New Leaf-Rolling Weevils (Coleoptera: Rhynchitidae, Attelabidae) from China

Baltic J. Coleopterol. 5 (2) 2005 ISSN 1407 - 8619

New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China

Andrei A. Legalov, Ning Liu

Legalov A.A., Liu N. 2005. New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China. Baltic J. Coleopterol., 5 (2): 99 - 132.

In this paper, 44 new taxa (5 new genera, 3 new subgenera and 36 new species) are described and 1 new combination is established.

Key words: Coleoptera, Curculionoidea, Rhynchitidae, Attelabidae, phylogeny, fauna, China

Andrei Legalov. Siberian Zoological Museum, Institute of Animal Systematics and Ecology, SB RAS, Frunze street-11, Novosibirsk 630091 Russia; e-mail: [email protected] Ning Liu. Institute of Zoology, Chinese Academy of Sciences, 25 Beisihuan Xilu, Beijing 100080, China; e-mail: [email protected]

INTRODUCTION Zoology, Academia Sinica, (Beijing); SZMN = Siberian Zoological Museum, Institute of Ani- mal Systematics and Ecology, SB RAS A growing interest in studies of the leaf-rolling (Novosibirsk). weevils (Legalov, 2003, 2004a, 2004b, 2004c, 2005) in the world fauna can be seen in recent years. The fauna of China is one of the richest in the world, with 168 species of Rhynchitidae and 193 RESULTS species of Attelabidae (Legalov, 2003b). This paper is based on the collections of the Institute of Zoology. 36 species are described here as new. Family Rhynchitidae Gistel, 1848 Thus, the fauna of China now totals 397 species. Tribe Isotheini Scudder, 1893 Genus Deporaus Samouelle, 1819

MATERIAL AND METHODS Deporaus ruber Legalov et Liu, sp.n. (Figs. 1-2) . Female (IZAS), China, Yunnan, Diqing, Type specimens are kept in the following collec- Holotype 3800 m, 01.VI.1984, Li Changfang. tions and museums: CKJU = Collection of P. Kresl (Janovice nad Uhlavou); IZAS = Institute of

99 Legalov A.A., Liu N.

Description. Female: Body red with sparse, light Points in them dense and fine. 9th stria merges semierect setae. Metathorax dark. Rostrum long, with 10th stria before 2nd ventrite. Prothorax lack- thick, weakly curved. Forehead wide, punctured. ing lustre, densely punctured. Mes-, Antennae located in the middle of rostrum, of metepisternum, meso-, metathorax densely and average size, reaching as far as first line of strongly punctured. Mesepisternum well visible pronotum. Pronotum lengthened, punctured. from above. Metepisternum very wide. Abdomen Elytra lengthened. Greatest width behind the mid- convex, weakly flattened in the middle, densely dle. Intervals convex, wide. Striae narrow and and finely punctured. Legs long. Tarsi long. 1st deep. Abdomen convex. Legs long. Femora wid- segment long and wide. 2nd segment wide - trian- ened. Tibiae narrow, weakly curved. Length of gular. 3rd segment bilobed. Caudal segment body: 3.4 mm. lengthened. Claws with long teeth. Length of body: 5.3 mm. Diagnosis. This new species is close to Deporaus bicolor Voss, 1938 but can be distinguished by Diagnosis. This new genus is close to genus the uniformly red colour of the body, a long ros- Byctiscophilus Voss, 1930 and can be distin- trum, weakly convex eyes and narrower body. guished by the bottom covered with dense, long, Etymology. The name is formed from the word appressed yellow setae which are concentrated for “red” – “ruber”. on the mesepisternum, metathorax and sides of ventrites, and a markedly lengthened and wider Tribe Byctiscini Voss, 1923 1st segment of tarsi.

Genus Baikovius Legalov, gen.n. (Figs. 3-4) Etymology. This new genus is named in honour of Konstantin Baikov (Novosibirsk). Type species: Baikovius unicus Legalov et Liu, sp.n. Baikovius unicus Legalov et Liu, sp.n. (Figs. 3- 4) Description. Body dark brown. Rostrum, clava of antennae, mesepisternum, mesothorax, Holotype. Female (IZAS), China, Yunnan, pygidium, first third of femora and tibiae red- Xishuangbanna, 620-650 m, 28.VIII.1959, Zhang brown. Metathorax, abdomen and metepisternum Facai. red. Body with densely appressed light setae. Bottom with densely long, appressed yellow se- Description. Female: Body dark brown. Rostrum, tae, which concentrate on mesepisternum, antennal clava, mesepisternum, mesothorax, metathorax and sides of ventrites. Rostrum al- pygidium, first third of femora, tibiae red-brown. most straight, thin, long, finely and densely punc- Metathorax, abdomen and metepisternum red. tured, not widening toward apex, with carina from Body with dense, appressed light setae. Bottom basis up to place of antennal attachment. Anten- with dense, long, appressed yellow setae, which nae located before the middle of rostrum. Fore- are concentrated on mesepisternum, metathorax head narrow, densely and roughly punctured. and sides of ventrites. Rostrum almost straight, Vertex convex, densely punctured. Temples not thin, long, 9 times longer than wide and 1.5 times lengthened. Eyes large, not protruding from con- longer than pronotum, finely and densely punc- tour of head. Antennae long, reaching as far as tured, not widening toward apex, with carina from the middle of pronotum. Pronotum almost trap- basis up to place of antennal attachment. Anten- ezoid, narrowed toward apex. Sides almost rec- nae located before the middle of rostrum. Fore- tangular. Disk convex, densely and strongly head narrow, densely and roughly punctured. punctured, with medial striae. Scutellum wide, Vertex convex, densely punctured. Temples not back trapezoid. Elytra wide. Humeri weakly lengthened. Eyes large, not protruding from con- smoothed. Intervals wide, densely, large and tour of head. Antennae long, reaching as far as roughly punctured. Striae narrow and deep. the middle of pronotum. Scapus, 1st-5th segments

100 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China

Figs. 1-13. Rhynchitidae gen. spp.: 1-2 – Deporaus ruber, 3-4 – Baikovius unicus, 5 – Byctiscus minimus (male), 6 – Aspidobyctiscus (Eobyctiscus) vossi (male), 7 – A. (E.) vossi (female), 8 – A. (E.) zhejiangensis, 9 – Byctiscus minimus (female), 10 – Aspidobyctiscus (Taiwanobyctiscus) cyanocuperus, 11 – A. (Eobyctiscus) pseudocoerulans, 12 – A. (Parabyctiscus) konoi, 13 – Nelistrobyctiscus (Listrobyctiscidius) pseudopatruelis. of funicle lengthened - oval. Scapus wider than times longer than wide. Humeri weakly smoothed. funicle. 1st segment more short than 2nd segment. Intervals wide, densely, large and roughly punc- 6th segment short - oval. 7th segment transversal, tured. Striae narrow and deep. Points in them similar to segments of clava. Clava short, much dense and fine. Scutellar striole advanced. 9th stria wider than funicle. 1st claval segment longer than merges with 10th stria before 2nd ventrite. Prothorax 2nd segment. 3rd segment shorter than previous lacking lustre, densely punctured. Mes-, segments, pointed. Pronotum almost trapezoid, metepisternum, meso-, metathorax densely and narrowed toward apex, 1.2 times wider than long. strongly punctured. Mesepisternum well visible Sides almost rectangular. Disk convex, densely from above. Metepisternum very wide. Abdomen and strongly punctured, with medial striae. convex, weakly flattened in the middle, densely Scutellum wide, back trapezoid. Elytra wide, 1.1 and finely punctured. 1st and 2nd ventrites wide.

101 Legalov A.A., Liu N.

3rd ventrite narrow. 4th ventrite narrower, rugose, thick. Antennae located in the middle of rostrum. punctured. 5th ventrite very narrow. Pygidium flat, Forehead wide, pressed, densely punctured. large and densely punctured. Legs long. Femora Eyes not protruding from contour of head. Ver- widened, densely punctured. Tibiae almost tex convex, densely punctured. Temples length- straight, thick, densely and finely punctured, ened. Antennae short, reaching as far as first line widening toward apex, little shorter than femora. of pronotum. Scapus and 1st segment of funicle Tarsi long. 1st segment long and wide. 2nd seg- wide - oval. 1st segment longer than 2nd segment. ment wide triangular. 3rd segment bilobed. Cau- 2nd and 3rd segments oval. 4th and 5th segments dal segment lengthened. Claws with long teeth. rounded. 6th segment weakly transversal. 7th seg- Length of body: 5.3 mm. ment strongly transversal, similar to segments of clava. Clava long, compact, wide, little shorter Diagnosis. This new species is similar to than funicle. 1st segment as long as 2nd seg- Byctiscophilus championi Voss, 1931 and can ment. 3rd segment longer than 2nd segment, be distinguished by the bottom covered with pointed. Pronotum weakly transversal, 1.1 times dense, long, appressed yellow setae which are wider than long, narrowed toward basis and apex. concentrated on mesepisternum, metathorax and Sides rounded. Disk convex, densely and finely sides of ventrites, almost square elytra, dark body, punctured, with medial striae. Grooves expressed. strongly lengthened and widening 1st segment Pronotal groove wrinkled. Scutellum wide, rec- of tarsi and antennae located before the middle tangular, densely punctured. Elytra almost rec- of rostrum. tangular, 1.1 times longer than wide. Humeri convex, widest in the middle of elytra. Intervals wide, Etymology. The name is formed from the word for weakly convex, densely punctured. Striae weak. “unique” – “unicus”. Points in them fine and dense. Prothorax with forward directed small teeth. Thorax, mes- and Genus Byctiscus C.G. Thomson, 1859 metepisternum with densely rugose-punctate. Abdomen convex, densely rugose-punctate, Byctiscus minimus Legalov et Liu, sp.n. (Figs. weakly flattened in the middle. 1st ventrite with 5, 9, 58) blades. 1-2nd ventrites wide. 3rd ventrite narrower. 4th ventrite narrow. 5th ventrite very nar- Holotype. Male (IZAS), China, Hebei, row. Pygidium convex, densely punctured, with Xiaowutaishan, Tatou, 1200-1600 m, 20.VIII.64, longitudinal carina in the middle. Legs long. Li Bingqian. Femora widened, densely punctured. Tibiae short, densely rugose-punctate. Protibiae straight Paratypes. 1 male (SZMN), idem; 1 female lines weakly widening toward apex. Meso- and (SZMN), Hebei, Xiaowutaishan, Huichuan, 1400 metatibiae weakly curved, more strongly widen- m, 14.VII.64, Han Yinheng; 2 females, (IZAS), ing toward apex. Tarsi long and wide. 1st seg- Hebei, Wulingshan, Liushuigou, 1200 m, ment triangular. 2nd segment wide - triangular. 04.VII.81, Yu Peiyu. 3rd segment bilobed. Caudal segment lengthened. Claws with long teeth. Length of body: Description. Pronotum and elytra green, with 4.6-4.7 mm. golden lustre. Head and pygidium golden-bronze. Edges of elytra violet. Rostrum, scutellum, tho- Female: Rostrum more weakly curved, 1.8-2.4 rax, legs, abdomen blue - green. Pronotum, elytra times longer than wide, dorsal surface of rostrum or rostrum sometimes bronze. Antennae black. weaker strongly bent. Antennae located before Body with dense, light appressed setae. the middle of rostrum. Pronotum 1.1-1.2 times wider than long. Sides of pronotum weaker Male: Rostrum short, 1.6-2 times longer than wide, rounded. Elytra 1.1-1.2 times longer than wide. widening toward apex, densely punctured, Greatest width behind the middle of elytra. curved, dorsal surface of rostrum strongly bent, Protibiae weakly curved. Length of body: 4.7-5.4 mm. 102 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China

Figs. 14-25. Leaf-rolling weevils gen.spp.: 14 – Aspidobyctiscus (Chinobyctiscus) mirabilis (male), 15 – A. (Ch.) mirabilis (female), 16 – Suniops (Neosuniops) menghunsis, 17 – Suniopsidius policoloratus (male), 18 – S. policoloratus (female), 19 – Macrosynaptopsis zhangi, 20 – Parasynatops (Pseudoeuops) tibetanus, 21 – P. (P.) pyralis (male), 22 – P. (s. str.) tzinpinensis, 23 – P. (Pseudoeuops) pyralis (female), 24 – P. (P.) lushuensis (male), 25 – P. (P.) lushuensis (female).

Diagnosis. This new species is close to Byctiscus Etymology. The name is formed from the word for fausti Sharp, 1889 but can be distinguished by “small” – “minimus”. the armament of endophallus, a densely punctured pronotum and dense setae on body. Genus Aspidobyctiscus Schilsky, 1903

Key to subgenera of genus Aspidobyctiscus Schilsky, 1903

1 (4) Pronotum punctured, as rule not wrinkled………...... ….2

2 (3) Points in striae of elytra fine, undeep, dense. Intervals of elytra flat or nearly flat. East and Southeast Asia……...... Eobyctiscus Legalov, sungen.n.

103 Legalov A.A., Liu N.

3 (2) Points in striae thin, deep, rough. Intervals of elytra convex

4 (1) Pronotum clearly wrinkled

5(6) Body dark blue. Points in striae large or fine, usually not merging. Southeast Asia…………...... …………Taiwanobyctiscus Kono

6(5) Body black, without metal lustre. Points in striae large, rough, merging. Southeast Asia…………...... …….Parabyctiscus Legalov

7 (8) Points in striae of elytra merging. Intervals of elytra densely and finely punctured. East, South- east Asia, Soenda isles……...... ………..Aspidobyctiscus s.str.

8(7) Points in striae of elytra not merging. Intervals of elytra wrinkled………...... …………..5

9(10) Body larger (6.5-7.8 mm), dark green. Nepal, Northern India…...... ….Nepalobyctiscus Legalov

10(9) Body smaller (4.9-6.5 mm). Top of body purple - red. China…Chinobyctiscus Legalov, sungen.n.

Subgenus Eobyctiscus Legalov, subgen.n. (Figs. elytra violet. Bottom of body blue - violet. Legs 6-8, 11, 60-61) with copper lustre. Antennae black.

