THE JEPSON GLOBE a Newsletter from the Friends of the Jepson Herbarium

THE JEPSON GLOBE A Newsletter from the Friends of The Jepson Herbarium

VOLUME 25 NUMBER 2, Fall 2015

Director’s Column: Sex and Grant Received to Digitize Our the Single Moss Historic Slide Collection of By Brent D. Mishler Botanical Images! Have you ever thought about how Through a generous award from difficult sexual reproduction is for a the California Digital Library, the Jep- desert moss? They retain the primitive son Herbarium has received funding reproductive system of their aquatic an- to digitize a significant portion of the cestors, with swimming male gametes University and Jepson Herbaria Slide that need to go many times their own Collection. Through the years, over length to find a female gamete—an egg 11,000 color 35mm slides of California cell, retained in the female sex organ plants were organized in family order (the archegonium). This requires a film and labeled with taxon name, date, and of liquid water, a rare occurrence in location. Amy Kasameyer, Archivist, the desert. If the male sex organ (the has transferred these slides into archi- antheridium) is produced on the same val boxes and they will be scanned in gametophyte (the green, haploid phase the coming year. The slides date from in the lifecycle) as the female one, the 1940s and include photos from termed monoicy, crossing is easier but notable herbaria scholars including results in a completely homozygous Introducing Carl Rothfels Bruce Baldwin, Annetta Carter, Lin- sporophyte (the diploid generation I’m very excited to join the Uni- coln Constance, Lauramay Dempster, formed after a successful fertilization), versity Herbarium and Department of Larry Heckard, James Hickman, Robert thus is essentially asexual reproduc- Integrative Biology as a faculty curator! Ornduff, and Alan Smith. The bulk of tion. If the male and female sex organs California is such a spectacular place the photographs were taken in the field are borne on different plants (termed for plants, and Berkeley is an extraor- and include slides showing growth dioicy), then crossing results in a new dinary place to study them. I’m a Ca- habit, landscape context, and close-ups genetic individual but is much more nadian, born and bred an hour north of of key taxonomic characters. These difficult to achieve. Toronto in rural southern Ontario. My (Continued on page 3) The desiccation tolerant desert interest in botany really took off when moss Syntrichia caninervis has been I got a summer job as an Interpretive ALSO IN THIS ISSUE investigated by a research group con- Naturalist at Algonquin Provincial Revisions of Geographic Subdivisions sisting of Lloyd Stark (University of Park, at the border of the boreal forest Undiscovered fern species Nevada, Las Vegas), Nick McLetchie and the broad-leaved hardwoods of the Protection for Livermore tarplant (University of Kentucky), myself, and Laurentian forest. My fellow naturalists Welcome new graduate students many others over a period of nearly 20 and I spent our working days delivering 2015 award recipients years. It is a dominant in the Mojave and interpretive presentations and guided New Jepson eFlora display Great Basin Deserts. Not just a domi- hikes for the park’s visitors, and our 2015 Workshop Year In Review nant moss, but a dominant plant with days off scouring the park for unusual perhaps the greatest amount of ground dragonflies, early migrant warblers, Cover photo: Carl Rothfels in southern cover of any plant in these communi- and, of course, rare plants! Two friends Mexico, with a handful of Cheiloplecton ties. One might think of Great Basin and I spent most of our free waking rigidum (Pteridaceae). Photo by Mónica Palacios-Rios. (Continued on page 8) (Continued on page 9) Collaborative Effort Produces Revisions for the Jepson Manual Geographic Subdivision Boundaries of the Klamath and Cascade Ranges of Northwestern California

