DOCSLIB.ORG

Functional Morphology of the Vertebral Column in Remingtonocetus (Mammalia, Cetacea) and the Evolution of Aquatic Locomotion in Early Archaeocetes

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Functional Morphology of the Vertebral Column in Remingtonocetus (Mammalia, Cetacea) and the Evolution of Aquatic Locomotion in Early Archaeocetes Functional Morphology of the Vertebral Column in Remingtonocetus (Mammalia, Cetacea) and the Evolution of Aquatic Locomotion in Early Archaeocetes by Ryan Matthew Bebej A dissertation submitted in partial fulfillment of the requirements for the degree of Doctor of Philosophy (Ecology and Evolutionary Biology) in The University of Michigan 2011 Doctoral Committee: Professor Philip D. Gingerich, Co-Chair Professor Philip Myers, Co-Chair Professor Daniel C. Fisher Professor Paul W. Webb © Ryan Matthew Bebej 2011 To my wonderful wife Melissa, for her infinite love and support ii Acknowledgments First, I would like to thank each of my committee members. I will be forever grateful to my primary mentor, Philip D. Gingerich, for providing me the opportunity of a lifetime, studying the very organisms that sparked my interest in evolution and paleontology in the first place. His encouragement, patience, instruction, and advice have been instrumental in my development as a scholar, and his dedication to his craft has instilled in me the importance of doing careful and solid research. I am extremely grateful to Philip Myers, who graciously consented to be my co-advisor and co-chair early in my career and guided me through some of the most stressful aspects of life as a Ph.D. student (e.g., preliminary examinations). I also thank Paul W. Webb, for his novel thoughts about living in and moving through water, and Daniel C. Fisher, for his insights into functional morphology, 3D modeling, and mammalian paleobiology. My research was almost entirely predicated on cetacean fossils collected through a collaboration of the University of Michigan and the Geological Survey of Pakistan before my arrival in Ann Arbor. Iyad S. Zalmout collected the new specimen of Remingtonocetus domandaensis described in Chapter 3 (as well as many others), and he generously permitted me to study this remarkable specimen for my dissertation work. Others who collected the fossils utilized in my work include Mohammad Anwar, Muhammad Arif, M. Akram Bhatti, William C. Clyde, Philip D. Gingerich, Munir ul-Haq, iii Intizar Hussain Khan, William J. Sanders, and Neil A. Wells. I especially thank William J. Sanders for skillfully preparing these fossils; patiently training me in the art of preparing, molding, and casting fossils; and for all of our stimulating and insightful discussions about vertebral biomechanics, the evolution of cetaceans in general, and, on occasion, the St. Louis Cardinals vs. the New York Mets. My research would also not have been possible without access to specimens of both modern and fossil mammals for comparison. I thank the following people for access to specimens in their care: Steve Hinshaw, Philip Myers, and Priscilla Tucker in the Mammal Division at the University of Michigan Museum of Zoology (Ann Arbor, MI); Judy Galkin and Eileen Westwig at the American Museum of Natural History (New York, NY); and David Bohaska, Linda Gordon, Jeremy Jacobs, and Charlie Potter at the National Museum of Natural History (Washington, DC). My dear friends Kyle and Michele Peterson graciously provided me with food, lodging, and wonderful tours of the District of Columbia during multiple research visits to the National Museum. Many faculty, staff members, post-docs, and fellow students contributed to making the University of Michigan Museum of Paleontology a wonderful and vibrant place to work, including Catherine Badgley, Tom Baumiller, Robyn Burnham, Devapriya Chattopadhyay, Mike Cherney, Mike D’Emic, Tom Eiting, Julia Fahlke, John Finarelli, Brady Foreman, John Graf, Gregg Gunnell, Amber Heard, Takehito Ikejiri, Alex Janevski, Cait Lamborne, Carly Manz, Amy McKeighan, Dan Miller, Chrissy Minor, Vlad Miskevich, Pieter Missiaen, Emile Moacdieh, Meegan Novara, Shanan Peters, Gerald Smith, Cindy Stauch, John Whitlock, Jeff Wilson, Miriam Zelditch, and the myriad of students working iv in the prep lab. Bonnie Miljour provided the beautiful reconstruction of R. domandaensis (see the frontispiece and Chapter 3), along with invaluable help in navigating Adobe Photoshop and Illustrator. Adam Rountrey was an invaluable resource on many topics, especially 3D scanning and modeling, and Katy Smith offered innumerable discussions about mammalian evolution, multivariate statistics, the latest episode of Lost, and Major League Baseball. I will forever be indebted to Aaron Wood for his depth of insight, his listening ear, his unwavering advice, and, most of all, his friendship. I am also grateful for many faculty, staff members, and students in the Department of Ecology and Evolutionary Biology (EEB), including Prosanta Chakrabarty, Clay Cressler, Deborah Goldberg, Aaron Iverson, Mandy Izzo, George Kling, Gail Kuhnlein, Jessica Middlemis Maher, the late Beverly Rathcke, Nancy Smith, and Rachel Vannette. Julia Eussen and Jane Sullivan were fabulous graduate coordinators, and I am thankful for their willingness to deal with and answer any question, no matter how inane. My experiences teaching with Marc Ammerlaan, Bill Fink, Phil Myers, and Ron Nussbaum were fantastic. But most of my favorite memories teaching are from my three semesters as co-graduate student instructor with Lucia Luna; she taught me so much, and her enthusiasm and humor always made my day. I also thank Chris Van Ee at Design Research Engineering (Novi, MI), whose time and expertise made it possible for me to develop the multibody dynamic models in Chapter 5; Brett Lyons and Eric Maslowski of the University of Michigan 3D Lab; my undergraduate mentors Ralph