Type species: Byctiscus coerulans Voss, 1929 Male: Rostrum long, 3 times longer than wide, widening toward apex, densely punctured, Diagnosis. This new subgenus is very close to weakly curved, thick. Antennae located in the the subgenera Taiwanobyctiscus Kâno, 1929 and middle of rostrum. Prementum with 1 long tooth. Parabyctiscus Legalov, 2003 but can be distin- Forehead wide, pressed, densely punctured. guished by the fine, shallow, dense points of Eyes not protruding from contour of head. Ver- elytral striae, and also flat or nearly flat intervals tex convex, densely rugose- punctate. Temples of elytra. lengthened. Antennae long, reaching as far as the middle of pronotum. Scapus and 1st segment Etymology. The name is formed by addition of of funicle wide - oval. 1st segment longer than the prefix “eo-” to “byctiscus”. 2nd segment. 2-4th segments oval. 5- and 6th segments rounded. 7th segment weakly transversal. Clava long, compact, little shorter than Aspidobyctiscus (Eobyctiscus) vossi Legalov et funicle. 1st segment longer than 2nd segment. Liu, sp.n. (Figs. 6-7, 60) 3rd segment longer than 2nd segment, pointed. Pronotum wide, 1.1 times wider than long. Sides Holotype. Male (IZAS), China, Zhejiang, rounded. Disk convex, densely and finely punc- Moganshan, 14.V.1935. tured, with medial striae. Grooves expressed. Postnotal groove well expressed. Pronotal Paratype. 1 female (SZMN), idem, 08.VI.1935. groove weak. Scutellum wide, rectangular, finely punctured. Elytra almost rectangular, wide, equal Description. Vertex, disk of pronotum and elytra length and width. Humeri weakly convex, widest green. Apex of rostrum, forehead, abdomen in the middle. Intervals wide, weakly convex, bronze. Sides of pronotum and elytra, pygidium densely punctured. Striae clear. Points in them golden - purple. Penultimate and last intervals of large and dense. 9th stria merges with 10th stria

104 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China

Figs. 26-41. Attelabidae gen. spp.: 26 – Riedeliops transasiaticus (male), 27 – R. transasiaticus (female), 28 – R. terminassianae, 29-30 – Orienteuops mirabilis, 31 – Sawadaeuops (s. str.) sitchuanensis (male), 32 – S. (s. str.) sitchuanensis (female), 33 – S. (s. str.) centralchinensis (male), 34 – S. (s. str.) centralchinensis (female), 35 – S. (s. str.) subelongatus (male), 36 – S. (s. str.) subelongatus (female), 37 – S. (Chinoeuops) hubeiensis (male), 38 – S. (Ch.) hubeiensis (female), 39 – S. (Ch.) nielamus (male), 40 – S. (Ch.) nielamus (female), 41 – S. (Sawadaeuopsis) punctatus. before 2nd ventrite. Prothorax with teeth forward ventrite narrower. 4th ventrite narrow. 5th ventrite directed. Mesothorax densely and strongly punc- very narrow. Pygidium convex, finely and densely tured. Mesepisternum very densely punctured. punctured. Legs long. Femora weakly widened, Metepisternum wide, strongly and densely punc- finely rugose-punctate. Protibiae almost straight, tured. Metathorax sparsely rugose-punctate. long, narrow, densely punctured. Meso- and Abdomen convex, finely and densely rugose- metatibiae weakly curved, more strongly widen- punctate, weakly flattened in the middle. 1st ing toward apex. Tarsi long. 1st segment triangu- ventrite with lamellae. 1-2nd ventrites wide. 3rd lar. 2nd segment wide - triangular. 3rd segment

105 Legalov A.A., Liu N. bilobed. Caudal segment lengthened. Claws with 1st segment longer than 2nd segment. 2-4th seg- long teeth. Length of body: 4.2 mm. ments oval. 5th and 6th segments rounded. 7th segment weakly transversal. Clava long, flat- Female: Rostrum shorter, 2.6 times longer than tened, compact, little shorter than funicle. 1st wide, almost straight, more strongly widening segment longer than 2nd segment. 3rd segment toward apex, thicker. Clava of antennae narrower. long, little shorter than 1st and 2nd segments Pronotum 1.1 times wider than long. Sides of taken together, pointed. Pronotum wide, 1.1 times pronotum weaker rounded. Elytra 1.1 times longer wider than long. Sides strongly rounded. Disk than wide. Greatest width of elytra behind the convex, sparsely and finely punctured, with very middle. Tibiae shorter and wide. Length of body: weak medial striae. Grooves clear, finely wrinkled. 4.5 mm. Scutellum wide, rectangular, finely punctured. Elytra almost rectangular, wide, 1.1 times longer Diagnosis. This new species is similar in colour- than wide. Humeri weakly smoothed. Greatest ing to Byctiscus potanini Legalov, 2004 but can width behind the middle. Intervals wide, espe- be distinguished by the merging of 9th and 10th cially near basis, weakly convex, sparsely and elytral striae, larger size and large points in striae finely punctured. Striae clear. Points in them large of elytra. It can be distinguished from and deep, decrease toward apex. 9th stria merges Aspidobyctiscus (Eobyctiscus) zhejiangensis with 10th stria before 2nd ventrite. Prothorax with- Legalov et Liu, sp.n. by the smaller punctured out teeth, wrinkled. Mesothorax densely punc- pronotum, flat, sparsely and finely punctured tured. Mesepisternum very densely punctured. forehead, eyes not protruding from contour of Metepisternum densely and finely punctured. head, long and thin rostrum, antennae located Metathorax sparsely rugose-punctate. Abdomen behind the middle of rostrum, less often punc- convex, finely rugose-punctate. 1st ventrite with tured metepisternum, strongly rounded by sides lamellae. 1-2nd ventrites wide. 3rd ventrite nar- of pronotum and large points in striae of elytra. rower. 4th ventrite narrow. 5th ventrite very nar- Etymology. This new species is named in hon- row. Pygidium convex, finely and densely punc- our of Eduard Voss. tured. Legs long. Femora weakly widened, finely rugose-punctate. Tibiae weakly curved, punctured. Tarsi long. 1st segment triangular. 2nd seg- Aspidobyctiscus (Eobyctiscus) zhejiangensis ment wide - triangular. 3rd segment bilobed. Cau- Legalov et Liu, sp.n. (Fig. 8) dal segment lengthened. Claws with long teeth. Length of body: 4.2 mm. Holotype. Female (IZAS), China, Zhejiang, Kotobuki, 11.VI.1935. Diagnosis. This new species is close to Aspidobyctiscus (Eobyctiscus) vossi Legalov et Description. Female: Body green. Apex and sides Liu, sp.n. but can be distinguished by the more of rostrum, part of 9th and 10th intervals of elytra, strongly punctured pronotum, pressed, roughly breast, abdomen and legs violet. Legs with green- punctured forehead, weakly convex eyes, short ish lustre. Rostrum long also thin, 3.7 times longer and thick rostrum, antennae located in the mid- than wide, finely and sparsely punctured, clearly dle of rostrum, thick punctured metepisternum, curved, and thinner from basis up to place of weakly rounded by sides of pronotum and fine antennal attachment, widening sharply behind points in striae of elytra. point of antennal attachment. Antennae located behind the middle of rostrum. Forehead wide, flat, Etymology. The name is formed from the name finely and sparsely punctured. Eyes do not pro- of province Zhejiang – “zhejiangensis”. trude from contour of head. Vertex convex, sparsely punctured. Temples lengthened. Anten- nae long, reaching as far as first line of pronotum. Scapus and 1st segment of funicle wide - oval.

106 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China

Figs. 42-54. Attelabidae gen. spp.: 42 – Sawadaeuops (Chinoeuops) australis (male), 43 – S. (Ch.) australis (female), 44-45 – Attelabus (s. str.) sitchuanensis, 46 – Humerilabus longulus, 47 – Lamprolabus tibetanus, 48 – Chinolabus bicolor (male), 49 – Ch. bicolor (female), 50 – Cupreuscelophilus mayongi, 51 – Lamprolabus pseudobispinosus, 52 – Henicolabus gigantinus (male), 53 – H. gigantinus (female), 54 – Chinolabus ningxianus (male).

Aspidobyctiscus (Eobyctiscus) wide, clearly curved, thick, widening toward apex, pseudocoerulans Legalov et Liu, sp.n. (Figs. 11, finely and sparsely punctured. Antennae located 61) behind the middle of rostrum. Forehead wide, weakly pressed, finely punctured. Eyes not pro- Holotype. Male (IZAS), China, Guangxi, truding from contour of head. Vertex convex, Longsheng, Baiyan, 1150 m, 21.VI.1963, Shi sparsely punctured. Temples lengthened. Anten- Yongshan. nae long, reaching the middle of pronotum. Scapus and 1st segment of funicle oval. 1st seg- Description. Male: Body dark blue, lacking lus- ment longer than 2nd segments. 2- and 3rd seg- tre naked. Rostrum long, 3.7 times longer than ments narrow - oval. 4th segment short - oval. 5-

107 Legalov A.A., Liu N. and 6th segments rounded. 7th segment trans- Paratype. 1 male (SZMN), idem, 20.VIII.1958, versal, similar to segments of clava. Clava long, Wang Shuyong. little longer than funicle, flatted. 1st segment longer than 2nd segment. 3rd segment little Description. Male: Body dark blue, lacking lus- shorter than previous segments. Pronotum wide, tre, naked. Rostrum long, 4.0-4.1 times longer than 1.2 times wider than long. Sides clearly rounded. wide, strongly curved, thick, widening toward Disk strongly convex, densely and finely punc- apex, finely and densely punctured. Antennae tured, with thin medial striae. Grooves well ex- located behind the middle of rostrum. Forehead pressed, wrinkled. Scutellum wide, rectangular, wide, weakly pressed in the middle, densely punc- densely punctured. Elytra almost rectangular, tured. Eyes not protruding from contour of head. wide, 1.2 times longer than wide. Humeri weakly Vertex convex, sparsely punctured. Temples convex, widest in the middle. Intervals wide, lengthened. Prementum almost straight. Anten- weakly convex, densely and weakly rugose-punc- nae long, reaching the middle of pronotum. tate. Striae clear. Points in them large and deep. Scapus oval, wider than first segments of funi- 9th stria merges with 10th stria before 2nd cle. 1st segment of funicle lengthened - oval, 2 ventrite. Prothorax densely punctured, with for- times longer than 2nd segment. 2nd segment short ward directed long teeth. Metepisternum and - oval. 3- and 4th segments oval. 5th segment metathorax densely and roughly rugose-punc- rounded. 6- and 7th segments transversal. Clava tate. Abdomen convex, rugose-punctate, weakly long, little longer than funicle, flattened. 1- and flattened in the middle. 1st ventrite with lamellae. 2- segments equal length. 3rd segment little 1-2nd ventrites wide. 3rd ventrite narrower. 4th shorter than previous segments, narrower, ventrite narrow. 5th ventrite very narrow. sharply asymmetric. Pronotum wide, 1.2 times Pygidium convex, densely punctured. Legs long. wider than long. Sides strongly rounded. Disk Femora weakly widened, finely rugose-punctate. strongly convex, densely and strongly punctured, Protibiae almost straight, long, narrow, densely with medial striae. Grooves well expressed, wrin- punctured. Meso- and metatibiae weakly bicon- kled. Scutellum wide, rectangular, finely punc- cave, more strongly widening toward apex. Tarsi tured. Elytra almost rectangular, of equal length long. 1st segment long - triangular. 2nd segment and width, weakly pressed for scutellum. Humeri wide - triangular. 3rd segment bilobed. Caudal weakly convex, widest in the middle. Intervals segment lengthened. Claws with long teeth. narrow, convex, finely punctured. Striae clear. Length of body: 5.6 mm. Points in them large, rough, deep. Some points in 8th stria merge. 9th stria merges with 10th stria Diagnosis. This new species is close to before 2nd ventrite. Prothorax sparsely punc- Aspidobyctiscus coerulans (Voss, 1929) but can tured, with forward directed long teeth. be distinguished by the weakly convex intervals Metepisternum and metathorax densely and of elytra, large points in striae, more sparsely roughly punctured. Abdomen convex, rugose- punctured head and wider basal sclerite of the punctate, weakly flattened in the middle. 1st endophallus. ventrite with lamellae. 1-2nd ventrites wide. 3rd ventrite narrower. 4th ventrite narrow. 5th ventrite Etymology. The name is formed by addition of very narrow. Pygidium convex, densely and the prefix “pseudo-” to “coeruleus”. strongly punctured. Legs long. Femora weakly widened, finely rugose-punctate. Protibiae almost straight, long, narrow, densely punctured. Meso- Aspidobyctiscus (Taiwanobyctiscus) and metatibiae weakly biconcave, more strongly nigrocyaneus Legalov et Liu, sp.n. widening toward apex. Tarsi long. 1st segment long - triangular. 2nd segment wide - triangular. Holotype. Male (IZAS), China, Yunnan, 3rd segment bilobed. Caudal segment length- Xishuangbanna, Meng’a, 1050-1080 m, ened. Claws with long teeth. Length of body: 09.VI.1958, Wang Shuyong. 5.3-6.0 mm.

108 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China

Figs. 55-73. Male genitalia: 55 – Chinolabus bicolor, 56 – Ch. ningxianus, 57 – Henicolabus gigantinus, 58 – Byctiscus micros, 59 – Aspidobyctiscus (Chinobyctiscus) mirabilis, 60 – A. (Eobyctiscus) vossi, 61 – A. (E.) pseudocoerulans, 62 – A. (Taiwanobyctiscus) cyanocuperus, 63-64 – Nelistrobyctiscus (Listrobyctiscidius) pseudopatruelis, 65 – Sawadaeuops (s. str.) sitchuanensis, 66 – Parasynatops (Pseudoeuops) tibetanus, 67 – Suniopsidius policoloratus, 68 – Sawadaeuops (s. str.) centralchinensis, 69 – Parasynatops (Pseudoeuops) pyralis, 70 – Riedeliops transasiaticus, 71 – Parasynatops (Pseudoeuops) lushuensis, 72 – Riedeliops terminassianae, 73 – Sawadaeuops (Chinoeuops) australis.

Diagnosis. This new species is close to Etymology. The name is formed from the words Aspidobyctiscus punctatostriatus Legalov, 2005 for “black” – “niger” and “dark blue” – but can be distinguished by the merging of points “cyaneus”. in 7- and 8th stria of elytra, and also armament of the endophallus.

