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About the Book the Format Acknowledgments
About the Book For more than ten years I have been working on a book on bryophyte ecology and was joined by Heinjo During, who has been very helpful in critiquing multiple versions of the chapters. But as the book progressed, the field of bryophyte ecology progressed faster. No chapter ever seemed to stay finished, hence the decision to publish online. Furthermore, rather than being a textbook, it is evolving into an encyclopedia that would be at least three volumes. Having reached the age when I could retire whenever I wanted to, I no longer needed be so concerned with the publish or perish paradigm. In keeping with the sharing nature of bryologists, and the need to educate the non-bryologists about the nature and role of bryophytes in the ecosystem, it seemed my personal goals could best be accomplished by publishing online. This has several advantages for me. I can choose the format I want, I can include lots of color images, and I can post chapters or parts of chapters as I complete them and update later if I find it important. Throughout the book I have posed questions. I have even attempt to offer hypotheses for many of these. It is my hope that these questions and hypotheses will inspire students of all ages to attempt to answer these. Some are simple and could even be done by elementary school children. Others are suitable for undergraduate projects. And some will take lifelong work or a large team of researchers around the world. Have fun with them! The Format The decision to publish Bryophyte Ecology as an ebook occurred after I had a publisher, and I am sure I have not thought of all the complexities of publishing as I complete things, rather than in the order of the planned organization. -
ARTHROPODA Subphylum Hexapoda Protura, Springtails, Diplura, and Insects
NINE Phylum ARTHROPODA SUBPHYLUM HEXAPODA Protura, springtails, Diplura, and insects ROD P. MACFARLANE, PETER A. MADDISON, IAN G. ANDREW, JOCELYN A. BERRY, PETER M. JOHNS, ROBERT J. B. HOARE, MARIE-CLAUDE LARIVIÈRE, PENELOPE GREENSLADE, ROSA C. HENDERSON, COURTenaY N. SMITHERS, RicarDO L. PALMA, JOHN B. WARD, ROBERT L. C. PILGRIM, DaVID R. TOWNS, IAN McLELLAN, DAVID A. J. TEULON, TERRY R. HITCHINGS, VICTOR F. EASTOP, NICHOLAS A. MARTIN, MURRAY J. FLETCHER, MARLON A. W. STUFKENS, PAMELA J. DALE, Daniel BURCKHARDT, THOMAS R. BUCKLEY, STEVEN A. TREWICK defining feature of the Hexapoda, as the name suggests, is six legs. Also, the body comprises a head, thorax, and abdomen. The number A of abdominal segments varies, however; there are only six in the Collembola (springtails), 9–12 in the Protura, and 10 in the Diplura, whereas in all other hexapods there are strictly 11. Insects are now regarded as comprising only those hexapods with 11 abdominal segments. Whereas crustaceans are the dominant group of arthropods in the sea, hexapods prevail on land, in numbers and biomass. Altogether, the Hexapoda constitutes the most diverse group of animals – the estimated number of described species worldwide is just over 900,000, with the beetles (order Coleoptera) comprising more than a third of these. Today, the Hexapoda is considered to contain four classes – the Insecta, and the Protura, Collembola, and Diplura. The latter three classes were formerly allied with the insect orders Archaeognatha (jumping bristletails) and Thysanura (silverfish) as the insect subclass Apterygota (‘wingless’). The Apterygota is now regarded as an artificial assemblage (Bitsch & Bitsch 2000). -
Amphiesmeno- Ptera: the Caddisflies and Lepidoptera
CY501-C13[548-606].qxd 2/16/05 12:17 AM Page 548 quark11 27B:CY501:Chapters:Chapter-13: 13Amphiesmeno-Amphiesmenoptera: The ptera:Caddisflies The and Lepidoptera With very few exceptions the life histories of the orders Tri- from Old English traveling cadice men, who pinned bits of choptera (caddisflies)Caddisflies and Lepidoptera (moths and butter- cloth to their and coats to advertise their fabrics. A few species flies) are extremely different; the former have aquatic larvae, actually have terrestrial larvae, but even these are relegated to and the latter nearly always have terrestrial, plant-feeding wet leaf litter, so many defining features of the order concern caterpillars. Nonetheless, the close relationship of these two