ferrusquía-Villafranca y colaboradores abordan la revisiónde los mamíferosdel Paleógenode una manerageneral, el cual estárestringi- do al Eocenoy estásólo registrado en tresfaunas locales. El registrode mamíferosdel Miocenoes analizadopor Jiménezy colaboradores,es escasopero está presenteen diferenteslatitudes del país,siendo las mejorestudiadas las del surestemexicano, las cuales dan una ideade la evoluciónde lasfaunas en laslatitudes bajas de NorteAmérica y sientan las basespara entender la diversidadde mamíferosantes del intercam- bio faunístico.México tiene una posición biogeográficainteresante que pone en contactodos regiones:ia Neotropica!y !a lJeáfica, por to c¡ue el estudiode la faunaen el pasacoaporta inÍcrrnacrón básica para enien- der el evento de intercambiofa.".:irístico entre Amérir--acjei Nc¡te y der sur a finalesdel TerciariotarCio. Lcs irabaics de üarra¡:za-Castane<iay ltrlilier dan un panorama general tanto de las far¡ras J',31T¡,¡ciaric tar,Jio. enÍai;- zando el centro de México,asi ccmc ce ics ilfx ia.ir,.ii.iiJS:rr-iijarrter;cano:i incluidosen elias.Arroyo-Cabraie: ,t i;,lial-,n,á,JLai':':Í. ircpc',jianan u¡ pi.i rner anáiisisde la base de datcs üi n-ra;.'rfi'-s r:,li l;!dt,'ri'r,a¡cr'aidio Cc México,que reflelacuales son lcs csia,ics ntct;tri:sir-iiiiail-s cómu-i¡st¿ fcrmado ei registropieisiccen,e:: v ios:rés!l''i:ii¡-.i ririsi¡;i,- ?arilcs s;itsti za de ríianeramuy completa e i.rUii,liccie r¡r¡míÍe;,ls.n¡:ri-,cs iósiies, ei cual es ün grupo generairre¡iiE¡c :cniadi, el ;Lrel-líry N,ic-Donaldhace lo propic con referenciaI io: xenai^iras,cJrr.-rr-1i, ce li'igen sudar¡rericano que llegé dr-¡ranteei evenic r-l*Inteicam!:iü f;iln,si'co V qle snriquecic, ias faunas rnexicanas. Ei territanc nacional es gia;idD. i'ta.y;:.;¡¡¡¡¿i&iFjc.;i t':i..¡o exi:i,jr"li \ pi'ospectar;falta pcr cc-litocüt"irn ,iif nili: rtriitili;, rj,r ti;llúitir,¡s,f,€ ic;s mamíferOSqUe viriierOnei ei p¡is¿C" ia cj;ve::::i:Ji.j,aüit-I.-l iran CamC,irilC y se han diversrficado,disiril¡',c,óri ge,rgráfrca y ;e.iact{,rít¿;tlrc i;lios. elf¡e ,,'i1,.,Í,..i otros.Esperenros que csle lltri v,_i :*,:r,-i,,i, lu il-.i..¿i;di'r €sie ...,a carnpo de la paieontologíay, sinraCe reÍe.€ncri !r:rd i.ls i.;uí: s;: rj:Cr quen a eila. Queremosagradecer a l s occtc:e: ;üariaTe ¡esa A,!berciy ciayrcr.i Raypor la lecturadel manuscl'iio,sr-:s airnállos corRentarios riteicraron ¡a redaccióny presentaciónde las apcrtac;cires 10 Panoramade los mamíferosrnesozoicos en México Mari so I Monte I lan o Bal/esferos i-"lastahace relativarnentepcr:o tremoo,e! ccnocrmientode la diversidad y distr"ibuciórrde le''s,"namlferos c..ie cenvivreronccn los dinosaurios duranie el l'-4ese,zoiccert escaso '- :iscurc A.Íinales del sigio XIX y prin- c!pics dei X).. rrQLrirospaleonri:cgüs comr E.D Cope. OC. Mai'sh. HenryF. O-cbcrn George G Sir¡c:¡n y Richa:clCwen, realizarondescu- brii"n¡entosy estL-:dic:;ije ir';ateriaipi'inctpairnenTe en Estados Unidos y Europa: y nc pcderrcs dela: '-Jere¡cionar el trabajo de Florentino Ar-neghinc.cn Arfientii-iaa p?'!lcin!;-isde ;igrc ci.jvasimpoi'tantes apoÉa- cro'resdaban r;na ',aija ;cie¡C:' ia ¡ivers rja'l :Jeeste grupo duranle la "Fi'a de los Reptilei",Fl¡3':asla Q:,e s€ r;iilizarcndiferentes técnrcas de recolecta,como gl tar--izada,que s* errtezaron a fcrmarcoleccicnes cje mamiferosmesozcicr¡s V su €sir.ldi,rse sisiei¡iat,zó.En 1979,Liliegraven, 'Mesozoic n¡eiair-Jawcrcivskay Clemens Dui:lrcarcnei iibro Mar¡mals:The frst iwc ll'lrrriscÍ n-r::r¡Ílalrarhrstcry" ccn;iciera,ic como el conocirnien- io i¡as avanzarfcce i,:s '-.