109 Legalov A.A., Liu N.

Aspidobyctiscus (Taiwanobyctiscus) long. 1st segment long - triangular. 2nd segment cyanocuperus Legalov et Liu, sp.n. (Figs. 10, 62) wide - triangular. 3rd segment bilobed. Caudal segment lengthened. Claws with long teeth. Holotype. Male (IZAS), China, Yunnan, Length of body: 7.2 mm. Xishuangbanna, Meng’a, 1050-1080 m, 05.VIII.1958, Chen Zhizi. Diagnosis. This new species is very close to Aspidobyctiscus nigrocyaneus Legalov et Liu, Description. Male: Body dark blue, with weak sp.n. but can be distinguished by the more bronze lustre, naked. Rostrum long, 3.8 times strongly and densely punctured pronotum, some longer than wide, strongly curved, thick, widen- points which merge in not clear striae, bronze ing toward apex, finely and densely punctured. lustre of top pointed toward aedeagus and arma- Antennae located behind the middle of rostrum. ment of the endophallus. Forehead wide, weakly pressed in the middle, densely punctured. Eyes not protruding from Etymology. The name is formed from the words contour of head. Vertex convex, sparsely punc- for “dark blue” – “cyaneus” and “copper”- tured. Temples lengthened. Prementum almost “cupreus”. straight. Antennae long, reaching the middle of pronotum. Scapus oval, wider than first segments of funicle. 1st segment of funicle lengthened - Subgenus Chinobyctiscus Legalov, subgen.n. oval, 2 times longer than 2nd segment. 2nd seg- (Figs. 14-15, 59) ment short - oval. 3rd and 4th segment oval. 5th segment rounded. 6th and 7th segments trans- Type species: Aspidobyctiscus mirabilis Legalov versal. Clava long, little longer than funicle, flat- et Liu, sp.n. tened. 1st and 2nd segments of equal length. 3rd segment little shorter than previous segments, Description. Body black. Pronotum and elytra narrower, sharply asymmetric. Pronotum wide, 1.1 purple - red. Forehead and vertex bronze. Body times wider than long. Sides strongly rounded. with dense, light, appressed setae. Rostrum long, Disk strongly convex, densely and strongly punc- shorter and thick in females, thin from basis up tured, some points merges in not clear striae, with to place of antennal attachment, densely punc- medial striae. Grooves well expressed, wrinkled. tured. Antennae located in the middle of rostrum. Scutellum wide, rectangular, finely punctured. Prementum semicircular, not pointed. Forehead Elytra almost rectangular, equal length and width, wide, strongly pressed, densely punctured. Ver- weakly pressed for scutellum. Humeri weakly tex convex, finely rugose-punctate. Temples convex, widest in the middle. Intervals narrow, lengthened. Antennae thick, long, reaching the convex, finely punctured. Striae clear. Points in middle of pronotum. Pronotum wide. Sides them large, rough, deep. Some points merges in rounded, less so in females. Disk strongly con- 8th stria. 9th stria merges with 10th stria before vex, densely rugose-punctate, with thin medial 2nd ventrite. Prothorax sparsely punctured, with striae. Grooves well expressed. Scutellum wide, forward-directed long teeth. Metepisternum and rectangular, densely punctured. Elytra almost metathorax densely and roughly punctured. Ab- rectangular, wide. Humeri weakly smoothed. In- domen convex, rugose-punctate, weakly flattened tervals wide, convex, very densely rugose-punc- in the middle. 1st ventrite with lamellae. 1-2nd tate. Striae clear. Points in them large and dense. ventrites wide. 3rd ventrite narrower. 4th ventrite Prothorax wrinkled, with long teeth forward di- narrow. 5th ventrite very narrow. Pygidium con- rected in males. Metepisternum and metathorax vex, densely and strongly punctured. Legs long. densely and roughly rugose-punctate. Abdomen Femora weakly widened, finely rugose-punctate. convex, thin rugose-punctate, weakly flattened Protibiae almost straight, long, narrow, densely in the middle. 1st ventrite with lamellae. Pygidium punctured. Meso- and metatibiae weakly bicon- convex, densely and strongly punctured. Legs cave, more strongly widening toward apex. Tarsi long. Femora weakly widened, finely rugose-

110 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China punctate. Protibiae almost direct in males and segment rounded. 6th segment transversal. 7th weakly curved in females, long, narrow, densely segment sharply transversal, similar to segments punctured. Meso- and metatibiae weakly bicon- of clava. Clava long, little shorter than funicle, cave, more strongly widening toward apex. Tarsi flattened. 1st and 2nd segments of equal length. long. 1st segment long - triangular. 2nd segment 3rd segment longer than 2nd segment, pointed, wide - triangular. 3rd segment bilobed. Caudal narrower. Pronotum wide, 1.1-1.2 times wider than segment lengthened. Claws with long teeth. long. Sides clearly rounded. Disk strongly con- Length of body: 4.9-6.5 mm. vex, densely rugose-punctate, with thin medial striae. Grooves well expressed. Scutellum wide, Diagnosis. This new subgenus is close to the rectangular, densely punctured. Elytra almost subgenus Nepalobyctiscus Legalov, 2003 but rectangular, wide, 1.1 times longer than wide. can be distinguished by the form of basal sclerite Humeri weakly smoothed, widest in the middle. of endophallus, purple - red top of body and Intervals wide, convex, very densely rugose- smaller size. From subgenus Aspidobyctiscus s. punctate. Striae clear. Points in them large and str. it differs by non-merging points in striae of dense. 9th stria merges with 10th stria before 2nd elytra and armament of the endophallus. ventrite. Prothorax wrinkled, with forward directed long teeth. Metepisternum and metathorax Etymology. The name is formed from the words densely and roughly rugose-punctate. Abdomen “Chinese” and “byctiscus”. convex, thin rugose-punctate, weakly flattened in the middle. 1st ventrite with lamellae. 1-2nd ventrites wide. 3rd ventrite narrower. 4th ventrite Aspidobyctiscus (Chinobyctiscus) mirabilis narrow. 5th ventrite very narrow. Pygidium con- Legalov et Liu, sp.n. (Figs. 14-15, 59) vex, densely and strongly punctured. Legs long. Femora weakly widened, finely rugose-punctate. Holotype. Male (IZAS), China, Sichuan, Wanxian, Protibiae almost straight, long, narrow, densely Wang’erbao, 1200 m, 29.IX.1994, Li Fasheng. punctured. Meso- and metatibiae weakly biconcave, more strongly widening toward apex. Tarsi Paratypes. 1 male (SZMN), idem; 1 male (IZAS), long. 1st segment long - triangular. 2nd segment Hubei, Shennongjia, Dajiuhu, 1800 m, wide - triangular. 3rd segment bilobed. Caudal 14.VIII.1981, Han Yinheng; 1 female (SZMN), segment lengthened. Claws with long teeth. Hubei, Shennongjia, 900-1200 m, 27.VI.1981, Han Length of body: 4.9-6.1 mm. Yinheng; 1 female (IZAS), “China”, V.1987. Female: Rostrum shorter and thick, 3 times longer Description. Body black. Pronotum and elytra than wide, more weakly curved. Antennae shorter purple - red. Forehead and vertex bronze. Body and thin. Pronotum 1.2 times wider than long. with dense, light, appressed setae. Sides of pronotum weaker rounded. Elytra wide, 1.2 times longer than wide. Greatest width of Male. Rostrum long, 3.3-4.7 times longer than elytra behind the middle. Protibiae weakly bicon- wide, curved, widening toward apex, thin from cave. Abdomen stronger convex. Length of body: basis up to place of antennal attachment, densely 6.2-6.5 mm. punctured. Antennae located in the middle of rostrum. Prementum semicircular, not pointed. Diagnosis. This new species is similar to Forehead wide, strongly pressed, densely punc- Aspidobyctiscus (s. str.) lacunipennis (Jekel, tured. Eyes not protruding from contour of head. 1860) but can be distinguished by non-merging Vertex convex, finely rugose-punctate. Temples points in striae of elytra, wrinkled intervals, and lengthened. Antennae thick, long, reaching the also armament of the endophallus. middle of pronotum. Scapus and 1-4th segments of funicle oval. 2nd and 3rd segments narrower. Etymology. The name is formed from the word for 1st segment little shorter than 2nd segment. 5th “surprising” – “mirabilis”.

111 Legalov A.A., Liu N.

Aspidobyctiscus (Parabyctiscus) konoi Legalov 2003 but can be distinguished by the sparsely et Liu, sp.n. (Fig. 12) strongly punctured metepisternum and points of pronotum merging in striae. Holotype. Female (IZAS), China, Yunnan, Yuelong, Zhibenshan, 2430 m, 24.VI.1981, Wang Etymology. This new species is named in honour Shuyong. of Hiromichi Kono.

Description. Female: Body black, lacking lustre, covered with dense, light, appressed setae. Ros- Nelistrobyctiscus (Listrobyctiscidius) trum long, 3 times longer than wide, widening pseudopatruelis Legalov et Liu, sp.n. (Figs. 13, toward apex, densely punctured, curved. Anten- 63-64) nae located in the middle of rostrum. Forehead wide, strongly pressed, densely and finely punc- Holotype. Male (IZAS), China, Liaoning, Changtu, tured. Eyes not protruding from contour of head. Quantou, 12.VI.1963, Li Hongxing. Vertex convex, finely rugose-punctate. Temples little lengthened. Antennae long, reaching as far Description. Male: Body black, naked, lacking as first line of pronotum, thick. Scapus and 1-3rd lustre. Rostrum short, 3.3 times longer than wide, segments of funicle oval. 1st segment as long as widening toward apex, roughly and densely 2nd segment. 4th segment short - oval. 5- and punctured, weakly curved, thick. Antennae lo- 6th segments rounded. 7th segment transversal, cated in the middle of rostrum. Forehead wide, similar to segments of clava. Clava long, wide, as flat, finely doubly punctured. Eyes not protrud- long as funicle.1st segment as long as 2nd seg- ing from contour of head. Vertex convex, finely ment. 3rd segment longer than 2nd segment, nar- doubly punctured. Temples little lengthened. rower, pointed. Pronotum wide, 1.1 times wider Antennae long, reaching as far as the middle of than long, narrowed toward apex. Sides rounded. pronotum. Scapus and 1st segment of funicle Grooves clear, wrinkled. Disk convex, densely and oval. 1st and 2nd segments of equal length. 2- roughly rugose-punctate, with deep medial striae 4th segments narrow - oval. 5- and 6th segments and 2 depression on each side. Scutellum wide, rounded. 7th segment transversal. Clava long, finely punctured. Elytra almost square, 1.1 times wide, little shorter than funicle. 1st segment as longer than wide. Greatest width behind the mid- long as 2nd segment. 3rd segment longer than dle. Humeri weakly smoothed. Intervals convex, 2nd segment. Pronotum lengthened, equal length narrow, finely punctured. Points in striae deep, and width. Sides weakly rounded. Disk strongly large, lengthened, from sides of elytra frequently convex, lacking lustre, densely and finely punc- merges with each other. 9th stria merges with 10th tured. Pronotal groove expressed. Scutellum stria in the middle. Prothorax finely and densely wide, rectangular, punctured. Elytra almost rec- wrinkled. Sides of prothorax without teeth. tangular, 1.3 times longer than wide. Humeri Metepisternum wide, sparsely and finely punc- weakly smoothed, widest in the middle. Intervals tured. Metathorax densely rugose-punctate. wide, weakly convex, very sparsely punctured. Abdomen convex, densely rugose-punctate, Striae deep. Points in them large and dense, de- weakly flattened in the middle. Pygidium convex, crease toward apex of elytra. 9th stria ends be- densely punctured. Legs long. Femora weakly fore abdomen and does not merge with 10th stria. widened. Tibiae weakly biconcave, widening to- Prothorax with forward directed small teeth. ward apex. Tarsi long. 1st segment triangular. 2nd Mesepisternum wrinkled. Metepisternum densely segment wide - triangular. 3rd segment bilobed. and strongly punctured. Metathorax sparsely Caudal segment lengthened. Claws with long and strongly punctured. Abdomen convex, thin teeth at basis. Length of body: 5.8 mm. rugose-punctate. 1st ventrite with lamellae. 1- and 2nd ventrites wide. 3rd ventrite narrower. 4th Diagnosis. This new species is close to ventrite narrow. 5th ventrite very narrow. Aspidobyctiscus (Parabyctiscus) niger Legalov, Pygidium convex, sparsely and strongly punc-

112 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China tured. Legs long. Femora widened, without teeth, Sides almost straight. Disk convex, with dense finely and sparsely wrinkled. Tibiae long, almost transversal rugose-punctate. Basis of pronotum straight, densely punctured, little widening to- extended. Scutellum almost rectangular, smooth. ward apex. Meso- and metatibiae shorter, than Elytra almost rectangular, 1.1 times longer than protibiae, more strongly widening toward apex. wide. Humeri convex. Greatest width in humeri Tarsi long. 1st segment lengthened - triangular. and in the middle. Intervals wide, convex, wrin- 2nd segment triangular. 3rd segment bilobed. kled. Striae clear. Points in them large and deep. Caudal segment long. Claws with long teeth. 9th stria merges with 10th stria before 1st Length of body: 2.6 mm. ventrite. Prothorax punctured. Mesothorax and mesepisternum matte. Metathorax and Diagnosis. This new species is close to metepisternum strongly and densely punctured. Nelistrobyctiscus (Listrobyctiscidius) patruelis Abdomen convex, densely rugose-punctate, (Voss, 1921) but can be distinguished by the more weakly flattened in the middle. 1-3rd ventrites strongly rounded sides of pronotum, flat inter- wide. 4th ventrite narrower. 5th ventrite narrow. vals of elytra and armament of the endophallus. 1-3rd ventrites with 2 lines of erecting setae. Etymology. The name is formed by addition of Pygidium convex, densely and strongly punc- the prefix “pseudo-“ to “patruelis”. tured. Legs long. Femora widened, with small teeth. Protibiae wide, weakly curved; biconcave at internal edge, with and uncus at apex. Family Attelabidae Billberg, 1820 Mesotibiae weakly biconcave, more strongly Subfamily Attelabinae Billberg, 1820 widening toward apex, shorter. Metatibiae bicon- Tribe Euopsini Voss, 1925 cave, longer. Tarsi lengthened, little shorter than Genus Suniops Voss, 1928 tibiae. 1st segment long. 2nd segment long - triangular, flat. 3rd segment wide - bilobed. Caudal Suniops menghunsis Legalov et Liu, sp.n. (Fig. segment lengthened. Claws long. Length of body: 16) 2.8 mm.

Holotype. Female (IZAS), China, Yunnan, Diagnosis. This new species is close to Suniops Xishuangbanna, Menghun, 750 m, 01.VI.1958, gorochovi Legalov, 2003 but can be distin- Hong Chunpei. guished by the different colouring of the body, rougher sculpture of the pronotum, strongly con- Description. Body brown. Head, pronotum and vex striae of the elytra, large points in elytral striae abdomen black - brown. and wide tarsi.