larval adaptations for an almost wholly aquatic lifestyle (Wig- orders hasLepidoptera essentially never been disputed and is supported gins, 1977, 1996). For example, larvae are apneustic (without by strong morphological (Kristensen, 1975, 1991), molecular spiracles) and respire through a thin, permeable cuticle, (Wheeler et al., 2001; Whiting, 2002), and paleontological evi- some of which have filamentous abdominal gills that are sim- dence. Synapomorphies linking these two orders include het- ple or intricately branched (Figure 13.3). Antennae and the erogametic females; a pair of glands on sternite V (found in tentorium of larvae are reduced, though functional signifi- Trichoptera and in basal moths); dense, long setae on the cance of these features is unknown. Larvae do not have pro- wing membrane (which are modified into scales in Lepi- legs on most abdominal segments, save for a pair of anal pro- doptera); forewing with the anal veins looping up to form a legs that have sclerotized hooks for anchoring the larva in its double “Y” configuration; larva with a fused hypopharynx case. -
Diversity and Ecosystem Services of Trichoptera
Review Diversity and Ecosystem Services of Trichoptera John C. Morse 1,*, Paul B. Frandsen 2,3, Wolfram Graf 4 and Jessica A. Thomas 5 1 Department of Plant & Environmental Sciences, Clemson University, E-143 Poole Agricultural Center, Clemson, SC 29634-0310, USA; [email protected] 2 Department of Plant & Wildlife Sciences, Brigham Young University, 701 E University Parkway Drive, Provo, UT 84602, USA; [email protected] 3 Data Science Lab, Smithsonian Institution, 600 Maryland Ave SW, Washington, D.C. 20024, USA 4 BOKU, Institute of Hydrobiology and Aquatic Ecology Management, University of Natural Resources and Life Sciences, Gregor Mendelstr. 33, A-1180 Vienna, Austria; [email protected] 5 Department of Biology, University of York, Wentworth Way, York Y010 5DD, UK; [email protected] * Correspondence: [email protected]; Tel.: +1-864-656-5049 Received: 2 February 2019; Accepted: 12 April 2019; Published: 1 May 2019 Abstract: The holometabolous insect order Trichoptera (caddisflies) includes more known species than all of the other primarily aquatic orders of insects combined. They are distributed unevenly; with the greatest number and density occurring in the Oriental Biogeographic Region and the smallest in the East Palearctic. Ecosystem services provided by Trichoptera are also very diverse and include their essential roles in food webs, in biological monitoring of water quality, as food for fish and other predators (many of which are of human concern), and as engineers that stabilize gravel bed sediment. They are especially important in capturing and using a wide variety of nutrients in many forms, transforming them for use by other organisms in freshwaters and surrounding riparian areas. -
Issue Information
Systematic Entomology (2017), 42, 240–266 DOI: 10.1111/syen.12209 Molecular phylogeny of Sericostomatoidea (Trichoptera) with the establishment of three new families KJELL ARNE JOHANSON1, TOBIAS MALM1 andMARIANNE ESPELAND2 1Department of Zoology, Swedish Museum of Natural History, Stockholm, Sweden and 2Arthropoda Department, Zoological Research Museum Alexander Koenig, Bonn, Germany Abstract. We inferred the phylogenetic relationships among 58 genera of Sericostom- atoidea, representing all previously accepted families as well as genera that were not placed in established families. The analyses were based on five fragments of the protein coding genes carbamoylphosphate synthetase (CPSase of CAD), isocitrate dehydroge- nase (IDH), Elongation factor 1a (EF-1a), RNA polymerase II (POL II) and cytochrome oxidase I (COI). The data set was analysed using Bayesian methods with a mixed model, raxml, and parsimony. The various methods generated slightly different results regarding relationships among families, but the shared results comprise support for: (i) a monophyletic Sericostomatoidea; (ii) a paraphyletic Parasericostoma due to inclusion of Myotrichia murina, leading to synonymization of Myotrichia with Parasericostoma; (iii) a polyphyletic Sericostomatidae, which is divided into two families, Sericostom- atidae sensu stricto and Parasericostomatidae fam.n.; (iv) a polyphyletic Helicophidae which is divided into Helicophidae sensu stricto and Heloccabucidae fam.n.; (v) hypoth- esized phylogenetic placement of the former incerta sedis genera Ngoya, Seselpsyche and Karomana; (vi) a paraphyletic Costora (Conoesucidae) that should be divided into several genera after more careful examination of morphological data; (vii) reinstatement of Gyrocarisa as a valid genus within Petrothrincidae. A third family, Ceylanopsychi- dae fam.n., is established based on morphological characters alone. A hypothesis of the relationship among 14 of the 15 families in the superfamily is presented. -