By Julie Kierstead Nelson and extend so far east. While relatively near maps to identify the “disputed area” Len Lindstrand III population centers and major transpor- requiring boundary revisions. Then On the way to nowhere, in the tation corridors, this area is functionally we took a hard look at the taxa docu- epicenter of poison-oakiness, steep very remote and characterized by steep mented from the disputed area to see if and rocky, with blast furnace heat in terrain, intense summer heat, and miles they merited geographic range updates summer and torrential rains in winter; of poison oak. Besides recreation at based on the new geographic subdivi- it’s no wonder the southeastern Klam- Shasta Lake, this area receives little sion boundaries. Using CCH records ath Ranges (KR) remain a mystery visitation and has largely been under- to identify these taxa, Jepson staff landscape explored by very few bota- explored botanically for years. developed a list of over 230 species nists. This is the area visited by John Why change the boundary now? for review. We reviewed each taxon O. Sawyer’s “Off the Beaten Path in Several large federal agency projects to identify those occurring in the KR, the Shasta-Trinity National Recreation in this area in the last two decades those that occur in both KR and CaR, Area” Jepson workshop in May of 2008; have produced an abundance of new and those that don’t occur in KR, and the oldest part of the Klamath Ranges. plant distribution and ecological data. submitted the results for appropriate When the 1993 Jepson Manual Additionally, digitized Consortium of updating. Meanwhile, Jepson associate was published, its map of Geographic California Herbaria (CCH) records with Heath Bartosh created a new KR GIS Subdivisions of California drew the KR coordinates now make floristic analysis boundary with our input. Following the boundary to exclude this landscape, much more feasible, and several new boundary change, we updated the exist- including it in the adjacent Cascade southeastern KR endemic species ing Jepson geographic subdivision de- Ranges (CaR). At its eastern bound- have been recently described or are scriptions for KR and CaR to reflect the ary the predominantly metamorphic in process. The time seemed right to boundary revisions. Following review KR meets the volcanic Cascade Range make much-needed changes to the KR and acceptance by the Jepson Curator, (CaR) in an area roughly interpreted by boundary. Bruce Baldwin, we declared victory! the 1993 Jepson map as the Interstate Since early 2014, a team has worked The result is more accurate KR and 5 corridor north of Redding, following together to address this issue and im- CaR boundaries, including the CaRF the upper Sacramento River canyon. prove the Jepson map by (Continued on page 3) This interpretation omits most of the revising the KR and CaR New Klamath Range southeastern Klamath Ranges, a sig- boundaries to better align nificant portion of the KR unique in its with current scientific con- MP own right, and incorrectly includes the sensus about the extents of (exc Wrn) area in CaR, resulting in large areas these geographic subdivi- of limestone and metasedimentary sions and published geol- CaRH geology being placed in CaR, which is ogy, soils, and vegetation inconsistent with the Jepson description maps. This team includes Mount Shasta for that volcanic region. Jepson staff and associates One outcome of the original KR/ Staci Markos, David Bax- KR CaR boundary that is problematic re- ter, Heath Bartosh, and sults when conducting floristic analysis three botanists familiar in this area; for example, Ageratina with the geography and shastensis, a rare KR endemic of lime- flora of the area: Shasta- Shasta Lake stone and metasedimentary rock out- Trinity National Forest NCo Redding crops, has its range described in the Botanist Julie Kierstead second edition of The Jepson Manual Nelson, Len Lindstrand III CaRF (TJM2) as “CaR” because of the mis- of North State Resources NCoRO NCoRI interpreted KR boundary. Other KR Inc., and botanical consul- Red Bluff endemics, such as Arnica venosa, are tant Dean W. Taylor. NCoRH currently described in TJM2 as occur- Our team started with ScV ring both in KR and CaR. our collective experience It’s certainly understandable that and existing ecological the KR region was not interpreted to subregion and geology Map courtesy of Len Lindstrand and Teri Mooney, North State Resources, Inc. 2 California Maidenhair Fern Eludes Discovery for Over a Century