Female: Rostrum short, straight, widening toward Etymology. The name is formed from the location apex, finely and sparsely punctured. Antennae “Menghun” – “menghunsis”. located at basis of rostrum. Forehead very narrow, matte. Eyes large, weakly convex. Vertex convex, gently wrinkled and sparsely strongly Genus Suniopsidius Legalov, gen.n. (Figs. 17- punctured. Temples weakly lengthened, wrinkled. 18, 67) Prementum with 3 teeth. Antennae long, reaching as far as the middle of pronotum. Scapus and Type species: Suniopsidius multicoloratus 1st segment wide - oval. 1st segment longer than Legalov et Liu, sp.n. 2nd segment. 2-3rd segments long - oval. 4-6th segments short - oval. 7th segment rounded. Description. Body brown. Pronotum bronze. Clava little shorter than funicle, pointed. 1st seg- Head, rostrum, scutellum dark. Femora, ment as long as 3rd segment. 2nd segment shorter mesothorax, metathorax, mes-, metepisternum than 1st segment. Pronotum almost companiform, green. Abdomen and pygidium golden-green. 1.1 times wider than long. Pronotal groove clear. Rostrum weakly curved, long in males and short

113 Legalov A.A., Liu N. in females, longer than head, smooth, widens Suniopsidius multicoloratus Legalov et Liu, toward apex. Antennae located near basis of ros- sp.n. (Figs. 17-18, 67) trum. Forehead very narrow, almost reduced. Eyes large, weakly convex. Vertex flattened, with Holotype. Male (IZAS), China, Yunnan, dense transversal rugose-punctate. Temples wrin- Xishuangbanna, Damenglong, 650 m, 19.IV.1958, kled, lengthened of males. Prementum with 3 Chen Zhizi. teeth. Antennae short, reaching as far as first line of pronotum. Clava long, little longer than Paratype. Female (SZMN), Yunnan, Nannoshan, funicle, pointed. 1st segment as long as 3rd seg- Fokhaja, 1400 m, 4.III.1957, Lju Da-xua. ment. 2nd segment shorter than 1st segment. Pronotum almost companiform. Pronotal groove Description. Body brown. Pronotum bronze. clear. Sides almost straight. Disk convex, with Head, rostrum, scutellum dark. Femora, dense transversal rugose-punctate. Basis of mesothorax, metathorax, mes-, metepisternum pronotum extended. Scutellum almost rectangu- green. Abdomen and pygidium golden - green. lar, impressed on centre, punctured. Elytra almost Male: Rostrum weakly curved, long, longer than rectangular. Humeri weakly smoothed. Greatest head, smooth, widens toward apex. Antennae width in humeri and in the middle. Intervals wide, located before basis of rostrum. Forehead very convex, smooth. Striae clear, deep. Points in them narrow, almost reduced. Eyes large, weakly con- large, in first third of elytra deep. 9th stria merges vex. Vertex flattened, with dense transversal ru- with 10th stria before metacoxae. Thorax doubly gose- punctate. Temples lengthened, wrinkled. punctured: finely, and also strongly and sparsely Prementum with 3 teeth. Antennae short, reach- punctured. Abdomen convex, finely and sparsely ing as far as first line of pronotum. Scapus and punctured, flattened in the middle of males. 1-3rd 1st segment wide - oval. 2-4th segments narrow - ventrites wide. 4th ventrite narrower. 5th ventrite oval. 5-7th segments short - oval. Clava long, narrow. 1-3rd ventrites with 2 lines of erect setae little longer than funicle, pointed. 1st segment as of females. Pygidium convex, densely and finely long as 3rd segment. 2nd segment shorter than punctured. Legs long. Forward legs lengthened 1st segment. Pronotum almost companiform, 1.1 in males. Femora widened, without teeth. times wider than long. Pronotal groove clear. Protibiae long in males, hardly shorter than Sides almost straight. Disk convex, with dense profemora, thin, apexes of tibiae with long mucro, transversal rugose-punctate. Basis of pronotum shorter in females, almost straight, very weakly extended. Scutellum almost rectangular, im- biconcave at internal edge. Tibiae with mucro pressed on centre, punctured. Elytra almost rec- and uncus in females. Meso- and metatibiae tangular, 1.1 times longer than wide. Humeri weakly biconcave, shorter. Tarsi lengthened, lit- weakly smoothed. Greatest width in humeri and tle shorter than tibiae. 1st segment long. 2nd seg- in the middle. Intervals wide, convex, smooth. ment long - triangular, flat. 3rd segment wide - Striae clear, deep. Points in them large; deep in bilobed. Caudal segment lengthened. Claws long. first third of elytra. 9th stria merges with 10th Length of body: 2.5-2.6 mm. stria before metacoxae. Thorax doubly punctured: finely, and also strongly and sparsely punctured. Diagnosis. This new genus is close to genus Abdomen convex, finely and sparsely punctured, Suniops Voss, 1928 but can be distinguished by flattened in the middle. 1-3rd ventrites wide. 4th the absence of teeth on femora. ventrite narrower. 5th ventrite narrow. Pygidium convex, densely and finely punctured. Legs long. Etymology. The name is formed by addition of Forward legs lengthened. Femora widened, with- the ending “-idius”to “suniops”. out teeth. Protibiae long, hardly shorter than profemora, thin, apexes of tibiae with long mucro. Meso- and metatibiae weakly biconcave, shorter. Tarsi lengthened, little shorter than tibiae. 1st

114 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China segment long. 2nd segment long - triangular, flat. densely punctured. Abdomen convex, rugose- 3rd segment wide - bilobed. Caudal segment punctate. 1st and 2nd ventrites wide. 3rd and 4th lengthened. Claws long. Length of body: 2.6 mm. ventrites narrower. 5th ventrite very narrow. 1- Female: Rostrum shorter, more strongly widen- 3rd ventrites with 2 lines of setae, 4th ventrite ing. Antennae located at basis of rostrum. Eyes with one line in females. Pygidium convex, finely hardly more small. Temples weakly lengthened. and densely punctured. Legs long. Femora wid- Pronotum 1.1 times wider than long. Elytra 1.3 ened, finely rugose-punctate. Protibiae narrow times longer than wide. Abdomen stronger con- in females, almost straight, at internal edge vex. 1-3rd ventrites with 2 lines of erecting setae. sharply biconcave, with long mucro and uncus Forward legs not lengthened. Tibiae with mucro at apex. Meso- and metatibiae biconcave, shorter and uncus. Protibiae shorter, almost straight, very and thick. Tarsi lengthened. Length of body: 4.7 weakly biconcave at internal edge. Length of mm. body: 2.5 mm. Diagnosis. This new genus is close to genus Diagnosis. This new species differs from other Parasynaptopsis Legalov, 2003 but can be dis- species of the subtribe Suniopsina Legalov, 2003 tinguished by the pretopmost protuberance of from Asia by femora without teeth, colouring of the elytra. the body and armament of the endophallus. Etymology. The name is formed by addition of Etymology. The name is formed from the word for the prefix “large” – “macro-” to “synaptopsis”. “multi-colour” – “multicolor”.

Macrosynaptopsis zhangi Legalov et Liu, sp.n. Genus Macrosynaptopsis Legalov, gen.n. (Fig. (Fig. 19) 19) Holotype. Female (IZAS), China, Fujian, Type species: Macrosynaptopsis zhangi Legalov Jianyang, Huangkeng, Guilin, 270-590 m, et Liu, sp.n. 07.IV.1960, Zuo Yong.

Description. Body darkly brown, with bronze lus- Description. Female: Body darkly brown, with tre. Rostrum short, straight, widening toward bronze lustre. Rostrum short, straight, widening apex, densely and strongly punctured, with weak toward apex, densely and strongly punctured, carina in first half. Forehead narrow, matte. Eyes with weak carina in first half. Antennae located large, weakly convex. Vertex flattened, with dense at basis of rostrum. Forehead narrow, matte. Eyes transversal rugosity. Temples weakly lengthened, large, weakly convex. Vertex flattened, with dense densely wrinkled. Antennae short. Clava long and transversal rugosity. Temples weakly lengthened, wide, little shorter than funicle, pointed. densely wrinkled. Antennae short. Scapus and Pronotum companiform, weakly narrowed toward 1st segment oval. 1st segment as long as 2nd apex. Grooves clear. Sides weakly rounded. Disk segment. 2-3rd segments narrow - oval. 4th seg- convex, densely rugose-punctate, with transver- ment short - oval. 5th segment shorter. 6th seg- sal striae and protuberance behind the middle. ment rounded. 7th segment transversal. Clava Scutellum almost rectangular, punctured. Elytra long and wide, little shorter than funicle, pointed. almost rectangular, lengthened, pressed against 1st segment as long as 2nd segment. 3rd seg- scutellum. Humeri convex. Greatest width in hu- ment longer than 2nd segments, pointed. meri. Intervals wide, convex, roughly wrinkled. Pronotum companiform, weakly narrowed toward 1st, 3rd, 5th and 6th intervals carinate at basis. apex, 1.1 times wider than long. Grooves clear. Protuberance at basis of 7th interval. 5th interval Sides weakly rounded. Disk convex, densely ru- with pretopmost protuberance. Striae clear. Tho- gose-punctate, with transversal striae and pro- rax and metepisternum roughly, strongly and tuberance behind the middle. Basis of pronotum

115 Legalov A.A., Liu N. extended. Scutellum almost rectangular, punc- convex. Vertex convex, sparsely and finely punctured. Elytra almost rectangular, lengthened, 1.1 tured. Temples weakly lengthened. Antennae times longer than wide. Humeri convex. Greatest long, reaching as far as the middle of pronotum. width in humeri. Intervals wide, convex, roughly Scapus and 1st segment wide - oval. 2-3rd seg- wrinkled. 1st, 3rd, 5th and 6th intervals carinate ments long - oval. 4-6th segments short - oval. at basis. Protuberance at basis of 7th interval. 7th segment rounded. Clava long, narrow, little 5th interval with pretopmost protuberance. Striae shorter than funicle, and pointed. 1st segment as clear. Points in them dense and deep. 9th stria long as 2nd segment. 3rd segment little shorter merges with 10th stria before abdomen. Thorax than previous segments. Pronotum companiform, and metepisternum roughly, strongly and densely length and width almost equal. Pronotal groove punctured. Abdomen convex, rugose-punctate. well expressed. Sides almost straight. Disk con- 1st and 2nd ventrites wide. 3rd and 4th ventrites vex, sparsely and finely punctured, with very weak narrower. 5th ventrite very narrow. 1-3rd ventrites medial transversal striae. Basis of pronotum ex- with 2 lines of setae. 4th ventrite with one line of tended. Scutellum back trapezoid, lengthened, setae. Pygidium convex, finely and densely punc- finely punctured. Elytra lengthened, 1.2 times tured. Legs long. Femora widened, finely rugose- longer than wide, almost rectangular. Humeri punctate. Protibiae narrow, almost straight, at slightly smoothed. Greatest width in humeri and internal edge sharply biconcave, with long mucro in the middle. Intervals wide, flat, smooth. Striae and uncus at apex. Meso- and metatibiae bicon- clear. Points in them large, decrease toward apex. cave, shorter and thick. Tarsi lengthened, little 9th stria merges with 10th stria before metacoxae. shorter than tibiae. 1st segment lengthened. 2nd Mesothorax and mesepisternum matte. segment long - triangular, flat. 3rd segment wide Metathorax densely and strongly punctured. - bilobed. Caudal segment lengthened. Claws Metepisternum sparsely and strongly punctured. long. Length of body: 4.7 mm. Abdomen convex, sparsely and finely punctured. 1st and 2nd ventrites wide. 3rd and 4th ventrites Diagnosis. This new species is similar to narrower. 5th ventrite very narrow. 1-3rd ventrites Parasynaptopsis cuprifulgens (Voss, 1942) but with 2 lines of setae. 4th ventrite with 1 line of can be distinguished by the pretopmost protu- setae. Pygidium convex, densely and strongly berance of the elytra, rough sculpture and large punctured. Legs long. Femora widened, finely size. and sparsely punctured. Protibiae almost straight, finely and sparsely punctured, with Etymology. This new species is named in honour mucro and uncus at apex. Meso- and metatibiae of Runzhi Zhang. biconcave, shorter and thick. Tarsi lengthened, little shorter than tibiae. 1st segment lengthened. 2nd segment long - triangular, flat. 3rd segment Genus Parasynatops Legalov, 2003 wide - bilobed. Caudal segment lengthened. Claws long. Length of body: 3.2 mm. Parasynatops (s. str.) tzinpinensis Legalov et Liu, sp.n. (Fig. 22) Diagnosis. This new species is close to Parasynatops (s. str.) politus (Roelofs, 1874) but Holotype. Female (IZAS), China, SE Yunnan, can be distinguished by the brown body, Tzinpin, 1700 m, 25.V.1956, Panfilov. pronotum without transversal wrinkles and a narrower forehead. Description. Female: Body brown. Head, pronotum, metepisternum, abdomen, pygidium Etymology. The name is formed from the location dark brown. Rostrum short, wide, straight, wid- “Tzinpin” – “tzinpinensis”. ening toward apex, sparsely and finely punctured. Antennae located at basis of rostrum. Prementum with 3 teeth. Forehead narrow. Eyes large, weakly