Functional Feeding Groups of Aquatic Insect Families in Latin America: a Critical Analysis and Review of Existing Literature
Functional feeding groups of aquatic insect families in Latin America: a critical analysis and review of existing literature Alonso Ramírez1 & Pablo E. Gutiérrez-Fonseca2 1. Department of Environmental Sciences, University of Puerto Rico, P.O. Box 190341, San Juan, Puerto Rico 00919; [email protected] 2. Department of Biology, University of Puerto Rico Rio Piedras, San Juan, Puerto Rico 00919; [email protected] Received 12-XII-2013. Corrected 20-I-2014. Accepted 13-II-2014. Abstract: Aquatic macroinvertebrates are involved in numerous processes within aquatic ecosystems. They often have important effects on ecosystem processes such as primary production (via grazing), detritus break- down, and nutrient mineralization and downstream spiraling. The functional feeding groups (FFG) classification was developed as a tool to facilitate the incorporation of macroinvertebrates in studies of aquatic ecosystems. This classification has the advantage of combining morphological characteristics (e.g., mouth part specializa- tion) and behavioral mechanisms (e.g., way of feeding) used by macroinvertebrates when consuming resources. Although recent efforts have greatly advanced our ability to identify aquatic macroinvertebrates, there is limited information on FFG assignment. Furthermore, there has been some variation in the use of the FFG classification, in part due to an emphasis on using gut content analysis to assign FFG, which is more appropriate for assigning trophic guilds. Thus, the main goals of this study are to (1) provide an overview of the value of using the FFG classification, (2) make an initial attempt to summarize available information on FFG for aquatic insects in Latin America, and (3) provide general guidelines on how to assign organisms to their FFGs. -
South, Tasmania
Biodiversity Summary for NRM Regions Guide to Users Background What is the summary for and where does it come from? This summary has been produced by the Department of Sustainability, Environment, Water, Population and Communities (SEWPC) for the Natural Resource Management Spatial Information System. It highlights important elements of the biodiversity of the region in two ways: • Listing species which may be significant for management because they are found only in the region, mainly in the region, or they have a conservation status such as endangered or vulnerable. • Comparing the region to other parts of Australia in terms of the composition and distribution of its species, to suggest components of its biodiversity which may be nationally significant. The summary was produced using the Australian Natural Natural Heritage Heritage Assessment Assessment Tool Tool (ANHAT), which analyses data from a range of plant and animal surveys and collections from across Australia to automatically generate a report for each NRM region. Data sources (Appendix 2) include national and state herbaria, museums, state governments, CSIRO, Birds Australia and a range of surveys conducted by or for DEWHA. Limitations • ANHAT currently contains information on the distribution of over 30,000 Australian taxa. This includes all mammals, birds, reptiles, frogs and fish, 137 families of vascular plants (over 15,000 species) and a range of invertebrate groups. The list of families covered in ANHAT is shown in Appendix 1. Groups notnot yet yet covered covered in inANHAT ANHAT are are not not included included in the in the summary. • The data used for this summary come from authoritative sources, but they are not perfect. -
Morse, JE 1993. a Checklist of the Trichoptera of North