Stearley, Randy Van Dragt, and Dave Warners of Calvin v College (Grand Rapids, MI) for their continued encouragement and support; Rolf Bouma and all of the Au Sable Graduate Fellows who have come and gone in the past six years; and Matt Remy and Paul Steen for their camaraderie, graduate life commiseration, and board game acumen. Lastly, I must express my sincere gratitude to my family, especially my parents and my in-laws, for all of their love and support over the years. Most of all, I am extremely thankful to my wife Melissa, who has given and sacrificed so much for me as I pursued my dream. Her love, encouragement, smile, and culinary skills are appreciated far more than she could ever know. And last, but not least, I am grateful for my wonderful little dog Duncan, who has never failed to greet me with a wagging tail, a belly to scratch, and/or a lick on the face after coming home from a long day at work— and who also never seemed to mind that I tried to gain insight into vertebral function by staring at his lumbar region while he gallivanted around our apartment. This research was supported by EEB Block Grants, the Peter Olaus Okkelberg Award, and the University of Michigan Museum of Paleontology. vi Table of Contents Dedication ............................................................................................................................ ii Acknowledgments............................................................................................................... iii List of Figures ..................................................................................................................... xii List of Tables ...................................................................................................................... xv List of Abbreviations ........................................................................................................ xvii Abstract ............................................................................................................................. xix Chapter 1: Introduction ...................................................................................................... 1 Background ................................................................................................................... 1 Evolution of Aquatic Locomotion ................................................................................. 3 Changes in the Function of the Vertebral Column ....................................................... 6 The Archaeocete Family Remingtonocetidae ............................................................... 8 Overview ..................................................................................................................... 10 Figures ......................................................................................................................... 14 Table ............................................................................................................................ 18 References .................................................................................................................. 19 vii Chapter 2: Systematic Review of the Remingtonocetidae (Mammalia, Cetacea) ........... 33 Introduction ................................................................................................................ 33 Age of Remingtonocetid-Bearing Formations ............................................................ 34 Systematic Paleontology ............................................................................................. 37 Discussion.................................................................................................................... 54 Are Dalanistes and Remingtonocetus males and females of a single species? .... 54 Differences in Size Between Homologous Postcranial Elements ................... 58 Differences in Size Between Canine Teeth ..................................................... 60 Stratigraphic
Load more

Recommended publications
  • JVP 26(3) September 2006—ABSTRACTS
    Neoceti Symposium, Saturday 8:45 acid-prepared osteolepiforms Medoevia and Gogonasus has offered strong support for BODY SIZE AND CRYPTIC TROPHIC SEPARATION OF GENERALIZED Jarvik’s interpretation, but Eusthenopteron itself has not been reexamined in detail. PIERCE-FEEDING CETACEANS: THE ROLE OF FEEDING DIVERSITY DUR- Uncertainty has persisted about the relationship between the large endoskeletal “fenestra ING THE RISE OF THE NEOCETI endochoanalis” and the apparently much smaller choana, and about the occlusion of upper ADAM, Peter, Univ. of California, Los Angeles, Los Angeles, CA; JETT, Kristin, Univ. of and lower jaw fangs relative to the choana. California, Davis, Davis, CA; OLSON, Joshua, Univ. of California, Los Angeles, Los A CT scan investigation of a large skull of Eusthenopteron, carried out in collaboration Angeles, CA with University of Texas and Parc de Miguasha, offers an opportunity to image and digital- Marine mammals with homodont dentition and relatively little specialization of the feeding ly “dissect” a complete three-dimensional snout region. We find that a choana is indeed apparatus are often categorized as generalist eaters of squid and fish. However, analyses of present, somewhat narrower but otherwise similar to that described by Jarvik. It does not many modern ecosystems reveal the importance of body size in determining trophic parti- receive the anterior coronoid fang, which bites mesial to the edge of the dermopalatine and tioning and diversity among predators. We established relationships between body sizes of is received by a pit in that bone. The fenestra endochoanalis is partly floored by the vomer extant cetaceans and their prey in order to infer prey size and potential trophic separation of and the dermopalatine, restricting the choana to the lateral part of the fenestra.