116 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China

Parasynatops (Pseudoeuops) tibetanus Legalov Diagnosis. This new species is close to et Liu, sp.n. (Figs. 20, 66) Parasynatops (Pseudoeuops) pyralis Legalov et Liu, sp.n. but can be distinguished by the green Holotype. Male (IZAS), China, Tibet, Yigong, body, narrower forehead, thick punctured ros- 2300 m, 16-17.VIII.1982, Lin Zai. trum, thick and rougher punctured pronotum, elytra widest in the middle, weakly smoothed Description. Male: Body bright green. Legs and humeri, intervals without lines of large points, abdomen with bronze lustre. Rostrum purple. longer tarsi and armament of the endophallus. Antennae and tarsi black. Rostrum short, straight, sparsely and finely punctured. Antennae located Etymology. The name is formed from the location at basis of rostrum. Forehead flat, narrow, densely “Tibet” – “tibetanus”. punctured. Eyes large, weakly convex. Vertex convex, densely doubly punctured. Temples weakly lengthened. Antennae long, reaching as Parasynatops (Pseudoeuops) pyralis Legalov et far as the middle of pronotum. Scapus and 1st Liu, sp.n. (Figs. 21, 23, 69) segment wide - oval. 1st segment longer than scapus. 2-3rd segments long narrow - oval. 1st Holotype. Male (IZAS), China, Yunnan, Weixi, segment as long as 2nd segment. 3rd segment 3400 m, 13.VIII.1984, Wang Shuyong. shorter than 2nd segment. 3-5th segments narrow - oval. 6th segment oval. 7th segment Paratypes. 4 males (IZAS), 2 females (IZAS), rounded. Clava wide, shorter than funicle, Yunnan, Deqin, Baimangxueshang, 3700 m, 25- pointed. 1st segment as long as 3rd segment and 26.VIII.1981, Fang Subai, Wang Shuyong; 3 males longer than 2nd segment. Pronotum almost (IZAS), 3 males (SZMN), 5 females (IZAS), 2 fe- companiform, 1.2 times wider than long. Grooves males (SZMN), Yunnan, Weixi, 3200-3400 m, 13- clear. Postnotal groove weak. Sides almost 17.VIII.1984, Wang Shuyong, Fan Jianguo; 1 fe- straight. Disk convex, densely and strongly dou- male (IZAS), Yunnan, Deqin, Meilixueshang, bly punctured, without medial striae. Basis of 3200-3500 m, 23.VII.1982, Wang Shuyong. pronotum extended. Scutellum back trapezoid, doubly punctured, smooth in the middle. Elytra Description. Body bronze. Antennae and tarsi lengthened, 1.1 times longer than wide. Humeri black. weakly smoothed, widest in the middle. Intervals wide, weakly convex, wrinkled and doubly punc- Male: Rostrum short, straight, sparsely and finely tured. Striae clear. Points in them rough and large, punctured. Antennae located at basis of rostrum. decrease toward apex. 9th stria merges with 10th Forehead flat, narrow, but wider than in stria before metacoxae. Prothorax and Parasynatops (Pseudoeuops) tibetanus Legalov metepisternum densely doubly punctured. et Liu, sp.n., densely punctured. Eyes large, Metathorax more sparsely doubly punctured. weakly convex. Vertex convex, strongly doubly Abdomen convex, densely and strongly doubly punctured. Temples weakly lengthened. Anten- punctured, weakly flattened and almost smooth nae long, reaching as far as the middle of in the middle. 1-4th ventrites wide. 5th ventrite pronotum. Scapus and 1st segment wide - oval. narrow, densely punctured. Pygidium densely 1st segment longer than scapus. 2-3rd segments and weakly doubly punctured. Legs long. Femora long narrow - oval. 1st segment as long as 2nd widened, doubly punctured. Protibiae almost segment. 3rd segment shorter than 2nd segment. straight, with sharp external edge, doubly punc- 3-5th segments narrow - oval. 6th segment oval. tured, shorter than femora, with mucro at apex. 7th segment rounded. Clava wide, shorter than Meso- and metatibiae biconcave, thicker. Tarsi funicle, pointed. 1st segment as long as 3rd seg- long, little shorter than tibiae. 1st segment long. ment and longer than 2nd segment. Pronotum 2nd segment long - triangular, flat. 3rd segment almost companiform, 1.3 times wider than long. wide - bilobed. Caudal segment lengthened. Claws Grooves clear. Postnotal groove weak. Sides long. Length of body: 3.1 mm. weakly rounded. Disk convex, strongly doubly 117 Legalov A.A., Liu N. punctured, points transversal lengthened, with- Parasynatops (Pseudoeuops) lushuensis Legalov out the medial striae, weakly pressed in the mid- et Liu, sp.n. (Figs. 24-25, 71) dle. Basis of pronotum extended. Scutellum back trapezoid, wide, doubly punctured, smooth in the Holotype. Male (IZAS), China, Yunnan, Lushui, middle. Elytra lengthened, 1.2-1.3 times longer 2400 m, 10.VI.1981, Liao Subai. than wide. Humeri strongly convex. Greatest width in humeri and in the middle. Intervals wide, Paratypes. 2 males (IZAS), 2 males (ZSMN), 1 weakly convex, wrinkled and doubly punctured, female (IZAS), Sichuan, Lushui, 2150-2400 m, 10- with line of large points. Striae clear. Points in 11.VI.1981, Wang Shuyong, Liao Subai. them rough and large, decrease toward apex. 9th stria merges with 10th stria before metacoxae. Description. Body bronze -golden, meso- and Prothorax and metepisternum densely doubly metathorax with episternums brown with golden punctured. Metathorax more sparsely doubly lustre, or body green with brown - bronze legs. punctured. Abdomen convex, densely and Male: Rostrum short, straight, strongly widen- strongly doubly punctured, weakly flattened and ing toward apex, sparsely and finely punctured. almost smooth in the middle. 1-4th ventrites wide. Antennae located at basis of rostrum. Forehead 5th ventrite narrow, densely punctured. Pygidium flat, very narrow. Eyes large, convex. Vertex flat- densely and finely doubly punctured. Legs long. tened, roughly punctured. Temples weakly Femora widened, doubly punctured. Protibiae lengthened. Antennae long, reaching as far as curved in topmost part, with sharp external edge, the middle of pronotum. Scapus and 1st segment doubly punctured, shorter than femora, with wide - oval. 1st segment longer than scapus. 2- mucro at apex. Meso- and metatibiae biconcave, 3rd segments long narrow - oval. 1st segment as thicker. Mesotibiae without appendages. Tarsi long as 2nd segment. 3rd segment shorter than lengthened, little shorter than tibiae. 1st segment 2nd segment. 3-5th segments narrow - oval. 6th long. 2nd segment long - triangular, flat. 3rd seg- segment oval. 7th segment rounded. Clava nar- ment wide - bilobed. Caudal segment lengthened. row, little shorter than funicle, pointed. 1st seg- Claws long. Length of body: 2.5-3.3 mm. ment longer than 2nd segment. 3rd segment longer than 1st segment, pointed. Pronotum al- Female: Pronotum 1.3 times wider than long. most companiform, weakly narrowed toward apex, Elytra 1.2 times longer than wide. 1-3rd ventrites 1.1-1.2 times wider than long. Postnotal groove with 2 lines of erecting setae. Tibiae with mucro not expressed. Pronotal groove weak. Sides al- and uncus. Protibiae shorter and wide. Length of most straight. Disk convex, with dense transver- body: 2.8-3.1 mm. sal rugose-punctate. Basis of pronotum extended. Scutellum almost rectangular, lengthened, Diagnosis. This new species is close to sparsely and finely punctured. Elytra lengthened, Parasynatops (Pseudoeuops) tibetanus Legalov 1.2 times longer than wide. Humeri strongly con- et Liu, sp.n. but can be distinguished by the vex. Greatest width in humeri. Intervals wide, flat, bronze body, wider forehead, less often punc- wrinkled, with line of points. Striae clear. Points tured rostrum, less often and more gently punc- in them dense and deep. 9th stria merges with tured pronotum, elytra widest at humeri and in 10th stria before metacoxae. Mesothorax and the middle, convex humeri, intervals with line of mesepisternum densely wrinkled. Metepisternum large points, shorter tarsi and armament of the narrow, strongly and densely punctured. Abdo- endophallus. From Parasynatops (Pseudoeuops) men convex, strongly and sparsely doubly punc- moanus Legalov, 2003, it differs by the metallic tured, weakly flattened and almost smooth in the colouring, narrower body and armament of the middle. 1-4th ventrites wide. 5th ventrite narrow, endophallus. densely punctured. Pygidium finely and not so densely punctured. Legs long. Femora widened, Etymology. The name is formed from the word for gently rugose-punctate. Tibiae narrow, thin, “fiery” – “pyra”. weakly curved, with long mucro at apex. Meso-

118 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China and metatibiae biconcave, thicker. Mesotibiae Pronotum almost trapezoid, equal length and with small appendages. Tarsi lengthened, little width. Sides very weakly rounded. Disk convex, shorter than tibiae. 1st segment long. 2nd seg- sparsely and finely punctured. Postnotal groove ment long - triangular, flat. 3rd segment wide - weak. Scutellum almost rectangular, small. Elytra bilobed. Caudal segment lengthened. Claws long. almost rectangular, 1.1-1.2 times longer than wide, Length of body: 2.9 mm weakly narrowed toward apex. Humeri well advanced. Intervals wide, flat, smooth. Points in Female: Rostrum wider. Antennae shorter. Eyes striae large and frequent. 9th stria merges with hardly more small. Pronotum 1.1 times wider than 10th stria before metacoxae. Metepisternum wide, long. Elytra 1.2 times longer than wide. 1-3rd lacking lustre, densely punctured. Abdomen con- ventrites with 2 lines of erecting setae. Tibiae vex, gently rugose-punctate, flatted, but without with mucro and uncus. Protibiae shorter and wide. lines of setae. 1-4th ventrites wide. 5th ventrite Length of body: 3.2 mm. narrow. Pygidium convex, punctured. Legs long, strong, forward not lengthened. Femora clavate. Diagnosis. This new species is close to Protibiae weakly curved, wide, with mucro. Parasynatops (Pseudoeuops) bicoloroides Mesotibiae biconcave, thick, from external edge Legalov, 2003 but can be distinguished by the in topmost third with short and wide append- colouring of body, armament of the endophallus, ages. Metatibiae biconcave. Tarsi long. 1st seg- not wrinkled intervals and smaller eyes. ment strongly lengthened. 2nd segment wide - triangular. 3rd segment bilobed. Caudal segment Etymology. The name is formed from the location lengthened. Length of body: 2.5-2.6 mm. “Lushui” – “lushuensis”. Female: Pronotum 1.1 times wider than long. Elytra 1.2 times longer than wide. Protibiae weakly Genus Riedeliops Alonso-Zarazaga et Lyal, 2002 curved, biconcave from internal edge. Mesotibiae without appendages. 1-3rd ventrites with 2 lines Riedeliops asiaticus Legalov et Liu, sp.n. (Figs. of setae. 4th ventrite with 1 line of setae. Length 26-27, 70) of body: 2.8 mm.

Holotype. Male (IZAS), China, Tibet, Nielamu, Diagnosis. This new species is very close to 1800 m, 06.V.1966, Wang Shuyong. Riedeliops vietnamensis Legalov, 2003 but can be distinguished by the armament of the Paratypes. 1 male (SZMN), Hubei, Shennongjia, endophallus, punctation of pronotum, densely 900 m, 12.VI.1981, Han Yinheng; 1 female (IZAS), punctured abdomen and mesotibiae with short Yunnan, Lanping, 2300 m, 25.VIII.1984, Wang and wide appendages of males. Shuyong. Etymology. The name is formed from the word for Description. Body dark, pronotum with weak “Asian” – “asiaticus”. copper lustre. Elytra near suture greenish, and their sides with violet lustre. Riedeliops terminassianae Legalov et Liu, sp.n. (Figs. 28, 72) Male: Rostrum short, straight, widening toward apex, finely punctured, place of attachment of Holotype. Male (IZAS), China, Yunnan, Menghai, antennae located at basis. Eyes of average size, 17.IV.1982, Jiang Shengqiao. weakly convex. Forehead very narrow. Vertex convex, smooth. Antennae of average size. Description. Male: Body black, lacking lustre, Scapus and 1st segment widely oval. Funicle thin. naked. Rostrum short, straight, widening toward 2- and 3rd segments lengthened. 4-7th segments apex, finely punctured. Antennae located at ba- oval. Clava narrow, compact, strongly pointed. sis of rostrum. Eyes large, not protruding from

119 Legalov A.A., Liu N. contour of head. Forehead very narrow. Vertex Description. Body brown red. Pronotum and convex, finely punctured. Antennae of average scutellum dark brown. Head and pygidium black. size. Scapus and 1st segment widely oval. Funi- Abdomen with bluish lustre. Vertex with weak cle thin. 2- and 3rd segments lengthened. 4-7th violet lustre. Elytra copper red. Female. Rostrum segments oval. Clava narrow, compact, weakly short, widening toward apex, straight line, and pointed. Pronotum companiform, weakly trans- smooth, lacking lustre. Antennae located before versal, 1.1 times wider than long. Sides weakly basis of rostrum. Forehead very narrow. Eyes rounded. Pronotal groove not clear. Postnotal large, weakly convex. Temples weakly length- groove weak. Disk convex, sparsely punctured, ened. Vertex convex, gently rugose-punctate. with weak transversal striae before the middle. Antennae long, reaching as far as the middle of Scutellum almost rectangular, small. Elytra almost pronotum. Scapus and 1st segment oval. 2-4th rectangular, wide, almost equal length and width segments narrow - oval. 5-6th segments short - weakly narrowed toward apex, widest at humeri. oval. 7th segment rounded. Clava long, little Humeri weakly smoothed. Intervals narrow, longer than funicle, pointed. 1st segment longer strongly convex, smooth. Striae deep. Points in than 2 segment, shorter than 3 segment. Pronotum them large and rough. 9th stria merges with 10- trapezoid, 1.2 times wider than long. Pronotal before metacoxae. Metepisternum wide, sparsely groove clear. Basis of pronotum extended to punctured. Abdomen convex, sparsely punc- scutellum. Sides almost straight. Disk convex, tured, weakly impressed in the middle. 1-4th lacking lustre, sparsely and finely punctured. ventrites with lines of setae. 1st ventrite wide. 2- Scutellum back trapezoid, smooth. Elytra wide, 4th ventrites narrower. 5th ventrite very narrow. equal length and width, widest at humeri and in Pygidium convex, punctured. Legs long. Femora the middle. Humeri weakly smoothed. Intervals widened. Protibiae weakly curved, wide, with wide, convex, sparsely and very finely punctured. mucro. Mesotibiae biconcave, thick, from exter- Striae clear. Points in them large and dense. 9th nal edge in topmost third with appendages. Back stria merges with 10th stria before metacoxae. tibiae biconcave. Tarsi long. 1st segment strongly Mesothorax and mesepisternum matte. lengthened. 2nd segment triangular. 3rd segment Metathorax and metepisternum doubly punc- bilobed. Caudal segment lengthened. Length of tured: finely, and also strongly and densely punc- body: 2.8 mm. tured. Abdomen convex, gently rugose-punctate, flatted, but without lines of setae. 1-3rd ventrites Diagnosis. This new species is close to wide. 4th ventrite narrower. 5th ventrite very nar- Riedeliops rasuwanus Legalov, 2003 but can be row. 1-3rd ventrites with 2 lines of setae. 4th distinguished by the narrow, strongly convex ventrite with 1 line of setae. Pygidium convex, intervals of elytra, deep elytral striae, points in densely and strongly punctured. Legs long. them large and rough, more sparsely punctured Femora widened, sparsely and finely punctured. pronotum, weak depression in abdomen and ar- Protibiae weakly curved, from internal edge bi- mament of the endophallus. concave. Mesotibiae without appendages. Metatibiae biconcave. Tarsi long. 1st segment Etymology. This new species is named in honour strongly lengthened. 2nd segment wide - trian- of M.E. Ter-Minassian. gular. 3rd segment bilobed. Caudal segment lengthened. Length of body: 2.6 mm.

Genus Orienteuops Legalov, 2003 Diagnosis. This new species is close to Orienteuops tonkinensis (Voss, 1933) but can be Orienteuops mirabilis Legalov et Liu, sp.n. (Figs. distinguished by the fine size of body, less often 29-30) punctured pronotum, prothorax and scutellum without greenish lustre, thick punctured abdo- Holotype. Female (IZAS), China, Yunnan, men, and more sparsely and strongly punctured Xishuangbanna, Menghai, 1200-1600 m, pygidium. 26.VII.1958, Pu Fuji. 120 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China

Etymology. The name is formed from the word for “surprising” – “mirabilis”.

Genus Sawadaeuops Legalov, 2003

Key of species of subgenus Sawadaeuops s. str.

1. Antennae red - brown. Gansu £ £ £ £ £ £ £ £ £ £ £ £ £ £ £ £ £ £ £ ¤ S. potanini (Legalov)

-. Antennae darkly brown or black ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¢ ¢ ¢ ¢ ¢ ¢ ¢ 2

2. Forehead wider. Body smaller (2.4 mm). Hebei. ¥ ¥ ¥ ¥ ¥ ¥ ¥ ¥ ¥ ¥ ¦ S. beijingensis (Legalov)

-. Forehead narrower. Body larger (2.6-3.8 mm) ¡ ¡ ¡ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ 3

3. Mesotibiae without appendages of males. Russia Far East, Japan..S. punctatostriatus (Motschulsky)

-. Mesotibiae with appendages of males ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ 4

4. Sides of elytra more direct. Pronotum gently punctured, with deeper transversal striae. Sichuan,

Yunnan ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ S. subelongatus Legalov et Liu , sp.n.