By Layne Huiet five-finger fern, is very easy to identify substrates. Surprisingly, A. shastense The California Floristic Province but A. jordanii and A. capillus-veneris isn’t rare, or found only in inacces- is one of the most intensively collected have a frond architecture that is very sible regions. In fact, a specimen had of the 35 global hotspots tracked by similar even though molecular data been collected in May of 1894 by Milo Conservation International, which shows that they are not closely related. Baker and Frank Nutting and was filed makes it an unlikely place to find an Juvenile and sterile forms of these two in the UC Herbarium, originally identi- undiscovered fern species. But, that’s taxa can be difficult to distinguish. The fied as A. jordanii. This serendipitous exactly what happened when a team most striking difference between them discovery only confirms that herbaria of botanists described Adiantum shas- is that A. jordanii is ephemeral, dying still are the most likely place to find tense, a new species of maidenhair fern back in mid-summer. undiscovered species! that is endemic to northern California To further investigate the identity and currently known only from Shasta of the specimen from Shasta Co., DNA County (Huiet et al., PhytoKeys 53:73- sequencing was employed and, shock- 81. 2015). ingly, it revealed that the plant was nei- While filing herbarium specimens ther A. capillus-veneris nor A. jordanii. at the Duke University Herbarium, I Collaborating with Alan Smith, Cura- noticed a recently received collection tor of Pteridophytes at the University identified as A. jordanii from Shasta and Jepson Herbaria, and Julie Nelson County. I was aware of several re- and Martin Lenz of the USDA Forest cords of A. capillus-veneris from the Service, we found that this new taxon county and thought that this specimen, occurred exclusively around Shasta labeled as A. jordanii, was misidenti- Lake. Appropriately, it has been named fied. Identification of Adiantum taxa A. shastense or the Shasta maidenhair can be tricky—there are three species fern. It is evergreen, in contrast to its of maidenhair fern found throughout sister species A. jordanii and is found Adiantum shastense, Shasta Co. Photo California; Adiantum aleuticum, the on both limestone and metasedimentary by Layne Huiet

(Klamath Range, continued from page 2) (Slide project, continued from page 1) and CaRH geographic subdivisions, photographs of living plants comple- updated geographic division descrip- ment the preserved specimens in our tions, and updated range descriptions collection and support students study- for selected taxa occurring in this area. ing the California flora. The new KR boundary now en- Historically the slide collection has compasses most of the lands east of been used as a teaching tool by faculty Interstate 5, south of Highway 89, and but use has significantly declined as north of Highway 299E to include digital presentations have replaced the Shasta Lake, the McCloud and Hos- slide carousel. We are excited to make selkus limestone formations, and Griz- this valuable educational collection zly Peak. The new CaR boundary now once again available for teaching and starts entirely east of Shasta Lake, and research use but this time, the im- abuts the Inner North Coast Ranges ages and the metadata (identification, subregion along Interstate 5 from west photographer, locality, date) from the of Redding south to Red Bluff. slide mount will be made available Our newly drawn boundary and online via our collections database analysis of CCH specimens have also (CollectionSpace) and the Jepson eFlo- shown clearly that the flora of the south- ra (ucjeps.berkeley.edu/IJM.html). eastern KR has more affinity with the Sierra Nevada than with the intervening CaR; but that is a story for another day. A sample of slides from our collection. Photo by Amy Kasameyer.