    [Show full text]
  • The World at the Time of Messel: Conference Volume
    T. Lehmann & S.F.K. Schaal (eds) The World at the Time of Messel - Conference Volume Time at the The World The World at the Time of Messel: Puzzles in Palaeobiology, Palaeoenvironment and the History of Early Primates 22nd International Senckenberg Conference 2011 Frankfurt am Main, 15th - 19th November 2011 ISBN 978-3-929907-86-5 Conference Volume SENCKENBERG Gesellschaft für Naturforschung THOMAS LEHMANN & STEPHAN F.K. SCHAAL (eds) The World at the Time of Messel: Puzzles in Palaeobiology, Palaeoenvironment, and the History of Early Primates 22nd International Senckenberg Conference Frankfurt am Main, 15th – 19th November 2011 Conference Volume Senckenberg Gesellschaft für Naturforschung IMPRINT The World at the Time of Messel: Puzzles in Palaeobiology, Palaeoenvironment, and the History of Early Primates 22nd International Senckenberg Conference 15th – 19th November 2011, Frankfurt am Main, Germany Conference Volume Publisher PROF. DR. DR. H.C. VOLKER MOSBRUGGER Senckenberg Gesellschaft für Naturforschung Senckenberganlage 25, 60325 Frankfurt am Main, Germany Editors DR. THOMAS LEHMANN & DR. STEPHAN F.K. SCHAAL Senckenberg Research Institute and Natural History Museum Frankfurt Senckenberganlage 25, 60325 Frankfurt am Main, Germany [email protected]; [email protected] Language editors JOSEPH E.B. HOGAN & DR. KRISTER T. SMITH Layout JULIANE EBERHARDT & ANIKA VOGEL Cover Illustration EVELINE JUNQUEIRA Print Rhein-Main-Geschäftsdrucke, Hofheim-Wallau, Germany Citation LEHMANN, T. & SCHAAL, S.F.K. (eds) (2011). The World at the Time of Messel: Puzzles in Palaeobiology, Palaeoenvironment, and the History of Early Primates. 22nd International Senckenberg Conference. 15th – 19th November 2011, Frankfurt am Main. Conference Volume. Senckenberg Gesellschaft für Naturforschung, Frankfurt am Main. pp. 203.
    [Show full text]
  • West Indian Manatee (Trichechus Manatus) Habitat Characterization Using Side-Scan Sonar
    Andrews University Digital Commons @ Andrews University Master's Theses Graduate Research 2017 West Indian Manatee (Trichechus Manatus) Habitat Characterization Using Side-Scan Sonar Mindy J. McLarty Andrews University, [email protected] Follow this and additional works at: https://digitalcommons.andrews.edu/theses Part of the Biology Commons Recommended Citation McLarty, Mindy J., "West Indian Manatee (Trichechus Manatus) Habitat Characterization Using Side-Scan Sonar" (2017). Master's Theses. 98. https://digitalcommons.andrews.edu/theses/98 This Thesis is brought to you for free and open access by the Graduate Research at Digital Commons @ Andrews University. It has been accepted for inclusion in Master's Theses by an authorized administrator of Digital Commons @ Andrews University. For more information, please contact [email protected]. ABSTRACT WEST INDIAN MANATEE (TRICHECHUS MANATUS) HABITAT CHARACTERIZATION USING SIDE-SCAN SONAR by Mindy J. McLarty Chair: Daniel Gonzalez-Socoloske ABSTRACT OF GRADUATE STUDENT RESEARCH Thesis Andrews University School of Arts and Sciences Title: WEST INDIAN MANATEE (TRICHECHUS MANATUS) HABITAT CHARACTERIZATION USING SIDE-SCAN SONAR Name of researcher: Mindy J. McLarty Name and degree of faculty chair: Daniel Gonzalez-Socoloske, Ph.D. Date completed: April 2017 In this study, the reliability of low cost side-scan sonar to accurately identify soft substrates such as grass and mud was tested. Benthic substrates can be hard to classify from the surface, necessitating an alternative survey approach. A total area of 11.5 km2 was surveyed with the sonar in a large, brackish mangrove lagoon system. Individual points were ground-truthed for comparison with the sonar recordings to provide a measure of accuracy.