751 NOMINA INSECTA NEARCTICA The Trichoptera list is taken from the very important Phryganeidae paper: Morse, J.E. 1993. A checklist of the Trichoptera (Phryganopsychidae not NA) of North America, including Greenland and Mexico. (Plectrotarsidae not NA) Transactions of the American Entomological Society, Polycentropodidae 119:47-93. The list was modified by the addition of Psychomyiidae generic synonomies, the reversion to original orthography Rhyacophilidae for the valid species group names, and the elimination of Sericostomatidae the purely Mexican species. (Stenopsychidae not NA) (Tasmiidae not NA) ALTERNATIVE FAMILY NAMES Uenoidae Xiphocentronidae The purpose of this section is to list some family names the user might encounter in the Trichoptera. This list is not an exhaustive compilation of family group names, but STATISTICS is included for the convenience of the users of the check list. The abbreviation (not NA) stands for not North The following statistics are offered without comment. America, ie. the family is not found north of the US- Mexican border. Family # Names # Valid Arctopsychidae 17 11 Beraeidae 3 3 Agrypniidae (see Phryganeidae) Brachycentridae 47 35 (Anomalopsychidae not NA) Calamoceratidae 8 5 (Antipodoeciidae not NA) Glossosomatidae 83 78 Goeridae 12 12 Arctopsychidae Helicopsychidae 12 9 (Atriplectididae not NA) Hydrobiosidae 3 3 (Barbarochthonidae not NA) Hydropsychidae 175 146 Beraeidae Hydroptilidae 268 245 Lepidostomatidae 89 68 Brachycentridae Leptoceridae 173 113 Calamoceratidae Limnephilidae 406 292 (Calocidae -
Trichoptera - Caddisflies Order Lepidoptera - Butterflies & Moths AMPHIESMENOPTERA - the “Garment Wings”
AMPHIESMENOPTERA - the “garment wings” The Major Groups of Holometabola Holometabola (=Endopterygota) Order Hymenoptera Superorder Neuropterida Order Raphidioptera Order Megaloptera Order Neuroptera Order Coleoptera Order Strepsiptera Superorder Mecopterida (= Panorpida) Antliophora Order Mecoptera - scorpionflies Order Siphonaptera - fleas Order Diptera - true flies Amphiesmenoptera Order Trichoptera - caddisflies Order Lepidoptera - butterflies & moths AMPHIESMENOPTERA - the “garment wings” SYNAPOMORPHIES • Adult prelabium fused with hypopharynx • Pronotum with paired setose “warts” • Wings with extensive covering of setae • Anal veins of the forewing apparently looping up into a double-Y configuration • Presence of paired glands opening on sternum V • Male genital segment IX with tergum and sternum fused, forming a closed ring Hypothetical amphiesmenopteran, from Kristensen (1984) TRICHOPTERA - caddisflies The Major Groups of Holometabola Holometabola (=Endopterygota) Order Hymenoptera Superorder Neuropterida Order Raphidioptera Order Megaloptera Order Neuroptera Order Coleoptera Order Strepsiptera Superorder Mecopterida (= Panorpida) Antliophora Order Mecoptera - scorpionflies Order Siphonaptera - fleas Order Diptera - true flies Amphiesmenoptera Order Trichoptera - caddisflies Order Lepidoptera - butterflies & moths Diversity of Recent hexapods as proportion of named species. (from Grimaldi & Engel 2005. Evolution of the Insects. Cambridge Univ. Press.) Trichoptera, a minor order! The 7th largest Hexapod order! Culicomorpha/Psychodomorpha -
Redalyc.Especies Del Orden Thichoptera (Insecta) En Colombia
Biota Colombiana ISSN: 0124-5376 [email protected] Instituto de Investigación de Recursos Biológicos "Alexander von Humboldt" Colombia Muñoz Quesada, Fernando Especies del Orden Thichoptera (Insecta) en Colombia Biota Colombiana, vol. 1, núm. 3, diciembre, 2000, pp. 267-288 Instituto de Investigación de Recursos Biológicos "Alexander von Humboldt" Bogotá, Colombia Disponible en: http://www.redalyc.org/articulo.oa?id=49110304 Cómo citar el artículo Número completo Sistema de Información Científica Más información del artículo Red de Revistas Científicas de América Latina, el Caribe, España y Portugal Página de la revista en redalyc.org Proyecto académico sin fines de lucro, desarrollado bajo la iniciativa de acceso abierto VarónBiota Colombiana 1 (3) 267 - 288, 2000 Eumastacids of Colombia - 267 Especies del Orden Trichoptera (Insecta) en Colombia Fernando Muñoz-Quesada Department of Entomology, University of Minnesota, 1980 Folwell Ave., St.Paul, MN 55108, U.S.A. [email protected]; [email protected] Palabras Clave: Trichoptera, Insectos Acuáticos, Listado de Especies, Colombia, Neotrópico Trichoptera es el orden de insectos cuyas alas, princi- Trichoptera poseen además de los tres pares de patas palmente las anteriores, están cubiertas por pelos (del grie- torácicas, un solo par de propatas anales con una uña en el go “trichos” = pelos, “pteron” = ala, “ala con pelos”), y último segmento abdominal, y no presentan espiráculos que presentan metamorfosis completa (holometábolos). Las abiertos (apneústicas). Las larvas tienen una glándula formas inmaduras (huevo, larva y pupa) son generalmente salival modificada que les permite producir seda para sos- acuáticas y se encuentran comúnmente en ríos, riachuelos, tenerse en el sustrato o para construir redes para filtrar cascadas y lagos. -