3 The Livermore tarplant steps toward official state protection By Heath Bartosh, Research Associate, the potential to be impacted by agri- the petition, CDFW staff determined University and Jepson Herbaria cultural or horse boarding operations there is sufficient scientific information This past spring, the Livermore based on current zoning designations. available at this time to indicate that the tarplant, Deinandra bacigalupii, offi- It is because of these serious threats petitioned action may be warranted and cially became a candidate species under that I decided to prepare a petition they recommended to the Commission the California Endangered Species Act to list the Livermore tarplant as en- that the petition be accepted for con- (CESA), and it is now afforded the same dangered under CESA. Following an sideration. The April 9, 2015, Fish and legal protection as an endangered or expert review from Bruce Baldwin and Game Commission meeting provided threatened species until a final decision Sue Bainbridge (also from the Jepson an opportunity to testify before the is made. Herbarium) the petition was submitted commissioners prior to them voting This conservation success story, a to the California Fish and Game Com- on whether listing was warranted. In collaborative effort of the Jepson Her- mission (Commission) in late August of addition to myself, representatives of barium and the California Native Plant 2014, triggering the process for listing CNPS, with whom I had worked closely Society, is centered in the Livermore consideration. during petition preparation, spoke in Valley, an interior portion of the East It can take up to two years to com- support of the Livermore tarplant. Bay where the habitat is more similar plete the listing process. The first step In front of nearly 200 attendees, the to the Carrizo Plain or southern San in this process is for California Depart- commissioners surprised themselves Joaquin Valley than ecosystems typical ment of Fish and Wildlife (CDFW) per- by voting unanimously to accept the of the Bay Area. The best example of sonnel to review the petition and make petition for consideration. The Liver- this alkaline influenced habitat is the a recommendation to the Commission. more tarplant has officially become a Springtown Alkali Sink. It is here that On September 19, 2014, CDFW person- candidate species! a taxon went undescribed for nearly nel Jeb Bjerke, Cherilyn Burton, and From this point, it may take up to a 40 years even after being collected by Marcia Grefsrud accompanied me on year and a half for a final decision to be Robert Hoover at the corner of Ames a tour of accessible populations. It was made on whether to list the Livermore Street and Raymond Road. Less than on this date that we observed the final tarplant as endangered under CESA. In a year later, the first Jepson Curator, deathblow being dealt to the Greenville the end, if the Commission designates Rimo Bacigalupi, wrote a note dated Road population, which, although dis- the Livermore tarplant as Endangered, April 26, 1967, on Hoover’s specimen heartening, demonstrated the serious it will have received the highest level of stating, “Does not seem to match any threats the Livermore tarplant faces. protection for a plant that lives off fed- thus far published species.” It wasn’t Based on this site visit and a review of eral and state owned lands. Stay tuned! until the late 1990s that investigations by tarplant expert and current Jepson Curator, Bruce Baldwin, took Rimo’s note further and resulted in describ- ing the Livermore tarplant (Deinandra bacigalupii) as a new species (Madroño 46: 55-57. 1999). Since the taxon was first collected in 1966, much of this regionally significant alkali sink has been lost to residential subdivisions and there are only a few populations of the Livermore tarplant that remain. The core population is mostly contained at the city- owned Springtown Wetlands Preserve and there are two others populations on private land. Sadly one of these populations has been extirpated by dumping clean fill on the population alongside Greenville Road. The other remaining population is located near the southeast Livermore tarplant (Deinandra bacigalupii) at the Springtown Wetlands Preserve corner of the Livermore Valley and has in Livermore, the largest population known. Photo by Heath Bartosh. 4 Welcome new graduate students Javier Jáuregui-Lazo Javier Jáuregui-Lazo is a Chil- ean native who recently joined Brent Mishler and the BryoLab as a first-year graduate student. Javier was born in Santiago, Chile, and started to make connections with plants when he studied horticulture for his undergraduate degree. He worked on the postharvest physiology of fruits and developed a system to identify the susceptibility of sweet cherry fruits to physical injury. He was later awarded a Chilean scholarship for graduate study at UC Davis. There, he changed his research interests to the study of botany and evolution of plants. Javier worked on the phylogeny and biogeography of the genus Acaena (Rosaceae), which is widely distributed in the Southern Javier Jáuregui-Lazo in Torres del Paine National Park, Region of Magallanes and Hemisphere, especially in Chile and Antártica Chilena, Chile. Photo by Dominique Dhainaut. New Zealand. At UC Berkeley, his work pleting his degree at UC Berkeley is to ing, drawing, playing soccer, and hav- will focus on measuring and comparing educate future generations of Chilean ing a good time with friends. Spiritually the biodiversity of bryophytes in Chile students and society about the benefits and personally, he is proud of starting and California, both of which have of doing research and communicating an independent life with his family in Mediterranean climates. science in an easy way for everyone. another country. Javier’s main objective after com- During his free time he enjoys hik- Jenna T. Baughman Jenna is a first-year Integrative ter- or cross-disciplinary Biology doctoral student in the Mishler approaches, potentially lab. Prior to this, she earned bachelors’ including genomics, geo- degrees in Biology from Purdue Uni- informatics, biogeochem- versity and Religious Studies from In- istry, morphometrics, citi- diana University, and a master’s degree zen science, and others. in Environmental Science, Biology op- In her free time, Jenna tion, from California State University, enjoys moss hunting, hik- Los Angeles. While at Cal State LA, ing, camping, painting, Jenna used next generation sequence animal training, and, re- data to investigate sex ratios and genetic cently, small-boat sailing. variation of the desiccation-tolerant Jenna lives in Fremont moss Syntrichia caninervis (Pottiaceae) with her partner Somi, her among Mojave Desert microenviron- cat Paws, and her bulldog ments and also characterized a newly mix Billy. discovered Mojave hypolithic moss community. Jenna Baughman on the While she is still exploring disser- Devil’s Backbone trail to the summit of Mount tation ideas, Jenna hopes to continue Baldy in the San Gabriel the theme of ecology and evolution of Mountains of Southern mosses in extreme environments. She California. Photo by Edith is interested in utilizing multiple in- Martinez.

