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(Mollusca, Gastropoda, Cypraeoidea) Description of a Species

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(Mollusca, Gastropoda, Cypraeoidea) Description of a Species Cainozoic Research, 8(1-2), pp. 29-34, December2011 Contributions to the knowledge of the Pediculariidae (Mollusca, Gastropoda, Cypraeoidea) 2. On the of the Eotrivia 1924 occurrence genus Schilder, in the Ukraine Eocene, with the description of a new species Dirk Fehse Nippeser Str. 3, D-12524 Berlin, Germany; e-mail: [email protected] Received 18 August 2010; revised version accepted 16 August 2011 known from West Eocene is confirmedfor the A The genus Eotrivia formerly European deposits now easternmost Paratethys. second, Eotrivia Eocene still unknown Eotrivia besides faracii is described from the of Ukraine as Eotrivia procera sp. nov. KEY WORDS:: Cypraeoidea, Eocypraeidae, Pediculariidae, Eotrivia, new species, new combinations, fossil, Eocene, Ukraine. Introduction clarified by geological studies. In the following the term ‘Mandrikovka beds, Eocene’ is therefore used. Recently some shells from Mandrikovka, Dnepropetrovsk region of Ukraine became available for study. Among In Part 1 (Fehse & Vician, 2008) of this small series of those shells several species of families Eocypraeidae papers on the Pediculariidae the identity of Projenneria Schilder, 1924 and PediculariidaeAdams & Adams, 1854 neumayri (Hilber, 1879) is discussed and Projenneria al- were found.These species formerly known from the Mid- bopunctata Fehse & Vician, 2008 is described. dle Eocene of Western Europe are unusually common at Mandrikovka. The species ofthe Eocypraeidae are Apiocy- Abbreviations - To denote the repositories of materialre- praea sellei (de Raincourt, 1874), Oxycypraea del- ferred to in the text, the following abbreviations are used: phinoides (Cossmann, 1886), Oxycypraea fourtaui (Op- penheim, 1906) comb, nov., Cyproglobina parvulorbis (de DFB collection Dirk Fehse, Berlin, Germany. Gregorio), 1880, Cyproglobina pisularia (de Gregorio, ZSM Zoological State Collection, Munich, Germany. and of the Pediculariidae 1880) are Projenneria pregnans (de Gregorio, 1880) and Eotrivia faracii (de Gregorio, L length ofthe shell including spire 1880). These species, described from the Lutetian and Bar- W width ofthe shell tonian of France and Italy, are very rare. Surprisingly, a H height ofthe shell second, hithertooverlooked Eotrivia is also found among ct number of columellar denticles the material. In the following this species, new to science, It number oflabral denticles is described as Eotriviaprocera sp. nov. A further pediculariid occurring at Mandrikovka might be The shell formula proposed by Schilder (1935, p. 327) is Projenneria ludoviciana (Johnson, 1899) formerly known used herein. This formula is derived from measurements from the Moodys Branch Formation (Late Eocene) of taken from all available fully mature and normally formed Louisiana, U.S.A. but this has to be clarified by further specimens. It consists ofthe following elements: [L (W-H) examination. LT:CT]. L: average length in mm, W: average width/ The age of the deposits at Mandrikovka is not yet fully length ratio in %, H: average height/ length ratio in %, LT: clarified. It is thought to be PriabonianLate Eocene (Ami- normalised numberoflabral teeth, CT: normalized number of trov, 2008). However, the occurrence so many Middle of columellar denticles. The normalised number of den- earlier - - Eocene species at Mandrikovka would suggest an ticles in relation to a shell of 25 mm length is calcu- such Bartonian Lutetian but this has be = age as or even to lated as follows; T 7 + [(c-7)*V (25/L)] T: normalized -30- numberof denticles, c: numberof denticles actually pres- Description — Shell large sized, solid, inflated and ovate. ent, L: length. Body whorl pyriform, inflatedand rounded, about80% of total height, with both terminals produced and separated from the dorsal elevation by a constriction. Terminal tips Systematic palaeontology posteriorly blunt, anteriorly protruded. Dorsum elevated with highest point at posterior third. Mid-dorsal sulcus Superfamily Cypraeoidea Troschel, 1863 obscured. Ventrum convex with outer margin rounded. PediculariidaeAdams 1854 widenedtowards fossular Family & Adams, Aperture fairly narrow, section, Subfamily JenneriinaeThiele, 1929 anteriorly straight, posteriorly curved. Labrum narrow, Genus EotriviaSchilder, 1924 widest at mid-portion, ventrally convex. Inner labral mar- gin rounded, curved, bears 19-24 fine denticles. Outer Type species - Trivia bouryi Cossmann, 1889 by mono- labralmargin roundly callused, shouldered.Denticles con- typy. Bartonian, France. tinued as folds onto labrum, becoming finer and more close-set onto dorsum and ventrum,terminateonto parietal lip as fine denticles. Parietallip slightly callused, bears 20- Eotriviaprocera