-. Sides of elytra more arched. Pronotum to thick and more rough punctured, with weaker transversal

striae ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¡ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ 5

5. Beetles larger (2.9-3.5 mm). Pronotum less often punctured. Hubei, Shaanx ¥ ¥ ¥ ¥ ¥ ¦ ¦ ¦ ¦ ¦

£ £ £ £ £ £ £ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ ¤ S. centralchinensis Legalov et Liu, sp.n.

-. Beetles smaller (2.6-3.2 mm). Pronotum to thick punctured. Sichuan ¦ ¦ ¦ ¦ ¦ £ £ £ £ £ £ £ £ £

¡ ¡ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ ¢ S. sitchuanensis Legalov et Liu, sp.n.

Sawadaeuops (s. str.) sichuanensis Legalov et Sichuan, Wolong, 2100 m, 24.VII.1983, Wang Liu, sp.n. (Figs. 31-32, 65) Shuyong; 1 male (SZMN), 2 females (IZAS), 1 female (SZMN), Sichuan, Wolong, 1600 m, Holotype. Male (IZAS), China, Sichuan, Lixian, 26.VII.1983, Wang Shuyong; 1 female (SZMN), 2800 m, 14.VIII.1983, Wang Shuyong. Sichuan, Gonggashan, 2500 m, 08.V.1983, Chen Yuanqing; 1 female (SZMN), Sichuan, Paratypes. 1 female (IZAS), idem; 4 males (IZAS), Gonggashan, 2500m, 10.VI.1983, Chen Yuanqing; 1 male (SZMN), Sichuan, Wolong, 2600 m, 1 male (SZMN), Sichuan, Wenchuan, Wolong, 08.VIII.1983, Wang Shuyong; 1 male (IZAS), 1 1600 m, 26.VII.1983, Wang Shuyong; 4 females female (IZAS), Sichuan, Lixian, 2800 m, (IZAS), Sichuan, Wenchuan, Wolong, 1900 m, 12.VIII.1983, Wang Shuyong; 2 males (IZAS), 1 27.VII.1983, Wang Shuyong. male (SZMN), 3 females (IZAS), Sichuan, Wolong, 2500 m, 06.VIII.1983, Wang Shuyong; 1 male Description. Body black with bluish or greenish (IZAS), Sichuan, Wenchuan, Wolong, 1900 m, lustre. Mucro and claws brown. Antennae black 27.VII.1983, Wang Shuyong; 1 male (IZAS), - brown.

121 Legalov A.A., Liu N.

Male: Rostrum short, straight, strongly widen- and uncus. Protibiae shorter and wide. Length of ing toward apex, strongly and sparsely punc- body: 2.6-3.2 mm. tured. Antennae located at basis of rostrum. Fore- head very narrow. Eyes large, weakly convex. Diagnosis. This new species is very close to Vertex convex, densely and strongly punctured. Sawadaeuops (s. str.) centralchinensis Legalov Temples weakly lengthened, wrinkled. et Liu, sp.n. but can be distinguished by the Prementum with 2-3 teeth. Antennae long, reach- smaller size and thick punctured pronotum. From ing as far as the middle of pronotum. Scapus and Sawadaeuops (s. str.) punctatostriatus 1st segment wide - oval. 2-3rd segments long - (Motschulsky, 1860) it differs by the mesotibiae oval. 4-5th segments short - oval. 6th segment with appendages at apex of males. rounded. 7th segment transversal. Clava little shorter than funicle, pointed, thick. 1st segment Etymology. The name is formed from the name of as long as 2nd segment. 3rd segment longer than province Sichuan – “sichuanensis”. 2nd segment. Pronotum almost companiform, 1.0- 1.2 times wider than long. Pronotal groove weak. Sides very weakly rounded. Disk convex, densely Sawadaeuops (s. str.) centralchinensis Legalov and strongly punctured, with medial transversal et Liu, sp.n. (Figs. 33-34, 68) striae. Basis of pronotum extended. Scutellum almost rectangular, sparsely punctured. Elytra Holotype. Male (IZAS), China, Hubei, Shennong, almost rectangular, 1.1-1.2 times longer than wide. 1600 m, 13.VII.1980, Yu Peiyu. Humeri weakly smoothed. Greatest width in humeri and in the middle. Intervals wide, flat, Paratypes. 1 male (IZAS), Hubei, Shennongjia, smooth. Striae clear. Points in them dense and 1660 m, 13.VII.1981, Yu Peiyu; 1 male (SZMN), small. 9th stria merges with 10th stria before Hubei, Shennongjia, 1640 m, 12.VII.1981, Han metacoxae. Prothorax strongly punctured. Yinheng; 1 female (IZAS), 1 female (SZMN), Mesothorax and mesepisternum matte. Shaanxi, Niangshan, 2300 m, 06.VIII.1979, Han Metathorax strongly and sparsely punctured. Yinheng; 1 female (IZAS), Hubei, Shennongjia, Metepisternum matte, strongly and sparsely 1660 m, 24.VII.1981, Han Yinheng; 2 females punctured. Abdomen convex, densely rugose- (IZAS), Hubei, Shennongjia, 1600 m, 13.VII.1980, punctate, weakly flattened in the middle. 1-3rd Yu Peiyu; 1 female (SZMN), China, Shaanxi prov., ventrites wide. 4th ventrite narrower. 5th ventrite Qing Ling Shan mts., road Baoji - Taibai vill, pass narrow. Pygidium convex, densely and finely 40 km S Baoji, 21-23.VI.1998, Zd. Jundra; 1 female punctured. Legs long. Femora widened, sparsely (CKJU), China, Shaanxi prov., Qing Ling Shan punctured. Tibiae narrow, almost straight, densely mts., 30 km SE Taibai Shan mt., Hoi Zen Zi vill, and finely punctured, hardly shorter than 1500 m, 25.VI.1998, Zd. Jundra. profemora, widening toward apex, with long mucro at apex. Meso- and metatibiae biconcave, Diagnosis. This new species is very close to shorter and wide. Mesotibiae with small append- Sawadaeuops (s. str.) sichuanensis Legalov et ages. Tarsi lengthened. 1st segment long. 2nd Liu, sp.n. but can be distinguished by the larger segment long - triangular, flat. 3rd segment wide size and less often punctured pronotum. - bilobed. Caudal segment lengthened. Claws Pronotum 1.1 times wider than long. Elytra 1.2- long. Length of body: 2.6-3.0 mm 1.3 times (males) and 1.2-1.4 times (females) longer than wide. Length of body: 3.1-3.4 mm Female: Rostrum wider. Eyes hardly more small. (males), 2.9-3.5 mm (females). Clava of antennae shorter and thin. Pronotum 1.1-1.2 times wider than long. Elytra 1.2 times Etymology. The name is formed from the words longer than wide. Abdomen stronger convex. 1- “central” and “Chinese”. 3rd ventrites with 2 lines of erecting setae. 4th ventrite with 1 line of setae. Tibiae with mucro

122 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China

Sawadaeuops (s. str.) subelongatus Legalov et punctured. Vertex convex, punctured. Prementum Liu, sp.n. (Figs. 35-36) with 2 teeth. Antennae of average size, thin. Scapus and 1st segment oval. 2-4th segments Holotype. Female (IZAS), China, Yunnan, narrowly and long - oval. 5-7th segments shortly Qiujiaba, 09.IX.92. oval. Clava narrow, compact, weakly pointed. Pronotum weakly transversal, 1.1 times wider than Paratypes. 1 male (IZAS),1 female (IZAS), long. Pronotal groove not clear. Postnotal groove Sichuan, Gonggashan, 2500 m, 10.VI.83, Chen weak. Sides weakly rounded. Disk convex, Yuanqing; 1 female (IZAS), 1 female (SZMN), strongly and densely punctured, with weak me- Yunnan, Qiujiaba, 10.IX.92; 1 female (IZAS), dial transversal striae. Scutellum narrow, almost Sichuan, Gonggashan, 8500 m, 10.VI.83, Chen rectangular. Elytra wide, 1.1 times longer than Yuanqing; 1 female (SZMN), Sichuan, Wanxian, wide, almost rectangular, narrowed toward apex. 1200 m, 22.V.93, Wang Shuyong; 1 female (IZAS), Humeri well advanced. Greatest width in humeri. Sichuan, Nanping, Jiuzhaigou, 2300 m, 06.IX.83, Intervals wide, smooth, flat. Points in striae Wang Shuyong. dense. 9th stria merges with 10th stria before 1st ventrite. Metepisternum wide, sparsely punc- Diagnosis. This new species is very close to tured. Abdomen convex, sparsely and strongly Sawadaeuops (s. str.) sichuanensis Legalov et punctured, flattened in the middle. 1-2nd ventrites Liu, sp.n. and S. (s. str.) centralchinensis Legalov wide. 3-4th ventrites narrower. 5th ventrite very et Liu, sp.n. but can be distinguished by the more narrow. Pygidium convex, punctured. Legs long. direct sides of elytra, and also gently punctured Femora widened. Protibiae hardly shorter than pronotum with deeper transversal striae. femora, almost straight, widening toward apex, Pronotum 1.1 times (males), 1.1-1.2 times (females) with mucro. Meso- and metatibiae biconcave. wider than long. Elytra 1.2 times (males), 1.3-1.4 Tarsi long. 1st segment lengthened. 2nd segment times (females) longer than wide. Length of body: triangular. 3rd segment bilobed. Caudal segment 2.8 mm (males), 3.2-3.8 mm (females). lengthened. Length of body: 2.9 mm.

Etymology. The name is formed from the word for Female: Pronotum stronger rounded, length and “a little lengthened” – “subelongatus”. width almost equal. Elytra 1.1 times longer than wide. Protibiae weakly curved, from inside weakly biconcave. Abdomen stronger convex. 1-4th Sawadaeuops (Chinoeuops) hubeiensis Legalov ventrites with 2 lines of setae. 4th ventrite with 1 et Liu, sp.n. (Figs. 37-38) line of setae. Length of body: 2.8 mm.

Holotype. Male (IZAS), China, Hubei, Xingshan, Diagnosis. This new species is close to Longmenhe, 1400 m, 23.VI.1993, Li Wenzhu. Sawadaeuops (Chinoeuops) davidiani Legalov, 2003 but can be distinguished by the densely Paratype. 1 female (SZMN), Hubei, Shennongjia, punctured pronotum, rostrum without carina from 1660 m, 16.VII.1981, Han Yinheng. basis up to place of attachment of antennae, narrower forehead, strongly punctured rostrum, and Description. Body pitch-black, lacking lustre, pronotum with very weak transversal striae. naked. Etymology. The name is formed from the name of Male: Rostrum thick, short, widening toward province Hubei – “hubeiensis”. apex, strongly punctured, straight. Rostrum from basis up to place of attachment of antennae without carina. Eyes of average size, not protruding from contour of head. Forehead convex, narrow,

123 Legalov A.A., Liu N.

Sawadaeuops (Chinoeuops) australis Legalov 4th ventrites narrower. 5th ventrite very narrow. et Liu, sp.n. (Figs. 42-43, 73) Pygidium convex, strongly and densely punctured. Legs long. Femora widened, sparsely and Holotype. Male (IZAS), China, Guizhou, Leishan, finely punctured. Protibiae almost straight, little 1000-1890 m, 06.VII.1988, Wang Shuyong. shorter than femora, finely and densely punctured, widening toward apex, with mucro. Paratypes. 3 males (IZAS), 1 male (SZMN), 3 fe- Mesotibiae shorter and thick, with obtuse ap- males (IZAS), 1 female (SZMN), idem; 2 males pendages at external edge in topmost third. (IZAS), Yunnan, Lanping, 2400 m, 24.VIII.1984, Metatibiae biconcave, thick. Tarsi long, hardly Wang Shuyong; 1 male (IZAS),China, SE Yunnan, shorter than tibiae. 1st segment lengthened. 2nd Tzinpin, 1700 m, 25.V.1956, Panfilov; 1 female segment triangular. 3rd segment bilobed. Caudal (IZAS), Yunnan, Weixi, 2400 m, 25.VII.1981, Wang segment lengthened. Length of body: 2.3-2.6 mm. Shuyong; 1 female (IZAS), Yunnan, Yunlong, Female: Pronotum 1.1 times wider than long. Zhibenshan, 2300 m, 21.VI.1981, Liao Subai; 2 Elytra 1.2 times longer than wide. Abdomen females (IZAS), 1 female (SZMN), Yunnan, stronger convex. 1-3rd ventrites with 2 lines of Lanping, 2300 m, 4-21.VIII.1984, Wang Shuyong; setae. 4th ventrite with 1 line of setae. Protibiae 1 male (SZMN), Yunnan, 1956. shorter and thick, weakly curved, from inside weakly biconcave. Mesotibiae without append- Description. Body brown with bronze lustre. ages. Length of body: 2.6-3.0 mm.

Male: Rostrum short, wide, widening toward apex, Diagnosis. This new species is close to finely and densely punctured. Antennae located Sawadaeuops (Chinoeuops) davidiani Legalov, at basis of rostrum. Forehead very narrow. Eyes 2003 but can be distinguished by the brown body large, very weakly convex. Vertex convex, with bronze lustre, mesotibiae with appendages sparsely and finely punctured. Prementum with at external apex of males, thick punctured 2 teeth. Temples weakly lengthened, wrinkled. pronotum and very narrow forehead. Antennae long, reaching as far as the middle of pronotum. Scapus and 1st segment wide - oval. Etymology. The name is formed from the word for 2-3rd segments narrow - oval. 4-5th segments “southern” – “australis”. short - oval. 6th segment oval. 7th segment rounded. Clava long, little shorter than funicle. 1st segment as long as 3rd segment. 2nd seg- Sawadaeuops (Chinoeuops) nielamus Legalov ment shorter than 1st segment. Pronotum et Liu, sp.n. (Figs. 39-40) companiform, length and width equal. Sides very weakly rounded. Disk convex, sparsely and finely Holotype. Male (IZAS), China, Tibet, Nielamu, punctured, with very weak medial transversal 2600 m, 06.V.1966, Wang Shuyong. striae. Basis extended to scutellum. Back of scutellum trapezoid. Elytra 1.1-1.2 times longer Paratype. 1 female (SZMN), Tibet, Nielamu, 2600 than wide, almost rectangular. Humeri convex. m, 06.V.1966, Wang Shuyong. Greatest width in humeri and in the middle. Inter- vals wide, almost flat, smooth. Striae clear. Points Diagnosis. This new species is close to in them large, dense, decrease toward apex. 9th Sawadaeuops (Chinoeuops) australis Legalov stria merges with 10th stria before metacoxae. et Liu, sp.n. but can be distinguished by the more Prothorax sparsely and strongly punctured. direct sides of pronotum, thick punctured Mesothorax and mesepisternum matte. metepisternum, mesotibiae with very weak ap- Metathorax doubly punctured. Metepisternum pendages of males. Pronotum 1.1 times wider than sparsely and strongly punctured. Abdomen con- long. Elytra 1.3 times (male) and 1.2 times (fe- vex, densely and finely punctured, smooth and male) longer than wide. Length of body: 2.8 mm flattened in the middle. 1-2nd ventrites wide. 3- (male), 2.6 mm (female).