5 Photo by Jack Owicki, TinyURL.com/OwickiPhotos Photo by Jeanne Marie Acceturo 2015 Workshop Year in Review This year’s highlights included (clockwise from top left:): blue skies and bristlecone pines in the White Mountains; intrepid agrostolo- gists at “Advanced Grasses;” Bursera, boojums, and cardons in the Cataviña boulder fields;Epilobium oreganum in the Trinity Alps; peel- ing fossil imprints from coal balls at “Lycophytes, Past and Present;” the Santa Rosa Island endemic Dudleya blochmaniae subsp. insularis; and trailside keying with Linda Beidleman at “50 Families in the Field.”

Photo by Alisha Kerschbaum Photo by Billy Hoyer

Photo by Jeanne Marie Acceturo Photo by Jeanne Marie Acceturo Photo by Aaron E. Sims and CNPS 6 New Jepson eFlora Display! We invite our Friends to visit the new Jepson eFlora display pages! Currently in beta (on the eFlora home page), these new pages will soon replace the old display. Enhanced features include illustrations from The Jepson Manual (2012), more photographs on each page, and a streamlined display of taxonomic information. See ucjeps.berkeley.edu/IJM.html. We welcome your feedback about additional photos that can be included and resolution of “yellow flags,” which represent incongruencies between taxon distributions in the Jepson eFlora and the georeferenced specimens in the Consortium of California Herbaria. Please email Staci Markos with your comments/suggestions ([email protected]).

7 (Director’s column, continued from page 1) sagebrush (Artemisia tridentata) as having the greatest cover in dry valleys of Nevada and Utah, but S. caninervis grows under every bush, and the space in between them, too. It covers a high percent of the soil surface as part of the cryptobiotic soil crust. How did a moss achieve such a smashing success in a harsh envi- ronment? Apparently not via sexual reproduction! It is a dioicious species, and our studies show that sporophytes are quite rare. It is a red letter day when you find one in most locations. Male plants themselves are quite rare—this species has one of the most skewed sex ratios known in plants—as many as 20 females for every male! We don’t yet Mojave Desert site near Phelan, California, Kirsten Fisher and Jenna Baughman gathering samples of Syntrichia caninervis along a transect (photo by Brent Mishler). know exactly the basis for this biased sex ratio. Maybe males are really there eration of S. caninervis aficionados!) in equal proportions, just less prone to are investigating this question using produce their sex organs (the “shy male” modern genomic tools. One of her Curatorial Volunteers hypothesis). Or, alternatively, the males students, Amber Paasch, led a study Needed at the University tend to die at some step in development using microsatellite DNA markers (now and Jepson Herbaria! (the “rare male” hypothesis). in press in the International Journal of My former PhD student, Kirsten Plant Sciences) that confirmed a highly Please help us with mounting, Fisher, now a professor at Cal State clonal population structure in these databasing, and filing herbarium LA, and her students (the third gen- mosses. Others of her students, Jenna specimens! We have group vol- Baughman (who is just unteer days listed below and also beginning PhD studies welcome volunteers during regular in my lab!) and Israel business hours. Weekday volun- Jimenez, have helped to teers help with routine tasks and develop sex-linked mo- specialist projects such as working lecular markers that will on specific collections or taxa. allow identification of any developmental stage 2015-2016 as male or female. Group Volunteer Schedule These molecular September 19 tools, coupled with care- October 10 ful ecological studies in November 7 the field and develop- December 5 mental studies in the lab, February 6 should allow us to finally March 5 understand just how this April 16 Cal Day moss is able to thrive May 7 in such an unexpected Closeup of a clump of the desert moss Syntrichia cani- habitat. nervis (photo by Kirsten Fisher). For more information about our volunteer opportunities or to be KQED public television released a video and story about desicca- added to the reminder list, please tion-tolerant mosses, featuring the Mishler Lab. See: contact Ana Penny 510-642-2465. kqed.org/science/2015/06/25/these-resurrection-plants-spring- No previous curatorial experience back-to-life-in-seconds/ necessary.