    [Show full text]
  • Wec01's SSSS Fossils Test 2019
    wec01’s SSSS Fossils Test 2019 Team Name: _________________KEY________________ Team Number: ___KEY___ Team Members: ____________KEY____________, ____________KEY____________ This test consists of 18 stations with a total of 200 points. Each answer is worth one point except where specified otherwise. You are only given 2 ½ minutes with the specimens at each station, however you can work on any station’s questions at any time. Scoring Station 1: ___10___ / 10 Station 10: ___12___ / 12 Station 2: ___10___ / 10 Station 11: ____9___ / 9 Station 3: ___11___ / 11 Station 12: ___11___ / 11 Station 4: ___10___ / 10 Station 13: ___10___ / 10 Station 5: ___10___ / 10 Station 14: ___10___ / 10 Station 6: ____9___ / 9 Station 15: ___12___ / 12 Station 7: ____9___ / 9 Station 16: ____9___ / 9 Station 8: ___10___ / 10 Station 17: ___10___ / 10 Station 9: ____9___ / 9 Station 18: ___29___ / 29 Total: __200___ / 200 Team Number: _KEY_ Station 1: Dinosaurs (10 pt) 1. Identify the genus of specimen A Tyrannosaurus (1 pt) 2. Identify the genus of specimen B Stegosaurus (1 pt) 3. Identify the genus of specimen C Allosaurus (1 pt) 4. Which specimen(s) (A, B, or C) are A, C (1 pt) Saurischians? 5. Which two specimens (A, B, or C) lived at B, C (1 pt) the same time? 6. Identify the genus of specimen D Velociraptor (1 pt) 7. Identify the genus of specimen E Coelophysis (1 pt) 8. Which specimen (D or E) is commonly E (1 pt) found in Ghost Ranch, New Mexico? 9. Which specimen (A, B, C, D, or E) would D (1 pt) specimen F have been found on? 10.
    [Show full text]
  • A New Middle Eocene Protocetid Whale (Mammalia: Cetacea: Archaeoceti) and Associated Biota from Georgia Author(S): Richard C
    A New Middle Eocene Protocetid Whale (Mammalia: Cetacea: Archaeoceti) and Associated Biota from Georgia Author(s): Richard C. Hulbert, Jr., Richard M. Petkewich, Gale A. Bishop, David Bukry and David P. Aleshire Source: Journal of Paleontology , Sep., 1998, Vol. 72, No. 5 (Sep., 1998), pp. 907-927 Published by: Paleontological Society Stable URL: https://www.jstor.org/stable/1306667 REFERENCES Linked references are available on JSTOR for this article: https://www.jstor.org/stable/1306667?seq=1&cid=pdf- reference#references_tab_contents You may need to log in to JSTOR to access the linked references. JSTOR is a not-for-profit service that helps scholars, researchers, and students discover, use, and build upon a wide range of content in a trusted digital archive. We use information technology and tools to increase productivity and facilitate new forms of scholarship. For more information about JSTOR, please contact [email protected]. Your use of the JSTOR archive indicates your acceptance of the Terms & Conditions of Use, available at https://about.jstor.org/terms SEPM Society for Sedimentary Geology and are collaborating with JSTOR to digitize, preserve and extend access to Journal of Paleontology This content downloaded from 131.204.154.192 on Thu, 08 Apr 2021 18:43:05 UTC All use subject to https://about.jstor.org/terms J. Paleont., 72(5), 1998, pp. 907-927 Copyright ? 1998, The Paleontological Society 0022-3360/98/0072-0907$03.00 A NEW MIDDLE EOCENE PROTOCETID WHALE (MAMMALIA: CETACEA: ARCHAEOCETI) AND ASSOCIATED BIOTA FROM GEORGIA RICHARD C. HULBERT, JR.,1 RICHARD M. PETKEWICH,"4 GALE A.