Are New Zealand's Marine Caddisflies a ‘Ghost of Gondwana’? Phylogenetic Placement of the Chathamiidae
I Acknowledgments This study would not have been possible without the considerable level of assistance and contribution others have provided to this study. Firstly I would like to thank my supervisor Dr Peter Ritchie who has helped and guided me through this thesis every step of the way, almost three years in the making. Pete willingly took me on long before caddisflies even crossed my mind as a research topic, and his patience with a student in ecology attempting a thesis on genetics has been extraordinary. My co-supervisor Dr Ian Henderson has also been of invaluable assistance, not only going out of his way to collect a considerable number of samples specifically for my study and donating his own specimens, but also willingly providing his formidable expertise to shape and steer this thesis. I am indebted to you both. I cannot thank enough those who were willing to help with my sampling in this study. I must give special thanks to Dr Alice Wells who provided me with samples collected from three different areas in New South Wales Australia. I also must thank Karen Baird for her readiness and willingness to collect for me samples from the almost untouchable Kermadec Islands, for which I am still immensely grateful to have had the fortune to have obtained! Michelle Jenkinson at Department of Conservation was of considerable help and patience for helping me to secure permission for use of the Kermadec material, of whom her and Karen both stepped out of protocol to make up for my own disorganisation! I must also extend my gratitude to the entomological society of New Zealand who provided me with a 21st anniversary grant to facilitate my travel to the Chatham Islands for collection. -
Redalyc.Functional Feeding Groups of Aquatic Insect Families in Latin
Revista de Biología Tropical ISSN: 0034-7744 [email protected] Universidad de Costa Rica Costa Rica Ramírez, Alonso; Gutiérrez-Fonseca, Pablo E. Functional feeding groups of aquatic insect families in Latin America: a critical analysis and review of existing literature Revista de Biología Tropical, vol. 62, núm. 2, abril, 2014, pp. 155-167 Universidad de Costa Rica San Pedro de Montes de Oca, Costa Rica Available in: http://www.redalyc.org/articulo.oa?id=44932430011 How to cite Complete issue Scientific Information System More information about this article Network of Scientific Journals from Latin America, the Caribbean, Spain and Portugal Journal's homepage in redalyc.org Non-profit academic project, developed under the open access initiative Functional feeding groups of aquatic insect families in Latin America: a critical analysis and review of existing literature Alonso Ramírez1 & Pablo E. Gutiérrez-Fonseca2 1. Department of Environmental Sciences, University of Puerto Rico, P.O. Box 190341, San Juan, Puerto Rico 00919; [email protected] 2. Department of Biology, University of Puerto Rico Rio Piedras, San Juan, Puerto Rico 00919; [email protected] Received 12-XII-2013. Corrected 20-I-2014. Accepted 13-II-2014. Abstract: Aquatic macroinvertebrates are involved in numerous processes within aquatic ecosystems. They often have important effects on ecosystem processes such as primary production (via grazing), detritus break- down, and nutrient mineralization and downstream spiraling. The functional feeding groups (FFG) classification was developed as a tool to facilitate the incorporation of macroinvertebrates in studies of aquatic ecosystems. This classification has the advantage of combining morphological characteristics (e.g., mouth part specializa- tion) and behavioral mechanisms (e.g., way of feeding) used by macroinvertebrates when consuming resources.