8 (Rothfels, continued from page 1) hours wading through bogs and si- fate of polyploids and in Cystopteri- Jepson Herbarium Resources dling along cliffs, trying to see each daceae—I just scratched the surface & Projects related to the of the 1100+ species of vascular plants of each during my dissertation, and California flora recorded from the park. I still remem- Cystopteris in particular contains a Approximately 400,000 plant specimens ber the first new record I found for the lifetime of thrilling frustration. I am from California park—an unassuming little weed by the also interested in almost any ques- Director: Brent D. Mishler name of Sanguisorba minor. And with tion that involves the inference of California Phylodiversity Project that, I was hooked on botany! evolutionary histories and their use in Systematics and ecology of Syntrichia I did a broad undergraduate degree understanding patterns of biodiversity Jepson Curator: Bruce G. Baldwin at McMaster University in Hamilton, and the processes that generated them. Jepson eFlora Systematics and evolution of Calif. tarweeds Ontario (Canada’s banana belt, full of If polyploidy and hybridization are and relatives (tribe Madieae, Compositae), exotic southern things like Lirioden- involved, all the better! I look forward Chaenactis (Chaenactidieae, Compositae), dron and Hamamelis), graduating with to continuing these studies at Berkeley, and Collinsia (Plantaginaceae). a Bachelor of Arts & Science and a home of the world’s best curated fern Curator of Ecology: David Ackerly combined honors in biology. After my collection. Please come by and say hi, Ecology and evolution of California flora undergraduate studies, I travelled for a and I’ll be looking for any available Climate change impacts and conservation year and then found myself with a real assistance in overcoming my terror at strategies job, at Royal Botanical Gardens (RBG), the prospect of learning the California Curator of Ferns and Lycophytes: Carl Ontario, where I would spend the next flora. Rothfels four years as Field Botanist, Herbarium Divergence and hybridization in Califor- Curator, and Natural Lands Steward nian ferns and lycophytes (especially (coincidentally, RBG would also play Notholaena, Cystopteris, Isoetes). an important role in David Baxter’s Curator of Monocots: Chelsea D. Specht path to botany, some years later). My Evolution and biogeography of Calif. monocots (including Allium, Nolina) responsibilities at RBG ranged from Population structure and floral color pat- ecological restoration, to rare species tern diversity in the Calochortus venustus inventory and management, herbarium complex curation, and interpretation and visitor Curator of W. N. Am. Botany: Barbara services, all in RBG’s >1000 hectares Ertter, Flora of Mount Diablo and flora of (~2500 acres) of nature sanctuaries. the East Bay, After RBG, I was drawn back to North American Potentilleae school, this time for my PhD, and it Asst. Director for Collections and Cura- was at this time that I turned to the dark tor of Cultivated plants: Andrew Doran side (i.e., ferns). I headed south, to Duke Asst. Director for Development & Out- University in North Carolina, where I reach: Staci Markos, Jepson eFlora, CCH Carl recreating an iconic image from & Globe editor studied fern phylogenetics under Kath- pteridology: that of Alan Smith’s advisor leen Pryer (Brent Mishler’s former PhD John T. Mickel pressing plants at Rancho Biodiversity Informatics Manager: student). My particular interest was in Tejas, Oaxaca. Photo by James Beck. David Baxter the effects of polyploidy (the state of Collections Staff & Plant