124 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China

Etymology. The name is formed from the location Sawadaeuops (Sawadaeuopsis) punctatus Nielamu - “nielamus“. Legalov et Liu, sp.n. (Fig. 41)

Subgenus Sawadaeuopsis Legalov, subgen.n. Holotype. Female (IZAS), China, Yunnan, (Fig. 41) Mengsong, 1600 m, 24.IV.1958, Pu Fuji.

Type species: Sawadaeuops punctatus Legalov Description. Female: Body brown. Head, rostrum, et Liu, sp.n. abdomen, and pygidium black - brown. Pronotum, scutellum dark brown. Rostrum short, Description. Body brown. Rostrum short, wide, wide, straight, widening toward apex, densely and straight, widening toward apex, densely and finely punctured. Antennae located at basis of finely punctured. Antennae located at basis of rostrum. Prementum with 2 teeth. Forehead very rostrum. Forehead very narrow. Eyes large, con- narrow. Eyes large, convex. Vertex convex, vex. Vertex convex, sparsely and strongly punc- sparsely and strongly punctured. Temples weakly tured. Temples weakly lengthened. Antennae lengthened. Antennae long, reaching as far as long, reaching as far as the middle of pronotum. the middle of pronotum. Scapus and 1st segment Pronotum companiform. Grooves weak. Sides al- wide - oval. 2-3rd segments long - oval. 4-6th most straight. Disk convex, densely and strongly segments short - oval. 7th segment rounded. punctured. Scutellum almost square, densely Clava long, little shorter than funicle, pointed. punctured. Elytra lengthened, almost rectangu- 1st segment as long as 3rd segment. 2nd seg- lar. Humeri slightly smoothed. Greatest width in ment shorter than 1st segment. Pronotum humeri and in the middle. Intervals wide, flat, with companiform, 1.1 times wider than long. Grooves 1 line of points. Striae clear. Points in them large, weak. Sides almost straight. Disk convex, densely decrease toward apex. Mesothorax and and strongly punctured. Basis of pronotum ex- mesepisternum matte. Metathorax densely and tended. Scutellum almost square, densely punc- strongly punctured. Metepisternum doubly tured. Elytra lengthened, 1.2 times longer than punctured. Abdomen convex, sparsely rugose- wide, almost rectangular. Humeri slightly punctate. 1st and 2nd ventrites wide. 3rd and 4th smoothed. Greatest width in humeri and in the ventrites narrower. 5th ventrite very narrow. middle. Intervals wide, flat, with 1 line of points. Pygidium convex, rough and strongly punctured. Striae clear. Points in them large, decrease to- Legs long. Femora widened, finely rugose-punc- ward apex. 9th stria merges with 10th stria before tate. Protibiae narrow, almost straight, at internal metacoxae. Mesothorax and mesepisternum edge weakly biconcave, with long mucro and matte. Metathorax densely and strongly punc- uncus at apex of female. Meso- and metatibiae tured. Metepisternum doubly punctated. Abdo- biconcave, shorter and thick. Tarsi lengthened, men convex, sparsely rugose-punctate. 1st and little shorter than tibiae. 1st segment lengthened. 2nd ventrites wide. 3rd and 4th ventrites nar- 2nd segment long - triangular, flat. 3rd segment rower. 5th ventrite very narrow. 1-3rd ventrites wide - bilobed. Caudal segment lengthened. Claws with 2 lines of setae. 4th ventrite with 1 line of long. Length of body: 3.2 mm. setae. Pygidium convex, rough and strongly punctured. Legs long. Femora widened, finely Diagnosis. This new subgenus differs from other rugose-punctate. Protibiae narrow, almost subgenera of this genus by the strongly and straight, at internal edge weakly biconcave, with roughly punctured pronotum. long mucro and uncus at apex. Meso- and metatibiae biconcave, shorter and thick. Tarsi Etymology. The name is formed by addition of lengthened, little shorter than tibiae. 1st segment the ending “-is” to “sawadaeuops”. lengthened. 2nd segment long - triangular, flat. 3rd segment wide - bilobed. Caudal segment lengthened. Claws long. Length of body: 3.2 mm.

125 Legalov A.A., Liu N. Diagnosis. This new species is similar to tured. Pygidium convex, finely and sparsely punc- Sawadaeuops (s. str.) sichuanensis Legalov et tured. Legs long. Femora thick, weakly squeezed Liu, sp.n. but can be distinguished by the strongly from sides, finely and sparsely punctured. Tibiae and roughly punctured pronotum, head strongly short, weakly curved, densely wrinkled, widen- and densely punctured, narrower intervals of ing toward apex. Tarsi long. 1st segment length- elytra with 1 line of large points and brown body. ened. 2nd segment triangular. 3rd segment wide - Etymology. The name is formed from the word bilobed. Caudal segment lengthened, narrow. for “punctured” – “punctatus”. Claws long. Length of body: 5.1 mm.

Diagnosis. This new species is similar to Tribe Attelabini Billberg, 1820 Attelabus (s. str.) cyanellus Voss, 1925 but can Genus Attelabus Linnaeus, 1758 be distinguished by the wider body, more strongly narrowed pronotum, and more strongly Attelabus (s. str.) sichuanensis Legalov et Liu, punctured metepisternum, more finely and less sp.n. (Figs. 44-45) often punctured intervals of elytra and strongly flattened femora. Holotype. Female (IZAS), China, Sichuan, Xiangcheng, 3500-3800 m, 05.VII.82, Wang Etymology. The name is formed from the name of Shuyong. province Sichuan – “sichuanensis”.

Description. Female: Body dark blue. Antennae black-brown. Mucro and uncus brown. Head Genus Lamprolabus Jekel, 1860 lengthened. Rostrum short and wide, widening toward apex, finely punctured. Antennae located Lamprolabus pseudobispinosus Legalov et Liu, at basis of rostrum. Forehead wide, flat, punc- sp.n. (Fig. 51) tured, with medial striae. Eyes small, weakly convex. Temples lengthened, finely rugose-punctate. Holotype. Female (IZAS), China, Yunnan, Vertex convex, finely and sparsely punctured. Xishuangbanna, Mengzhe, 1750 m, 24.VI.58, Pu Antennae short, reaching as far as the middle of Fuji. pronotum. Scapus and 1st segment oval. 2-5th segments lengthened - oval. 6th and 7th seg- Paratype. 1 female (SZMN), idem. ments short - oval. Clava much shorter than funicle. 1st segment longer than 2nd segment. 3rd Description. Female: Body red-brown. Bottom segment longer than 1st segment, weakly and antennae yellow. Teeth on elytra black. Head pointed. Pronotum trapezoid, 1.3 times wider than conical. Rostrum short, widening toward apex, long, strongly narrowed toward apex. Sides sparsely punctured. Antennae located near ba- weakly rounded. Disk convex, finely and sparsely sis of rostrum. Forehead wide, convex, smooth, punctured, with weak medial line, transversal with 2 striae on each side, forming sharp angle. wrinkled at basis. Scutellum lengthened, trap- Eyes not big and strongly convex. Temples ezoid. Elytra wide, 1.1 times longer than wide, lengthened. Vertex convex, smooth, with striae. weakly pressed for scutellum. Greatest width Antennae short, reaching as far as the middle of behind the middle. Intervals wide, flat, finely punc- pronotum. Scapus and 1st segment oval. 2-5th tured. Penultimate interval convex. Striae shal- segments lengthened. 6th and 7th segments short low. Points in them dense. Mesothorax matte. - oval. Clava lengthened, little shorter than funi- Metathorax punctured. Mesepisternum densely cle. 1st and 2nd segments of equal length. 3rd punctured. Metepisternum sparsely and strongly segment shorter than 2nd segment, weakly punctured. Abdomen convex. 1-3rd ventrites with pointed. Pronotum trapezoid, 1.3 times wider than lateral and medial edges. 1-3rd ventrites wide. long. Sides almost straight. Pronotal groove 4th ventrite narrower. 5th ventrite narrow, punc- sharp. Disk convex, smooth. Basis convex to

126 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China scutellum. Scutellum almost triangular. Elytra oval. Clava lengthened, little shorter than funi- lengthened, 1.3 times longer than wide, widest at cle, pointed. 1st segment long, as long as 3rd humeri. Long teeth in first third of elytra. Inter- segment, longer than 2nd segment. Pronotum vals wide, convex, smooth. Striae weak. Points in trapezoid, 1.3-1.4 times wider than long. Sides them dense, especial in topmost half of elytra. weakly rounded. Pronotal groove sharp. Disk Metepisternum narrow, sparsely punctured. Ab- convex, smooth. Basis convex to scutellum. domen convex, smooth. 1-3rd ventrites wide. 4th Scutellum wide, almost triangular. Elytra length- ventrite narrower. 5th ventrite narrow, densely ened, 1.2 times longer than wide, widest at hu- punctured. Pygidium convex, sparsely punctured. meri. Long wide teeth in first third of elytra. Inter- Legs long. Femora widened, without teeth. Tibiae vals narrow, convex, smooth. Striae weak, wide. biconcave, narrow, their internal edge denticu- Points in them dense, especially in topmost half late with erect setae. Protibiae longer than aver- of elytra. Thorax strongly punctured. age and back. Apex of tibiae with mucro and un- Metepisternum narrow, sparsely and strongly cus. Tarsi long. 1st segment lengthened. 2nd seg- punctured. Abdomen convex, sparsely rugose- ment triangular. 3rd segment bilobed. Caudal seg- punctate. Sutures between ventrites deep. 1-3 ment lengthened. Claws long. Length of body: ventrites wide. 4th ventrite narrower. 5th ventrite 5.4-5.7 mm. narrow, densely punctured. Pygidium convex, densely punctured. Legs long. Coxae densely Diagnosis. This new species is close to punctured. Femora widened, without teeth. Tibiae Lamprolabus bispinosus (Gyllenhal, 1833) but biconcave, wide, their internal edge finely den- can be distinguished by the longer and sharp ticulate. Protibiae longer than meso- and teeth on elytra, yellow tibiae, and elytra without metatibiae. Apex of tibiae with mucro and uncus. dark spots, more strongly widening and more Tarsi long. 1st segment lengthened. 2nd segment finely punctured rostrum. wide - triangular. 3rd segment bilobed. Caudal segment lengthened. Claws long. Length of body: Etymology. The name is formed by addition of 7.5-8.1 mm. the prefix “pseudo-“ to “bispinosus”. Diagnosis. This new species is close to Lamprolabus spiculatus (Boheman, 1845) but can Lamprolabus tibetanus Legalov et Liu, sp.n. be distinguished by the more light body, in part (Fig. 47) black, sharper teeth on elytra, and weaker convex intervals in first third of elytra. Holotype. Female (IZAS), China, Tibet, Motuo, 800-1000 m, 15.V.83, Han Yinheng. Etymology. The name is formed from the location “Tibet” – “tibetanus”. Paratype. 1 female (SZMN), idem, 850m, 04.VII.83, Han Yinheng; Genus Henicolabus Voss, 1925 Description. Female: Body red-brown. Bottom half of teeth on elytra black. Head conical. Ros- Henicolabus gigantinus Legalov et Liu, sp.n. trum short, widening toward apex, sparsely punc- (Figs. 52-53, 57) tured. Antennae located near basis of rostrum. Forehead wide, convex, smooth, with 2 striae on Holotype. Male (IZAS), China, Yunnan, each side, forming sharp angle and medial striae. Xishuangbanna, Mengzhe, 1750 m, 25.VI.1958, Eyes not big and strongly convex. Temples Pu Fuji. lengthened. Vertex convex, smooth, with striae. Antennae short, reaching as far as the middle of Paratypes. 1 male (SZMN), Yunnan, pronotum. Scapus and 1st segment oval. 2-4th Xishuangbanna, Xiaomengyang, 850 m, segments lengthened. 5-7th segments short - 25.X.1957, Wang Shuyong; 1 female (IZAS), idem,

127 Legalov A.A., Liu N.

18.VI.1957, Wang Shuyong; 1 female (IZAS), 1 Diagnosis. This new species is close to female (SZMN), idem, 25.X.1957, Wang Shuyong. Henicolabus giganteus (Faust, 1882) but can be distinguished by the shorter weakly curved for- Description. Body yellowish-brown. Apex of ros- ward tibiae, wider head, narrower pronotum, more trum, femora, tibiae, tarsi and antennae black. strongly elytra widening toward apex, and form of aedeagus. Male: Head almost cylindrical. Rostrum very short and wide, densely and finely punctured, with Etymology. The name is formed from the word for striae in the middle. Antennae located at basis of “giant” – “gigas”. rostrum. Forehead wide, convex, smooth, with medial striae in first half and 2 deep striae on each side. Eyes large, convex. Temples length- Genus Humerilabus Legalov, 2003 ened. Vertex convex, punctured, with medial line. Antennae short, reaching as far as first line of Humerilabus longulus Legalov et Liu, sp.n. (Fig. pronotum. Funicle thick. Scapus and 1-4th seg- 46) ments wide - oval. 5-7th segments rounded. Clava long, little shorter than funicle. 1st and 2nd seg- Holotype. Female (IZAS), China, Yunnan, ments of equal length. 3rd segment longer, little Xishuangbanna, Xiaomengyang, 850 m, 25.X.57, shorter than previous segments. Pronotum trans- Wang Shuyong. versal, 1.5 times wider than long, with marked grooves. Sides very weakly rounded. Disk con- Paratype. 1 female (SZMN), Yunnan, Binbjan, vex, very finely and sparsely punctured. Deveishan, 1500 m, 22.VI.1956, Xuan Ke-zhen. Scutellum almost triangular. Elytra wide, 1.0-1.1 times longer than wide. Humeri convex. Greatest Description. Female: Body red-brown. Elytra and width behind the middle. Intervals wide, sparsely sometimes pronotum yellowish-brown. Antennae punctured. Striae weak. Points in them fine. Tho- yellow. Head lengthened. Rostrum straight line rax and metepisternum roughly and strongly widening toward apex, sparsely punctured. An- punctured. Abdomen convex, rugose-punctate. tennae located before basis of rostrum. Forehead 1-3rd ventrites wide. 4th ventrite narrower. 5th wide, convex, smooth, with 2 weak striae on each ventrite narrow. Pygidium convex, finely and side. Eyes large, strongly convex. Temples sparsely punctured. Legs long. Forward legs strongly lengthened. Vertex convex, finely ru- strongly lengthened. Profemora strongly wid- gose-punctate. Antennae short, reaching as far ened, with teeth. Meso- and metafemora widened, as first line of pronotum. Scapus and 1st seg- with tooth at basis. Protibiae long, weakly curved, ment oval. 2-5th segments lengthened. 6th seg- with long mucro at apex and with long teeth on ment short - oval. 7th segment rounded. Clava internal edge. Meso- and metatibiae short, almost lengthened, little shorter than funicle. 1st and straight, with weak teeth on internal edge. Tarsi 2nd segments of equal length. 3rd segment long. 1st segment lengthened. 2nd segment long lengthened, weakly pointed, little shorter than - triangular. 3rd segment bilobed. Caudal segment 1st and 2nd segments taken together. Pronotum lengthened. Length of body: 8.2-8.3 mm. transversal, 1.2-1.3 times wider than long, with well advanced grooves. Sides weakly rounded. Female: 3rd segment of clava little longer than Disk convex, sparsely punctured, with weak me- 2nd segment. Pronotum 1.5-1.7 times wider than dial line. Scutellum lengthened, wrinkled, almost long. Protibiae shorter and wide. Apex of tibiae triangular. Elytra lengthened, 1.4 times longer than with mucro and uncus. Length of body: 7.6-8.2 wide, widest in the middle. Humeri strongly con- mm. vex. Intervals weakly convex, smooth. Striae weak. Points in them fine, especial in topmost half of elytra. Mesothorax wrinkled. Metathorax sparsely punctured. Metepisternum roughly and densely