    [Show full text]
  • Energetic Tradeoffs Control the Size Distribution of Aquatic Mammals William Gearty
    University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Faculty Publications in the Biological Sciences Papers in the Biological Sciences 4-17-2018 Energetic tradeoffs control the size distribution of aquatic mammals William Gearty Craig R. McClain Jonathan Payne Follow this and additional works at: https://digitalcommons.unl.edu/bioscifacpub Part of the Biology Commons, Evolution Commons, and the Terrestrial and Aquatic Ecology Commons This Article is brought to you for free and open access by the Papers in the Biological Sciences at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Faculty Publications in the Biological Sciences by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. Energetic tradeoffs control the size distribution of aquatic mammals William Geartya,1, Craig R. McClainb, and Jonathan L. Paynea aDepartment of Geological Sciences, Stanford University, Stanford, CA 94305; and bLouisiana Universities Marine Consortium, Chauvin, LA 70344 Edited by Nicholas D. Pyenson, Smithsonian Institution, Washington, DC, and accepted by Editorial Board Member David Jablonski February 23, 2018 (received for review August 8, 2017) Four extant lineages of mammals have invaded and diversified in the entering the water will increase in average size, these theories differ in water: Sirenia, Cetacea, Pinnipedia, and Lutrinae. Most of these aquatic their predictions for how such a size change is achieved. More spe- clades are larger bodied, on average, than their closest land-dwelling cifically, they differ in their predictions both about the rate of evo- relatives, but the extent to which potential ecological, biomechanical, lution toward the new, larger average size as well as the variance of and physiological controls contributed to this pattern remains untested the aquatic size distribution relative to its terrestrial sister group (22).
    [Show full text]
  • The Genome of the Pyrenean Desman and the Effects of Bottlenecks and Inbreeding on the Genomic Landscape of an Endangered Species
    Received: 18 August 2020 | Revised: 19 March 2021 | Accepted: 27 April 2021 DOI: 10.1111/eva.13249 ORIGINAL ARTICLE The genome of the Pyrenean desman and the effects of bottlenecks and inbreeding on the genomic landscape of an endangered species Lídia Escoda | Jose Castresana Institute of Evolutionary Biology (CSIC- Universitat Pompeu Fabra), Barcelona, Spain Abstract The Pyrenean desman (Galemys pyrenaicus) is a small semiaquatic mammal endemic Correspondence Jose Castresana, Institute of Evolutionary to the Iberian Peninsula. Despite its limited range, this species presents a strong ge- Biology (CSIC- Universitat Pompeu Fabra), netic structure due to past isolation in glacial refugia and subsequent bottlenecks. Passeig Marítim de la Barceloneta 37, 08003 Barcelona, Spain. Additionally, some populations are highly fragmented today as a consequence of river Email: [email protected] barriers, causing substantial levels of inbreeding. These features make the Pyrenean Funding information desman a unique model in which to study the genomic footprints of differentiation, Plan Nacional I+D+i del Ministerio de bottlenecks and extreme isolation in an endangered species. To understand these Ciencia e Innovación, Grant/Award Number: CGL2017- 84799- P (MINECO/AEI/FEDER, UE) processes, the complete genome of the Pyrenean desman was sequenced and as- sembled using a Bloom filter- based approach. An analysis of the 1.83 Gb reference genome and the sequencing of five additional individuals from different evolutionary units allowed us to detect its main genomic characteristics. The population differen- tiation of the species was reflected in highly distinctive demographic trajectories. In addition, a severe population bottleneck during the postglacial recolonization of the eastern Pyrenees created one of the lowest genomic heterozygosity values recorded in a mammal.