Identification: having more than two entire sets of Kim Kersh, Clare Loughran, Ana Penny, and Margriet Wetherwax chromosomes) on the evolutionary fate Archivist and Librarian: Amy Kasameyer of lineages (do polyploids tend to speci- Public Programs: Jeanne Marie Acceturo ate or go extinct more quickly than their Membership, workshop enrollment, non-polyploid relatives?). To study this and Globe design: Edith Summers question, I focused on the most ferny Staff Research Associate: Bridget Wessa of ferns, the genus Cystopteris and Trustees: its relatives in the Cystopteridaceae. Vice Chancellor Emeritus Beth Burnside; After graduating from Duke, I spent UC Botanical Garden Director Paul Licht; two years at the University of British Cathy Park; Professor Emeritus Russell Columbia with Sally Otto studying Jones; Professors John Taylor and Brent D. methods for inferring diversification Notholaena ochracea (Pteridaceae), a Mishler (ex officio) rates from evolutionary trees. xeric-adapted fern, showing its abaxial Constancea: UC Publications in Botany covering of orange farina. Photo by Carl I maintain a strong interest in the Rothfels. (online) 9 Reunion of Herbaria Alumni In August, current and former members of the Baldwin and Mishler labs met in Roberts Park (Oakland, Calif.) for a reunion and to introduce new faculty, students, and post-docs. Many of our alumni now hold prestigious positions around the world and in this issue, we share an update about two herbaria alumni that hold faculty positions at California State Universities. Ben Carter (PhD, Mishler lab, 2012) recently accepted a position as Assistant Professor and Director of the Carl W. Sharsmith Herbarium at San Jose State University. Since finishing his PhD in bryophyte systematics, he Herbaria alumni reunion and new introductions (Roberts Park, August 2015). Left has worked as a research scientist on to right, Back row: Tracy Misiewicz, Somi Ekwealor, Tom Madsen, Naomi Brydon, Santa Catalina Island and as a postdoc- Andrew Thornhill, Carl Rothfels, Brent Mishler, Kirsten Fisher, Eric Harris (with toral scholar at the Duke University son Henry), Dean Kelch, Andrew Murdock, Susan Tremblay, Staci Markos. Middle Herbarium. Although broadly interested row: Abby Wall, Bruce Baldwin, Jenna Baughman, Dennis Wall (with son Tanner), in plant systematics, Ben is particularly Cathy Rushworth, Sophia Khan, Will Freyman (with daughter Aliya Freyman), Lisa interested in the California bryophyte Schultheis, Patricia Sanchez (with son Pablo), Danica Harbaugh (with daughters flora. He maintains active research pro- Taylor and Indigo), Ben Carter (with daughter Cayenne), and Cleo (Andy’s daughter). Front row: Ty and Nate Norvell. Photo by Craig Norvell. jects in floristics, molecular systematics, and biogeography of California bryo- logical Sciences at California State Uni- methods to understanding cryptic diver- phytes, and is particularly interested in versity, Los Angeles where she has been sity in mosses. She and her lab group are understanding how the bryophytes, with a faculty member since 2008. Kirsten’s investigating the potential link between their unique physiology and ecology, research interests include plant molecu- physiological specialization and cryptic contribute to a richer understanding of lar systematics, phylogeography, and diversification in mosses, focusing on plant evolution in California. the use of phylogenetic trees for explor- the desiccation-tolerant model moss Kirsten Fisher (PhD, Mishler lab, ing evolutionary questions. In particu- Syntrichia ruralis. 2004) is an Associate Professor of Bio- lar, she applies molecular phylogenetic