sp. nov. 25 denticles, anterior denticles coarser, more prominent. Figures 2.1, 23-2.5, 3.1, 3.2 Siphonal and anal canals indented.Columellaslightly con- vex, relatively narrow and tapering steeply inwards. Inner adaxial carinalridge absent. Fossula less developed, short, Locus typicus - Mandrikovka, Dnepropetrovsk region, indentedjust above inner fossular margin, not clearly de- Ukraine. limited from the rest ofthe columella. Inner fossular mar- gin callused, slightly protruded, roundly edged. Juvenile - Mandrikovka Eocene. shell visible columella. Stratum typicum beds, grid usually on Holotype- Zoological State Collection, Munich, Germany, Shellformula - [14 (69-57) 27:27] No. 20070571 Range of variation - Shell inflation varies considerably. Distribution - Known only from the type locality so far. Dorsal sulcus sometimes indistinctbut mostly obsolete. Materialstudied- five from the Derivatio nominis: From the Latin Altogether specimens type adjective procerus, -a, locality. meaning beautiful. Dimensions- = = = Holotype: L 14.0 mm, W 9.7 mm, D 8.0 mm, ct 20, It 19 (ZSM, No. 20070571) = = = It Paratype 1: L 15.5 mm, W 10.5 mm, D 8.7 mm, ct 25, 24 (DFB, No. 8525-1) 2: L = 14.1 W = 9.6 D = 7.8 ct 20 It 22 No. Paratype mm, mm, mm, , (DFB, 8525-2) = W = D = ct It Paratype 3: L 14.0 mm, 10.0 mm, 8.5 mm, 22, 23 (DFB, No. 8525-3) = = 12.8 W = 8.7 D 7.2 It 23 No. Paratype 4: L mm, mm, mm, ct 23, (DFB, 8525-4) Discussion - The following species belong toto the genus Eotrivia Schilder, 1924 - Eotrivia baronensis (Dolin, Dolin & Le Renard, 1980)1980) comb. nov. Lutetian, M. Eocene Cotentin Basin - Eotrivia bouryi (Cossmann, 1889) Bartonian, M. Eocene Paris Basin - Eotriviafaracii (de Gregorio, 1880) Lutetian, M. Eocene Italy - EotriviapalumbellaEotrivia palumbella (de Gregorio, 1880) Lutetian, M. Eocene ItalyItaly - Eotrivia pedicularis (Deshayes in Deshayes & Milne Edwards, 1844) Lutetian, M. Eocene Paris Basin - Eotrivia recluzi (Cossmann, 1897) comb. nov. (=Cypraea (LuponiaLuponia ) æquipartitaoequipartita Cossmann, 1902) Lutetian, M. Eocene Contentin Basin Schilder & Schilder (1971, p. 71) assigned Trivia recluzi has a rounded, inflated and ribbed shell similar to E. to Cossmann, 1897 to the genus Cypropterina de Gregorio, bouryi. Therefore, T. recluzi is here assigned Eotrivia. - ceciliae de is that all known of Eo- 1880 type species: Cypropterina Gregorio, It noteworthy previously species 1880. Their decision is inexplicable because there is no trivia were restricted to the Middle Eocene of northern resemblance between C. ceciliae and T. recluzi. The first France and northern Italy. Therefore, the existence of at smooth and flattenedshell least two Eotrivia in the Ukraine Mandrikovka possesses a completely laterally species with only the central dorsum elevated. In contrast T. recluzi beds is unexpected. -31 - 1. Figure Appearance ofthejuvenile shell grid examined on columella:left sideEotriviaprocera nov. sp.; right sideEotriviafaracii (de Gregorio, 1880). It is far from the original geographical range ofthe genus, not be located in any Italian Museum. which is now extended considerably. But the age of the The new species differsfrom E. pedicularis by the differ- Mandrikovka beds is most probably Middleto Late Eocene ent shell morphology and especially the much finer shell - either Lutetian or Bartonian. ribbing. Similarly, E. bouryi has much coarser ribs and a Eotriviaprocera nov. sp. differs from its congenerE.fara- distinct dorsal sulcus. Eotrivia baronensis has again cii especially by the differentjuvenile shell grid (Figure 1). coarser, more distantribs and a sharply keeled outer labral However, both are also distinguishable by theiradultshell margin. Eotrivia recluzi has a completely differently The dorsal ribs the labral anal morphology: are coarser, outer shaped canal and deeply incised dorsal sulcus. Eo- shoulder and keeled and innerfossular more prominent the triviaprocera nov. sp. differs from E. palumbella immedi- less in show de margin projects E. faracii. It was hoped to ately by the much narrower labrum and the position ofthe ofEotrivia could Gregorio’s types but, unfortunately, they aperture. Figure 2. Eotrivia from the Eocene Mandrikovkabeds, Mandrikovka, Dnepropetrovsk region, Ukraine. 1. Eotrivia No. 20070571. procera nov. sp. Holotype, ZSM, 2. Eotrivia faracii (de Gregorio, 1880). DFB, No. 7677-1. L= 10.7 mm. 3. Eotrivia No. the shell dorsum. procera nov. sp. Holotype, ZSM, 20070571. Enlarged view on grid on 4. 4. No. 8525-4. Eotrivia procera nov. sp. Paratype DFB, 5. Eotrivia No. procera nov. sp. Paratype 3, DFB, 8525-3. -32- Figure 2 (explanation on foregoing page). -33 - 3 Figure (explanation on next page). -34- References d'Information des Geologues du Bassin de Paris, 17. 26-48. Fehse, D. & Vician, Z. (2008): On the identitiy of Projenneria Amitrov, O.V. 2008. The
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