¿¡¡;Í3i'csr'¡esczciccs nasta ese momento En i! se c:n*"ideraei reUi:ir; rr--,sr:l:ei gru[.i rir el espacrc y tienrpo.así aorilo la clasif¡cació¡Ce i¡;sl'r-:¿:':tricrls V las.tieas o hipótesisqUe Se pi.o- ^^'.,"'* -.t^i^ ñ1;rT|l!)raq ¡ Clugl \rC il-* ll lil\rrif..¡¡ir¡t: J9::: ric. ¡; l:-rJi,rr ::l-¿l i I'J Uv- nrimillvnslll lll¡vv9, HL'lllLll Pi talgs CO,"nOSU reproduccion. El Mesozoicose considerabacorno ia edad de oscuridadde ios mamíferosdebido ai pob'ev c:rpeisLrregrsirc q-e setenla, pero a medl- cjaque ha pasadoel tiempc)1 que el registrofósil de mamíferosmeso- zcicosse ha incrementado.nos hemog percatado que Ia historia es oti'a. ''adquirieron" Estaépoca es sumamenteinnponante ya qijeen ellase las característicasCei ser mamífero,entendiendo c,or esto la presenciade lreshueseciilos en el oido interno.que icrmanla afiiculacióndentario- sscamosoentre cránec y mandÍbula(para c;{e rentes puntos de vrstade ia diagrrosisde Mammalia,ver Fov¡e.i988:S/ible '1991;Lucas, 1992). Losfósiles más antiguosconsiderados rnamiÍeros, han sidodescubier- tes en depósitosdel T¡'iásicoTardío i210 ma) y JurásicoTemprano (205 ma)de Europaocciclenial (Alemanla, Francia, e lnglaterra),Africa del Sur n [)epto. Paieontología.Insiliuto d. Geo]L)cia tji!4¡,1 1l '1994; y China(Sigogneau-Russell, Clemens,1986). Estos registros con- sistenen dientesreferidos a tres diferentesgrupos: harámidos, morga- nucodontesy kuehneotéridos.Descubrimientos recientes han provoca- do cambiosen la asignaciónde la edadde lasfaunas locales, y los nue- vos análisissistemáticos han modificadonuestra percepción de la evo- luciónde mamíferosy sinápsidosdesde el Triásicotardío al Jurásico temprano.Se observaque no sólo existióun linajede mamíferos,sino que hubouna variedad de linajescon característicasdentales diferentes, lo que significaque los mamíferospresentaban diferentes adaptaciones y que ocup,abanuna granvariedad de nichos.Un puntoque es impor- tantees que su distribuciónno estárestringida sólo a un continente,sino que es mundial.Lo que sí es comúnen el grupoes su tallapequeña, el mamíferomás grandefue del tamañode un perrofox-terrier. A finalesdel Mesozoico,en el periodoCretácico, se registrael origen de los dos grandesgrupos que actualmentevemos a nuestroalrededor: los marsupialeso metaterios,y los placentadoso euterios.Después de la extinciónde los dinosauriosy ya entrandoen la era Cenozoica,se observauna gran radiaciónadaptativa de los mamíferostanto marsupia- lescomo placentados.La gran explosión de formasde losmarsupiales se "islas dio en lo que se ha denominado continentes",tierras que quedaron aisladasdurante gran parte del Cenozoico,lo que permitióque estosani- malesdesarrollaran gran cantidadde adaptaciones,asi tenemosformas saltadoras(kanguros), arborícolas (koala), cavadores (wombats), planea- doras,insectivoras o carnívoras de grandestallas. Los placentados por su parte,también tuvieron una gran explosiónsobre todo en el nichode la herbivoría,pero también vemos formas que regresaronal
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