Lifetime Member’s Annual Event On May 17, 2015, Brent Mishler and Bruce Baldwin met with Lifetime Members on the Tiburon peninsula for a morning of botanical walks and discussion. Guests were treated to breakfast at The Lodge at Tiburon, and then everyone joined in walks at Old St. Hilary’s Open Space Preserve and Ring Mountain. Highlights seen during the walks were Streptanthus niger, Hesperolinon congestum, and Calochortus tiburonensis.

Lifetime Members provide sig- Bruce Baldwin (above) answering questions for nificant support for the Herbarium Lifetime Members during the hike in Old St. Hil- and its programs. ary’s Preserve. Calochortus tiburonensis (left) at We are grateful for their Ring Mountain. Photos by Staci Markos. commitment! 10 The Importance of Endowment Funds Larry R. Heckard, former Jepson Curator, and Robert Ornduff, former Director and Professor, each established an endowment fund with a gift from their estate. These endowment funds support graduate students with funding for research expenses and travel to scientific meetings. 2015 Award Recipients The Lawrence R. Heckard Fund of the Jepson Herbarium The Robert Ornduff Fellowship of the University and Will Freyman, Graduate Student, Baldwin Laboratory, Jepson Herbaria UC/JEPS and Department of Integrative Biology. Funds Jeff Benca, Graduate Student, Looy Laboratory, UC were awarded for research titled: Chylismia (Onagraceae) Museum of Paleontology and Department of Integrative phylogenomics and reticulate evolution. Biology. Funds were awarded for Jeff to attend the Next Prahlada Papper, Graduate Student, Ackerly Laboratory, Generation Pteridology Conference at the Smithsonian Department of Integrative Biology. Funds were awarded for National Museum of Natural History in Washington DC. research titled: Species concepts in Californian white oaks Andrew P. Weitz, Graduate Student, Ackerly Laboratory, (Quercus, Fagaceae) Department of Integrative Biology. Funds were awarded for Andrew to attend the annual meeting of the Ecological Society of America in Baltimore, Maryland.

With an estate gift of $50,000 or more, you too could have a lasting legacy on California botany by establishing an endowment fund that will support work that is of particular interest to you. Emerging leaves of Quercus sp. Photo by Prahlada Papper.

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Basic Membership ($35 individual, $50 family) Basic members receive The Jepson Globe and discounts on Weekend Workshops. Sustaining Membership ($200) Receive basic membership benefits plus acknowledgment in the Jepson eFlora. Lifetime Membership ($5,000 total, or pledge a minimum of $250/year) Demonstrate your dedication and commitment to the Jepson Herbarium with a lifetime membership. Gain recog- nition for your support in The Jepson Globe and the Jepson eFlora. Share your ideas with the Director and Curator at special, invitation-only events. My or my spouse’s employer will match this gift. (Please enclose company form.) This gift is ___ in honor of / ___ in memory of ______Please send me information about including the Herbarium in my will or estate plan. Please make your check payable to the UC Regents, charge your gift, or give online at: give.berkeley.edu/fund/?f=FU0840000 Jepson Herbarium, 1001 Valley Life Sciences Building #2465, University of California, Berkeley, CA 94720-2465 Thank you for supporting the Herbarium and its programs!

11 The Jepson Globe, Vol. 25 No. 2 Nonprofit Organization University of California, Berkeley U.S. Postage PAID Friends of the Jepson Herbarium 1001 Valley Life Sciences Building # 2465 University of California, Berkeley, CA 94720-2465 Berkeley

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Spaces Available in Jepson Workshops! There are a few spaces left in the following Berkeley-based Jepson Workshops: October 17-18 Restoration Ecology This workshop will approach work in modified systems through the lens of water, soil, and people. We’ll use local and national case studies as a catalyst for group discussion. Sunday, we’ll visit a large-scale restoration in an urban context. December 11-13 Evolution and Diversity of Mushrooms This workshop will provide an introduction to the biology and identification of California’s mushrooms. Through a combination of lec- tures and discussions, workshop participants will learn about the evolutionary history of fungi and the ecological role of fungi in nature. Advanced Registration is required: information is available at: ucjeps. berkeley.edu/workshops/2015/regform_2015.html

Please call the Herbarium at (510) 643-7008, if you have additional Amamus sebastopol. Photo by questions. Else Vellinga. Registration for 2016 The 2016 Jepson Herbarium Workshop Program schedule will be announced in late November. Current members of the Friends of the Jepson Herbarium can register for workshops one week before the general public. So, if your membership recently expired, or if it will expire soon, be sure to renew – check your address label for your expiration date.

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