128 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China punctured. Abdomen convex, finely rugose-punc- sharp. Striae clear. Points in them shallow and tate. 1-3rd ventrites wide. 4th ventrite narrower. dense. 9th stria merges with 10th stria before 5th ventrite narrow, densely punctured. Pygidium metacoxae. Prothorax wrinkled, with blunt teeth convex, finely and sparsely punctured. Legs on first line. Precoxal part short. Meso- and long. Forward legs strongly lengthened. metathorax gently wrinkled. Metepisternum Profemora large, strongly widened, with weak smooth. Abdomen convex, wrinkled. Pygidium tooth at apex, sparsely both finely wrinkled. convex, sparsely punctured. Legs long. Forward Meso- and metafemora widened, without teeth. legs strongly lengthened. Profemora strongly Protibiae long, weakly curved, densely punc- widened, without teeth. Meso- and metafemora tured, with mucro and uncus. Meso- and widened. Protibiae long, weakly curved, wide, metatibiae short, flattened, almost straight. Tarsi with long mucro at apex and with 7 teeth on inter- long. 1st segment lengthened. 2nd segment tri- nal edge. Meso- and metatibiae short, biconcave. angular. 3rd segment bilobed. Caudal segment Tarsi long, little shorter than tibiae. 1st segment lengthened, longer than previous segments. lengthened - triangular. 2nd segment triangular. Claws long. Length of body: 7.9-8.5 mm. 3rd segment bilobed. Caudal segment lengthened. Length of body: 4.7-5.6 mm. Diagnosis. This new species is close to Humerilabus fausti (Voss, 1925) but can be dis- Diagnosis. This new genus is close to genus tinguished by the more poorly convex Humeri, Henicolabus Voss, 1925 but can be distinguished more strongly lengthened elytra, more strongly by the sharp pronotal groove, femora without lengthened 3rd segment of clava, weak tooth on teeth, shorter protibiae and armament of the profemora and narrower protibiae. endophallus.

Etymology. The name is formed from the word for Etymology. The name is formed from the words “lengthened” – “longulus”. “Chinese” and “labus”.

Genus Chinolabus Legalov, gen.n.(Figs. 48-49, Chinolabus bicolor Legalov et Liu, sp.n. (Figs. 54-56) 48-49, 55)

Type species: Chinolabus ningxianus Legalov Holotype. Male (IZAS), China, Qinghai, Maixiu, et Liu, sp.n. 14.V.1984.

Description. Body black. Elytra red or yellowy- Paratype. 1 female (SZMN), idem, 08.VII.1982. brown. Head lengthened. Rostrum short, Description. Body black. Elytra red. Claws, mucro strongly widening, sparsely punctured. Anten- and uncus brown. nae located before basis of rostrum. Forehead wide, pressed, finely punctured, with 2 deep striae Male: Head lengthened. Rostrum short, strongly on each side, merge in sharp angle. Eyes large, widening, sparsely punctured. Antennae located convex. Temples lengthened. Vertex convex, before basis of rostrum. Forehead wide, pressed, smooth, with medial striae. Antennae short, finely punctured, with 2 deep striae on each side, reaching as far as first line of pronotum. Pronotum merge in sharp angle. Eyes large, convex. Tem- trapezoid, narrowed toward apex. Disk convex, ples lengthened. Vertex convex, smooth, with smooth, with 2 deep impression in the middle, medial striae. Antennae short, reaching as far as without medial striae. Pronotal groove clear. first line of pronotum. Scapus and 1st segment Postnotal groove weak. Scutellum convex, almost oval. 2nd segment short - oval, much shorter than rectangular. Elytra wide. Humeri convex. Great- 1st segment. 3rd and 4th segments narrow - oval. est width behind the middle. Intervals wide, flat, 5-6th segments tear-shaped. 7th segment trap- weakly rugose-punctate. Edge of elytra carinate, ezoid, wide, similar to segments of clava. Clava

129 Legalov A.A., Liu N. long, shorter than funicle. 1st segment longer Holotype. Male (IZAS), China, Ningxia, Longtan, than 2nd segment. 3rd segment narrower, long, 14.VII.1986. little shorter previous, pointed. Pronotum trapezoid, 1.4 times wider than long, narrowed to- Paratypes. 2 males (IZAS), 1 male (SZMN), 1 fe- ward apex. Sides almost straight. Disk convex, male (SZMN), idem; 1 female (IZAS), Ningxia, smooth, with 2 deep depressions in the middle, Liupanshan, VI.1964. without medial striae. Pronotal groove clear. Postnotal groove weak. Scutellum convex, almost Diagnosis. This new species is very close to rectangular. Elytra wide, length and width equal. Chinolabus bicolor Legalov et Liu, sp.n. but can Humeri convex. Greatest width behind the mid- be distinguished by the weak transversal striae dle. Intervals wide, flat, weakly rugose-punctate. on vertex and armament of the endophallus. Edge of elytra carinate, sharp. Striae clear. Points Pronotum 1.3-1.4 times (males) and 1.3-1.5 times in them shallow and dense. 9th stria merges with (females) wider than long. Length and width of 10th stria before metacoxae. Prothorax wrinkled, elytra approximately equal. Length of body: 5.1- with blunt teeth on first line. Precoxal part short. 5.6 mm (males), 4.7-4.8 mm (females). Meso- and metathorax gently wrinkled. Metepisternum smooth. Abdomen convex, wrin- Etymology. The name is formed from the name of kled. 1-3rd ventrites wide. 4th ventrite narrower. province Ningxia – “ningxianus“. 5th ventrite narrow. Pygidium convex, sparsely punctured. Legs long. Forward legs strongly lengthened. Profemora strongly widened, with- Tribe Euscelophilini Voss, 1925 out teeth. Meso- and metafemora widened. Protibiae long, weakly curved, wide, with long Genus Cupreuscelophilus Legalov, gen.n. (Fig. mucro at apex and with 7 teeth on internal edge. 50) Meso- and metatibiae short, biconcave. Tarsi long, little shorter than tibiae. 1st segment length- Type species: Cupreuscelophilus mayongi ened - triangular. 2nd segment triangular. 3rd seg- Legalov et Liu, sp.n. ment bilobed. Caudal segment lengthened. Length of body: 5.2 mm. Description. Most parts of body with copper lustre. Body with light, dense appressed setae. Ros- Female: Antennae shorter. Clava shorter. trum short, wide, densely punctured. Antennae Pronotum 1.3 times wider than long. Elytra located near basis of rostrum. Eyes not big, stronger widening toward apex. Protibiae shorter strongly convex. Forehead wide. Temples length- and wide. Apex of tibiae with mucro and uncus. ened, weakly narrowed to pronotum. Antennae Length of body: 5.1 mm. short, first line of pronotum reach. Pronotum companiform. Postnotal groove clear. Pronotal Diagnosis. This new species is close to groove very weak. Sides weakly rounded. Disk Chinolabus ningxianus Legalov et Liu, sp.n. but convex, densely rugose-punctate, sometimes can be distinguished by the deep transversal with protuberance. Scutellum wide, back trap- striae on vertex and form of sclerites an ezoid. Elytra lengthened, sometimes with protu- endophallus. berance, widest in the middle. Intervals weakly convex, wide. Striae clear. Points in them large Etymology. The name is formed from the word for and dense. 9th stria merges with 10th stria near “bicoloured” – “bicolor”. apex of elytra. Thorax densely rugose-punctate. Metepisternum wide. Abdomen convex, densely rugose-punctate. 1-3rd ventrites wide. 4th Chinolabus ningxianus Legalov et Liu, sp.n. ventrite narrow. 5th ventrite very narrow. (Figs. 54, 56) Pygidium weakly convex, densely punctured. Legs long. Femora widened, with 1 tooth. Tibiae

130 New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China wide, almost straight, densely punctured, from carinate in topmost half of elytra. Striae clear. internal edge weakly biconcave. Tarsi long, little Points in them large and dense. 9th stria merges shorter than tibiae. 1st segment lengthened. 2nd with 10th stria near apex of elytra. Thorax densely segment long - triangular. 3rd segment bilobed. rugose-punctate. Metepisternum wide. Abdomen Caudal segment long. Claws long. Length of convex, densely rugose-punctate. 1-3rd ventrites body: 3.5-4.6 mm. wide. 4th ventrite narrow. 5th ventrite very narrow. Pygidium weakly convex, densely punctured. Diagnosis. This new genus is close to genus Legs long. Femora widened, with 1 tooth. Tibiae Euscelophilus Voss, 1925 but can be distin- wide, almost straight, densely punctured, from guished by the copper lustre of body, short tem- internal edge weakly biconcave. Tarsi long, little ples and dense setae on the body. shorter than tibiae. 1st segment lengthened. 2nd segment long - triangular. 3rd segment bilobed. Etymology. The name is formed from the word for Caudal segment long. Claws long. Length of “copper” – “cupreus”, and “euscelophilus”. body: 4.6 mm.

Diagnosis. This new species is close to Cupreuscelophilus mayongi Legalov et Liu, sp.n. Cupreuscelophilus kunmingensis (Liang, 1994) (Fig. 50) but can be distinguished by the absence of a protuberance on the pronotum and elytra, and Holotype. Female (IZAS), China, Zhejiang, Anji, also a smaller size of the body. Longwangshan, 13.V.1996, Wu Hong. Etymology. This new species is named in honour Description. Female: Body black. Head, rostrum, of Ma Yong. pronotum, elytra, scutellum, abdomen, thorax copper. Body with light, dense, appressed setae. Rostrum short, wide, densely punctured. Anten- Cupreuscelophilus kunmingensis (Liang, 1994), nae located near basis of rostrum. Eyes not big, comb.n. strongly convex. Forehead wide, strongly pressed in the middle, densely rugose-punctate, Euscelophilus kunmingensis Liang, 1994: 490 with medial carina. Temples lengthened, weakly narrowed toward pronotum, rugose-punctate. Distribution. China (YUN). Antennae short, reaching first line of pronotum. Scapus and 1st segment wide - oval. 2nd seg- Remarks. On the basis of the description, Fig- ment narrow - oval, as long as 1st segment. 3rd ures and photo in Liang (1994) these species rep- segment short - oval. 4th segment rounded. 5th resent a new genus. and 6th segments transversal. 7th segment sharply transversal, similar to segments of clava. Clava short, much shorter than funicle, compact. ACNOWLEDGEMENT 1st segment as long as 3rd segment. 2nd segment hardly shorter than 1st segment, pointed. This work was supported by a grant of the Rus- Pronotum companiform, 1.4 times wider than long. sian Science Support Foundation and a grant of Postnotal groove clear. Pronotal groove very the Russian Federal Property Fund No 04-04- weak. Sides weakly rounded. Disk convex, 48727-a. densely rugose-punctate, with medial striae and 2 depressions on each side. Scutellum wide, back I wish to thank A.-L.-L. Friedman (Tel Aviv), J. trapezoid, densely punctured. Elytra lengthened, Frisch (Berlin), O. Jaeger (Dresden), O.N. 1.8 times longer than wide. Humeri weakly Kabakov (Saint-Petersburg), B.A. Korotyaev smoothed, widest in the middle. Intervals weakly (Saint-Petersburg), P. Kresl (Janovice nad convex, wide, densely punctured. 9th interval Uhlavou), O. Merkl (Budapest), Dr. Y. Sawada

131 Legalov A.A., Liu N.

(Sanda), M. Schmitt (Bonn), R. Zhang (Beijing), (Coleoptera: Rhynchitidae, Attelabidae) Ma Yong (Beijing), Lu Xiu Xia (Beijing), for help using the Synap method. Report 1 // in the work. Zoologicheskii zhurnal 83 (12): 1427-1432. (in Russian)

REFERENCES Legalov A.A. 2004c. Experience of construction of phylogenetic systems of the beetles Legalov A.A. 2003a. Taxonomy, classification and (Coleoptera) by the example of the leaf- phylogeny of the leaf-rolling weevils rolling weevils (families Rhynchitidae and (Coleoptera: Rhynchitidae, Attelabidae) of Attelabidae) // Herald of Tomsk State the world fauna. Novosibirsk. CD-R. ¹ University. Series “Natural sciences”. 0320301200. 733+350 p. (641 Mb.) (In Appendix 11: 46-51. (in Russian) Russian with English diagnosis) Legalov A.A. 2005. Reconstruction of phylogeny Legalov A.A. 2003b. Checklist of the leaf-rolling in leaf-rolling weevils (Coleoptera: weevils (Coleoptera: Rhynchitidae, Rhynchitidae, Attelabidae) using the Attelabidae) of China (24 pp.) // Taxonomy, Synap method. Report 2 // Zoologicheskii classification and phylogeny of the leaf- zhurnal 84 (2): 190-194. (in Russian) rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) of the world fauna. Liang X. 1994. Seven new species of the Novosibirsk. CD-R. (641 Mb.) Euscelophilus (Coleoptera, Attelabidae) from Yunnan, South-west China // Japa- Legalov A.A. 2004a. New data of the leaf-rolling nese Journal of Entomology 62 (3): 483- weevils (Coleoptera: Rhynchitidae, 496. Attelabidae) of the world fauna with description of 35 new taxons // Baltic Journal of Coleopterology 4 (1): 63-88.

Legalov A.A. 2004b. Reconstruction of Received: 23.03.2005. phylogeny in leaf-rolling weevils Accepted: 23.09.2005.

132