    [Show full text]
  • Muscles of the Thorax, Back & Abdomen
    MUSCLES OF THE THORAX, BACK & ABDOMEN Muscles of the Thorax Thoracic Muscles Origin Insertion Action Innervation M. pectoralis clavicula pars clavicularis major (medial ½ ) manubrium sterni et adduction, internal M. pectoralis pars crista tuberculi cartilagines costae rotation, arm flexion; major sternocostalis majoris (2nd-7th) auxiliary inspiratory m. M. pectoralis vagina musculi recti pars abdominalis major abdominis Plexus brachialis processus pulls the clavicle; M. pectoralis minor 3rd - 5th rib coracoideus auxiliary inspiration m scapulae pulls clavicule → clavicula indirectly the shoulder M. subclavius first rib (inferior surface) distoventrally; auxiliary inspiration m. pulls the clavicle from scapula the backbone; pulls M. serratus anterior cranial 9 ribs (margo medialis et inferior angle laterally → angulus inferior) rotates scapula; auxiliary respirat. m. Thoracic Muscles Origin Insertion Action Innervation inferior margin of ribs - superior margin of elevation of lower ribs, from the costal tubercle Mm. intercostales externi ribs immediately thorax expansion → to the beginning of rib below inspiratory m. cartilage inferior margin of adduction of cranial superior margin of ribs - Nn. Mm. intercostales interni ribs immediately ribs to caudal ribs → intercostales costal angle to sternum above expiratory m. internal surface of cartilagines costae M. transversus thoracis xiphoid process and expiratory muscle verae body of sternum inner surface of xiphoid Diaphragma sternal part process inner surface of Diaphragma costal part cartilage of ribs 7-12 main inspiratory Plexus central tendon muscle; abdominal ligamentum cervicalis lumbar part, press Diaphragma longitudinale anterius medial crus (vertebrae lumbales) ligaments jump over the lumbar part, Diaphragma psoas and quadratus lateral crus muscles Muscles of the Back Superficial muscles . functionally belong to the upper limb Intermediate muscles .
    [Show full text]
  • Checklist of Rodents and Insectivores of the Mordovia, Russia
    ZooKeys 1004: 129–139 (2020) A peer-reviewed open-access journal doi: 10.3897/zookeys.1004.57359 RESEARCH ARTICLE https://zookeys.pensoft.net Launched to accelerate biodiversity research Checklist of rodents and insectivores of the Mordovia, Russia Alexey V. Andreychev1, Vyacheslav A. Kuznetsov1 1 Department of Zoology, National Research Mordovia State University, Bolshevistskaya Street, 68. 430005, Saransk, Russia Corresponding author: Alexey V. Andreychev ([email protected]) Academic editor: R. López-Antoñanzas | Received 7 August 2020 | Accepted 18 November 2020 | Published 16 December 2020 http://zoobank.org/C127F895-B27D-482E-AD2E-D8E4BDB9F332 Citation: Andreychev AV, Kuznetsov VA (2020) Checklist of rodents and insectivores of the Mordovia, Russia. ZooKeys 1004: 129–139. https://doi.org/10.3897/zookeys.1004.57359 Abstract A list of 40 species is presented of the rodents and insectivores collected during a 15-year period from the Republic of Mordovia. The dataset contains more than 24,000 records of rodent and insectivore species from 23 districts, including Saransk. A major part of the data set was obtained during expedition research and at the biological station. The work is based on the materials of our surveys of rodents and insectivo- rous mammals conducted in Mordovia using both trap lines and pitfall arrays using traditional methods. Keywords Insectivores, Mordovia, rodents, spatial distribution Introduction There is a need to review the species composition of rodents and insectivores in all regions of Russia, and the work by Tovpinets et al. (2020) on the Crimean Peninsula serves as an example of such research. Studies of rodent and insectivore diversity and distribution have a long history, but there are no lists for many regions of Russia of Copyright A.V.
    [Show full text]
  • Active Release Techniques Spine Level 2
    Active Release Techniques Spine Level 2 Dates of program- Montvale, NJ February 18-21, 2021 Colorado Springs, CO March 4-7, 2021 Orlando, FL June 10-13, 2021 Chicago, IL September 30 – October 3, 2021 Total Hours: 24 Summary: Active Release Techniques® Spine Level 2 offers intense training in 75 manual treatment protocols of the cervical, thoracic, and lumbar spine. ART® treatment utilizes manual techniques to move tissues and joints while under tension. The system allows for relative motion between the tissues and articulations. This seminar emphasizes the manipulation of the neuromusculoskeletal system to diagnose and correct alterations in tissue texture, tension, movement, and function between tissues. Evaluation and treatment occur simultaneously. Learning Outcomes: 1. By the end of the seminar, learners will be able to correctly identify (palpate) 75 facial seams of soft-tissue structures within the spine. 2. By the end of the seminars, learners will be able to correctly state the muscle actions of two adjacent spinal muscles. 3. By the end of the seminar, learners will be able to effectively recognize common symptom patterns of spinal neuromuscular injuries and disorders. 4. By the end of the seminar, learners will correctly identify the structure treated and associated concentric and eccentric muscle actions via video presentations. 5. By the end of the seminar, the learner will correctly move the muscle from its shortened position to elongated position using two-hand placement techniques. 6. By the end of the seminar, the learner can successfully differentiate between healthy and unhealthy tissue utilizing hands-on palpation techniques. 7. By the end of the seminar, the learner will proficiently palpate 75 anatomical soft-tissue structures within the spine, using an appropriate tension, depth, and motion to properly perform the treatment protocol.
    [Show full text]
  • Whale Evolution: a Whale of a Tale
    Creation Research Society Quarterly 2012. 49:122–134. 122 Creation Research Society Quarterly Whale Evolution: A Whale of a Tale Jerry Bergman* Abstract review of the evolution of whales from terrestrial land animals finds A that the evidence used to support the current theory is either wrong or very questionable. A focus is on the hip bone and fetal teeth evidence because they are commonly used as proof for the land mammal-to-whale evolution theory. The putative fossil evidence for whale evolution from terrestrial animals is also evaluated, concluding that the examples used are likely all extinct mammals and not transitional forms. Introduction be only about 20 feet long, but baleen A major problem has been deter- The term “whale” is a common noun and blue whales can grow up to 100 feet mining which terrestrial animal whales that can refer to all marine mammals long. Toothed whales are, on average, evolved from. Charles Darwin proposed called cetaceans (members of order smaller then baleen whales, ranging one of the first theories of whale evolu- cetacea), including dolphins and por- from 3 to 32 feet long, although most tion, suggesting they evolved from bears. poises. In this paper the term “whales” are from 10 to 30 feet long. Blue whales He wrote, “I can see no difficulty in a excludes both dolphins and porpoises. can weigh up to 150 tons. race of bears being rendered, by natural Classification of whales divides them selection, more and more aquatic in into two groups; toothed whales and their structure and habits, with larger baleen whales, the latter of which use The Origin of Whales and larger mouths, till a creature was large brush-like structures to filter food The evolution of whales is one of the produced as monstrous as a whale” from the ocean.
    [Show full text]
  • Currently Zygorhiza Kochii; Mammalia, Cetacea): Proposed Replacement of the Holotype by a Neotype
    Case 3611Basilosaurus kochii Reichenbach, 1847 (currently Zygorhiza kochii; Mammalia, Cetacea): proposed replacement of the holotype by a neotype Author: Uhen, Mark D. Source: The Bulletin of Zoological Nomenclature, 70(2) : 103-107 Published By: International Commission on Zoological Nomenclature URL: https://doi.org/10.21805/bzn.v70i2.a14 BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access titles in the biological, ecological, and environmental sciences published by nonprofit societies, associations, museums, institutions, and presses. Your use of this PDF, the BioOne Complete website, and all posted and associated content indicates your acceptance of BioOne’s Terms of Use, available at www.bioone.org/terms-of-use. Usage of BioOne Complete content is strictly limited to personal, educational, and non - commercial use. Commercial inquiries or rights and permissions requests should be directed to the individual publisher as copyright holder. BioOne sees sustainable scholarly publishing as an inherently collaborative enterprise connecting authors, nonprofit publishers, academic institutions, research libraries, and research funders in the common goal of maximizing access to critical research. Downloaded From: https://bioone.org/journals/The-Bulletin-of-Zoological-Nomenclature on 08 Apr 2021 Terms of Use: https://bioone.org/terms-of-use Access provided by Auburn University Bulletin of Zoological Nomenclature 70(2) June 2013 103 Case 3611 Basilosaurus kochii Reichenbach, 1847 (currently Zygorhiza kochii; Mammalia, Cetacea): proposed replacement of the holotype by a neotype Mark D. Uhen Department of Atmospheric, Oceanic, and Earth Sciences, George Mason University, MS 5F2, Fairfax, VA 22030, U.S.A. (e-mail: [email protected]) Abstract.
    [Show full text]